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<mods:title id="CE28DAA8EEE59F2F6599CA61B0F9B9B2">Type designations for Neotropical water beetles in the genus Hydrochus Leach 1817 (Coleoptera: Hydrochidae)</mods:title>
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<treatment id="C17B87FF3C3B1231C0E6FC88FA14F7CB" ID-GBIF-Taxon="160605447" LSID="urn:lsid:plazi:treatment:C17B87FF3C3B1231C0E6FC88FA14F7CB" httpUri="http://treatment.plazi.org/id/C17B87FF3C3B1231C0E6FC88FA14F7CB" lastPageId="7" lastPageNumber="155" pageId="5" pageNumber="154">
<subSubSection id="01C865623C3B1233C0E6FC88FE48F1EC" box="[151,474,1730,1757]" pageId="5" pageNumber="154" type="nomenclature">
<paragraph id="496D36E93C3B1233C0E6FC88FE48F1EC" blockId="5.[151,474,1730,1793]" box="[151,474,1730,1757]" pageId="5" pageNumber="154">
<heading id="122581853C3B1233C0E6FC88FE48F1EC" bold="true" box="[151,474,1730,1757]" fontSize="11" level="1" pageId="5" pageNumber="154" reason="1">
<taxonomicName id="8ED24D6A3C3B1233C0E6FC88FE48F1EC" ID-CoL="3N67D" authority="Knisch" authorityName="Knisch" box="[151,474,1730,1757]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="7BA6EAFB3C3B1233C0E6FC88FE48F1EC" bold="true" box="[151,474,1730,1757]" pageId="5" pageNumber="154">
<emphasis id="7BA6EAFB3C3B1233C0E6FC88FEE8F1ED" bold="true" box="[151,378,1730,1757]" italics="true" pageId="5" pageNumber="154">Hydrochus ducalis</emphasis>
Knisch
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="01C865623C3B1233C0E6FCACFEAFF031" box="[151,317,1766,1793]" pageId="5" pageNumber="154" type="description">
<paragraph id="496D36E93C3B1233C0E6FCACFEAFF031" blockId="5.[151,474,1730,1793]" box="[151,317,1766,1793]" pageId="5" pageNumber="154">
<figureCitation id="D1E92A6C3C3B1233C0E6FCACFF77F031" box="[151,229,1766,1793]" captionStart="FIGURE 4" captionStartId="6.[151,250,1780,1805]" captionTargetBox="[166,1432,326,1745]" captionTargetId="figure@6.[151,1435,309,1755]" captionTargetPageId="6" captionText="FIGURE 4. Hydrochus ducalis Knisch, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3557935" httpUri="https://zenodo.org/record/3557935/files/figure.png" pageId="5" pageNumber="154">Figs. 4</figureCitation>
,
<figureCitation id="D1E92A6C3C3B1233C080FCACFE93F031" box="[241,257,1766,1793]" captionStart="FIGURE 5" captionStartId="7.[151,250,1550,1575]" captionTargetBox="[172,1435,309,1510]" captionTargetId="figure@7.[151,1435,273,1525]" captionTargetPageId="7" captionText="FIGURE 5. Hydrochus ducalis Knisch, habitus and male genitalia of paralectotype." figureDoi="http://doi.org/10.5281/zenodo.3557937" httpUri="https://zenodo.org/record/3557937/files/figure.png" pageId="5" pageNumber="154">5</figureCitation>
,
<figureCitation id="D1E92A6C3C3B1233C17CFCADFEAFF031" box="[269,317,1767,1793]" captionStart="FIGURES 12" captionStartId="15.[151,264,1980,2005]" captionTargetBox="[215,1415,381,1942]" captionTargetId="figure@15.[158,1430,381,1955]" captionTargetPageId="15" captionText="FIGURES 12AE. Labels of lectotypes. (A) Hydrochus ducalis Knisch; (B) Hydrochus drakei Knisch; (C) Hydrochus variabilis Knisch; (D) Hydrochus corruscans Bruch; (E) Hydrochus argutus Knisch." figureDoi="http://doi.org/10.5281/zenodo.3557951" httpUri="https://zenodo.org/record/3557951/files/figure.png" pageId="5" pageNumber="154">12A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="01C865623C3B1233C0E6FD61FD9BF073" box="[151,521,1834,1859]" pageId="5" pageNumber="154" type="reference_group">
<paragraph id="496D36E93C3B1233C0E6FD61FD9BF073" blockId="5.[151,663,1834,1890]" box="[151,521,1834,1859]" pageId="5" pageNumber="154">
<treatmentCitationGroup id="69C211C73C3B1233C0E6FD61FD9BF073" box="[151,521,1834,1859]" pageId="5" pageNumber="154">
<taxonomicName id="8ED24D6A3C3B1233C0E6FD61FD96F073" authority="Knisch 1921: 1" authorityName="Knisch" authorityPageNumber="1" authorityYear="1921" box="[151,516,1834,1859]" class="Aves" family="Apodidae" genus="Hydrochous" kingdom="Animalia" order="Apodiformes" pageId="5" pageNumber="154" phylum="Chordata" rank="species" species="ducalis">
<emphasis id="7BA6EAFB3C3B1233C0E6FD61FEF3F073" box="[151,353,1835,1859]" italics="true" pageId="5" pageNumber="154">Hydrochous ducalis</emphasis>
<treatmentCitation id="C87310F83C3B1233C119FD60FD96F073" author="Knisch, A." box="[360,516,1834,1859]" page="1" pageId="5" pageNumber="154" year="1921">
<bibRefCitation id="2D434B183C3B1233C119FD60FD96F073" author="Knisch, A." box="[360,516,1834,1859]" pageId="5" pageNumber="154" pagination="1 - 24" refId="ref5602" refString="Knisch, A. (1921) Hydrophiliden aus Matto Grosso (Erster Teil). Archiv fur Naturegeschichte, 87 A, 1 - 24." type="journal article" year="1921">Knisch 1921: 1</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<paragraph id="496D36E93C3B1233C0E6FD03FD05F052" blockId="5.[151,663,1834,1890]" box="[151,663,1865,1890]" pageId="5" pageNumber="154">
<taxonomicName id="8ED24D6A3C3B1233C0E6FD03FD96F051" ID-CoL="3N677" authority="Makhan 1992" authorityName="Makhan" authorityYear="1992" box="[151,516,1865,1890]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="desenderi">
<emphasis id="7BA6EAFB3C3B1233C0E6FD03FEFCF051" box="[151,366,1865,1889]" italics="true" pageId="5" pageNumber="154">Hydrochus desenderi</emphasis>
<bibRefCitation id="2D434B183C3B1233C104FD03FD96F051" author="Makhan, D." box="[373,516,1865,1890]" pageId="5" pageNumber="154" pagination="95 - 103" refId="ref5661" refString="Makhan, D. (1992) Twelve new Hydrochus - species from South America. Phegea, 20 (2), 95 - 103." type="journal article" year="1992">Makhan 1992</bibRefCitation>
</taxonomicName>
;
<bibRefCitation id="2D434B183C3B1233C27FFD03FD06F052" author="Oliva, A." box="[526,660,1865,1890]" pageId="5" pageNumber="154" pagination="301 - 341" refId="ref6127" refString="Oliva, A. (1996) The genus Hydrochus Leach (Coleoptera: Hydrophiloidea: Hydrochidae) in South America, with special reference to Argentina. Bulletin et annales de la Societe royale belge d'entomologie, 132, 301 - 341." type="journal article" year="1996">Oliva (1996)</bibRefCitation>
.
</paragraph>
<paragraph id="496D36E93C3B1230C0E6FDDAFE46F62F" blockId="5.[151,1437,1936,2034]" lastBlockId="6.[151,1437,152,287]" lastPageId="6" lastPageNumber="155" pageId="5" pageNumber="154">
The RBINS kindly provided four
<typeStatus id="9669884B3C3B1233C265FDDAFDE4F09A" box="[532,630,1936,1962]" pageId="5" pageNumber="154" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="8ED24D6A3C3B1233C2ECFDDAFC5BF09A" ID-CoL="3N67D" authority="Knisch" authorityName="Knisch" box="[669,969,1936,1962]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="7BA6EAFB3C3B1233C2ECFDDAFCFDF09A" box="[669,879,1936,1962]" italics="true" pageId="5" pageNumber="154">Hydrochus ducalis</emphasis>
Knisch
</taxonomicName>
, which are part of the Orchymont/Knisch types collection. All four
<typeStatus id="9669884B3C3B1233C1B3FDFEFDB6F0FE" box="[450,548,1972,1998]" pageId="5" pageNumber="154" type="syntype">syntypes</typeStatus>
are conspecific males. I have selected one to serve as
<typeStatus id="9669884B3C3B1233C4D9FDFEFA9DF0FE" box="[1192,1295,1972,1998]" pageId="5" pageNumber="154" type="lectotype">lectotype</typeStatus>
and labeled it as such, and provide images of the habitus, male genitalia, and labels of this specimen (
<figureCitation id="D1E92A6C3C3B1233C4D8FD92FB69F0C2" box="[1193,1275,2008,2034]" captionStart="FIGURE 4" captionStartId="6.[151,250,1780,1805]" captionTargetBox="[166,1432,326,1745]" captionTargetId="figure@6.[151,1435,309,1755]" captionTargetPageId="6" captionText="FIGURE 4. Hydrochus ducalis Knisch, habitus and male genitalia of lectotype." figureDoi="http://doi.org/10.5281/zenodo.3557935" httpUri="https://zenodo.org/record/3557935/files/figure.png" pageId="5" pageNumber="154">Figs. 4</figureCitation>
,
<figureCitation id="D1E92A6C3C3B1233C578FD92FAAFF0C2" box="[1289,1341,2008,2034]" captionStart="FIGURES 12" captionStartId="15.[151,264,1980,2005]" captionTargetBox="[215,1415,381,1942]" captionTargetId="figure@15.[158,1430,381,1955]" captionTargetPageId="15" captionText="FIGURES 12AE. Labels of lectotypes. (A) Hydrochus ducalis Knisch; (B) Hydrochus drakei Knisch; (C) Hydrochus variabilis Knisch; (D) Hydrochus corruscans Bruch; (E) Hydrochus argutus Knisch." figureDoi="http://doi.org/10.5281/zenodo.3557951" httpUri="https://zenodo.org/record/3557951/files/figure.png" pageId="5" pageNumber="154">12A</figureCitation>
). Lecto-
<subSubSection id="01C865623C381230C0E6FAD3FE46F62F" pageId="6" pageNumber="155" type="description">
type labels (
<figureCitation id="D1E92A6C3C381230C152FAD3FE1FF783" box="[291,397,153,179]" captionStart="FIGURES 12" captionStartId="15.[151,264,1980,2005]" captionTargetBox="[215,1415,381,1942]" captionTargetId="figure@15.[158,1430,381,1955]" captionTargetPageId="15" captionText="FIGURES 12AE. Labels of lectotypes. (A) Hydrochus ducalis Knisch; (B) Hydrochus drakei Knisch; (C) Hydrochus variabilis Knisch; (D) Hydrochus corruscans Bruch; (E) Hydrochus argutus Knisch." figureDoi="http://doi.org/10.5281/zenodo.3557951" httpUri="https://zenodo.org/record/3557951/files/figure.png" pageId="6" pageNumber="155">Fig. 12A</figureCitation>
): “Bras.Corumba [Matto Grosso] // Coll. A.Kniž COTYPUS // det. Kniž
<taxonomicName id="8ED24D6A3C381230C4A5FAD3FAB7F783" box="[1236,1317,153,179]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="155" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="7BA6EAFB3C381230C4A5FAD3FAB7F783" box="[1236,1317,153,179]" italics="true" pageId="6" pageNumber="155">ducalis</emphasis>
</taxonomicName>
m. // coll. A.dOrchymont:
<taxonomicName id="8ED24D6A3C381230C126FAF7FE43F7E7" authorityName="Leach" authorityYear="1817" box="[343,465,189,215]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="7BA6EAFB3C381230C126FAF7FE43F7E7" box="[343,465,189,215]" italics="true" pageId="6" pageNumber="155">Hydrochus</emphasis>
</taxonomicName>
R.M.H.N.B. 15.962 // HABITUS/GENIT. DIGITAL IMAGES captured 2019 P.D. Perkins // [red]
<typeStatus id="9669884B3C381230C150FAABFE53F7CB" box="[289,449,225,251]" pageId="6" pageNumber="155" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="8ED24D6A3C381230C1B9FAABFD62F7CB" authority="Knisch" authorityName="Knisch" box="[456,752,225,251]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="155" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="7BA6EAFB3C381230C1B9FAABFD08F7CB" box="[456,666,225,251]" italics="true" pageId="6" pageNumber="155">Hydrochus ducalis</emphasis>
Knisch
</taxonomicName>
desig. P. D. Perkins.” The three non-lectotype
<typeStatus id="9669884B3C381230C48FFAABFAF2F7CB" box="[1278,1376,225,251]" pageId="6" pageNumber="155" type="syntype">syntypes</typeStatus>
have been labeled
<typeStatus id="9669884B3C381230C15AFB4FFE42F62F" box="[299,464,261,287]" pageId="6" pageNumber="155" type="paralectotype">paralectotypes</typeStatus>
.
</subSubSection>
</paragraph>
<caption id="1DAD66613C381230C0E6FCBEFC53F03D" ID-DOI="http://doi.org/10.5281/zenodo.3557935" ID-Zenodo-Dep="3557935" box="[151,961,1780,1805]" httpUri="https://zenodo.org/record/3557935/files/figure.png" pageId="6" pageNumber="155" startId="6.[151,250,1780,1805]" targetBox="[166,1432,326,1745]" targetPageId="6">
<paragraph id="496D36E93C381230C0E6FCBEFC53F03D" blockId="6.[151,961,1780,1805]" box="[151,961,1780,1805]" pageId="6" pageNumber="155">
<emphasis id="7BA6EAFB3C381230C0E6FCBEFE81F03D" bold="true" box="[151,275,1780,1805]" pageId="6" pageNumber="155">FIGURE 4.</emphasis>
<taxonomicName id="8ED24D6A3C381230C168FCBFFDB5F03D" authority="Knisch" authorityName="Knisch" box="[281,551,1780,1805]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="155" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="7BA6EAFB3C381230C168FCBFFE45F03D" box="[281,471,1781,1805]" italics="true" pageId="6" pageNumber="155">Hydrochus ducalis</emphasis>
Knisch
</taxonomicName>
, habitus and male genitalia of lectotype.
</paragraph>
</caption>
<subSubSection id="01C865623C381231C0B6FD76FA14F7CB" lastPageId="7" lastPageNumber="156" pageId="6" pageNumber="155" type="discussion">
<paragraph id="496D36E93C381231C0B6FD76FA14F7CB" blockId="6.[151,1436,1851,2022]" lastBlockId="7.[151,1436,153,251]" lastPageId="7" lastPageNumber="156" pageId="6" pageNumber="155">
<emphasis id="7BA6EAFB3C381230C0B6FD76FEA9F066" bold="true" box="[199,315,1852,1878]" pageId="6" pageNumber="155">Remarks.</emphasis>
One of the male
<typeStatus id="9669884B3C381230C186FD71FD0BF065" box="[503,665,1851,1877]" pageId="6" pageNumber="155" type="paralectotype">paralectotypes</typeStatus>
has a distinctly more ovoid basal piece, while the structures of the aedeagus and the parameres do not significantly differ from those of the
<typeStatus id="9669884B3C381230C3DBFD15FB86F049" box="[938,1044,1887,1913]" pageId="6" pageNumber="155" type="lectotype">lectotype</typeStatus>
. Images of this specimen are given as an example of some of the infraspecific variation to be expected in species of
<taxonomicName id="8ED24D6A3C381230C458FDC9FB31F0AD" authorityName="Leach" authorityYear="1817" box="[1065,1187,1923,1949]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="7BA6EAFB3C381230C458FDC9FB31F0AD" box="[1065,1187,1923,1949]" italics="true" pageId="6" pageNumber="155">Hydrochus</emphasis>
</taxonomicName>
(
<figureCitation id="D1E92A6C3C381230C4C2FDC9FB6AF0AD" box="[1203,1272,1922,1949]" captionStart="FIGURE 5" captionStartId="7.[151,250,1550,1575]" captionTargetBox="[172,1435,309,1510]" captionTargetId="figure@7.[151,1435,273,1525]" captionTargetPageId="7" captionText="FIGURE 5. Hydrochus ducalis Knisch, habitus and male genitalia of paralectotype." figureDoi="http://doi.org/10.5281/zenodo.3557937" httpUri="https://zenodo.org/record/3557937/files/figure.png" pageId="6" pageNumber="155">Fig. 5</figureCitation>
). The drawing of
<taxonomicName id="8ED24D6A3C381230C0C9FDEDFE76F0F1" authority="Knisch" authorityName="Knisch" box="[184,484,1959,1985]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="155" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="7BA6EAFB3C381230C0C9FDEDFE1EF0F1" box="[184,396,1959,1985]" italics="true" pageId="6" pageNumber="155">Hydrochus ducalis</emphasis>
Knisch
</taxonomicName>
male genitalia by
<bibRefCitation id="2D434B183C381230C2CCFDEDFCDDF0F1" author="Oliva, A." box="[701,847,1959,1985]" pageId="6" pageNumber="155" pagination="87 - 104" refId="ref6088" refString="Oliva, A. (1992) The species of Hydrochus (Coleoptera; Hydrophiloidea; Hydrochidae) described from South America. Bulletin et annales de la Societe royale belge d'entomologie, 128, 87 - 104." type="journal article" year="1992">Oliva (1992)</bibRefCitation>
is unfortunately very inaccurate in several aspects, including general proportions, shapes of the parameres, size of the basal piece, etc. I have also examined the holo- type of
<taxonomicName id="8ED24D6A3C391231C09AFAD3FDEAF783" authority="Makhan 1992" authorityName="Makhan" authorityYear="1992" box="[235,632,153,179]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="desenderi">
<emphasis id="7BA6EAFB3C391231C09AFAD3FE4AF783" box="[235,472,153,179]" italics="true" pageId="7" pageNumber="156">Hydrochus desenderi</emphasis>
<bibRefCitation id="2D434B183C391231C1AFFAD3FDEAF783" author="Makhan, D." box="[478,632,153,179]" pageId="7" pageNumber="156" pagination="95 - 103" refId="ref5661" refString="Makhan, D. (1992) Twelve new Hydrochus - species from South America. Phegea, 20 (2), 95 - 103." type="journal article" year="1992">Makhan 1992</bibRefCitation>
</taxonomicName>
(which is one of the
<typeStatus id="9669884B3C391231C311FAD3FC50F783" box="[864,962,153,179]" pageId="7" pageNumber="156" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="8ED24D6A3C391231C397FAD3FB25F783" box="[998,1207,153,179]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="7BA6EAFB3C391231C397FAD3FB25F783" box="[998,1207,153,179]" italics="true" pageId="7" pageNumber="156">Hydrochus ducalis</emphasis>
</taxonomicName>
), including the male genitalia. I concur with
<bibRefCitation id="2D434B183C391231C1D2FAF7FDA6F7E8" author="Oliva, A." box="[419,564,189,216]" pageId="7" pageNumber="156" pagination="301 - 341" refId="ref6127" refString="Oliva, A. (1996) The genus Hydrochus Leach (Coleoptera: Hydrophiloidea: Hydrochidae) in South America, with special reference to Argentina. Bulletin et annales de la Societe royale belge d'entomologie, 132, 301 - 341." type="journal article" year="1996">Oliva (1996)</bibRefCitation>
that
<taxonomicName id="8ED24D6A3C391231C21DFAF7FC2CF7E7" authority="Makhan" authorityName="Makhan" authorityYear="1992" box="[620,958,189,215]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="desenderi">
<emphasis id="7BA6EAFB3C391231C21DFAF7FCC9F7E7" box="[620,859,189,215]" italics="true" pageId="7" pageNumber="156">Hydrochus desenderi</emphasis>
Makhan
</taxonomicName>
is a junior synonym of
<taxonomicName id="8ED24D6A3C391231C4BBFAF7FEB2F7CB" authority="Knisch. The" authorityName="Knisch. The" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="ducalis">
<emphasis id="7BA6EAFB3C391231C4BBFAF7FA0EF7E7" box="[1226,1436,189,215]" italics="true" pageId="7" pageNumber="156">Hydrochus ducalis</emphasis>
Knisch. The
</taxonomicName>
<typeStatus id="9669884B3C391231C156FAABFE1BF7CB" box="[295,393,225,251]" pageId="7" pageNumber="156" type="holotype">holotype</typeStatus>
of
<taxonomicName id="8ED24D6A3C391231C1DFFAA8FD31F7CB" authority="Makhan" authorityName="Makhan" authorityYear="1992" box="[430,675,225,251]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="desenderi">
<emphasis id="7BA6EAFB3C391231C1DFFAA8FDAFF7CB" box="[430,573,225,251]" italics="true" pageId="7" pageNumber="156">H. desenderi</emphasis>
Makhan
</taxonomicName>
, deposited in the RBINS, has the same label data as the
<typeStatus id="9669884B3C391231C566FAABFA13F7CB" box="[1303,1409,225,251]" pageId="7" pageNumber="156" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>