plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/85/69/BD/8569BD6CFF96126031A91C47FDC2F972.xml
maintenance a6fc63282e added 85
2024-06-21 12:42:32 +02:00

1083 lines
164 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="43C76E5C982953DCBBF4327372E32AAA" ID-CLB-Dataset="9031" ID-DOI="10.11646/zootaxa.4861.4.2" ID-GBIF-Dataset="a15b3af7-16c7-492b-bf7b-d08a36b7f6a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4416659" ID-ZooBank="8B330C85-5B40-48EF-8C17-48332637C1C9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609800668329" checkinUser="plazi" docAuthor="Phillips, Gary, Moulton, John K. &amp; Bernard, Ernest C." docDate="2020" docId="8569BD6CFF96126031A91C47FDC2F972" docLanguage="en" docName="zootaxa.4861.4.2.pdf" docOrigin="Zootaxa 4861 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heth pivari Phillips &amp; Moulton &amp; Bernard 2020, n. sp." docType="treatment" docVersion="9" lastPageNumber="498" masterDocId="7950C514FF90126C313E1B02FFC3FFDF" masterDocTitle="Heth pivari n. sp. (Nematoda: Ransomnematoidea: Hethidae) from the indigenous North American millipede Narceus gordanus (Spirobolida: Spirobolidae), with keys for worldwide Heth spp." masterLastPageNumber="514" masterPageNumber="486" pageNumber="492" updateTime="1698852234366" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="679F8D346B812ABA241B7C0C3F7E0C2C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F077651CF6B8F82E386C4A4C2C33A384">
<mods:title id="93830C2B04CE9ACDB93F6D804CE2E744">Heth pivari n. sp. (Nematoda: Ransomnematoidea: Hethidae) from the indigenous North American millipede Narceus gordanus (Spirobolida: Spirobolidae), with keys for worldwide Heth spp.</mods:title>
</mods:titleInfo>
<mods:name id="710D02A6FD5C11F3724051F181664CB2" type="personal">
<mods:role id="C52BFA33383D8C3EBD02B476E897C95C">
<mods:roleTerm id="0286D488CD8E98DD04748AF405303F11">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="13878100004843FD40097E63FE760DBE">Phillips, Gary</mods:namePart>
<mods:nameIdentifier id="FD700EE81329DEDB90D3A645EBE9C2D9" type="ORCID">0000-0003-1930-1009</mods:nameIdentifier>
<mods:affiliation id="482E808CF0B0F46585EB8FAF08796250">Department of Entomology and Plant Pathology, University of Tennessee, 370 Plant Biotechnology Building, 2505 E J Chapman Drive, Knoxville, TN 37996 - 4560 USA. &amp; https: // orcid. org / 0000 - 0003 - 1930 - 1009</mods:affiliation>
</mods:name>
<mods:name id="5CE8A92F904F736210E1FC39FEEF9128" type="personal">
<mods:role id="849EA6856C316789DE052F913E8DFE46">
<mods:roleTerm id="4F565469AC21B30397CF14746C1DC20A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F3AF6FD2E37F1A4597BEFC4B90EA11C2">Moulton, John K.</mods:namePart>
<mods:nameIdentifier id="A5D8FC44614A9C38AA593EB99E559006" type="ORCID">0000-0001-8760-3274</mods:nameIdentifier>
<mods:affiliation id="FC3AE5481AA38118D74688CB3A0CB78F">Department of Entomology and Plant Pathology, University of Tennessee, 370 Plant Biotechnology Building, 2505 E J Chapman Drive, Knoxville, TN 37996 - 4560 USA. &amp; https: // orcid. org / 0000 - 0001 - 8760 - 3274</mods:affiliation>
</mods:name>
<mods:name id="3763C3AC2964C286919A3D9E8C195A2D" type="personal">
<mods:role id="A9F1247DB19F6FBB4FC6E4B0F539F1CB">
<mods:roleTerm id="8C96AE19C22778D04DDA442229458DB6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="416F05BE6D72DC8327FE6FB3269D7AA3">Bernard, Ernest C.</mods:namePart>
<mods:nameIdentifier id="69456672E9798C000E47C728E8CF6012" type="ORCID">0000-0003-3359-1516</mods:nameIdentifier>
<mods:affiliation id="347A702DFE6A78818E61997481FA7211">Department of Entomology and Plant Pathology, University of Tennessee, 370 Plant Biotechnology Building, 2505 E J Chapman Drive, Knoxville, TN 37996 - 4560 USA. &amp; https: // orcid. org / 0000 - 0003 - 3359 - 1516</mods:affiliation>
</mods:name>
<mods:typeOfResource id="08B1D49F6D4526E352A6E3D57BA2828B">text</mods:typeOfResource>
<mods:relatedItem id="EB792D0140C7D56AC19E6EACFCEC3195" type="host">
<mods:titleInfo id="217E72BB2C2DB85B1BFE5BD6398B3868">
<mods:title id="191D4AE934BB123F92A314C119C0CEFF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="065092E9E411F4C5D1296885E13F788D">
<mods:date id="8D4562952AB647BFA50994BFD7E11732">2020</mods:date>
<mods:detail id="782C06CDEB664BA9BC69E5F39C6A03AC" type="pubDate">
<mods:number id="BC084D64E1092D99F30496B5D6D509F3">2020-10-20</mods:number>
</mods:detail>
<mods:detail id="FF7DD9AD32636AB346E396DB74E097EC" type="volume">
<mods:number id="6E001F2D8FDDB16059B8EE7878D71FB4">4861</mods:number>
</mods:detail>
<mods:detail id="E0D70E2F2902A6EB571C8270A7D5AB1F" type="issue">
<mods:number id="4EC6743D20807849330B1AC611F6E26F">4</mods:number>
</mods:detail>
<mods:extent id="026D90C566D6652D7758AD1884325473" unit="page">
<mods:start id="4E9D64670CFE84A641E817CF70DAA274">486</mods:start>
<mods:end id="5736CFB131B1DAEDA900AD05C0FB4EB1">514</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="0C53AC45A51B782A75F426234E0D504E">journal article</mods:classification>
<mods:identifier id="A4CDA9B54023C03E09F0DBB678C53BF7" type="CLB-Dataset">9031</mods:identifier>
<mods:identifier id="BC5EDD93B5942BF24BBF67036EE1C49C" type="DOI">10.11646/zootaxa.4861.4.2</mods:identifier>
<mods:identifier id="B8FDA46954CACC14B0BD2557374993FB" type="GBIF-Dataset">a15b3af7-16c7-492b-bf7b-d08a36b7f6a2</mods:identifier>
<mods:identifier id="421FA7B9CEBD974DCE5E6F1688E9FEF6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="64099B298F7913D664EDB70BC604E5CF" type="Zenodo-Dep">4416659</mods:identifier>
<mods:identifier id="7063896FA7703E497F7E3081330324D7" type="ZooBank">8B330C85-5B40-48EF-8C17-48332637C1C9</mods:identifier>
</mods:mods>
<treatment id="8569BD6CFF96126031A91C47FDC2F972" ID-DOI="http://doi.org/10.5281/zenodo.4416770" ID-GBIF-Taxon="176440189" ID-Zenodo-Dep="4416770" LSID="urn:lsid:plazi:treatment:8569BD6CFF96126031A91C47FDC2F972" httpUri="http://treatment.plazi.org/id/8569BD6CFF96126031A91C47FDC2F972" lastPageId="12" lastPageNumber="498" pageId="6" pageNumber="492">
<subSubSection id="45DA5FF1FF96126A31A91C47FE5FF85B" pageId="6" pageNumber="492" type="nomenclature">
<paragraph id="0D7F0C7AFF96126A31A91C47FEA8F8BF" blockId="6.[151,412,1861,1924]" box="[151,363,1861,1888]" pageId="6" pageNumber="492">
<heading id="5637BB16FF96126A31A91C47FEA8F8BF" bold="true" box="[151,363,1861,1888]" fontSize="11" level="1" pageId="6" pageNumber="492" reason="1">
<emphasis id="3FB4D068FF96126A31A91C47FEA8F8BF" bold="true" box="[151,363,1861,1888]" pageId="6" pageNumber="492">
<taxonomicName id="CAC077F9FF96126A31A91C47FEE2F8BF" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[151,289,1861,1888]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="6" pageNumber="492" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF96126A31A91C47FEE2F8BF" bold="true" box="[151,289,1861,1888]" italics="true" pageId="6" pageNumber="492">Heth pivari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24876D13FF96126A30161C44FEA8F8BF" box="[296,363,1862,1888]" pageId="6" pageNumber="492" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="0D7F0C7AFF96126A31A91C68FE5FF85B" blockId="6.[151,412,1861,1924]" box="[151,412,1898,1924]" pageId="6" pageNumber="492">
(
<tableCitation id="404239C1FF96126A31A11C68FEE3F85B" box="[159,288,1898,1924]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStart-2="TABLE 4" captionStartId-0="8.[151,239,260,284]" captionStartId-1="10.[151,241,151,175]" captionStartId-2="16.[151,239,477,503]" captionTargetPageId-0="8" captionTargetPageId-1="10" captionTargetPageId-2="16" captionText-0="TABLE 2. Morphometric measurements of Heth pivari n. sp. females." captionText-1="TABLE 3. Morphometric measurements of Heth pivari n. sp. males." captionText-2="TABLE 4. Selected Heth male characteristics." httpUri-0="http://table.plazi.org/id/59BF5CF2FF98126431A91A06FC4FFEC3" httpUri-1="http://table.plazi.org/id/59BF5CF2FF9A126631A91B95FCBAFF70" httpUri-2="http://table.plazi.org/id/59BF5CF2FF80127C31A91ADFFD44FE28" pageId="6" pageNumber="492" tableUuid-0="59BF5CF2FF98126431A91A06FC4FFEC3" tableUuid-1="59BF5CF2FF9A126631A91B95FCBAFF70" tableUuid-2="59BF5CF2FF80127C31A91ADFFD44FE28">Tables 2–4</tableCitation>
;
<figureCitation id="95FB10FFFF96126A30151C68FE56F85B" box="[299,405,1898,1924]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="7.[151,250,1201,1225]" captionStartId-1="9.[151,250,1497,1521]" captionStartId-2="11.[151,250,1896,1920]" captionStartId-3="12.[151,250,1361,1385]" captionStartId-4="13.[151,250,1904,1928]" captionStartId-5="14.[151,250,1613,1637]" captionTargetBox-0="[156,1432,457,1173]" captionTargetBox-1="[155,1429,349,1466]" captionTargetBox-2="[160,1426,354,1861]" captionTargetBox-3="[155,1433,184,1334]" captionTargetBox-4="[154,1433,672,1879]" captionTargetBox-5="[155,1432,185,1583]" captionTargetId-0="figure-135@7.[151,1436,453,1177]" captionTargetId-1="figure-113@9.[151,1436,345,1473]" captionTargetId-2="figure-116@11.[151,1436,345,1872]" captionTargetId-3="figure-18@12.[151,1436,181,1337]" captionTargetId-4="figure-214@13.[151,1435,669,1881]" captionTargetId-5="figure-18@14.[151,1436,181,1589]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionTargetPageId-2="11" captionTargetPageId-3="12" captionTargetPageId-4="13" captionTargetPageId-5="14" captionText-0="FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov &amp; Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm." captionText-1="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." captionText-2="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." captionText-3="FIGURE 5. SEM micrographs of Heth pivari n. sp. males. A) Anterior end, stoma slit-like. B) Anterior end, stoma open. C) Closeup of stoma showing striae and minute amphid aperture (a). D) Posterior region showing ventral sucker (S), papillae 1,2 and 4, and tail. E) Posterior region showing arrangement of posterior papillae 2–7. F) Closeup of cloacal region. Scale bars: A, B, E, 30 µm; C, F, 20 µm; D, 200 µm." captionText-4="FIGURE 6. DIC micrographs of Heth pivari n. sp., female and male structures. A) Lateral ornamentation in anterior region of female. B) Male, lateral somal extensions of posterior arcade surrounding anterior esophagus. C) Female, secretory-excretory pore (arrow). D. Female, somatic papillae anterior to vulva. E) Fusiform to comet-shaped sperm in male. All scale bars 20 µm." captionText-5="FIGURE 7. Heth pivari n. sp. A) Live female with single egg. B–E) Female enlargement and increasing numbers of eggs. The uterus is capacious but the vagina holds just one egg at a time (B, D). F) Female, dense trichomycete attachment in anterior region. G) Male, arthropod seta lodged in anteriormost region of intestine. H) Male, lateral chord (arrows). Scale bars: A, 500 µm; B–F, 250 µm; G, 100 µm; H, 50 µm." figureDoi-0="http://doi.org/10.5281/zenodo.4416665" figureDoi-1="http://doi.org/10.5281/zenodo.4416668" figureDoi-2="http://doi.org/10.5281/zenodo.4416670" figureDoi-3="http://doi.org/10.5281/zenodo.4416674" figureDoi-4="http://doi.org/10.5281/zenodo.4416676" figureDoi-5="http://doi.org/10.5281/zenodo.4416680" httpUri-0="https://zenodo.org/record/4416665/files/figure.png" httpUri-1="https://zenodo.org/record/4416668/files/figure.png" httpUri-2="https://zenodo.org/record/4416670/files/figure.png" httpUri-3="https://zenodo.org/record/4416674/files/figure.png" httpUri-4="https://zenodo.org/record/4416676/files/figure.png" httpUri-5="https://zenodo.org/record/4416680/files/figure.png" pageId="6" pageNumber="492">Figs. 2–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="45DA5FF1FF96126B31A91CB0FD88FEBA" lastPageId="7" lastPageNumber="493" pageId="6" pageNumber="492" type="materials_examined">
<paragraph id="0D7F0C7AFF96126B31A91CB0FAD4FF0A" blockId="6.[151,1437,1969,2032]" lastBlockId="7.[151,1437,150,429]" lastPageId="7" lastPageNumber="493" pageId="6" pageNumber="492">
<materialsCitation id="BDA80627FF96126A31A91CB0FBA2F82F" ID-GBIF-Occurrence="3012561305" county="Marion County" latitude="29.257727" location="Ocala National Forest" longLatPrecision="1" longitude="-81.7787" municipality="Silver Springs" pageId="6" pageNumber="492" specimenCount="1" stateProvince="Florida" typeStatus="holotype">
<emphasis id="3FB4D068FF96126A31A91CB0FE01F814" bold="true" box="[151,450,1969,1996]" pageId="6" pageNumber="492">
<typeStatus id="D27BB2D8FF96126A31A91CB0FF17F813" box="[151,212,1970,1996]" pageId="6" pageNumber="492">Type</typeStatus>
locality and habitat.
</emphasis>
<collectingRegion id="CF04C298FF96126A30F81CB0FDD8F813" box="[454,539,1970,1996]" country="United States of America" name="Florida" pageId="6" pageNumber="492">Florida</collectingRegion>
:
<collectingCounty id="E41E74F6FF96126A331D1CB0FD0DF813" box="[547,718,1970,1996]" pageId="6" pageNumber="492">Marion County</collectingCounty>
,
<collectingMunicipality id="ED1B9600FF96126A33E91CB0FCB5F813" box="[727,886,1970,1996]" pageId="6" pageNumber="492">Silver Springs</collectingMunicipality>
,
<location id="081F5AA1FF96126A32411CB0FBACF813" LSID="urn:lsid:plazi:treatment:8569BD6CFF96126031A91C47FDC2F972:081F5AA1FF96126A32411CB0FBACF813" box="[895,1135,1970,1996]" county="Marion County" latitude="29.257727" longLatPrecision="1" longitude="-81.7787" municipality="Silver Springs" name="Ocala National Forest" pageId="6" pageNumber="492" stateProvince="Florida">Ocala National Forest</location>
(
<geoCoordinate id="68F46ABDFF96126A35431CB3FACCF813" box="[1149,1295,1969,1996]" degrees="29.257726" direction="north" orientation="latitude" pageId="6" pageNumber="492" precision="1" value="29.257727">29.257726 N</geoCoordinate>
, –
<geoCoordinate id="68F46ABDFF96126A341B1CB0FF72F82F" degrees="81.778702" direction="west" orientation="longitude" pageId="6" pageNumber="492" precision="1" value="-81.7787">81.778702 W</geoCoordinate>
), xeric scrub located near
<location id="081F5AA1FF96126A30E91CD4FD28F82F" LSID="urn:lsid:plazi:treatment:8569BD6CFF96126031A91C47FDC2F972:081F5AA1FF96126A30E91CD4FD28F82F" box="[471,747,2006,2032]" county="Marion County" latitude="29.257727" longLatPrecision="1" longitude="-81.7787" municipality="Silver Springs" name="Mount Dora Sand Ridge" pageId="6" pageNumber="492" stateProvince="Florida">Mount Dora Sand Ridge</location>
, in intestine of
<taxonomicName id="CAC077F9FF96126A32AA1CD4FBA2F82F" authorityName="Chamberlin" authorityYear="1943" box="[916,1121,2006,2032]" class="Diplopoda" family="Spirobolidae" genus="Narceus" kingdom="Animalia" order="Spirobolida" pageId="6" pageNumber="492" phylum="Arthropoda" rank="species" species="gordanus">
<emphasis id="3FB4D068FF96126A32AA1CD4FBA2F82F" box="[916,1121,2006,2032]" italics="true" pageId="6" pageNumber="492">Narceus gordanus</emphasis>
</taxonomicName>
</materialsCitation>
.
<materialsCitation id="BDA80627FF96126B35501CD4FAD3FF0A" ID-GBIF-Occurrence="3012561301" county="Hernando County" lastPageId="7" lastPageNumber="493" latitude="28.707008" location="Citrus Springs" longLatPrecision="1" longitude="-89.39525" municipality="Ridge Manor" pageId="6" pageNumber="492" specimenCount="1" stateProvince="Florida">
Other specimens collected:
<collectingMunicipality id="ED1B9600FF97126B31A91B95FEF1FF6E" box="[151,306,151,177]" pageId="7" pageNumber="493">Ridge Manor</collectingMunicipality>
,
<collectingCounty id="E41E74F6FF97126B307E1B95FDC9FF6E" box="[320,522,151,177]" pageId="7" pageNumber="493">Hernando County</collectingCounty>
(
<geoCoordinate id="68F46ABDFF97126B33221B94FD75FF6E" box="[540,694,150,177]" degrees="28.509794" direction="north" orientation="latitude" pageId="7" pageNumber="493" precision="1" value="28.509794">28.509794 N</geoCoordinate>
, –
<geoCoordinate id="68F46ABDFF97126B33EC1B95FCB1FF6E" box="[722,882,151,177]" degrees="82.186692" direction="west" orientation="longitude" pageId="7" pageNumber="493" precision="1" value="-82.18669">82.186692 W</geoCoordinate>
);
<location id="081F5AA1FF97126B32B71B95FBECFF6E" LSID="urn:lsid:plazi:treatment:8569BD6CFF96126031A91C47FDC2F972:081F5AA1FF97126B32B71B95FBECFF6E" box="[905,1071,151,177]" county="Hernando County" latitude="28.707008" longLatPrecision="1" longitude="-89.39525" municipality="Ridge Manor" name="Citrus Springs" pageId="7" pageNumber="493" stateProvince="Florida">Citrus Springs</location>
,
<location id="081F5AA1FF97126B35001B95FB1DFF6E" LSID="urn:lsid:plazi:treatment:8569BD6CFF96126031A91C47FDC2F972:081F5AA1FF97126B35001B95FB1DFF6E" box="[1086,1246,151,177]" county="Hernando County" latitude="28.707008" longLatPrecision="1" longitude="-89.39525" municipality="Ridge Manor" name="Citrus County" pageId="7" pageNumber="493" stateProvince="Florida">Citrus County</location>
(
<geoCoordinate id="68F46ABDFF97126B35CE1B95FA5BFF6E" box="[1264,1432,151,177]" degrees="29.9786111" direction="north" orientation="latitude" pageId="7" pageNumber="493" precision="1" value="29.978611">29.9786111 N</geoCoordinate>
, –
<geoCoordinate id="68F46ABDFF97126B319B1BB9FE93FF0A" box="[165,336,187,213]" degrees="82.4327777" direction="west" orientation="longitude" pageId="7" pageNumber="493" precision="1" value="-82.43278">82.4327777 W</geoCoordinate>
); and
<location id="081F5AA1FF97126B30AD1BB9FDD6FF0A" LSID="urn:lsid:plazi:treatment:8569BD6CFF96126031A91C47FDC2F972:081F5AA1FF97126B30AD1BB9FDD6FF0A" box="[403,533,187,213]" county="Hernando County" latitude="28.707008" longLatPrecision="1" longitude="-89.39525" municipality="Ridge Manor" name="Gainesville" pageId="7" pageNumber="493" stateProvince="Florida">Gainesville</location>
,
<location id="081F5AA1FF97126B331F1BB9FD14FF0A" LSID="urn:lsid:plazi:treatment:8569BD6CFF96126031A91C47FDC2F972:081F5AA1FF97126B331F1BB9FD14FF0A" box="[545,727,187,213]" county="Hernando County" latitude="28.707008" longLatPrecision="1" longitude="-89.39525" municipality="Ridge Manor" name="Alachua County" pageId="7" pageNumber="493" stateProvince="Florida">Alachua County</location>
(
<geoCoordinate id="68F46ABDFF97126B33D81BB9FCBEFF0A" box="[742,893,187,213]" degrees="28.707009" direction="north" orientation="latitude" pageId="7" pageNumber="493" precision="1" value="28.707008">28.707009 N</geoCoordinate>
, –
<geoCoordinate id="68F46ABDFF97126B32A81BB8FBF1FF0A" box="[918,1074,186,213]" degrees="89.395251" direction="west" orientation="longitude" pageId="7" pageNumber="493" precision="1" value="-89.39525">89.395251 W</geoCoordinate>
), all in
<taxonomicName id="CAC077F9FF97126B35BB1BB9FAD3FF0A" authorityName="Chamberlin" authorityYear="1943" box="[1157,1296,187,213]" class="Diplopoda" family="Spirobolidae" genus="Narceus" kingdom="Animalia" order="Spirobolida" pageId="7" pageNumber="493" phylum="Arthropoda" rank="species" species="gordanus">
<emphasis id="3FB4D068FF97126B35BB1BB9FAD3FF0A" box="[1157,1296,187,213]" italics="true" pageId="7" pageNumber="493">N. gordanus</emphasis>
</taxonomicName>
</materialsCitation>
.
</paragraph>
<paragraph id="0D7F0C7AFF97126B31F91BDDFD88FEBA" blockId="7.[151,1437,150,429]" pageId="7" pageNumber="493">
<emphasis id="3FB4D068FF97126B31F91BDDFD8CFF27" bold="true" box="[199,591,222,249]" pageId="7" pageNumber="493">Type designation and deposition</emphasis>
.
<materialsCitation id="BDA80627FF97126B33601BDDFD14FEC2" ID-GBIF-Occurrence="3012561303" collectionCode="USDA" location="Beltsville" municipality="Nematode Collection" pageId="7" pageNumber="493" specimenCount="2" stateProvince="Maryland" typeStatus="holotype">
<typeStatus id="D27BB2D8FF97126B33601BDDFD05FF26" box="[606,710,223,249]" pageId="7" pageNumber="493" type="holotype">Holotype</typeStatus>
female (T-
<quantity id="CA38A19FFF97126B32791BDDFCB8FF26" box="[839,891,223,249]" metricMagnitude="5" metricUnit="kg" metricValue="7.08" pageId="7" pageNumber="493" unit="t" value="708.0">708t</quantity>
) and male
<typeStatus id="D27BB2D8FF97126B32C21BDDFB9FFF26" box="[1020,1116,223,249]" pageId="7" pageNumber="493" type="paratype">paratype</typeStatus>
(T-6937p), deposited in the
<collectionCode id="6BD194BFFF97126B31A91A01FF20FEC2" box="[151,227,259,285]" country="USA" httpUri="http://grbio.org/cool/e3b9-k2i3" name="United States Department of Agriculture" pageId="7" pageNumber="493">USDA</collectionCode>
<collectingMunicipality id="ED1B9600FF97126B31D21A01FE1CFEC2" box="[236,479,259,285]" pageId="7" pageNumber="493">Nematode Collection</collectingMunicipality>
,
<location id="081F5AA1FF97126B30D51A01FD9AFEC2" LSID="urn:lsid:plazi:treatment:8569BD6CFF96126031A91C47FDC2F972:081F5AA1FF97126B30D51A01FD9AFEC2" box="[491,601,259,285]" municipality="Nematode Collection" name="Beltsville" pageId="7" pageNumber="493" stateProvince="Maryland">Beltsville</location>
,
<collectingRegion id="CF04C298FF97126B33581A01FD14FEC2" box="[614,727,259,285]" country="United States of America" name="Maryland" pageId="7" pageNumber="493">Maryland</collectingRegion>
</materialsCitation>
.
<materialsCitation id="BDA80627FF97126B33DC1A01FD84FEBA" ID-GBIF-Occurrence="3012561304" collectorName="Control Numbers B" location="Tennessee" pageId="7" pageNumber="493" specimenCount="1" stateProvince="Tennessee">
Additional specimens deposited in the nematode collection in the Entomology and Plant Pathology Department, University of
<collectingRegion id="CF04C298FF97126B32471A25FC32FE9E" box="[889,1009,295,321]" country="United States of America" name="Tennessee" pageId="7" pageNumber="493">Tennessee</collectingRegion>
, Knoxville,
<collectingRegion id="CF04C298FF97126B35401A25FB30FE9E" box="[1150,1267,295,321]" country="United States of America" name="Tennessee" pageId="7" pageNumber="493">Tennessee</collectingRegion>
(University of
<collectingRegion id="CF04C298FF97126B31A91A49FECFFEBA" box="[151,268,331,357]" country="United States of America" name="Tennessee" pageId="7" pageNumber="493">Tennessee</collectingRegion>
<collectorName id="A03569ACFF97126B302D1A49FE30FEBA" box="[275,499,331,357]" pageId="7" pageNumber="493">Control Numbers B</collectorName>
1–B24)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="45DA5FF1FF97126531F91A6DFBC7F87D" lastPageId="9" lastPageNumber="495" pageId="7" pageNumber="493" type="etymology">
<paragraph id="0D7F0C7AFF97126B31F91A6DFB14FE72" blockId="7.[151,1437,150,429]" pageId="7" pageNumber="493">
<emphasis id="3FB4D068FF97126B31F91A6DFE8CFE56" bold="true" box="[199,335,367,393]" pageId="7" pageNumber="493">Etymology.</emphasis>
We are pleased to dedicate this new species to Dr. Robert J. Pivar, outstanding former graduate student in the Entomology and Plant Pathology Department, University of
<collectingRegion id="CF04C298FF97126B32E31A91FB96FE72" box="[989,1109,403,429]" country="United States of America" name="Tennessee" pageId="7" pageNumber="493">Tennessee</collectingRegion>
, Knoxville.
</paragraph>
<caption id="59BF5CF2FF97126B31A91FB3FD70FAEA" ID-DOI="http://doi.org/10.5281/zenodo.4416665" ID-Zenodo-Dep="4416665" httpUri="https://zenodo.org/record/4416665/files/figure.png" pageId="7" pageNumber="493" startId="7.[151,250,1201,1225]" targetBox="[156,1432,457,1173]" targetPageId="7">
<paragraph id="0D7F0C7AFF97126B31A91FB3FD70FAEA" blockId="7.[151,1437,1201,1333]" pageId="7" pageNumber="493">
<emphasis id="3FB4D068FF97126B31A91FB3FED0FB16" bold="true" box="[151,275,1201,1225]" pageId="7" pageNumber="493">FIGURE 2.</emphasis>
<taxonomicName id="CAC077F9FF97126B30241FB3FE4CFB16" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[282,399,1201,1225]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="7" pageNumber="493" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF97126B30241FB3FE4CFB16" box="[282,399,1201,1225]" italics="true" pageId="7" pageNumber="493">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF97126B30A81FB3FE0CFB16" bold="true" box="[406,463,1201,1225]" pageId="7" pageNumber="493">
<taxonomicNameLabel id="24876D13FF97126B30A81FB3FE0CFB16" box="[406,463,1201,1225]" pageId="7" pageNumber="493" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (
<bibRefCitation id="6951718BFF97126B34311FD7FEDFFACE" author="Spiridonov, S. E. &amp; Yushin, V. V." pageId="7" pageNumber="493" pagination="417 - 424" refId="ref20512" refString="Spiridonov, S. E. &amp; Yushin, V. V. (2000) Ultrastructure of the stoma in Heth mauriesi Adamson, 1982 (Rhigonematida: Hethidae). Nematology, 2, 417 - 424. https: // doi. org / 10.1163 / 156854100509277" type="journal article" year="2000">Spiridonov &amp; Yushin 2000</bibRefCitation>
). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm.
</paragraph>
</caption>
<paragraph id="0D7F0C7AFF97126B31F91E61FEABF8F2" blockId="7.[151,1437,1379,2018]" pageId="7" pageNumber="493">
<emphasis id="3FB4D068FF97126B31F91E61FEF3FAA2" bold="true" box="[199,304,1379,1405]" pageId="7" pageNumber="493">Females.</emphasis>
Measurements and ratios given in
<tableCitation id="404239C1FF97126B33EE1E61FCEFFAA2" box="[720,812,1379,1405]" captionStart="TABLE 2" captionStartId="8.[151,239,260,284]" captionTargetPageId="8" captionText="TABLE 2. Morphometric measurements of Heth pivari n. sp. females." httpUri="http://table.plazi.org/id/59BF5CF2FF98126431A91A06FC4FFEC3" pageId="7" pageNumber="493" tableUuid="59BF5CF2FF98126431A91A06FC4FFEC3">Table 2</tableCitation>
. Body cylindrical, white in life, robust, anterior head region broad; maximum diameter near mid-body, then tapering posteriorly and terminating in filiform tail (
<figureCitation id="95FB10FFFF97126B345A1E85FF7FFA1A" captionStart="FIGURE 2" captionStartId="7.[151,250,1201,1225]" captionTargetBox="[156,1432,457,1173]" captionTargetId="figure-135@7.[151,1436,453,1177]" captionTargetPageId="7" captionText="FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov &amp; Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm." figureDoi="http://doi.org/10.5281/zenodo.4416665" httpUri="https://zenodo.org/record/4416665/files/figure.png" pageId="7" pageNumber="493">Figs. 2A</figureCitation>
,
<figureCitation id="95FB10FFFF97126B31F61EA9FF2DFA1A" box="[200,238,1451,1477]" captionStart="FIGURE 7" captionStartId="14.[151,250,1613,1637]" captionTargetBox="[155,1432,185,1583]" captionTargetId="figure-18@14.[151,1436,181,1589]" captionTargetPageId="14" captionText="FIGURE 7. Heth pivari n. sp. A) Live female with single egg. B–E) Female enlargement and increasing numbers of eggs. The uterus is capacious but the vagina holds just one egg at a time (B, D). F) Female, dense trichomycete attachment in anterior region. G) Male, arthropod seta lodged in anteriormost region of intestine. H) Male, lateral chord (arrows). Scale bars: A, 500 µm; B–F, 250 µm; G, 100 µm; H, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416680" httpUri="https://zenodo.org/record/4416680/files/figure.png" pageId="7" pageNumber="493">7A</figureCitation>
). Differentiated lateral field absent. Annules
<quantity id="CA38A19FFF97126B33D31EA9FC9AFA19" box="[749,857,1451,1478]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="7" pageNumber="493" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 µm</quantity>
wide posterior to cervical collar, each annule from neck to anal region with numerous minute longitudinal striae (
<figureCitation id="95FB10FFFF97126B32731ECDFC65FA36" box="[845,934,1487,1513]" captionStart="FIGURE 3" captionStartId="9.[151,250,1497,1521]" captionTargetBox="[155,1429,349,1466]" captionTargetId="figure-113@9.[151,1436,345,1473]" captionTargetPageId="9" captionText="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="7" pageNumber="493">Fig. 3D</figureCitation>
); neck region finely annulated but with weak or no transverse striations (
<figureCitation id="95FB10FFFF97126B30F81EF1FDD1F9D2" box="[454,530,1523,1549]" captionStart="FIGURE 3" captionStartId="9.[151,250,1497,1521]" captionTargetBox="[155,1429,349,1466]" captionTargetId="figure-113@9.[151,1436,345,1473]" captionTargetPageId="9" captionText="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="7" pageNumber="493">Figs. 3</figureCitation>
A–C). Buccal cavity tubular, expanded anteriorly, much detail obscured by other head structures; anterior and posterior parts divided by circular ring of short, longitudinal striae; posterior part with extremely fine transverse striae about
<quantity id="CA38A19FFF97126B33781D39FD55F989" box="[582,662,1595,1622]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="493" unit="mm" value="0.5">0.5 µm</quantity>
apart (
<figureCitation id="95FB10FFFF97126B33D81D39FCA8F98A" box="[742,875,1595,1621]" captionStart="FIGURE 2" captionStartId="7.[151,250,1201,1225]" captionTargetBox="[156,1432,457,1173]" captionTargetId="figure-135@7.[151,1436,453,1177]" captionTargetPageId="7" captionText="FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov &amp; Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm." figureDoi="http://doi.org/10.5281/zenodo.4416665" httpUri="https://zenodo.org/record/4416665/files/figure.png" pageId="7" pageNumber="493">Figs. 2B, C</figureCitation>
). Amphid apertures circular,
<quantity id="CA38A19FFF97126B35881D39FACDF989" box="[1206,1294,1595,1622]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="7" pageNumber="493" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 µm</quantity>
in diameter, located on convex lateral aspect of each pseudolabium (
<figureCitation id="95FB10FFFF97126B32361D5DFC90F9A6" box="[776,851,1631,1657]" captionStart="FIGURE 3" captionStartId="9.[151,250,1497,1521]" captionTargetBox="[155,1429,349,1466]" captionTargetId="figure-113@9.[151,1436,345,1473]" captionTargetPageId="9" captionText="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="7" pageNumber="493">Figs. 3</figureCitation>
A–C). Pseudolabia rectangular in dorso-lateral view (
<figureCitation id="95FB10FFFF97126B319E1D81FF3BF942" box="[160,248,1667,1693]" captionStart="FIGURE 3" captionStartId="9.[151,250,1497,1521]" captionTargetBox="[155,1429,349,1466]" captionTargetId="figure-113@9.[151,1436,345,1473]" captionTargetPageId="9" captionText="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="7" pageNumber="493">Fig. 3A</figureCitation>
), rounded on outer margin in dorsal view (
<figureCitation id="95FB10FFFF97126B33F11D81FCDAF942" box="[719,793,1667,1693]" captionStart="FIGURE 3" captionStartId="9.[151,250,1497,1521]" captionTargetBox="[155,1429,349,1466]" captionTargetId="figure-113@9.[151,1436,345,1473]" captionTargetPageId="9" captionText="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="7" pageNumber="493">Figs. 3</figureCitation>
A-C), ornamented with combs and spines; small pectinate combs with 2–3 µm-long bristles on interior lateral, median and cleft margins, and larger combs with 5–7 µm-long bristles continuing along dorsoventral aspect of each pseudolabium; dorsal and ventral bristles rounded distally; length of bristles on each margin approximately equal (
<figureCitation id="95FB10FFFF97126B33C41DEDFC91F8D6" box="[762,850,1775,1801]" captionStart="FIGURE 3" captionStartId="9.[151,250,1497,1521]" captionTargetBox="[155,1429,349,1466]" captionTargetId="figure-113@9.[151,1436,345,1473]" captionTargetPageId="9" captionText="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="7" pageNumber="493">Fig. 3C</figureCitation>
). Anterior stomal region often filled with amorphous material (
<figureCitation id="95FB10FFFF97126B303C1C11FE99F8F2" box="[258,346,1811,1837]" captionStart="FIGURE 3" captionStartId="9.[151,250,1497,1521]" captionTargetBox="[155,1429,349,1466]" captionTargetId="figure-113@9.[151,1436,345,1473]" captionTargetPageId="9" captionText="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="7" pageNumber="493">Fig. 3B</figureCitation>
).
</paragraph>
<paragraph id="0D7F0C7AFF97126431F91C35FC8BFF09" blockId="7.[151,1437,1379,2018]" lastBlockId="8.[151,1437,151,214]" lastPageId="8" lastPageNumber="494" pageId="7" pageNumber="493">
Neck region with four folds anterior to cervical collar, with second, third and fourth folds each containing one smooth, knob-like cervical sensory papilla per quadrant (
<figureCitation id="95FB10FFFF97126B322B1C59FC5AF8AA" box="[789,921,1883,1909]" captionStart="FIGURE 3" captionStartId="9.[151,250,1497,1521]" captionTargetBox="[155,1429,349,1466]" captionTargetId="figure-113@9.[151,1436,345,1473]" captionTargetPageId="9" captionText="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="7" pageNumber="493">Figs. 3B, C</figureCitation>
); papillae about
<quantity id="CA38A19FFF97126B356D1C59FB69F8A9" box="[1107,1194,1883,1910]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="7" pageNumber="493" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 µm</quantity>
in diameter. Cervical collar with about 72 spines,
<quantity id="CA38A19FFF97126B30E61C7DFDECF845" box="[472,559,1919,1946]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="7" pageNumber="493" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 µm</quantity>
long; collar interrupted laterally on each side by shield, edge spines often bi- or trifurcate (
<figureCitation id="95FB10FFFF97126B30111CA1FE7AF861" box="[303,441,1955,1982]" captionStart="FIGURE 3" captionStartId="9.[151,250,1497,1521]" captionTargetBox="[155,1429,349,1466]" captionTargetId="figure-113@9.[151,1436,345,1473]" captionTargetPageId="9" captionText="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="7" pageNumber="493">Figs. 3A, D</figureCitation>
); shield wider than long (
<figureCitation id="95FB10FFFF97126B33DF1CA1FC84F862" box="[737,839,1955,1981]" captionStart="FIGURE 2" captionStartId="7.[151,250,1201,1225]" captionTargetBox="[156,1432,457,1173]" captionTargetId="figure-135@7.[151,1436,453,1177]" captionTargetPageId="7" captionText="FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov &amp; Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm." figureDoi="http://doi.org/10.5281/zenodo.4416665" httpUri="https://zenodo.org/record/4416665/files/figure.png" pageId="7" pageNumber="493">Figs. 2D</figureCitation>
,
<figureCitation id="95FB10FFFF97126B326A1CA1FCBBF862" box="[852,888,1955,1981]" captionStart="FIGURE 3" captionStartId="9.[151,250,1497,1521]" captionTargetBox="[155,1429,349,1466]" captionTargetId="figure-113@9.[151,1436,345,1473]" captionTargetPageId="9" captionText="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="7" pageNumber="493">3A</figureCitation>
,
<figureCitation id="95FB10FFFF97126B32B81CA1FC6FF861" box="[902,940,1955,1982]" captionStart="FIGURE 6" captionStartId="13.[151,250,1904,1928]" captionTargetBox="[154,1433,672,1879]" captionTargetId="figure-214@13.[151,1435,669,1881]" captionTargetPageId="13" captionText="FIGURE 6. DIC micrographs of Heth pivari n. sp., female and male structures. A) Lateral ornamentation in anterior region of female. B) Male, lateral somal extensions of posterior arcade surrounding anterior esophagus. C) Female, secretory-excretory pore (arrow). D. Female, somatic papillae anterior to vulva. E) Fusiform to comet-shaped sperm in male. All scale bars 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416676" httpUri="https://zenodo.org/record/4416676/files/figure.png" pageId="7" pageNumber="493">6A</figureCitation>
). Two lateral pairs of large, acute spines in tandem,
<quantity id="CA38A19FFF97126B31C41CC5FEACF83D" box="[250,367,1991,2018]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="7" pageNumber="493" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 µm</quantity>
long; anterior pair sometimes connected by a shallow ridge, posterior pair less frequently so (
<figureCitation id="95FB10FFFF98126431A11B95FEE7FF6E" box="[159,292,151,177]" captionStart="FIGURE 3" captionStartId="9.[151,250,1497,1521]" captionTargetBox="[155,1429,349,1466]" captionTargetId="figure-113@9.[151,1436,345,1473]" captionTargetPageId="9" captionText="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="8" pageNumber="494">Figs. 3A, B</figureCitation>
). Annulation between each pair of spines irregular and areolated. Knob-like somatic papillae scattered along length of body, denser anterior to vulva (
<figureCitation id="95FB10FFFF98126433981BB9FCC9FF0A" box="[678,778,187,213]" captionStart="FIGURE 3" captionStartId="9.[151,250,1497,1521]" captionTargetBox="[155,1429,349,1466]" captionTargetId="figure-113@9.[151,1436,345,1473]" captionTargetPageId="9" captionText="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="8" pageNumber="494">Figs. 3D</figureCitation>
,
<figureCitation id="95FB10FFFF98126432281BB9FCFFFF09" box="[790,828,187,214]" captionStart="FIGURE 6" captionStartId="13.[151,250,1904,1928]" captionTargetBox="[154,1433,672,1879]" captionTargetId="figure-214@13.[151,1435,669,1881]" captionTargetPageId="13" captionText="FIGURE 6. DIC micrographs of Heth pivari n. sp., female and male structures. A) Lateral ornamentation in anterior region of female. B) Male, lateral somal extensions of posterior arcade surrounding anterior esophagus. C) Female, secretory-excretory pore (arrow). D. Female, somatic papillae anterior to vulva. E) Fusiform to comet-shaped sperm in male. All scale bars 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416676" httpUri="https://zenodo.org/record/4416676/files/figure.png" pageId="8" pageNumber="494">6D</figureCitation>
).
</paragraph>
<caption id="59BF5CF2FF98126431A91A06FC4FFEC3" ID-Table-UUID="59BF5CF2FF98126431A91A06FC4FFEC3" box="[151,908,260,285]" httpUri="http://table.plazi.org/id/59BF5CF2FF98126431A91A06FC4FFEC3" pageId="8" pageNumber="494" targetBox="[166,1349,300,1023]" targetIsTable="true" targetPageId="8">
<paragraph id="0D7F0C7AFF98126431A91A06FC4FFEC3" blockId="8.[151,908,260,285]" box="[151,908,260,285]" pageId="8" pageNumber="494">
<emphasis id="3FB4D068FF98126431A91A06FEC9FEC3" bold="true" box="[151,266,260,284]" pageId="8" pageNumber="494">TABLE 2.</emphasis>
Morphometric measurements of
<taxonomicName id="CAC077F9FF98126433501A06FD2BFEC3" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[622,744,260,284]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="8" pageNumber="494" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF98126433501A06FD2BFEC3" box="[622,744,260,284]" italics="true" pageId="8" pageNumber="494">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF98126433D11A07FCE8FEC2" bold="true" box="[751,811,261,285]" pageId="8" pageNumber="494">
<taxonomicNameLabel id="24876D13FF98126433D11A07FCE8FEC2" box="[751,811,261,285]" pageId="8" pageNumber="494" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
females.
</paragraph>
</caption>
<paragraph id="0D7F0C7AFF98126433171A2EFAFAFC20" pageId="8" pageNumber="494">
<table id="7FC0FEDAFF98ED9331981A2EFA86FC20" box="[166,1349,300,1023]" gridcols="6" gridrows="19" pageId="8" pageNumber="494">
<tr id="B3F00E38FF98ED9331981A2EFA86FE9B" box="[166,1349,300,324]" gridrow="0" pageId="8" pageNumber="494" rowspan-0="1" rowspan-4="1" rowspan-5="1">
<th id="F0216744FF98ED9333171A2EFD44FE9B" box="[553,647,300,324]" gridcol="1" gridrow="0" pageId="8" pageNumber="494">Holotype</th>
<th id="F0216744FF98ED9333DC1A2EFC2BFE9B" box="[738,1000,300,324]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="8" pageNumber="494">
Paratype females (
<emphasis id="3FB4D068FF98126432A11A2FFC6FFE9B" box="[927,940,301,324]" italics="true" pageId="8" pageNumber="494">n</emphasis>
= 71)
</th>
</tr>
<tr id="B3F00E38FF98ED9331981A52FA86FEB4" box="[166,1349,336,363]" gridrow="1" pageId="8" pageNumber="494" rowspan-0="1">
<td id="F0216744FF98ED9333171A52FD44FEB4" box="[553,647,336,363]" gridcol="1" gridrow="1" pageId="8" pageNumber="494">female</td>
<td id="F0216744FF98ED9333DC1A52FCDFFEB4" box="[738,796,336,363]" gridcol="2" gridrow="1" pageId="8" pageNumber="494">Mean</td>
<td id="F0216744FF98ED9332BF1A52FC2BFEB4" box="[897,1000,336,363]" gridcol="3" gridrow="1" pageId="8" pageNumber="494">SD</td>
<td id="F0216744FF98ED93351E1A52FB5CFEB4" box="[1056,1183,336,363]" gridcol="4" gridrow="1" pageId="8" pageNumber="494">Range</td>
<td id="F0216744FF98ED9334271A52FA86FEB4" box="[1305,1349,336,363]" gridcol="5" gridrow="1" pageId="8" pageNumber="494">CV</td>
</tr>
<tr id="B3F00E38FF98ED9331981A78FA86FE4D" box="[166,1349,378,402]" gridrow="2" pageId="8" pageNumber="494" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="F0216744FF98ED9331981A78FE4DFE4D" box="[166,398,378,402]" gridcol="0" gridrow="2" pageId="8" pageNumber="494">Measurements (µm)</th>
</tr>
<tr id="B3F00E38FF98ED9331981AA2FA86FE67" box="[166,1349,416,440]" gridrow="3" pageId="8" pageNumber="494">
<th id="F0216744FF98ED9331981AA2FE4DFE67" box="[166,398,416,440]" gridcol="0" gridrow="3" pageId="8" pageNumber="494">Length</th>
<td id="F0216744FF98ED9333171AA2FD44FE67" box="[553,647,416,440]" gridcol="1" gridrow="3" pageId="8" pageNumber="494">2,800</td>
<td id="F0216744FF98ED9333DC1AA2FCDFFE67" box="[738,796,416,440]" gridcol="2" gridrow="3" pageId="8" pageNumber="494">3,011</td>
<td id="F0216744FF98ED9332BF1AA2FC2BFE67" box="[897,1000,416,440]" gridcol="3" gridrow="3" pageId="8" pageNumber="494">406</td>
<td id="F0216744FF98ED93351E1AA2FB5CFE67" box="[1056,1183,416,440]" gridcol="4" gridrow="3" pageId="8" pageNumber="494">2,190–4,483</td>
<td id="F0216744FF98ED9334271AA2FA86FE67" box="[1305,1349,416,440]" gridcol="5" gridrow="3" pageId="8" pageNumber="494">13.5</td>
</tr>
<tr id="B3F00E38FF98ED9331981AC5FA86FE00" box="[166,1349,455,479]" gridrow="4" pageId="8" pageNumber="494">
<th id="F0216744FF98ED9331981AC5FE4DFE00" box="[166,398,455,479]" gridcol="0" gridrow="4" pageId="8" pageNumber="494">Maximum width</th>
<td id="F0216744FF98ED9333171AC5FD44FE00" box="[553,647,455,479]" gridcol="1" gridrow="4" pageId="8" pageNumber="494">182</td>
<td id="F0216744FF98ED9333DC1AC5FCDFFE00" box="[738,796,455,479]" gridcol="2" gridrow="4" pageId="8" pageNumber="494">191</td>
<td id="F0216744FF98ED9332BF1AC5FC2BFE00" box="[897,1000,455,479]" gridcol="3" gridrow="4" pageId="8" pageNumber="494">52</td>
<td id="F0216744FF98ED93351E1AC5FB5CFE00" box="[1056,1183,455,479]" gridcol="4" gridrow="4" pageId="8" pageNumber="494">131–380</td>
<td id="F0216744FF98ED9334271AC5FA86FE00" box="[1305,1349,455,479]" gridcol="5" gridrow="4" pageId="8" pageNumber="494">27.2</td>
</tr>
<tr id="B3F00E38FF98ED9331981AECFA86FDD9" box="[166,1349,494,518]" gridrow="5" pageId="8" pageNumber="494">
<th id="F0216744FF98ED9331981AECFE4DFDD9" box="[166,398,494,518]" gridcol="0" gridrow="5" pageId="8" pageNumber="494">Esophagus length</th>
<td id="F0216744FF98ED9333171AECFD44FDD9" box="[553,647,494,518]" gridcol="1" gridrow="5" pageId="8" pageNumber="494">382</td>
<td id="F0216744FF98ED9333DC1AECFCDFFDD9" box="[738,796,494,518]" gridcol="2" gridrow="5" pageId="8" pageNumber="494">371</td>
<td id="F0216744FF98ED9332BF1AECFC2BFDD9" box="[897,1000,494,518]" gridcol="3" gridrow="5" pageId="8" pageNumber="494">17</td>
<td id="F0216744FF98ED93351E1AECFB5CFDD9" box="[1056,1183,494,518]" gridcol="4" gridrow="5" pageId="8" pageNumber="494">327–407</td>
<td id="F0216744FF98ED9334271AECFA86FDD9" box="[1305,1349,494,518]" gridcol="5" gridrow="5" pageId="8" pageNumber="494">4.6</td>
</tr>
<tr id="B3F00E38FF98ED9331981916FA86FDF3" box="[166,1349,532,556]" gridrow="6" pageId="8" pageNumber="494">
<th id="F0216744FF98ED9331981916FE4DFDF3" box="[166,398,532,556]" gridcol="0" gridrow="6" pageId="8" pageNumber="494">Basal bulb length</th>
<td id="F0216744FF98ED9333171916FD44FDF3" box="[553,647,532,556]" gridcol="1" gridrow="6" pageId="8" pageNumber="494">140</td>
<td id="F0216744FF98ED9333DC1916FCDFFDF3" box="[738,796,532,556]" gridcol="2" gridrow="6" pageId="8" pageNumber="494">148</td>
<td id="F0216744FF98ED9332BF1916FC2BFDF3" box="[897,1000,532,556]" gridcol="3" gridrow="6" pageId="8" pageNumber="494">7.1</td>
<td id="F0216744FF98ED93351E1916FB5CFDF3" box="[1056,1183,532,556]" gridcol="4" gridrow="6" pageId="8" pageNumber="494">135–176</td>
<td id="F0216744FF98ED9334271916FA86FDF3" box="[1305,1349,532,556]" gridcol="5" gridrow="6" pageId="8" pageNumber="494">4.8</td>
</tr>
<tr id="B3F00E38FF98ED9331981939FA86FD8B" box="[166,1349,571,596]" gridrow="7" pageId="8" pageNumber="494">
<th id="F0216744FF98ED9331981939FE4DFD8B" box="[166,398,571,596]" gridcol="0" gridrow="7" pageId="8" pageNumber="494">Egg length</th>
<td id="F0216744FF98ED9333171939FD44FD8B" box="[553,647,571,596]" gridcol="1" gridrow="7" pageId="8" pageNumber="494">210</td>
<td id="F0216744FF98ED9333DC1939FCDFFD8B" box="[738,796,571,596]" gridcol="2" gridrow="7" pageId="8" pageNumber="494">209</td>
<td id="F0216744FF98ED9332BF1939FC2BFD8B" box="[897,1000,571,596]" gridcol="3" gridrow="7" pageId="8" pageNumber="494">11</td>
<td id="F0216744FF98ED93351E1939FB5CFD8B" box="[1056,1183,571,596]" gridcol="4" gridrow="7" pageId="8" pageNumber="494">180–234</td>
<td id="F0216744FF98ED9334271939FA86FD8B" box="[1305,1349,571,596]" gridcol="5" gridrow="7" pageId="8" pageNumber="494">5.3</td>
</tr>
<tr id="B3F00E38FF98ED9331981960FA86FDA5" box="[166,1349,610,634]" gridrow="8" pageId="8" pageNumber="494">
<th id="F0216744FF98ED9331981960FE4DFDA5" box="[166,398,610,634]" gridcol="0" gridrow="8" pageId="8" pageNumber="494">Egg width</th>
<td id="F0216744FF98ED9333171960FD44FDA5" box="[553,647,610,634]" gridcol="1" gridrow="8" pageId="8" pageNumber="494">111</td>
<td id="F0216744FF98ED9333DC1960FCDFFDA5" box="[738,796,610,634]" gridcol="2" gridrow="8" pageId="8" pageNumber="494">94</td>
<td id="F0216744FF98ED9332BF1960FC2BFDA5" box="[897,1000,610,634]" gridcol="3" gridrow="8" pageId="8" pageNumber="494">10.7</td>
<td id="F0216744FF98ED93351E1960FB5CFDA5" box="[1056,1183,610,634]" gridcol="4" gridrow="8" pageId="8" pageNumber="494">59–116</td>
<td id="F0216744FF98ED9334271960FA86FDA5" box="[1305,1349,610,634]" gridcol="5" gridrow="8" pageId="8" pageNumber="494">11.4</td>
</tr>
<tr id="B3F00E38FF98ED933198198AFA86FD7E" box="[166,1349,648,673]" gridrow="9" pageId="8" pageNumber="494">
<th id="F0216744FF98ED933198198AFE4DFD7E" box="[166,398,648,673]" gridcol="0" gridrow="9" pageId="8" pageNumber="494">Head to excretory pore</th>
<td id="F0216744FF98ED933317198AFD44FD7E" box="[553,647,648,673]" gridcol="1" gridrow="9" pageId="8" pageNumber="494">167</td>
<td id="F0216744FF98ED9333DC198AFCDFFD7E" box="[738,796,648,673]" gridcol="2" gridrow="9" pageId="8" pageNumber="494">195</td>
<td id="F0216744FF98ED9332BF198AFC2BFD7E" box="[897,1000,648,673]" gridcol="3" gridrow="9" pageId="8" pageNumber="494">20</td>
<td id="F0216744FF98ED93351E198AFB5CFD7E" box="[1056,1183,648,673]" gridcol="4" gridrow="9" pageId="8" pageNumber="494">167–234</td>
<td id="F0216744FF98ED933427198AFA86FD7E" box="[1305,1349,648,673]" gridcol="5" gridrow="9" pageId="8" pageNumber="494">10.3</td>
</tr>
<tr id="B3F00E38FF98ED93319819B3FA86FD16" box="[166,1349,689,713]" gridrow="10" pageId="8" pageNumber="494">
<th id="F0216744FF98ED93319819B3FE4DFD16" box="[166,398,689,713]" gridcol="0" gridrow="10" pageId="8" pageNumber="494">Head to vulva</th>
<td id="F0216744FF98ED93331719B3FD44FD16" box="[553,647,689,713]" gridcol="1" gridrow="10" pageId="8" pageNumber="494">2,189</td>
<td id="F0216744FF98ED9333DC19B3FCDFFD16" box="[738,796,689,713]" gridcol="2" gridrow="10" pageId="8" pageNumber="494">2,337</td>
<td id="F0216744FF98ED9332BF19B3FC2BFD16" box="[897,1000,689,713]" gridcol="3" gridrow="10" pageId="8" pageNumber="494">326</td>
<td id="F0216744FF98ED93351E19B3FB5CFD16" box="[1056,1183,689,713]" gridcol="4" gridrow="10" pageId="8" pageNumber="494">1,664–3,644</td>
<td id="F0216744FF98ED93342719B3FA86FD16" box="[1305,1349,689,713]" gridcol="5" gridrow="10" pageId="8" pageNumber="494">13.9</td>
</tr>
<tr id="B3F00E38FF98ED93319819DAFA86FD2F" box="[166,1349,728,752]" gridrow="11" pageId="8" pageNumber="494">
<th id="F0216744FF98ED93319819DAFE4DFD2F" box="[166,398,728,752]" gridcol="0" gridrow="11" pageId="8" pageNumber="494">Head to anus</th>
<td id="F0216744FF98ED93331719DAFD44FD2F" box="[553,647,728,752]" gridcol="1" gridrow="11" pageId="8" pageNumber="494">2,394</td>
<td id="F0216744FF98ED9333DC19DAFCDFFD2F" box="[738,796,728,752]" gridcol="2" gridrow="11" pageId="8" pageNumber="494">2,585</td>
<td id="F0216744FF98ED9332BF19DAFC2BFD2F" box="[897,1000,728,752]" gridcol="3" gridrow="11" pageId="8" pageNumber="494">370</td>
<td id="F0216744FF98ED93351E19DAFB5CFD2F" box="[1056,1183,728,752]" gridcol="4" gridrow="11" pageId="8" pageNumber="494">1,743–3,938</td>
<td id="F0216744FF98ED93342719DAFA86FD2F" box="[1305,1349,728,752]" gridcol="5" gridrow="11" pageId="8" pageNumber="494">14.3</td>
</tr>
<tr id="B3F00E38FF98ED93319819FCFA86FCC8" box="[166,1349,766,791]" gridrow="12" pageId="8" pageNumber="494">
<th id="F0216744FF98ED93319819FCFE4DFCC8" box="[166,398,766,791]" gridcol="0" gridrow="12" pageId="8" pageNumber="494">Tail length</th>
<td id="F0216744FF98ED93331719FCFD44FCC8" box="[553,647,766,791]" gridcol="1" gridrow="12" pageId="8" pageNumber="494">406</td>
<td id="F0216744FF98ED9333DC19FCFCDFFCC8" box="[738,796,766,791]" gridcol="2" gridrow="12" pageId="8" pageNumber="494">426</td>
<td id="F0216744FF98ED9332BF19FCFC2BFCC8" box="[897,1000,766,791]" gridcol="3" gridrow="12" pageId="8" pageNumber="494">50.2</td>
<td id="F0216744FF98ED93351E19FCFB5CFCC8" box="[1056,1183,766,791]" gridcol="4" gridrow="12" pageId="8" pageNumber="494">318–545</td>
<td id="F0216744FF98ED93342719FCFA86FCC8" box="[1305,1349,766,791]" gridcol="5" gridrow="12" pageId="8" pageNumber="494">11.8</td>
</tr>
<tr id="B3F00E38FF98ED9331981827FA86FCE2" box="[166,1349,805,829]" gridrow="13" pageId="8" pageNumber="494" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="F0216744FF98ED9331981827FE4DFCE2" box="[166,398,805,829]" gridcol="0" gridrow="13" pageId="8" pageNumber="494">Ratios</th>
</tr>
<tr id="B3F00E38FF98ED933198184EFA86FCBB" box="[166,1349,844,868]" gridrow="14" pageId="8" pageNumber="494">
<th id="F0216744FF98ED933198184EFE4DFCBB" box="[166,398,844,868]" gridcol="0" gridrow="14" pageId="8" pageNumber="494">a</th>
<td id="F0216744FF98ED933317184EFD44FCBB" box="[553,647,844,868]" gridcol="1" gridrow="14" pageId="8" pageNumber="494">15.4</td>
<td id="F0216744FF98ED9333DC184EFCDFFCBB" box="[738,796,844,868]" gridcol="2" gridrow="14" pageId="8" pageNumber="494">16.3</td>
<td id="F0216744FF98ED9332BF184EFC2BFCBB" box="[897,1000,844,868]" gridcol="3" gridrow="14" pageId="8" pageNumber="494">2</td>
<td id="F0216744FF98ED93351E184EFB5CFCBB" box="[1056,1183,844,868]" gridcol="4" gridrow="14" pageId="8" pageNumber="494">10.9–21.7</td>
<td id="F0216744FF98ED933427184EFA86FCBB" box="[1305,1349,844,868]" gridcol="5" gridrow="14" pageId="8" pageNumber="494">13.5</td>
</tr>
<tr id="B3F00E38FF98ED9331981871FA86FC54" box="[166,1349,883,907]" gridrow="15" pageId="8" pageNumber="494">
<th id="F0216744FF98ED9331981871FE4DFC54" box="[166,398,883,907]" gridcol="0" gridrow="15" pageId="8" pageNumber="494">b</th>
<td id="F0216744FF98ED9333171871FD44FC54" box="[553,647,883,907]" gridcol="1" gridrow="15" pageId="8" pageNumber="494">5.4</td>
<td id="F0216744FF98ED9333DC1871FCDFFC54" box="[738,796,883,907]" gridcol="2" gridrow="15" pageId="8" pageNumber="494">5.8</td>
<td id="F0216744FF98ED9332BF1871FC2BFC54" box="[897,1000,883,907]" gridcol="3" gridrow="15" pageId="8" pageNumber="494">0.74</td>
<td id="F0216744FF98ED93351E1871FB5CFC54" box="[1056,1183,883,907]" gridcol="4" gridrow="15" pageId="8" pageNumber="494">4.4–8</td>
<td id="F0216744FF98ED9334271871FA86FC54" box="[1305,1349,883,907]" gridcol="5" gridrow="15" pageId="8" pageNumber="494">27.2</td>
</tr>
<tr id="B3F00E38FF98ED933198189BFA86FC6D" box="[166,1349,921,946]" gridrow="16" pageId="8" pageNumber="494">
<th id="F0216744FF98ED933198189BFE4DFC6D" box="[166,398,921,946]" gridcol="0" gridrow="16" pageId="8" pageNumber="494">c</th>
<td id="F0216744FF98ED933317189BFD44FC6D" box="[553,647,921,946]" gridcol="1" gridrow="16" pageId="8" pageNumber="494">6.9</td>
<td id="F0216744FF98ED9333DC189BFCDFFC6D" box="[738,796,921,946]" gridcol="2" gridrow="16" pageId="8" pageNumber="494">7.1</td>
<td id="F0216744FF98ED9332BF189BFC2BFC6D" box="[897,1000,921,946]" gridcol="3" gridrow="16" pageId="8" pageNumber="494">0.64</td>
<td id="F0216744FF98ED93351E189BFB5CFC6D" box="[1056,1183,921,946]" gridcol="4" gridrow="16" pageId="8" pageNumber="494">4.8–9.4</td>
<td id="F0216744FF98ED933427189BFA86FC6D" box="[1305,1349,921,946]" gridcol="5" gridrow="16" pageId="8" pageNumber="494">4.6</td>
</tr>
<tr id="B3F00E38FF98ED93319818C2FA86FC07" box="[166,1349,960,984]" gridrow="17" pageId="8" pageNumber="494">
<th id="F0216744FF98ED93319818C2FE4DFC07" box="[166,398,960,984]" gridcol="0" gridrow="17" pageId="8" pageNumber="494">V (%)</th>
<td id="F0216744FF98ED93331718C2FD44FC07" box="[553,647,960,984]" gridcol="1" gridrow="17" pageId="8" pageNumber="494">78.2</td>
<td id="F0216744FF98ED9333DC18C2FCDFFC07" box="[738,796,960,984]" gridcol="2" gridrow="17" pageId="8" pageNumber="494">78</td>
<td id="F0216744FF98ED9332BF18C2FC2BFC07" box="[897,1000,960,984]" gridcol="3" gridrow="17" pageId="8" pageNumber="494">1.6</td>
<td id="F0216744FF98ED93351E18C2FB5CFC07" box="[1056,1183,960,984]" gridcol="4" gridrow="17" pageId="8" pageNumber="494">73–83</td>
<td id="F0216744FF98ED93342718C2FA86FC07" box="[1305,1349,960,984]" gridcol="5" gridrow="17" pageId="8" pageNumber="494">4.8</td>
</tr>
<tr id="B3F00E38FF98ED93319818E5FA86FC20" box="[166,1349,999,1023]" gridrow="18" pageId="8" pageNumber="494">
<th id="F0216744FF98ED93319818E5FE4DFC20" box="[166,398,999,1023]" gridcol="0" gridrow="18" pageId="8" pageNumber="494">V’</th>
<td id="F0216744FF98ED93331718E5FD44FC20" box="[553,647,999,1023]" gridcol="1" gridrow="18" pageId="8" pageNumber="494">0.91</td>
<td id="F0216744FF98ED9333DC18E5FCDFFC20" box="[738,796,999,1023]" gridcol="2" gridrow="18" pageId="8" pageNumber="494">0.91</td>
<td id="F0216744FF98ED9332BF18E5FC2BFC20" box="[897,1000,999,1023]" gridcol="3" gridrow="18" pageId="8" pageNumber="494">0.02</td>
<td id="F0216744FF98ED93351E18E5FB5CFC20" box="[1056,1183,999,1023]" gridcol="4" gridrow="18" pageId="8" pageNumber="494">0.87–0.96</td>
<td id="F0216744FF98ED93342718E5FA86FC20" box="[1305,1349,999,1023]" gridcol="5" gridrow="18" pageId="8" pageNumber="494">5.3</td>
</tr>
</table>
</paragraph>
<paragraph id="0D7F0C7AFF98126431F91F2CFC2AFBB3" blockId="8.[151,1437,1070,2032]" pageId="8" pageNumber="494">
Neck region dorsally and ventrally with retractor muscles attached to body wall at each major fold, integrated with strong longitudinal muscle associated with large nucleus (
<figureCitation id="95FB10FFFF98126432661F50FC1FFBB3" box="[856,988,1106,1132]" captionStart="FIGURE 3" captionStartId="9.[151,250,1497,1521]" captionTargetBox="[155,1429,349,1466]" captionTargetId="figure-113@9.[151,1436,345,1473]" captionTargetPageId="9" captionText="FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="8" pageNumber="494">Figs. 3B, C</figureCitation>
).
</paragraph>
<paragraph id="0D7F0C7AFF98126431F91F74FDDBFAB7" blockId="8.[151,1437,1070,2032]" pageId="8" pageNumber="494">
Two-part esophagus consisting of procorpus and basal bulb with grinding valve, surrounded anteriorly by six prominent, uninucleate, amber-colored glands (
<figureCitation id="95FB10FFFF981264339B1F98FD3FFB6B" box="[677,764,1178,1204]" captionStart="FIGURE 2" captionStartId="7.[151,250,1201,1225]" captionTargetBox="[156,1432,457,1173]" captionTargetId="figure-135@7.[151,1436,453,1177]" captionTargetPageId="7" captionText="FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov &amp; Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm." figureDoi="http://doi.org/10.5281/zenodo.4416665" httpUri="https://zenodo.org/record/4416665/files/figure.png" pageId="8" pageNumber="494">Fig. 2A</figureCitation>
) identical in number and arrangement to those of male (
<figureCitation id="95FB10FFFF981264345A1F98FF1FFB07" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1920]" captionTargetBox="[160,1426,354,1861]" captionTargetId="figure-116@11.[151,1436,345,1872]" captionTargetPageId="11" captionText="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="8" pageNumber="494">Figs. 4A, B</figureCitation>
,
<figureCitation id="95FB10FFFF98126431D91FBCFECBFB06" box="[231,264,1214,1241]" captionStart="FIGURE 6" captionStartId="13.[151,250,1904,1928]" captionTargetBox="[154,1433,672,1879]" captionTargetId="figure-214@13.[151,1435,669,1881]" captionTargetPageId="13" captionText="FIGURE 6. DIC micrographs of Heth pivari n. sp., female and male structures. A) Lateral ornamentation in anterior region of female. B) Male, lateral somal extensions of posterior arcade surrounding anterior esophagus. C) Female, secretory-excretory pore (arrow). D. Female, somatic papillae anterior to vulva. E) Fusiform to comet-shaped sperm in male. All scale bars 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416676" httpUri="https://zenodo.org/record/4416676/files/figure.png" pageId="8" pageNumber="494">6B</figureCitation>
) and one or more small colorless glands (
<figureCitation id="95FB10FFFF98126433E61FBCFCECFB07" box="[728,815,1214,1240]" captionStart="FIGURE 2" captionStartId="7.[151,250,1201,1225]" captionTargetBox="[156,1432,457,1173]" captionTargetId="figure-135@7.[151,1436,453,1177]" captionTargetPageId="7" captionText="FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov &amp; Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm." figureDoi="http://doi.org/10.5281/zenodo.4416665" httpUri="https://zenodo.org/record/4416665/files/figure.png" pageId="8" pageNumber="494">Fig. 2B</figureCitation>
). Secretory-excretory (SE) pore minute, inconspicuous,
<quantity id="CA38A19FFF98126431A91FE0FEE4FB23" box="[151,295,1250,1276]" metricMagnitude="-1" metricUnit="m" metricValue="2.005" metricValueMax="2.34" metricValueMin="1.67" pageId="8" pageNumber="494" unit="mm" value="200.5" valueMax="234.0" valueMin="167.0">167–234 µm</quantity>
from anterior head (
<figureCitation id="95FB10FFFF981264332F1FE0FDB5FB23" box="[529,630,1250,1276]" captionStart="FIGURE 2" captionStartId="7.[151,250,1201,1225]" captionTargetBox="[156,1432,457,1173]" captionTargetId="figure-135@7.[151,1436,453,1177]" captionTargetPageId="7" captionText="FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov &amp; Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm." figureDoi="http://doi.org/10.5281/zenodo.4416665" httpUri="https://zenodo.org/record/4416665/files/figure.png" pageId="8" pageNumber="494">Figs. 2A</figureCitation>
,
<figureCitation id="95FB10FFFF98126433BC1FE0FD65FB22" box="[642,678,1250,1277]" captionStart="FIGURE 6" captionStartId="13.[151,250,1904,1928]" captionTargetBox="[154,1433,672,1879]" captionTargetId="figure-214@13.[151,1435,669,1881]" captionTargetPageId="13" captionText="FIGURE 6. DIC micrographs of Heth pivari n. sp., female and male structures. A) Lateral ornamentation in anterior region of female. B) Male, lateral somal extensions of posterior arcade surrounding anterior esophagus. C) Female, secretory-excretory pore (arrow). D. Female, somatic papillae anterior to vulva. E) Fusiform to comet-shaped sperm in male. All scale bars 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416676" httpUri="https://zenodo.org/record/4416676/files/figure.png" pageId="8" pageNumber="494">6C</figureCitation>
); SE complex fused with nerve ring and anchored dorsally to body wall, identical to that of male (
<figureCitation id="95FB10FFFF98126430CD1E04FD8FFAFF" box="[499,588,1286,1312]" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1920]" captionTargetBox="[160,1426,354,1861]" captionTargetId="figure-116@11.[151,1436,345,1872]" captionTargetPageId="11" captionText="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="8" pageNumber="494">Fig. 4B</figureCitation>
). Four coelomocytes: one in esophageal region, two next to anterior region of intestine or near anterior regions of ovaries, one near spermathecae (
<figureCitation id="95FB10FFFF98126432921E28FBC6FA9B" box="[940,1029,1322,1348]" captionStart="FIGURE 2" captionStartId="7.[151,250,1201,1225]" captionTargetBox="[156,1432,457,1173]" captionTargetId="figure-135@7.[151,1436,453,1177]" captionTargetPageId="7" captionText="FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov &amp; Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm." figureDoi="http://doi.org/10.5281/zenodo.4416665" httpUri="https://zenodo.org/record/4416665/files/figure.png" pageId="8" pageNumber="494">Fig. 2A</figureCitation>
). Lateral nerve chord clearly visible, identical to that of male (
<figureCitation id="95FB10FFFF981264308E1E4CFDC8FAB7" box="[432,523,1358,1384]" captionStart="FIGURE 7" captionStartId="14.[151,250,1613,1637]" captionTargetBox="[155,1432,185,1583]" captionTargetId="figure-18@14.[151,1436,181,1589]" captionTargetPageId="14" captionText="FIGURE 7. Heth pivari n. sp. A) Live female with single egg. B–E) Female enlargement and increasing numbers of eggs. The uterus is capacious but the vagina holds just one egg at a time (B, D). F) Female, dense trichomycete attachment in anterior region. G) Male, arthropod seta lodged in anteriormost region of intestine. H) Male, lateral chord (arrows). Scale bars: A, 500 µm; B–F, 250 µm; G, 100 µm; H, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416680" httpUri="https://zenodo.org/record/4416680/files/figure.png" pageId="8" pageNumber="494">Fig. 7H</figureCitation>
).
</paragraph>
<paragraph id="0D7F0C7AFF98126431F91E70FEDDF9C3" blockId="8.[151,1437,1070,2032]" pageId="8" pageNumber="494">
Two gonads, pro-didelphic, reflexed; spermathecae present, filled with large, fusiform or comet-shaped motile sperm (
<figureCitation id="95FB10FFFF98126431D61E94FEA6FA6F" box="[232,357,1430,1456]" captionStart="FIGURE 2" captionStartId="7.[151,250,1201,1225]" captionTargetBox="[156,1432,457,1173]" captionTargetId="figure-135@7.[151,1436,453,1177]" captionTargetPageId="7" captionText="FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov &amp; Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm." figureDoi="http://doi.org/10.5281/zenodo.4416665" httpUri="https://zenodo.org/record/4416665/files/figure.png" pageId="8" pageNumber="494">Figs.2A, E</figureCitation>
,
<figureCitation id="95FB10FFFF98126430501E94FE52FA6E" box="[366,401,1430,1457]" captionStart="FIGURE 6" captionStartId="13.[151,250,1904,1928]" captionTargetBox="[154,1433,672,1879]" captionTargetId="figure-214@13.[151,1435,669,1881]" captionTargetPageId="13" captionText="FIGURE 6. DIC micrographs of Heth pivari n. sp., female and male structures. A) Lateral ornamentation in anterior region of female. B) Male, lateral somal extensions of posterior arcade surrounding anterior esophagus. C) Female, secretory-excretory pore (arrow). D. Female, somatic papillae anterior to vulva. E) Fusiform to comet-shaped sperm in male. All scale bars 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416676" httpUri="https://zenodo.org/record/4416676/files/figure.png" pageId="8" pageNumber="494">6E</figureCitation>
). Muscular, spherical sphincter at junction of gonads with oviduct. Uterus prominent, distinctly muscled (
<figureCitation id="95FB10FFFF98126430381EB8FEAAFA0B" box="[262,361,1466,1492]" captionStart="FIGURE 2" captionStartId="7.[151,250,1201,1225]" captionTargetBox="[156,1432,457,1173]" captionTargetId="figure-135@7.[151,1436,453,1177]" captionTargetPageId="7" captionText="FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov &amp; Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm." figureDoi="http://doi.org/10.5281/zenodo.4416665" httpUri="https://zenodo.org/record/4416665/files/figure.png" pageId="8" pageNumber="494">Figs. 2E</figureCitation>
,
<figureCitation id="95FB10FFFF98126430491EB8FE7EFA0B" box="[375,445,1466,1492]" captionStart="FIGURE 7" captionStartId="14.[151,250,1613,1637]" captionTargetBox="[155,1432,185,1583]" captionTargetId="figure-18@14.[151,1436,181,1589]" captionTargetPageId="14" captionText="FIGURE 7. Heth pivari n. sp. A) Live female with single egg. B–E) Female enlargement and increasing numbers of eggs. The uterus is capacious but the vagina holds just one egg at a time (B, D). F) Female, dense trichomycete attachment in anterior region. G) Male, arthropod seta lodged in anteriormost region of intestine. H) Male, lateral chord (arrows). Scale bars: A, 500 µm; B–F, 250 µm; G, 100 µm; H, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416680" httpUri="https://zenodo.org/record/4416680/files/figure.png" pageId="8" pageNumber="494">7C, E</figureCitation>
). Eggs large, few to moderate in number, eggshells thin, without ornamentation (
<figureCitation id="95FB10FFFF981264345A1EB8FF67FA27" captionStart="FIGURE 7" captionStartId="14.[151,250,1613,1637]" captionTargetBox="[155,1432,185,1583]" captionTargetId="figure-18@14.[151,1436,181,1589]" captionTargetPageId="14" captionText="FIGURE 7. Heth pivari n. sp. A) Live female with single egg. B–E) Female enlargement and increasing numbers of eggs. The uterus is capacious but the vagina holds just one egg at a time (B, D). F) Female, dense trichomycete attachment in anterior region. G) Male, arthropod seta lodged in anteriormost region of intestine. H) Male, lateral chord (arrows). Scale bars: A, 500 µm; B–F, 250 µm; G, 100 µm; H, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416680" httpUri="https://zenodo.org/record/4416680/files/figure.png" pageId="8" pageNumber="494">Figs. 7</figureCitation>
A–E). Vulva near anus. Phasmid apertures minute, pores located just posterior to anus. Tail filiform, attenuated to a fine point.
</paragraph>
<paragraph id="0D7F0C7AFF98126431F91D24FE00F8E3" blockId="8.[151,1437,1070,2032]" pageId="8" pageNumber="494">
<emphasis id="3FB4D068FF98126431F91D24FED5F99F" bold="true" box="[199,278,1574,1600]" pageId="8" pageNumber="494">Males.</emphasis>
Measurements given in
<tableCitation id="404239C1FF98126433061D24FD50F99F" box="[568,659,1574,1600]" captionStart="TABLE 3" captionStartId="10.[151,241,151,175]" captionTargetPageId="10" captionText="TABLE 3. Morphometric measurements of Heth pivari n. sp. males." httpUri="http://table.plazi.org/id/59BF5CF2FF9A126631A91B95FCBAFF70" pageId="8" pageNumber="494" tableUuid="59BF5CF2FF9A126631A91B95FCBAFF70">Table 3</tableCitation>
. Body robust, smaller than females, white, head and neck without ornamentation (
<figureCitation id="95FB10FFFF98126430761D48FE6FF9BB" box="[328,428,1610,1636]" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1920]" captionTargetBox="[160,1426,354,1861]" captionTargetId="figure-116@11.[151,1436,345,1872]" captionTargetPageId="11" captionText="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="8" pageNumber="494">Figs. 4A</figureCitation>
,
<figureCitation id="95FB10FFFF98126430861D48FDDCF9BB" box="[440,543,1610,1637]" captionStart="FIGURE 5" captionStartId="12.[151,250,1361,1385]" captionTargetBox="[155,1433,184,1334]" captionTargetId="figure-18@12.[151,1436,181,1337]" captionTargetPageId="12" captionText="FIGURE 5. SEM micrographs of Heth pivari n. sp. males. A) Anterior end, stoma slit-like. B) Anterior end, stoma open. C) Closeup of stoma showing striae and minute amphid aperture (a). D) Posterior region showing ventral sucker (S), papillae 1,2 and 4, and tail. E) Posterior region showing arrangement of posterior papillae 2–7. F) Closeup of cloacal region. Scale bars: A, B, E, 30 µm; C, F, 20 µm; D, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4416674" httpUri="https://zenodo.org/record/4416674/files/figure.png" pageId="8" pageNumber="494">5A, B, C</figureCitation>
); lateral field absent; cuticle finely annulated, each annule
<quantity id="CA38A19FFF981264358D1D48FADCF9BB" box="[1203,1311,1610,1637]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="8" pageNumber="494" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 µm</quantity>
wide, with scattered, smooth papillae (
<figureCitation id="95FB10FFFF98126430F71D6CFD88F957" box="[457,587,1646,1673]" captionStart="FIGURE 5" captionStartId="12.[151,250,1361,1385]" captionTargetBox="[155,1433,184,1334]" captionTargetId="figure-18@12.[151,1436,181,1337]" captionTargetPageId="12" captionText="FIGURE 5. SEM micrographs of Heth pivari n. sp. males. A) Anterior end, stoma slit-like. B) Anterior end, stoma open. C) Closeup of stoma showing striae and minute amphid aperture (a). D) Posterior region showing ventral sucker (S), papillae 1,2 and 4, and tail. E) Posterior region showing arrangement of posterior papillae 2–7. F) Closeup of cloacal region. Scale bars: A, B, E, 30 µm; C, F, 20 µm; D, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4416674" httpUri="https://zenodo.org/record/4416674/files/figure.png" pageId="8" pageNumber="494">Figs. 5B, E</figureCitation>
). Stomatal opening longer than wide to slit-like (
<figureCitation id="95FB10FFFF98126435531D6CFB2CF957" box="[1133,1263,1646,1672]" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1920]" captionTargetBox="[160,1426,354,1861]" captionTargetId="figure-116@11.[151,1436,345,1872]" captionTargetPageId="11" captionText="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="8" pageNumber="494">Figs. 4B, C</figureCitation>
,
<figureCitation id="95FB10FFFF98126435C51D6CFACBF956" box="[1275,1288,1646,1673]" captionStart="FIGURE 5" captionStartId="12.[151,250,1361,1385]" captionTargetBox="[155,1433,184,1334]" captionTargetId="figure-18@12.[151,1436,181,1337]" captionTargetPageId="12" captionText="FIGURE 5. SEM micrographs of Heth pivari n. sp. males. A) Anterior end, stoma slit-like. B) Anterior end, stoma open. C) Closeup of stoma showing striae and minute amphid aperture (a). D) Posterior region showing ventral sucker (S), papillae 1,2 and 4, and tail. E) Posterior region showing arrangement of posterior papillae 2–7. F) Closeup of cloacal region. Scale bars: A, B, E, 30 µm; C, F, 20 µm; D, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4416674" httpUri="https://zenodo.org/record/4416674/files/figure.png" pageId="8" pageNumber="494">5</figureCitation>
A–C). Stoma tubular, with three feather-like appendages (
<figureCitation id="95FB10FFFF98126433AC1D90FD2EF972" box="[658,749,1682,1709]" captionStart="FIGURE 5" captionStartId="12.[151,250,1361,1385]" captionTargetBox="[155,1433,184,1334]" captionTargetId="figure-18@12.[151,1436,181,1337]" captionTargetPageId="12" captionText="FIGURE 5. SEM micrographs of Heth pivari n. sp. males. A) Anterior end, stoma slit-like. B) Anterior end, stoma open. C) Closeup of stoma showing striae and minute amphid aperture (a). D) Posterior region showing ventral sucker (S), papillae 1,2 and 4, and tail. E) Posterior region showing arrangement of posterior papillae 2–7. F) Closeup of cloacal region. Scale bars: A, B, E, 30 µm; C, F, 20 µm; D, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4416674" httpUri="https://zenodo.org/record/4416674/files/figure.png" pageId="8" pageNumber="494">Fig. 5B</figureCitation>
); stomatal structure partially obscured by tapering, partially striated esophageal collar. Lips finely striated, slightly elevated with four inconspicuous cephalic papillae (
<figureCitation id="95FB10FFFF98126434081DB4FF6EF92B" captionStart="FIGURE 5" captionStartId="12.[151,250,1361,1385]" captionTargetBox="[155,1433,184,1334]" captionTargetId="figure-18@12.[151,1436,181,1337]" captionTargetPageId="12" captionText="FIGURE 5. SEM micrographs of Heth pivari n. sp. males. A) Anterior end, stoma slit-like. B) Anterior end, stoma open. C) Closeup of stoma showing striae and minute amphid aperture (a). D) Posterior region showing ventral sucker (S), papillae 1,2 and 4, and tail. E) Posterior region showing arrangement of posterior papillae 2–7. F) Closeup of cloacal region. Scale bars: A, B, E, 30 µm; C, F, 20 µm; D, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4416674" httpUri="https://zenodo.org/record/4416674/files/figure.png" pageId="8" pageNumber="494">Figs. 5B, C</figureCitation>
). Amphid apertures inconspicuous, circular, on inner slope of raised lip region, approximately
<quantity id="CA38A19FFF98126435E41DD8FAD5F92B" box="[1242,1302,1754,1780]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="494" unit="mm" value="3.0">3 µm</quantity>
in diameter (
<figureCitation id="95FB10FFFF981264319E1DFCFF3AF8C6" box="[160,249,1790,1817]" captionStart="FIGURE 5" captionStartId="12.[151,250,1361,1385]" captionTargetBox="[155,1433,184,1334]" captionTargetId="figure-18@12.[151,1436,181,1337]" captionTargetPageId="12" captionText="FIGURE 5. SEM micrographs of Heth pivari n. sp. males. A) Anterior end, stoma slit-like. B) Anterior end, stoma open. C) Closeup of stoma showing striae and minute amphid aperture (a). D) Posterior region showing ventral sucker (S), papillae 1,2 and 4, and tail. E) Posterior region showing arrangement of posterior papillae 2–7. F) Closeup of cloacal region. Scale bars: A, B, E, 30 µm; C, F, 20 µm; D, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4416674" httpUri="https://zenodo.org/record/4416674/files/figure.png" pageId="8" pageNumber="494">Fig. 5C</figureCitation>
). Head region with pairs of subdorsal and subventral muscles attached to stoma and extending obliquely to body wall (
<figureCitation id="95FB10FFFF981264300C1C20FE75F8E3" box="[306,438,1826,1852]" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1920]" captionTargetBox="[160,1426,354,1861]" captionTargetId="figure-116@11.[151,1436,345,1872]" captionTargetPageId="11" captionText="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="8" pageNumber="494">Figs. 4B, C</figureCitation>
).
</paragraph>
<paragraph id="0D7F0C7AFF98126531F91C44FBD7FE9E" blockId="8.[151,1437,1070,2032]" lastBlockId="9.[151,1436,151,321]" lastPageId="9" lastPageNumber="495" pageId="8" pageNumber="494">
Six prominent amber-brown, uninucleate gland-like somatic extensions of posterior arcade syncytium surrounding anterior part of esophagus and extending to nerve ring (
<figureCitation id="95FB10FFFF98126432441C68FC1AF85B" box="[890,985,1898,1924]" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1920]" captionTargetBox="[160,1426,354,1861]" captionTargetId="figure-116@11.[151,1436,345,1872]" captionTargetPageId="11" captionText="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="8" pageNumber="494">Figs. 4B</figureCitation>
,
<figureCitation id="95FB10FFFF98126432DD1C68FBC5F85A" box="[995,1030,1898,1925]" captionStart="FIGURE 6" captionStartId="13.[151,250,1904,1928]" captionTargetBox="[154,1433,672,1879]" captionTargetId="figure-214@13.[151,1435,669,1881]" captionTargetPageId="13" captionText="FIGURE 6. DIC micrographs of Heth pivari n. sp., female and male structures. A) Lateral ornamentation in anterior region of female. B) Male, lateral somal extensions of posterior arcade surrounding anterior esophagus. C) Female, secretory-excretory pore (arrow). D. Female, somatic papillae anterior to vulva. E) Fusiform to comet-shaped sperm in male. All scale bars 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416676" httpUri="https://zenodo.org/record/4416676/files/figure.png" pageId="8" pageNumber="494">6B</figureCitation>
); at least two much smaller, colorless extensions of the anterior arcade syncytium located near base of stoma (
<figureCitation id="95FB10FFFF98126432F11C8CFBEAF877" box="[975,1065,1934,1960]" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1920]" captionTargetBox="[160,1426,354,1861]" captionTargetId="figure-116@11.[151,1436,345,1872]" captionTargetPageId="11" captionText="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="8" pageNumber="494">Fig. 4B</figureCitation>
). Esophagus approximately onethird of body length, procorpus muscled; broadly pyriform basal bulb containing grinding valve. Intestine dilated at anterior end, then attenuating to a uniform diameter and in ventral position, crossing testis near ventral sucker and becoming dorsal, then narrowing at cloaca (
<figureCitation id="95FB10FFFF99126533401B95FCC7FF6E" box="[638,772,151,177]" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1920]" captionTargetBox="[160,1426,354,1861]" captionTargetId="figure-116@11.[151,1436,345,1872]" captionTargetPageId="11" captionText="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="9" pageNumber="495">Figs. 4A, D</figureCitation>
). Secretory-excretory (SE) complex and nerve ring forming a single arching structure (
<figureCitation id="95FB10FFFF99126530851BB9FDD1FF0A" box="[443,530,187,213]" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1920]" captionTargetBox="[160,1426,354,1861]" captionTargetId="figure-116@11.[151,1436,345,1872]" captionTargetPageId="11" captionText="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="9" pageNumber="495">Fig. 4B</figureCitation>
); SE pore inconspicuous, connected to sinuous tube in the SE complex; nerve ring thin, with many associated nuclei, fused with connective tissue extending to dorsal cuticular wall (
<figureCitation id="95FB10FFFF99126535C61BDDFA91FF26" box="[1272,1362,223,249]" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1920]" captionTargetBox="[160,1426,354,1861]" captionTargetId="figure-116@11.[151,1436,345,1872]" captionTargetPageId="11" captionText="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="9" pageNumber="495">Fig. 4B</figureCitation>
). Four coelomocytes, one each in esophageal region, in anterior area of intestine, near flexure of testis and near junction of seminal vesicle and vas deferens. Lateral nerve chords clearly visible (
<figureCitation id="95FB10FFFF99126532931A25FBC4FE9E" box="[941,1031,295,321]" captionStart="FIGURE 7" captionStartId="14.[151,250,1613,1637]" captionTargetBox="[155,1432,185,1583]" captionTargetId="figure-18@14.[151,1436,181,1589]" captionTargetPageId="14" captionText="FIGURE 7. Heth pivari n. sp. A) Live female with single egg. B–E) Female enlargement and increasing numbers of eggs. The uterus is capacious but the vagina holds just one egg at a time (B, D). F) Female, dense trichomycete attachment in anterior region. G) Male, arthropod seta lodged in anteriormost region of intestine. H) Male, lateral chord (arrows). Scale bars: A, 500 µm; B–F, 250 µm; G, 100 µm; H, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416680" httpUri="https://zenodo.org/record/4416680/files/figure.png" pageId="9" pageNumber="495">Fig. 7H</figureCitation>
).
</paragraph>
<caption id="59BF5CF2FF99126531A91EDBFAA8F9CA" ID-DOI="http://doi.org/10.5281/zenodo.4416668" ID-Zenodo-Dep="4416668" httpUri="https://zenodo.org/record/4416668/files/figure.png" pageId="9" pageNumber="495" startId="9.[151,250,1497,1521]" targetBox="[155,1429,349,1466]" targetPageId="9">
<paragraph id="0D7F0C7AFF99126531A91EDBFAA8F9CA" blockId="9.[151,1436,1497,1557]" pageId="9" pageNumber="495">
<emphasis id="3FB4D068FF99126531A91EDBFED6FA2E" bold="true" box="[151,277,1497,1521]" pageId="9" pageNumber="495">FIGURE 3.</emphasis>
SEM micrographs of
<taxonomicName id="CAC077F9FF99126530C31EDBFDB0FA2E" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[509,627,1497,1521]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="9" pageNumber="495" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF99126530C31EDBFDB0FA2E" box="[509,627,1497,1521]" italics="true" pageId="9" pageNumber="495">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF99126533451EDBFD75FA2E" bold="true" box="[635,694,1497,1521]" pageId="9" pageNumber="495">
<taxonomicNameLabel id="24876D13FF99126533451EDBFD75FA2E" box="[635,694,1497,1521]" pageId="9" pageNumber="495" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm.
</paragraph>
</caption>
<paragraph id="0D7F0C7AFF99126531F91D41FBC7F87D" blockId="9.[151,1437,1603,1954]" pageId="9" pageNumber="495">
Reproductive system monorchic, with short flexure in germinal zone. Sperm comet to fusiform-shaped, about
<quantity id="CA38A19FFF99126531A91D65FF21F95D" box="[151,226,1639,1666]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" pageId="9" pageNumber="495" unit="mm" value="88.0">88 µm</quantity>
long ×
<quantity id="CA38A19FFF99126530001D65FE4AF95D" box="[318,393,1639,1666]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="495" unit="mm" value="15.0">15 µm</quantity>
at widest point (
<figureCitation id="95FB10FFFF99126533721D65FD66F95D" box="[588,677,1639,1666]" captionStart="FIGURE 6" captionStartId="13.[151,250,1904,1928]" captionTargetBox="[154,1433,672,1879]" captionTargetId="figure-214@13.[151,1435,669,1881]" captionTargetPageId="13" captionText="FIGURE 6. DIC micrographs of Heth pivari n. sp., female and male structures. A) Lateral ornamentation in anterior region of female. B) Male, lateral somal extensions of posterior arcade surrounding anterior esophagus. C) Female, secretory-excretory pore (arrow). D. Female, somatic papillae anterior to vulva. E) Fusiform to comet-shaped sperm in male. All scale bars 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4416676" httpUri="https://zenodo.org/record/4416676/files/figure.png" pageId="9" pageNumber="495">Fig. 6E</figureCitation>
). Ventral sucker present,
<quantity id="CA38A19FFF99126532F41D65FB98F95D" box="[970,1115,1639,1666]" metricMagnitude="-1" metricUnit="m" metricValue="2.66" metricValueMax="3.3" metricValueMin="2.02" pageId="9" pageNumber="495" unit="mm" value="266.0" valueMax="330.0" valueMin="202.0">202–330 µm</quantity>
anterior to cloacal opening, flanked by first pair of large pre-cloacal papillae (
<figureCitation id="95FB10FFFF99126533F61D89FCEEF97A" box="[712,813,1675,1701]" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1920]" captionTargetBox="[160,1426,354,1861]" captionTargetId="figure-116@11.[151,1436,345,1872]" captionTargetPageId="11" captionText="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="9" pageNumber="495">Figs. 4D</figureCitation>
,
<figureCitation id="95FB10FFFF99126532071D89FC9DF979" box="[825,862,1675,1702]" captionStart="FIGURE 5" captionStartId="12.[151,250,1361,1385]" captionTargetBox="[155,1433,184,1334]" captionTargetId="figure-18@12.[151,1436,181,1337]" captionTargetPageId="12" captionText="FIGURE 5. SEM micrographs of Heth pivari n. sp. males. A) Anterior end, stoma slit-like. B) Anterior end, stoma open. C) Closeup of stoma showing striae and minute amphid aperture (a). D) Posterior region showing ventral sucker (S), papillae 1,2 and 4, and tail. E) Posterior region showing arrangement of posterior papillae 2–7. F) Closeup of cloacal region. Scale bars: A, B, E, 30 µm; C, F, 20 µm; D, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4416674" httpUri="https://zenodo.org/record/4416674/files/figure.png" pageId="9" pageNumber="495">5D</figureCitation>
). Region between sucker and cloaca with scattered somatic papillae of
<quantity id="CA38A19FFF99126530BE1DADFE18F915" box="[384,475,1711,1738]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="9" pageNumber="495" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 µm</quantity>
diameter. Spicules equal, fused for distal three-fourths; each spicule with distinct capitulum, apical fused portion arcuate ventrad, tapering to a point; gubernaculum with pointed proximal end, distal end broad and bilobed (
<figureCitation id="95FB10FFFF99126530D61DF5FDACF8CD" box="[488,623,1783,1810]" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1920]" captionTargetBox="[160,1426,354,1861]" captionTargetId="figure-116@11.[151,1436,345,1872]" captionTargetPageId="11" captionText="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="9" pageNumber="495">Figs. 4D, E</figureCitation>
). Cloacal lips smooth. Seven pairs of genital papillae, each with several conspicuous internal striae or minute tubes; one median ventral papilla. Caudal papilla pattern as follows: first pair slightly posterior to ventral sucker; second pair about
<quantity id="CA38A19FFF99126533C81C3DFC83F885" box="[758,832,1855,1882]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="495" unit="mm" value="30.0">30 µm</quantity>
anterior to cloaca, distant from third pair; pairs 3 and 4 slightly anterior to cloaca, pair 3 ventral, pair 4 lateral; single medial papilla (5) on anterior cloacal lip; posterior to cloaca, two ventral pairs (6, 7) and one subdorsal pair (8) (
<figureCitation id="95FB10FFFF991265327A1C85FC6BF87E" box="[836,936,1927,1953]" captionStart="FIGURE 4" captionStartId="11.[151,250,1896,1920]" captionTargetBox="[160,1426,354,1861]" captionTargetId="figure-116@11.[151,1436,345,1872]" captionTargetPageId="11" captionText="FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="9" pageNumber="495">Figs. 4D</figureCitation>
,
<figureCitation id="95FB10FFFF991265328D1C85FC03F87D" box="[947,960,1927,1954]" captionStart="FIGURE 5" captionStartId="12.[151,250,1361,1385]" captionTargetBox="[155,1433,184,1334]" captionTargetId="figure-18@12.[151,1436,181,1337]" captionTargetPageId="12" captionText="FIGURE 5. SEM micrographs of Heth pivari n. sp. males. A) Anterior end, stoma slit-like. B) Anterior end, stoma open. C) Closeup of stoma showing striae and minute amphid aperture (a). D) Posterior region showing ventral sucker (S), papillae 1,2 and 4, and tail. E) Posterior region showing arrangement of posterior papillae 2–7. F) Closeup of cloacal region. Scale bars: A, B, E, 30 µm; C, F, 20 µm; D, 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4416674" httpUri="https://zenodo.org/record/4416674/files/figure.png" pageId="9" pageNumber="495">5</figureCitation>
D–F).
</paragraph>
</subSubSection>
<caption id="59BF5CF2FF9A126631A91B95FCBAFF70" ID-Table-UUID="59BF5CF2FF9A126631A91B95FCBAFF70" box="[151,889,151,176]" httpUri="http://table.plazi.org/id/59BF5CF2FF9A126631A91B95FCBAFF70" pageId="10" pageNumber="496" targetBox="[166,1367,196,976]" targetIsTable="true" targetPageId="10">
<paragraph id="0D7F0C7AFF9A126631A91B95FCBAFF70" blockId="10.[151,889,151,176]" box="[151,889,151,176]" pageId="10" pageNumber="496">
<emphasis id="3FB4D068FF9A126631A91B95FECFFF70" bold="true" box="[151,268,151,175]" pageId="10" pageNumber="496">TABLE 3.</emphasis>
Morphometric measurements of
<taxonomicName id="CAC077F9FF9A1266334E1B95FD29FF70" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[624,746,151,175]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A1266334E1B95FD29FF70" box="[624,746,151,175]" italics="true" pageId="10" pageNumber="496">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126633CF1B9AFCEEFF6F" bold="true" box="[753,813,152,176]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126633CF1B9AFCEEFF6F" box="[753,813,152,176]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
males.
</paragraph>
</caption>
<paragraph id="0D7F0C7AFF9A126633551BD5FA88FC0F" pageId="10" pageNumber="496">
<table id="7FC0FEDAFF9AED9331981BC6FA94FC0F" box="[166,1367,196,976]" gridcols="6" gridrows="19" pageId="10" pageNumber="496">
<tr id="B3F00E38FF9AED9331981BC6FA94FEDD" box="[166,1367,196,258]" gridrow="0" pageId="10" pageNumber="496" rowspan-0="1">
<th id="F0216744FF9AED9333551BC6FD07FEDD" box="[619,708,196,258]" gridcol="1" gridrow="0" pageId="10" pageNumber="496">Allotype</th>
<th id="F0216744FF9AED9332251BC6FBCCFEDD" box="[795,1039,196,258]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="10" pageNumber="496">
Paratype Males (
<emphasis id="3FB4D068FF9A126632F81BC7FC10FF03" box="[966,979,197,220]" italics="true" pageId="10" pageNumber="496">n</emphasis>
= 30) Mean SD
</th>
<th id="F0216744FF9AED9335131BC6FB6FFEDD" box="[1069,1196,196,258]" gridcol="4" gridrow="0" pageId="10" pageNumber="496">Range</th>
<th id="F0216744FF9AED9334151BC6FA94FEDD" box="[1323,1367,196,258]" gridcol="5" gridrow="0" pageId="10" pageNumber="496">CV</th>
</tr>
<tr id="B3F00E38FF9AED9331981A34FA94FE91" box="[166,1367,310,334]" gridrow="1" pageId="10" pageNumber="496" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="F0216744FF9AED9331981A34FE5AFE91" box="[166,409,310,334]" gridcol="0" gridrow="1" pageId="10" pageNumber="496">Measurements (µm)</th>
</tr>
<tr id="B3F00E38FF9AED9331981A59FA94FEAB" box="[166,1367,347,372]" gridrow="2" pageId="10" pageNumber="496">
<th id="F0216744FF9AED9331981A59FE5AFEAB" box="[166,409,347,372]" gridcol="0" gridrow="2" pageId="10" pageNumber="496">Length</th>
<td id="F0216744FF9AED9333551A59FD07FEAB" box="[619,708,347,372]" gridcol="1" gridrow="2" pageId="10" pageNumber="496">2,075</td>
<td id="F0216744FF9AED9332251A59FC97FEAB" box="[795,852,347,372]" gridcol="2" gridrow="2" pageId="10" pageNumber="496">2,253</td>
<td id="F0216744FF9AED9332931A59FBCCFEAB" box="[941,1039,347,372]" gridcol="3" gridrow="2" pageId="10" pageNumber="496">280</td>
<td id="F0216744FF9AED9335131A59FB6FFEAB" box="[1069,1196,347,372]" gridcol="4" gridrow="2" pageId="10" pageNumber="496">1,897–2,997</td>
<td id="F0216744FF9AED9334151A59FA94FEAB" box="[1323,1367,347,372]" gridcol="5" gridrow="2" pageId="10" pageNumber="496">12.4</td>
</tr>
<tr id="B3F00E38FF9AED9331981A83FA94FE46" box="[166,1367,385,409]" gridrow="3" pageId="10" pageNumber="496">
<th id="F0216744FF9AED9331981A83FE5AFE46" box="[166,409,385,409]" gridcol="0" gridrow="3" pageId="10" pageNumber="496">Maximum width</th>
<td id="F0216744FF9AED9333551A83FD07FE46" box="[619,708,385,409]" gridcol="1" gridrow="3" pageId="10" pageNumber="496">130</td>
<td id="F0216744FF9AED9332251A83FC97FE46" box="[795,852,385,409]" gridcol="2" gridrow="3" pageId="10" pageNumber="496">137</td>
<td id="F0216744FF9AED9332931A83FBCCFE46" box="[941,1039,385,409]" gridcol="3" gridrow="3" pageId="10" pageNumber="496">29</td>
<td id="F0216744FF9AED9335131A83FB6FFE46" box="[1069,1196,385,409]" gridcol="4" gridrow="3" pageId="10" pageNumber="496">99–198</td>
<td id="F0216744FF9AED9334151A83FA94FE46" box="[1323,1367,385,409]" gridcol="5" gridrow="3" pageId="10" pageNumber="496">21.2</td>
</tr>
<tr id="B3F00E38FF9AED9331981AA5FA94FE60" box="[166,1367,423,447]" gridrow="4" pageId="10" pageNumber="496">
<th id="F0216744FF9AED9331981AA5FE5AFE60" box="[166,409,423,447]" gridcol="0" gridrow="4" pageId="10" pageNumber="496">Esophagus length</th>
<td id="F0216744FF9AED9333551AA5FD07FE60" box="[619,708,423,447]" gridcol="1" gridrow="4" pageId="10" pageNumber="496">596</td>
<td id="F0216744FF9AED9332251AA5FC97FE60" box="[795,852,423,447]" gridcol="2" gridrow="4" pageId="10" pageNumber="496">622</td>
<td id="F0216744FF9AED9332931AA5FBCCFE60" box="[941,1039,423,447]" gridcol="3" gridrow="4" pageId="10" pageNumber="496">28</td>
<td id="F0216744FF9AED9335131AA5FB6FFE60" box="[1069,1196,423,447]" gridcol="4" gridrow="4" pageId="10" pageNumber="496">567–681</td>
<td id="F0216744FF9AED9334151AA5FA94FE60" box="[1323,1367,423,447]" gridcol="5" gridrow="4" pageId="10" pageNumber="496">4.5</td>
</tr>
<tr id="B3F00E38FF9AED9331981ACFFA94FE3A" box="[166,1367,461,485]" gridrow="5" pageId="10" pageNumber="496">
<th id="F0216744FF9AED9331981ACFFE5AFE3A" box="[166,409,461,485]" gridcol="0" gridrow="5" pageId="10" pageNumber="496">Basal bulb length</th>
<td id="F0216744FF9AED9333551ACFFD07FE3A" box="[619,708,461,485]" gridcol="1" gridrow="5" pageId="10" pageNumber="496">138</td>
<td id="F0216744FF9AED9332251ACFFC97FE3A" box="[795,852,461,485]" gridcol="2" gridrow="5" pageId="10" pageNumber="496">135</td>
<td id="F0216744FF9AED9332931ACFFBCCFE3A" box="[941,1039,461,485]" gridcol="3" gridrow="5" pageId="10" pageNumber="496">3.1</td>
<td id="F0216744FF9AED9335131ACFFB6FFE3A" box="[1069,1196,461,485]" gridcol="4" gridrow="5" pageId="10" pageNumber="496">130–144</td>
<td id="F0216744FF9AED9334151ACFFA94FE3A" box="[1323,1367,461,485]" gridcol="5" gridrow="5" pageId="10" pageNumber="496">2.3</td>
</tr>
<tr id="B3F00E38FF9AED9331981AF0FA94FDD4" box="[166,1367,498,523]" gridrow="6" pageId="10" pageNumber="496">
<th id="F0216744FF9AED9331981AF0FE5AFDD4" box="[166,409,498,523]" gridcol="0" gridrow="6" pageId="10" pageNumber="496">Head to excretory pore</th>
<td id="F0216744FF9AED9333551AF0FD07FDD4" box="[619,708,498,523]" gridcol="1" gridrow="6" pageId="10" pageNumber="496">250</td>
<td id="F0216744FF9AED9332251AF0FC97FDD4" box="[795,852,498,523]" gridcol="2" gridrow="6" pageId="10" pageNumber="496">264</td>
<td id="F0216744FF9AED9332931AF0FBCCFDD4" box="[941,1039,498,523]" gridcol="3" gridrow="6" pageId="10" pageNumber="496">19</td>
<td id="F0216744FF9AED9335131AF0FB6FFDD4" box="[1069,1196,498,523]" gridcol="4" gridrow="6" pageId="10" pageNumber="496">236–290</td>
<td id="F0216744FF9AED9334151AF0FA94FDD4" box="[1323,1367,498,523]" gridcol="5" gridrow="6" pageId="10" pageNumber="496">7.2</td>
</tr>
<tr id="B3F00E38FF9AED933198191AFA94FDEF" box="[166,1367,536,560]" gridrow="7" pageId="10" pageNumber="496">
<th id="F0216744FF9AED933198191AFE5AFDEF" box="[166,409,536,560]" gridcol="0" gridrow="7" pageId="10" pageNumber="496">Ventral sucker length</th>
<td id="F0216744FF9AED933355191AFD07FDEF" box="[619,708,536,560]" gridcol="1" gridrow="7" pageId="10" pageNumber="496">20</td>
<td id="F0216744FF9AED933225191AFC97FDEF" box="[795,852,536,560]" gridcol="2" gridrow="7" pageId="10" pageNumber="496">22.5</td>
<td id="F0216744FF9AED933293191AFBCCFDEF" box="[941,1039,536,560]" gridcol="3" gridrow="7" pageId="10" pageNumber="496">3</td>
<td id="F0216744FF9AED933513191AFB6FFDEF" box="[1069,1196,536,560]" gridcol="4" gridrow="7" pageId="10" pageNumber="496">18–29</td>
<td id="F0216744FF9AED933415191AFA94FDEF" box="[1323,1367,536,560]" gridcol="5" gridrow="7" pageId="10" pageNumber="496">13.3</td>
</tr>
<tr id="B3F00E38FF9AED933198193CFA94FD89" box="[166,1367,574,598]" gridrow="8" pageId="10" pageNumber="496">
<th id="F0216744FF9AED933198193CFE5AFD89" box="[166,409,574,598]" gridcol="0" gridrow="8" pageId="10" pageNumber="496">Ventral sucker width</th>
<td id="F0216744FF9AED933355193CFD07FD89" box="[619,708,574,598]" gridcol="1" gridrow="8" pageId="10" pageNumber="496">15</td>
<td id="F0216744FF9AED933225193CFC97FD89" box="[795,852,574,598]" gridcol="2" gridrow="8" pageId="10" pageNumber="496">15.9</td>
<td id="F0216744FF9AED933293193CFBCCFD89" box="[941,1039,574,598]" gridcol="3" gridrow="8" pageId="10" pageNumber="496">2.7</td>
<td id="F0216744FF9AED933513193CFB6FFD89" box="[1069,1196,574,598]" gridcol="4" gridrow="8" pageId="10" pageNumber="496">9–22</td>
<td id="F0216744FF9AED933415193CFA94FD89" box="[1323,1367,574,598]" gridcol="5" gridrow="8" pageId="10" pageNumber="496">17</td>
</tr>
<tr id="B3F00E38FF9AED9331981966FA94FDA3" box="[166,1367,612,636]" gridrow="9" pageId="10" pageNumber="496">
<th id="F0216744FF9AED9331981966FE5AFDA3" box="[166,409,612,636]" gridcol="0" gridrow="9" pageId="10" pageNumber="496">Ventral sucker to cloaca</th>
<td id="F0216744FF9AED9333551966FD07FDA3" box="[619,708,612,636]" gridcol="1" gridrow="9" pageId="10" pageNumber="496">219</td>
<td id="F0216744FF9AED9332251966FC97FDA3" box="[795,852,612,636]" gridcol="2" gridrow="9" pageId="10" pageNumber="496">248</td>
<td id="F0216744FF9AED9332931966FBCCFDA3" box="[941,1039,612,636]" gridcol="3" gridrow="9" pageId="10" pageNumber="496">32</td>
<td id="F0216744FF9AED9335131966FB6FFDA3" box="[1069,1196,612,636]" gridcol="4" gridrow="9" pageId="10" pageNumber="496">202–330</td>
<td id="F0216744FF9AED9334151966FA94FDA3" box="[1323,1367,612,636]" gridcol="5" gridrow="9" pageId="10" pageNumber="496">12.9</td>
</tr>
<tr id="B3F00E38FF9AED9331981988FA94FD7D" box="[166,1367,650,674]" gridrow="10" pageId="10" pageNumber="496">
<th id="F0216744FF9AED9331981988FE5AFD7D" box="[166,409,650,674]" gridcol="0" gridrow="10" pageId="10" pageNumber="496">Spicule arc</th>
<td id="F0216744FF9AED9333551988FD07FD7D" box="[619,708,650,674]" gridcol="1" gridrow="10" pageId="10" pageNumber="496">102</td>
<td id="F0216744FF9AED9332251988FC97FD7D" box="[795,852,650,674]" gridcol="2" gridrow="10" pageId="10" pageNumber="496">109</td>
<td id="F0216744FF9AED9332931988FBCCFD7D" box="[941,1039,650,674]" gridcol="3" gridrow="10" pageId="10" pageNumber="496">6.4</td>
<td id="F0216744FF9AED9335131988FB6FFD7D" box="[1069,1196,650,674]" gridcol="4" gridrow="10" pageId="10" pageNumber="496">96–124</td>
<td id="F0216744FF9AED9334151988FA94FD7D" box="[1323,1367,650,674]" gridcol="5" gridrow="10" pageId="10" pageNumber="496">5.9</td>
</tr>
<tr id="B3F00E38FF9AED93319819ADFA94FD17" box="[166,1367,687,712]" gridrow="11" pageId="10" pageNumber="496">
<th id="F0216744FF9AED93319819ADFE5AFD17" box="[166,409,687,712]" gridcol="0" gridrow="11" pageId="10" pageNumber="496">Gubernaculum arc</th>
<td id="F0216744FF9AED93335519ADFD07FD17" box="[619,708,687,712]" gridcol="1" gridrow="11" pageId="10" pageNumber="496">50</td>
<td id="F0216744FF9AED93322519ADFC97FD17" box="[795,852,687,712]" gridcol="2" gridrow="11" pageId="10" pageNumber="496">66</td>
<td id="F0216744FF9AED93329319ADFBCCFD17" box="[941,1039,687,712]" gridcol="3" gridrow="11" pageId="10" pageNumber="496">5.1</td>
<td id="F0216744FF9AED93351319ADFB6FFD17" box="[1069,1196,687,712]" gridcol="4" gridrow="11" pageId="10" pageNumber="496">53–79</td>
<td id="F0216744FF9AED93341519ADFA94FD17" box="[1323,1367,687,712]" gridcol="5" gridrow="11" pageId="10" pageNumber="496">7.7</td>
</tr>
<tr id="B3F00E38FF9AED93319819D7FA94FD32" box="[166,1367,725,749]" gridrow="12" pageId="10" pageNumber="496">
<th id="F0216744FF9AED93319819D7FE5AFD32" box="[166,409,725,749]" gridcol="0" gridrow="12" pageId="10" pageNumber="496">Cloaca head</th>
<td id="F0216744FF9AED93335519D7FD07FD32" box="[619,708,725,749]" gridcol="1" gridrow="12" pageId="10" pageNumber="496">1,916</td>
<td id="F0216744FF9AED93322519D7FC97FD32" box="[795,852,725,749]" gridcol="2" gridrow="12" pageId="10" pageNumber="496">2,086</td>
<td id="F0216744FF9AED93329319D7FBCCFD32" box="[941,1039,725,749]" gridcol="3" gridrow="12" pageId="10" pageNumber="496">271</td>
<td id="F0216744FF9AED93351319D7FB6FFD32" box="[1069,1196,725,749]" gridcol="4" gridrow="12" pageId="10" pageNumber="496">1,752–2,815</td>
<td id="F0216744FF9AED93341519D7FA94FD32" box="[1323,1367,725,749]" gridcol="5" gridrow="12" pageId="10" pageNumber="496">13</td>
</tr>
<tr id="B3F00E38FF9AED93319819F9FA94FCCC" box="[166,1367,763,787]" gridrow="13" pageId="10" pageNumber="496">
<th id="F0216744FF9AED93319819F9FE5AFCCC" box="[166,409,763,787]" gridcol="0" gridrow="13" pageId="10" pageNumber="496">Tail length</th>
<td id="F0216744FF9AED93335519F9FD07FCCC" box="[619,708,763,787]" gridcol="1" gridrow="13" pageId="10" pageNumber="496">159</td>
<td id="F0216744FF9AED93322519F9FC97FCCC" box="[795,852,763,787]" gridcol="2" gridrow="13" pageId="10" pageNumber="496">167</td>
<td id="F0216744FF9AED93329319F9FBCCFCCC" box="[941,1039,763,787]" gridcol="3" gridrow="13" pageId="10" pageNumber="496">15</td>
<td id="F0216744FF9AED93351319F9FB6FFCCC" box="[1069,1196,763,787]" gridcol="4" gridrow="13" pageId="10" pageNumber="496">137–202</td>
<td id="F0216744FF9AED93341519F9FA94FCCC" box="[1323,1367,763,787]" gridcol="5" gridrow="13" pageId="10" pageNumber="496">9</td>
</tr>
<tr id="B3F00E38FF9AED9331981823FA94FCE6" box="[166,1367,801,825]" gridrow="14" pageId="10" pageNumber="496">
<th id="F0216744FF9AED9331981823FA94FCE6" box="[166,1367,801,825]" colspan="6" colspanRight="5" gridcol="0" gridrow="14" pageId="10" pageNumber="496">Ratios</th>
</tr>
<tr id="B3F00E38FF9AED9331981845FA94FC80" box="[166,1367,839,863]" gridrow="15" pageId="10" pageNumber="496">
<th id="F0216744FF9AED9331981845FE5AFC80" box="[166,409,839,863]" gridcol="0" gridrow="15" pageId="10" pageNumber="496">a</th>
<td id="F0216744FF9AED9333551845FD07FC80" box="[619,708,839,863]" gridcol="1" gridrow="15" pageId="10" pageNumber="496">16</td>
<td id="F0216744FF9AED9332251845FC97FC80" box="[795,852,839,863]" gridcol="2" gridrow="15" pageId="10" pageNumber="496">16.8</td>
<td id="F0216744FF9AED9332931845FBCCFC80" box="[941,1039,839,863]" gridcol="3" gridrow="15" pageId="10" pageNumber="496">2.5</td>
<td id="F0216744FF9AED9335131845FB6FFC80" box="[1069,1196,839,863]" gridcol="4" gridrow="15" pageId="10" pageNumber="496">13–24</td>
<td id="F0216744FF9AED9334151845FA94FC80" box="[1323,1367,839,863]" gridcol="5" gridrow="15" pageId="10" pageNumber="496">14.9</td>
</tr>
<tr id="B3F00E38FF9AED933198186EFA94FC5A" box="[166,1367,876,901]" gridrow="16" pageId="10" pageNumber="496">
<th id="F0216744FF9AED933198186EFE5AFC5A" box="[166,409,876,901]" gridcol="0" gridrow="16" pageId="10" pageNumber="496">b</th>
<td id="F0216744FF9AED933355186EFD07FC5A" box="[619,708,876,901]" gridcol="1" gridrow="16" pageId="10" pageNumber="496">3.9</td>
<td id="F0216744FF9AED933225186EFC97FC5A" box="[795,852,876,901]" gridcol="2" gridrow="16" pageId="10" pageNumber="496">3</td>
<td id="F0216744FF9AED933293186EFBCCFC5A" box="[941,1039,876,901]" gridcol="3" gridrow="16" pageId="10" pageNumber="496">0.35</td>
<td id="F0216744FF9AED933513186EFB6FFC5A" box="[1069,1196,876,901]" gridcol="4" gridrow="16" pageId="10" pageNumber="496">2.6–3.9</td>
<td id="F0216744FF9AED933415186EFA94FC5A" box="[1323,1367,876,901]" gridcol="5" gridrow="16" pageId="10" pageNumber="496">11.7</td>
</tr>
<tr id="B3F00E38FF9AED9331981890FA94FC75" box="[166,1367,914,938]" gridrow="17" pageId="10" pageNumber="496">
<th id="F0216744FF9AED9331981890FE5AFC75" box="[166,409,914,938]" gridcol="0" gridrow="17" pageId="10" pageNumber="496">c</th>
<td id="F0216744FF9AED9333551890FD07FC75" box="[619,708,914,938]" gridcol="1" gridrow="17" pageId="10" pageNumber="496">13</td>
<td id="F0216744FF9AED9332251890FC97FC75" box="[795,852,914,938]" gridcol="2" gridrow="17" pageId="10" pageNumber="496">13.5</td>
<td id="F0216744FF9AED9332931890FBCCFC75" box="[941,1039,914,938]" gridcol="3" gridrow="17" pageId="10" pageNumber="496">1.4</td>
<td id="F0216744FF9AED9335131890FB6FFC75" box="[1069,1196,914,938]" gridcol="4" gridrow="17" pageId="10" pageNumber="496">11–17</td>
<td id="F0216744FF9AED9334151890FA94FC75" box="[1323,1367,914,938]" gridcol="5" gridrow="17" pageId="10" pageNumber="496">10.4</td>
</tr>
<tr id="B3F00E38FF9AED93319818BAFA94FC0F" box="[166,1367,952,976]" gridrow="18" pageId="10" pageNumber="496">
<th id="F0216744FF9AED93319818BAFE5AFC0F" box="[166,409,952,976]" gridcol="0" gridrow="18" pageId="10" pageNumber="496">c’</th>
<td id="F0216744FF9AED93335518BAFD07FC0F" box="[619,708,952,976]" gridcol="1" gridrow="18" pageId="10" pageNumber="496">92</td>
<td id="F0216744FF9AED93322518BAFC97FC0F" box="[795,852,952,976]" gridcol="2" gridrow="18" pageId="10" pageNumber="496">93</td>
<td id="F0216744FF9AED93329318BAFBCCFC0F" box="[941,1039,952,976]" gridcol="3" gridrow="18" pageId="10" pageNumber="496">0.73</td>
<td id="F0216744FF9AED93351318BAFB6FFC0F" box="[1069,1196,952,976]" gridcol="4" gridrow="18" pageId="10" pageNumber="496">91–94</td>
<td id="F0216744FF9AED93341518BAFA94FC0F" box="[1323,1367,952,976]" gridcol="5" gridrow="18" pageId="10" pageNumber="496">0.8</td>
</tr>
</table>
</paragraph>
<subSubSection id="45DA5FF1FF9A126031F918FAFDC2F972" lastPageId="12" lastPageNumber="498" pageId="10" pageNumber="496" type="diagnosis">
<paragraph id="0D7F0C7AFF9A126631F918FAFB0EFBA0" blockId="10.[151,1437,1016,2015]" pageId="10" pageNumber="496">
<emphasis id="3FB4D068FF9A126631F918FAFE11FBCD" bold="true" box="[199,466,1016,1042]" pageId="10" pageNumber="496">Differential diagnosis.</emphasis>
Females of
<taxonomicName id="CAC077F9FF9A1266335918FBFD28FBCC" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[615,747,1017,1043]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A1266335918FBFD28FBCC" box="[615,747,1017,1043]" italics="true" pageId="10" pageNumber="496">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126633CB18FBFCFBFBCC" bold="true" box="[757,824,1017,1043]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126633CB18FBFCFBFBCC" box="[757,824,1017,1043]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are characterized by smooth, knob-like cervical and somatic papillae, shallow cuticular shields, continuous cuticular collars with approximately 72 subequal spines, and two pairs of paired anterior and posterior lateral spines. Males lack ornate cuticular ornamentation, have a narrowed stomal opening and smooth somatic papillae, but otherwise are similar to other described males.
</paragraph>
<paragraph id="0D7F0C7AFF9A126631F91F8BFE9AF98B" blockId="10.[151,1437,1016,2015]" pageId="10" pageNumber="496">
The morphology of females of
<taxonomicName id="CAC077F9FF9A1266331B1F88FD49FB7C" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[549,650,1162,1187]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A1266331B1F88FD49FB7C" box="[549,650,1162,1187]" italics="true" pageId="10" pageNumber="496">H. pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126633AF1F8BFD13FB7C" bold="true" box="[657,720,1161,1187]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126633AF1F8BFD13FB7C" box="[657,720,1161,1187]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is most similar to those
<taxonomicName id="CAC077F9FF9A126632DD1F8BFBDAFB7C" authorityName="Cobb" authorityYear="1898" box="[995,1049,1161,1187]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="genus">
<emphasis id="3FB4D068FF9A126632DD1F8BFBDAFB7C" box="[995,1049,1161,1187]" italics="true" pageId="10" pageNumber="496">Heth</emphasis>
</taxonomicName>
spp. having a continuous collar of cervical spines with a spiny lateral shield.
<taxonomicName id="CAC077F9FF9A126633481FAFFD34FB18" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[630,759,1197,1223]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A126633481FAFFD34FB18" box="[630,759,1197,1223]" italics="true" pageId="10" pageNumber="496">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126633C01FAFFCFDFB18" bold="true" box="[766,830,1197,1223]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126633C01FAFFCFDFB18" box="[766,830,1197,1223]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is distinct from
<taxonomicName id="CAC077F9FF9A126632C61FACFAFBFB18" authority="Adamson, 1982" authorityName="Adamson" authorityYear="1982" box="[1016,1336,1197,1223]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="mauriesi">
<emphasis id="3FB4D068FF9A126632C61FACFBBEFB18" box="[1016,1149,1198,1223]" italics="true" pageId="10" pageNumber="496">H. mauriesi</emphasis>
<bibRefCitation id="6951718BFF9A126635BD1FAFFAFBFB18" author="Adamson, M. L." box="[1155,1336,1197,1223]" pageId="10" pageNumber="496" pagination="405 - 418" refId="ref18074" refString="Adamson, M. L. (1982) Two new species of Heth Cobb, 1898 (Nematoda, Rhigonematidae), from South American diplopods. Bulletin de Museum National de Histoire Naturelle 4 e Serie Section, A 4, 405 - 418." type="journal article" year="1982">Adamson, 1982</bibRefCitation>
</taxonomicName>
, its closest geographic neighbor, in female length (
<quantity id="CA38A19FFF9A126633431FD3FCF9FB34" box="[637,826,1233,1259]" metricMagnitude="0" metricUnit="m" metricValue="3.3364999999999996" metricValueMax="4.483" metricValueMin="2.19" pageId="10" pageNumber="496" unit="mm" value="3336.5" valueMax="4483.0" valueMin="2190.0">
<geoCoordinate id="68F46ABDFF9A126633431FD3FD7EFB34" box="[637,701,1233,1259]" degrees="2,190" direction="north" orientation="latitude" pageId="10" pageNumber="496" precision="55" value="2.19">2,190</geoCoordinate>
<geoCoordinate id="68F46ABDFF9A126633831FD3FCC8FB34" box="[701,779,1233,1259]" degrees="4,483" direction="west" orientation="longitude" pageId="10" pageNumber="496" precision="55" value="-4.483">–4,483</geoCoordinate>
µm
</quantity>
vs.
<quantity id="CA38A19FFF9A126632541FD3FBE7FB34" box="[874,1060,1233,1260]" metricMagnitude="0" metricUnit="m" metricValue="1.7875" metricValueMax="2.0" metricValueMin="1.575" pageId="10" pageNumber="496" unit="mm" value="1787.5" valueMax="2000.0" valueMin="1575.0">
<geoCoordinate id="68F46ABDFF9A126632541FD3FC64FB33" box="[874,935,1233,1260]" degrees="1,575" direction="north" orientation="latitude" pageId="10" pageNumber="496" precision="55" value="1.575">1,575</geoCoordinate>
<geoCoordinate id="68F46ABDFF9A126632991FD3FC35FB33" box="[935,1014,1233,1260]" degrees="2,000" direction="west" orientation="longitude" pageId="10" pageNumber="496" precision="55" value="-2.0">–2,000</geoCoordinate>
µm
</quantity>
in
<taxonomicName id="CAC077F9FF9A126635751FD0FB13FB34" authorityName="Adamson" authorityYear="1982" box="[1099,1232,1234,1259]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="mauriesi">
<emphasis id="3FB4D068FF9A126635751FD0FB13FB34" box="[1099,1232,1234,1259]" italics="true" pageId="10" pageNumber="496">H. mauriesi</emphasis>
</taxonomicName>
), smooth cervical and somatic papillae (multi-cusped in
<taxonomicName id="CAC077F9FF9A126633781FF4FD09FAD0" authorityName="Adamson" authorityYear="1982" box="[582,714,1270,1295]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="mauriesi">
<emphasis id="3FB4D068FF9A126633781FF4FD09FAD0" box="[582,714,1270,1295]" italics="true" pageId="10" pageNumber="496">H. mauriesi</emphasis>
</taxonomicName>
), and a trapezoidal shield interrupting the cervical collar (shield replaced by two stout spines in line with the smaller collar spines in
<taxonomicName id="CAC077F9FF9A1266329C1E18FBEBFAEC" authorityName="Adamson" authorityYear="1982" box="[930,1064,1306,1331]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="mauriesi">
<emphasis id="3FB4D068FF9A1266329C1E18FBEBFAEC" box="[930,1064,1306,1331]" italics="true" pageId="10" pageNumber="496">H. mauriesi</emphasis>
</taxonomicName>
) (
<figureCitation id="95FB10FFFF9A1266357F1E1BFB0BFAEC" box="[1089,1224,1305,1331]" captionStart="FIGURE 1" captionStartId="6.[151,250,1323,1347]" captionTargetBox="[166,1425,424,1300]" captionTargetId="figure-129@6.[161,1425,417,1300]" captionTargetPageId="6" captionText="FIGURE 1. Heth mauriesi female, terminology used to describe the anterior ornamentation of Heth spp. A) Drawings of H. mauriesi from Adamson (1982), courtesy of Dr. Martin Adamson. B) SEM of H. mauriesi, previously unpublished image courtesy of Dr. Ramon Carreno. Scale bars 50 µm." figureDoi="http://doi.org/10.5281/zenodo.4416663" httpUri="https://zenodo.org/record/4416663/files/figure.png" pageId="10" pageNumber="496">Figs. 1A, B</figureCitation>
). Among the other subtropical and tropical North American species,
<taxonomicName id="CAC077F9FF9A126633841E3CFC86FA88" authorityName="Mejia-Madrid" authorityYear="2014" box="[698,837,1341,1367]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="tuxtlensis">
<emphasis id="3FB4D068FF9A126633841E3CFC86FA88" box="[698,837,1341,1367]" italics="true" pageId="10" pageNumber="496">H. tuxtlensis</emphasis>
</taxonomicName>
(
<figureCitation id="95FB10FFFF9A1266326D1E3FFC76FA88" box="[851,949,1341,1367]" captionStart="FIGURE 12" captionStartId="23.[151,250,899,923]" captionTargetBox="[155,1432,186,869]" captionTargetId="figure-30@23.[151,1436,181,875]" captionTargetPageId="23" captionText="FIGURE 12. Worldwide Heth spp. Group 2 (continued). A) H. tonkinensis (after Malysheva &amp; Spiridonov 2010). B) H. tuxtlensis (after Mejía-Madrid 2014). C) H. vietnamensis (after Malysheva &amp; Spiridonov 2010). D) H. xaniophora (after Hunt 1994), en face, lateral and dorso-ventral views." figureDoi="http://doi.org/10.5281/zenodo.4416693" httpUri="https://zenodo.org/record/4416693/files/figure.png" pageId="10" pageNumber="496">Fig. 12B</figureCitation>
) is most similar to
<taxonomicName id="CAC077F9FF9A126635B91E3CFB28FA88" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[1159,1259,1342,1367]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A126635B91E3CFB28FA88" box="[1159,1259,1342,1367]" italics="true" pageId="10" pageNumber="496">H. pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126635CF1E3FFAEDFA88" bold="true" box="[1265,1326,1341,1367]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126635CF1E3FFAEDFA88" box="[1265,1326,1341,1367]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
in having a continuous cervical collar, the presence of lappets that usually do not overlap the anterior lateral spines, absence of lateral alae, and a finely striated body.
<taxonomicName id="CAC077F9FF9A1266335C1E87FD21FA40" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[610,738,1413,1439]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A1266335C1E87FD21FA40" box="[610,738,1413,1439]" italics="true" pageId="10" pageNumber="496">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126633D61E87FCE4FA40" bold="true" box="[744,807,1413,1439]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126633D61E87FCE4FA40" box="[744,807,1413,1439]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="CAC077F9FF9A126632621E84FC2BFA40" authorityName="Mejia-Madrid" authorityYear="2014" box="[860,1000,1413,1439]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="tuxtlensis">
<emphasis id="3FB4D068FF9A126632621E84FC2BFA40" box="[860,1000,1413,1439]" italics="true" pageId="10" pageNumber="496">H. tuxtlensis</emphasis>
</taxonomicName>
are easily differentiated by the number of cervical spines (
<emphasis id="3FB4D068FF9A126630541EA8FE4FFA1C" box="[362,396,1450,1475]" italics="true" pageId="10" pageNumber="496">ca.</emphasis>
72 for
<taxonomicName id="CAC077F9FF9A126630E21EA8FD82FA1C" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[476,577,1450,1475]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A126630E21EA8FD82FA1C" box="[476,577,1450,1475]" italics="true" pageId="10" pageNumber="496">H. pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126633791EABFD46FA1C" bold="true" box="[583,645,1449,1475]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126633791EABFD46FA1C" box="[583,645,1449,1475]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="3FB4D068FF9A126633AD1EA8FD76FA1C" box="[659,693,1450,1475]" italics="true" pageId="10" pageNumber="496">ca.</emphasis>
100 for
<taxonomicName id="CAC077F9FF9A1266322D1EA8FC5CFA1C" authorityName="Mejia-Madrid" authorityYear="2014" box="[787,927,1449,1475]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="tuxtlensis">
<emphasis id="3FB4D068FF9A1266322D1EA8FC5CFA1C" box="[787,927,1449,1475]" italics="true" pageId="10" pageNumber="496">H. tuxtlensis</emphasis>
</taxonomicName>
), a shallow cervical shield in
<taxonomicName id="CAC077F9FF9A126635D51EA8FA93FA1C" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[1259,1360,1450,1475]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A126635D51EA8FA93FA1C" box="[1259,1360,1450,1475]" italics="true" pageId="10" pageNumber="496">H. pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126634691EABFA56FA1C" bold="true" box="[1367,1429,1449,1475]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126634691EABFA56FA1C" box="[1367,1429,1449,1475]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, (more pronounced in
<taxonomicName id="CAC077F9FF9A126630B11ECCFDDEFA38" authorityName="Mejia-Madrid" authorityYear="2014" box="[399,541,1485,1511]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="tuxtlensis">
<emphasis id="3FB4D068FF9A126630B11ECCFDDEFA38" box="[399,541,1485,1511]" italics="true" pageId="10" pageNumber="496">H. tuxtlensis</emphasis>
</taxonomicName>
), more bifurcated lappet spines in
<taxonomicName id="CAC077F9FF9A126632901ECCFBFFFA38" authorityName="Mejia-Madrid" authorityYear="2014" box="[942,1084,1485,1511]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="tuxtlensis">
<emphasis id="3FB4D068FF9A126632901ECCFBFFFA38" box="[942,1084,1485,1511]" italics="true" pageId="10" pageNumber="496">H. tuxtlensis</emphasis>
</taxonomicName>
as compared to fewer cervical spine bifurcations in
<taxonomicName id="CAC077F9FF9A126630BC1EF0FE24F9D4" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[386,487,1522,1547]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A126630BC1EF0FE24F9D4" box="[386,487,1522,1547]" italics="true" pageId="10" pageNumber="496">H. pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126630D01EF3FDEFF9D4" bold="true" box="[494,556,1521,1547]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126630D01EF3FDEFF9D4" box="[494,556,1521,1547]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and smaller anterior and posterior lateral spines (
<quantity id="CA38A19FFF9A1266356B1EF3FB0AF9D4" box="[1109,1225,1521,1547]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="10" pageNumber="496" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 µm</quantity>
long) compared to the larger and more robust spines of
<taxonomicName id="CAC077F9FF9A126633121D14FD7BF9F0" authorityName="Mejia-Madrid" authorityYear="2014" box="[556,696,1557,1583]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="tuxtlensis">
<emphasis id="3FB4D068FF9A126633121D14FD7BF9F0" box="[556,696,1557,1583]" italics="true" pageId="10" pageNumber="496">H. tuxtlensis</emphasis>
</taxonomicName>
(anterior spine length
<quantity id="CA38A19FFF9A1266328C1D17FBE4F9F0" box="[946,1063,1557,1584]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="6.7" metricValueMin="5.8" pageId="10" pageNumber="496" unit="mm" value="62.5" valueMax="67.0" valueMin="58.0">58–67 µm</quantity>
, posterior spine length
<quantity id="CA38A19FFF9A126634141D17FA5FF9F0" box="[1322,1436,1557,1583]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" metricValueMax="9.4" metricValueMin="8.2" pageId="10" pageNumber="496" unit="mm" value="88.0" valueMax="94.0" valueMin="82.0">82–94 µm</quantity>
posterior spines).
</paragraph>
<paragraph id="0D7F0C7AFF9A126631F91D5FFB28F8F4" blockId="10.[151,1437,1016,2015]" pageId="10" pageNumber="496">
Females of
<taxonomicName id="CAC077F9FF9A126630741D5CFE6CF9A8" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[330,431,1630,1655]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A126630741D5CFE6CF9A8" box="[330,431,1630,1655]" italics="true" pageId="10" pageNumber="496">H. pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126630881D5FFE36F9A8" bold="true" box="[438,501,1629,1655]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126630881D5FFE36F9A8" box="[438,501,1629,1655]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
resemble those of
<taxonomicName id="CAC077F9FF9A126633F61D5CFC76F9A7" authority="(Vietnam)" baseAuthorityName="Vietnam" box="[712,949,1629,1656]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="taybaci">
<emphasis id="3FB4D068FF9A126633F61D5CFCFEF9A8" box="[712,829,1629,1655]" italics="true" pageId="10" pageNumber="496">H. taybaci</emphasis>
(
<collectingCountry id="75D74CEAFF9A126632751D5FFC6DF9A7" box="[843,942,1629,1656]" name="Vietnam" pageId="10" pageNumber="496">Vietnam</collectingCountry>
)
</taxonomicName>
in that the shields have similar shapes. The posteriormost spines of the cuticular collar nearly reach the base of the anterior lateral spines, and both have smooth somatic papillae.
<taxonomicName id="CAC077F9FF9A126630611DA7FE22F960" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[351,481,1701,1727]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A126630611DA7FE22F960" box="[351,481,1701,1727]" italics="true" pageId="10" pageNumber="496">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126630D41DA7FDE9F960" bold="true" box="[490,554,1701,1727]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126630D41DA7FDE9F960" box="[490,554,1701,1727]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has fewer collar spines (approximately 72) compared to about 88 around the circumference of
<taxonomicName id="CAC077F9FF9A126630601DC8FE10F93C" box="[350,467,1737,1763]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="taybaci">
<emphasis id="3FB4D068FF9A126630601DC8FE10F93C" box="[350,467,1737,1763]" italics="true" pageId="10" pageNumber="496">H. taybaci</emphasis>
</taxonomicName>
.
<taxonomicName id="CAC077F9FF9A126630DC1DCBFDA0F93C" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[482,611,1737,1763]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A126630DC1DCBFDA0F93C" box="[482,611,1737,1763]" italics="true" pageId="10" pageNumber="496">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126633551DCBFD68F93C" bold="true" box="[619,683,1737,1763]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126633551DCBFD68F93C" box="[619,683,1737,1763]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
also differs in that the anterior and posterior lateral spine pairs are smaller than those of
<emphasis id="3FB4D068FF9A126630B81DECFDC1F8D8" box="[390,514,1773,1799]" italics="true" pageId="10" pageNumber="496">
<taxonomicName id="CAC077F9FF9A126630B81DECFE3DF8D8" box="[390,510,1773,1799]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="taybaci">H. taybaci</taxonomicName>
.
</emphasis>
Males of
<taxonomicName id="CAC077F9FF9A1266334F1DECFD16F8D8" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[625,725,1774,1799]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A1266334F1DECFD16F8D8" box="[625,725,1774,1799]" italics="true" pageId="10" pageNumber="496">H. pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126633E21DEFFCD9F8D8" bold="true" box="[732,794,1773,1799]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126633E21DEFFCD9F8D8" box="[732,794,1773,1799]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
have smaller spicule arcs (mean
<quantity id="CA38A19FFF9A126635B51DEFFB20F8D8" box="[1163,1251,1773,1799]" metricMagnitude="-1" metricUnit="m" metricValue="1.09" pageId="10" pageNumber="496" unit="mm" value="109.0">109 µm</quantity>
) than
<taxonomicName id="CAC077F9FF9A126634161DECFA5FF8D8" box="[1320,1436,1773,1799]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="taybaci">
<emphasis id="3FB4D068FF9A126634161DECFA5FF8D8" box="[1320,1436,1773,1799]" italics="true" pageId="10" pageNumber="496">H. taybaci</emphasis>
</taxonomicName>
(
<quantity id="CA38A19FFF9A126631A11C13FF3AF8F4" box="[159,249,1809,1835]" metricMagnitude="-1" metricUnit="m" metricValue="1.3599999999999999" pageId="10" pageNumber="496" unit="mm" value="136.0">136 µm</quantity>
) but have longer bodies (
<taxonomicName id="CAC077F9FF9A126633281C13FD54F8F4" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[534,663,1809,1835]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A126633281C13FD54F8F4" box="[534,663,1809,1835]" italics="true" pageId="10" pageNumber="496">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126633A01C13FD1FF8F4" bold="true" box="[670,732,1809,1835]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126633A01C13FD1FF8F4" box="[670,732,1809,1835]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
males
<quantity id="CA38A19FFF9A126632121C13FC24F8F4" box="[812,999,1809,1835]" metricMagnitude="0" metricUnit="m" metricValue="2.253" metricValueMax="2.6089999999999995" metricValueMin="1.8969999999999998" pageId="10" pageNumber="496" unit="mm" value="2253.0" valueMax="2609.0" valueMin="1897.0">
<geoCoordinate id="68F46ABDFF9A126632121C13FCAAF8F4" box="[812,873,1809,1835]" degrees="1,897" direction="north" orientation="latitude" pageId="10" pageNumber="496" precision="55" value="1.897">1,897</geoCoordinate>
<geoCoordinate id="68F46ABDFF9A126632571C13FC7BF8F4" box="[873,952,1809,1835]" degrees="2,609" direction="west" orientation="longitude" pageId="10" pageNumber="496" precision="55" value="-2.609">–2,609</geoCoordinate>
µm
</quantity>
,
<taxonomicName id="CAC077F9FF9A126632CD1C10FBA4F8F4" box="[1011,1127,1809,1835]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="taybaci">
<emphasis id="3FB4D068FF9A126632CD1C10FBA4F8F4" box="[1011,1127,1809,1835]" italics="true" pageId="10" pageNumber="496">H. taybaci</emphasis>
</taxonomicName>
<quantity id="CA38A19FFF9A126635511C13FB1CF8F4" box="[1135,1247,1809,1836]" metricMagnitude="0" metricUnit="m" metricValue="1.52" pageId="10" pageNumber="496" unit="mm" value="1520.0">1,520 µm</quantity>
).
</paragraph>
<paragraph id="0D7F0C7AFF9A126731F91C37FF26FE9E" blockId="10.[151,1437,1016,2015]" lastBlockId="11.[151,1436,151,321]" lastPageId="11" lastPageNumber="497" pageId="10" pageNumber="496">
Two additional
<taxonomicName id="CAC077F9FF9A126630491C37FE6EF890" authorityName="Cobb" authorityYear="1898" box="[375,429,1845,1871]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="genus">
<emphasis id="3FB4D068FF9A126630491C37FE6EF890" box="[375,429,1845,1871]" italics="true" pageId="10" pageNumber="496">Heth</emphasis>
</taxonomicName>
species with a continuous shield-like collar occur in the western hemisphere:
<taxonomicName id="CAC077F9FF9A126634261C34FF2DF8AB" authority="(Brazil)" authorityName="Kloss" authorityYear="1965" baseAuthorityName="Brazil" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="insularis">
<emphasis id="3FB4D068FF9A126634261C34FA5FF890" box="[1304,1436,1845,1871]" italics="true" pageId="10" pageNumber="496">H. insularis</emphasis>
(
<collectingCountry id="75D74CEAFF9A126631A11C5BFF25F8AB" box="[159,230,1881,1908]" name="Brazil" pageId="10" pageNumber="496">Brazil</collectingCountry>
)
</taxonomicName>
and
<taxonomicName id="CAC077F9FF9A1266301C1C58FDF0F8AC" authority="(Paraguay)" authorityName="Adamson" authorityYear="1987" baseAuthorityName="Paraguay" box="[290,563,1881,1907]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="orthopori">
<emphasis id="3FB4D068FF9A1266301C1C58FE6DF8AC" box="[290,430,1881,1907]" italics="true" pageId="10" pageNumber="496">H. orthopori</emphasis>
(
<collectingCountry id="75D74CEAFF9A126630821C5BFDE8F8AC" box="[444,555,1881,1907]" name="Paraguay" pageId="10" pageNumber="496">Paraguay</collectingCountry>
)
</taxonomicName>
. In both of those species the lateral spine pairs are broadly joined at their bases. All other
<taxonomicName id="CAC077F9FF9A1266303B1C7FFEF8F848" authorityName="Cobb" authorityYear="1898" box="[261,315,1917,1943]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="genus">
<emphasis id="3FB4D068FF9A1266303B1C7FFEF8F848" box="[261,315,1917,1943]" italics="true" pageId="10" pageNumber="496">Heth</emphasis>
</taxonomicName>
species in this group are located in
<collectingCountry id="75D74CEAFF9A126633E81C7FFC83F848" box="[726,832,1917,1943]" name="Australia" pageId="10" pageNumber="496">Australia</collectingCountry>
,
<collectingCountry id="75D74CEAFF9A126632721C7FFC25F848" box="[844,998,1917,1943]" name="New Zealand" pageId="10" pageNumber="496">New Zealand</collectingCountry>
, and Asia.
<taxonomicName id="CAC077F9FF9A1266355A1C7FFB25F848" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[1124,1254,1917,1943]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9A1266355A1C7FFB25F848" box="[1124,1254,1917,1943]" italics="true" pageId="10" pageNumber="496">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9A126635D01C7FFAEDF848" bold="true" box="[1262,1326,1917,1943]" pageId="10" pageNumber="496">
<taxonomicNameLabel id="24876D13FF9A126635D01C7FFAEDF848" box="[1262,1326,1917,1943]" pageId="10" pageNumber="496" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has short lappets whose posteriormost spines do not reach the anterior lateral spines, whereas in
<taxonomicName id="CAC077F9FF9A1266354A1CA0FB2BF864" authorityName="Hunt" authorityYear="1994" box="[1140,1256,1954,1979]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="costata">
<emphasis id="3FB4D068FF9A1266354A1CA0FB2BF864" box="[1140,1256,1954,1979]" italics="true" pageId="10" pageNumber="496">H. costata</emphasis>
</taxonomicName>
,
<taxonomicName id="CAC077F9FF9A126635C61CA0FF76F800" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="impalutiensis">
<emphasis id="3FB4D068FF9A126635C61CA0FF76F800" italics="true" pageId="10" pageNumber="496">H. impalutiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="CAC077F9FF9A126631FA1CC4FEA4F800" authorityName="Hunt" authorityYear="1994" box="[196,359,1989,2015]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="sutherlandi">
<emphasis id="3FB4D068FF9A126631FA1CC4FEA4F800" box="[196,359,1989,2015]" italics="true" pageId="10" pageNumber="496">H. sutherlandi</emphasis>
</taxonomicName>
,
<taxonomicName id="CAC077F9FF9A126630481CC4FDD7F800" box="[374,532,1990,2015]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="taynguyeni">
<emphasis id="3FB4D068FF9A126630481CC4FDD7F800" box="[374,532,1990,2015]" italics="true" pageId="10" pageNumber="496">H. taynguyeni</emphasis>
</taxonomicName>
, and
<taxonomicName id="CAC077F9FF9A1266336D1CC4FCCBF800" authorityName="Malysheva &amp; Spiridonov" authorityYear="2010" box="[595,776,1990,2015]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="10" pageNumber="496" phylum="Nematoda" rank="species" species="vietnamensis">
<emphasis id="3FB4D068FF9A1266336D1CC4FCCBF800" box="[595,776,1990,2015]" italics="true" pageId="10" pageNumber="496">H. vietnamensis</emphasis>
</taxonomicName>
the spines of the much longer lappets overlap the anterior lateral spines.
<taxonomicName id="CAC077F9FF9B126730061B95FE0FFF6E" authorityName="Malysheva &amp; Cribb" authorityYear="2012" box="[312,460,151,177]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="11" pageNumber="497" phylum="Nematoda" rank="species" species="baudini">
<emphasis id="3FB4D068FF9B126730061B95FE0FFF6E" box="[312,460,151,177]" italics="true" pageId="11" pageNumber="497">Heth baudini</emphasis>
</taxonomicName>
and
<taxonomicName id="CAC077F9FF9B1267333C1B95FD65FF6E" authorityName="Chitwood" authorityYear="1935" box="[514,678,151,177]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="11" pageNumber="497" phylum="Nematoda" rank="species" species="dimorphum">
<emphasis id="3FB4D068FF9B1267333C1B95FD65FF6E" box="[514,678,151,177]" italics="true" pageId="11" pageNumber="497">H. dimorphum</emphasis>
</taxonomicName>
have shields in which length and width are about equal, whereas in
<taxonomicName id="CAC077F9FF9B126731A91BB9FF3FFF0B" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[151,252,187,212]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="11" pageNumber="497" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9B126731A91BB9FF3FFF0B" box="[151,252,187,212]" italics="true" pageId="11" pageNumber="497">H. pivari</emphasis>
</taxonomicName>
<taxonomicNameLabel id="24876D13FF9B1267303D1BB9FE81FF0A" box="[259,322,187,213]" pageId="11" pageNumber="497" rank="species">
<emphasis id="3FB4D068FF9B1267303D1BB9FEF8FF0A" bold="true" box="[259,315,187,213]" pageId="11" pageNumber="497">n. sp</emphasis>
.
</taxonomicNameLabel>
the shallow shield width is about twice its length. The
<typeStatus id="D27BB2D8FF9B126732921BB9FC1FFF0A" box="[940,988,187,213]" pageId="11" pageNumber="497">type</typeStatus>
species,
<taxonomicName id="CAC077F9FF9B1267357C1BB9FB4AFF0A" authorityName="Cobb" authorityYear="1898" box="[1090,1161,187,213]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="11" pageNumber="497" phylum="Nematoda" rank="species" species="juli">
<emphasis id="3FB4D068FF9B1267357C1BB9FB4AFF0A" box="[1090,1161,187,213]" italics="true" pageId="11" pageNumber="497">H. juli</emphasis>
</taxonomicName>
, has a “W” shape as opposed to a trapezoidal shape in
<taxonomicName id="CAC077F9FF9B126730CC1BDDFD94FF27" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[498,599,223,248]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="11" pageNumber="497" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9B126730CC1BDDFD94FF27" box="[498,599,223,248]" italics="true" pageId="11" pageNumber="497">H. pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9B126733631BDDFD58FF26" bold="true" box="[605,667,223,249]" pageId="11" pageNumber="497">
<taxonomicNameLabel id="24876D13FF9B126733631BDDFD58FF26" box="[605,667,223,249]" pageId="11" pageNumber="497" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Finally,
<taxonomicName id="CAC077F9FF9B126733C31BDDFC5CFF26" authorityName="Hunt" authorityYear="1994" box="[765,927,223,249]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="11" pageNumber="497" phylum="Nematoda" rank="species" species="xaniophora">
<emphasis id="3FB4D068FF9B126733C31BDDFC5CFF26" box="[765,927,223,249]" italics="true" pageId="11" pageNumber="497">H. xaniophora</emphasis>
</taxonomicName>
has prominent lateral alae and robust anterior and posterior lateral spines, whereas
<taxonomicName id="CAC077F9FF9B126733051A01FD62FEC3" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[571,673,259,284]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="11" pageNumber="497" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9B126733051A01FD62FEC3" box="[571,673,259,284]" italics="true" pageId="11" pageNumber="497">H. pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9B126733941A01FD29FEC2" bold="true" box="[682,746,259,285]" pageId="11" pageNumber="497">
<taxonomicNameLabel id="24876D13FF9B126733941A01FD29FEC2" box="[682,746,259,285]" pageId="11" pageNumber="497" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
lacks lateral alae and has small anterior and posterior lateral spines.
</paragraph>
<caption id="59BF5CF2FF9B126731A91C6AFEB5F817" ID-DOI="http://doi.org/10.5281/zenodo.4416670" ID-Zenodo-Dep="4416670" httpUri="https://zenodo.org/record/4416670/files/figure.png" pageId="11" pageNumber="497" startId="11.[151,250,1896,1920]" targetBox="[160,1426,354,1861]" targetPageId="11">
<paragraph id="0D7F0C7AFF9B126731A91C6AFEB5F817" blockId="11.[151,1436,1896,1992]" pageId="11" pageNumber="497">
<emphasis id="3FB4D068FF9B126731A91C6AFED7F85F" bold="true" box="[151,276,1896,1920]" pageId="11" pageNumber="497">FIGURE 4.</emphasis>
<taxonomicName id="CAC077F9FF9B126730251C6AFE53F85F" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[283,400,1896,1920]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="11" pageNumber="497" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9B126730251C6AFE53F85F" box="[283,400,1896,1920]" italics="true" pageId="11" pageNumber="497">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9B126730A91C6AFE12F85F" bold="true" box="[407,465,1896,1920]" pageId="11" pageNumber="497">
<taxonomicNameLabel id="24876D13FF9B126730A91C6AFE12F85F" box="[407,465,1896,1920]" pageId="11" pageNumber="497" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm.
</paragraph>
</caption>
<caption id="59BF5CF2FF9C126031A91E53FDD9FA0A" ID-DOI="http://doi.org/10.5281/zenodo.4416674" ID-Zenodo-Dep="4416674" httpUri="https://zenodo.org/record/4416674/files/figure.png" pageId="12" pageNumber="498" startId="12.[151,250,1361,1385]" targetBox="[155,1433,184,1334]" targetPageId="12">
<paragraph id="0D7F0C7AFF9C126031A91E53FDD9FA0A" blockId="12.[151,1437,1361,1493]" pageId="12" pageNumber="498">
<emphasis id="3FB4D068FF9C126031A91E53FED7FAB6" bold="true" box="[151,276,1361,1385]" pageId="12" pageNumber="498">FIGURE 5.</emphasis>
SEM micrographs of
<taxonomicName id="CAC077F9FF9C126030C41E53FDB3FAB6" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[506,624,1361,1385]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="12" pageNumber="498" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9C126030C41E53FDB3FAB6" box="[506,624,1361,1385]" italics="true" pageId="12" pageNumber="498">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9C126033491E53FD72FAB6" bold="true" box="[631,689,1361,1385]" pageId="12" pageNumber="498">
<taxonomicNameLabel id="24876D13FF9C126033491E53FD72FAB6" box="[631,689,1361,1385]" pageId="12" pageNumber="498" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
males. A) Anterior end, stoma slit-like. B) Anterior end, stoma open. C) Closeup of stoma showing striae and minute amphid aperture (a). D) Posterior region showing ventral sucker (S), papillae 1,2 and 4, and tail. E) Posterior region showing arrangement of posterior papillae 2–7. F) Closeup of cloacal region. Scale bars: A, B, E, 30 µm; C, F, 20 µm; D, 200 µm.
</paragraph>
</caption>
<paragraph id="0D7F0C7AFF9C126031F91D01FDC2F972" blockId="12.[151,1437,1539,1710]" pageId="12" pageNumber="498">
<emphasis id="3FB4D068FF9C126031F91D01FE6DF9C2" bold="true" box="[199,430,1539,1565]" pageId="12" pageNumber="498">Molecular analysis.</emphasis>
With the addition of
<taxonomicName id="CAC077F9FF9C126033A71D06FD3EF9C2" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[665,765,1540,1565]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="12" pageNumber="498" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9C126033A71D06FD3EF9C2" box="[665,765,1540,1565]" italics="true" pageId="12" pageNumber="498">H. pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9C1260323D1D01FC82F9C2" bold="true" box="[771,833,1539,1565]" pageId="12" pageNumber="498">
<taxonomicNameLabel id="24876D13FF9C1260323D1D01FC82F9C2" box="[771,833,1539,1565]" pageId="12" pageNumber="498" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="CAC077F9FF9C126032481D06FC3AF9C2" authorityName="Adamson" authorityYear="1982" box="[886,1017,1540,1565]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="12" pageNumber="498" phylum="Nematoda" rank="species" species="mauriesi">
<emphasis id="3FB4D068FF9C126032481D06FC3AF9C2" box="[886,1017,1540,1565]" italics="true" pageId="12" pageNumber="498">H. mauriesi</emphasis>
</taxonomicName>
, nine
<taxonomicName id="CAC077F9FF9C126035031D01FBB0F9C2" authorityName="Cobb" authorityYear="1898" box="[1085,1139,1539,1565]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="12" pageNumber="498" phylum="Nematoda" rank="genus">
<emphasis id="3FB4D068FF9C126035031D01FBB0F9C2" box="[1085,1139,1539,1565]" italics="true" pageId="12" pageNumber="498">Heth</emphasis>
</taxonomicName>
species were included in a 28S rDNA tree (
<figureCitation id="95FB10FFFF9C126030711D25FE50F99E" box="[335,403,1575,1601]" captionStart="FIGURE 8" captionStartId="15.[151,250,1932,1956]" captionTargetBox="[162,1417,241,1905]" captionTargetId="figure-66@15.[162,1425,237,1909]" captionTargetPageId="15" captionText="FIGURE 8. Molecular relationships among Heth spp. and other Rhigonematomorpha and Oxyuridomorpha using partial sequences of the D2-D3 expansion segment of 28S LSU rDNA. Bootstrap and posterior probability scores are placed above and below, respectively, corresponding nodes. XXXXXXXDNA sequence of H.mauriesi unpublished." figureDoi="http://doi.org/10.5281/zenodo.4416684" httpUri="https://zenodo.org/record/4416684/files/figure.png" pageId="12" pageNumber="498">Fig. 8</figureCitation>
).
<taxonomicName id="CAC077F9FF9C126030971D25FDEAF99E" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[425,553,1575,1601]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="12" pageNumber="498" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9C126030971D25FDEAF99E" box="[425,553,1575,1601]" italics="true" pageId="12" pageNumber="498">Heth pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9C1260330E1D25FDACF99E" bold="true" box="[560,623,1575,1601]" pageId="12" pageNumber="498">
<taxonomicNameLabel id="24876D13FF9C1260330E1D25FDACF99E" box="[560,623,1575,1601]" pageId="12" pageNumber="498" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and two Mexican species,
<taxonomicName id="CAC077F9FF9C126032A11D2AFBCCF99E" authorityName="Mejia-Madrid" authorityYear="2014" box="[927,1039,1575,1601]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="12" pageNumber="498" phylum="Nematoda" rank="species" species="gordae">
<emphasis id="3FB4D068FF9C126032A11D2AFBCCF99E" box="[927,1039,1575,1601]" italics="true" pageId="12" pageNumber="498">H. gordae</emphasis>
</taxonomicName>
and
<taxonomicName id="CAC077F9FF9C126035781D2AFB0CF99E" authorityName="Mejia-Madrid" authorityYear="2014" box="[1094,1231,1575,1601]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="12" pageNumber="498" phylum="Nematoda" rank="species" species="xarochae">
<emphasis id="3FB4D068FF9C126035781D2AFB0CF99E" box="[1094,1231,1575,1601]" italics="true" pageId="12" pageNumber="498">H. xarochae</emphasis>
</taxonomicName>
, grouped together with high support. The other six, including
<taxonomicName id="CAC077F9FF9C126033431D4EFCC1F9BA" authorityName="Adamson" authorityYear="1982" box="[637,770,1612,1637]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="12" pageNumber="498" phylum="Nematoda" rank="species" species="mauriesi">
<emphasis id="3FB4D068FF9C126033431D4EFCC1F9BA" box="[637,770,1612,1637]" italics="true" pageId="12" pageNumber="498">H. mauriesi</emphasis>
</taxonomicName>
, had no support except for
<emphasis id="3FB4D068FF9C1260350B1D4EFAA0F9BA" box="[1077,1379,1611,1638]" italics="true" pageId="12" pageNumber="498">
<taxonomicName id="CAC077F9FF9C1260350B1D4EFB08F9BA" box="[1077,1227,1612,1637]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="12" pageNumber="498" phylum="Nematoda" rank="species" species="initiaensis">H. initiaensis</taxonomicName>
-
<taxonomicName id="CAC077F9FF9C126035EA1D4EFAA0F9BA" authorityName="Malysheva &amp; Sumaya" authorityYear="2017" box="[1236,1379,1611,1637]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="12" pageNumber="498" phylum="Nematoda" rank="species" species="konoplevi">H. konoplevi</taxonomicName>
</emphasis>
. The molecular relationship of
<taxonomicName id="CAC077F9FF9C126030841D72FDE3F956" authority="Phillips &amp; Moulton &amp; Bernard, 2020" authorityName="Phillips &amp; Moulton &amp; Bernard" authorityYear="2020" box="[442,544,1648,1673]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="12" pageNumber="498" phylum="Nematoda" rank="species" species="pivari" status="sp. nov.">
<emphasis id="3FB4D068FF9C126030841D72FDE3F956" box="[442,544,1648,1673]" italics="true" pageId="12" pageNumber="498">H. pivari</emphasis>
</taxonomicName>
<emphasis id="3FB4D068FF9C126033161D6DFDABF956" bold="true" box="[552,616,1647,1673]" pageId="12" pageNumber="498">
<taxonomicNameLabel id="24876D13FF9C126033161D6DFDABF956" box="[552,616,1647,1673]" pageId="12" pageNumber="498" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
to
<taxonomicName id="CAC077F9FF9C126033B01D72FD3CF956" authorityName="Mejia-Madrid" authorityYear="2014" box="[654,767,1647,1673]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="12" pageNumber="498" phylum="Nematoda" rank="species" species="gordae">
<emphasis id="3FB4D068FF9C126033B01D72FD3CF956" box="[654,767,1647,1673]" italics="true" pageId="12" pageNumber="498">H. gordae</emphasis>
</taxonomicName>
and
<taxonomicName id="CAC077F9FF9C126032091D72FC02F956" authorityName="Mejia-Madrid" authorityYear="2014" box="[823,961,1647,1673]" class="Secernentea" family="Thelastomatidae" genus="Heth" kingdom="Animalia" order="Oxyurida" pageId="12" pageNumber="498" phylum="Nematoda" rank="species" species="xarochae">
<emphasis id="3FB4D068FF9C126032091D72FC02F956" box="[823,961,1647,1673]" italics="true" pageId="12" pageNumber="498">H. xarochae</emphasis>
</taxonomicName>
is supported by geographical location but not by morphological similarity.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink