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<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.97.115682" ID-Pensoft-Pub="2747-8106-97-53" ID-Pensoft-UUID="4A3318772B185D35B1E7EC9578EF4A47" ID-ZooBank="933EC356F21C45AF9CFA563E64D27953" ModsDocID="2747-8106-97-53" checkinTime="1702462880200" checkinUser="pensoft" docAuthor="Neubauer, Thomas A. &amp; Wesselingh, Frank P." docDate="2023" docId="4A27F5D770E05EE4B31764A66CBEDF82" docLanguage="en" docName="Zitteliana 97: 53-88" docOrigin="Zitteliana 97" docPubDate="2023-12-12" docSource="http://dx.doi.org/10.3897/zitteliana.97.115682" docTitle="Corymbina elegans" docType="treatment" docVersion="1" id="4A3318772B185D35B1E7EC9578EF4A47" lastPageNumber="53" masterDocId="4A3318772B185D35B1E7EC9578EF4A47" masterDocTitle="The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms" masterLastPageNumber="88" masterPageNumber="53" pageNumber="53" updateTime="1702462880200" updateUser="pensoft">
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<mods:title id="F848AD7125D01A162B4CE23FDD7660AA">The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms</mods:title>
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<mods:namePart id="2C147D83B23CDCF5997FAFCE0D813EB2">Neubauer, Thomas A.</mods:namePart>
<mods:nameIdentifier id="299047608C073099105D4A1C6676C6C8" type="ORCID">https://orcid.org/0000-0002-1398-9941</mods:nameIdentifier>
<mods:affiliation id="D11C0753B7D592146393359CA928CCEC">SNSB - Bavarian State Collection for Palaeontology and Geology, Richard-Wagner-Strasse 10, 80333 Munich, Germany &amp; Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
<mods:nameIdentifier id="8967A06AE0500411C169091E962BC594" type="email">neubauer@snsb.de</mods:nameIdentifier>
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<mods:namePart id="5B23DADE478B11E3281BD4536E4133F9">Wesselingh, Frank P.</mods:namePart>
<mods:nameIdentifier id="4B735685D1205895447EB9EF1C01B7B6" type="ORCID">https://orcid.org/0000-0003-3655-0701</mods:nameIdentifier>
<mods:affiliation id="DF42022B50E80D5B64D5AE64BCF9AFCA">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands &amp; Department of Earth Sciences, Utrecht University, Princetonlaan 8 a, 3584 CB Utrecht, Netherlands</mods:affiliation>
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<mods:title id="9D327DFEB82FE85100B01970A79F1D41">Zitteliana</mods:title>
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<mods:date id="C19F42473073C83902FD3042F072948A">2023</mods:date>
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<mods:number id="2C4DBAD33125915718168124883B0B64">97</mods:number>
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<treatment id="4A27F5D770E05EE4B31764A66CBEDF82" LSID="urn:lsid:plazi:treatment:4A27F5D770E05EE4B31764A66CBEDF82" httpUri="http://treatment.plazi.org/id/4A27F5D770E05EE4B31764A66CBEDF82" lastPageNumber="53" pageId="0" pageNumber="53">
<subSubSection id="A3C1009CB026F812B8F7280BE2243FEB" pageId="0" pageNumber="53" type="nomenclature">
<paragraph id="5912BC9448B2D9839C6BE87623BDECB1" pageId="0" pageNumber="53">
<taxonomicName id="D3BEFDCA59460830A1EC160970E96D47" ID-CoL="YQDD" LSID="4A27F5D7-70E0-5EE4-B317-64A66CBEDF82" authority="(Cantraine, 1841)" baseAuthorityName="Cantraine" baseAuthorityYear="1841" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina elegans" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="elegans">Corymbina elegans (Cantraine, 1841)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="6460D9A65B64183C17B95BD4CC99CE96" pageId="0" pageNumber="53" type="description">
<paragraph id="C3B76471844558B34BE724309B5D7E7B" pageId="0" pageNumber="53">
<figureCitation id="CEE0431C15EFB0D9F0631D0DB55BDED9" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Lymnaeidae of the Kolankaya I fauna. A-C. Corymbina elegans (Cantraine, 1841), RGM 962696, sample 2. D-F. Radix sp., RGM 1310798, sample 2. Scale bars: 1 mm." figureDoi="10.3897/zitteliana.97.115682.figure16" httpUri="https://binary.pensoft.net/fig/950753" pageId="0" pageNumber="53">Fig. 16A-C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F4FE5B72F75F2B68F8CC2563D7A91336" pageId="0" pageNumber="53" type="reference_group">
<paragraph id="7A5A638751BF9D21CADD0B4334A0DF64" pageId="0" pageNumber="53">
<taxonomicName id="27A59D97962A13C9B0DBDFE1E23237FE" authorityName="Bukowski" authorityYear="1892" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">*1841 Ad. Corymbina</taxonomicName>
[
<taxonomicName id="D77181614DCC63DAFAF8308E0723E262" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina elina" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="elina">
<emphasis id="5B6D9749013E530FF25487582E34BCFD" italics="true" pageId="0" pageNumber="53">Corymbina elina</emphasis>
</taxonomicName>
]
<taxonomicName id="D978DB4CEEBDD62E20F9AA69B4DF492B" baseAuthorityName="Cantraine" baseAuthorityYear="1841" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina elegans" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="elegans">Corymbina elegans</taxonomicName>
. Nob. - Cantraine: 156, pl. 5, fig. 12, 12a.
</paragraph>
<paragraph id="ADED7557E65582DF4CADEFB5832A39FC" pageId="0" pageNumber="53">
<taxonomicName id="438AC527B91BB03616B7FEBCA86DE341" class="Gastropoda" family="Lymnaeidae" genus="Limneus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limneus" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">1847 Limneus</taxonomicName>
Adelina - Spratt and Forbes: 177, textfig. a.
</paragraph>
<paragraph id="4F7A33E90BB0668F10AF9B8D9FB03C7E" pageId="0" pageNumber="53">
<taxonomicName id="03431BC305144A113E75E5E86FD90C00" class="Gastropoda" family="Lymnaeidae" genus="Lymnaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lymnaeus" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">? 1877 Lymnaeus</taxonomicName>
<taxonomicName id="86A428A50F40A434CC6AF5F2ED74F727" authorityName="Bruand" authorityYear="1851" family="Lymnaeidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Hygrophila" pageId="0" pageNumber="53" rank="subFamily" subFamily="Adelinae">Adelinae</taxonomicName>
[sic] Forbes [sic] - Fuchs: 5, pl. 1, fig. 4.
</paragraph>
<paragraph id="5E967A8A074B84CF9AF1D1C24052EBAB" pageId="0" pageNumber="53">
<taxonomicName id="396C3B50FAB75F1F0B0C8FEB4D9BE79E" class="Gastropoda" family="Lymnaeidae" genus="Lymnaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lymnaeus" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">? 1877 Lymnaeus</taxonomicName>
<taxonomicName id="E435BD98F0B94D418F136AB2C97723D0" authorityName="Bruand" authorityYear="1851" family="Lymnaeidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Hygrophila" pageId="0" pageNumber="53" rank="subFamily" subFamily="Adelinae">Adelinae</taxonomicName>
[sic] Cantr. - Fuchs: 37, pl. 4, figs 1-6.
</paragraph>
<paragraph id="84F27F9D46499D4F4B0C13768B368F38" pageId="0" pageNumber="53">
<taxonomicName id="63A6D5CD40A56DEAAC4DD11BEA816434" baseAuthorityName="Cantraine" baseAuthorityYear="1841" class="Gastropoda" family="Lymnaeidae" genus="Radix" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Radix (Adelinella) elegans" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="elegans" subGenus="Adelinella">1923 Radix (Adelinella) elegans</taxonomicName>
(Cantraine) - Wenz: 1319-1320.
</paragraph>
<paragraph id="393659D7EA0D8943140056DFE0A7034F" pageId="0" pageNumber="53">
<taxonomicName id="AA2833476C19913A45AC7190639806DF" baseAuthorityName="Cantraine" baseAuthorityYear="1841" class="Gastropoda" family="Lymnaeidae" genus="Radix" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Radix (Adelinella) elegans" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="elegans" subGenus="Adelinella">1973 Radix (Adelinella) elegans</taxonomicName>
(Cantraine) -
<normalizedToken id="12A2A9981CBF5BA5079AECD152449D10" originalValue="Schütt">Schuett</normalizedToken>
and Besenecker: 16, pl. 1, figs 19, 20.
</paragraph>
<paragraph id="A7C613DEE4DB460F3119A8FB8B464EFE" pageId="0" pageNumber="53">
<taxonomicName id="D3B101106FEAB9983136BF587EDAEDA2" baseAuthorityName="Cantraine" baseAuthorityYear="1841" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina elegans" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="elegans">1981 Corymbina elegans</taxonomicName>
(Cantraine) - Willmann: pl. 13, figs 18-20.
</paragraph>
<paragraph id="0407206597EDB633C8EC971702248B34" pageId="0" pageNumber="53">
<taxonomicName id="9E01E1C90613C44D4DD1402532BB3152" baseAuthorityName="Cantraine" baseAuthorityYear="1841" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina elegans" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="elegans">? 2022 Corymbina elegans</taxonomicName>
(Cantraine, 1841) - Neubauer: fig. 6.7k, l.
</paragraph>
</subSubSection>
<subSubSection id="4DF64CACC8EDCE7BF5D130C1C99C3EFD" pageId="0" pageNumber="53" type="materials_examined">
<paragraph id="4D305E2AA2D3D486BC1A4F7E40DBB2E3" pageId="0" pageNumber="53">Material.</paragraph>
<paragraph id="4439113DD63BAC1233ADCF91C78433EC" pageId="0" pageNumber="53">
<specimenCount id="8EEA9DD3F43A79F69F8DE4AA1B7052B4" type="juvenile">1 juvenile</specimenCount>
specimen (RGM 962696) from sample 2.
</paragraph>
</subSubSection>
<subSubSection id="64B92EAA633B3A3E77737977E12656C7" pageId="0" pageNumber="53" type="type locality">
<paragraph id="54A6DA39302C8925A42BE253FD2E62EF" pageId="0" pageNumber="53">Type locality.</paragraph>
<paragraph id="EDEB832412793F096933E4FD263B0BF1" pageId="0" pageNumber="53">
Italy; no further details known-Cantraine wrote: &quot;Je ne connais cette
<normalizedToken id="F8823A4247C52DE0161CBA27AB8ED34D" originalValue="espèce">espece</normalizedToken>
<normalizedToken id="B59A66CA7B7CFCCC09616365F0FBA6B0" originalValue="qu’à">qu'a</normalizedToken>
<normalizedToken id="FDB8B5FC82A249AA88E4C48BDED08EA3" originalValue="l’état">l'etat</normalizedToken>
fossile, le seul individu que je
<normalizedToken id="4BBFEDEB90628162E018E2954FC3C326" originalValue="possède">possede</normalizedToken>
<normalizedToken id="4D4032D2D59234B97AB61FA259D77F8C" originalValue="m’a">m'a</normalizedToken>
<normalizedToken id="16CAB9C27EF009C39010233568B1D7CC" originalValue="été">ete</normalizedToken>
<normalizedToken id="5654C3B0128AE9574F0C1B7D71A3E5DC" originalValue="donnée">donnee</normalizedToken>
par mon ami M. Rossi de Livourne, qui
<normalizedToken id="0C26CB319D2BB81AFEB7550D96F8B6C7" originalValue="a’en">a'en</normalizedToken>
connaissait pas exactement la provenarco.&quot; [I only know this species in the fossil state, the only individual I have was given to me by my friend Mr. Rossi from Livorno, who
<normalizedToken id="1FEC6749EA3C71AF2FF2D32F74A75036" originalValue="didn’t">didn't</normalizedToken>
know exactly where it came from.].
</paragraph>
</subSubSection>
<subSubSection id="2AE2EB6C89DDAA7965BF9B96036D5CCC" pageId="0" pageNumber="53" type="description">
<paragraph id="D22C726BB90DC5327087391C3B426F1D" pageId="0" pageNumber="53">Description.</paragraph>
<paragraph id="A55C4737D9DD6180ED11DEACEED13E28" pageId="0" pageNumber="53">Shell comprises protoconch and about 0.75 teleoconch whorls. Initial part of protoconch smooth, followed by quarter whorl bearing faint riblets. P/T boundary not clearly visible, but probably coincides with weak angulation and onset of growth lines. Soon distinct, sharp ribs develop, which soon become broader and bulgier, leading to an irregular, wavy surface. Aperture fragmented, parts of outer lip missing; originally apparently elliptical. Umbilicus narrow.</paragraph>
</subSubSection>
<subSubSection id="EF6567A546679056BF218C6E535B23F6" pageId="0" pageNumber="53" type="remarks">
<paragraph id="50D758FE33C5307A42029A4A678B550D" pageId="0" pageNumber="53">Remarks.</paragraph>
<paragraph id="915257BA665B815BE6A4B8E960F4FD8A" pageId="0" pageNumber="53">
<bibRefCitation id="0EBD53D37ACA8686C0C557B1B57B7894" author="Spratt, TAB" journalOrPublisher="Zeitschrift fuer wissenschaftliche Zoologie" pageId="0" pageNumber="53" publicationUrl="http://reader.digitale-sammlungen.de/de/fs1/object/display/bsb10468250_00009.html" refId="B146" refString="Spratt, TAB, Forbes, E, 1847. Travels in Lycia, Milyas, and the Cibyratis, in company with the late Rev. E. T. Daniell. 2 volumes. John van Voorst, Paternoster Row, London. [Volume 1:] http://reader.digitale-sammlungen.de/de/fs1/object/display/bsb10468250_00009.html" title="Travels in Lycia, Milyas, and the Cibyratis, in company with the late Rev. E. T. Daniell. 2 volumes. John van Voorst, Paternoster Row, London. [Volume 1:]" url="http://reader.digitale-sammlungen.de/de/fs1/object/display/bsb10468250_00009.html" year="1847">Spratt and Forbes (1847)</bibRefCitation>
mentioned the name
<taxonomicName id="434FDE701238D48E8C0227613576E3B9" class="Gastropoda" family="Lymnaeidae" genus="Limneus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Limneus" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="1C236F82DA0802CE58A670A493DC796F" italics="true" pageId="0" pageNumber="53">Limneus</emphasis>
</taxonomicName>
[sic]
<taxonomicName id="78A04BDD411636670C35CED6A6D09042" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lymnaea adelina" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="adelina">
<emphasis id="28A5E56DF0ECFC1A1D185444ECC4A44B" italics="true" pageId="0" pageNumber="53">Lymnaea adelina</emphasis>
</taxonomicName>
along with an illustration, making the name available. However, they clearly referred it to be the same species as
<emphasis id="3702796BD2CDE72403DE3E3D90F0E9D4" italics="true" pageId="0" pageNumber="53">
Adelina
<taxonomicName id="E766363F1A3E1F1077AF90D4D1FD243A" class="Gastropoda" family="Dreissenidae" genus="Prososthenia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prososthenia elegans" order="Cardiida" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="elegans">Prososthenia elegans</taxonomicName>
</emphasis>
Cantraine, 1841, rendering
<taxonomicName id="E2261CA61ED2CC13F118E837E22BC489" class="Gastropoda" family="Lymnaeidae" genus="Lymnaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lymnaea adelina" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="adelina">
<emphasis id="81AB6910FBF892B9366E5156A7FAFD66" italics="true" pageId="0" pageNumber="53">Lymnaea adelina</emphasis>
</taxonomicName>
a junior objective synonym of
<taxonomicName id="C7D53E972F6D88785C596A94186B5852" baseAuthorityName="Cantraine" baseAuthorityYear="1841" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina elegans" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="elegans">
<emphasis id="A13E7E45965FAE3B36156B25379CDD2E" italics="true" pageId="0" pageNumber="53">Corymbina elegans</emphasis>
</taxonomicName>
. The specimen they illustrated has a much larger aperture, but the species has an extremely variable shell (see also
<bibRefCitation id="40F3597420C9447A0997BC04085AD88A" author="Willmann, R" journalOrPublisher="A" pageId="0" pageNumber="53" pagination="10 - 235" refId="B179" refString="Willmann, R, 1981. Evolution, Systematik und stratigraphische Bedeutung der neogenen Suesswassergastropoden von Rhodos und Kos/Aegaeis. Palaeontographica Abt. A 174: 10 - 235" title="Evolution, Systematik und stratigraphische Bedeutung der neogenen Suesswassergastropoden von Rhodos und Kos / Aegaeis. Palaeontographica Abt." volume="174" year="1981">Willmann 1981</bibRefCitation>
).
</paragraph>
<paragraph id="3FC8BB30FD9FD46665F12EAD84650C20" pageId="0" pageNumber="53">
<taxonomicName id="F71D683DA7EE7126B86D4709D9C7FBE5" baseAuthorityName="T.Fuchs" baseAuthorityYear="1877" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina bicarinata" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="bicarinata">
<emphasis id="7A5F7D14A086FB1E45BEF4F8279F60A3" italics="true" pageId="0" pageNumber="53">Corymbina bicarinata</emphasis>
</taxonomicName>
(Fuchs, 1877) from the Early Pleistocene of Livanates (Phthiotis, Greece) has a
<taxonomicName id="921503B11CF632638699B3ED6BB5B93F" authorityName="Montfort" authorityYear="1810" class="Gastropoda" family="Lymnaeidae" genus="Radix" kingdom="Animalia" lsidName="Radix" order="Cardiida" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="ECBB7BE7A1BD771901C5812134E548AF" italics="true" pageId="0" pageNumber="53">Radix</emphasis>
</taxonomicName>
-like appearance, with broad ovoid, attached aperture, an angulated last whorl, slender ribs, and small, pointy spire (Neubauer 2023b). Specimens from the same deposits identified by
<bibRefCitation id="D32B8A41F2D7EC60148A13325DBE5133" author="Fuchs, T" journalOrPublisher="Denkschriften der mathematisch-naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften" pageId="0" pageNumber="53" pagination="1 - 42" publicationUrl="https://www.biodiversitylibrary.org/page/7221662" refId="B57" refString="Fuchs, T, 1877. Studien ueber die juengeren Tertiaerbildungen Griechenlands. Denkschriften der mathematisch-naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften 37: 1 - 42, https://www.biodiversitylibrary.org/page/7221662" title="Studien ueber die juengeren Tertiaerbildungen Griechenlands." url="https://www.biodiversitylibrary.org/page/7221662" volume="37" year="1877">Fuchs (1877)</bibRefCitation>
as
<taxonomicName id="EF8D5C55F14C2A1A9A46EABBD1D9595B" lsidName="L. adelinae" pageId="0" pageNumber="53" rank="species" species="adelinae">
<emphasis id="A2282FB3878473817D77A86D300541DE" italics="true" pageId="0" pageNumber="53">L. adelinae</emphasis>
</taxonomicName>
[=
<taxonomicName id="CEBBC91FBBA466354DB7EDB05298A113" lsidName="C. elegans" pageId="0" pageNumber="53" rank="species" species="elegans">
<emphasis id="D5D175DE77B50F4550FD51DBC1E419E0" italics="true" pageId="0" pageNumber="53">C. elegans</emphasis>
</taxonomicName>
] differ considerably from the phenotypes shown by
<bibRefCitation id="8DB40ED78CBA963B80785BD514C0BFA3" author="Cantraine, F" journalOrPublisher="Nouveaux Memoires de l'Academie Royale des Sciences et Belles-Lettres de Bruxelles" pageId="0" pageNumber="53" pagination="1 - 173" publicationUrl="http://www.digizeitschriften.de/dms/img/?PPN=PPN129323640_0013&amp;DMDID=dmdlog24" refId="B34" refString="Cantraine, F, 1841. Malacologie mediterraneenne et littorale, ou description des mollusques qui vivent dans la Mediterranee ou sur le continent de l'Italie, ainsi que des coquilles qui se trouvent dans les terrains tertiaires italiens, avec des observations sur leur anatomie, leurs moeurs, leur analogie et leur gisement. Ouvrage servant de faune malacologique italienne et de complement a la Conchiologia fossile subapennina de Brocchi. Nouveaux Memoires de l'Academie Royale des Sciences et Belles-Lettres de Bruxelles 13: 1 - 173, http://www.digizeitschriften.de/dms/img/?PPN=PPN129323640_0013&amp;DMDID=dmdlog24" title="Malacologie mediterraneenne et littorale, ou description des mollusques qui vivent dans la Mediterranee ou sur le continent de l'Italie, ainsi que des coquilles qui se trouvent dans les terrains tertiaires italiens, avec des observations sur leur anatomie, leurs moeurs, leur analogie et leur gisement. Ouvrage servant de faune malacologique italienne et de complement a la Conchiologia fossile subapennina de Brocchi." url="http://www.digizeitschriften.de/dms/img/?PPN=PPN129323640_0013&amp;DMDID=dmdlog24" volume="13" year="1841">Cantraine (1841)</bibRefCitation>
,
<bibRefCitation id="E1062E50DE06515215F277F6261602DB" author="Spratt, TAB" journalOrPublisher="Zeitschrift fuer wissenschaftliche Zoologie" pageId="0" pageNumber="53" publicationUrl="http://reader.digitale-sammlungen.de/de/fs1/object/display/bsb10468250_00009.html" refId="B146" refString="Spratt, TAB, Forbes, E, 1847. Travels in Lycia, Milyas, and the Cibyratis, in company with the late Rev. E. T. Daniell. 2 volumes. John van Voorst, Paternoster Row, London. [Volume 1:] http://reader.digitale-sammlungen.de/de/fs1/object/display/bsb10468250_00009.html" title="Travels in Lycia, Milyas, and the Cibyratis, in company with the late Rev. E. T. Daniell. 2 volumes. John van Voorst, Paternoster Row, London. [Volume 1:]" url="http://reader.digitale-sammlungen.de/de/fs1/object/display/bsb10468250_00009.html" year="1847">Spratt and Forbes (1847)</bibRefCitation>
, and
<bibRefCitation id="C54C5ED31DD04420CB467337C2F1E218" author="Willmann, R" journalOrPublisher="A" pageId="0" pageNumber="53" pagination="10 - 235" refId="B179" refString="Willmann, R, 1981. Evolution, Systematik und stratigraphische Bedeutung der neogenen Suesswassergastropoden von Rhodos und Kos/Aegaeis. Palaeontographica Abt. A 174: 10 - 235" title="Evolution, Systematik und stratigraphische Bedeutung der neogenen Suesswassergastropoden von Rhodos und Kos / Aegaeis. Palaeontographica Abt." volume="174" year="1981">Willmann (1981)</bibRefCitation>
. They rather resemble
<taxonomicName id="8FFDF172FB75E1C32CFE92A02EB6CC17" lsidName="C. bicarinata" pageId="0" pageNumber="53" rank="species" species="bicarinata">
<emphasis id="D3F27E0CD60F85200E637CFC40D98ABA" italics="true" pageId="0" pageNumber="53">C. bicarinata</emphasis>
</taxonomicName>
, yet with a keel at the angulation. More data on the variability of both species is needed to disentangle their taxonomy and synonymies. We tentatively associate our single juvenile specimen with
<taxonomicName id="0D5FAF33762FB26A4A8FC7CF594F93EC" lsidName="C. elegans" pageId="0" pageNumber="53" rank="species" species="elegans">
<emphasis id="BA62BA896A05AA7D29E7461B24B1182D" italics="true" pageId="0" pageNumber="53">C. elegans</emphasis>
</taxonomicName>
given the general match in shell shape and sculpture as well as the known stratigraphic and geographic distribution of the species.
</paragraph>
<paragraph id="6F0DF037E7238604AB3177B34E52A623" pageId="0" pageNumber="53">
The Late Miocene
<taxonomicName id="472F4C1E4F018148E489B8012D7912FE" authorityName="Bukowski" authorityYear="1892" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina rhodensis" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="rhodensis">
<emphasis id="806F69A04C7A8288DB138AB716094255" italics="true" pageId="0" pageNumber="53">Corymbina rhodensis</emphasis>
</taxonomicName>
Bukowski, 1892 is smaller and has a more slender shell, often with reduced sculpture (
<bibRefCitation id="41FED898C82551E62E5401DD1FDF7CCD" author="Willmann, R" journalOrPublisher="A" pageId="0" pageNumber="53" pagination="10 - 235" refId="B179" refString="Willmann, R, 1981. Evolution, Systematik und stratigraphische Bedeutung der neogenen Suesswassergastropoden von Rhodos und Kos/Aegaeis. Palaeontographica Abt. A 174: 10 - 235" title="Evolution, Systematik und stratigraphische Bedeutung der neogenen Suesswassergastropoden von Rhodos und Kos / Aegaeis. Palaeontographica Abt." volume="174" year="1981">Willmann 1981</bibRefCitation>
). In both
<taxonomicName id="482207CD75A9F1924C65EC6E78FD05B9" lsidName="C. elegans" pageId="0" pageNumber="53" rank="species" species="elegans">
<emphasis id="DFEE8131FA09B2F4BBFF054070307F80" italics="true" pageId="0" pageNumber="53">C. elegans</emphasis>
</taxonomicName>
and
<taxonomicName id="9906BC2575C4B6DC2F9032BFB5B99341" lsidName="C. rhodensis" pageId="0" pageNumber="53" rank="species" species="rhodensis">
<emphasis id="D9CB133E58D397719B7BC8FECE5A4663" italics="true" pageId="0" pageNumber="53">C. rhodensis</emphasis>
</taxonomicName>
specimens occur with the last whorl partly or entirely uncoiled (
<bibRefCitation id="BCFB7C3FF8C73D397A427F99573711C8" author="Willmann, R" journalOrPublisher="A" pageId="0" pageNumber="53" pagination="10 - 235" refId="B179" refString="Willmann, R, 1981. Evolution, Systematik und stratigraphische Bedeutung der neogenen Suesswassergastropoden von Rhodos und Kos/Aegaeis. Palaeontographica Abt. A 174: 10 - 235" title="Evolution, Systematik und stratigraphische Bedeutung der neogenen Suesswassergastropoden von Rhodos und Kos / Aegaeis. Palaeontographica Abt." volume="174" year="1981">Willmann 1981</bibRefCitation>
).
<taxonomicName id="6888DDB28EDF639B0937D97D0A0FE21C" baseAuthorityName="Oppenheim" baseAuthorityYear="1919" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina aegaea" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="aegaea">
<emphasis id="2104E0CBA61FF952041D135C037CA1DB" italics="true" pageId="0" pageNumber="53">Corymbina aegaea</emphasis>
</taxonomicName>
(Oppenheim, 1919) from Upper Miocene (?) deposits near
<normalizedToken id="4D46EF529C0B1A395B7E7FC3F7C798EB" originalValue="Harmancık">Harmancik</normalizedToken>
(NW Turkey) has a more elongated shell (
<bibRefCitation id="5731D5FFAFAAE551D968F5CDB54ED17C" author="Schuett, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="53" pagination="217 - 229" refId="B138" refString="Schuett, H, Kavusan, G, 1984. Mollusken der miozaenen Suesswasserablagerungen in der Umgebung von Harmancik bei Kuetahya-Bursa in Nordwestanatolien. Archiv fuer Molluskenkunde 114: 217 - 229" title="Mollusken der miozaenen Suesswasserablagerungen in der Umgebung von Harmancik bei Kuetahya-Bursa in Nordwestanatolien." volume="114" year="1984">
<normalizedToken id="99DFF6A4DF182E5A3A3FD0AB1030B518" originalValue="Schütt">Schuett</normalizedToken>
and Kavusan 1984
</bibRefCitation>
). The Early Sarmatian (Serravallian, Middle Miocene)
<taxonomicName id="834C82EF7D4A014C9ADAFC315CA24ACB" baseAuthorityName="Marinescu" baseAuthorityYear="1992" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina coronata" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="coronata">
<emphasis id="54BB99C0EE714597BB580A74B70356F5" italics="true" pageId="0" pageNumber="53">Corymbina coronata</emphasis>
</taxonomicName>
(Marinescu, 1992) from Romania differs from
<taxonomicName id="0A4EE4CC2A63444402AC99778AEDE04F" lsidName="C. elegans" pageId="0" pageNumber="53" rank="species" species="elegans">
<emphasis id="7AAF9E921047145389CC902D396C7AF2" italics="true" pageId="0" pageNumber="53">C. elegans</emphasis>
</taxonomicName>
in the more angulated body whorl and the ribs being reduced to elongated knobs at the uppermost whorl portion.
</paragraph>
<paragraph id="0C76BB101A1430DD1FCEBB70656960ED" pageId="0" pageNumber="53">
<taxonomicName id="3111CD24B7259FDFFCE7FC99C5108BDB" authorityName="Inal" authorityYear="1975" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina rhodensis subsp. senestris" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="subSpecies" species="rhodensis" subSpecies="senestris">
<emphasis id="B9445F67176790CAE4110831DBDB2AEB" italics="true" pageId="0" pageNumber="53">Corymbina rhodensis senestris</emphasis>
</taxonomicName>
<normalizedToken id="FF4B1570376ECD96D20B7399DCBEC04A" originalValue="İnal">Inal</normalizedToken>
, 1975 from supposedly Pliocene sediments at
<normalizedToken id="E7C0B988EE923444B961E24BD4A3E8DD" originalValue="Gürleyik">Guerleyik</normalizedToken>
SE Afyonkarahisar (c. 170 km ENE Denizli) shows a similar type of sculpture, but it is a rare case of a sinistral representative (
<bibRefCitation id="505120635D22FAAB79A49F5D7F9ADC99" author="Inal, A" journalOrPublisher="Bulletin of the Geological Society of Turkey" pageId="0" pageNumber="53" pagination="161 - 164" refId="B71" refString="Inal, A, 1975. Description of two new Gastropod species and one sub-species from the lacustrine sediments of Neogene age Afyon, Turkey. Bulletin of the Geological Society of Turkey 18: 161 - 164" title="Description of two new Gastropod species and one sub-species from the lacustrine sediments of Neogene age Afyon, Turkey." volume="18" year="1975">
<normalizedToken id="D7113CCBCBFCE88E809F44BD616EB1AA" originalValue="İnal">Inal</normalizedToken>
1975
</bibRefCitation>
). It is questionable, however, if it represents a distinct species or simply an aberrant morphotype of an existing species.
</paragraph>
</subSubSection>
<subSubSection id="7E76F9023096DD196CF0D9EDAFFC0081" pageId="0" pageNumber="53" type="distribution">
<paragraph id="9958E9FDB7A51240B1AADE6329784405" pageId="0" pageNumber="53">Distribution.</paragraph>
<paragraph id="AD4130776260AB3D6667F636F1A1262B" pageId="0" pageNumber="53">
<taxonomicName id="DCC84BD6EDFA295096BB472FD64D2349" baseAuthorityName="Cantraine" baseAuthorityYear="1841" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina elegans" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="elegans">
<emphasis id="7EBFB855534EDE68752BA65457D7706E" italics="true" pageId="0" pageNumber="53">Corymbina elegans</emphasis>
</taxonomicName>
has been reported from the Pliocene of the Denizli,
<normalizedToken id="E9F709D079F021248CA4AE5239A27297" originalValue="Eşen">Esen</normalizedToken>
, and
<normalizedToken id="CF38D360B4E5F97E661131B012A2C7EC" originalValue="Çameli">Cameli</normalizedToken>
basins in Turkey (
<bibRefCitation id="EF0ACEE95CCC2D67C2D542100B8E5C23" author="Spratt, TAB" journalOrPublisher="Zeitschrift fuer wissenschaftliche Zoologie" pageId="0" pageNumber="53" publicationUrl="http://reader.digitale-sammlungen.de/de/fs1/object/display/bsb10468250_00009.html" refId="B146" refString="Spratt, TAB, Forbes, E, 1847. Travels in Lycia, Milyas, and the Cibyratis, in company with the late Rev. E. T. Daniell. 2 volumes. John van Voorst, Paternoster Row, London. [Volume 1:] http://reader.digitale-sammlungen.de/de/fs1/object/display/bsb10468250_00009.html" title="Travels in Lycia, Milyas, and the Cibyratis, in company with the late Rev. E. T. Daniell. 2 volumes. John van Voorst, Paternoster Row, London. [Volume 1:]" url="http://reader.digitale-sammlungen.de/de/fs1/object/display/bsb10468250_00009.html" year="1847">Spratt and Forbes 1847</bibRefCitation>
;
<bibRefCitation id="C9158DA69B0D62303AB9D51DA9742F9D" author="Oppenheim, P" journalOrPublisher="Zeitschrift der Deutschen Geologischen Gesellschaft" pageId="0" pageNumber="53" pagination="1 - 210" publicationUrl="https://www.biodiversitylibrary.org/page/43778447" refId="B102" refString="Oppenheim, P, 1919. Das Neogen in Kleinasien. Zeitschrift der Deutschen Geologischen Gesellschaft 70: 1 - 210, https://www.biodiversitylibrary.org/page/43778447" title="Das Neogen in Kleinasien." url="https://www.biodiversitylibrary.org/page/43778447" volume="70" year="1919">Oppenheim 1919</bibRefCitation>
; Wenz 1923), the Early Pliocene of central Italy (
<bibRefCitation id="6090A70ED4154F52ACE0DC276AAB6A1B" DOI="https://doi.org/10.1127/arch.moll/147/049-054" author="Esu, D" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="53" pagination="49 - 54" refId="B48" refString="Esu, D, Girotti, O, 2018. Valvata mathiasi n. sp. (Gastropoda: Heterobranchia: Valvatidae) from the Lower Pliocene of the Val di Pesa (Tuscany, Central Italy). Archiv fuer Molluskenkunde 147: 49 - 54, DOI: https://doi.org/10.1127/arch.moll/147/049-054" title="Valvata mathiasi n. sp. (Gastropoda: Heterobranchia: Valvatidae) from the Lower Pliocene of the Val di Pesa (Tuscany, Central Italy)." url="https://doi.org/10.1127/arch.moll/147/049-054" volume="147" year="2018">Esu and Girotti 2018</bibRefCitation>
), and the Pliocene-Pleistocene of Greece (Wenz 1923;
<bibRefCitation id="8B38A753A0615FE06B3771F98C4EAAF8" author="Schuett, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="53" pagination="1 - 29" refId="B137" refString="Schuett, H, Besenecker, H, 1973. Eine Molluskenfauna aus dem Neogen von Chios (Aegaeis). Archiv fuer Molluskenkunde 103: 1 - 29" title="Eine Molluskenfauna aus dem Neogen von Chios (Aegaeis)." volume="103" year="1973">
<normalizedToken id="F78923933EDEE1CAEDB54B409001DED1" originalValue="Schütt">Schuett</normalizedToken>
and Besenecker 1973
</bibRefCitation>
;
<bibRefCitation id="5981F856B75D12092EEA7CCD22512737" author="Gillet, S" journalOrPublisher="Annales Geologiques des Pays Helleniques" pageId="0" pageNumber="53" pagination="554 - 580" refId="B59" refString="Gillet, S, Sauvage, J, Keraudren, B, 1979. Etude paleontologique du plio-pleistocene de Locride et d'Eubee occidentale (Grece centrale): Malacologie et palynologie. Annales Geologiques des Pays Helleniques 29: 554 - 580" title="Etude paleontologique du plio-pleistocene de Locride et d'Eubee occidentale (Grece centrale): Malacologie et palynologie." volume="29" year="1979">Gillet et al. 1979</bibRefCitation>
). Records from the Late Miocene (Chios Island;
<bibRefCitation id="8A2B2DFEAEE468FBD1D19EA1B25FA0C7" author="Schuett, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="53" pagination="1 - 29" refId="B137" refString="Schuett, H, Besenecker, H, 1973. Eine Molluskenfauna aus dem Neogen von Chios (Aegaeis). Archiv fuer Molluskenkunde 103: 1 - 29" title="Eine Molluskenfauna aus dem Neogen von Chios (Aegaeis)." volume="103" year="1973">
<normalizedToken id="BA4B8F9B935F9AE7BBCC6E814D7C535D" originalValue="Schütt">Schuett</normalizedToken>
and Besenecker 1973
</bibRefCitation>
) are probably based on an outdated stratigraphy.
</paragraph>
<caption id="23745532A6074B027F20DA4FDAEF9236" doi="10.3897/zitteliana.97.115682.figure16" httpUri="https://binary.pensoft.net/fig/950753" pageId="0" pageNumber="53" start="Figure 16" startId="F16">
<paragraph id="DB3A8FB769A40F4156223249BD833435" pageId="0" pageNumber="53">
<emphasis id="A8AAFCA9EA8AEFAA1AD2DB23454FA8DC" bold="true" pageId="0" pageNumber="53">Figure 16.</emphasis>
<taxonomicName id="987AD94FFFAE8B0861E77655A38BB66C" class="Gastropoda" family="Lymnaeidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="family">Lymnaeidae</taxonomicName>
of the Kolankaya I fauna.
<emphasis id="F315C5CF6151122B6E2BF995406B5D5F" bold="true" pageId="0" pageNumber="53">A-C.</emphasis>
<taxonomicName id="78F783C10A2025EAEC548C776D16F8D1" baseAuthorityName="Cantraine" baseAuthorityYear="1841" class="Gastropoda" family="Lymnaeidae" genus="Corymbina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Corymbina elegans" order="Hygrophila" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="elegans">
<emphasis id="FADDE25B73CF6C7CBE5A3DBC44138DFA" italics="true" pageId="0" pageNumber="53">Corymbina elegans</emphasis>
</taxonomicName>
(Cantraine, 1841), RGM 962696, sample 2.
<emphasis id="78E0A9DFCFC314698DE483FE46560833" bold="true" pageId="0" pageNumber="53">D-F.</emphasis>
<taxonomicName id="4D521D37837BC7FF54EB515C2244CC68" authorityName="Montfort" authorityYear="1810" class="Gastropoda" family="Lymnaeidae" genus="Radix" kingdom="Animalia" lsidName="Radix" order="Cardiida" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
<emphasis id="0CEF450435DD048CA161F5D81DC0061B" italics="true" pageId="0" pageNumber="53">Radix</emphasis>
</taxonomicName>
sp., RGM 1310798, sample 2. Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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