treatments-xml/data/50/3B/8E/503B8E366769B14AFE65A01AECAF5D28.xml

<document id="AF8E14A4341CAD80F3A1F1BFACBE06BB" ID-CLB-Dataset="254448" ID-DOI="10.1080/00222933.2023.2192431" ID-GBIF-Dataset="64f4ca9e-4556-45ac-8a57-00fc3a16e837" ID-ISSN="1464-5262" ID-Zenodo-Dep="7859817" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1682060567496" checkinUser="plazi" docAuthor="Cumberlidge, Neil, Daniels, Savel R., Soma, Julia B. &amp; Leever, Ellen M." docDate="2023" docId="503B8E366769B14AFE65A01AECAF5D28" docLanguage="en" docName="JNATHIST.57.5-8.463-474.pdf" docOrigin="Journal of Natural History 57 (5 - 8)" docSource="http://dx.doi.org/10.1080/00222933.2023.2192431" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Vichai cyanalepou Cumberlidge &amp; Daniels &amp; Soma &amp; Leever 2023, sp. nov." docType="treatment" docVersion="7" lastPageNumber="473" masterDocId="AC02F64E676EB140FF8DA501ED46586F" masterDocTitle="Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar" masterLastPageNumber="474" masterPageNumber="463" pageNumber="470" updateTime="1702003164460" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="49C9A5A862FA9D100147B9EC48535D74">Vichai cyanapelou gen. et sp. nov. (Crustacea: Deckeniidae: Hydrothelphusinae), a new genus and new species of freshwater crab from northern Madagascar</mods:title>
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<mods:namePart id="1582A9C534E29075008E92E82E95B583">Cumberlidge, Neil</mods:namePart>
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<mods:namePart id="DAA2099F2E973CF009EDE9F40439FF21">Daniels, Savel R.</mods:namePart>
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<mods:namePart id="E0A3AFE1C5726F5BF34113B2E191C30C">Soma, Julia B.</mods:namePart>
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<mods:namePart id="561891AFBAAA6822F0C508E8F45DE962">Leever, Ellen M.</mods:namePart>
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<mods:title id="26292F936E6332860836C499B97002FE">Journal of Natural History</mods:title>
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<mods:date id="C38248A5DB2DDB0F8E1AA400B9616083">2023</mods:date>
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<mods:number id="130A77757FB3C4DF18E51313535BEBDE">2023-04-20</mods:number>
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<mods:number id="F66CF371BFB3A86372684E3045B0EA2C">57</mods:number>
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<mods:number id="9578878A39AE69DEF8D551FF7E1A4682">5 - 8</mods:number>
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<treatment id="503B8E366769B14AFE65A01AECAF5D28" ID-DOI="http://doi.org/10.5281/zenodo.7859823" ID-GBIF-Taxon="209569086" ID-Zenodo-Dep="7859823" LSID="urn:lsid:plazi:treatment:503B8E366769B14AFE65A01AECAF5D28" httpUri="http://treatment.plazi.org/id/503B8E366769B14AFE65A01AECAF5D28" lastPageId="10" lastPageNumber="473" pageId="7" pageNumber="470">
<subSubSection id="90886CAB6769B147FE65A01AEE7D5D59" box="[488,827,1307,1334]" pageId="7" pageNumber="470" type="nomenclature">
<paragraph id="D82D3F206769B147FE65A01AEE7D5D59" blockId="7.[488,827,1307,1334]" box="[488,827,1307,1334]" pageId="7" pageNumber="470">
<heading id="8365884C6769B147FE65A01AEE7D5D59" box="[488,827,1307,1334]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="470" reason="5">
<taxonomicName id="1F9244A36769B147FE65A01AEF955D59" authority="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever, 2023" authorityName="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever" authorityYear="2023" box="[488,723,1307,1334]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
<emphasis id="EAE6E3326769B147FE65A01AEF955D59" bold="true" box="[488,723,1307,1334]" italics="true" pageId="7" pageNumber="470">Vichai cyanalepou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1D55E496769B147FD51A01AEE7D5D59" box="[732,827,1307,1334]" pageId="7" pageNumber="470" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="90886CAB6769B147FE26A057EF895E7F" pageId="7" pageNumber="470" type="description">
<paragraph id="D82D3F206769B147FE26A057EE3E5D1E" blockId="7.[427,888,1366,1393]" box="[427,888,1366,1393]" pageId="7" pageNumber="470">
(
<figureCitation id="40A923A56769B147FE39A057EF595D1E" box="[436,543,1366,1393]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="7" pageNumber="470">Figures 1</figureCitation>
(a,b), 2(a–i), 3(a–d); 
<tableCitation id="95100A9B6769B147FC98A057EE365D1E" box="[789,880,1366,1393]" captionStart="Table 1" captionStartId="1.[160,216,1316,1339]" captionTargetPageId="1" captionText="Table 1. Vichai cyanalepou sp. nov. male holotype (FMNH 7589). Length in mm of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5), and the leg length in relation to the carapace width (CW). The sum of the length of each article of the four ambulatory legs divided by the carapace width is also shown (Total of 5 articles/CW), as is the sum of the relative length of the four ambulatory legs (ΣP2–5/CW = 5.35), placing this species in the short ambulatory leg size class (short: 5.0–5.9; medium: 6.0–6.9; long:&gt;7.0)." httpUri="http://table.plazi.org/id/8CED6FA8676FB141FF2DA025EE1C5DA2" pageId="7" pageNumber="470" tableUuid="8CED6FA8676FB141FF2DA025EE1C5DA2">Table 1</tableCitation>
)
</paragraph>
<paragraph id="D82D3F206769B147FF2DA0B0EC895DA4" blockId="7.[160,1156,1457,1552]" box="[160,463,1457,1483]" pageId="7" pageNumber="470">
<emphasis id="EAE6E3326769B147FF2DA0B0EC895DA4" bold="true" box="[160,463,1457,1483]" italics="true" pageId="7" pageNumber="470">Nomenclatural statement</emphasis>
</paragraph>
<paragraph id="D82D3F206769B147FF2DA0D5EF895E7F" blockId="7.[160,1156,1457,1552]" pageId="7" pageNumber="470">
A life science identifier (LSID) number was obtained for the new species: 
<uri id="AC0333226769B147FC4BA0D5EF895E7F" pageId="7" pageNumber="470">
urn:lsid:zoobank. org:act: 
<uuid id="AC3405F56769B147FF66A0F6EF895E7F" box="[235,719,1527,1552]" pageId="7" pageNumber="470">B5985969-B677-47E4-BE59-37C1308B7A66</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="90886CAB6769B148FF2DA33FE9C25A6D" lastPageId="8" lastPageNumber="471" pageId="7" pageNumber="470" type="materials_examined">
<paragraph id="D82D3F206769B147FF2DA33FEC395E37" blockId="7.[160,1156,1598,1728]" box="[160,383,1598,1624]" pageId="7" pageNumber="470">
<emphasis id="EAE6E3326769B147FF2DA33FEC395E37" bold="true" box="[160,383,1598,1624]" italics="true" pageId="7" pageNumber="470">Material examined</emphasis>
</paragraph>
<paragraph id="D82D3F206769B148FF2DA363E90858A5" blockId="7.[160,1156,1598,1728]" lastBlockId="8.[160,1156,142,514]" lastPageId="8" lastPageNumber="471" pageId="7" pageNumber="470">
<emphasis id="EAE6E3326769B147FF2DA363EC705E14" box="[160,310,1634,1659]" italics="true" pageId="7" pageNumber="470">Type material</emphasis>
. 
<materialsCitation id="68FA357D6769B148FEC9A363E90858A5" ID-GBIF-Occurrence="4089470301" collectingDate="2001-10-16" collectionCode="FW, ASL" collectorName="Bemanivy River. &amp; S. M. Goodman" country="Madagascar" county="Bemanivy River" elevation="810" lastPageId="8" lastPageNumber="471" latitude="-14.4267" location="Sava Region" longLatPrecision="5" longitude="49.60833" municipality="National Park" pageId="7" pageNumber="470" specimenCode="CW 38.5, CL 27.7, FMNH 7589" specimenCount="1" typeStatus="holotype">
<typeStatus id="072981826769B147FEC9A363ECEB5E14" box="[324,429,1634,1659]" pageId="7" pageNumber="470" type="holotype">Holotype</typeStatus>
: 
<collectingCountry id="A0857FB06769B147FE34A363EF045E14" box="[441,578,1634,1659]" name="Madagascar" pageId="7" pageNumber="470">Madagascar</collectingCountry>
: adult J (
<specimenCode id="8834975B6769B147FD3CA363EE555E14" box="[689,787,1634,1659]" collectionCode="CW" pageId="7" pageNumber="470">CW 38.5</specimenCode>
, 
<specimenCode id="8834975B6769B147FC92A363EE325E14" box="[799,884,1634,1659]" collectionCode="CL" pageId="7" pageNumber="470">CL 27.7</specimenCode>
, CH 12.5, 
<collectionCode id="BE83A7E56769B147FC6BA363E94C5E14" box="[998,1034,1634,1659]" pageId="7" pageNumber="470">FW</collectionCode>
<quantity id="1F6A92C56769B147FB9FA363E93F5E14" box="[1042,1145,1634,1659]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="7" pageNumber="470" unit="mm" value="12.4">12.4 mm</quantity>
), 
<location id="DD4D69FB6769B147FF2DA384EC6A5EF1" LSID="urn:lsid:plazi:treatment:503B8E366769B14AFE65A01AECAF5D28:DD4D69FB6769B147FF2DA384EC6A5EF1" box="[160,300,1669,1694]" country="Madagascar" county="Bemanivy River" latitude="-14.4267" longLatPrecision="5" longitude="49.60833" municipality="National Park" name="Sava Region" pageId="7" pageNumber="470">Sava Region</location>
, 
<taxonomicName id="1F9244A36769B147FEB8A384ECD05EF1" box="[309,406,1669,1694]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="470" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
<collectingMunicipality id="3849A55A6769B147FE10A384EF735EF1" box="[413,565,1669,1694]" pageId="7" pageNumber="470">National Park</collectingMunicipality>
, 
<locationDeviation id="C99F26026769B147FDB2A384EE605EF1" box="[575,806,1669,1694]" location="Doany" pageId="7" pageNumber="470">
<quantity id="1F6A92C56769B147FDB2A384EFD15EF1" box="[575,663,1669,1694]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="7" pageNumber="470" unit="km" value="11.5">11.5 km</quantity>
SE of Doany
</locationDeviation>
(
<geoCoordinate id="BDA659E76769B147FCBFA384EEE25EF1" box="[818,932,1669,1694]" degrees="14.4267" direction="south" orientation="latitude" pageId="7" pageNumber="470" precision="5" value="-14.4267">14.4267°S</geoCoordinate>
, 
<geoCoordinate id="BDA659E76769B147FC23A384E9685EF1" box="[942,1070,1669,1694]" degrees="49.60833" direction="east" orientation="longitude" pageId="7" pageNumber="470" precision="1" value="49.60833">49.60833°E</geoCoordinate>
), 
<quantity id="1F6A92C56769B147FBB0A384E9C55EF1" box="[1085,1155,1669,1694]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="7" pageNumber="470" unit="m" value="810.0">
<elevation id="53BFD8136769B147FBB0A384E9C55EF1" box="[1085,1155,1669,1694]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="7" pageNumber="470" unit="m" value="810.0">810 m</elevation>
</quantity>
<collectionCode id="BE83A7E56769B147FF2DA3A6ED8B5EAF" box="[160,205,1703,1728]" pageId="7" pageNumber="470">ASL</collectionCode>
, in mostly undisturbed upper lowland forest, in tributary of 
<collectorName id="75675AF66769B147FCEEA3A6E9535EAF" box="[867,1045,1703,1728]" pageId="7" pageNumber="470">
<collectingCounty id="314C47AC6769B147FCEEA3A6E9575EAF" box="[867,1041,1703,1728]" pageId="7" pageNumber="470">Bemanivy River</collectingCounty>
.
</collectorName>
Collected by hand during day on river bed, mostly rocky, with some open basins with sandy alluvium, coll. 
<collectorName id="75675AF66766B148FECCA5B0ECB658A5" box="[321,496,177,202]" pageId="8" pageNumber="471">S. M. Goodman</collectorName>
, 
<date id="AC2C19E06766B148FE77A5B0EFF158A5" box="[506,695,177,202]" pageId="8" pageNumber="471" value="2001-10-16">
<collectingDate id="BC68E0086766B148FE77A5B0EFF158A5" box="[506,695,177,202]" pageId="8" pageNumber="471" value="2001-10-16">16 October 2001</collectingDate>
</date>
(
<specimenCode id="8834975B6766B148FD4EA5B0EE0C58A5" box="[707,842,177,202]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="471" type="Museum">FMNH 7589</specimenCode>
) (GenBank MT749742).
</materialsCitation>
</paragraph>
<paragraph id="D82D3F206766B148FF4DA5D5E9C25A6D" blockId="8.[160,1156,142,514]" pageId="8" pageNumber="471">
<materialsCitation id="68FA357D6766B148FF4DA5D5EC9D595D" ID-GBIF-Occurrence="4089470308" collectingDate="2002-02-08" collectionCode="CL, CW, FMNH" pageId="8" pageNumber="471" specimenCode="CW 28.4, CL 19.7, FMNH 7579" typeStatus="paratype">
<typeStatus id="072981826766B148FF4DA5D5EC765882" box="[192,304,212,237]" pageId="8" pageNumber="471" type="paratype">Paratypes</typeStatus>
: 
<collectingCountry id="A0857FB06766B148FEB0A5D5EC805882" box="[317,454,212,237]" name="Madagascar" pageId="8" pageNumber="471">Madagascar</collectingCountry>
: subadult J (
<specimenCode id="8834975B6766B148FDEFA5D5EF835882" box="[610,709,212,237]" collectionCode="CW" pageId="8" pageNumber="471">CW 28.4</specimenCode>
, 
<specimenCode id="8834975B6766B148FD5FA5D5EE6F5882" box="[722,809,212,237]" collectionCode="CL" pageId="8" pageNumber="471">CL 19.7</specimenCode>
, CH 11.0, 
<collectionCode id="BE83A7E56766B148FC12A5D5EE855882" box="[927,963,212,237]" pageId="8" pageNumber="471">FW</collectionCode>
11.0 mm), same locality as for holotype, coll. V. Soarimalala and M. Raheriarisena, 
<date id="AC2C19E06766B148FCF7A5F7E9695960" box="[890,1071,246,271]" pageId="8" pageNumber="471" value="2002-02-08">
<collectingDate id="BC68E0086766B148FCF7A5F7E9695960" box="[890,1071,246,271]" pageId="8" pageNumber="471" value="2002-02-08">8 February 2002</collectingDate>
</date>
(
<specimenCode id="8834975B6766B148FBB1A5F7ED9E595D" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="471" type="Museum">FMNH 7579</specimenCode>
) (
<location id="DD4D69FB6766B148FF60A418EC15595D" LSID="urn:lsid:plazi:treatment:503B8E366769B14AFE65A01AECAF5D28:DD4D69FB6766B148FF60A418EC15595D" box="[237,339,281,306]" country="Switzerland" county="GenBank" municipality="GenBank" name="GenBank" pageId="8" pageNumber="471">GenBank</location>
MT749741)
</materialsCitation>
; 
<materialsCitation id="68FA357D6766B148FE6BA418EE6D5918" ID-GBIF-Occurrence="4089470307" collectionCode="CW, CL, FMNH" pageId="8" pageNumber="471" specimenCode="CW 35.5, CL 25.9, CW 30.6, CL 21.3, CW 27.5, CL 21.6, FMNH 7589.2" specimenCount="1" specimenCount-female="1">
subadult 
<specimenCount id="CE94F4A96766B148FDDCA418EF3F595F" box="[593,633,281,304]" pageId="8" pageNumber="471" type="female">♀♀</specimenCount>
(
<specimenCode id="8834975B6766B148FD05A418EFAC595D" box="[648,746,281,306]" collectionCode="CW" pageId="8" pageNumber="471">CW 35.5</specimenCode>
, 
<specimenCode id="8834975B6766B148FD7BA418EE0A595D" box="[758,844,281,306]" collectionCode="CL" pageId="8" pageNumber="471">CL 25.9</specimenCode>
, CH 11.4, 
<collectionCode id="BE83A7E56766B148FC33A418EEA4595D" box="[958,994,281,306]" pageId="8" pageNumber="471">FW</collectionCode>
<quantity id="1F6A92C56766B148FC66A418E917595D" box="[1003,1105,281,306]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="8" pageNumber="471" unit="mm" value="11.4">11.4 mm</quantity>
, 
<specimenCode id="8834975B6766B148FBD0A418ED97593A" collectionCode="CW" pageId="8" pageNumber="471">CW 30.6</specimenCode>
, 
<specimenCode id="8834975B6766B148FF51A43DEC76593A" box="[220,304,316,341]" collectionCode="CL" pageId="8" pageNumber="471">CL 21.3</specimenCode>
, CH 9.8, 
<collectionCode id="BE83A7E56766B148FE1FA43DECF0593A" box="[402,438,316,341]" pageId="8" pageNumber="471">FW</collectionCode>
<quantity id="1F6A92C56766B148FE30A43DEF52593A" box="[445,532,316,341]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="8" pageNumber="471" unit="mm" value="9.8">9.8 mm</quantity>
, 
<specimenCode id="8834975B6766B148FD92A43DEF38593A" box="[543,638,316,341]" collectionCode="CW" pageId="8" pageNumber="471">CW 27.5</specimenCode>
, 
<specimenCode id="8834975B6766B148FD04A43DEF9B593A" box="[649,733,316,341]" collectionCode="CL" pageId="8" pageNumber="471">CL 21.6</specimenCode>
, CH 9.3, 
<collectionCode id="BE83A7E56766B148FCB2A43DEE25593A" box="[831,867,316,341]" pageId="8" pageNumber="471">FW</collectionCode>
<quantity id="1F6A92C56766B148FCE7A43DEE87593A" box="[874,961,316,341]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="8" pageNumber="471" unit="mm" value="9.3">9.3 mm</quantity>
), same collection data as for holotype (
<specimenCode id="8834975B6766B148FE1CA45FEF6D5918" box="[401,555,350,375]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="471" type="Museum">FMNH 7589.2</specimenCode>
) (
<collectingMunicipality id="3849A55A6766B148FDB3A45FEFE25918" box="[574,676,350,375]" pageId="8" pageNumber="471">GenBank</collectingMunicipality>
MT749745)
</materialsCitation>
; 
<materialsCitation id="68FA357D6766B148FCB8A45FE9C759F5" ID-GBIF-Occurrence="4089470305" collectionCode="FMNH, CW, CL" pageId="8" pageNumber="471" specimenCode="CW 35.7, CL 25.6, FMNH 7589.3" typeStatus="holotype">
subadult J (
<specimenCode id="8834975B6766B148FC4DA45FE9675918" box="[960,1057,350,375]" collectionCode="CW" pageId="8" pageNumber="471">CW 35.7</specimenCode>
, 
<specimenCode id="8834975B6766B148FBA6A45FE9C65918" box="[1067,1152,350,375]" collectionCode="CL" pageId="8" pageNumber="471">CL 25.6</specimenCode>
, CH 10.8, 
<collectionCode id="BE83A7E56766B148FE84A480EC6B59F5" box="[265,301,385,410]" pageId="8" pageNumber="471">FW</collectionCode>
<quantity id="1F6A92C56766B148FEBBA480ECD859F5" box="[310,414,385,410]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="8" pageNumber="471" unit="mm" value="10.8">10.8 mm</quantity>
), same details as 
<typeStatus id="072981826766B148FDE6A480EF8959F5" box="[619,719,385,410]" pageId="8" pageNumber="471" type="holotype">holotype</typeStatus>
(
<specimenCode id="8834975B6766B148FD52A480EE3A59F5" box="[735,892,385,410]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="471" type="Museum">FMNH 7589.3</specimenCode>
) (
<collectingCounty id="314C47AC6766B148FC1CA480EEB159F5" box="[913,1015,385,410]" pageId="8" pageNumber="471">GenBank</collectingCounty>
MT749746)
</materialsCitation>
; 
<materialsCitation id="68FA357D6766B148FF2DA4A5ECD759B0" ID-GBIF-Occurrence="4089470303" collectingDate="2001-10-16" collectionCode="FMNH" pageId="8" pageNumber="471" specimenCode="FMNH 7590" typeStatus="holotype">
same locality as for 
<typeStatus id="072981826766B148FE1AA4A5EF4759D2" box="[407,513,420,445]" pageId="8" pageNumber="471" type="holotype">holotype</typeStatus>
, coll. S. M. Goodman, 
<date id="AC2C19E06766B148FC99A4A5EE9B59D2" box="[788,989,420,445]" pageId="8" pageNumber="471" value="2001-10-16">
<collectingDate id="BC68E0086766B148FC99A4A5EE9B59D2" box="[788,989,420,445]" pageId="8" pageNumber="471" value="2001-10-16">16 October 2001</collectingDate>
</date>
(
<specimenCode id="8834975B6766B148FC62A4A5E93859D2" box="[1007,1150,420,445]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="471" type="Museum">FMNH 7590</specimenCode>
) (
<location id="DD4D69FB6766B148FF2BA4C7EC4A59B0" LSID="urn:lsid:plazi:treatment:503B8E366769B14AFE65A01AECAF5D28:DD4D69FB6766B148FF2BA4C7EC4A59B0" box="[166,268,454,479]" country="Switzerland" municipality="Cl" name="GenBank" pageId="8" pageNumber="471">GenBank</location>
MT749744)
</materialsCitation>
; 
<materialsCitation id="68FA357D6766B148FE14A4C7E9C25A6D" ID-GBIF-Occurrence="4089470304" collectingDate="2001-10-13" collectionCode="FMNH" pageId="8" pageNumber="471" specimenCode="FMNH 7592" specimenCount="1" specimenCount-female="1">
subadult 
<specimenCount id="CE94F4A96766B148FD8CA4C6EF5359B1" box="[513,533,455,478]" pageId="8" pageNumber="471" type="female">♀</specimenCount>
(
<collectionCode id="BE83A7E56766B148FDADA4C7EF0E59B0" box="[544,584,454,479]" pageId="8" pageNumber="471">CW</collectionCode>
22.0, 
<collectingMunicipality id="3849A55A6766B148FD05A4C7EFD859B0" box="[648,670,454,479]" pageId="8" pageNumber="471">Cl</collectingMunicipality>
<quantity id="1F6A92C56766B148FD29A4C7EE4159B0" box="[676,775,454,479]" metricMagnitude="-2" metricUnit="m" metricValue="1.71" pageId="8" pageNumber="471" unit="mm" value="17.1">17.1 mm</quantity>
; CH 7.5, 
<collectionCode id="BE83A7E56766B148FCEEA4C7EEC159B0" box="[867,903,454,479]" pageId="8" pageNumber="471">FW</collectionCode>
<quantity id="1F6A92C56766B148FC01A4C7EEA459B0" box="[908,994,454,479]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="471" unit="mm" value="7.5">7.5 mm</quantity>
), same locality as for holotype, coll. S. M. Goodman, 
<date id="AC2C19E06766B148FDB4A4E8EFB55A6D" box="[569,755,489,514]" pageId="8" pageNumber="471" value="2001-10-13">
<collectingDate id="BC68E0086766B148FDB4A4E8EFB55A6D" box="[569,755,489,514]" pageId="8" pageNumber="471" value="2001-10-13">13 October 2001</collectingDate>
</date>
(
<specimenCode id="8834975B6766B148FD73A4E8EEC25A6D" box="[766,900,489,514]" collectionCode="FMNH" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="8" pageNumber="471" type="Museum">FMNH 7592</specimenCode>
) (
<location id="DD4D69FB6766B148FC18A4E8EEBD5A6D" LSID="urn:lsid:plazi:treatment:503B8E366769B14AFE65A01AECAF5D28:DD4D69FB6766B148FC18A4E8EEBD5A6D" box="[917,1019,489,514]" country="Switzerland" name="GenBank" pageId="8" pageNumber="471">GenBank</location>
MT749743).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="90886CAB6766B148FF2DA72FEC265A05" box="[160,352,558,618]" pageId="8" pageNumber="471" type="diagnosis">
<paragraph id="D82D3F206766B148FF2DA72FEC525A27" blockId="8.[160,352,558,618]" box="[160,276,558,584]" pageId="8" pageNumber="471">
<emphasis id="EAE6E3326766B148FF2DA72FEC525A27" bold="true" box="[160,276,558,584]" italics="true" pageId="8" pageNumber="471">Diagnosis</emphasis>
</paragraph>
<paragraph id="D82D3F206766B148FF2DA750EC265A05" blockId="8.[160,352,558,618]" box="[160,352,593,618]" pageId="8" pageNumber="471">As for the genus.</paragraph>
</subSubSection>
<subSubSection id="90886CAB6766B149FF2DA797EE315B49" lastPageId="9" lastPageNumber="472" pageId="8" pageNumber="471" type="description">
<paragraph id="D82D3F206766B148FF2DA797EC605ADF" blockId="8.[160,1156,662,1728]" box="[160,294,662,688]" pageId="8" pageNumber="471">
<emphasis id="EAE6E3326766B148FF2DA797EC605ADF" bold="true" box="[160,294,662,688]" italics="true" pageId="8" pageNumber="471">Description</emphasis>
</paragraph>
<paragraph id="D82D3F206766B149FF2DA7BBEF465882" blockId="8.[160,1156,662,1728]" lastBlockId="9.[160,1156,142,549]" lastPageId="9" lastPageNumber="472" pageId="8" pageNumber="471">
Based on 
<typeStatus id="072981826766B148FE80A7BBEC325ABC" box="[269,372,698,723]" pageId="8" pageNumber="471" type="holotype">holotype</typeStatus>
, adult J. Carapace outline transversely oval, moderate height (CH/FW 1.0) gently inflated, CH equal to FW; front broad (FW/CW = 0.3), deflexed covering part of antennulular fossae; dorsal surface with faint carinae in anterolateral, posterolateral regions, otherwise smooth. Anterolateral margin between exorbital, epibranchial teeth convex, granulated, lacking intermediate tooth; epibranchial tooth reduced to small granule, close to exorbital tooth, positioned in line with postorbital margin; lateral margin evenly curved outward, lined with granules, posterior end curving inward, not continuous with posterolateral margin; postfrontal crest faint, incomplete, not traversing entire carapace, ending before meeting epibranchial teeth, epigastric crests faint, in advanced position, in line with postorbital margin; postorbital crests, cardiac, urogastric sulci faint, semicircular sulcus deep, cervical sulcus faint, short, not meeting postorbital crest (
<figureCitation id="40A923A56766B148FBB1A114EDEF5C3F" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="8" pageNumber="471">Figure 1</figureCitation>
(a)). Suborbital, subhepatic regions of branchiostegite with faint granules; pterygostomial region smooth except for granules along epimeral (longitudinal) sulcus; vertical sulcus on branchiostegite curved, granular, running downward from base of epibranchial tooth to epimeral sulcus (
<figureCitation id="40A923A56766B148FE36A19EEF535CD7" box="[443,533,1183,1208]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="8" pageNumber="471">Figure 2</figureCitation>
(a)). Epistomial tooth triangular, deflexed, edges smooth (
<figureCitation id="40A923A56766B148FF25A1C3EC405CB4" box="[168,262,1218,1243]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="8" pageNumber="471">Figure 2</figureCitation>
(a)). Mandibular palp terminal article bilobed; anterior lobe on terminal article conspicuous, medium-sized (MPAL/MPTA = 0.4) (
<figureCitation id="40A923A56766B148FD40A1E4EE6B5C91" box="[717,813,1253,1278]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="8" pageNumber="471">Figure 2</figureCitation>
(i)). Exopod of third maxilliped reaching to lower half of merus, exopod with short flagellum (less than half merus length), ischium of third maxilliped with faint shallow vertical sulcus (
<figureCitation id="40A923A56766B148FCCAA02BEEE25D2C" box="[839,932,1322,1347]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="8" pageNumber="471">Figure 2</figureCitation>
(g,h)). Sternal suture S1/2 short, faint; S2/3 deep, completely traversing sternum; S3/4 broad, U-shaped, deepest at edges, not meeting anterior margin of sternopleonal cavity; S4/5 meeting pleon at level of telson/PL6 suture; S6/7 meeting midpoint of lateral margin of PL6; episternal sulci S4/E4, S5/E5, S6/E6, S7/E7 absent, smooth (
<figureCitation id="40A923A56766B148FD09A0B4EF9A5DA1" box="[644,732,1461,1486]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="8" pageNumber="471">Figure 1</figureCitation>
(b)). Left chela larger, proximal half of cutting edge of fixed finger (pollex of propodus) with one large molar flanked by small teeth, lined with small teeth distally; cutting edge of movable finger (dactylus) lined with small teeth, lacking large teeth (
<figureCitation id="40A923A56766B148FD9EA31CEF375E59" box="[531,625,1565,1590]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="8" pageNumber="471">Figure 2</figureCitation>
(c)); right chela slightly smaller than left chela, dentition same as left chela (
<figureCitation id="40A923A56766B148FE70A33EEF195E37" box="[509,607,1599,1624]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="8" pageNumber="471">Figure 2</figureCitation>
(b)); lower margin of propodus of both chelae conspicuously indented medially (
<figureCitation id="40A923A56766B148FDA0A363EFC85E14" box="[557,654,1634,1659]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="8" pageNumber="471">Figure 2</figureCitation>
(b,c)). Inner margin of cheliped carpus flattened, broad, distal tooth on inner margin small, pointed; proximal tooth smaller, acute, small granules between teeth, and following proximal tooth (
<figureCitation id="40A923A56766B148FCC1A3A6EEEF5EAF" box="[844,937,1703,1728]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="8" pageNumber="471">Figure 2</figureCitation>
(f)). Inferior margins of cheliped merus both lined with small, rounded teeth, distal tooth largest; cheliped ischium margins smooth, rounded (
<figureCitation id="40A923A56767B149FDB1A5B0EFDA58A5" box="[572,668,177,202]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="9" pageNumber="472">Figure 2</figureCitation>
(d,e)). Ambulatory legs P2–5 short (ΣP2–5/ CW = 5.4) (
<figureCitation id="40A923A56767B149FE91A5D5EC325882" box="[284,372,212,237]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="9" pageNumber="472">Figure 1</figureCitation>
(a); 
<tableCitation id="95100A9B6767B149FE2CA5D5ECB25882" box="[417,500,212,237]" captionStart="Table 1" captionStartId="1.[160,216,1316,1339]" captionTargetPageId="1" captionText="Table 1. Vichai cyanalepou sp. nov. male holotype (FMNH 7589). Length in mm of the five articles (ischium, merus, carpus, propodus, dactylus) of the cheliped (P1) and the four ambulatory legs (P2–5), and the leg length in relation to the carapace width (CW). The sum of the length of each article of the four ambulatory legs divided by the carapace width is also shown (Total of 5 articles/CW), as is the sum of the relative length of the four ambulatory legs (ΣP2–5/CW = 5.35), placing this species in the short ambulatory leg size class (short: 5.0–5.9; medium: 6.0–6.9; long:&gt;7.0)." httpUri="http://table.plazi.org/id/8CED6FA8676FB141FF2DA025EE1C5DA2" pageId="9" pageNumber="472" tableUuid="8CED6FA8676FB141FF2DA025EE1C5DA2">Table 1</tableCitation>
).
</paragraph>
<paragraph id="D82D3F206767B149FF4DA5F7E90E5A4A" blockId="9.[160,1156,142,549]" pageId="9" pageNumber="472">
Pleon slim, lateral margins taper distally, widest at PL3, narrowest at telson/PL6 suture; PL6 relatively broad; telson outline with straight lateral margins, apex rounded (
<figureCitation id="40A923A56767B149FB8DA418E912595D" box="[1024,1108,281,306]" captionStart="Figure 1" captionStartId="3.[160,225,1754,1777]" captionTargetBox="[172,1144,142,1722]" captionTargetId="figure-3@3.[172,1144,142,1722]" captionTargetPageId="3" captionText="Figure 1. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park,Sava region,northernMadagascar,whole animal:(a) dorsal view;(b) cephalothorax,ventral view.Scale bar: a = 10.8 mm; b = 5.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7859821" httpUri="https://zenodo.org/record/7859821/files/figure.png" pageId="9" pageNumber="472">Figure 1</figureCitation>
(b)). G1TA medium length (G1TA/G1SA = 0.3), slim, basal two-thirds directed outward,distal onethird curving upward, tapering to blunt open tip; G1TA ventral side with faint sulcus between G1TA and G1SA (
<figureCitation id="40A923A56767B149FE54A480EF7F59F5" box="[473,569,385,410]" captionStart="Figure 3" captionStartId="6.[160,225,1101,1124]" captionTargetBox="[163,1153,142,1069]" captionTargetId="figure-190@6.[163,1153,142,1069]" captionTargetPageId="6" captionText="Figure 3. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) right G1, ventral view; (b) right G1, dorsal view; (c) right G1, superior view; (d) right G2, ventral view. Scale bar: a, b = 1.4 mm; c = 1.1 mm; d = 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7859827" httpUri="https://zenodo.org/record/7859827/files/figure.png" pageId="9" pageNumber="472">Figure 3</figureCitation>
(a)); G1TA dorsal side with broad, trapezoid dorsal membrane (DM) at G1TA-G1SA junction; DM superior margin arrowhead-shaped, inferior margin U-shaped, lateral margin short, mesial margin elongated (
<figureCitation id="40A923A56767B149FCE7A4C7EE8259B0" box="[874,964,454,479]" captionStart="Figure 3" captionStartId="6.[160,225,1101,1124]" captionTargetBox="[163,1153,142,1069]" captionTargetId="figure-190@6.[163,1153,142,1069]" captionTargetPageId="6" captionText="Figure 3. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) right G1, ventral view; (b) right G1, dorsal view; (c) right G1, superior view; (d) right G2, ventral view. Scale bar: a, b = 1.4 mm; c = 1.1 mm; d = 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7859827" httpUri="https://zenodo.org/record/7859827/files/figure.png" pageId="9" pageNumber="472">Figure 3</figureCitation>
(b)); G1SA lacking raised rounded shoulder on external margin near G1TA-G1SA junction (
<figureCitation id="40A923A56767B149FC2BA4E8EEB95A6D" box="[934,1023,489,514]" captionStart="Figure 3" captionStartId="6.[160,225,1101,1124]" captionTargetBox="[163,1153,142,1069]" captionTargetId="figure-190@6.[163,1153,142,1069]" captionTargetPageId="6" captionText="Figure 3. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) right G1, ventral view; (b) right G1, dorsal view; (c) right G1, superior view; (d) right G2, ventral view. Scale bar: a, b = 1.4 mm; c = 1.1 mm; d = 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7859827" httpUri="https://zenodo.org/record/7859827/files/figure.png" pageId="9" pageNumber="472">Figure 3</figureCitation>
(a–b)); G2TA distinctly elongated, flagellum-like (G2TA/G2SA = 0.85) with pointed tip (
<figureCitation id="40A923A56767B149FC4EA70DE9585A4A" box="[963,1054,524,549]" captionStart="Figure 3" captionStartId="6.[160,225,1101,1124]" captionTargetBox="[163,1153,142,1069]" captionTargetId="figure-190@6.[163,1153,142,1069]" captionTargetPageId="6" captionText="Figure 3. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) right G1, ventral view; (b) right G1, dorsal view; (c) right G1, superior view; (d) right G2, ventral view. Scale bar: a, b = 1.4 mm; c = 1.1 mm; d = 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7859827" httpUri="https://zenodo.org/record/7859827/files/figure.png" pageId="9" pageNumber="472">Figure 3</figureCitation>
(d)).
</paragraph>
<paragraph id="D82D3F206767B149FF2DA756ED885A1E" blockId="9.[160,1156,599,694]" box="[160,206,599,625]" pageId="9" pageNumber="472">
<emphasis id="EAE6E3326767B149FF2DA756ED885A1E" bold="true" box="[160,206,599,625]" italics="true" pageId="9" pageNumber="472">Size</emphasis>
</paragraph>
<paragraph id="D82D3F206767B149FF2DA77AECC55AD9" blockId="9.[160,1156,599,694]" pageId="9" pageNumber="472">
Medium-sized species, largest known specimen CW 
<quantity id="1F6A92C56767B149FD75A77AEE195AFB" box="[760,863,635,660]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" pageId="9" pageNumber="472" unit="mm" value="38.5">38.5 mm</quantity>
, pubertal moult between CWs 36 and 
<quantity id="1F6A92C56767B149FEA0A79CEC395AD9" box="[301,383,669,694]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="9" pageNumber="472" unit="mm" value="38.0">38 mm</quantity>
.
</paragraph>
<paragraph id="D82D3F206767B149FF2DA7E8EDA85B6C" blockId="9.[160,887,745,806]" box="[160,238,745,771]" pageId="9" pageNumber="472">
<emphasis id="EAE6E3326767B149FF2DA7E8EDA85B6C" bold="true" box="[160,238,745,771]" italics="true" pageId="9" pageNumber="472">Colour</emphasis>
</paragraph>
<paragraph id="D82D3F206767B149FF2DA60CEE315B49" blockId="9.[160,887,745,806]" box="[160,887,781,806]" pageId="9" pageNumber="472">Unknown in life. Preserved specimens are uniformly pale brown.</paragraph>
</subSubSection>
<subSubSection id="90886CAB6767B149FF2DA659EF815BD8" pageId="9" pageNumber="472" type="distribution">
<paragraph id="D82D3F206767B149FF2DA659EC6A5B1D" blockId="9.[160,1155,856,951]" box="[160,300,856,882]" pageId="9" pageNumber="472">
<emphasis id="EAE6E3326767B149FF2DA659EC6A5B1D" bold="true" box="[160,300,856,882]" italics="true" pageId="9" pageNumber="472">Distribution</emphasis>
</paragraph>
<paragraph id="D82D3F206767B149FF2DA67DEF815BD8" blockId="9.[160,1155,856,951]" pageId="9" pageNumber="472">
<taxonomicName id="1F9244A36767B149FF2DA67DEC215BFA" authority="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever, 2023" authorityName="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever" authorityYear="2023" box="[160,359,892,917]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
<emphasis id="EAE6E3326767B149FF2DA67DEC215BFA" box="[160,359,892,917]" italics="true" pageId="9" pageNumber="472">Vichai cyanalepou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1D55E496767B149FEFDA67DEC8E5BFA" box="[368,456,892,917]" pageId="9" pageNumber="472" rank="species">sp. nov.</taxonomicNameLabel>
<materialsCitation id="68FA357D6767B149FE42A67DEF855BD8" ID-GBIF-Occurrence="4089470302" country="Madagascar" location="National Park" municipality="Sava Region" pageId="9" pageNumber="472" specimenCount="1">
is only known from the 
<typeStatus id="072981826767B149FD6DA67DEE545BFA" box="[736,786,892,917]" pageId="9" pageNumber="472">type</typeStatus>
locality in the 
<taxonomicName id="1F9244A36767B149FC31A67DE95B5BFA" box="[956,1053,892,917]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
<location id="DD4D69FB6767B149FBA8A67DED965BD8" LSID="urn:lsid:plazi:treatment:503B8E366769B14AFE65A01AECAF5D28:DD4D69FB6767B149FBA8A67DED965BD8" country="Madagascar" municipality="Sava Region" name="National Park" pageId="9" pageNumber="472">National Park</location>
in the 
<collectingMunicipality id="3849A55A6767B149FEACA69FECEC5BD8" box="[289,426,926,951]" pageId="9" pageNumber="472">Sava Region</collectingMunicipality>
of northern 
<collectingCountry id="A0857FB06767B149FDB7A69FEF855BD8" box="[570,707,926,951]" name="Madagascar" pageId="9" pageNumber="472">Madagascar</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="90886CAB6767B149FF2DA6EBEFEF5EAF" pageId="9" pageNumber="472" type="materials_examined">
<paragraph id="D82D3F206767B149FF2DA6EBEC705C6B" blockId="9.[160,1155,1002,1097]" box="[160,310,1002,1028]" pageId="9" pageNumber="472">
<emphasis id="EAE6E3326767B149FF2DA6EBEC705C6B" bold="true" box="[160,310,1002,1028]" italics="true" pageId="9" pageNumber="472">Type locality</emphasis>
</paragraph>
<paragraph id="D82D3F206767B149FF2DA10CE9325C26" blockId="9.[160,1155,1002,1097]" pageId="9" pageNumber="472">
<materialsCitation id="68FA357D6767B149FF2DA10CE9375C26" ID-GBIF-Occurrence="4089470306" collectionCode="ASL" country="Madagascar" county="Bemanavy River" elevation="810" latitude="-14.4267" location="Sava Region" longLatPrecision="5" longitude="49.60833" municipality="Bemanivy River" pageId="9" pageNumber="472" specimenCount="1">
<collectingCountry id="A0857FB06767B149FF2DA10CEC6F5C49" box="[160,297,1037,1062]" name="Madagascar" pageId="9" pageNumber="472">Madagascar</collectingCountry>
, 
<location id="DD4D69FB6767B149FEB8A10CEC825C49" LSID="urn:lsid:plazi:treatment:503B8E366769B14AFE65A01AECAF5D28:DD4D69FB6767B149FEB8A10CEC825C49" box="[309,452,1037,1062]" country="Madagascar" county="Bemanavy River" latitude="-14.4267" longLatPrecision="5" longitude="49.60833" municipality="Bemanivy River" name="Sava Region" pageId="9" pageNumber="472">Sava Region</location>
, 
<taxonomicName id="1F9244A36767B149FE5DA10CEF775C49" box="[464,561,1037,1062]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
<location id="DD4D69FB6767B149FDB6A10CEF905C49" LSID="urn:lsid:plazi:treatment:503B8E366769B14AFE65A01AECAF5D28:DD4D69FB6767B149FDB6A10CEF905C49" box="[571,726,1037,1062]" country="Madagascar" county="Bemanavy River" latitude="-14.4267" longLatPrecision="5" longitude="49.60833" municipality="Bemanivy River" name="National Park" pageId="9" pageNumber="472">National Park</location>
, 
<locationDeviation id="C99F26026767B149FD6FA10CEE915C49" box="[738,983,1037,1062]" location="Doany" pageId="9" pageNumber="472">
<quantity id="1F6A92C56767B149FD6FA10CEE785C49" box="[738,830,1037,1062]" metricMagnitude="4" metricUnit="m" metricValue="1.15" pageId="9" pageNumber="472" unit="km" value="11.5">11.5 km</quantity>
SE of Doany
</locationDeviation>
, in tributary of 
<collectingMunicipality id="3849A55A6767B149FF2DA131EC0A5C26" box="[160,332,1072,1097]" pageId="9" pageNumber="472">Bemanivy River</collectingMunicipality>
near junction with 
<collectingCounty id="314C47AC6767B149FDA7A131EF995C26" box="[554,735,1072,1097]" pageId="9" pageNumber="472">Bemanavy River</collectingCounty>
, 
<quantity id="1F6A92C56767B149FD67A131EE765C26" box="[746,816,1072,1097]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="9" pageNumber="472" unit="m" value="810.0">
<elevation id="53BFD8136767B149FD67A131EE765C26" box="[746,816,1072,1097]" metricMagnitude="2" metricUnit="m" metricValue="8.1" pageId="9" pageNumber="472" unit="m" value="810.0">810 m</elevation>
</quantity>
<collectionCode id="BE83A7E56767B149FCB5A131EE245C26" box="[824,866,1072,1097]" pageId="9" pageNumber="472">ASL</collectionCode>
(
<geoCoordinate id="BDA659E76767B149FCE2A131EEA75C26" box="[879,993,1072,1097]" degrees="14.4267" direction="south" orientation="latitude" pageId="9" pageNumber="472" precision="5" value="-14.4267">14.4267°S</geoCoordinate>
, 
<geoCoordinate id="BDA659E76767B149FC66A131E92D5C26" box="[1003,1131,1072,1097]" degrees="49.60833" direction="east" orientation="longitude" pageId="9" pageNumber="472" precision="1" value="49.60833">49.60833°E</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="D82D3F206767B149FF2DA17DECD45CF9" blockId="9.[160,1157,1148,1728]" box="[160,402,1148,1174]" pageId="9" pageNumber="472">
<emphasis id="EAE6E3326767B149FF2DA17DECD45CF9" bold="true" box="[160,402,1148,1174]" italics="true" pageId="9" pageNumber="472">Habitat and ecology</emphasis>
</paragraph>
<paragraph id="D82D3F206767B149FF2DA19EEFEF5EAF" blockId="9.[160,1157,1148,1728]" pageId="9" pageNumber="472">
The rainforests of 
<taxonomicName id="1F9244A36767B149FEFBA19EEC9E5CD7" box="[374,472,1183,1208]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
National Park in Sava Province are the northernmost part of this World Heritage Site, which comprises a chain of six national parks distributed in the highlands of eastern 
<collectingCountry id="A0857FB06767B149FE62A1E4EF315C91" box="[495,631,1253,1278]" name="Madagascar" pageId="9" pageNumber="472">Madagascar</collectingCountry>
stretching from 
<taxonomicName id="1F9244A36767B149FCCEA1E4EEE35C91" box="[835,933,1253,1278]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
to as far south as the Anosyennes Mountains in the south-east of the island. This extensive World Heritage Site protects some of the last intact primary rainforest in 
<collectingCountry id="A0857FB06767B149FC32A02BE90C5D2C" box="[959,1098,1322,1347]" name="Madagascar" pageId="9" pageNumber="472">Madagascar</collectingCountry>
, and each park within it supports a rich biodiversity, including threatened species. The eastern mountains in 
<collectingCountry id="A0857FB06767B149FE2CA06EEF6F5DE7" box="[417,553,1391,1416]" name="Madagascar" pageId="9" pageNumber="472">Madagascar</collectingCountry>
receive a high annual rainfall that supports humid rainforest at lower elevations (
<quantity id="1F6A92C56767B149FD8EA093EFD55DC4" box="[515,659,1426,1451]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="9" pageNumber="472" unit="m" value="850.0" valueMax="1200.0" valueMin="500.0">500–1200 m</quantity>
<collectionCode id="BE83A7E56767B149FD11A093EF8A5DC4" box="[668,716,1426,1451]" pageId="9" pageNumber="472">ASL</collectionCode>
), drier sclerophyllous mountain forest (
<quantity id="1F6A92C56767B149FF2BA0B4EC005DA1" box="[166,326,1461,1486]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="9" pageNumber="472" unit="m" value="1400.0" valueMax="1600.0" valueMin="1200.0">1200–1600 m</quantity>
<collectionCode id="BE83A7E56767B149FEDFA0B4ECC45DA1" box="[338,386,1461,1486]" pageId="9" pageNumber="472">ASL</collectionCode>
), and montane forest on the highest slopes (
<quantity id="1F6A92C56767B149FC12A0B4E9795DA1" box="[927,1087,1461,1486]" metricMagnitude="3" metricUnit="m" metricValue="1.832" metricValueMax="2.064" metricValueMin="1.6" pageId="9" pageNumber="472" unit="m" value="1832.0" valueMax="2064.0" valueMin="1600.0">1600–2064 m</quantity>
<collectionCode id="BE83A7E56767B149FBC6A0B4E93D5DA1" box="[1099,1147,1461,1486]" pageId="9" pageNumber="472">ASL</collectionCode>
). 
<taxonomicName id="1F9244A36767B149FF2DA0D6EC445D9F" box="[160,258,1495,1520]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
National Park is a part of the 
<collectingCountry id="A0857FB06767B149FD1DA0D6EE5E5D9F" box="[656,792,1495,1520]" name="Madagascar" pageId="9" pageNumber="472">Madagascar</collectingCountry>
Eastern Highlands freshwater ecoregion (
<collectionCode id="BE83A7E56767B149FEA8A0FBEC2D5E7C" box="[293,363,1530,1555]" pageId="9" pageNumber="472">FEOW</collectionCode>
581; 
<bibRefCitation id="BC0342D16767B149FE38A0FBEF385E7C" author="Abell R &amp; Thieme ML &amp; Revenga C &amp; Bryer M &amp; Kottelat M &amp; Bogutskaya N &amp; Coad B &amp; Mandrak N &amp; Balderas SC &amp; Bussing W" box="[437,638,1530,1555]" pageId="9" pageNumber="472" pagination="403 - 414" refId="ref5869" refString="Abell R, Thieme ML, Revenga C, Bryer M, Kottelat M, Bogutskaya N, Coad B, Mandrak N, Balderas SC, Bussing W. 2008. Freshwater ecoregions of the world: a new map of biogeographic units for freshwater biodiversity conservation. BioScience. 58 (5): 403 - 414. doi: 10.1641 / B 580507." type="journal article" year="2008">Abell et al. 2008</bibRefCitation>
), and the new species was collected from under rocks on the bed of a tributary of the Bemanivy River in the mostly undisturbed upper reaches of the lowland forest belt of the park. Interestingly, 
<taxonomicName id="1F9244A36767B149FB97A33EED9B5E14" authority="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever, 2023" authorityName="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever" authorityYear="2023" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
<emphasis id="EAE6E3326767B149FB97A33EED9B5E14" italics="true" pageId="9" pageNumber="472">V. cyanalepou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1D55E496767B149FF6AA363EC045E14" box="[231,322,1634,1659]" pageId="9" pageNumber="472" rank="species">sp. nov.</taxonomicNameLabel>
was collected together with 
<specimenCount id="CE94F4A96767B149FD11A363EE615E14" box="[668,807,1634,1659]" pageId="9" pageNumber="472" type="subadult">six subadult</specimenCount>
specimens of 
<taxonomicName id="1F9244A36767B149FC59A363EFCD5EF1" authority="(A. Milne-Edwards, 1872)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1872" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="EAE6E3326767B149FC59A363EC235EF1" italics="true" pageId="9" pageNumber="472">Hydrothelphusa madagascariensis</emphasis>
(A. Milne-Edwards, 1872)
</taxonomicName>
(
<collectionCode id="BE83A7E56767B149FD14A384EFA75EF1" box="[665,737,1669,1694]" country="USA" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="472" type="Museum">FMNH</collectionCode>
7589.1, Genbank MT749717), which is, therefore, syntopic with the new species.
</paragraph>
</subSubSection>
<subSubSection id="90886CAB6764B14AFF2DA58FEFDA595C" pageId="10" pageNumber="473" type="etymology">
<paragraph id="D82D3F206764B14AFF2DA58FEC5B58C7" blockId="10.[160,1156,142,307]" box="[160,285,142,168]" pageId="10" pageNumber="473">
<emphasis id="EAE6E3326764B14AFF2DA58FEC5B58C7" bold="true" box="[160,285,142,168]" italics="true" pageId="10" pageNumber="473">Etymology</emphasis>
</paragraph>
<paragraph id="D82D3F206764B14AFF2DA5B3EFDA595C" blockId="10.[160,1156,142,307]" pageId="10" pageNumber="473">The species name ′cyanalepou̍ is derived from Greek and is a combination of two words ′cyan̍ for dark blue and ′alepou̍ for fox, a reference to the dark blue shirts that Leicester City play in, and the club mascot the fox. The specific epithet is used as a Latin noun in genitive singular and is treated as masculine.</paragraph>
</subSubSection>
<subSubSection id="90886CAB6764B14AFF2DA462ECAF5D28" pageId="10" pageNumber="473" type="discussion">
<paragraph id="D82D3F206764B14AFF2DA462EC405912" blockId="10.[160,1156,355,1351]" box="[160,262,355,381]" pageId="10" pageNumber="473">
<emphasis id="EAE6E3326764B14AFF2DA462EC405912" bold="true" box="[160,262,355,381]" italics="true" pageId="10" pageNumber="473">Remarks</emphasis>
</paragraph>
<paragraph id="D82D3F206764B14AFF2DA487E9C55A95" blockId="10.[160,1156,355,1351]" pageId="10" pageNumber="473">
<taxonomicName id="1F9244A36764B14AFF2DA487EC2E59F0" authority="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever, 2023" authorityName="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever" authorityYear="2023" box="[160,360,390,415]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
<emphasis id="EAE6E3326764B14AFF2DA487EC2E59F0" box="[160,360,390,415]" italics="true" pageId="10" pageNumber="473">Vichai cyanalepou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1D55E496764B14AFEFCA487EC8F59F0" box="[369,457,390,415]" pageId="10" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
was referred to as ′New genus E̍ in the phylogenetic tree of 
<bibRefCitation id="BC0342D16764B14AFF2DA4A8ECF459AD" author="Cumberlidge N &amp; Soma JB &amp; Leever EM &amp; Daniels SR" box="[160,434,425,450]" pageId="10" pageNumber="473" pagination="584 - 599" refId="ref6294" refString="Cumberlidge N, Soma JB, Leever EM, Daniels SR. 2020. New lineages within the Malagasy freshwater crab fauna: agora n. gen. for Thelphusa goudoti H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya Cumberlidge &amp; Sternberg, 2002 (Brachyura: Potamonautidae: Deckeniidae). J Crust Biol. 40 (5): 584 - 599. doi: 10.1093 / jcbiol / ruaa 050." type="journal article" year="2020">Cumberlidge et al. (2020</bibRefCitation>
, fig. 1). The 
<specimenCount id="CE94F4A96764B14AFDBAA4A8EFB259AD" box="[567,756,425,450]" pageId="10" pageNumber="473" type="generic">seven specimens</specimenCount>
sequenced in that study (described here and listed under Material and methods above) are all closely related to each other (with short branch lengths). These specimens comprise a genetically separate lineage from the nearest sister clade consisting of specimens labelled ′New genus C̍, ′New genus D̍, ′FMNH 13935̍ and ′ 
<taxonomicName id="1F9244A36764B14AFE28A735EF055A22" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[421,579,564,589]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="raimundi">
<emphasis id="EAE6E3326764B14AFE28A735EF055A22" box="[421,579,564,589]" italics="true" pageId="10" pageNumber="473">Foza raimundi</emphasis>
</taxonomicName>
̍ in 
<bibRefCitation id="BC0342D16764B14AFDE2A735EECE5A22" author="Cumberlidge N &amp; Soma JB &amp; Leever EM &amp; Daniels SR" box="[623,904,564,589]" pageId="10" pageNumber="473" pagination="584 - 599" refId="ref6294" refString="Cumberlidge N, Soma JB, Leever EM, Daniels SR. 2020. New lineages within the Malagasy freshwater crab fauna: agora n. gen. for Thelphusa goudoti H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya Cumberlidge &amp; Sternberg, 2002 (Brachyura: Potamonautidae: Deckeniidae). J Crust Biol. 40 (5): 584 - 599. doi: 10.1093 / jcbiol / ruaa 050." type="journal article" year="2020">Cumberlidge et al. (2020</bibRefCitation>
, fig. 1). Subsequently, ′New genus C̍ was identified using molecular and morphological evidence as 
<taxonomicName id="1F9244A36764B14AFB86A757EF085AFD" authority="Cumberlidge et al. (2021)" authorityName="Cumberlidge" authorityYear="2021" class="Malacostraca" family="Potamonautidae" genus="Crosnautes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="ranomafana">
<emphasis id="EAE6E3326764B14AFB86A757EC6E5AFD" italics="true" pageId="10" pageNumber="473">Crosnautes ranomafana</emphasis>
<bibRefCitation id="BC0342D16764B14AFEA2A778EF085AFD" author="Cumberlidge N &amp; Soma J &amp; Leever EM &amp; Clark PF &amp; Daniels SR" box="[303,590,633,658]" pageId="10" pageNumber="473" pagination="1 - 10" refId="ref6232" refString="Cumberlidge N, Soma J, Leever EM, Clark PF, Daniels SR. 2021. Description of a new genus and two new species of freshwater crabs (Decapoda: Brachyura: Potamonautidae) from southeastern Madagascar. J Crust Biol. 41 (3): 1 - 10. doi: 10.1093 / jcbiol / ruab 034." type="journal article" year="2021">Cumberlidge et al. (2021)</bibRefCitation>
</taxonomicName>
(see 
<bibRefCitation id="BC0342D16764B14AFD07A778EEDE5AFD" author="Cumberlidge N &amp; Soma J &amp; Leever EM &amp; Clark PF &amp; Daniels SR" box="[650,920,633,658]" pageId="10" pageNumber="473" pagination="1 - 10" refId="ref6232" refString="Cumberlidge N, Soma J, Leever EM, Clark PF, Daniels SR. 2021. Description of a new genus and two new species of freshwater crabs (Decapoda: Brachyura: Potamonautidae) from southeastern Madagascar. J Crust Biol. 41 (3): 1 - 10. doi: 10.1093 / jcbiol / ruab 034." type="journal article" year="2021">Cumberlidge et al. 2021</bibRefCitation>
), ′New genus D̍ was identified as a second species of this genus, 
<taxonomicName id="1F9244A36764B14AFD23A79DE90A5ADA" authority="Cumberlidge et al. (2021)" authorityName="Cumberlidge" authorityYear="2021" box="[686,1100,668,693]" class="Malacostraca" family="Potamonautidae" genus="Crosnautes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="alainus">
<emphasis id="EAE6E3326764B14AFD23A79DEE5D5ADA" box="[686,795,668,693]" italics="true" pageId="10" pageNumber="473">C. alainus</emphasis>
<bibRefCitation id="BC0342D16764B14AFCABA79DE90A5ADA" author="Cumberlidge N &amp; Soma J &amp; Leever EM &amp; Clark PF &amp; Daniels SR" box="[806,1100,668,693]" pageId="10" pageNumber="473" pagination="1 - 10" refId="ref6232" refString="Cumberlidge N, Soma J, Leever EM, Clark PF, Daniels SR. 2021. Description of a new genus and two new species of freshwater crabs (Decapoda: Brachyura: Potamonautidae) from southeastern Madagascar. J Crust Biol. 41 (3): 1 - 10. doi: 10.1093 / jcbiol / ruab 034." type="journal article" year="2021">Cumberlidge et al. (2021)</bibRefCitation>
</taxonomicName>
(see 
<bibRefCitation id="BC0342D16764B14AFF2DA7BFECE95AB8" author="Cumberlidge N &amp; Soma J &amp; Leever EM &amp; Clark PF &amp; Daniels SR" box="[160,431,702,727]" pageId="10" pageNumber="473" pagination="1 - 10" refId="ref6232" refString="Cumberlidge N, Soma J, Leever EM, Clark PF, Daniels SR. 2021. Description of a new genus and two new species of freshwater crabs (Decapoda: Brachyura: Potamonautidae) from southeastern Madagascar. J Crust Biol. 41 (3): 1 - 10. doi: 10.1093 / jcbiol / ruab 034." type="journal article" year="2021">Cumberlidge et al. 2021</bibRefCitation>
), and ′ 
<taxonomicName id="1F9244A36764B14AFE71A7BFEFDF5AB8" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[508,665,702,727]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="raimundi">
<emphasis id="EAE6E3326764B14AFE71A7BFEFDF5AB8" box="[508,665,702,727]" italics="true" pageId="10" pageNumber="473">Foza raimundi</emphasis>
</taxonomicName>
̍ (FMNH 7438) and FMNH 13935 were both confirmed as 
<taxonomicName id="1F9244A36764B14AFEB8A7E0EE675A95" authority="Reed and Cumberlidge (2006)" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[309,801,737,762]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="raimundi">
<emphasis id="EAE6E3326764B14AFEB8A7E0EC965A95" box="[309,464,737,762]" italics="true" pageId="10" pageNumber="473">Foza raimundi</emphasis>
<bibRefCitation id="BC0342D16764B14AFE5BA7E0EE675A95" author="Reed SK &amp; Cumberlidge N" box="[470,801,737,762]" pageId="10" pageNumber="473" pagination="58 - 66" refId="ref6666" refString="Reed SK, Cumberlidge N. 2006. Foza raimundi, a new genus and species of potamonautid freshwater crab (Crustacea: Decapoda: Potamoidea) from northern Madagascar. Proc Biol Soc Wash. 119 (1): 58 - 66. doi: 10.2988 / 0006 - 324 X (2006) 119 [58: FRANGA] 2.0. CO; 2." type="journal article" year="2006">Reed and Cumberlidge (2006)</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="BC0342D16764B14AFCA1A7E0E9705A95" author="Cumberlidge N &amp; Soma JB &amp; Leever EM &amp; Daniels SR" box="[812,1078,737,762]" pageId="10" pageNumber="473" pagination="584 - 599" refId="ref6294" refString="Cumberlidge N, Soma JB, Leever EM, Daniels SR. 2020. New lineages within the Malagasy freshwater crab fauna: agora n. gen. for Thelphusa goudoti H. Milne Edwards, 1853, and a phytotelmic new species of Malagasya Cumberlidge &amp; Sternberg, 2002 (Brachyura: Potamonautidae: Deckeniidae). J Crust Biol. 40 (5): 584 - 599. doi: 10.1093 / jcbiol / ruaa 050." type="journal article" year="2020">Cumberlidge et al. 2020</bibRefCitation>
, fig. 1).
</paragraph>
<paragraph id="D82D3F206764B14AFF4DA605E94F5C3A" blockId="10.[160,1156,355,1351]" pageId="10" pageNumber="473">
<taxonomicName id="1F9244A36764B14AFF4DA605ECC05B72" authority="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever, 2023" authorityName="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever" authorityYear="2023" box="[192,390,772,797]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
<emphasis id="EAE6E3326764B14AFF4DA605ECC05B72" box="[192,390,772,797]" italics="true" pageId="10" pageNumber="473">Vichai cyanalepou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1D55E496764B14AFE03A605ECA35B72" box="[398,485,772,797]" pageId="10" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
is the fourth genus and fourth species to be recorded from 
<taxonomicName id="1F9244A36764B14AFF2DA627EC475B50" box="[160,257,806,831]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
National Park, the other taxa being 
<emphasis id="EAE6E3326764B14AFD2FA627E9585B50" box="[674,1054,806,831]" italics="true" pageId="10" pageNumber="473">
<taxonomicName id="1F9244A36764B14AFD2FA627E95F5B50" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1872" box="[674,1049,806,831]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="madagascariensis">Hydrothelphusa madagascariensis</taxonomicName>
,
</emphasis>
<taxonomicName id="1F9244A36764B14AFBABA627EF7D5B0D" authority="Cumberlidge et al. (2002)" authorityName="Cumberlidge" authorityYear="2002" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="longimerus">
<emphasis id="EAE6E3326764B14AFBABA627EC5E5B0D" italics="true" pageId="10" pageNumber="473">Marojejy longimerus</emphasis>
Cumberlidge et al. (2002)
</taxonomicName>
and 
<taxonomicName id="1F9244A36764B14AFDFEA648EE495B0D" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[627,783,841,866]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="raimundi">
<emphasis id="EAE6E3326764B14AFDFEA648EE495B0D" box="[627,783,841,866]" italics="true" pageId="10" pageNumber="473">Foza raimundi</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="BC0342D16764B14AFCC7A648ED915BEA" author="Cumberlidge N &amp; Sternberg R" pageId="10" pageNumber="473" pagination="41 - 79" refId="ref6374" refString="Cumberlidge N, Sternberg R. 2002. The freshwater crabs of Madagascar (Crustacea, Decapoda, Potamoidea). Zoosystema. 24: 41 - 79." type="journal article" year="2002">Cumberlidge and Sternberg 2002</bibRefCitation>
; 
<bibRefCitation id="BC0342D16764B14AFF6AA66DEF475BEA" author="Cumberlidge N &amp; Reed SK &amp; Boyko CB" box="[231,513,876,901]" pageId="10" pageNumber="473" pagination="1133 - 1157" refId="ref6190" refString="Cumberlidge N, Reed SK, Boyko CB. 2004. Distribution patterns of the Malagasy freshwater crabs (Crustacea: Decapoda: Brachyura). J Nat Hist. 38: 1133 - 1157. doi: 10.1080 / 0022293031000068419." type="journal article" year="2004">Cumberlidge et al. 2004</bibRefCitation>
; 
<bibRefCitation id="BC0342D16764B14AFD9CA66DEE185BEA" author="Reed SK &amp; Cumberlidge N" box="[529,862,876,901]" pageId="10" pageNumber="473" pagination="58 - 66" refId="ref6666" refString="Reed SK, Cumberlidge N. 2006. Foza raimundi, a new genus and species of potamonautid freshwater crab (Crustacea: Decapoda: Potamoidea) from northern Madagascar. Proc Biol Soc Wash. 119 (1): 58 - 66. doi: 10.2988 / 0006 - 324 X (2006) 119 [58: FRANGA] 2.0. CO; 2." type="journal article" year="2006">Reed and Cumberlidge 2006</bibRefCitation>
). Two of these four taxa (
<taxonomicName id="1F9244A36764B14AFF25A68FECC05BC8" box="[168,390,910,935]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="longimerus">
<emphasis id="EAE6E3326764B14AFF25A68FECC05BC8" box="[168,390,910,935]" italics="true" pageId="10" pageNumber="473">Marojejy longimerus</emphasis>
</taxonomicName>
and 
<taxonomicName id="1F9244A36764B14AFE48A68FEFC85BC8" authority="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever, 2023" authorityName="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever" authorityYear="2023" box="[453,654,910,935]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
<emphasis id="EAE6E3326764B14AFE48A68FEFC85BC8" box="[453,654,910,935]" italics="true" pageId="10" pageNumber="473">Vichai cyanalepou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1D55E496764B14AFD15A68FEFB45BC8" box="[664,754,910,935]" pageId="10" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
) are endemic to 
<taxonomicName id="1F9244A36764B14AFC36A68FE95A5BC8" box="[955,1052,910,935]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
National Park. Earlier reports of 
<taxonomicName id="1F9244A36764B14AFE23A6B0EEFF5BA5" authority="Ng and Takeda (1994)" authorityName="Ng and Takeda" authorityYear="1994" box="[430,953,945,970]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="eumeces">
<emphasis id="EAE6E3326764B14AFE23A6B0EFEF5BA5" box="[430,681,945,970]" italics="true" pageId="10" pageNumber="473">Skelosophusa eumeces</emphasis>
<bibRefCitation id="BC0342D16764B14AFD38A6B0EEFF5BA5" author="Ng PKL &amp; Takeda M" box="[693,953,945,970]" pageId="10" pageNumber="473" pagination="161 - 172" refId="ref6616" refString="Ng PKL, Takeda M. 1994. Skelosophusa (Crustacea, Decapoda, Brachyura), a new genus of potamonautid freshwater crab from Madagascar, with descriptions of two new species. Bull Nat Sci Mus Ser A (Zool). 20: 161 - 172." type="journal article" year="1994">Ng and Takeda (1994)</bibRefCitation>
</taxonomicName>
, occurring in the 
<taxonomicName id="1F9244A36764B14AFF2DA6D5EC475B82" box="[160,257,980,1005]" class="Malacostraca" family="Potamonautidae" genus="Marojejy" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="genus">Marojejy</taxonomicName>
National Park by 
<bibRefCitation id="BC0342D16764B14AFE46A6D5EFAC5B82" author="Cumberlidge N &amp; Reed SK &amp; Boyko CB" box="[459,746,980,1005]" pageId="10" pageNumber="473" pagination="1133 - 1157" refId="ref6190" refString="Cumberlidge N, Reed SK, Boyko CB. 2004. Distribution patterns of the Malagasy freshwater crabs (Crustacea: Decapoda: Brachyura). J Nat Hist. 38: 1133 - 1157. doi: 10.1080 / 0022293031000068419." type="journal article" year="2004">Cumberlidge et al. (2004)</bibRefCitation>
were misidentifications and are not supported by the present study, as those specimens (FMNH 7579, FMNH 7590, and FMNH 7592) are reidentified here as belonging to 
<taxonomicName id="1F9244A36764B14AFD28A118EE365C5D" authority="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever, 2023" authorityName="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever" authorityYear="2023" box="[677,880,1049,1074]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
<emphasis id="EAE6E3326764B14AFD28A118EE365C5D" box="[677,880,1049,1074]" italics="true" pageId="10" pageNumber="473">Vichai cyanalepou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1D55E496764B14AFCF6A118EE905C5D" box="[891,982,1049,1074]" pageId="10" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
See above for comparisons of 
<taxonomicName id="1F9244A36764B14AFED9A13DECAA5C3A" authority="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever, 2023" authorityName="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever" authorityYear="2023" box="[340,492,1084,1109]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
<emphasis id="EAE6E3326764B14AFED9A13DECAA5C3A" box="[340,492,1084,1109]" italics="true" pageId="10" pageNumber="473">V. cyanalepou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1D55E496764B14AFE79A13DEF0D5C3A" box="[500,587,1084,1109]" pageId="10" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
with 
<taxonomicName id="1F9244A36764B14AFD06A13DEE585C3A" authorityName="Ng and Takeda" authorityYear="1994" box="[651,798,1084,1109]" class="Malacostraca" family="Potamonautidae" genus="Skelosophusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="genus">
<emphasis id="EAE6E3326764B14AFD06A13DEE585C3A" box="[651,798,1084,1109]" italics="true" pageId="10" pageNumber="473">Skelosophusa</emphasis>
</taxonomicName>
and 
<taxonomicName id="1F9244A36764B14AFCDAA13DE9455C3A" authorityName="A. Milne-Edwards" authorityYear="1872" box="[855,1027,1084,1109]" class="Malacostraca" family="Potamonautidae" genus="Hydrothelphusa" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="genus">
<emphasis id="EAE6E3326764B14AFCDAA13DE9455C3A" box="[855,1027,1084,1109]" italics="true" pageId="10" pageNumber="473">Hydrothelphusa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D82D3F206764B14AFF4DA15FECAF5D28" blockId="10.[160,1156,355,1351]" pageId="10" pageNumber="473">
<taxonomicName id="1F9244A36764B14AFF4DA15FECD65C18" authority="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever, 2023" authorityName="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever" authorityYear="2023" box="[192,400,1118,1143]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
<emphasis id="EAE6E3326764B14AFF4DA15FECD65C18" box="[192,400,1118,1143]" italics="true" pageId="10" pageNumber="473">Vichai cyanalepou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1D55E496764B14AFE11A15FECBC5C18" box="[412,506,1118,1143]" pageId="10" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from 
<taxonomicName id="1F9244A36764B14AFCC0A15FEE965C18" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[845,976,1118,1143]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="raimundi">
<emphasis id="EAE6E3326764B14AFCC0A15FEE965C18" box="[845,976,1118,1143]" italics="true" pageId="10" pageNumber="473">F. raimundi</emphasis>
</taxonomicName>
mainly by the G1TA which is glabrous on the dorsal side in 
<taxonomicName id="1F9244A36764B14AFD3FA180EE095CF5" authority="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever, 2023" authorityName="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever" authorityYear="2023" box="[690,847,1153,1178]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
<emphasis id="EAE6E3326764B14AFD3FA180EE095CF5" box="[690,847,1153,1178]" italics="true" pageId="10" pageNumber="473">V. cyanalepou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1D55E496764B14AFCD5A180EEF45CF5" box="[856,946,1153,1178]" pageId="10" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="40A923A56764B14AFC4CA180E9645CF5" box="[961,1058,1153,1178]" captionStart="Figure 3" captionStartId="6.[160,225,1101,1124]" captionTargetBox="[163,1153,142,1069]" captionTargetId="figure-190@6.[163,1153,142,1069]" captionTargetPageId="6" captionText="Figure 3. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) right G1, ventral view; (b) right G1, dorsal view; (c) right G1, superior view; (d) right G2, ventral view. Scale bar: a, b = 1.4 mm; c = 1.1 mm; d = 1.7 mm." figureDoi="http://doi.org/10.5281/zenodo.7859827" httpUri="https://zenodo.org/record/7859827/files/figure.png" pageId="10" pageNumber="473">Figure 3</figureCitation>
(b)), vs a G1TA that is heavily setose on the dorsal side in 
<taxonomicName id="1F9244A36764B14AFD51A1A5EE1A5CD2" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[732,860,1188,1213]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="raimundi">
<emphasis id="EAE6E3326764B14AFD51A1A5EE1A5CD2" box="[732,860,1188,1213]" italics="true" pageId="10" pageNumber="473">F. raimundi</emphasis>
</taxonomicName>
(
<bibRefCitation id="BC0342D16764B14AFCE6A1A5E97D5CD2" author="Leever EM &amp; Daniels SR &amp; Soma JB &amp; Cumberlidge N" box="[875,1083,1188,1213]" pageId="10" pageNumber="473" pagination="241 - 263" refId="ref6477" refString="Leever EM, Daniels SR, Soma JB, Cumberlidge N. 2022. Two new genera and a new species of freshwater crabs from northern Madagascar: vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae). J Nat Hist. 56 (1 - 4): 241 - 263. doi: 10.1080 / 00222933.2022.2049389." type="journal article" year="2022">Leever et al. 2022</bibRefCitation>
, fig. 7 (h)). Furthermore, a lack of setae on the superior part of the pterygostomial region and on the anterior subpleonal cavity in 
<taxonomicName id="1F9244A36764B14AFD1DA1E8EE765D6D" authority="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever, 2023" authorityName="Cumberlidge &amp; Daniels &amp; Soma &amp; Leever" authorityYear="2023" box="[656,816,1257,1282]" class="Malacostraca" family="Potamonautidae" genus="Vichai" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="cyanalepou" status="sp. nov.">
<emphasis id="EAE6E3326764B14AFD1DA1E8EE765D6D" box="[656,816,1257,1282]" italics="true" pageId="10" pageNumber="473">V. cyanalepou</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F1D55E496764B14AFCB6A1E8EEDE5D6D" box="[827,920,1257,1282]" pageId="10" pageNumber="473" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="40A923A56764B14AFC27A1E8E94A5D6D" box="[938,1036,1257,1282]" captionStart="Figure 2" captionStartId="4.[160,225,1736,1759]" captionTargetBox="[163,1153,142,1704]" captionTargetId="figure-3@4.[163,1153,142,1704]" captionTargetPageId="4" captionText="Figure 2. Vichai cyanalepou sp. nov., holotype, adult J CW 38.5 mm, FMNH 7589, from Marojejy National Park, Sava region, northern Madagascar: (a) frontal view of carapace; (b) frontal view of right chela; (c) frontal view of left chela; (d) right cheliped merus, underside; (e) right cheliped merus, dorsal view; (f) right cheliped carpus, dorsal view; (g) third maxilliped, frontal view; (h) third maxilliped, posterior view; (i) mandible frontal view. Scale bar: a = 5.3 mm; b, c = 8.0 mm; d = 5.3 mm; e, f = 6.7 mm; g, h = 3.3 mm; i = 2.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7859825" httpUri="https://zenodo.org/record/7859825/files/figure.png" pageId="10" pageNumber="473">Figure 2</figureCitation>
(a)) distinguishes it from 
<taxonomicName id="1F9244A36764B14AFEEEA00DECA15D4A" authorityName="Reed and Cumberlidge" authorityYear="2006" box="[355,487,1292,1317]" class="Malacostraca" family="Potamonautidae" genus="Foza" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="raimundi">
<emphasis id="EAE6E3326764B14AFEEEA00DECA15D4A" box="[355,487,1292,1317]" italics="true" pageId="10" pageNumber="473">F. raimundi</emphasis>
</taxonomicName>
where both of these regions are distinctly setose (see 
<bibRefCitation id="BC0342D16764B14AFF2DA02FEC355D28" author="Leever EM &amp; Daniels SR &amp; Soma JB &amp; Cumberlidge N" box="[160,371,1326,1351]" pageId="10" pageNumber="473" pagination="241 - 263" refId="ref6477" refString="Leever EM, Daniels SR, Soma JB, Cumberlidge N. 2022. Two new genera and a new species of freshwater crabs from northern Madagascar: vahatra gen. nov. for Foza ambohitra Cumberlidge and Meyer, 2009, and Toamasina gen. nov. for Toamasina clarki sp. nov. (Brachyura: Potamoidea: Deckeniidae). J Nat Hist. 56 (1 - 4): 241 - 263. doi: 10.1080 / 00222933.2022.2049389." type="journal article" year="2022">Leever et al. 2022</bibRefCitation>
, fig. 7(c)).
</paragraph>
</subSubSection>
</treatment>
</document>