1036 lines
114 KiB
XML
1036 lines
114 KiB
XML
<document id="294B2D698878A2497F05910FFB3E53A2" ID-CLB-Dataset="45691" ID-DOI="10.5281/zenodo.212548" ID-GBIF-Dataset="78d2027b-125a-47be-adf9-b250221340be" ID-ISSN="1175-5326" ID-Zenodo-Dep="212548" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460425492893" checkinUser="plazi" docAuthor="Matsui, Masafumi, Nishikawa, Kanto, Belabut, Daicus M., Norhayati, Ahmad & Yong, Hoi Sen" docDate="2012" docId="3B3E633AFFA6FF8CFF40FB20FD64F999" docLanguage="en" docName="zt03155p046.pdf" docOrigin="Zootaxa 3155" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Kalophrynus limbooliati Matsui, Nishikawa, Belabut, Norhayati & Yong, 2012, sp. nov." docType="treatment" docVersion="9" lastPageNumber="44" masterDocId="C7071B42FFA7FF8AFFD7FFCDFF91FFFD" masterDocTitle="A new species of Kalophrynus (Amphibia, Anura, Microhylidae) from Southern Peninsular Malaysia" masterLastPageNumber="46" masterPageNumber="38" pageNumber="39" updateTime="1698330612502" updateUser="ExternalLinkService">
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<mods:mods id="D51BCDC622AEF5E0150176EFE50A3AB8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6E4F2DAE10ABF940F9D7B4544A7F416E">
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<mods:title id="C94203CD43838D0CCE986389CBDB6268">A new species of Kalophrynus (Amphibia, Anura, Microhylidae) from Southern Peninsular Malaysia</mods:title>
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</mods:titleInfo>
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<mods:name id="85C54E7E96CD3A34898E4C531E969CD1" type="personal">
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<mods:role id="F70834AC8CAC2E5DC368B845CA7C223A">
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<mods:roleTerm id="9193B0498A6950F39BF40446C8E7D428">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="75A5E29EC491A689986C42AFF9A7BF1C">Matsui, Masafumi</mods:namePart>
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</mods:name>
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<mods:name id="5D95EB083CBA8A67627F7DC18860770A" type="personal">
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<mods:role id="D110D8F78E5CA0613CA7AE522C68F793">
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<mods:roleTerm id="FF46DA46CB6AC367557046F8A474F071">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0B6F172B548E9F64E0B54818EE9334DE">Nishikawa, Kanto</mods:namePart>
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</mods:name>
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<mods:name id="E835C5716C3B1758030F2D87844E5B06" type="personal">
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<mods:roleTerm id="C8557D713EA9CFC457ED16BF373A4094">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CB788E5AF95E61B366F29D3A9F1FEA7E">Belabut, Daicus M.</mods:namePart>
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</mods:name>
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<mods:name id="DD1C487E088195C9CD49C67A233B5596" type="personal">
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<mods:roleTerm id="8A78B53E826826712A7D3D70CEDE81F4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F1BD63F1E1EF6B9CA5D6479372C60937">Norhayati, Ahmad</mods:namePart>
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</mods:name>
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<mods:name id="5A9AAE25195A0AEFC687980EFB36C7DF" type="personal">
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<mods:role id="EF393C807F60343AE29B6CBFAC633DB9">
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<mods:roleTerm id="C18F661A4A1A15B976A096D5831BD9CD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="18033A2EDB646E398587275CA1C827B7">Yong, Hoi Sen</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="41B3F75DFB98A59096EA393FB48C70A1">text</mods:typeOfResource>
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<mods:title id="ED6BC253A33C0577B2D737353FEF5E1D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="86F534EE0E81D029CDD54C8EE3D13E76">
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<mods:date id="2A0A57D725635B376F629719B42800D2">2012</mods:date>
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<mods:detail id="2FE5A46C1395E76E3E846056750FA494" type="volume">
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<mods:number id="C310F63B968208FA1B7B8CDC746082FD">3155</mods:number>
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<mods:end id="77C84F742D79EB893422A5CD1596E201">46</mods:end>
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<mods:classification id="6D9D82EE3DEEF5FB0F4A9ADF5E5F1160">journal article</mods:classification>
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<mods:identifier id="F80E0B4CCB14A581AEF5CBA1CB215F95" type="CLB-Dataset">45691</mods:identifier>
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<mods:identifier id="ADEB953A35C99A79D89F74D548B7BE98" type="DOI">10.5281/zenodo.212548</mods:identifier>
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<mods:identifier id="FBEAE42C3719F2328672334D271C37C9" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="AC6D122E2E3510F12A82E7612A18E1AC" type="Zenodo-Dep">212548</mods:identifier>
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<treatment id="3B3E633AFFA6FF8CFF40FB20FD64F999" ID-DOI="http://doi.org/10.5281/zenodo.6166891" ID-GBIF-Taxon="119428279" ID-Zenodo-Dep="6166891" LSID="urn:lsid:plazi:treatment:3B3E633AFFA6FF8CFF40FB20FD64F999" httpUri="http://treatment.plazi.org/id/3B3E633AFFA6FF8CFF40FB20FD64F999" lastPageId="6" lastPageNumber="44" pageId="1" pageNumber="39">
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<subSubSection id="FB8D81A7FFA6FF8BFF40FB20FBAFFA93" pageId="1" pageNumber="39" type="nomenclature">
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<paragraph id="B328D22CFFA6FF8BFF40FB20FDB0FAFA" blockId="1.[151,545,1261,1319]" box="[151,545,1261,1287]" pageId="1" pageNumber="39">
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<heading id="E8606540FFA6FF8BFF40FB20FDB0FAFA" bold="true" box="[151,545,1261,1287]" fontSize="11" level="1" pageId="1" pageNumber="39" reason="1">
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<emphasis id="81E30E3EFFA6FF8BFF40FB20FDB0FAFA" bold="true" box="[151,545,1261,1287]" pageId="1" pageNumber="39">
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<taxonomicName id="7497A9AFFFA6FF8BFF40FB20FE2CFAFA" ID-CoL="6NK56" box="[151,445,1261,1287]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="39" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA6FF8BFF40FB20FE2CFAFA" bold="true" box="[151,445,1261,1287]" italics="true" pageId="1" pageNumber="39">Kalophrynus limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA6FF8BFE13FB20FDB0FAFA" box="[452,545,1261,1287]" pageId="1" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA6FF8BFF40FAC2FE83FADA" blockId="1.[151,545,1261,1319]" box="[151,274,1295,1319]" pageId="1" pageNumber="39">
|
||
(
|
||
<figureCitation id="2BACCEA9FFA6FF8BFF48FAC2FE98FADA" box="[159,265,1295,1319]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,249,1447,1470]" captionStartId-1="3.[151,249,830,853]" captionStartId-2="3.[151,249,1654,1677]" captionTargetBox-0="[450,1136,367,1425]" captionTargetBox-1="[415,1171,183,808]" captionTargetBox-2="[340,1247,932,1632]" captionTargetId-0="figure@2.[450,1136,367,1426]" captionTargetId-1="figure@3.[415,1171,183,809]" captionTargetId-2="figure@3.[340,1248,932,1632]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="FIGURE 1. (A) Dorsal and (B) ventral views of the holotype of Kalophrynus limbooliati sp. nov. (male, UKMHC 705). Scale bar, 10 mm." captionText-1="FIGURE 2. Ventral views of (A) hand and (B) foot of the holotype of Kalophrynus limbooliati sp. nov. (male, UKMHC 705). Scale bar, 5 mm." captionText-2="FIGURE 3. Dorsolateral view of the holotype of Kalophrynus limbooliati sp. nov. (male, UKMHC 705) from Gunung Pulai, Johor." httpUri-0="https://zenodo.org/record/212549/files/figure.png" httpUri-1="https://zenodo.org/record/212550/files/figure.png" httpUri-2="https://zenodo.org/record/212551/files/figure.png" pageId="1" pageNumber="39">Figs. 1–3</figureCitation>
|
||
)
|
||
</paragraph>
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||
<paragraph id="B328D22CFFA6FF8BFF40FA9AFBAFFA93" blockId="1.[151,1086,1367,1390]" box="[151,1086,1367,1390]" pageId="1" pageNumber="39">
|
||
<emphasis id="81E30E3EFFA6FF8BFF40FA9AFE83FA93" bold="true" box="[151,274,1367,1390]" pageId="1" pageNumber="39">Synonymy:</emphasis>
|
||
<taxonomicName id="7497A9AFFFA6FF8BFEC0FA95FD6BFA90" authority="Lim and Lim, 1992" authorityName="Lim and Lim" authorityYear="1992" box="[279,762,1368,1390]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="39" phylum="Chordata" rank="species" species="pleurostigma">
|
||
<emphasis id="81E30E3EFFA6FF8BFEC0FA95FDB7FA93" box="[279,550,1368,1390]" italics="true" pageId="1" pageNumber="39">Kalophrynus pleurostigma</emphasis>
|
||
:
|
||
<bibRefCitation id="D706AFDDFFA6FF8BFDE4FA95FD6BFA90" author="Lim" box="[563,762,1368,1390]" pageId="1" pageNumber="39" refString="Lim, K. K. P. & Lim, F. L. K. (1992) A Guide to the Amphibians and Reptiles of Singapore. Singapore Science Centre, Singapore. 160 pp." type="book" year="1992">Lim and Lim, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
, p. 40, Bukit Timah,
|
||
<collectingCountry id="CB8092BCFFA6FF8BFC07FA95FBABFA93" box="[976,1082,1368,1390]" name="Singapore" pageId="1" pageNumber="39">Singapore</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FB8D81A7FFA6FF8BFF40FA56FD24F96E" pageId="1" pageNumber="39" type="materials_examined">
|
||
<paragraph id="B328D22CFFA6FF8BFF40FA56FD9DFA00" blockId="1.[151,1436,1434,2007]" pageId="1" pageNumber="39">
|
||
<emphasis id="81E30E3EFFA6FF8BFF40FA56FE9AFA4E" bold="true" box="[151,267,1435,1459]" pageId="1" pageNumber="39">
|
||
<typeStatus id="6C2C6C8EFFA6FF8BFF40FA56FE97FA4E" box="[151,262,1435,1459]" pageId="1" pageNumber="39" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="D5864AE9FFA6FF8BFEC4FA57FEEAFA4F" box="[275,379,1434,1458]" pageId="1" pageNumber="39">UKMHC</collectionCode>
|
||
705 (formerly
|
||
<collectionCode id="D5864AE9FFA6FF8BFDFFFA56FDE7FA4F" box="[552,630,1435,1458]" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="1" pageNumber="39">KUHE</collectionCode>
|
||
53284), adult male from Gunung (= Mt.) Pulai, Kpg. (Kampung = village) Sri, Kulai, State of Johor, Peninsular
|
||
<collectingCountry id="CB8092BCFFA6FF8BFD44FA0CFD6BFA24" box="[659,762,1473,1497]" name="Malaysia" pageId="1" pageNumber="39">Malaysia</collectingCountry>
|
||
(
|
||
<geoCoordinate id="D6A3B4EBFFA6FF8BFCD9FA0CFCEBFA24" box="[782,890,1470,1497]" direction="north" orientation="latitude" pageId="1" pageNumber="39" precision="925" value="1.6">01o36’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D6A3B4EBFFA6FF8BFC5EFA0CFB9BFA24" box="[905,1034,1470,1497]" direction="east" orientation="longitude" pageId="1" pageNumber="39" precision="925" value="103.53333">103o 32’ E</geoCoordinate>
|
||
,
|
||
<quantity id="746F7FC9FFA6FF8BFBCDFA0CFBF4FA24" box="[1050,1125,1473,1497]" metricMagnitude="2" metricUnit="m" metricValue="4.57" pageId="1" pageNumber="39" unit="m" value="457.0">457 m</quantity>
|
||
a.s.l.), collected by Kanto Nishikawa on
|
||
<date id="C729F4ECFFA6FF8BFEEEFA28FD99FA00" box="[313,520,1509,1533]" pageId="1" pageNumber="39" value="2009-09-06">6 September 2009</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA6FF8BFF12F9C0FD24F96E" blockId="1.[151,1436,1434,2007]" pageId="1" pageNumber="39">
|
||
<emphasis id="81E30E3EFFA6FF8BFF12F9C0FED5F9D8" bold="true" box="[197,324,1549,1573]" pageId="1" pageNumber="39">
|
||
<typeStatus id="6C2C6C8EFFA6FF8BFF12F9C0FEAEF9D8" box="[197,319,1549,1573]" pageId="1" pageNumber="39" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="D5864AE9FFA6FF8BFE9BF9C0FE0BF9D9" box="[332,410,1549,1572]" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="1" pageNumber="39">KUHE</collectionCode>
|
||
53314, 53315, two adult males from Gunung Lambak (
|
||
<geoCoordinate id="D6A3B4EBFFA6FF8BFBCBF9C1FB17F9D9" box="[1052,1158,1545,1572]" direction="north" orientation="latitude" pageId="1" pageNumber="39" precision="925" value="2.0">02o00’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D6A3B4EBFFA6FF8BFB44F9C1FA97F9D9" box="[1171,1286,1545,1572]" direction="east" orientation="longitude" pageId="1" pageNumber="39" precision="925" value="103.36667">103o22’ E</geoCoordinate>
|
||
,
|
||
<quantity id="746F7FC9FFA6FF8BFAC3F9C1FADFF9D9" box="[1300,1358,1548,1572]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="39" unit="m" value="75.0">75 m</quantity>
|
||
a.s.l.), Kluang, Johor, collected
|
||
<date id="C729F4ECFFA6FF8BFE65F9FDFD11F9B5" box="[434,640,1584,1608]" pageId="1" pageNumber="39" value="2009-09-09">9 September 2009</date>
|
||
by Kanto Nishikawa and Masafumi Matsui;
|
||
<collectionCode id="D5864AE9FFA6FF8BFB55F9FCFB41F9B5" box="[1154,1232,1585,1608]" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="1" pageNumber="39">KUHE</collectionCode>
|
||
52061, a juvenile from Jalan Lombong, Panti (
|
||
<geoCoordinate id="D6A3B4EBFFA6FF8BFE0DF99AFDD3F992" box="[474,578,1620,1647]" direction="north" orientation="latitude" pageId="1" pageNumber="39" precision="925" value="1.8166666">01o49’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D6A3B4EBFFA6FF8BFD99F99AFD2EF992" box="[590,703,1620,1647]" direction="east" orientation="longitude" pageId="1" pageNumber="39" precision="925" value="103.833336">103o50’ E</geoCoordinate>
|
||
,
|
||
<quantity id="746F7FC9FFA6FF8BFD1CF99AFC95F992" box="[715,772,1623,1647]" metricMagnitude="1" metricUnit="m" metricValue="5.7" pageId="1" pageNumber="39" unit="m" value="57.0">57 m</quantity>
|
||
a.s.l.), Kota Tinggi, Johor, collected
|
||
<date id="C729F4ECFFA6FF8BFB72F99AFAA7F992" box="[1189,1334,1623,1647]" pageId="1" pageNumber="39" value="2008-07-30">30 July 2008</date>
|
||
by Amir Hamidy, Daicus Belabut, and Masafumi Matsui.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FB8D81A7FFA6FF8BFF12F952FD0AF926" pageId="1" pageNumber="39" type="etymology">
|
||
<paragraph id="B328D22CFFA6FF8BFF12F952FD0AF926" blockId="1.[151,1436,1434,2007]" pageId="1" pageNumber="39">
|
||
<emphasis id="81E30E3EFFA6FF8BFF12F952FEDAF94A" bold="true" box="[197,331,1695,1719]" pageId="1" pageNumber="39">Etymology.</emphasis>
|
||
The specific name is dedicated to Dr. Lim Boo Liat, Fellow of the Academy of Sciences
|
||
<collectingCountry id="CB8092BCFFA6FF8BFAF9F952FA06F94A" box="[1326,1431,1695,1719]" name="Malaysia" pageId="1" pageNumber="39">Malaysia</collectingCountry>
|
||
, who is a pioneer of field zoology in
|
||
<collectingCountry id="CB8092BCFFA6FF8BFDFAF90EFD06F926" box="[557,663,1731,1755]" name="Malaysia" pageId="1" pageNumber="39">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FB8D81A7FFA6FF88FF12F92BFBD9FEC2" lastPageId="2" lastPageNumber="40" pageId="1" pageNumber="39" type="diagnosis">
|
||
<paragraph id="B328D22CFFA6FF8BFF12F92BFF78F872" blockId="1.[151,1436,1434,2007]" pageId="1" pageNumber="39">
|
||
<emphasis id="81E30E3EFFA6FF8BFF12F92BFEAEF902" bold="true" box="[197,319,1766,1791]" pageId="1" pageNumber="39">Diagnosis.</emphasis>
|
||
The new species is assigned to a member of
|
||
<taxonomicName id="7497A9AFFFA6FF8BFC91F92AFC46F902" box="[838,983,1767,1791]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="1" pageNumber="39" phylum="Chordata" rank="genus">
|
||
<emphasis id="81E30E3EFFA6FF8BFC91F92AFC46F902" box="[838,983,1767,1791]" italics="true" pageId="1" pageNumber="39">Kalophrynus</emphasis>
|
||
</taxonomicName>
|
||
through mitochondrial DNA genealogy (
|
||
<bibRefCitation id="D706AFDDFFA6FF8BFF49F8C6FEE1F8DE" author="Matsui" box="[158,368,1803,1827]" pageId="1" pageNumber="39" refString="Matsui, M., Hamidy, A., Daicus, M. B., Norhayati, A., Panha, S., Sudin, A., Khonsue, W., Oh, H. - S., Yong, H. - S., Jiang, J. - P. & Nishikawa, K. (2011) Systematic relationships of Oriental tiny frogs of the family Microhylidae (Amphibia, Anura) as revealed by mtDNA genealogy. Molecular Phylogenetics and Evolution, 61, 167 - 176." type="journal article" year="2011">
|
||
Matsui
|
||
<emphasis id="81E30E3EFFA6FF8BFF21F8C1FEBEF8DE" box="[246,303,1803,1827]" italics="true" pageId="1" pageNumber="39">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
) together with the following morphological characteristics: No spine-like projection of skin at heel and elbow; belly without a network pattern; tips of fingers not expanded; underside of fingers without greatly enlarged tubercles; snout less than twice diameter of eye; inner metatarsal tubercle low, not shovel-like; tympanum visible.
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA6FF88FF12F856FBD9FEC2" blockId="1.[151,1436,1434,2007]" lastBlockId="2.[151,1437,151,320]" lastPageId="2" lastPageNumber="40" pageId="1" pageNumber="39">
|
||
A medium-sized species that can be distinguished from all congeneric species by the following combination of characters; adult males
|
||
<quantity id="746F7FC9FFA6FF8BFE49F872FDD1F82A" box="[414,576,1983,2007]" metricMagnitude="-2" metricUnit="m" metricValue="2.745" metricValueMax="2.87" metricValueMin="2.62" pageId="1" pageNumber="39" unit="mm" value="27.45" valueMax="28.7" valueMin="26.2">26.2–28.7 mm</quantity>
|
||
SVL (mean = 27.3, n = 3); snout pointed, directed downwards; third toe longer than fifth; projection of fourth finger from palm shorter than length of terminal phalanx of third finger; two subarticular tubercles under fourth finger; no subarticular tubercles under fifth toe; indistinct gland dorsal to arm insertion; no nuptial pads or asperities; forelimb not very stout, without a humeral spine in males; outer metatarsal tubercle absent; light lateral stripe; usually with distinct inguinal dark spot without white rim or spotting; calls similar to
|
||
<taxonomicName id="7497A9AFFFA5FF88FF37FEE5FE66FEBD" box="[224,503,296,320]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="2" pageNumber="40" phylum="Chordata" rank="species" species="laterimaculata">
|
||
<emphasis id="81E30E3EFFA5FF88FF37FEE5FE66FEBD" box="[224,503,296,320]" italics="true" pageId="2" pageNumber="40">Hylarana laterimaculata</emphasis>
|
||
</taxonomicName>
|
||
with a long series of high-pitched whistle like notes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="E7E882A4FFA5FF88FF40FA6AFE80FA21" httpUri="https://zenodo.org/record/212549/files/figure.png" pageId="2" pageNumber="40" targetBox="[450,1136,367,1425]" targetPageId="2">
|
||
<paragraph id="B328D22CFFA5FF88FF40FA6AFE80FA21" blockId="2.[151,1436,1447,1500]" pageId="2" pageNumber="40">
|
||
<emphasis id="81E30E3EFFA5FF88FF40FA6AFE82FA40" bold="true" box="[151,275,1447,1470]" pageId="2" pageNumber="40">FIGURE 1.</emphasis>
|
||
(A) Dorsal and (B) ventral views of the holotype of
|
||
<taxonomicName id="7497A9AFFFA5FF88FCFEFA6AFBB0FA40" box="[809,1057,1447,1469]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="2" pageNumber="40" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA5FF88FCFEFA6AFBB0FA40" box="[809,1057,1447,1469]" italics="true" pageId="2" pageNumber="40">Kalophrynus limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA5FF88FBFFFA65FBE9FA40" box="[1064,1144,1448,1469]" pageId="2" pageNumber="40" rank="species">
|
||
<emphasis id="81E30E3EFFA5FF88FBFFFA65FBE9FA40" bold="true" box="[1064,1144,1448,1469]" pageId="2" pageNumber="40">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(male, UKMHC 705). Scale bar, 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="FB8D81A7FFA5FF8FFF12F9CEFE20FCCA" lastPageId="5" lastPageNumber="43" pageId="2" pageNumber="40" type="description">
|
||
<paragraph id="B328D22CFFA5FF88FF12F9CEFD60F8C1" blockId="2.[151,1436,1539,2032]" pageId="2" pageNumber="40">
|
||
<emphasis id="81E30E3EFFA5FF88FF12F9CEFE4CF9E1" bold="true" box="[197,477,1539,1564]" pageId="2" pageNumber="40">
|
||
Description of
|
||
<typeStatus id="6C2C6C8EFFA5FF88FEA1F9C9FE4CF9E1" box="[374,477,1540,1564]" pageId="2" pageNumber="40" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(measurements in mm). Medium sized
|
||
<emphasis id="81E30E3EFFA5FF88FC4AF9CEFC37F9E1" bold="true" box="[925,934,1539,1564]" pageId="2" pageNumber="40">(</emphasis>
|
||
SVL 26.2); habitus moderately stocky (
|
||
<figureCitation id="2BACCEA9FFA5FF88FAB4F9C9FF34F9C2" captionStart="FIGURE 1" captionStartId="2.[151,249,1447,1470]" captionTargetBox="[450,1136,367,1425]" captionTargetId="figure@2.[450,1136,367,1426]" captionTargetPageId="2" captionText="FIGURE 1. (A) Dorsal and (B) ventral views of the holotype of Kalophrynus limbooliati sp. nov. (male, UKMHC 705). Scale bar, 10 mm." httpUri="https://zenodo.org/record/212549/files/figure.png" pageId="2" pageNumber="40">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="2BACCEA9FFA5FF88FF63F9EAFF52F9C2" box="[180,195,1575,1599]" captionStart="FIGURE 3" captionStartId="3.[151,249,1654,1677]" captionTargetBox="[340,1247,932,1632]" captionTargetId="figure@3.[340,1248,932,1632]" captionTargetPageId="3" captionText="FIGURE 3. Dorsolateral view of the holotype of Kalophrynus limbooliati sp. nov. (male, UKMHC 705) from Gunung Pulai, Johor." httpUri="https://zenodo.org/record/212551/files/figure.png" pageId="2" pageNumber="40">3</figureCitation>
|
||
); head triangular, slightly shorter (8.6) than wide (9.0); snout pointed, directed downwards in profile (
|
||
<figureCitation id="2BACCEA9FFA5FF88FA9EF9EAFA1CF9C2" box="[1353,1421,1575,1599]" captionStart="FIGURE 3" captionStartId="3.[151,249,1654,1677]" captionTargetBox="[340,1247,932,1632]" captionTargetId="figure@3.[340,1248,932,1632]" captionTargetPageId="3" captionText="FIGURE 3. Dorsolateral view of the holotype of Kalophrynus limbooliati sp. nov. (male, UKMHC 705) from Gunung Pulai, Johor." httpUri="https://zenodo.org/record/212551/files/figure.png" pageId="2" pageNumber="40">Fig. 3</figureCitation>
|
||
), projecting beyond lower jaw; eye moderate, as long (3.5) as snout (3.5); canthus rostralis not sharply defined, straight; loreal region vertical, very slightly concave; nostril below canthus rostralis, directed laterally, much closer to tip of snout (1.1) than to eye (2.0); interorbital distance (3.7) much wider than internarial distance (2.1), the latter subequal to upper eyelid (2.2); pineal spot absent; tympanum distinct, diameter (2.3) more than three-fifths that of eye, and very close to eye (0.3); upper jaw edentate; tongue entire, without papillae; a crenulated ridge of skin on palate posterior to eye, preceded by a shorter, similarly notched one between posterior halves of eye; a median, subgular vocal sac; vocal openings posterior to rictus.
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA5FF88FF12F88AFE1AF80D" blockId="2.[151,1436,1539,2032]" pageId="2" pageNumber="40">
|
||
Forelimb long (18.0, 72% of SVL) and slender; fingers thick, basally slightly webbed; tips rounded, not dilated; relative length of fingers: IV<I<II<III; portion of fourth finger projecting from palm (0.9) shorter than length of terminal phalanx of third finger (1.4); outer palmar tubercle large and fleshy, inner indistinct; subarticular tubercles between finger tip and palmer tubercle rounded, two on fingers I, II and IV, three on III (
|
||
<figureCitation id="2BACCEA9FFA5FF88FAE7F879FAE9F831" box="[1328,1400,1972,1996]" captionStart="FIGURE 2" captionStartId="3.[151,249,830,853]" captionTargetBox="[415,1171,183,808]" captionTargetId="figure@3.[415,1171,183,809]" captionTargetPageId="3" captionText="FIGURE 2. Ventral views of (A) hand and (B) foot of the holotype of Kalophrynus limbooliati sp. nov. (male, UKMHC 705). Scale bar, 5 mm." httpUri="https://zenodo.org/record/212550/files/figure.png" pageId="2" pageNumber="40">Fig. 2</figureCitation>
|
||
A); humeral spine absent.
|
||
</paragraph>
|
||
<caption id="E7E882A4FFA4FF89FF40FCF3FED3FC8E" httpUri="https://zenodo.org/record/212550/files/figure.png" pageId="3" pageNumber="41" targetBox="[415,1171,183,808]" targetPageId="3">
|
||
<paragraph id="B328D22CFFA4FF89FF40FCF3FED3FC8E" blockId="3.[151,1437,830,883]" pageId="3" pageNumber="41">
|
||
<emphasis id="81E30E3EFFA4FF89FF40FCF3FE82FCAE" bold="true" box="[151,275,830,853]" pageId="3" pageNumber="41">FIGURE 2.</emphasis>
|
||
Ventral views of (A) hand and (B) foot of the holotype of
|
||
<taxonomicName id="7497A9AFFFA4FF89FCB2FCF3FBCEFCA9" box="[869,1119,830,852]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="41" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA4FF89FCB2FCF3FBCEFCA9" box="[869,1119,830,852]" italics="true" pageId="3" pageNumber="41">Kalophrynus limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA4FF89FBB3FCF2FB24FCA9" box="[1124,1205,831,852]" pageId="3" pageNumber="41" rank="species">
|
||
<emphasis id="81E30E3EFFA4FF89FBB3FCF2FB24FCA9" bold="true" box="[1124,1205,831,852]" pageId="3" pageNumber="41">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(male, UKMHC 705). Scale bar, 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E7E882A4FFA4FF89FF40F9BBFF44F956" httpUri="https://zenodo.org/record/212551/files/figure.png" pageId="3" pageNumber="41" targetBox="[340,1247,932,1632]" targetPageId="3">
|
||
<paragraph id="B328D22CFFA4FF89FF40F9BBFF44F956" blockId="3.[151,1436,1654,1707]" pageId="3" pageNumber="41">
|
||
<emphasis id="81E30E3EFFA4FF89FF40F9BBFE82F971" bold="true" box="[151,275,1654,1677]" pageId="3" pageNumber="41">FIGURE 3.</emphasis>
|
||
Dorsolateral view of the holotype of
|
||
<taxonomicName id="7497A9AFFFA4FF89FD42F9BAFC1FF970" box="[661,910,1655,1677]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="3" pageNumber="41" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA4FF89FD42F9BAFC1FF970" box="[661,910,1655,1677]" italics="true" pageId="3" pageNumber="41">Kalophrynus limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA4FF89FC42F9BAFC76F971" box="[917,999,1655,1676]" pageId="3" pageNumber="41" rank="species">
|
||
<emphasis id="81E30E3EFFA4FF89FC42F9BAFC76F971" bold="true" box="[917,999,1655,1676]" pageId="3" pageNumber="41">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(male, UKMHC 705) from Gunung Pulai, Johor.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B328D22CFFA4FF89FF12F915FCD3F811" blockId="3.[151,1436,1752,2028]" pageId="3" pageNumber="41">
|
||
Hindlimb moderately long (35.5, 135% of SVL) and slender; tibia moderately long (10.9, 42% of SVL), heels not overlapping when folded limbs held at right angles to body; tibiotarsal articulation of adpressed limb reaching to the center of tympanum when held alongside body; foot (8.7) much shorter than tibia (FL, 80% of TL); toe tips rounded; relative length of toes: I<V<II<III<IV; webbing poorly developed (
|
||
<figureCitation id="2BACCEA9FFA4FF89FC24F889FBA6F8A1" box="[1011,1079,1860,1884]" captionStart="FIGURE 2" captionStartId="3.[151,249,830,853]" captionTargetBox="[415,1171,183,808]" captionTargetId="figure@3.[415,1171,183,809]" captionTargetPageId="3" captionText="FIGURE 2. Ventral views of (A) hand and (B) foot of the holotype of Kalophrynus limbooliati sp. nov. (male, UKMHC 705). Scale bar, 5 mm." httpUri="https://zenodo.org/record/212550/files/figure.png" pageId="3" pageNumber="41">Fig. 2</figureCitation>
|
||
B), formula:
|
||
<date id="C729F4ECFFA4FF89FB10F889FF6AF87D" pageId="3" pageNumber="41">
|
||
<emphasis id="81E30E3EFFA4FF89FB10F889FB43F8A1" bold="true" box="[1223,1234,1860,1884]" pageId="3" pageNumber="41">I</emphasis>
|
||
1–2
|
||
<emphasis id="81E30E3EFFA4FF89FADDF889FAB1F8A1" bold="true" box="[1290,1312,1860,1884]" pageId="3" pageNumber="41">II</emphasis>
|
||
2–3
|
||
<emphasis id="81E30E3EFFA4FF89FA8FF889FAE8F8A1" bold="true" box="[1368,1401,1860,1884]" pageId="3" pageNumber="41">III</emphasis>
|
||
2– 3
|
||
<emphasis id="81E30E3EFFA4FF89FF7CF8A5FF5BF87D" bold="true" box="[171,202,1896,1920]" pageId="3" pageNumber="41">IV</emphasis>
|
||
3–1
|
||
</date>
|
||
<emphasis id="81E30E3EFFA4FF89FED6F8A5FE82F87D" bold="true" box="[257,275,1896,1920]" pageId="3" pageNumber="41">V</emphasis>
|
||
; subarticular tubercles numbering one on first and second toes, two on third toe, and three on fourth toe; distal tubercle on third toe and distal and middle ones on fourth toe prominent and rounded, others small and less distinct; no subarticular tubercles on fifth toe; inner metatarsal tubercle oval, length (1.0) more than half of first toe length (1.5); an indistinct outer metatarsal tubercle (
|
||
<figureCitation id="2BACCEA9FFA4FF89FD0AF819FCB0F811" box="[733,801,2004,2028]" captionStart="FIGURE 2" captionStartId="3.[151,249,830,853]" captionTargetBox="[415,1171,183,808]" captionTargetId="figure@3.[415,1171,183,809]" captionTargetPageId="3" captionText="FIGURE 2. Ventral views of (A) hand and (B) foot of the holotype of Kalophrynus limbooliati sp. nov. (male, UKMHC 705). Scale bar, 5 mm." httpUri="https://zenodo.org/record/212550/files/figure.png" pageId="3" pageNumber="41">Fig. 2</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA3FF8EFF12FF5AFAA3FEC2" blockId="4.[151,1436,151,789]" pageId="4" pageNumber="42">Skin coarsely granular dorsally, with small tubercles scattered posteriorly from behind upper eyelid to vent; tips of tubercles forming white asperities; an indistinct gland on side of head behind tympanum, not delimited by a sinuous groove; upper lateral surfaces of body with scattered minute tubercles; lower lateral surfaces of body, posterior half of chin, abdomen, and posterior side of thighs with large, flattened glandules; skin of gular region not modified for vocal sac; inner and outer margins of fourth finger without skin fringes; nuptial pads absent.</paragraph>
|
||
<paragraph id="B328D22CFFA3FF8EFF12FE86FD26FDC1" blockId="4.[151,1436,151,789]" pageId="4" pageNumber="42">
|
||
<emphasis id="81E30E3EFFA3FF8EFF12FE86FE9FFE99" bold="true" box="[197,270,331,356]" pageId="4" pageNumber="42">Color.</emphasis>
|
||
In life, ground color of dorsum highly variable from light orange brown to dark chocolate brown, with several small dark brown spots and obscure hour-glass shaped marking from upper eyelid to shoulder; a light dorsolateral line extending from snout tip through margin of upper eyelid to groin, forming a boundary between lighter dorsum and darker sides of head and shoulder; tips of dorsal granules whitish; limbs dorsally brown with an obscure darker bar (
|
||
<figureCitation id="2BACCEA9FFA3FF8EFEA1FE11FE51FE09" box="[374,448,476,500]" captionStart="FIGURE 1" captionStartId="2.[151,249,1447,1470]" captionTargetBox="[450,1136,367,1425]" captionTargetId="figure@2.[450,1136,367,1426]" captionTargetPageId="2" captionText="FIGURE 1. (A) Dorsal and (B) ventral views of the holotype of Kalophrynus limbooliati sp. nov. (male, UKMHC 705). Scale bar, 10 mm." httpUri="https://zenodo.org/record/212549/files/figure.png" pageId="4" pageNumber="42">Figs. 1</figureCitation>
|
||
A, 3); ventrum pinkish gray, with scattered white flecks (
|
||
<figureCitation id="2BACCEA9FFA3FF8EFB95FE11FB10FE09" box="[1090,1153,476,500]" captionStart="FIGURE 1" captionStartId="2.[151,249,1447,1470]" captionTargetBox="[450,1136,367,1425]" captionTargetId="figure@2.[450,1136,367,1426]" captionTargetPageId="2" captionText="FIGURE 1. (A) Dorsal and (B) ventral views of the holotype of Kalophrynus limbooliati sp. nov. (male, UKMHC 705). Scale bar, 10 mm." httpUri="https://zenodo.org/record/212549/files/figure.png" pageId="4" pageNumber="42">Fig. 1</figureCitation>
|
||
B); inguinal spot entirely black without lighter borders or pattern; iris golden with black pigmentation. In preservative, color and pattern have generally faded but not obviously changed.
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA3FF8EFF12FD8AFC80FCE8" blockId="4.[151,1436,151,789]" pageId="4" pageNumber="42">
|
||
<emphasis id="81E30E3EFFA3FF8EFF12FD8AFEA8FD9D" bold="true" box="[197,313,583,608]" pageId="4" pageNumber="42">Variation.</emphasis>
|
||
Individuals of the
|
||
<typeStatus id="6C2C6C8EFFA3FF8EFDCBFD85FDDDFD9D" box="[540,588,584,608]" pageId="4" pageNumber="42">type</typeStatus>
|
||
series are generally similar in morphology (
|
||
<tableCitation id="FE15E797FFA3FF8EFB84FD8AFB38FDA2" box="[1107,1193,583,607]" captionStart="TABLE 1" captionStartId="4.[151,237,836,857]" captionText="TABLE 1. Measurements of 18 characters (SVL in mm and medians of % ratios [R] of other characters to SVL) in the holotype and two paratypes (all adult males) of Kalophrynus limbooliati. See text for character abbreviations." pageId="4" pageNumber="42">Table 1</tableCitation>
|
||
). In both adult male
|
||
<typeStatus id="6C2C6C8EFFA3FF8EFF40FDA0FE93FD78" box="[151,258,621,645]" pageId="4" pageNumber="42" type="paratype">paratypes</typeStatus>
|
||
(KUHE 53314, 53315), internarial distance is larger than upper eyelid width. They have darker throat and more developed asperities on dorsum than the
|
||
<typeStatus id="6C2C6C8EFFA3FF8EFD0FFD42FCAFFD5A" box="[728,830,655,679]" pageId="4" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
. The inguinal black spot varies in size, and is lacking on the left side in one individual (KUHE 53315). The single juvenile (KUHE 52061) has smooth dorsum, with several black spots scattered from shoulder, some continuing to the inguinal marking.
|
||
<collectingCountry id="CB8092BCFFA3FF8EFB9FFD1AFBEFFD12" box="[1096,1150,727,751]" name="China" pageId="4" pageNumber="42">Chin</collectingCountry>
|
||
and anterior part of ventrum are dotted with dark spots in this juvenile
|
||
<typeStatus id="6C2C6C8EFFA3FF8EFD7FFD30FC9DFCE8" box="[680,780,765,789]" pageId="4" pageNumber="42" type="paratype">paratype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<caption id="E7E882A4FFA3FF8EFF40FC89FB7CFC85" pageId="4" pageNumber="42">
|
||
<paragraph id="B328D22CFFA3FF8EFF40FC89FB7CFC85" blockId="4.[151,1436,835,1062]" pageId="4" pageNumber="42">
|
||
<emphasis id="81E30E3EFFA3FF8EFF40FC89FE9BFCA4" bold="true" box="[151,266,836,857]" pageId="4" pageNumber="42">TABLE 1.</emphasis>
|
||
Measurements of 18 characters (SVL in mm and medians of % ratios [R] of other characters to SVL) in the holotype and two paratypes (all adult males) of
|
||
<taxonomicName id="7497A9AFFFA3FF8EFDADFCAFFCE3FC85" box="[634,882,866,888]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="4" pageNumber="42" phylum="Chordata" rank="species" species="limbooliati">
|
||
<emphasis id="81E30E3EFFA3FF8EFDADFCAFFCE3FC85" box="[634,882,866,888]" italics="true" pageId="4" pageNumber="42">Kalophrynus limbooliati</emphasis>
|
||
</taxonomicName>
|
||
. See text for character abbreviations.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B328D22CFFA3FF8EFD83FC59FA9FFC2F" blockId="4.[151,1436,835,1062]" pageId="4" pageNumber="42">
|
||
SVL RHL RHW RIND RIOD RUEW UKMHC 705 (
|
||
<typeStatus id="6C2C6C8EFFA3FF8EFEE1FC70FE03FC2E" box="[310,402,957,979]" pageId="4" pageNumber="42" type="holotype">holotype</typeStatus>
|
||
) 26.2 32.6 34.3 8 13.9 8.2
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA3FF8EFF48FC2AFA9CFC01" blockId="4.[151,1436,835,1062]" box="[159,1293,998,1021]" pageId="4" pageNumber="42">
|
||
KUHE 53314 (
|
||
<typeStatus id="6C2C6C8EFFA3FF8EFEEFFC2AFE03FC00" box="[312,402,999,1021]" pageId="4" pageNumber="42" type="paratype">paratype</typeStatus>
|
||
) 27 32.6 32.2 9.6 12.2 7.8
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA3FF8EFF48FBDCFA9CFBD8" blockId="4.[151,1436,835,1062]" box="[159,1293,1040,1062]" pageId="4" pageNumber="42">
|
||
KUHE 53315 (
|
||
<typeStatus id="6C2C6C8EFFA3FF8EFEEFFBDDFE03FBDB" box="[312,402,1040,1062]" pageId="4" pageNumber="42" type="paratype">paratype</typeStatus>
|
||
) 28.7 31.9 32.6 9.1 11.7 7.3
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA3FF8EFF48FBF7FA9CFA11" pageId="4" pageNumber="42">
|
||
<table id="C197208CFFA30075FF40FBF4FA0DFA11" box="[151,1436,1081,1516]" gridcols="6" gridrows="4" pageId="4" pageNumber="42">
|
||
<tr id="0DA7D06EFFA30075FF40FBF4FA0DFBB2" box="[151,1436,1081,1103]" gridrow="0" pageId="4" pageNumber="42">
|
||
<th id="4E76B912FFA30075FF40FBF4FE08FBB2" box="[151,409,1081,1103]" gridcol="0" gridrow="0" pageId="4" pageNumber="42">mean+SD/median</th>
|
||
<th id="4E76B912FFA30075FD84FBF4FC82FBB2" box="[595,787,1081,1103]" gridcol="1" gridrow="0" pageId="4" pageNumber="42">27.3+1.28 32.6</th>
|
||
<th id="4E76B912FFA30075FCB7FBF4FC3AFBB2" box="[864,939,1081,1103]" gridcol="2" gridrow="0" pageId="4" pageNumber="42">32.6</th>
|
||
<th id="4E76B912FFA30075FC30FBF4FBB8FBB2" box="[999,1065,1081,1103]" gridcol="3" gridrow="0" pageId="4" pageNumber="42">9.1</th>
|
||
<th id="4E76B912FFA30075FBB7FBF4FB3FFBB2" box="[1120,1198,1081,1103]" gridcol="4" gridrow="0" pageId="4" pageNumber="42">12.2</th>
|
||
<th id="4E76B912FFA30075FB39FBF4FA0DFBB2" box="[1262,1436,1081,1103]" gridcol="5" gridrow="0" pageId="4" pageNumber="42">7.8</th>
|
||
</tr>
|
||
<tr id="0DA7D06EFFA30075FF40FBAEFA0DFB5C" box="[151,1436,1123,1185]" gridrow="1" pageId="4" pageNumber="42">
|
||
<th id="4E76B912FFA30075FF40FBAEFE08FB5C" box="[151,409,1123,1185]" gridcol="0" gridrow="1" pageId="4" pageNumber="42">UKMHC 705 (holotype)</th>
|
||
<td id="4E76B912FFA30075FD84FBAEFC82FB5C" box="[595,787,1123,1185]" gridcol="1" gridrow="1" pageId="4" pageNumber="42">RSL REL 13.4 13.4</td>
|
||
<td id="4E76B912FFA30075FCB7FBAEFC3AFB5C" box="[864,939,1123,1185]" gridcol="2" gridrow="1" pageId="4" pageNumber="42">RN-EL 7.6</td>
|
||
<td id="4E76B912FFA30075FC30FBAEFBB8FB5C" box="[999,1065,1123,1185]" gridcol="3" gridrow="1" pageId="4" pageNumber="42">RTD 8.8</td>
|
||
<td id="4E76B912FFA30075FBB7FBAEFB3FFB5C" box="[1120,1198,1123,1185]" gridcol="4" gridrow="1" pageId="4" pageNumber="42">RT-EL 1</td>
|
||
<td id="4E76B912FFA30075FB39FBAEFA0DFB5C" box="[1262,1436,1123,1185]" gridcol="5" gridrow="1" pageId="4" pageNumber="42">RLAL 47.3</td>
|
||
</tr>
|
||
<tr id="0DA7D06EFFA30075FF40FB78FA0DFAE3" box="[151,1436,1205,1310]" gridrow="2" pageId="4" pageNumber="42">
|
||
<th id="4E76B912FFA30075FF40FB78FE08FAE3" box="[151,409,1205,1310]" gridcol="0" gridrow="2" pageId="4" pageNumber="42">KUHE 53314 (paratype) KUHE 53315 (paratype) median</th>
|
||
<td id="4E76B912FFA30075FD84FB78FC82FAE3" box="[595,787,1205,1310]" gridcol="1" gridrow="2" pageId="4" pageNumber="42">12.8 17.4 12 13.4 12.8 13.4</td>
|
||
<td id="4E76B912FFA30075FCB7FB78FC3AFAE3" box="[864,939,1205,1310]" gridcol="2" gridrow="2" pageId="4" pageNumber="42">6.4 6.8 6.8</td>
|
||
<td id="4E76B912FFA30075FC30FB78FBB8FAE3" box="[999,1065,1205,1310]" gridcol="3" gridrow="2" pageId="4" pageNumber="42">8.1 9.4 8.8</td>
|
||
<td id="4E76B912FFA30075FBB7FB78FB3FFAE3" box="[1120,1198,1205,1310]" gridcol="4" gridrow="2" pageId="4" pageNumber="42">0.4 0.7 0.7</td>
|
||
<td id="4E76B912FFA30075FB39FB78FA0DFAE3" box="[1262,1436,1205,1310]" gridcol="5" gridrow="2" pageId="4" pageNumber="42">46.7 46 46.7</td>
|
||
</tr>
|
||
<tr id="0DA7D06EFFA30075FF40FAFFFA0DFA11" box="[151,1436,1330,1516]" gridrow="3" pageId="4" pageNumber="42">
|
||
<th id="4E76B912FFA30075FF40FAFFFE08FA11" box="[151,409,1330,1516]" gridcol="0" gridrow="3" pageId="4" pageNumber="42">UKMHC 705 (holotype) KUHE 53314 (paratype) KUHE 53315 (paratype) median</th>
|
||
<td id="4E76B912FFA30075FD84FAFFFC82FA11" box="[595,787,1330,1516]" gridcol="1" gridrow="3" pageId="4" pageNumber="42">RFLL RTL 68 41.4 63 41.3 63.4 41.5 63.4 41.4</td>
|
||
<td id="4E76B912FFA30075FCB7FAFFFC3AFA11" box="[864,939,1330,1516]" gridcol="2" gridrow="3" pageId="4" pageNumber="42">RFL 33.2 35.6 35.2 35.2</td>
|
||
<td id="4E76B912FFA30075FC30FAFFFBB8FA11" box="[999,1065,1330,1516]" gridcol="3" gridrow="3" pageId="4" pageNumber="42">RHLL 135.5 134.1 128.6 134.1</td>
|
||
<td id="4E76B912FFA30075FBB7FAFFFB3FFA11" box="[1120,1198,1330,1516]" gridcol="4" gridrow="3" pageId="4" pageNumber="42">RIMTL 3.8 3.3 3 3.3</td>
|
||
<td id="4E76B912FFA30075FB39FAFFFA0DFA11" box="[1262,1436,1330,1516]" gridcol="5" gridrow="3" pageId="4" pageNumber="42">R1TL 5.7 5.6 5.7 5.7</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA3FF8EFF12F9E6FB35F92F" blockId="4.[151,1436,1578,2035]" pageId="4" pageNumber="42">
|
||
<emphasis id="81E30E3EFFA3FF8EFF12F9E6FE89F9BE" bold="true" box="[197,280,1579,1603]" pageId="4" pageNumber="42">Range.</emphasis>
|
||
Southern part of Western (Peninsular)
|
||
<collectingCountry id="CB8092BCFFA3FF8EFD1AF9E7FCA9F9BF" box="[717,824,1578,1602]" name="Malaysia" pageId="4" pageNumber="42">Malaysia</collectingCountry>
|
||
: Johor (Panti, Kota Tinggi; Gunung Pulai, Kulai [
|
||
<quantity id="746F7FC9FFA3FF8EFAB6F9E7FF71F99B" metricMagnitude="2" metricUnit="m" metricValue="4.775" metricValueMax="6.05" metricValueMin="3.5" pageId="4" pageNumber="42" unit="m" value="477.5" valueMax="605.0" valueMin="350.0">350– 605 m</quantity>
|
||
a.s.l.]; Gunung Lambak, Kluang [
|
||
<geoCoordinate id="D6A3B4EBFFA3FF8EFDBCF983FD43F99B" box="[619,722,1614,1638]" direction="east" orientation="longitude" pageId="4" pageNumber="42" precision="925" value="2.0">02°00' E</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D6A3B4EBFFA3FF8EFD36F983FCC5F99B" box="[737,852,1614,1638]" direction="east" orientation="longitude" pageId="4" pageNumber="42" precision="925" value="103.36667">103°22' E</geoCoordinate>
|
||
,
|
||
<quantity id="746F7FC9FFA3FF8EFCB5F983FC44F99B" box="[866,981,1614,1638]" metricMagnitude="2" metricUnit="m" metricValue="1.3" metricValueMax="1.85" metricValueMin="0.75" pageId="4" pageNumber="42" unit="m" value="130.0" valueMax="185.0" valueMin="75.0">75–185 m</quantity>
|
||
a.s.l.]; Trail Lagenda, Gunung Ledang, Tangkak [
|
||
<geoCoordinate id="D6A3B4EBFFA3FF8EFED1F9BFFEFEF977" box="[262,367,1650,1674]" direction="north" orientation="latitude" pageId="4" pageNumber="42" precision="925" value="2.3666668">02°22' N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D6A3B4EBFFA3FF8EFEABF9BFFE7CF977" box="[380,493,1650,1674]" direction="east" orientation="longitude" pageId="4" pageNumber="42" precision="925" value="102.61667">102°37' E</geoCoordinate>
|
||
,
|
||
<quantity id="746F7FC9FFA3FF8EFE2DF9BFFDE8F977" box="[506,633,1650,1674]" metricMagnitude="2" metricUnit="m" metricValue="8.225" metricValueMax="8.95" metricValueMin="7.5" pageId="4" pageNumber="42" unit="m" value="822.5" valueMax="895.0" valueMin="750.0">750–895 m</quantity>
|
||
a.s.l.: recorded by calls]) and Negeri Sembilan (Sungai Kenaboi, Kenaboi: recorded by calls).
|
||
<collectingCountry id="CB8092BCFFA3FF8EFE7AF95BFDB2F953" box="[429,547,1686,1710]" name="Singapore" pageId="4" pageNumber="42">Singapore</collectingCountry>
|
||
: Bukit Timah Nature Reserve (
|
||
<bibRefCitation id="D706AFDDFFA3FF8EFCA9F95BFAD4F953" author="Lim" box="[894,1349,1686,1710]" pageId="4" pageNumber="42" refString="Lim, K. K. P. & Lim, F. L. K. (1992) A Guide to the Amphibians and Reptiles of Singapore. Singapore Science Centre, Singapore. 160 pp." type="book" year="1992">Lim & Lim 1992: see below, discussion</bibRefCitation>
|
||
). Possibly Janda Baik, Pahang, Western (Peninsular)
|
||
<collectingCountry id="CB8092BCFFA3FF8EFD4AF977FC95F92F" box="[669,772,1722,1746]" name="Malaysia" pageId="4" pageNumber="42">Malaysia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="D706AFDDFFA3FF8EFCC4F977FB08F92F" author="Dring" box="[787,1177,1722,1746]" pageId="4" pageNumber="42" refString="Dring, J. C. M. (1979) Amphibians and reptiles from northern Trengganu, Malaysia, with descriptions of two new geckos: Cnemaspis and Cyrtodactylus. Bulletin of the British Museum of Natural History (Zoology), 34, 181 - 241." type="journal article" year="1979">Dring 1979: see below, discussion</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA3FF8EFF12F912FAD9F87B" blockId="4.[151,1436,1578,2035]" pageId="4" pageNumber="42">
|
||
<emphasis id="81E30E3EFFA3FF8EFF12F912FE13F90A" bold="true" box="[197,386,1758,1783]" pageId="4" pageNumber="42">Natural history.</emphasis>
|
||
On Gunung Pulai, Gunung Ledang, and Gunung Lambak, males were found calling in widely scattered choruses at dry nights (from 1930 h) in early September in the dense secondary broad-leaf forest. Calling males hid themselves among dead leaves and were very difficult to locate, but responded to playback of recorded calls and/or whistles imitating their calls. There were no large bodies of water at the calling sites. The associated anuran species observed at the nearest waters were
|
||
<taxonomicName id="7497A9AFFFA3FF8EFD01F8A3FC7CF87B" box="[726,1005,1902,1926]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="4" pageNumber="42" phylum="Chordata" rank="species" species="laterimaculata">
|
||
<emphasis id="81E30E3EFFA3FF8EFD01F8A3FC7CF87B" box="[726,1005,1902,1926]" italics="true" pageId="4" pageNumber="42">Hylarana laterimaculata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7497A9AFFFA3FF8EFBF3F8A3FABBF87B" authority="Vo" authorityName="Vo" box="[1060,1322,1902,1926]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="4" pageNumber="42" phylum="Chordata" rank="species" species="heymonsi">
|
||
<emphasis id="81E30E3EFFA3FF8EFBF3F8A3FA99F87B" box="[1060,1288,1902,1926]" italics="true" pageId="4" pageNumber="42">Microhyla heymonsi</emphasis>
|
||
Vo
|
||
</taxonomicName>
|
||
g t.
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA3FF8EFF12F85EFAC6F80F" blockId="4.[151,1436,1578,2035]" pageId="4" pageNumber="42">
|
||
At Sungai Kenaboi, calls were heard in secondary and bamboo forests on dry night.
|
||
<taxonomicName id="7497A9AFFFA3FF8EFB7BF85EFA04F856" box="[1196,1429,1939,1963]" class="Amphibia" family="Microhylidae" genus="Microhyla" kingdom="Animalia" order="Anura" pageId="4" pageNumber="42" phylum="Chordata" rank="species" species="heymonsi">
|
||
<emphasis id="81E30E3EFFA3FF8EFB7BF85EFA04F856" box="[1196,1429,1939,1963]" italics="true" pageId="4" pageNumber="42">Microhyla heymonsi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA3FF8EFF40F87AFDC8F833" authority="Gravenhorst" authorityName="Gravenhorst" box="[151,601,1974,1998]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="42" phylum="Chordata" rank="species" species="limnocharis">
|
||
<emphasis id="81E30E3EFFA3FF8EFF40F87AFE3BF833" box="[151,426,1974,1998]" italics="true" pageId="4" pageNumber="42">Fejervarya limnocharis</emphasis>
|
||
(Gravenhorst)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA3FF8EFDBAF875FC70F833" authority="Boulenger" authorityName="Boulenger" box="[621,993,1974,1999]" class="Amphibia" family="Ranidae" genus="Humerana" kingdom="Animalia" order="Anura" pageId="4" pageNumber="42" phylum="Chordata" rank="species" species="miopus">
|
||
<emphasis id="81E30E3EFFA3FF8EFDBAF875FCD7F833" box="[621,838,1975,1999]" italics="true" pageId="4" pageNumber="42">Humerana miopus</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA3FF8EFC22F87BFACEF833" authority="Boulenger" authorityName="Boulenger" box="[1013,1375,1974,1998]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="4" pageNumber="42" phylum="Chordata" rank="species" species="labialis">
|
||
<emphasis id="81E30E3EFFA3FF8EFC22F87BFB54F833" box="[1013,1221,1974,1998]" italics="true" pageId="4" pageNumber="42">Hylarana labialis</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="7497A9AFFFA3FF8EFF40F816FDC4F80F" authority="Gravenhorst" authorityName="Gravenhorst" box="[151,597,2010,2035]" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="4" pageNumber="42" phylum="Chordata" rank="species" species="leucomystax">
|
||
<emphasis id="81E30E3EFFA3FF8EFF40F816FE20F80E" box="[151,433,2011,2035]" italics="true" pageId="4" pageNumber="42">Polypedates leucomystax</emphasis>
|
||
(Gravenhorst)
|
||
</taxonomicName>
|
||
were observed at the same locality around pools along a forest road.
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA2FF8FFF12FF5AFB1BFEE1" blockId="5.[151,1437,151,2037]" pageId="5" pageNumber="43">
|
||
The single juvenile
|
||
<typeStatus id="6C2C6C8EFFA2FF8FFE70FF55FD96FF4D" box="[423,519,152,176]" pageId="5" pageNumber="43" type="paratype">paratype</typeStatus>
|
||
from Panti (KUHE 52061) was found walking at a dry night at the edge of secondary forest in late July. In early September, no further specimens of this species or calls were detected at this locality. Associated anuran species observed were
|
||
<taxonomicName id="7497A9AFFFA2FF8FFD0FFF2DFBC0FF0A" authority="Gravenhorst" authorityName="Gravenhorst" box="[728,1105,223,248]" class="Amphibia" family="Bufonidae" genus="Phrynoidis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="aspera">
|
||
<emphasis id="81E30E3EFFA2FF8FFD0FFF2DFC39FF05" box="[728,936,224,248]" italics="true" pageId="5" pageNumber="43">Phrynoidis aspera</emphasis>
|
||
(Gravenhorst)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA2FF8FFB89FF2DFACBFF0A" authority="Gray" authorityName="Gray" box="[1118,1370,223,248]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="pulchra">
|
||
<emphasis id="81E30E3EFFA2FF8FFB89FF2DFA89FF05" box="[1118,1304,224,248]" italics="true" pageId="5" pageNumber="43">Kaloula pulchra</emphasis>
|
||
Gray
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA2FF8FFAB0FF2CFE48FEE1" authority="Boulenger" authorityName="Boulenger" class="Amphibia" family="Microhylidae" genus="Micryletta" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="inornata">
|
||
<emphasis id="81E30E3EFFA2FF8FFAB0FF2CFED6FEE1" italics="true" pageId="5" pageNumber="43">Micryletta inornata</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA2FF8FFE33FEC8FCC0FEE1" authority="Boulenger" authorityName="Boulenger" box="[484,849,260,284]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="blythii">
|
||
<emphasis id="81E30E3EFFA2FF8FFE33FEC8FD51FEE1" box="[484,704,260,284]" italics="true" pageId="5" pageNumber="43">Limnonectes blythii</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA2FF8FFC8BFEC8FC5EFEE1" box="[860,975,261,284]" class="Amphibia" family="Ranidae" genus="Humerana" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="miopus">
|
||
<emphasis id="81E30E3EFFA2FF8FFC8BFEC8FC5EFEE1" box="[860,975,261,284]" italics="true" pageId="5" pageNumber="43">H. miopus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="7497A9AFFFA2FF8FFBDBFEC8FB12FEE1" box="[1036,1155,260,284]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="labialis">
|
||
<emphasis id="81E30E3EFFA2FF8FFBDBFEC8FB12FEE1" box="[1036,1155,260,284]" italics="true" pageId="5" pageNumber="43">H. labialis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA2FF8FFF12FEEAFBC0FDA2" blockId="5.[151,1437,151,2037]" pageId="5" pageNumber="43">
|
||
<emphasis id="81E30E3EFFA2FF8FFF12FEEAFE22FEBD" bold="true" box="[197,435,295,320]" pageId="5" pageNumber="43">Call characteristics.</emphasis>
|
||
We recorded the calls of
|
||
<taxonomicName id="7497A9AFFFA2FF8FFD05FEE4FCFCFEBD" box="[722,877,296,320]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="limbooliati">
|
||
<emphasis id="81E30E3EFFA2FF8FFD05FEE4FCFCFEBD" box="[722,877,296,320]" italics="true" pageId="5" pageNumber="43">K. limbooliati</emphasis>
|
||
</taxonomicName>
|
||
at Gunung Pulai, Kulai on
|
||
<date id="C729F4ECFFA2FF8FFB76FEEAFAFCFEC2" box="[1185,1389,295,319]" pageId="5" pageNumber="43" value="2009-09-06">6 September 2009</date>
|
||
(air temperature 23.9 C), Gunung Lambak, Kluang on
|
||
<date id="C729F4ECFFA2FF8FFD35FE80FC22FE99" box="[738,947,332,356]" pageId="5" pageNumber="43" value="2009-09-07">7 September 2009</date>
|
||
(air temperature 27.2 C), and at Kenaboi, Negeri Sembilan on
|
||
<date id="C729F4ECFFA2FF8FFEABFEA2FD37FE7A" box="[380,678,367,391]" pageId="5" pageNumber="43" value="2009-09-10" valueMax="2009-09-11" valueMin="2009-09-10">10 and 11 September 2009</date>
|
||
(air temperature 26.4 C). The advertisement call (
|
||
<figureCitation id="2BACCEA9FFA2FF8FFB03FEA2FA86FE7A" box="[1236,1303,367,391]" captionStart="FIGURE 4" captionStartId="6.[151,249,1196,1219]" captionTargetBox="[175,1392,348,1153]" captionTargetId="figure@6.[151,1436,326,1175]" captionTargetPageId="6" captionText="FIGURE 4. Sonagram (A) and wave form (B) showing a nine note portion of an advertisement call of Kalophrynus limbooliati sp. nov. from Gunung Lambak, Kluang, Johor, recorded at an air temperature 27.2 ° C." httpUri="https://zenodo.org/record/212552/files/figure.png" pageId="5" pageNumber="43">Fig. 4</figureCitation>
|
||
) consists of a very long series of 24–61 (mean±SD = 41.1±11.2, n = 62/seven males) unpulsed notes and lasts about 6.3–15.2 (mean±SD = 10.9±2.8) s. The note repetition rate is 3.31–4.20 (mean±SD =3.72±0.30) notes per s. Each note lasts about 61–76 (mean±SD = 70.0±6.0) ms, and time interval between two notes varies from 238–310 (mean±SD = 271.7±25.4) ms. The dominant frequency lies at
|
||
<date id="C729F4ECFFA2FF8FFD06FE32FCDEFDEA" box="[721,847,511,535]" pageId="5" pageNumber="43" value="1632" valueMax="2008">1632–2008</date>
|
||
(mean±SD = 1909±156) hz, and harmonics are at about 3000–4000 and 7000–8000 hz (
|
||
<figureCitation id="2BACCEA9FFA2FF8FFD96FDE9FD14FDC1" box="[577,645,548,572]" captionStart="FIGURE 4" captionStartId="6.[151,249,1196,1219]" captionTargetBox="[175,1392,348,1153]" captionTargetId="figure@6.[151,1436,326,1175]" captionTargetPageId="6" captionText="FIGURE 4. Sonagram (A) and wave form (B) showing a nine note portion of an advertisement call of Kalophrynus limbooliati sp. nov. from Gunung Lambak, Kluang, Johor, recorded at an air temperature 27.2 ° C." httpUri="https://zenodo.org/record/212552/files/figure.png" pageId="5" pageNumber="43">Fig. 4</figureCitation>
|
||
A). The call has marked frequency and intensity modulations, and frequencies abruptly increase to the middle and then decrease towards the end of a note.
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA2FF8FFF12FDA6FE20FCCA" blockId="5.[151,1437,151,2037]" pageId="5" pageNumber="43">
|
||
<emphasis id="81E30E3EFFA2FF8FFF12FDA6FEFBFD79" bold="true" box="[197,362,619,644]" pageId="5" pageNumber="43">Comparisons.</emphasis>
|
||
Males of
|
||
<taxonomicName id="7497A9AFFFA2FF8FFE08FDA0FDEDFD79" box="[479,636,620,644]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA2FF8FFE08FDA0FDEDFD79" box="[479,636,620,644]" italics="true" pageId="5" pageNumber="43">K. limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA2FF8FFD53FDA1FD4EFD79" box="[644,735,620,644]" pageId="5" pageNumber="43" rank="species">
|
||
<emphasis id="81E30E3EFFA2FF8FFD53FDA1FD4EFD79" bold="true" box="[644,735,620,644]" pageId="5" pageNumber="43">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
with a SVL of
|
||
<quantity id="746F7FC9FFA2FF8FFC44FDA1FBA9FD79" box="[915,1080,620,644]" metricMagnitude="-2" metricUnit="m" metricValue="2.745" metricValueMax="2.87" metricValueMin="2.62" pageId="5" pageNumber="43" unit="mm" value="27.45" valueMax="28.7" valueMin="26.2">26.2–28.7 mm</quantity>
|
||
SVL (mean =
|
||
<quantity id="746F7FC9FFA2FF8FFB34FDA1FADBFD79" box="[1251,1354,620,644]" metricMagnitude="-2" metricUnit="m" metricValue="2.73" pageId="5" pageNumber="43" unit="mm" value="27.3">27.3 mm</quantity>
|
||
) are of similar size to
|
||
<taxonomicName id="7497A9AFFFA2FF8FFEEDFD5CFDF0FD5A" authority="Boulenger" authorityName="Boulenger" box="[314,609,655,680]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="heterochirus">
|
||
<emphasis id="81E30E3EFFA2FF8FFEEDFD5CFEDCFD55" box="[314,333,657,680]" italics="true" pageId="5" pageNumber="43">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA2FF8FFE8DFD5DFE77FD55" box="[346,486,656,680]" italics="true" pageId="5" pageNumber="43">heterochirus</emphasis>
|
||
Boulenger
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="746F7FC9FFA2FF8FFDA7FD42FC87FD5A" box="[624,790,655,679]" metricMagnitude="-2" metricUnit="m" metricValue="2.565" metricValueMax="2.7199999999999998" metricValueMin="2.41" pageId="5" pageNumber="43" unit="mm" value="25.65" valueMax="27.2" valueMin="24.1">24.1–27.2 mm</quantity>
|
||
, mean =
|
||
<quantity id="746F7FC9FFA2FF8FFCACFD42FC73FD5A" box="[891,994,655,679]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="5" pageNumber="43" unit="mm" value="26.3">26.3 mm</quantity>
|
||
),
|
||
<taxonomicName id="7497A9AFFFA2FF8FFC22FD5CFB58FD5A" authority="Das & Haas" authorityName="Das & Haas" box="[1013,1225,655,680]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="eok">
|
||
<emphasis id="81E30E3EFFA2FF8FFC22FD5CFBADFD55" box="[1013,1084,656,680]" italics="true" pageId="5" pageNumber="43">K. eok</emphasis>
|
||
Das & Haas
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="746F7FC9FFA2FF8FFB0FFD42FAD0FD5A" box="[1240,1345,655,679]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="5" pageNumber="43" unit="mm" value="26.3">26.3 mm</quantity>
|
||
), and
|
||
<taxonomicName id="7497A9AFFFA2FF8FFA55FD5CFEC1FD31" authority="Peters" authorityName="Peters" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="punctatus">
|
||
<emphasis id="81E30E3EFFA2FF8FFA55FD5CFE95FD31" italics="true" pageId="5" pageNumber="43">K. punctatus</emphasis>
|
||
Peters
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="746F7FC9FFA2FF8FFEB6FD79FE5DFD31" box="[353,460,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="5" pageNumber="43" unit="mm" value="28.3">28.3 mm</quantity>
|
||
); it can be readily differentiated from these species as follows: white spots in a black inguinal marking absent (present in
|
||
<taxonomicName id="7497A9AFFFA2FF8FFDEFFD14FD79FD0D" box="[568,744,728,752]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="heterochirus">
|
||
<emphasis id="81E30E3EFFA2FF8FFDEFFD14FD79FD0D" box="[568,744,728,752]" italics="true" pageId="5" pageNumber="43">K. heterochirus</emphasis>
|
||
</taxonomicName>
|
||
); second finger with two subarticular tubercles (with single subarticular tubercle in
|
||
<taxonomicName id="7497A9AFFFA2FF8FFE75FD30FE7BFCE9" box="[418,490,764,788]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="eok">
|
||
<emphasis id="81E30E3EFFA2FF8FFE75FD30FE7BFCE9" box="[418,490,764,788]" italics="true" pageId="5" pageNumber="43">K. eok</emphasis>
|
||
</taxonomicName>
|
||
); fifth toe is not projecting as far as or farther than the third toe unlike
|
||
<taxonomicName id="7497A9AFFFA2FF8FFAD9FD30FA0DFCE9" box="[1294,1436,765,788]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="punctatus">
|
||
<emphasis id="81E30E3EFFA2FF8FFAD9FD30FA0DFCE9" box="[1294,1436,765,788]" italics="true" pageId="5" pageNumber="43">K. punctatus</emphasis>
|
||
</taxonomicName>
|
||
(data from
|
||
<bibRefCitation id="D706AFDDFFA2FF8FFEC4FCD2FE34FCCA" author="Matsui" box="[275,421,799,823]" pageId="5" pageNumber="43" refString="Matsui, M. (2009) A new species of Kalophrynus with a unique male humeral spine from Peninsular Malaysia (Amphibia, Anura, Microhylidae). Zoological Science, 26, 579 - 585." type="journal article" year="2009">Matsui 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FB8D81A7FFA2FF8CFF12FC89FD64F999" lastPageId="6" lastPageNumber="44" pageId="5" pageNumber="43" type="reference_group">
|
||
<paragraph id="B328D22CFFA2FF8FFF12FC89FC8DFBA5" blockId="5.[151,1437,151,2037]" pageId="5" pageNumber="43">
|
||
<taxonomicName id="7497A9AFFFA2FF8FFF12FC89FE46FCA1" ID-CoL="6NK56" box="[197,471,836,860]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA2FF8FFF12FC89FE46FCA1" box="[197,471,836,860]" italics="true" pageId="5" pageNumber="43">Kalophrynus limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA2FF8FFE0AFC89FDA6FCA1" box="[477,567,836,860]" pageId="5" pageNumber="43" rank="species">
|
||
<emphasis id="81E30E3EFFA2FF8FFE0AFC89FDA6FCA1" bold="true" box="[477,567,836,860]" pageId="5" pageNumber="43">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is smaller in male body size than
|
||
<taxonomicName id="7497A9AFFFA2FF8FFC66FC88FB9EFCA1" ID-CoL="3QZF6" box="[945,1039,837,860]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="yongi">
|
||
<emphasis id="81E30E3EFFA2FF8FFC66FC88FB9EFCA1" box="[945,1039,837,860]" italics="true" pageId="5" pageNumber="43">K. yongi</emphasis>
|
||
</taxonomicName>
|
||
(28.8–31.0 mm, mean =
|
||
<quantity id="746F7FC9FFA2FF8FFAFFFC89FA01FCA1" box="[1320,1424,836,860]" metricMagnitude="-2" metricUnit="m" metricValue="3.03" pageId="5" pageNumber="43" unit="mm" value="30.3">30.3 mm</quantity>
|
||
),
|
||
<taxonomicName id="7497A9AFFFA2FF8FFF40FCA4FE0AFC82" ID-CoL="6NK4Z" authority="Dehling" authorityName="Dehling" box="[151,411,871,896]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="calciphilus">
|
||
<emphasis id="81E30E3EFFA2FF8FFF40FCA4FEA7FC7D" box="[151,310,872,896]" italics="true" pageId="5" pageNumber="43">K. calciphilus</emphasis>
|
||
Dehling
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="746F7FC9FFA2FF8FFE79FCAAFDCBFC82" box="[430,602,871,895]" metricMagnitude="-2" metricUnit="m" metricValue="2.9899999999999998" metricValueMax="3.0100000000000002" metricValueMin="2.9699999999999998" pageId="5" pageNumber="43" unit="mm" value="29.9" valueMax="30.1" valueMin="29.7">29.7–30.1 mm</quantity>
|
||
),
|
||
<taxonomicName id="7497A9AFFFA2FF8FFDA6FCA4FCEFFC82" ID-CoL="6NK4X" authority="Iskandar" authorityName="Iskandar" box="[625,894,871,896]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="minusculus">
|
||
<emphasis id="81E30E3EFFA2FF8FFDA6FCA4FC82FC7D" box="[625,787,872,896]" italics="true" pageId="5" pageNumber="43">K. minusculus</emphasis>
|
||
Iskandar
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="746F7FC9FFA2FF8FFC46FCAAFC6EFC82" box="[913,1023,871,895]" metricMagnitude="-2" metricUnit="m" metricValue="3.22" pageId="5" pageNumber="43" unit="mm" value="32.2">32.2 mm</quantity>
|
||
),
|
||
<taxonomicName id="7497A9AFFFA2FF8FFBC2FCA4FB5EFC7D" box="[1045,1231,872,896]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="palmatissimu">
|
||
<emphasis id="81E30E3EFFA2FF8FFBC2FCA4FB5EFC7D" box="[1045,1231,872,896]" italics="true" pageId="5" pageNumber="43">K. palmatissimu</emphasis>
|
||
</taxonomicName>
|
||
s (
|
||
<quantity id="746F7FC9FFA2FF8FFB3AFCAAFA06FC7D" box="[1261,1431,871,896]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="3.88" metricValueMin="3.12" pageId="5" pageNumber="43" unit="mm" value="35.0" valueMax="38.8" valueMin="31.2">31.2–38.8 mm</quantity>
|
||
, mean =
|
||
<quantity id="746F7FC9FFA2FF8FFF25FC41FECBFC59" box="[242,346,908,932]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" pageId="5" pageNumber="43" unit="mm" value="34.5">34.5 mm</quantity>
|
||
),
|
||
<taxonomicName id="7497A9AFFFA2FF8FFEBAFC40FD6EFC59" ID-CoL="6NJSW" authority="Dutta, Ahmed & Das" authorityName="Dutta, Ahmed & Das" box="[365,767,908,932]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="orangensis">
|
||
<emphasis id="81E30E3EFFA2FF8FFEBAFC40FD98FC59" box="[365,521,909,932]" italics="true" pageId="5" pageNumber="43">K. orangensis</emphasis>
|
||
Dutta, Ahmed & Das
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="746F7FC9FFA2FF8FFCD9FC41FC1DFC59" box="[782,908,908,932]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="5" pageNumber="43" unit="mm" value="36.5" valueMax="38.0" valueMin="35.0">35–38 mm</quantity>
|
||
),
|
||
<taxonomicName id="7497A9AFFFA2FF8FFC48FC40FB16FC58" ID-CoL="6NK57" authority="Inger" authorityName="Inger" box="[927,1159,908,933]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="intermedius">
|
||
<emphasis id="81E30E3EFFA2FF8FFC48FC40FBD5FC59" box="[927,1092,908,932]" italics="true" pageId="5" pageNumber="43">K. intermedius</emphasis>
|
||
Inger
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="746F7FC9FFA2FF8FFB41FC41FAADFC59" box="[1174,1340,908,932]" metricMagnitude="-2" metricUnit="m" metricValue="3.92" metricValueMax="4.05" metricValueMin="3.79" pageId="5" pageNumber="43" unit="mm" value="39.2" valueMax="40.5" valueMin="37.9">37.9–40.5 mm</quantity>
|
||
, mean =
|
||
<quantity id="746F7FC9FFA2FF8FFF40FC62FF6FFC3A" box="[151,254,943,967]" metricMagnitude="-2" metricUnit="m" metricValue="3.92" pageId="5" pageNumber="43" unit="mm" value="39.2">39.2 mm</quantity>
|
||
),
|
||
<taxonomicName id="7497A9AFFFA2FF8FFEC6FC7CFE4EFC3A" ID-CoL="6NJRB" authority="Kiew" authorityName="Kiew" box="[273,479,943,968]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="baluensis">
|
||
<emphasis id="81E30E3EFFA2FF8FFEC6FC7CFE0AFC35" box="[273,411,944,968]" italics="true" pageId="5" pageNumber="43">K. baluensis</emphasis>
|
||
Kiew
|
||
</taxonomicName>
|
||
(34.8–39.0 mm),
|
||
<taxonomicName id="7497A9AFFFA2FF8FFD7FFC7CFC04FC3A" ID-CoL="3QZF3" authority="Stejneger" authorityName="Stejneger" box="[680,917,943,968]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="stellatus">
|
||
<emphasis id="81E30E3EFFA2FF8FFD7FFC7CFCB4FC35" box="[680,805,944,968]" italics="true" pageId="5" pageNumber="43">K. stellatus</emphasis>
|
||
Stejneger
|
||
</taxonomicName>
|
||
(35.0–45.0 mm, mean =
|
||
<quantity id="746F7FC9FFA2FF8FFB67FC62FA86FC3A" box="[1200,1303,943,967]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" pageId="5" pageNumber="43" unit="mm" value="39.3">39.3 mm</quantity>
|
||
),
|
||
<taxonomicName id="7497A9AFFFA2FF8FFAFDFC7CFF73FC11" ID-CoL="6NK4W" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="pleurostigma">
|
||
<emphasis id="81E30E3EFFA2FF8FFAFDFC7CFF73FC11" italics="true" pageId="5" pageNumber="43">K. pleurostigma</emphasis>
|
||
</taxonomicName>
|
||
(35.0–
|
||
<quantity id="746F7FC9FFA2FF8FFEE7FC19FE07FC11" box="[304,406,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="5.04" pageId="5" pageNumber="43" unit="mm" value="50.4">50.4 mm</quantity>
|
||
, mean =
|
||
<quantity id="746F7FC9FFA2FF8FFE2BFC19FDF5FC11" box="[508,612,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="4.22" pageId="5" pageNumber="43" unit="mm" value="42.2">42.2 mm</quantity>
|
||
), and
|
||
<taxonomicName id="7497A9AFFFA2FF8FFD71FC18FC08FC11" ID-CoL="6NK4V" authority="Blyth" authorityName="Blyth" box="[678,921,980,1004]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="interlineatus">
|
||
<emphasis id="81E30E3EFFA2FF8FFD71FC18FCC5FC11" box="[678,852,980,1004]" italics="true" pageId="5" pageNumber="43">K. interlineatus</emphasis>
|
||
Blyth
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="746F7FC9FFA2FF8FFC7FFC19FBC1FC11" box="[936,1104,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="4.255" metricValueMax="4.77" metricValueMin="3.7399999999999998" pageId="5" pageNumber="43" unit="mm" value="42.55" valueMax="47.7" valueMin="37.4">37.4–47.7 mm</quantity>
|
||
), and larger than
|
||
<taxonomicName id="7497A9AFFFA2FF8FFAC6FC18FA0DFC11" ID-CoL="6NJSR" box="[1297,1436,980,1004]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="robinsoni">
|
||
<emphasis id="81E30E3EFFA2FF8FFAC6FC18FA0DFC11" box="[1297,1436,980,1004]" italics="true" pageId="5" pageNumber="43">K. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="746F7FC9FFA2FF8FFF48FC3AFE99FBF2" box="[159,264,1015,1039]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="5" pageNumber="43" unit="mm" value="16.8">16.8 mm</quantity>
|
||
),
|
||
<taxonomicName id="7497A9AFFFA2FF8FFECBFC34FE76FBF2" ID-CoL="6NK59" authority="Dring" authorityName="Dring" box="[284,487,1015,1040]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="nubicola">
|
||
<emphasis id="81E30E3EFFA2FF8FFECBFC34FE0FFBED" box="[284,414,1016,1040]" italics="true" pageId="5" pageNumber="43">K. nubicola</emphasis>
|
||
Dring
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="746F7FC9FFA2FF8FFE21FC3AFD0EFBF2" box="[502,671,1015,1039]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" metricValueMax="2.44" metricValueMin="1.44" pageId="5" pageNumber="43" unit="mm" value="19.4" valueMax="24.4" valueMin="14.4">14.4–24.4 mm</quantity>
|
||
),
|
||
<taxonomicName id="7497A9AFFFA2FF8FFD65FC34FC78FBF2" ID-CoL="6NJT9" authority="Yang & Su" authorityName="Yang & Su" box="[690,1001,1015,1041]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="menglienicus">
|
||
<emphasis id="81E30E3EFFA2FF8FFD65FC34FCF6FBED" box="[690,871,1016,1040]" italics="true" pageId="5" pageNumber="43">K. menglienicus</emphasis>
|
||
Yang & Su
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="746F7FC9FFA2FF8FFC2EFC3AFB0EFBED" box="[1017,1183,1015,1040]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" metricValueMax="2.34" metricValueMin="1.98" pageId="5" pageNumber="43" unit="mm" value="21.6" valueMax="23.4" valueMin="19.8">19.8–23.4 mm</quantity>
|
||
, mean =
|
||
<quantity id="746F7FC9FFA2FF8FFAD1FC3AFAFEFBF2" box="[1286,1391,1015,1039]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" pageId="5" pageNumber="43" unit="mm" value="21.2">21.2 mm</quantity>
|
||
),
|
||
<taxonomicName id="7497A9AFFFA2FF8FFA55FC34FE02FBC9" ID-CoL="6NJSZ" authority="Gunther" authorityName="Gunther" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="bunguranus">
|
||
<emphasis id="81E30E3EFFA2FF8FFA55FC34FE8CFBC9" italics="true" pageId="5" pageNumber="43">K. bunguranus</emphasis>
|
||
(Günther)
|
||
</taxonomicName>
|
||
(
|
||
<quantity id="746F7FC9FFA2FF8FFE76FBD1FDD8FBC9" box="[417,585,1052,1076]" metricMagnitude="-2" metricUnit="m" metricValue="2.175" metricValueMax="2.28" metricValueMin="2.07" pageId="5" pageNumber="43" unit="mm" value="21.75" valueMax="22.8" valueMin="20.7">20.7–22.8 mm</quantity>
|
||
, mean =
|
||
<quantity id="746F7FC9FFA2FF8FFD66FBD1FC88FBC9" box="[689,793,1052,1076]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="5" pageNumber="43" unit="mm" value="21.8">21.8 mm</quantity>
|
||
), and
|
||
<taxonomicName id="7497A9AFFFA2FF8FFC8BFBD0FBDCFBC8" ID-CoL="3QZF4" authority="Inger" authorityName="Inger" box="[860,1101,1052,1077]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="subterrestris">
|
||
<emphasis id="81E30E3EFFA2FF8FFC8BFBD0FB9BFBC9" box="[860,1034,1052,1076]" italics="true" pageId="5" pageNumber="43">K. subterrestris</emphasis>
|
||
Inger
|
||
</taxonomicName>
|
||
(21.0–
|
||
<quantity id="746F7FC9FFA2FF8FFB4BFBD1FA92FBC9" box="[1180,1283,1052,1076]" metricMagnitude="-2" metricUnit="m" metricValue="2.34" pageId="5" pageNumber="43" unit="mm" value="23.4">23.4 mm</quantity>
|
||
, mean =
|
||
<quantity id="746F7FC9FFA2FF8FFABCFBD1FF55FBA5" metricMagnitude="-2" metricUnit="m" metricValue="2.26" pageId="5" pageNumber="43" unit="mm" value="22.6">22.6 mm</quantity>
|
||
) (data from
|
||
<bibRefCitation id="D706AFDDFFA2FF8FFE98FB8CFE5DFBA5" author="Inger" box="[335,460,1088,1113]" pageId="5" pageNumber="43" refString="Inger, R. F. (1954) Systematics and zoogeography of Philippine Amphibia. Fieldiana: Zoology, 33, 183 - 531." type="journal article" year="1954">Inger 1954</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D706AFDDFFA2FF8FFE0FFB8DFDF9FBA5" author="Matsui" box="[472,616,1088,1112]" pageId="5" pageNumber="43" refString="Matsui, M. (2009) A new species of Kalophrynus with a unique male humeral spine from Peninsular Malaysia (Amphibia, Anura, Microhylidae). Zoological Science, 26, 579 - 585." type="journal article" year="2009">Matsui 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D706AFDDFFA2FF8FFDA4FB8DFC81FBA5" author="Dehling" box="[627,784,1088,1112]" pageId="5" pageNumber="43" refString="Dehling, J. M. (2011) A new karst-dwelling species of Kalophrynus (Anura: Microhylidae) from Gunung Mulu National Park, Borneo, Malaysia. Zootaxa, 2737, 49 - 60." type="journal article" year="2011">Dehling 2011</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA2FF8FFF12FBA9FEF8FAA9" blockId="5.[151,1437,151,2037]" pageId="5" pageNumber="43">
|
||
Other than the body size differences,
|
||
<taxonomicName id="7497A9AFFFA2FF8FFDB3FBA8FD6EFB81" ID-CoL="6NK56" box="[612,767,1124,1148]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA2FF8FFDB3FBA8FD6EFB81" box="[612,767,1124,1148]" italics="true" pageId="5" pageNumber="43">K. limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA2FF8FFCD2FBA9FCCFFB81" box="[773,862,1124,1148]" pageId="5" pageNumber="43" rank="species">
|
||
<emphasis id="81E30E3EFFA2FF8FFCD2FBA9FCCFFB81" bold="true" box="[773,862,1124,1148]" pageId="5" pageNumber="43">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be differentiated from these species as follows: from
|
||
<taxonomicName id="7497A9AFFFA2FF8FFF03FB44FEA2FB5D" ID-CoL="3QZF6" box="[212,307,1161,1184]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="yongi">
|
||
<emphasis id="81E30E3EFFA2FF8FFF03FB44FEA2FB5D" box="[212,307,1161,1184]" italics="true" pageId="5" pageNumber="43">K. yongi</emphasis>
|
||
</taxonomicName>
|
||
by having a normal humerus (strongly developed terminal ridges on humerus, and related skin modification in
|
||
<taxonomicName id="7497A9AFFFA2FF8FFEC0FB60FEE7FB39" ID-CoL="3QZF6" box="[279,374,1197,1220]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="yongi">
|
||
<emphasis id="81E30E3EFFA2FF8FFEC0FB60FEE7FB39" box="[279,374,1197,1220]" italics="true" pageId="5" pageNumber="43">K. yongi</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="7497A9AFFFA2FF8FFE1BFB60FDF6FB39" ID-CoL="6NK4Z" box="[460,615,1196,1220]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="calciphilus">
|
||
<emphasis id="81E30E3EFFA2FF8FFE1BFB60FDF6FB39" box="[460,615,1196,1220]" italics="true" pageId="5" pageNumber="43">K. calciphilus</emphasis>
|
||
</taxonomicName>
|
||
by the absence of outer metatarsal tubercle, presence of thinner dorsolateral stripe, without black margin dorsally, and dorsally more pointed snout (prominent tubercle present, dorsolateral stripe wide, bordered by black on both margins, and dorsally obtusely pointed in
|
||
<taxonomicName id="7497A9AFFFA2FF8FFB6AFB38FACBFAF1" ID-CoL="6NK4Z" box="[1213,1370,1268,1292]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="calciphilus">
|
||
<emphasis id="81E30E3EFFA2FF8FFB6AFB38FACBFAF1" box="[1213,1370,1268,1292]" italics="true" pageId="5" pageNumber="43">K. calciphilus</emphasis>
|
||
</taxonomicName>
|
||
); and from
|
||
<taxonomicName id="7497A9AFFFA2FF8FFF03FAD4FEE8FACD" ID-CoL="6NK57" box="[212,377,1304,1328]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="intermedius">
|
||
<emphasis id="81E30E3EFFA2FF8FFF03FAD4FEE8FACD" box="[212,377,1304,1328]" italics="true" pageId="5" pageNumber="43">K. intermedius</emphasis>
|
||
</taxonomicName>
|
||
by the possession of light lateral stripe and inguinal black marking (stripe and marking absent in
|
||
<taxonomicName id="7497A9AFFFA2FF8FFF63FAF0FEC9FAA9" ID-CoL="6NK57" box="[180,344,1340,1364]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="intermedius">
|
||
<emphasis id="81E30E3EFFA2FF8FFF63FAF0FEC9FAA9" box="[180,344,1340,1364]" italics="true" pageId="5" pageNumber="43">K. intermedius</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA2FF8FFF12FAACFE24F989" blockId="5.[151,1437,151,2037]" pageId="5" pageNumber="43">
|
||
In
|
||
<taxonomicName id="7497A9AFFFA2FF8FFF34FAACFEECFA85" ID-CoL="6NK56" box="[227,381,1376,1400]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA2FF8FFF34FAACFEECFA85" box="[227,381,1376,1400]" italics="true" pageId="5" pageNumber="43">K. limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA2FF8FFE53FAADFE4CFA85" box="[388,477,1376,1400]" pageId="5" pageNumber="43" rank="species">
|
||
<emphasis id="81E30E3EFFA2FF8FFE53FAADFE4CFA85" bold="true" box="[388,477,1376,1400]" pageId="5" pageNumber="43">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, the fourth finger from palm is shorter than length of the terminal phalanx of third finger, unlike
|
||
<taxonomicName id="7497A9AFFFA2FF8FFECDFA48FE2DFA61" ID-CoL="6NK4X" box="[282,444,1412,1436]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="minusculus">
|
||
<emphasis id="81E30E3EFFA2FF8FFECDFA48FEBCFA61" box="[282,301,1413,1436]" italics="true" pageId="5" pageNumber="43">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA2FF8FFEE9FA49FE2DFA61" box="[318,444,1412,1436]" italics="true" pageId="5" pageNumber="43">minusculus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA2FF8FFE1BFA48FD01FA61" ID-CoL="6NJT8" box="[460,656,1412,1436]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="palmatissimus">
|
||
<emphasis id="81E30E3EFFA2FF8FFE1BFA48FE4EFA61" box="[460,479,1413,1436]" italics="true" pageId="5" pageNumber="43">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA2FF8FFE27FA49FD01FA61" box="[496,656,1412,1436]" italics="true" pageId="5" pageNumber="43">palmatissimus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA2FF8FFD76FA48FCAEFA61" ID-CoL="6NJSW" box="[673,831,1413,1436]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="orangensis">
|
||
<emphasis id="81E30E3EFFA2FF8FFD76FA48FD25FA61" box="[673,692,1413,1436]" italics="true" pageId="5" pageNumber="43">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA2FF8FFD13FA48FCAEFA61" box="[708,831,1413,1436]" italics="true" pageId="5" pageNumber="43">orangensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA2FF8FFC87FA48FB90FA61" ID-CoL="6NK4V" box="[848,1025,1412,1436]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="interlineatus">
|
||
<emphasis id="81E30E3EFFA2FF8FFC87FA48FCF2FA61" box="[848,867,1413,1436]" italics="true" pageId="5" pageNumber="43">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA2FF8FFCA3FA49FB90FA61" box="[884,1025,1412,1436]" italics="true" pageId="5" pageNumber="43">interlineatus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="7497A9AFFFA2FF8FFB93FA48FB6AFA61" ID-CoL="6NK4W" box="[1092,1275,1412,1436]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="pleurostigma">
|
||
<emphasis id="81E30E3EFFA2FF8FFB93FA48FBC6FA61" box="[1092,1111,1413,1436]" italics="true" pageId="5" pageNumber="43">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA2FF8FFBBFFA49FB6AFA61" box="[1128,1275,1412,1436]" italics="true" pageId="5" pageNumber="43">pleurostigma</emphasis>
|
||
</taxonomicName>
|
||
, in which the fourth finger is longer. In addition,
|
||
<taxonomicName id="7497A9AFFFA2FF8FFDE2FA64FD45FA3D" ID-CoL="6NK56" box="[565,724,1448,1472]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA2FF8FFDE2FA64FD45FA3D" box="[565,724,1448,1472]" italics="true" pageId="5" pageNumber="43">K. limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA2FF8FFD09FA65FCAAFA3D" box="[734,827,1448,1472]" pageId="5" pageNumber="43" rank="species">
|
||
<emphasis id="81E30E3EFFA2FF8FFD09FA65FCAAFA3D" bold="true" box="[734,827,1448,1472]" pageId="5" pageNumber="43">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
has poorly developed webbing on fourth toe, which barely reaches median subarticular tubercle, sharply contrasting with the well-developed webbing in
|
||
<taxonomicName id="7497A9AFFFA2FF8FFB28FA00FF55F9F5" ID-CoL="6NJT8" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="palmatissimus">
|
||
<emphasis id="81E30E3EFFA2FF8FFB28FA00FA83FA19" box="[1279,1298,1485,1508]" italics="true" pageId="5" pageNumber="43">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA2FF8FFAC8FA01FF55F9F5" italics="true" pageId="5" pageNumber="43">palmatissimus</emphasis>
|
||
</taxonomicName>
|
||
; lacks distinct dorsal marking unlike
|
||
<taxonomicName id="7497A9AFFFA2FF8FFDBBFA3CFC9AF9F5" ID-CoL="6NK4X" box="[620,779,1520,1544]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="minusculus">
|
||
<emphasis id="81E30E3EFFA2FF8FFDBBFA3CFC9AF9F5" box="[620,779,1520,1544]" italics="true" pageId="5" pageNumber="43">K. minusculus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7497A9AFFFA2FF8FFC94FA3CFC5FF9F5" ID-CoL="6NJRB" box="[835,974,1520,1544]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="baluensis">
|
||
<emphasis id="81E30E3EFFA2FF8FFC94FA3CFC5FF9F5" box="[835,974,1520,1544]" italics="true" pageId="5" pageNumber="43">K. baluensis</emphasis>
|
||
</taxonomicName>
|
||
, which usually have clear dark markings on dorsum; has black inguinal marking (yellow in
|
||
<taxonomicName id="7497A9AFFFA2FF8FFD0CF9D8FCF6F9D1" ID-CoL="6NJRB" box="[731,871,1556,1580]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="baluensis">
|
||
<emphasis id="81E30E3EFFA2FF8FFD0CF9D8FCF6F9D1" box="[731,871,1556,1580]" italics="true" pageId="5" pageNumber="43">K. baluensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="7497A9AFFFA2FF8FFC57F9D9FB05F9D1" ID-CoL="6NK56" box="[896,1172,1556,1580]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="limbooliati">
|
||
<emphasis id="81E30E3EFFA2FF8FFC57F9D9FB05F9D1" box="[896,1172,1556,1580]" italics="true" pageId="5" pageNumber="43">Kalophrynus limbooliati</emphasis>
|
||
</taxonomicName>
|
||
further differs from
|
||
<taxonomicName id="7497A9AFFFA2FF8FFA55F9D8FF65F9AD" ID-CoL="3QZF3" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="stellatus">
|
||
<emphasis id="81E30E3EFFA2FF8FFA55F9D8FF65F9AD" italics="true" pageId="5" pageNumber="43">K. stellatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7497A9AFFFA2FF8FFEFCF9F4FE4EF9AD" ID-CoL="6NK4W" box="[299,479,1592,1616]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="pleurostigma">
|
||
<emphasis id="81E30E3EFFA2FF8FFEFCF9F4FE4EF9AD" box="[299,479,1592,1616]" italics="true" pageId="5" pageNumber="43">K. pleurostigma</emphasis>
|
||
</taxonomicName>
|
||
by the lack of nuptial pads in males; and from
|
||
<taxonomicName id="7497A9AFFFA2FF8FFC27F9F4FBEBF9AD" ID-CoL="6NJSR" box="[1008,1146,1592,1616]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="robinsoni">
|
||
<emphasis id="81E30E3EFFA2FF8FFC27F9F4FBEBF9AD" box="[1008,1146,1592,1616]" italics="true" pageId="5" pageNumber="43">K. robinsoni</emphasis>
|
||
</taxonomicName>
|
||
in the absence of spinous nuptial pads in the males.
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA2FF8FFF12F94DFEC4F8B1" blockId="5.[151,1437,151,2037]" pageId="5" pageNumber="43">
|
||
<taxonomicName id="7497A9AFFFA2FF8FFF12F94DFE48F965" ID-CoL="6NK56" box="[197,473,1664,1688]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA2FF8FFF12F94DFE48F965" box="[197,473,1664,1688]" italics="true" pageId="5" pageNumber="43">Kalophrynus limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA2FF8FFE36F94DFDADF965" box="[481,572,1664,1688]" pageId="5" pageNumber="43" rank="species">
|
||
<emphasis id="81E30E3EFFA2FF8FFE36F94DFDADF965" bold="true" box="[481,572,1664,1688]" pageId="5" pageNumber="43">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="7497A9AFFFA2FF8FFD03F94CFCC9F965" ID-CoL="6NK59" box="[724,856,1664,1688]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="nubicola">
|
||
<emphasis id="81E30E3EFFA2FF8FFD03F94CFCC9F965" box="[724,856,1664,1688]" italics="true" pageId="5" pageNumber="43">K. nubicola</emphasis>
|
||
</taxonomicName>
|
||
by the possession of distinct subarticular tubercles of fingers and toes (indistinct or absent in
|
||
<taxonomicName id="7497A9AFFFA2FF8FFDA4F968FD67F941" ID-CoL="6NK59" box="[627,758,1700,1724]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="nubicola">
|
||
<emphasis id="81E30E3EFFA2FF8FFDA4F968FD17F941" box="[627,646,1701,1724]" italics="true" pageId="5" pageNumber="43">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA2FF8FFD43F969FD67F941" box="[660,758,1700,1724]" italics="true" pageId="5" pageNumber="43">nubicola</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="7497A9AFFFA2FF8FFC9BF968FB90F941" ID-CoL="6NJT9" box="[844,1025,1700,1724]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="menglienicus">
|
||
<emphasis id="81E30E3EFFA2FF8FFC9BF968FB90F941" box="[844,1025,1700,1724]" italics="true" pageId="5" pageNumber="43">K. menglienicus</emphasis>
|
||
</taxonomicName>
|
||
by the possession of distinct tympanum and toe web (tympanum concealed and toe web absent in
|
||
<taxonomicName id="7497A9AFFFA2FF8FFC87F904FB95F91D" ID-CoL="6NJT9" box="[848,1028,1736,1760]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="menglienicus">
|
||
<emphasis id="81E30E3EFFA2FF8FFC87F904FCF2F91D" box="[848,867,1737,1760]" italics="true" pageId="5" pageNumber="43">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA2FF8FFCA7F905FB95F91D" box="[880,1028,1736,1760]" italics="true" pageId="5" pageNumber="43">menglienicus</emphasis>
|
||
</taxonomicName>
|
||
); from
|
||
<taxonomicName id="7497A9AFFFA2FF8FFB8FF904FB6FF91D" ID-CoL="6NJSZ" box="[1112,1278,1736,1760]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="bunguranus">
|
||
<emphasis id="81E30E3EFFA2FF8FFB8FF904FB6FF91D" box="[1112,1278,1736,1760]" italics="true" pageId="5" pageNumber="43">K. bunguranus</emphasis>
|
||
</taxonomicName>
|
||
by the lack of light marking surrounding black inguinal mark (black inguinal marking in a light area in
|
||
<taxonomicName id="7497A9AFFFA2FF8FFBAFF920FA8EF8F9" ID-CoL="6NJSZ" box="[1144,1311,1772,1796]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="bunguranus">
|
||
<emphasis id="81E30E3EFFA2FF8FFBAFF920FA8EF8F9" box="[1144,1311,1772,1796]" italics="true" pageId="5" pageNumber="43">K. bunguranus</emphasis>
|
||
</taxonomicName>
|
||
); and from
|
||
<taxonomicName id="7497A9AFFFA2FF8FFF40F8DCFED5F8D5" ID-CoL="3QZF4" box="[151,324,1808,1832]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="subterrestris">
|
||
<emphasis id="81E30E3EFFA2FF8FFF40F8DCFF3BF8D5" box="[151,170,1809,1832]" italics="true" pageId="5" pageNumber="43">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA2FF8FFF6FF8DDFED5F8D5" box="[184,324,1808,1832]" italics="true" pageId="5" pageNumber="43">subterrestris</emphasis>
|
||
</taxonomicName>
|
||
by the presence of two subarticular tubercles on the fourth finger (a single subarticular tubercle in
|
||
<taxonomicName id="7497A9AFFFA2FF8FFF40F8F8FED2F8B1" ID-CoL="3QZF4" box="[151,323,1844,1868]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="subterrestris">
|
||
<emphasis id="81E30E3EFFA2FF8FFF40F8F8FF3BF8B1" box="[151,170,1845,1868]" italics="true" pageId="5" pageNumber="43">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA2FF8FFF60F8F9FED2F8B1" box="[183,323,1844,1868]" italics="true" pageId="5" pageNumber="43">subterrestris</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA2FF8CFF10F895FBB0FEE9" blockId="5.[151,1437,151,2037]" lastBlockId="6.[151,1436,151,277]" lastPageId="6" lastPageNumber="44" pageId="5" pageNumber="43">
|
||
The calls of six species of
|
||
<taxonomicName id="7497A9AFFFA2FF8FFE22F895FD17F88D" ID-CoL="62W7Y" box="[501,646,1880,1904]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="genus">
|
||
<emphasis id="81E30E3EFFA2FF8FFE22F895FD17F88D" box="[501,646,1880,1904]" italics="true" pageId="5" pageNumber="43">Kalophrynus</emphasis>
|
||
</taxonomicName>
|
||
have been analyzed (
|
||
<taxonomicName id="7497A9AFFFA2FF8FFCABF894FB99F88D" ID-CoL="6NJRB" box="[892,1032,1880,1904]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="baluensis">
|
||
<emphasis id="81E30E3EFFA2FF8FFCABF894FB99F88D" box="[892,1032,1880,1904]" italics="true" pageId="5" pageNumber="43">K. baluensis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="7497A9AFFFA2FF8FFBC0F894FB22F88D" ID-CoL="6NK4Z" box="[1047,1203,1880,1904]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="calciphilus">
|
||
<emphasis id="81E30E3EFFA2FF8FFBC0F894FB22F88D" box="[1047,1203,1880,1904]" italics="true" pageId="5" pageNumber="43">K. calciphilus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="7497A9AFFFA2FF8FFB14F894FAE3F88D" ID-CoL="6NK4V" box="[1219,1394,1880,1904]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="interlineatus">
|
||
<emphasis id="81E30E3EFFA2FF8FFB14F894FAE3F88D" box="[1219,1394,1880,1904]" italics="true" pageId="5" pageNumber="43">K. interlineatus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="7497A9AFFFA2FF8FFA55F894FF68F86C" ID-CoL="6NK59" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="nubicola">
|
||
<emphasis id="81E30E3EFFA2FF8FFA55F894FF68F86C" italics="true" pageId="5" pageNumber="43">K. nubicola</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="7497A9AFFFA2FF8FFEDBF8B7FE55F86C" ID-CoL="6NK4W" box="[268,452,1913,1937]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="pleurostigma">
|
||
<emphasis id="81E30E3EFFA2FF8FFEDBF8B7FE55F86C" box="[268,452,1913,1937]" italics="true" pageId="5" pageNumber="43">K. pleurostigma</emphasis>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="7497A9AFFFA2FF8FFDDAF8B7FC8BF86C" ID-CoL="3QZF6" authority="Matsui 2009" authorityName="Matsui" authorityYear="2009" box="[525,794,1913,1937]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="yongi">
|
||
<emphasis id="81E30E3EFFA2FF8FFDDAF8B7FDE0F86C" box="[525,625,1914,1937]" italics="true" pageId="5" pageNumber="43">K. yongi</emphasis>
|
||
:
|
||
<bibRefCitation id="D706AFDDFFA2FF8FFD53F8B4FC8BF86C" author="Matsui" box="[644,794,1913,1937]" pageId="5" pageNumber="43" refString="Matsui, M. (2009) A new species of Kalophrynus with a unique male humeral spine from Peninsular Malaysia (Amphibia, Anura, Microhylidae). Zoological Science, 26, 579 - 585." type="journal article" year="2009">Matsui 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="D706AFDDFFA2FF8FFCFDF8B4FC5DF86C" author="Dehling" box="[810,972,1913,1937]" pageId="5" pageNumber="43" refString="Dehling, J. M. (2011) A new karst-dwelling species of Kalophrynus (Anura: Microhylidae) from Gunung Mulu National Park, Borneo, Malaysia. Zootaxa, 2737, 49 - 60." type="journal article" year="2011">Dehling 2011</bibRefCitation>
|
||
). The call of
|
||
<taxonomicName id="7497A9AFFFA2FF8FFBA5F8B7FA82F86C" ID-CoL="6NK56" box="[1138,1299,1913,1937]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA2FF8FFBA5F8B7FB14F86C" box="[1138,1157,1914,1937]" italics="true" pageId="5" pageNumber="43">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA2FF8FFB4FF8B4FA82F86C" box="[1176,1299,1913,1937]" italics="true" pageId="5" pageNumber="43">limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="81E30E3EFFA2FF8FFAC8F8B7FAECF86F" bold="true" box="[1311,1405,1914,1938]" pageId="5" pageNumber="43">
|
||
<taxonomicNameLabel id="9AD0B345FFA2FF8FFAC8F8B7FAECF86F" box="[1311,1405,1914,1938]" pageId="5" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is unpulsed unlike the well pulsed call of
|
||
<taxonomicName id="7497A9AFFFA2FF8FFD81F851FCE9F84F" ID-CoL="6NK59" authority="Dring 1984" authorityName="Dring" authorityYear="1984" box="[598,888,1946,1971]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="nubicola">
|
||
<emphasis id="81E30E3EFFA2FF8FFD81F851FD4BF84E" box="[598,730,1947,1971]" italics="true" pageId="5" pageNumber="43">K. nubicola</emphasis>
|
||
(
|
||
<bibRefCitation id="D706AFDDFFA2FF8FFD3DF857FCE0F84F" author="Dring" box="[746,881,1946,1970]" pageId="5" pageNumber="43" refString="Dring, J. C. M. (1984 " 1983 ") Some new frogs from Sarawak. Amphibia-Reptilia, 4, 103 - 115." type="journal article" year="1984">Dring 1984</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and is composed of successive notes unlike the short, single note in
|
||
<taxonomicName id="7497A9AFFFA2FF8FFE5FF870FD73F829" ID-CoL="6NK4Z" authority="Dehling 2011" authorityName="Dehling" authorityYear="2011" box="[392,738,1980,2004]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="calciphilus">
|
||
<emphasis id="81E30E3EFFA2FF8FFE5FF870FDB6F829" box="[392,551,1980,2004]" italics="true" pageId="5" pageNumber="43">K. calciphilus</emphasis>
|
||
(
|
||
<bibRefCitation id="D706AFDDFFA2FF8FFDEEF871FD4BF829" author="Dehling" box="[569,730,1980,2004]" pageId="5" pageNumber="43" refString="Dehling, J. M. (2011) A new karst-dwelling species of Kalophrynus (Anura: Microhylidae) from Gunung Mulu National Park, Borneo, Malaysia. Zootaxa, 2737, 49 - 60." type="journal article" year="2011">Dehling 2011</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA2FF8FFD26F870FCEEF829" ID-CoL="6NJRB" box="[753,895,1980,2004]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="baluensis">
|
||
<emphasis id="81E30E3EFFA2FF8FFD26F870FCEEF829" box="[753,895,1980,2004]" italics="true" pageId="5" pageNumber="43">K. baluensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA2FF8FFC46F870FBD0F829" box="[913,1089,1980,2004]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="heterochirus">
|
||
<emphasis id="81E30E3EFFA2FF8FFC46F870FBD0F829" box="[913,1089,1980,2004]" italics="true" pageId="5" pageNumber="43">K. heterochirus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="7497A9AFFFA2FF8FFB52F870FA09F829" ID-CoL="3QZF6" authority="Matsui 2009" authorityName="Matsui" authorityYear="2009" box="[1157,1432,1980,2004]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="yongi">
|
||
<emphasis id="81E30E3EFFA2FF8FFB52F870FB79F829" box="[1157,1256,1981,2004]" italics="true" pageId="5" pageNumber="43">K. yongi</emphasis>
|
||
(
|
||
<bibRefCitation id="D706AFDDFFA2FF8FFB2DF871FA01F829" author="Matsui" box="[1274,1424,1980,2004]" pageId="5" pageNumber="43" refString="Matsui, M. (2009) A new species of Kalophrynus with a unique male humeral spine from Peninsular Malaysia (Amphibia, Anura, Microhylidae). Zoological Science, 26, 579 - 585." type="journal article" year="2009">Matsui 2009</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="7497A9AFFFA2FF8FFF40F810FE38F808" ID-CoL="6NK56" box="[151,425,2013,2037]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA2FF8FFF40F810FE38F808" box="[151,425,2013,2037]" italics="true" pageId="5" pageNumber="43">Kalophrynus limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA2FF8FFE67F810FD9BF808" box="[432,522,2013,2037]" pageId="5" pageNumber="43" rank="species">
|
||
<emphasis id="81E30E3EFFA2FF8FFE67F810FD9BF808" bold="true" box="[432,522,2013,2037]" pageId="5" pageNumber="43">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
has a call with successive notes like
|
||
<taxonomicName id="7497A9AFFFA2FF8FFC7FF813FAD2F808" ID-CoL="6NK4W" authority="Matsui et al. 1996" authorityName="Matsui et al." authorityYear="1996" box="[936,1347,2013,2037]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="5" pageNumber="43" phylum="Chordata" rank="species" species="pleurostigma">
|
||
<emphasis id="81E30E3EFFA2FF8FFC7FF813FBCDF808" box="[936,1116,2013,2037]" italics="true" pageId="5" pageNumber="43">K. pleurostigma</emphasis>
|
||
(
|
||
<bibRefCitation id="D706AFDDFFA2FF8FFBBDF810FAADF808" author="Matsui" box="[1130,1340,2013,2037]" pageId="5" pageNumber="43" refString="Matsui, M., Chan-ard, T. & Nabhitabhata, J. (1996) Distinct specific status of Kalophrynus pleurostigma interlineatus (Anura, Microhylidae). Copeia, 1996, 440 - 445." type="journal article" year="1996">
|
||
Matsui
|
||
<emphasis id="81E30E3EFFA2FF8FFB16F813FB63F808" box="[1217,1266,2013,2037]" italics="true" pageId="5" pageNumber="43">et al</emphasis>
|
||
. 1996
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but the note length is shorter (61–76 vs. 170–487 ms), the note repetition rate is larger (3.3–4.2 vs. 0.9–2.7 notes per s), and the dominant frequency is much higher (
|
||
<date id="C729F4ECFFA1FF8CFD5EFF74FC99FF2C" box="[649,776,185,209]" pageId="6" pageNumber="44" value="1632" valueMax="2008">1632–2008</date>
|
||
vs. 438–575 hz) than
|
||
<taxonomicName id="7497A9AFFFA1FF8CFC28FF77FB23FF2C" ID-CoL="6NK4W" box="[1023,1202,185,209]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="pleurostigma">
|
||
<emphasis id="81E30E3EFFA1FF8CFC28FF77FB83FF2C" box="[1023,1042,186,209]" italics="true" pageId="6" pageNumber="44">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA1FF8CFBC8FF74FB23FF2C" box="[1055,1202,185,209]" italics="true" pageId="6" pageNumber="44">pleurostigma</emphasis>
|
||
</taxonomicName>
|
||
. The call of
|
||
<taxonomicName id="7497A9AFFFA1FF8CFAE9FF77FF61FF0E" ID-CoL="6NK4V" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="interlineatus">
|
||
<emphasis id="81E30E3EFFA1FF8CFAE9FF77FF61FF0E" italics="true" pageId="6" pageNumber="44">K. interlineatus</emphasis>
|
||
</taxonomicName>
|
||
is a long trill unlike that of
|
||
<taxonomicName id="7497A9AFFFA1FF8CFDF2FF11FC36FF0F" ID-CoL="6NK56" authority="Matsui et al. 1996" authorityName="Matsui et al." authorityYear="1996" box="[549,935,218,243]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="limbooliati">
|
||
<emphasis id="81E30E3EFFA1FF8CFDF2FF11FDA9FF0E" box="[549,568,220,243]" italics="true" pageId="6" pageNumber="44">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA1FF8CFD91FF16FD50FF0E" box="[582,705,219,243]" italics="true" pageId="6" pageNumber="44">limbooliati</emphasis>
|
||
(
|
||
<bibRefCitation id="D706AFDDFFA1FF8CFD18FF17FC0EFF0F" author="Matsui" box="[719,927,218,243]" pageId="6" pageNumber="44" refString="Matsui, M., Chan-ard, T. & Nabhitabhata, J. (1996) Distinct specific status of Kalophrynus pleurostigma interlineatus (Anura, Microhylidae). Copeia, 1996, 440 - 445." type="journal article" year="1996">
|
||
Matsui
|
||
<emphasis id="81E30E3EFFA1FF8CFCF2FF11FCC4FF0E" box="[805,853,219,243]" italics="true" pageId="6" pageNumber="44">et al</emphasis>
|
||
. 1996
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and that of
|
||
<taxonomicName id="7497A9AFFFA1FF8CFBE7FF11FB60FF0E" ID-CoL="6NJT8" box="[1072,1265,219,243]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="palmatissimus">
|
||
<emphasis id="81E30E3EFFA1FF8CFBE7FF11FB60FF0E" box="[1072,1265,219,243]" italics="true" pageId="6" pageNumber="44">K. palmatissimus</emphasis>
|
||
</taxonomicName>
|
||
(a soft “ko-koko”:
|
||
<bibRefCitation id="D706AFDDFFA1FF8CFF19FF31FEDFFEE9" author="Kiew" box="[206,334,252,276]" pageId="6" pageNumber="44" refString="Kiew, B. H. (1984) A new species of sticky frog (Kalophrynus palmatissimus n. sp.) from Peninsular Malaysia. Malayan Nature Journal, 37, 145 - 152." type="journal article" year="1984">Kiew 1984</bibRefCitation>
|
||
:150) is clearly very different from that of
|
||
<taxonomicName id="7497A9AFFFA1FF8CFCF2FF30FC51FEE9" ID-CoL="6NK56" box="[805,960,252,276]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA1FF8CFCF2FF30FCA9FEE9" box="[805,824,253,276]" italics="true" pageId="6" pageNumber="44">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA1FF8CFC92FF31FC51FEE9" box="[837,960,252,276]" italics="true" pageId="6" pageNumber="44">limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA1FF8CFC10FF31FBB0FEE9" box="[967,1057,252,276]" pageId="6" pageNumber="44" rank="species">
|
||
<emphasis id="81E30E3EFFA1FF8CFC10FF31FBB0FEE9" bold="true" box="[967,1057,252,276]" pageId="6" pageNumber="44">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<caption id="E7E882A4FFA1FF8CFF40FB61FB92FB1C" httpUri="https://zenodo.org/record/212552/files/figure.png" pageId="6" pageNumber="44" targetBox="[175,1392,348,1153]" targetPageId="6">
|
||
<paragraph id="B328D22CFFA1FF8CFF40FB61FB92FB1C" blockId="6.[151,1435,1196,1249]" pageId="6" pageNumber="44">
|
||
<emphasis id="81E30E3EFFA1FF8CFF40FB61FE83FB3C" bold="true" box="[151,274,1196,1219]" pageId="6" pageNumber="44">FIGURE 4.</emphasis>
|
||
Sonagram (A) and wave form (B) showing a nine note portion of an advertisement call of
|
||
<taxonomicName id="7497A9AFFFA1FF8CFB73FB61FA0AFB3F" ID-CoL="6NK56" box="[1188,1435,1196,1218]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA1FF8CFB73FB61FA0AFB3F" box="[1188,1435,1196,1218]" italics="true" pageId="6" pageNumber="44">Kalophrynus limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA1FF8CFF40FB06FF76FB1D" box="[151,231,1227,1248]" pageId="6" pageNumber="44" rank="species">
|
||
<emphasis id="81E30E3EFFA1FF8CFF40FB06FF76FB1D" bold="true" box="[151,231,1227,1248]" pageId="6" pageNumber="44">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from Gunung Lambak, Kluang, Johor, recorded at an air temperature 27.2 °C.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B328D22CFFA1FF8CFF12FAC4FACAFA05" blockId="6.[151,1436,1288,1637]" pageId="6" pageNumber="44">
|
||
A ranid,
|
||
<taxonomicName id="7497A9AFFFA1FF8CFEFEFAC5FDD0FADD" ID-CoL="6MMSM" box="[297,577,1288,1312]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="laterimaculata">
|
||
<emphasis id="81E30E3EFFA1FF8CFEFEFAC5FDD0FADD" box="[297,577,1288,1312]" italics="true" pageId="6" pageNumber="44">Hylarana laterimaculata</emphasis>
|
||
</taxonomicName>
|
||
was found syntopic with
|
||
<taxonomicName id="7497A9AFFFA1FF8CFCB3FAC4FB91FADD" ID-CoL="6NK56" box="[868,1024,1288,1312]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA1FF8CFCB3FAC4FCE6FADD" box="[868,887,1289,1312]" italics="true" pageId="6" pageNumber="44">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA1FF8CFC52FAC5FB91FADD" box="[901,1024,1288,1312]" italics="true" pageId="6" pageNumber="44">limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="81E30E3EFFA1FF8CFBDFFAC4FBF3FADC" bold="true" box="[1032,1122,1289,1313]" pageId="6" pageNumber="44">
|
||
<taxonomicNameLabel id="9AD0B345FFA1FF8CFBDFFAC4FBF3FADC" box="[1032,1122,1289,1313]" pageId="6" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in Pulai and Lambak. They emitted long calls very similar to the human ear, but details clearly differ between the two species. Calls of
|
||
<taxonomicName id="7497A9AFFFA1FF8CFA99FAE3FF7FFA95" ID-CoL="6NK56" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="limbooliati" status="sp. nov.">
|
||
<emphasis id="81E30E3EFFA1FF8CFA99FAE3FAF0FAB8" box="[1358,1377,1326,1349]" italics="true" pageId="6" pageNumber="44">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA1FF8CFAB8FAE0FF7FFA95" italics="true" pageId="6" pageNumber="44">limbooliati</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9AD0B345FFA1FF8CFF22FA9CFEDEFA95" box="[245,335,1361,1384]" pageId="6" pageNumber="44" rank="species">
|
||
<emphasis id="81E30E3EFFA1FF8CFF22FA9CFEDEFA95" bold="true" box="[245,335,1361,1384]" pageId="6" pageNumber="44">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
were longer than
|
||
<taxonomicName id="7497A9AFFFA1FF8CFDCCFA9CFD73FA95" ID-CoL="6MMSM" box="[539,738,1360,1384]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="laterimaculata">
|
||
<emphasis id="81E30E3EFFA1FF8CFDCCFA9CFD73FA95" box="[539,738,1360,1384]" italics="true" pageId="6" pageNumber="44">H. laterimaculata</emphasis>
|
||
</taxonomicName>
|
||
recorded at Lambak (mean±SD =10.9±2.8 vs. 6.0+
|
||
<geoCoordinate id="D6A3B4EBFFA1FF8CFAF0FA9DFACDFA95" box="[1319,1372,1360,1384]" direction="south" orientation="latitude" pageId="6" pageNumber="44" precision="5555" value="-0.2">0.2 s</geoCoordinate>
|
||
in
|
||
<taxonomicName id="7497A9AFFFA1FF8CFA56FA9CFEADFA70" ID-CoL="6MMSM" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="laterimaculata">
|
||
<emphasis id="81E30E3EFFA1FF8CFA56FA9CFEADFA70" italics="true" pageId="6" pageNumber="44">H. laterimaculata</emphasis>
|
||
</taxonomicName>
|
||
) and included larger number of notes (mean±SD = 41.1±11.2 vs. 12.5±0.6). Each note lasted shorter (mean±SD = 70.6±6.0 vs. 109.4±4.4 ms), with shorter time interval between two notes (mean±SD = 271.7± 25.4 vs. 520.3±18.2 ms) and larger note repetition rate (mean = 3.72±0.30 vs. 1.96 notes per s). In addition, the dominant frequency was evidently lower (mean±SD = 1909±156 vs. 2897±123 hz) than that of
|
||
<taxonomicName id="7497A9AFFFA1FF8CFB5AFA2CFAC5FA05" ID-CoL="6MMSM" box="[1165,1364,1504,1528]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="laterimaculata">
|
||
<emphasis id="81E30E3EFFA1FF8CFB5AFA2CFAC5FA05" box="[1165,1364,1504,1528]" italics="true" pageId="6" pageNumber="44">H. laterimaculata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B328D22CFFA1FF8CFF12F9C9FD64F999" blockId="6.[151,1436,1288,1637]" pageId="6" pageNumber="44">
|
||
Finally, the uncorrected pairwise sequence divergences of mitochondrial 16S rRNA gene between
|
||
<taxonomicName id="7497A9AFFFA1FF8CFACBF9CBFF2CF9BD" ID-CoL="6NK56" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="limbooliati">
|
||
<emphasis id="81E30E3EFFA1FF8CFACBF9CBFF2CF9BD" italics="true" pageId="6" pageNumber="44">K. limbooliati</emphasis>
|
||
</taxonomicName>
|
||
and the seven congeners (
|
||
<taxonomicName id="7497A9AFFFA1FF8CFE3FF9E4FD04F9BD" box="[488,661,1576,1600]" class="Amphibia" family="Microhylidae" genus="Kaloula" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="heterochirus">
|
||
<emphasis id="81E30E3EFFA1FF8CFE3FF9E4FE6AF9BD" box="[488,507,1577,1600]" italics="true" pageId="6" pageNumber="44">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA1FF8CFDDEF9E5FD04F9BD" box="[521,661,1576,1600]" italics="true" pageId="6" pageNumber="44">heterochirus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA1FF8CFD73F9E4FCC3F9BD" ID-CoL="6NK4V" box="[676,850,1576,1600]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="interlineatus">
|
||
<emphasis id="81E30E3EFFA1FF8CFD73F9E4FD26F9BD" box="[676,695,1577,1600]" italics="true" pageId="6" pageNumber="44">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA1FF8CFD12F9E5FCC3F9BD" box="[709,850,1576,1600]" italics="true" pageId="6" pageNumber="44">interlineatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA1FF8CFCB5F9E4FBB2F9BD" ID-CoL="6NJT8" box="[866,1059,1576,1600]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="palmatissimus">
|
||
<emphasis id="81E30E3EFFA1FF8CFCB5F9E4FCE4F9BD" box="[866,885,1577,1600]" italics="true" pageId="6" pageNumber="44">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA1FF8CFC54F9E5FBB2F9BD" box="[899,1059,1576,1600]" italics="true" pageId="6" pageNumber="44">palmatissimus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA1FF8CFBE5F9E4FB77F9BD" ID-CoL="6NK4W" box="[1074,1254,1576,1600]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="pleurostigma">
|
||
<emphasis id="81E30E3EFFA1FF8CFBE5F9E4FBD4F9BD" box="[1074,1093,1577,1600]" italics="true" pageId="6" pageNumber="44">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA1FF8CFB83F9E5FB77F9BD" box="[1108,1254,1576,1600]" italics="true" pageId="6" pageNumber="44">pleurostigma</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA1FF8CFB22F9E4FAE2F9BD" ID-CoL="3QZF3" box="[1269,1395,1576,1600]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="stellatus">
|
||
<emphasis id="81E30E3EFFA1FF8CFB22F9E4FA99F9BD" box="[1269,1288,1577,1600]" italics="true" pageId="6" pageNumber="44">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA1FF8CFAC1F9E5FAE2F9BD" box="[1302,1395,1576,1600]" italics="true" pageId="6" pageNumber="44">stellatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7497A9AFFFA1FF8CFA55F9E4FEB2F998" ID-CoL="3QZF4" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="subterrestris">
|
||
<emphasis id="81E30E3EFFA1FF8CFA55F9E4FA04F9BD" box="[1410,1429,1577,1600]" italics="true" pageId="6" pageNumber="44">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA1FF8CFF40F980FEB2F998" box="[151,291,1613,1637]" italics="true" pageId="6" pageNumber="44">subterrestris</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="7497A9AFFFA1FF8CFEB6F983FE2EF998" ID-CoL="3QZF6" box="[353,447,1613,1637]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="6" pageNumber="44" phylum="Chordata" rank="species" species="yongi">
|
||
<emphasis id="81E30E3EFFA1FF8CFEB6F983FEE5F998" box="[353,372,1614,1637]" italics="true" pageId="6" pageNumber="44">K</emphasis>
|
||
.
|
||
<emphasis id="81E30E3EFFA1FF8CFE56F983FE2EF998" box="[385,447,1614,1637]" italics="true" pageId="6" pageNumber="44">yongi</emphasis>
|
||
</taxonomicName>
|
||
) are as large as 5.2–10.5%.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |