treatments-xml/data/03/C2/87/03C287E8FFEFFFDEFF4BFB40FB48E5C7.xml
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<document id="68113B9335955BB8468ED8A692876A01" ID-CLB-Dataset="182444" ID-DOI="10.11646/zootaxa.5202.1.1" ID-GBIF-Dataset="45ba25c1-47a9-4001-93d9-294815bf37e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="7285600" ID-ZooBank="BF4831D2-D98B-4265-9138-03DB8607B826" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1667554680289" checkinUser="plazi" docAuthor="Sánchez-Vialas, Alberto, Calvo-Revuelta, Marta &amp; Riva, Ignacio De La" docDate="2022" docId="03C287E8FFEFFFDEFF4BFB40FB48E5C7" docLanguage="en" docName="zootaxa.5202.1.1.pdf" docOrigin="Zootaxa 5202 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5202.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Philothamnus nitidus" docType="treatment" docVersion="6" lastPageNumber="78" masterDocId="FFFBFF90FFA2FF93FFDCFF9DFFB5E232" masterDocTitle="Synopsis of the terrestrial Reptiles of Equatorial Guinea" masterLastPageNumber="197" masterPageNumber="1" pageNumber="78" updateTime="1699409742602" updateUser="ExternalLinkService">
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<mods:title id="36988B2786C30E1D80C98CB8330229B3">Synopsis of the terrestrial Reptiles of Equatorial Guinea</mods:title>
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<heading id="D09C8192FFEFFFDEFF4BFB40FDDFE6C5" bold="true" box="[151,618,1245,1272]" fontSize="11" level="2" pageId="77" pageNumber="78" reason="4">
<taxonomicName id="4C6B4D7DFFEFFFDEFF4BFB40FDDFE6C5" authority="(Gunther, 1863)" baseAuthorityName="Gunther" baseAuthorityYear="1863" box="[151,618,1245,1272]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="nitidus">
<emphasis id="B91FEAECFFEFFFDEFF4BFB40FDDFE6C5" bold="true" box="[151,618,1245,1272]" pageId="77" pageNumber="78">
<emphasis id="B91FEAECFFEFFFDEFF4BFB40FE28E6CA" bold="true" box="[151,413,1245,1272]" italics="true" pageId="77" pageNumber="78">Philothamnus nitidus</emphasis>
(Günther, 1863)
</emphasis>
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<subSubSection id="C3716575FFEFFFDEFF4BFA9CFEA3E729" box="[151,278,1281,1307]" pageId="77" pageNumber="78" type="description">
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<figureCitation id="13502A7BFFEFFFDEFF4BFA9CFEA3E729" box="[151,278,1281,1307]" captionStart="FIGURE 19" captionStartId="80.[151,250,1910,1934]" captionTargetBox="[169,1433,196,1867]" captionTargetId="figure-19@80.[154,1433,181,1886]" captionTargetPageId="80" captionText="FIGURE 19. From left to right: lateral, dorsal and ventral views of the head of: Philothamnus girardi (A) (EBD 27758), Philothamnus heterodermus (B) (MNCN 22864), and Philothamnus nitidus (C) (EBD 27534). Photos: AS-V." figureDoi="http://doi.org/10.5281/zenodo.7285679" httpUri="https://zenodo.org/record/7285679/files/figure.png" pageId="77" pageNumber="78">Figure 19C</figureCitation>
</paragraph>
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<subSubSection id="C3716575FFEFFFDEFF4BFADBFA3FE7EB" pageId="77" pageNumber="78" type="reference_group">
<paragraph id="8BD436FEFFEFFFDEFF4BFADBFEA7E74F" blockId="77.[151,1437,1350,1497]" pageId="77" pageNumber="78">
<treatmentCitationGroup id="AB7B11D0FFEFFFDEFF4BFADBFEA7E74F" pageId="77" pageNumber="78">
<treatmentCitation id="0ACA10EFFFEFFFDEFF4BFADBFDA4E76C" author="Gunther, A." box="[151,529,1350,1374]" page="286" pageId="77" pageNumber="78" year="1863">
<taxonomicName id="4C6B4D7DFFEFFFDEFF4BFADBFDA4E76C" authority="Gunther, 1863 c: 286" authorityName="Gunther" authorityPageNumber="286" authorityYear="1863" box="[151,529,1350,1374]" class="Reptilia" family="Colubridae" genus="Ahaetulla" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="nitida">
<emphasis id="B91FEAECFFEFFFDEFF4BFADBFE8FE76C" box="[151,314,1350,1374]" italics="true" pageId="77" pageNumber="78">Ahaetulla nitida</emphasis>
<bibRefCitation id="EFFA4B0FFFEFFFDEFE9CFADBFDA4E76C" author="Gunther, A." box="[320,529,1350,1374]" pageId="77" pageNumber="78" pagination="283 - 287" refId="ref78995" refString="Gunther, A. (1863 c) On some species of tree snakes (Ahaetulla). Annals &amp; Magazine of Natural History, Series 3, 11 (64), 283 - 287. https: // doi. org / 10.1080 / 00222936308681421" type="journal article" year="1863">Günther, 1863c: 286</bibRefCitation>
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.
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locality: “
<collectingCountry id="F37C766EFFEFFFDEFD6FFADBFCA9E76C" box="[691,796,1350,1374]" name="Guyana" pageId="77" pageNumber="78">Demerara</collectingCountry>
<collectingCountry id="F37C766EFFEFFFDEFCF6FADBFCCCE76C" box="[810,889,1350,1374]" name="Guyana" pageId="77" pageNumber="78">Guyana</collectingCountry>
(in error), corrected to “
<collectingRegion id="49AFF81CFFEFFFDEFBB0FADBFB1AE76C" box="[1132,1199,1350,1374]" country="Nigeria" name="Lagos" pageId="77" pageNumber="78">Lagos</collectingRegion>
”,
<collectingCountry id="F37C766EFFEFFFDEFB1EFADBFABBE76C" box="[1218,1294,1350,1374]" name="Nigeria" pageId="77" pageNumber="78">Nigeria</collectingCountry>
by
<bibRefCitation id="EFFA4B0FFFEFFFDEFAEFFADBFEBBE74F" author="Boulenger, G. A." pageId="77" pageNumber="78" refId="ref71762" refString="Boulenger, G. A. (1894) Catalogue of the Snakes in the British Museum (Natural History). Vol. 2. Trustees, London, xi + 382 pp., 20 pls." type="book" year="1894">Boulenger (1894)</bibRefCitation>
.
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</paragraph>
<paragraph id="8BD436FEFFEFFFDEFF4BFA1EFCC4E7A9" blockId="77.[151,1437,1350,1497]" box="[151,881,1411,1436]" pageId="77" pageNumber="78">
<treatmentCitationGroup id="AB7B11D0FFEFFFDEFF4BFA1EFCC4E7A9" box="[151,881,1411,1436]" pageId="77" pageNumber="78">
<treatmentCitation id="0ACA10EFFFEFFFDEFF4BFA1EFD94E7A9" author="Gunther, A." box="[151,545,1411,1435]" page="26" pageId="77" pageNumber="78" year="1872">
<taxonomicName id="4C6B4D7DFFEFFFDEFF4BFA1EFD94E7A9" authority="Gunther, 1872: 26" authorityName="Gunther" authorityPageNumber="26" authorityYear="1872" box="[151,545,1411,1435]" class="Reptilia" family="Colubridae" genus="Ahaetulla" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="lagoensis">
<emphasis id="B91FEAECFFEFFFDEFF4BFA1EFED5E7A9" box="[151,352,1411,1435]" italics="true" pageId="77" pageNumber="78">Ahaetulla lagoensis</emphasis>
<bibRefCitation id="EFFA4B0FFFEFFFDEFEBAFA1EFD94E7A9" author="Gunther, A." box="[358,545,1411,1435]" pageId="77" pageNumber="78" pagination="13 - 37" refId="ref79093" refString="Gunther, A. (1872) Seventh account of new species of snakes in the collection of the British Museum. Annals &amp; Magazine of Natural History, Series 4, 9 (49), 13 - 37. https: // doi. org / 10.1080 / 002229372011951771" type="journal article" year="1872">Günther, 1872: 26</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54D0885CFFEFFFDEFDF6FA19FDE9E7AE" box="[554,604,1412,1436]" pageId="77" pageNumber="78">Type</typeStatus>
locality: “
<collectingRegion id="49AFF81CFFEFFFDEFD1AFA1EFCBCE7A9" box="[710,777,1411,1435]" country="Nigeria" name="Lagos" pageId="77" pageNumber="78">Lagos</collectingRegion>
”,
<collectingCountry id="F37C766EFFEFFFDEFCC1FA1EFCD9E7A9" box="[797,876,1411,1435]" name="Nigeria" pageId="77" pageNumber="78">Nigeria</collectingCountry>
.
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</paragraph>
<paragraph id="8BD436FEFFEFFFDEFF4BFA3FFCB7E788" blockId="77.[151,1437,1350,1497]" box="[151,770,1442,1466]" pageId="77" pageNumber="78">
<treatmentCitationGroup id="AB7B11D0FFEFFFDEFF4BFA3FFCB7E788" box="[151,770,1442,1466]" pageId="77" pageNumber="78">
<taxonomicName id="4C6B4D7DFFEFFFDEFF4BFA3FFDA9E788" authority="(Gunther, 1863)" baseAuthorityName="Gunther" baseAuthorityYear="1863" box="[151,540,1442,1466]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="nitidus">
<emphasis id="B91FEAECFFEFFFDEFF4BFA3FFEDAE788" box="[151,367,1442,1466]" italics="true" pageId="77" pageNumber="78">Philothamnus nitidus</emphasis>
(Günther, 1863)
</taxonomicName>
:
<treatmentCitation id="0ACA10EFFFEFFFDEFDF9FA3FFD4BE788" author="Boulenger, G. A." box="[549,766,1442,1466]" page="100" pageId="77" pageNumber="78" year="1894">
<bibRefCitation id="EFFA4B0FFFEFFFDEFDF9FA3FFD4BE788" author="Boulenger, G. A." box="[549,766,1442,1466]" pageId="77" pageNumber="78" refId="ref71762" refString="Boulenger, G. A. (1894) Catalogue of the Snakes in the British Museum (Natural History). Vol. 2. Trustees, London, xi + 382 pp., 20 pls." type="book" year="1894">Boulenger 1894: 100</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD436FEFFEFFFDEFF4BFA5DFA3FE7EB" blockId="77.[151,1437,1350,1497]" box="[151,1418,1472,1497]" pageId="77" pageNumber="78">
<treatmentCitationGroup id="AB7B11D0FFEFFFDEFF4BFA5DFA3FE7EB" box="[151,1418,1472,1497]" pageId="77" pageNumber="78">
<treatmentCitation id="0ACA10EFFFEFFFDEFF4BFA5DFD2FE7EB" author="Laurent, R. F." box="[151,666,1472,1497]" page="40" pageId="77" pageNumber="78" year="1960">
<taxonomicName id="4C6B4D7DFFEFFFDEFF4BFA5DFD2FE7EB" authority="Laurent, 1960: 40" authorityName="Laurent" authorityPageNumber="40" authorityYear="1960" box="[151,666,1472,1497]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="subSpecies" species="nitidus" subSpecies="loveridgei">
<emphasis id="B91FEAECFFEFFFDEFF4BFA5DFE68E7EA" box="[151,477,1472,1496]" italics="true" pageId="77" pageNumber="78">Philothamnus nitidus loveridgei</emphasis>
<bibRefCitation id="EFFA4B0FFFEFFFDEFE3FFA5CFD2FE7EB" author="Laurent, R. F." box="[483,666,1472,1497]" pageId="77" pageNumber="78" pagination="1 - 86" refId="ref81959" refString="Laurent, R. F. (1960) Notes complementaires sur les cheloniens et les ophidiens du Congo Oriental. Annales Du Musee Royal Du Congo Belge (Sciences Zoologiques), 84, 1 - 86." type="journal article" year="1960">Laurent, 1960: 40</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54D0885CFFEFFFDEFD78FA5CFD63E7EB" box="[676,726,1473,1497]" pageId="77" pageNumber="78">Type</typeStatus>
locality:
<location id="8EB46025FFEFFFDEFCEAFA5DFCD2E7EA" LSID="urn:lsid:plazi:treatment:03C287E8FFEFFFDEFF4BFB40FB48E5C7:8EB46025FFEFFFDEFCEAFA5DFCD2E7EA" box="[822,871,1472,1496]" country="Democratic Republic of the Congo" county="Kivu" municipality="Shabunda" name="Itula" pageId="77" pageNumber="78">Itula</location>
,
<collectingMunicipality id="6BB0AC84FFEFFFDEFCADFA5DFC6DE7EA" box="[881,984,1472,1496]" pageId="77" pageNumber="78">Shabunda</collectingMunicipality>
,
<collectingCounty id="62B54E72FFEFFFDEFC3DFA5DFBA3E7EA" box="[993,1046,1472,1496]" pageId="77" pageNumber="78">Kivu</collectingCounty>
,
<collectingCountry id="F37C766EFFEFFFDEFBFDFA5DFA32E7EB" box="[1057,1415,1472,1497]" name="Democratic Republic of the Congo" pageId="77" pageNumber="78">Democratic Republic of the Congo</collectingCountry>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFEFFFDEFF4BF99BFDEEE483" pageId="77" pageNumber="78" type="distribution">
<paragraph id="8BD436FEFFEFFFDEFF4BF99BFDEEE483" blockId="77.[151,1437,1542,2037]" pageId="77" pageNumber="78">
<emphasis id="B91FEAECFFEFFFDEFF4BF99BFE9CE412" bold="true" box="[151,297,1542,1568]" pageId="77" pageNumber="78">Distribution</emphasis>
. This species extends from
<collectingCountry id="F37C766EFFEFFFDEFD5FF99AFCACE413" box="[643,793,1543,1569]" name="Sierra Leone" pageId="77" pageNumber="78">Sierra Leone</collectingCountry>
and
<collectingCountry id="F37C766EFFEFFFDEFC81F99AFC1BE413" box="[861,942,1543,1569]" name="Guinea-Bissau" pageId="77" pageNumber="78">Guinea</collectingCountry>
to
<collectingCountry id="F37C766EFFEFFFDEFC03F99AFB99E413" box="[991,1068,1543,1569]" name="Democratic Republic of the Congo" pageId="77" pageNumber="78">Congo</collectingCountry>
, and southward to
<collectingCountry id="F37C766EFFEFFFDEFAC6F99AFE2DE477" name="Democratic Republic of the Congo" pageId="77" pageNumber="78">Democratic Republic of the Congo</collectingCountry>
and
<collectingCountry id="F37C766EFFEFFFDEFE13F9B6FD90E477" box="[463,549,1579,1605]" name="Angola" pageId="77" pageNumber="78">Angola</collectingCountry>
. In
<collectingCountry id="F37C766EFFEFFFDEFD93F9B6FCA9E477" box="[591,796,1579,1605]" name="Equatorial Guinea" pageId="77" pageNumber="78">Equatorial Guinea</collectingCountry>
it has been recorded in Bioko at San Carlos de Luba [as
<taxonomicName id="4C6B4D7DFFEFFFDEFF4BF9D2FEE8E45A" baseAuthorityName="Smith" baseAuthorityYear="1847" box="[151,349,1615,1640]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="semivariegatus">
<emphasis id="B91FEAECFFEFFFDEFF4BF9D2FEE8E45A" box="[151,349,1615,1640]" italics="true" pageId="77" pageNumber="78">P. semivariegatus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFFA4B0FFFEFFFDEFEB7F9D2FDBAE45B" author="Bocage, J. V. B." box="[363,527,1614,1641]" pageId="77" pageNumber="78" pagination="1 - 15" refId="ref70929" refString="Bocage, J. V. B. (1895 a) Subsidios para a fauna da Ilha de Fernao do Po. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 4 (13), 1 - 15." type="journal article" year="1895">
Bocage 1895
<bibRefCitation id="EFFA4B0FFFEFFFDEFDDEF9D3FDBAE45B" author="Boulenger, G. A." box="[514,527,1614,1641]" pageId="77" pageNumber="78" pagination="350 - 354" refId="ref72138" refString="Boulenger, G. A. (1903) Descriptions of new snakes in the collection of the British Museum. Annals &amp; Magazine of Natural History, Series 7, 12 (69), 350 - 354. https: // doi. org / 10.1080 / 00222930308678866" type="journal article" year="1903">a</bibRefCitation>
</bibRefCitation>
, 1903;
<bibRefCitation id="EFFA4B0FFFEFFFDEFDBDF9D2FCA2E45B" author="Loveridge, A." box="[609,791,1614,1641]" pageId="77" pageNumber="78" pagination="1 - 198" refId="ref83221" refString="Loveridge, A. (1958) Revision of five african snake genera. Bulletin of the Museum of Comparative Zoology, 119, 1 - 198." type="journal article" year="1958">Loveridge 1958</bibRefCitation>
)] and in
<collectingCountry id="F37C766EFFEFFFDEFCA5F9D2FC53E45B" box="[889,998,1615,1641]" name="Equatorial Guinea" pageId="77" pageNumber="78">Río Muni</collectingCountry>
at Monte Alén National Park (
<bibRefCitation id="EFFA4B0FFFEFFFDEFAE2F9D2FF4DE4BF" author="Lasso, C. A. &amp; Rial, A. I. &amp; Castroviejo, J. &amp; De la Riva, I." pageId="77" pageNumber="78" pagination="21 - 34" refId="ref81678" refString="Lasso, C. A., Rial, A. I., Castroviejo, J. &amp; De la Riva, I. (2002) Herpetofauna del Parque Nacional de Monte Alen (Rio Muni, Guinea Ecuatorial). Graellsia, 58, 21 - 34. https: // doi. org / 10.3989 / graellsia. 2002. v 58. i 2.276" type="journal article" year="2002">
Lasso
<emphasis id="B91FEAECFFEFFFDEFA54F9D2FF01E4BF" italics="true" pageId="77" pageNumber="78">et al.</emphasis>
2002
</bibRefCitation>
).
<bibRefCitation id="EFFA4B0FFFEFFFDEFED7F9EEFE65E4BF" author="Loveridge, A." box="[267,464,1650,1677]" pageId="77" pageNumber="78" pagination="1 - 198" refId="ref83221" refString="Loveridge, A. (1958) Revision of five african snake genera. Bulletin of the Museum of Comparative Zoology, 119, 1 - 198." type="journal article" year="1958">Loveridge (1958)</bibRefCitation>
suggested that
<bibRefCitation id="EFFA4B0FFFEFFFDEFD5CF9EEFCF0E4BF" author="Bocage, J. V. B." box="[640,837,1650,1677]" pageId="77" pageNumber="78" pagination="1 - 15" refId="ref70929" refString="Bocage, J. V. B. (1895 a) Subsidios para a fauna da Ilha de Fernao do Po. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 4 (13), 1 - 15." type="journal article" year="1895">Bocages (1895a)</bibRefCitation>
record of
<taxonomicName id="4C6B4D7DFFEFFFDEFC66F9EEFB35E4BE" baseAuthorityName="Smith" baseAuthorityYear="1847" box="[954,1152,1651,1676]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="semivariegatus">
<emphasis id="B91FEAECFFEFFFDEFC66F9EEFB35E4BE" box="[954,1152,1651,1676]" italics="true" pageId="77" pageNumber="78">P. semivariegatus</emphasis>
</taxonomicName>
probably belongs to
<emphasis id="B91FEAECFFEFFFDEFAB2F9EEFF56E483" italics="true" pageId="77" pageNumber="78">
P. s.
<taxonomicName id="4C6B4D7DFFEFFFDEFF4BF90AFF56E483" baseAuthorityName="Gunther" baseAuthorityYear="1863" box="[151,227,1687,1713]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="nitidus">nitidus</taxonomicName>
</emphasis>
(currently
<taxonomicName id="4C6B4D7DFFEFFFDEFEBCF90AFE72E483" baseAuthorityName="Gunther" baseAuthorityYear="1863" box="[352,455,1687,1713]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="nitidus">
<emphasis id="B91FEAECFFEFFFDEFEBCF90AFE72E483" box="[352,455,1687,1713]" italics="true" pageId="77" pageNumber="78">P. nitidus</emphasis>
</taxonomicName>
) (
<figureCitation id="13502A7BFFEFFFDEFE03F90AFDFBE483" box="[479,590,1687,1713]" captionStart="MAPS 22" captionStartId="81.[151,223,1943,1967]" captionTargetBox="[202,1385,192,1908]" captionTargetId="figure-18@81.[159,1429,181,1919]" captionTargetPageId="81" captionText="MAPS 22 AC. Distribution maps for Equatorial Guinean records of (A) Philothamnus heterodermus; (B) Philothamnus nitidus; (C) Thelotornis kirtlandii." figureDoi="http://doi.org/10.5281/zenodo.7304729" httpUri="https://zenodo.org/record/7304729/files/figure.png" pageId="77" pageNumber="78">Map 22B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3716575FFEFFFDEFF1BF927FB48E5C7" pageId="77" pageNumber="78" type="discussion">
<paragraph id="8BD436FEFFEFFFDEFF1BF927FB1FE573" blockId="77.[151,1437,1542,2037]" pageId="77" pageNumber="78">
<emphasis id="B91FEAECFFEFFFDEFF1BF927FEFAE4E6" bold="true" box="[199,335,1722,1748]" pageId="77" pageNumber="78">Comments.</emphasis>
Previous reports on
<taxonomicName id="4C6B4D7DFFEFFFDEFDF3F926FD47E4E6" baseAuthorityName="Smith" baseAuthorityYear="1847" box="[559,754,1723,1748]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="semivariegatus">
<emphasis id="B91FEAECFFEFFFDEFDF3F926FD47E4E6" box="[559,754,1723,1748]" italics="true" pageId="77" pageNumber="78">P. semivariegatus</emphasis>
</taxonomicName>
in Bioko are herein treated as
<taxonomicName id="4C6B4D7DFFEFFFDEFBE6F926FB15E4E7" baseAuthorityName="Gunther" baseAuthorityYear="1863" box="[1082,1184,1723,1749]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="nitidus">
<emphasis id="B91FEAECFFEFFFDEFBE6F926FB15E4E7" box="[1082,1184,1723,1749]" italics="true" pageId="77" pageNumber="78">P. nitidus</emphasis>
</taxonomicName>
following
<bibRefCitation id="EFFA4B0FFFEFFFDEFACAF926FF57E4CB" author="Loveridge, A." pageId="77" pageNumber="78" pagination="1 - 198" refId="ref83221" refString="Loveridge, A. (1958) Revision of five african snake genera. Bulletin of the Museum of Comparative Zoology, 119, 1 - 198." type="journal article" year="1958">Loveridges (1958)</bibRefCitation>
assumption on the identity of the Biokoan population [“
<collectingCountry id="F37C766EFFEFFFDEFCB3F942FBBFE4CB" box="[879,1034,1759,1785]" name="Equatorial Guinea" pageId="77" pageNumber="78">Fernando Po</collectingCountry>
: San Carlos (
<bibRefCitation id="EFFA4B0FFFEFFFDEFB7AF942FAE0E4CB" author="Bocage, J. V. B." box="[1190,1365,1758,1785]" pageId="77" pageNumber="78" pagination="1 - 15" refId="ref70929" refString="Bocage, J. V. B. (1895 a) Subsidios para a fauna da Ilha de Fernao do Po. Jornal de Sciencias Mathematicas, Physicas e Naturaes, Segunda Serie, 4 (13), 1 - 15." type="journal article" year="1895">Bocage, 1895a</bibRefCitation>
: more probably
<emphasis id="B91FEAECFFEFFFDEFEDDF89EFED0E52F" box="[257,357,1795,1821]" italics="true" pageId="77" pageNumber="78">
s.
<taxonomicName id="4C6B4D7DFFEFFFDEFEC5F89EFED0E52F" baseAuthorityName="Gunther" baseAuthorityYear="1863" box="[281,357,1795,1821]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="nitidus">nitidus</taxonomicName>
</emphasis>
)”
<emphasis id="B91FEAECFFEFFFDEFE5EF89EFE19E52F" box="[386,428,1795,1821]" italics="true" pageId="77" pageNumber="78">fide</emphasis>
<bibRefCitation id="EFFA4B0FFFEFFFDEFE6EF89EFDCFE52F" author="Loveridge, A." box="[434,634,1794,1821]" pageId="77" pageNumber="78" pagination="1 - 198" refId="ref83221" refString="Loveridge, A. (1958) Revision of five african snake genera. Bulletin of the Museum of Comparative Zoology, 119, 1 - 198." type="journal article" year="1958">Loveridge (1958)</bibRefCitation>
: 119].
<taxonomicName id="4C6B4D7DFFEFFFDEFD18F89EFC07E52F" baseAuthorityName="Gunther" baseAuthorityYear="1863" box="[708,946,1795,1821]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="nitidus">
<emphasis id="B91FEAECFFEFFFDEFD18F89EFC07E52F" box="[708,946,1795,1821]" italics="true" pageId="77" pageNumber="78">Philothamnus nitidus</emphasis>
</taxonomicName>
is the sister taxon to
<taxonomicName id="4C6B4D7DFFEFFFDEFB40F89EFA9AE52F" baseAuthorityName="Bocage" baseAuthorityYear="1866" box="[1180,1327,1795,1821]" class="Reptilia" family="Typhlopidae" genus="Afrotyphlops" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="B91FEAECFFEFFFDEFB40F89EFA9AE52F" box="[1180,1327,1795,1821]" italics="true" pageId="77" pageNumber="78">P. angolensis</emphasis>
</taxonomicName>
, and both are nested within the paraphyletic assemblage of
<taxonomicName id="4C6B4D7DFFEFFFDEFD62F8BAFB13E573" authority="(Engelbrecht et al. 2019)" baseAuthorityName="Engelbrecht" baseAuthorityYear="2019" box="[702,1190,1831,1857]" class="Reptilia" family="Colubridae" genus="Philothamnus" kingdom="Animalia" order="Squamata" pageId="77" pageNumber="78" phylum="Chordata" rank="species" species="semivariegatus">
<emphasis id="B91FEAECFFEFFFDEFD62F8BAFC31E572" box="[702,900,1831,1856]" italics="true" pageId="77" pageNumber="78">P. semivariegatus</emphasis>
(
<bibRefCitation id="EFFA4B0FFFEFFFDEFC4EF8BAFB2BE573" author="Engelbrecht, H. M. &amp; Branch, W. R. &amp; Greenbaum, E. &amp; Alexander, G. J. &amp; Jackson, K. &amp; Burger, M. &amp; Conradie, W. &amp; Kusamba, C. &amp; ZassiBoulou, A-G. &amp; Tolley, K. A." box="[914,1182,1831,1857]" pageId="77" pageNumber="78" pagination="357 - 365" refId="ref76556" refString="Engelbrecht, H. M., Branch, W. R., Greenbaum, E., Alexander, G. J., Jackson, K., Burger, M., Conradie, W., Kusamba, C., ZassiBoulou, A-G. &amp; Tolley, K. A. (2019) Diversifying into the branches: species boundaries in African green and bush snakes, Philothamnus (Serpentes: Colubridae). Molecular Phylogenetics and Evolution, 130, 357 - 365. https: // doi. org / 10.1016 / j. ympev. 2018.10.023" type="journal article" year="2019">
Engelbrecht
<emphasis id="B91FEAECFFEFFFDEFBFEF8BAFBEFE573" box="[1058,1114,1831,1857]" italics="true" pageId="77" pageNumber="78">et al.</emphasis>
2019
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BD436FEFFEFFFDEFF1BF8D7FB48E5C7" blockId="77.[151,1437,1542,2037]" pageId="77" pageNumber="78">
<emphasis id="B91FEAECFFEFFFDEFF1BF8D7FE73E556" bold="true" box="[199,454,1866,1892]" pageId="77" pageNumber="78">Specimens examined.</emphasis>
<materialsCitation id="3B033CA3FFEFFFDEFE17F8D6FAAAE557" ID-GBIF-Occurrence="3962324795" accessionNumber="EBD 20931, EBD 20789" box="[459,1311,1866,1893]" collectorName="Ten" country="Equatorial Guinea" location="Miboman" pageId="77" pageNumber="78" specimenCount="1">
<collectorName id="269E5328FFEFFFDEFE17F8D6FE40E557" box="[459,501,1867,1893]" pageId="77" pageNumber="78">Ten</collectorName>
specimens.
<collectingCountry id="F37C766EFFEFFFDEFDA1F8D6FD59E557" box="[637,748,1867,1893]" name="Equatorial Guinea" pageId="77" pageNumber="78">Río Muni</collectingCountry>
:
<location id="8EB46025FFEFFFDEFD2AF8D6FCD3E557" LSID="urn:lsid:plazi:treatment:03C287E8FFEFFFDEFF4BFB40FB48E5C7:8EB46025FFEFFFDEFD2AF8D6FCD3E557" box="[758,870,1867,1893]" country="Equatorial Guinea" name="Miboman" pageId="77" pageNumber="78">Miboman</location>
, 19851986 (EBD 20931, EBD 20789)
</materialsCitation>
,
<materialsCitation id="3B033CA3FFEFFFDEFAF5F8D6FDBEE5BB" ID-GBIF-Occurrence="3962324446" accessionNumber="EBD 27734, EBD 27774" collectingDate="1987-12" country="Equatorial Guinea" pageId="77" pageNumber="78" specimenCount="1">
<date id="FFD5103EFFEFFFDEFAF5F8D6FF7AE5BB" pageId="77" pageNumber="78" value="1987-12">
<collectingDate id="EF91E9D6FFEFFFDEFAF5F8D6FF7AE5BB" pageId="77" pageNumber="78" value="1987-12">December 1987</collectingDate>
</date>
(EBD 27734, EBD 27774)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFEFFFDEFDC4F8F2FA96E5BB" ID-GBIF-Occurrence="3962324416" accessionNumber="EBD 27536" box="[536,1315,1902,1929]" collectingDate="1987-08" collectionCode="EBD" collectorName="Miboman" country="Equatorial Guinea" location="Movo" municipality="Niefang" pageId="77" pageNumber="78" specimenCount="1">
<collectorName id="269E5328FFEFFFDEFDC4F8F2FD30E5BB" box="[536,645,1903,1929]" pageId="77" pageNumber="78">Miboman</collectorName>
(road from
<location id="8EB46025FFEFFFDEFCCCF8F2FCE6E5BB" LSID="urn:lsid:plazi:treatment:03C287E8FFEFFFDEFF4BFB40FB48E5C7:8EB46025FFEFFFDEFCCCF8F2FCE6E5BB" box="[784,851,1903,1929]" country="Equatorial Guinea" municipality="Niefang" name="Movo" pageId="77" pageNumber="78">Movo</location>
to
<collectingMunicipality id="6BB0AC84FFEFFFDEFCA6F8F2FC6CE5BB" box="[890,985,1903,1929]" pageId="77" pageNumber="78">Niefang</collectingMunicipality>
),
<date id="FFD5103EFFEFFFDEFC30F8F2FBC8E5BB" box="[1004,1149,1903,1929]" pageId="77" pageNumber="78" value="1987-08">
<collectingDate id="EF91E9D6FFEFFFDEFC30F8F2FBC8E5BB" box="[1004,1149,1903,1929]" pageId="77" pageNumber="78" value="1987-08">August 1987</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFEFFFDEFB51F8F2FB72E5BB" box="[1165,1223,1903,1929]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="77" pageNumber="78">EBD</collectionCode>
27536)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFEFFFDEFAF3F8F2FCE7E59F" ID-GBIF-Occurrence="3962324735" accessionNumber="EBD 20791" collectingDate="1985-07-29" collectionCode="EBD" collectorName="Miboman &amp; Km" country="Equatorial Guinea" location="Bata" municipality="Movo" pageId="77" pageNumber="78" specimenCount="1">
<collectorName id="269E5328FFEFFFDEFAF3F8F2FA29E5BB" box="[1327,1436,1903,1929]" pageId="77" pageNumber="78">Miboman</collectorName>
(
<collectorName id="269E5328FFEFFFDEFF43F80EFF7EE59F" box="[159,203,1939,1965]" pageId="77" pageNumber="78">Km</collectorName>
27, road from
<location id="8EB46025FFEFFFDEFEACF80EFE16E59F" LSID="urn:lsid:plazi:treatment:03C287E8FFEFFFDEFF4BFB40FB48E5C7:8EB46025FFEFFFDEFEACF80EFE16E59F" box="[368,419,1939,1965]" country="Equatorial Guinea" municipality="Movo" name="Bata" pageId="77" pageNumber="78">Bata</location>
to
<collectingMunicipality id="6BB0AC84FFEFFFDEFE1BF80EFDBBE59F" box="[455,526,1939,1965]" pageId="77" pageNumber="78">Movo</collectingMunicipality>
),
<date id="FFD5103EFFEFFFDEFDFDF80EFD04E59F" box="[545,689,1938,1965]" pageId="77" pageNumber="78" value="1985-07-29">
<collectingDate id="EF91E9D6FFEFFFDEFDFDF80EFD04E59F" box="[545,689,1938,1965]" pageId="77" pageNumber="78" value="1985-07-29">29 July 1985</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFEFFFDEFD63F80EFD4CE59F" box="[703,761,1939,1965]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="77" pageNumber="78">EBD</collectionCode>
20791)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFEFFFDEFC81F80EFE80E5E3" ID-GBIF-Occurrence="3962324805" accessionNumber="EBD 25021" collectingDate="1987-02-25" collectionCode="EBD" collectorName="Miboman" country="Equatorial Guinea" location="Niefang" pageId="77" pageNumber="78" specimenCount="1">
<collectorName id="269E5328FFEFFFDEFC81F80EFC7FE59F" box="[861,970,1939,1965]" pageId="77" pageNumber="78">Miboman</collectorName>
(
<quantity id="4C939B1BFFEFFFDEFC05F80EFB94E59F" box="[985,1057,1939,1965]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="77" pageNumber="78" unit="km" value="30.0">30 km</quantity>
from
<location id="8EB46025FFEFFFDEFBB9F80EFB71E59F" LSID="urn:lsid:plazi:treatment:03C287E8FFEFFFDEFF4BFB40FB48E5C7:8EB46025FFEFFFDEFBB9F80EFB71E59F" box="[1125,1220,1939,1965]" country="Equatorial Guinea" name="Niefang" pageId="77" pageNumber="78">Niefang</location>
),
<date id="FFD5103EFFEFFFDEFB0AF80FFA29E59F" box="[1238,1436,1938,1965]" pageId="77" pageNumber="78" value="1987-02-25">
<collectingDate id="EF91E9D6FFEFFFDEFB0AF80FFA29E59F" box="[1238,1436,1938,1965]" pageId="77" pageNumber="78" value="1987-02-25">25 February 1987</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFEFFFDEFF43F82AFF6CE5E3" box="[159,217,1975,2001]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="77" pageNumber="78">EBD</collectionCode>
25021)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFEFFFDEFE9FF82AFDA0E5C7" ID-GBIF-Occurrence="3962324642" accessionNumber="EBD 27534" collectingDate="1987-07-28" collectionCode="EBD" collectorName="Bata" country="Equatorial Guinea" county="Bata" location="Colegio La Salle" municipality="San Joaquin de Ndyiacon" pageId="77" pageNumber="78" specimenCount="1">
<location id="8EB46025FFEFFFDEFE9FF82AFDBDE5E3" LSID="urn:lsid:plazi:treatment:03C287E8FFEFFFDEFF4BFB40FB48E5C7:8EB46025FFEFFFDEFE9FF82AFDBDE5E3" box="[323,520,1975,2001]" country="Equatorial Guinea" county="Bata" municipality="San Joaquin de Ndyiacon" name="Colegio La Salle" pageId="77" pageNumber="78">Colegio La Salle</location>
,
<collectorName id="269E5328FFEFFFDEFDCAF82AFDF9E5E3" box="[534,588,1975,2001]" pageId="77" pageNumber="78">Bata</collectorName>
, 1967 (B2859, without
<collectionCode id="ED7AAE3BFFEFFFDEFCBCF82AFC2DE5E3" box="[864,920,1975,2001]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="77" pageNumber="78">EBD</collectionCode>
number);
<collectingMunicipality id="6BB0AC84FFEFFFDEFBCEF82AFA8DE5E3" box="[1042,1336,1975,2001]" pageId="77" pageNumber="78">San Joaquín de Ndyiacon</collectingMunicipality>
(road to
<collectingCounty id="62B54E72FFEFFFDEFF4BF846FF7BE5C7" box="[151,206,2011,2037]" pageId="77" pageNumber="78">Bata</collectingCounty>
),
<date id="FFD5103EFFEFFFDEFF3EF846FEC6E5C7" box="[226,371,2011,2037]" pageId="77" pageNumber="78" value="1987-07-28">
<collectingDate id="EF91E9D6FFEFFFDEFF3EF846FEC6E5C7" box="[226,371,2011,2037]" pageId="77" pageNumber="78" value="1987-07-28">28 July 1987</collectingDate>
</date>
(
<collectionCode id="ED7AAE3BFFEFFFDEFE5DF846FE0EE5C7" box="[385,443,2011,2037]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="77" pageNumber="78">EBD</collectionCode>
27534)
</materialsCitation>
;
<materialsCitation id="3B033CA3FFEFFFDEFDFCF846FB4FE5C7" ID-GBIF-Occurrence="3962324803" box="[544,1274,2010,2037]" collectionCode="EBD" country="Equatorial Guinea" location="San Joaquin de Ndyiacom" pageId="77" pageNumber="78" specimenCount="1">
<location id="8EB46025FFEFFFDEFDFCF846FCFDE5C7" LSID="urn:lsid:plazi:treatment:03C287E8FFEFFFDEFF4BFB40FB48E5C7:8EB46025FFEFFFDEFDFCF846FCFDE5C7" box="[544,840,2011,2037]" country="Equatorial Guinea" name="San Joaquin de Ndyiacom" pageId="77" pageNumber="78">San Joaquín de Ndyiacom</location>
, 1990 (B 9580, without
<collectionCode id="ED7AAE3BFFEFFFDEFB84F846FB25E5C7" box="[1112,1168,2011,2037]" country="Spain" httpUri="http://grbio.org/cool/ah5f-9iii" name="Estacion Biologica de Donana" pageId="77" pageNumber="78">EBD</collectionCode>
number)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>