183 lines
23 KiB
XML
183 lines
23 KiB
XML
<document id="E1EE44D12671293991CA351ED573982C" ID-DOI="http://dx.doi.org/10.3897/zookeys.719.21253" ID-GBIF-Dataset="b92acfc0-9d67-42c4-9e37-ebd5a53d24b5" ID-Pensoft-Pub="1313-2970-719-75" ID-ZooBank="36C4E2C8E07D4CC9A1D696B0FCE92CCF" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-719-75" ModsDocOrigin="ZooKeys 719" ModsDocTitle="A taxonomic revision of the subfamily Tillinae Leach sensu lato (Coleoptera, Cleridae) in the New World" checkinTime="1555319501509" checkinUser="pensoft" docAuthor="Burke, Alan & Zolnerowich, Gregory" docDate="2017" docId="6BF531F08F470C164182A08914FB8A1F" docLanguage="en" docName="ZooKeys 719: 75-157" docOrigin="ZooKeys 719" docSource="http://dx.doi.org/10.3897/zookeys.719.21253" docTitle="Lecontella striatopunctata Chevrolat 1876, comb. n." docType="treatment" docVersion="5" lastPageNumber="122" masterDocId="5C2D8118751DD235197A6569FFE59A6E" masterDocTitle="A taxonomic revision of the subfamily Tillinae Leach sensu lato (Coleoptera, Cleridae) in the New World" masterLastPageNumber="157" masterPageNumber="75" pageNumber="120" updateTime="1701317054126" updateUser="plazi">
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<mods:title id="7C036B156BC674C560B5CBA0A75EFFD0">A taxonomic revision of the subfamily Tillinae Leach sensu lato (Coleoptera, Cleridae) in the New World</mods:title>
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<mods:namePart id="CE3A76DF6E7098338EDE59DACB67C532">Burke, Alan</mods:namePart>
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<mods:namePart id="A778DEBC0DB135321141E061FA59C59C">Zolnerowich, Gregory</mods:namePart>
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<mods:date id="B5B3702464DA0E6414F19747DD81E0C2">2017</mods:date>
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<mods:identifier id="41B5C4336ABED3F242A6688A0E76F7C3" type="DOI">http://dx.doi.org/10.3897/zookeys.719.21253</mods:identifier>
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<treatment id="6BF531F08F470C164182A08914FB8A1F" ID-GBIF-Taxon="156200926" LSID="urn:lsid:plazi:treatment:6BF531F08F470C164182A08914FB8A1F" httpUri="http://treatment.plazi.org/id/6BF531F08F470C164182A08914FB8A1F" lastPageId="47" lastPageNumber="122" pageId="45" pageNumber="120">
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<subSubSection id="11918317918A78687FA3D92CF413C7DB" pageId="45" pageNumber="120" type="nomenclature">
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<paragraph id="2EC2934C2E9FA4F9BDE5AED5D08EC799" pageId="45" pageNumber="120">
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<taxonomicName id="7C31A948F72DB7E1E598D5D87ADAA205" authority="Chevrolat, 1876" authorityName="Chevrolat" authorityYear="1876" class="Insecta" family="Cleridae" genus="Lecontella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lecontella striatopunctata" order="Coleoptera" pageId="45" pageNumber="120" phylum="Arthropoda" rank="species" species="striatopunctata">Lecontella striatopunctata (Chevrolat, 1876)</taxonomicName>
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<taxonomicNameLabel id="E569BE0D65414E5F29D4A37A1C81FFF2" pageId="45" pageNumber="120">comb. n.</taxonomicNameLabel>
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Figs 4A, 10B
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</paragraph>
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</subSubSection>
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<subSubSection id="712DD2C6254B1E06CAAE30CA72121599" pageId="45" pageNumber="120" type="synonyms">
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<paragraph id="8A62996AEBAF10080B48383A8A1CBDC3" pageId="45" pageNumber="120">Synonyms.</paragraph>
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<paragraph id="3B4B98523736D6512399900A232EBC68" pageId="45" pageNumber="120">
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<taxonomicName id="BC0343ACD26EEF5FBC4F184C9511353D" class="Insecta" family="Cleridae" genus="Cymatodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatodera striatopunctata" order="Coleoptera" pageId="45" pageNumber="120" phylum="Arthropoda" rank="species" species="striatopunctata">Cymatodera striatopunctata</taxonomicName>
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Chevrolat, 1876,
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<normalizedToken id="BA4EAA82EB97FA8FEE481AA8E8DD5557" originalValue="Mémoire">Memoire</normalizedToken>
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sur la famille des
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<normalizedToken id="3B655A6AB1AA4205B7BB32E79AFE338D" originalValue="Clérites">Clerites</normalizedToken>
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. Buquet, Paris, 51 p.
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</paragraph>
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</subSubSection>
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<subSubSection id="6DA12E34AB94238D086C9C83708804F0" pageId="45" pageNumber="120" type="type material">
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<paragraph id="8008A2902DD0F1D17B5EAEE3F37C6230" pageId="45" pageNumber="120">Type material not examined.</paragraph>
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</subSubSection>
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<subSubSection id="8811F071E5220D1E888E2F8BD3962281" pageId="45" pageNumber="120" type="type locality">
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<paragraph id="1910A95D21825F304162A22575D571D8" pageId="45" pageNumber="120">Type locality.</paragraph>
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<paragraph id="D5B6E8C64F6DD1403CE7945DE587D6D1" pageId="45" pageNumber="120">
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<normalizedToken id="F02444D6BFBE4CFCD4780108274DF440" originalValue="“Mexique”">"Mexique"</normalizedToken>
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. Type depository:
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<normalizedToken id="EB9A35DEA6E988BB534DC3F90ED97212" originalValue="Muséum">Museum</normalizedToken>
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National
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<normalizedToken id="DBEF52A884C125C6A4ACAACA04AF3487" originalValue="d’Histoire">d'Histoire</normalizedToken>
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Naturelle, Paris (MNHN).
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</paragraph>
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</subSubSection>
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<subSubSection id="EEEADAE7E0C6AA8EE252FE3660EC4FCD" pageId="45" pageNumber="120" type="distribution">
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<paragraph id="4C29F2BF7382EC5A897D91F5982035F6" pageId="45" pageNumber="120">Distribution.</paragraph>
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<paragraph id="C665E03D2F0BCC32FA672CBDB7472729" pageId="45" pageNumber="120">Mexico: Jalisco, Morelos, Nayarit, Oaxaca; Central America: Guatemala.</paragraph>
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</subSubSection>
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<subSubSection id="1E6C97819174ADAC14C2D8C394FB77BC" pageId="45" pageNumber="120" type="differential diagnosis">
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<paragraph id="97D0FA4A256B3C86A87912329E07ED31" pageId="45" pageNumber="120">Differential diagnosis.</paragraph>
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<paragraph id="6156F48BD9EE8827C8443FA95757C2C5" pageId="45" pageNumber="120">
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<taxonomicName id="33E91D16F4C6417F1F388F648F3D38ED" class="Insecta" family="Cleridae" genus="Lecontella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lecontella striatopunctata" order="Coleoptera" pageId="45" pageNumber="120" phylum="Arthropoda" rank="species" species="striatopunctata">Lecontella striatopunctata</taxonomicName>
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is most similar to
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<taxonomicName id="376ADAD2908F1B43E8F616A383AF8B26" lsidName="L. brunnea" pageId="45" pageNumber="120" rank="species" species="brunnea">L. brunnea</taxonomicName>
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. The two species are parapatric in distribution and can be misidentified as each other; however, males of
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<taxonomicName id="C245F94D0D39A7B960FF42295CE38BC4" lsidName="L. striatopunctata" pageId="45" pageNumber="120" rank="species" species="striatopunctata">L. striatopunctata</taxonomicName>
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have the antennomeres 2-10 conspicuously compacted and robust, and the eleventh antennomere is 3-4
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<normalizedToken id="F8B4B539E2AFEA5DD293F7ECCEF88F0F" originalValue="×">x</normalizedToken>
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the length of tenth antennomere (Fig. 10B). Males of
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<taxonomicName id="E6992128BB4B124E975BB19509A91875" lsidName="L. brunnea" pageId="45" pageNumber="120" rank="species" species="brunnea">L. brunnea</taxonomicName>
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have the antennae compacted, antennomeres 3-10 gradually increasing in width toward the distal end, and the eleventh antennomere is 4-5
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<normalizedToken id="56ADD3634F09DE5ECA7CB013D647E203" originalValue="×">x</normalizedToken>
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the length of the tenth antennomere (Fig. 9F). Males and females of these species can also be differentiated based on the shape of the epipleural fold.
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<taxonomicName id="1AAFC22AF717D0FBF2BA4A4DC95E432A" class="Insecta" family="Cleridae" genus="Lecontella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lecontella striatopunctata" order="Coleoptera" pageId="45" pageNumber="120" phylum="Arthropoda" rank="species" species="striatopunctata">Lecontella striatopunctata</taxonomicName>
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has the posterior portion of the epipleural fold smooth, while
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<taxonomicName id="2B3C15E59D879FE5E581D95C09E2EA55" lsidName="L. brunnea" pageId="45" pageNumber="120" rank="species" species="brunnea">L. brunnea</taxonomicName>
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has the same portion of the epipleural fold moderately crenulate.
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</paragraph>
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</subSubSection>
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<subSubSection id="778A58D9050F17D1C537B84AAA32006B" lastPageId="47" lastPageNumber="122" pageId="45" pageNumber="120" type="redescription">
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<paragraph id="942F73AA53FCCC1E5550EB77D8386834" pageId="45" pageNumber="120">Redescription.</paragraph>
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<paragraph id="9AD4B0C072903549DACCA0BE36E40373" pageId="45" pageNumber="120">Male. Form: Small to large individuals, moderately robust. Color: Head, pronotum, thorax, scutellum, legs, antennae and elytra light testaceous to dark brown; mouthparts fuscous, last fourth of mandibles piceous to black; abdominal segments testaceous to piceous; elytral disc devoid of any bands or fasciae (Fig. 4A).</paragraph>
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<caption id="FA82F5282DC54A4BA7538FFFB4A54DA7" pageId="45" pageNumber="120">
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<paragraph id="DD981FE19B9A5A2BA291981022AFC714" pageId="45" pageNumber="120">
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Figure 4. Habitus of: A
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<taxonomicName id="E2A0DDCA692EF4E1991E7A9C195D12B4" class="Insecta" family="Cleridae" genus="Lecontella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lecontella striatopunctata" order="Coleoptera" pageId="45" pageNumber="120" phylum="Arthropoda" rank="species" species="striatopunctata">Lecontella striatopunctata</taxonomicName>
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comb. n. B
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<taxonomicName id="046C9A41ABA63940BED1EFEB5E79D72C" class="Insecta" family="Cleridae" genus="Monophylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monophylla californica" order="Coleoptera" pageId="45" pageNumber="120" phylum="Arthropoda" rank="species" species="californica">Monophylla californica</taxonomicName>
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C
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<taxonomicName id="DA199D730ACA55AABCFA85D84B7AD04A" class="Insecta" family="Cleridae" genus="Monophylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monophylla pallipes" order="Coleoptera" pageId="45" pageNumber="120" phylum="Arthropoda" rank="species" species="pallipes">Monophylla pallipes</taxonomicName>
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D
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<taxonomicName id="8F9B2BD47C95087F636717BD24395B3E" class="Insecta" family="Cleridae" genus="Monophylla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Monophylla terminata" order="Coleoptera" pageId="45" pageNumber="120" phylum="Arthropoda" rank="species" species="terminata">Monophylla terminata</taxonomicName>
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E
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<taxonomicName id="652CC3106FEA475073A0E06ED60BB9A0" class="Insecta" family="Cleridae" genus="Teloclerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Teloclerus compresicornis" order="Coleoptera" pageId="45" pageNumber="120" phylum="Arthropoda" rank="species" species="compresicornis">Teloclerus compresicornis</taxonomicName>
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F
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<taxonomicName id="4F6AB0A0787699BD6A40BA7F5B463B66" class="Insecta" family="Cleridae" genus="Cymatodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatodera bogcioides" order="Coleoptera" pageId="45" pageNumber="120" phylum="Arthropoda" rank="species" species="bogcioides">Cymatodera bogcioides</taxonomicName>
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.
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</paragraph>
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</caption>
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<paragraph id="F84A4DE805B66E0D61BBB29EEA725714" lastPageId="46" lastPageNumber="121" pageId="45" pageNumber="120">
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Head: Including eyes wider than pronotum; eyes large, taller than wide, bulging laterally, coarsely faceted, emarginate posteriorly; antennal notch located in front of emargination; frons bi-impressed; integument coarsely, conspicuously, shallowly punc
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<pageBreakToken id="3DF732F404059134ED8FFA4495AAFB56" pageId="46" pageNumber="121" start="start">tate</pageBreakToken>
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; clothed with fine, pale, recumbent setae intermixed with some erect, pale setae; antennae consisting of 11 antennomeres; second antennomere moderately robust, slightly shorter than third antennomere; antennomeres 3-10 serrate, conspicuously robust and compacted, about the same length; last antennomere of males sexually dimorphic, conspicuously elongate, somewhat robust, parallel, cylindrical, posterior portion rounded 4-5
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<normalizedToken id="0A23C1DAA8E7F189425305BD7C3E3E70" originalValue="×">x</normalizedToken>
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longer than length of tenth antennomere.
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</paragraph>
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<paragraph id="76B4E1EBC189F82D476CFBBE0A06DF2C" pageId="46" pageNumber="121">Thorax: Pronotum bisinuate, widest at middle, slightly short in length; sides constricted subapically, more strongly constricted behind middle and feebly constricted in front of middle; surface conspicuously punctate, elytral disc with punctations small, shallow; clothed with fine, short, pale, recumbent setae interspersed with some long, semierect, fine, pale setae; long setae more abundant on anterior and lateral area of pronotum; anterior transverse depression present; subbasal tumescence absent; posterior margin of pronotum feebly compressed. Prosternum conspicuously wider than long; moderately to strongly punctate, punctation fine, deep; surface vested to glabrous. Mesoventrite surface smooth, vested with fine, pale, semi-erect setae; moderately to coarsely punctate, punctations wide, deep. Metaventrite surface smooth to finely rugulose, convex; numerously, rather punctate, punctations coarse, shallow; clothed with fine, pale, recumbent setae; longitudinal depression and metaventral process present. Scutellum wide, clothed with pale, fine, semirecumbent setae, compressed medially.</paragraph>
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<paragraph id="E3FEDCD318211296317C46E34414BB33" pageId="46" pageNumber="121">
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Elytra: Broader than pronotum; elongate; humeri indicated, rounded; sides inconspicuously broadening toward distal end, broadest on posterior 1/4, then abruptly narrowing toward apex at posterior 1/4; surface rugose to rugulose at interstices; elytral apices subtriangular; inconspicuously dehiscent; elytral declivity moderately steep; surface vested with fine, short, pale, recumbent setae and some pale, fine, long, erect setae; conspicuously, coarsely punctate, punctations arranged in regular striae; sculpturing consisting of coarse, deep, wide punctations arranged in regular striae that decrease in size in posterior fourth, punctation reaching elytral apex; interstices at elytral base 0.5
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<normalizedToken id="C0BCA46B57C5FD28016AF2589268EDDC" originalValue="×">x</normalizedToken>
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the width of punctation; interstices smooth. Epipleural fold gradually narrowing toward apex, not crenulate.
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</paragraph>
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<paragraph id="589B94CA2D70142AD1629CDF485523F8" pageId="46" pageNumber="121">Legs: Femora rugose; slightly swollen on distal end; clothed with some pale, fine, semirecumbent setae mixed with some semi-erect setae; surface conspicuously punctate; punctations small, shallow. Tibiae rugulose, punctate; punctations shallow and small; vestiture consisting of fine, recumbent and semirecumbent setae.</paragraph>
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<paragraph id="043E9230A58DB99E118BCBD763AF8765" lastPageId="47" lastPageNumber="122" pageId="46" pageNumber="121">
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Abdomen: Six visible ventrites. First ventrite rugulose. Ventrites 2-4 moderately to strongly rugulose, convex, subquadrate, punctate, vested with fine, long, pale, recumbent setae; not compressed laterally; posterior margins truncate. Posterior margin of first and second visible ventrites elevated with a transverse carina, this carina originating next to posterolateral angles producing a broad, elevated, arcuate emargination. Fifth visible ventrite subtriangular; surface rugulose, convex, moderately punctate, punctations shallow, small; vested with fine, pale, recumbent setae; lateral margins oblique, feebly arcuate; posterior margin broadly, shallowly emarginate. Sixth visible ventrite small, shape subtriangular; rugulose to rugose; surface convex; finely punctate; clothed with short, pale, fine, recumbent setae; as broad as long; lateral margins strongly oblique, arcuate; posterior margin short, somewhat acuminate, very shallowly emarginate. Fifth
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<pageBreakToken id="BCB83FFA277B3EE12C76EE27B22CA413" pageId="47" pageNumber="122" start="start">tergite</pageBreakToken>
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subquadrate, surface convex, rugulose; posterior margin truncate. Sixth tergite subtriangular; finely to moderately rugulose; surface moderately convex; clothed with fine, pale, recumbent setae; lateral margins oblique; posterior margin truncate. Sixth tergite extending beyond apical margin of sixth visible ventrite, covering sixth ventrite in dorsal view.
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</paragraph>
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<paragraph id="2DAA6B212C64A7C3EB0F3E4A04F0BB84" pageId="47" pageNumber="122">Aedeagus: Phallobasic apodeme present; phallus with copulatory piece swollen at apex; phallic plate unarmed, devoid of denticles; intraspicular plate present, elongate; phallobasic apodeme long, expanded distally; phallobase subparallel; parameres free; tegmen complete, covering phallus; parameres pointed distally; endophallic struts long, slender distally (Fig. 20A).</paragraph>
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<paragraph id="97BBC73F5294BBFA5F00F0628081003E" pageId="47" pageNumber="122">
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Sexual dimorphism: Females of
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<taxonomicName id="084D0949AD4357942211C2698C4100C5" lsidName="L. striatopunctata" pageId="47" pageNumber="122" rank="species" species="striatopunctata">L. striatopunctata</taxonomicName>
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differ from males in the shape of the last abdominal segment. Females have the lateral and posterior margins of the sixth abdominal segment broadly rounded. Males have the lateral margins of the sixth abdominal segment subtriangular in shape, strongly oblique, and the posterior margin is short, almost acuminate and shallowly emarginate. In addition, females have the eleventh antennomere short, robust, obtusely rounded, and approximately 2
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<normalizedToken id="BBD2DC58E7E6C21E9675AE168A639EE0" originalValue="×">x</normalizedToken>
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longer than tenth antennomere, while males have the same antennomere cylindrical, not compressed medially and 4-5
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<normalizedToken id="5451C5D9C477C457C38DF75C384637AD" originalValue="×">x</normalizedToken>
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longer than the tenth antennomere.
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</paragraph>
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</subSubSection>
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<subSubSection id="CAC89A29064AD81F84776927532B3264" pageId="47" pageNumber="122" type="materials_examined">
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<paragraph id="4967B420FB920944DE7AA99E9D4D2A44" pageId="47" pageNumber="122">Material examined.</paragraph>
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<paragraph id="D119E4AD72220F5578D0189B2D914FE1" pageId="47" pageNumber="122">
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1 male, 1 female: Morelos, Mexico, Tepalcingo, N El
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<normalizedToken id="711915C0087F249E29305AD7404C8F71" originalValue="Limón">Limon</normalizedToken>
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,
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<geoCoordinate id="BABB0A050C21B59602C60FC89443A112" direction="north" orientation="latitude" precision="1" value="18.538416">18°32'18.3"N</geoCoordinate>
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<geoCoordinate id="D0D9780018B0F8AD534D50691A8280E4" direction="west" orientation="longitude" precision="1" value="-98.93381">98°56'01.7"W</geoCoordinate>
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, 1272 m, Selva baja caducifolia, trampa de luz, VI-6-2008, M. De
|
||
<normalizedToken id="98E528429F88D9D5F949ED3788C88FA5" originalValue="León">Leon</normalizedToken>
|
||
; 1 male: Mexico, Jalisco, Estacion Biologica Chamela, VIII-1-2-1991, E. Giesbert; 1 female: Guerrero, Mexico, Highway 95, 5.6 km S Milpillas, [no collector data]; 3 females: Jalisco, Mexico, Estacion Biologica Chamela, VII-10-20-1985, E. Giesbert; 1 male, 1 female: Jalisco, Mexico, Estacion Biologica Chamela, X-1-2-1991, E. Giesbert; 1 male: El Progreso, Guatemala, Highway 17, vic.
|
||
<normalizedToken id="88B2D1DE2F15EA92BB3E278F49577E39" originalValue="Morazán">Morazan</normalizedToken>
|
||
, 1700', V-29 to VI-2-1989, E. Giesbert; 1 female: Zacapa, Guatemala, 12-14 km S San Lorenzo, 1000-2000', VI-3-6-1989, E. Giesbert; 1 male: Jalisco, Mexico, Estacion Biologica Chamela, X-15-21-1987, E. Giesbert; 1 male, 1 female: Mex., Jalisco, Mexico, Chamela Estacion UNAM, X-1-2-1991, J. E. Wappes; 2 males: Jalisco, Mexico, Chamela vic. E. B. UNAM, VII-9-19-1993, J. E. Wappes; 3 females: Jalisco, Mexico, vic Chamela, 15-VII-1990, J. E. Wappes; 1 female: Zacapa, Guatemala, 12-14 km S San Lorenzo, 1-2000', VI-3-6-1989; 1 female: Mexico, Jalisco, 17.6 km N Chamela, VII-16-1987, R. Turnbow, 2 males: Mexico, Jalisco, vic Estacion de Biologia Chamela UNAM, VII-9-14-1993, Black Light, Morris, Huether, Wappes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4CC4578731A2FC00C1D805D7553B1FB5" pageId="47" pageNumber="122" type="remarks">
|
||
<paragraph id="4E40C7C7935F42AFEFDC85CF85FE2C86" pageId="47" pageNumber="122">Remarks.</paragraph>
|
||
<paragraph id="DD0CF95F99878AEFC9CFBA041D2E1E6C" pageId="47" pageNumber="122">
|
||
<bibRefCitation id="D349EF28B6967485C9B38CA53DEFF7BB" author="Chevrolat, MA" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="75" pageNumber="150" pagination="1 - 51" title="Memoire sur la famille des Clerites." volume="5" year="1876">Chevrolat (1876)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="DCD9C6B5D7D40803439FB7FEBA84C086" class="Insecta" family="Cleridae" genus="Cymatodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatodera striatopunctata" order="Coleoptera" pageId="47" pageNumber="122" phylum="Arthropoda" rank="species" species="striatopunctata">Cymatodera striatopunctata</taxonomicName>
|
||
from material collected in Mexico (type locality not specified). This species is here transferred to
|
||
<taxonomicName id="40380A48829DF3C7F51E446A6020025F" class="Insecta" family="Cleridae" genus="Lecontella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lecontella" order="Coleoptera" pageId="47" pageNumber="122" phylum="Arthropoda" rank="genus">Lecontella</taxonomicName>
|
||
based on the compacted and serrate condition of the antennae (Fig. 10B), the broadly and deeply excavated elytral punctations that reach the elytral apex (Fig. 4A), and the overall similarities observed in the aedeagus of
|
||
<taxonomicName id="B8D08D2E84DDF34BCC080BA7DEC0B4B6" lsidName="C. striatopunctata" pageId="47" pageNumber="122" rank="species" species="striatopunctata">C. striatopunctata</taxonomicName>
|
||
(Fig. 20A),
|
||
<taxonomicName id="8E197C5271F4A497C9EDC2EB165256A7" lsidName="L. brunnea" pageId="47" pageNumber="122" rank="species" species="brunnea">L. brunnea</taxonomicName>
|
||
(Fig. 19E) and
|
||
<taxonomicName id="7191038BC44C3362E1AB422D1137066D" lsidName="L. gnara" pageId="47" pageNumber="122" rank="species" species="gnara">L. gnara</taxonomicName>
|
||
(Fig. 19F). Other characters similar in these species are the uniformly brown to dark brown integument color, the moderately to strongly punctate pronotum, and the male pygidium subtriangular in shape.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |