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<document id="4D809ACCAC3821FE1335799E143798D2" ID-DOI="10.5281/zenodo.13245522" ID-ISSN="2345-7600" ID-Zenodo-Dep="13245522" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722970765904" checkinUser="felipe" docAuthor="Shokita, Yixiong Cai Shigemitsu" docDate="2006" docId="03EC878C550FCD522AFB69D4FE1E2F88" docLanguage="en" docName="RafflesBZool.54.2.245-270.pdf" docOrigin="Raffles Bulletin of Zoology 54 (2)" docStyle="DocumentStyle:C6443723295B2E4C6F22B9B23F08089C.5:RafflesBZool.2006-2013.journal_article" docStyleId="C6443723295B2E4C6F22B9B23F08089C" docStyleName="RafflesBZool.2006-2013.journal_article" docStyleVersion="5" docTitle="Caridina palawanensis Shokita, 2006, new species" docType="treatment" docVersion="2" lastPageNumber="259" masterDocId="FFD5FFF45504CD5C29586B35FFAE2970" masterDocTitle="Report On A Collection Of Freshwater Shrimps (Crustacea: Decapoda: Caridea) From The Philippines, With Descriptions Of Four New Species" masterLastPageNumber="270" masterPageNumber="245" pageNumber="256" updateTime="1722977171740" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="81832E1B610090C06733AF3BBCA263C6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="06243A34155517B49244B473E9876D00">
<mods:title id="EA1DFC2BA4298E7FA084BE311078CF3F">Report On A Collection Of Freshwater Shrimps (Crustacea: Decapoda: Caridea) From The Philippines, With Descriptions Of Four New Species</mods:title>
</mods:titleInfo>
<mods:name id="E9B52FA1B3C7C038CE8DA4942CFBD27D" type="personal">
<mods:role id="CFF526624EA000A84666124220527CE0">
<mods:roleTerm id="AE7950609051D56D67A7B2AF8587316A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5E288D017E7C4EF9A0501CDBA9737DFD">Shokita, Yixiong Cai Shigemitsu</mods:namePart>
</mods:name>
<mods:typeOfResource id="F37411E7B76A72A857FF9F61CCD6137F">text</mods:typeOfResource>
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<mods:title id="610E501847DDAC74FFCB5806E3B3C72D">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="65CAD707DC927370A1D192BD634AFACD">
<mods:date id="949FC3DAFC143F3029E90D29BC9D2CBD">2006</mods:date>
<mods:detail id="76BE125FF112F8C03E68561310E47750" type="pubDate">
<mods:number id="ACCD75B8EC8DA8A95B25E0A245A5B3D1">2006-08-31</mods:number>
</mods:detail>
<mods:detail id="29A636D324259BDE9AB6FCFDAD4EF0AE" type="volume">
<mods:number id="2AF3E0249C5CB90E608A6B99C1169BB9">54</mods:number>
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<mods:detail id="C4BE9B2FB72A740FB827D06BE8C3AFDE" type="issue">
<mods:number id="A5454FE5897D0A5BD1DEEA8A9B0C06D7">2</mods:number>
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<mods:classification id="81D3318D2D378993F76FD453A1FDB2F3">journal article</mods:classification>
<mods:identifier id="14F67F2B2D24647C5B4EE4D2A9D7B971" type="DOI">10.5281/zenodo.13245522</mods:identifier>
<mods:identifier id="837A884FC8C5E189946C5AEA7ED67B07" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="E5CBE87E64CA84E342157F2708C86C8B" type="Zenodo-Dep">13245522</mods:identifier>
</mods:mods>
<treatment id="03EC878C550FCD522AFB69D4FE1E2F88" LSID="urn:lsid:plazi:treatment:03EC878C550FCD522AFB69D4FE1E2F88" httpUri="http://treatment.plazi.org/id/03EC878C550FCD522AFB69D4FE1E2F88" lastPageId="14" lastPageNumber="259" pageId="11" pageNumber="256">
<subSubSection id="C35F6511550FCD572AFB69D4FA9B2B88" box="[931,1333,736,761]" pageId="11" pageNumber="256" type="nomenclature">
<paragraph id="8BFA369A550FCD572AFB69D4FA9B2B88" blockId="11.[931,1333,736,792]" box="[931,1333,736,761]" pageId="11" pageNumber="256">
<heading id="D0B281F6550FCD572AFB69D4FA9B2B88" bold="true" box="[931,1333,736,761]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="256" reason="2">
<emphasis id="B931EA88550FCD572AFB69D4FA9B2B88" bold="true" box="[931,1333,736,761]" pageId="11" pageNumber="256">
<emphasis id="B931EA88550FCD572AFB69D4FB062B89" bold="true" box="[931,1192,737,761]" italics="true" pageId="11" pageNumber="256">
<taxonomicName id="4C454D19550FCD572AFB69D4FB0C2B89" ID-CoL="8VKCM" authorityName="Shokita" authorityYear="2006" box="[931,1186,737,761]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="256" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">Caridina palawanensis</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A20257F3550FCD572DE869D5FA9B2B88" box="[1200,1333,736,760]" pageId="11" pageNumber="256" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35F6511550FCD572D4E6834FB6F2A68" box="[1046,1217,769,792]" pageId="11" pageNumber="256" type="description">
<paragraph id="8BFA369A550FCD572D4E6834FB6F2A68" blockId="11.[931,1333,736,792]" box="[1046,1217,769,792]" pageId="11" pageNumber="256">
(
<figureCitation id="137E2A1F550FCD572D466834FBC42A68" box="[1054,1130,769,792]" captionStart="Fig" captionStartId="12.[136,171,2006,2027]" captionTargetBox="[244,1371,222,1931]" captionTargetId="figure-103@12.[211,1422,191,1960]" captionTargetPageId="12" captionText="Fig. 5. Caridina palawanensis, new species, ovig. females, cl 4.1 mm, paratype, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island Philippines:A, cephalothorax and cephalic appendages, lateral view; B, telson;C, distal portion of telson;D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scale bars: A, B, D = 1 mm; E-G, I, K = 0.5 mm; C = 0.2 mm; H, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13245534" httpUri="https://zenodo.org/record/13245534/files/figure.png" pageId="11" pageNumber="256">Figs. 5</figureCitation>
,
<figureCitation id="137E2A1F550FCD572D2F6834FB172A68" box="[1143,1209,769,792]" captionStart="Fig" captionStartId="13.[113,148,1950,1971]" captionTargetBox="[244,1310,216,1897]" captionTargetId="figure-129@13.[192,1396,180,1922]" captionTargetPageId="13" captionText="Fig. 6. Caridina palawanensis, new species, male, cl 2.8 mm, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island, Philippines: A, male second pleopod; B, male first pleopod. Caridina sumatrensis, male, cl 3.2 mm, ZRC, Santacruz River, Palawan Island, Philippines: C, cephalothorax and cephalic appendages, lateral view; D, first pereiopod; E, second pereiopod; F, third pereiopod; G, preanal carina. Potamalpheops miyai, male, cl 3.0 mm, ZRC, Surigao mangrove, Mindanao Island, Philippines:H, cephalothorax and cephalic appendages, lateral view; I, second pereiopod; J, propodus and dactylus of third pereiopod; K, fifth pereiopod; L, uropodal diaeresis. Scale bars: A, B = 0.2 mm, C, H = 1 mm, D-G, I-L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13245536" httpUri="https://zenodo.org/record/13245536/files/figure.png" pageId="11" pageNumber="256">6A, B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35F6511550FCD572A746875FA892D72" pageId="11" pageNumber="256" type="materials_examined">
<paragraph id="8BFA369A550FCD572A746875FBC92AFF" blockId="11.[812,1452,832,911]" pageId="11" pageNumber="256">
<emphasis id="B931EA88550FCD572A746875FC532A25" bold="true" box="[812,1021,832,853]" pageId="11" pageNumber="256">
<emphasis id="B931EA88550FCD572A746875FC432A25" bold="true" box="[812,1005,832,853]" italics="true" pageId="11" pageNumber="256">Material examined.</emphasis>
–
</emphasis>
<materialsCitation id="3B2D3CC7550FCD572D5A6874FBC92AFF" collectingDate="1985-08-06" collectionCode="NSMT" country="Philippines" location="Panitian River" municipality="Island" pageId="11" pageNumber="256" specimenCount="1" stateProvince="Palawan" typeStatus="holotype">
<typeStatus id="54FE8838550FCD572D5A6874FBF12A26" box="[1026,1119,833,854]" pageId="11" pageNumber="256" type="holotype">Holotype</typeStatus>
: 1 ovig. female, cl
<quantity id="4CBD9B7F550FCD572C486874FAF42A26" box="[1296,1370,833,854]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="11" pageNumber="256" unit="mm" value="4.1">4.1 mm</quantity>
,
<collectionCode id="ED54AE5F550FCD572C3B6874FA062A26" box="[1379,1448,833,854]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="11" pageNumber="256">NSMT</collectionCode>
,
<location id="8E9A6041550FCD572A746868FC172A02" LSID="urn:lsid:plazi:treatment:03EC878C550FCD522AFB69D4FE1E2F88:8E9A6041550FCD572A746868FC172A02" box="[812,953,861,882]" country="Philippines" municipality="Island" name="Panitian River" pageId="11" pageNumber="256" stateProvince="Palawan">Panitian River</location>
, upper stream, ca
<quantity id="4CBD9B7F550FCD572D3E6868FB0D2A02" box="[1126,1187,861,882]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="256" unit="km" value="30.0">30 km</quantity>
from river mouth,
<collectingRegion id="4981F878550FCD572C026868FA052A02" box="[1370,1451,861,882]" country="Philippines" name="Palawan" pageId="11" pageNumber="256">Palawan</collectingRegion>
<collectingMunicipality id="6B9EACE0550FCD572A74684FFCC72AFF" box="[812,873,890,911]" pageId="11" pageNumber="256">Island</collectingMunicipality>
,
<collectingCountry id="F352760A550FCD572A2C684FFC4D2AFF" box="[884,995,890,911]" name="Philippines" pageId="11" pageNumber="256">Philippines</collectingCountry>
,
<date id="FFFB105A550FCD572AB5684FFBCD2AFF" box="[1005,1123,890,911]" pageId="11" pageNumber="256" value="1985-08-06">
<collectingDate id="EFBFE9B2550FCD572AB5684FFBCD2AFF" box="[1005,1123,890,911]" pageId="11" pageNumber="256" value="1985-08-06">6 Aug.1985</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BFA369A550FCD572A746881FA892D72" blockId="11.[812,1452,948,1026]" pageId="11" pageNumber="256">
<materialsCitation id="3B2D3CC7550FCD572A746881FCF32A96" collectionCode="ZRC" pageId="11" pageNumber="256" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="54FE8838550FCD572A746881FC232AB9" box="[812,909,948,969]" pageId="11" pageNumber="256" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D43FD13550FCD572ACE6881FC4C2AB9" box="[918,994,948,969]" count="2" pageId="11" pageNumber="256" type="male">2 males</specimenCount>
, cl
<quantity id="4CBD9B7F550FCD572D5C6881FBD82AB9" box="[1028,1142,948,969]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.9" metricValueMin="2.6" pageId="11" pageNumber="256" unit="mm" value="2.75" valueMax="2.9" valueMin="2.6">2.6-2.9 mm</quantity>
, 2 ovig. females, cl
<quantity id="4CBD9B7F550FCD572C6E6881FA062AB9" box="[1334,1448,948,969]" metricMagnitude="-3" metricUnit="m" metricValue="3.9499999999999997" metricValueMax="4.1" metricValueMin="3.8" pageId="11" pageNumber="256" unit="mm" value="3.9499999999999997" valueMax="4.1" valueMin="3.8">3.8-4.1 mm</quantity>
,
<collectionCode id="ED54AE5F550FCD572A7468E4FCF32A96" box="[812,861,977,998]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="11" pageNumber="256">ZRC</collectionCode>
</materialsCitation>
,
<specimenCount id="9D43FD13550FCD572A3168E4FC162A96" box="[873,952,977,998]" count="3" pageId="11" pageNumber="256" type="male">3 males</specimenCount>
, cl
<quantity id="4CBD9B7F550FCD572A8668E4FBFD2A96" box="[990,1107,977,998]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="11" pageNumber="256" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.2-2.8 mm</quantity>
, 2 ovig. females, cl
<quantity id="4CBD9B7F550FCD572C4768E4FA3D2A96" box="[1311,1427,977,998]" metricMagnitude="-3" metricUnit="m" metricValue="4.15" metricValueMax="4.2" metricValueMin="4.1" pageId="11" pageNumber="256" unit="mm" value="4.15" valueMax="4.2" valueMin="4.1">4.1-4.2 mm</quantity>
,
<materialsCitation id="3B2D3CC7550FCD572CC768E4FA8D2D72" collectingDate="1985-08-06" collectionCode="NSMT" country="Philippines" location="Panitian River" municipality="Island" pageId="11" pageNumber="256" specimenCount="2" specimenCount-female="2" stateProvince="Palawan" typeStatus="paratype">
<specimenCount id="9D43FD13550FCD572CC768E4FCD72D72" count="2" pageId="11" pageNumber="256" type="female">2 females</specimenCount>
, cl
<quantity id="4CBD9B7F550FCD572AC568D8FBBE2D72" box="[925,1040,1005,1026]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="11" pageNumber="256" unit="mm" value="2.7" valueMax="2.8" valueMin="2.6">2.6-2.8 mm</quantity>
, UR, data same as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35F6511550FCD522A746F1FFE9D2D88" lastPageId="14" lastPageNumber="259" pageId="11" pageNumber="256" type="description">
<paragraph id="8BFA369A550FCD572A746F1FFB652C71" blockId="11.[812,1452,1066,1281]" pageId="11" pageNumber="256">
<emphasis id="B931EA88550FCD572A746F1FFC682D32" bold="true" box="[812,966,1066,1090]" pageId="11" pageNumber="256">
<emphasis id="B931EA88550FCD572A746F1FFC1C2D32" bold="true" box="[812,946,1066,1090]" italics="true" pageId="11" pageNumber="256">Description.</emphasis>
–
</emphasis>
Rostrum straight (5A), reaching to or beyond end of antennualr peduncle, or slight beyond end of scaphocerite, armed dorsally with 21 to 29 teeth throughout the dorsal margin, including 2 to 3 posterior to postorbital margin; armed ventrally with 5 to 10 teeth on anterior half; antennal spine distinctly lower than sub-orbital angle. Pterygostomian angle sub-rectangular.
</paragraph>
<paragraph id="8BFA369A550FCD572A746E1FFB812F51" blockId="11.[812,1452,1322,1569]" pageId="11" pageNumber="256">
Third abdominal somite with moderately convex dorsal profile. Sixth abdominal somite 0.6 times as long as carapace, 1.8 times as long as fifth somite, slightly longer than telson. Telson (
<figureCitation id="137E2A1F550FCD572ADB6EBFFC5D2CD1" box="[899,1011,1418,1441]" captionStart="Fig" captionStartId="12.[136,171,2006,2027]" captionTargetBox="[244,1371,222,1931]" captionTargetId="figure-103@12.[211,1422,191,1960]" captionTargetPageId="12" captionText="Fig. 5. Caridina palawanensis, new species, ovig. females, cl 4.1 mm, paratype, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island Philippines:A, cephalothorax and cephalic appendages, lateral view; B, telson;C, distal portion of telson;D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scale bars: A, B, D = 1 mm; E-G, I, K = 0.5 mm; C = 0.2 mm; H, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13245534" httpUri="https://zenodo.org/record/13245534/files/figure.png" pageId="11" pageNumber="256">Fig. 5B, C</figureCitation>
) 3.4 times as long as wide, terminating in a small projection, with four pairs of dorsal spinules and one pair of dorsolateral spinules; lateral pair of distal spines slightly longer than intermediate pairs. Preanal carina small, lacking spine (
<figureCitation id="137E2A1F550FCD572A936D3FFB8D2F51" box="[971,1059,1546,1569]" captionStart="Fig" captionStartId="12.[136,171,2006,2027]" captionTargetBox="[244,1371,222,1931]" captionTargetId="figure-103@12.[211,1422,191,1960]" captionTargetPageId="12" captionText="Fig. 5. Caridina palawanensis, new species, ovig. females, cl 4.1 mm, paratype, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island Philippines:A, cephalothorax and cephalic appendages, lateral view; B, telson;C, distal portion of telson;D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scale bars: A, B, D = 1 mm; E-G, I, K = 0.5 mm; C = 0.2 mm; H, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13245534" httpUri="https://zenodo.org/record/13245534/files/figure.png" pageId="11" pageNumber="256">Fig. 5K</figureCitation>
).
</paragraph>
<paragraph id="8BFA369A550FCD572A746D7FFC132E31" blockId="11.[812,1451,1610,1857]" pageId="11" pageNumber="256">
Eyes well developed, small, anterior end reaching to 0.7 length of basal segment of antennular peduncle. Antennular peduncle 0.7 times as long as carapace; basal segment of antennular peduncle longer than sum of second and third segment length; second segment distinctly longer than third segment. Stylocerite reaching to 0.7 length of basal segment of antennular peduncle. Scaphocerite (
<figureCitation id="137E2A1F550FCD572D906C3FFAB32E51" box="[1224,1309,1802,1825]" captionStart="Fig" captionStartId="12.[136,171,2006,2027]" captionTargetBox="[244,1371,222,1931]" captionTargetId="figure-103@12.[211,1422,191,1960]" captionTargetPageId="12" captionText="Fig. 5. Caridina palawanensis, new species, ovig. females, cl 4.1 mm, paratype, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island Philippines:A, cephalothorax and cephalic appendages, lateral view; B, telson;C, distal portion of telson;D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scale bars: A, B, D = 1 mm; E-G, I, K = 0.5 mm; C = 0.2 mm; H, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13245534" httpUri="https://zenodo.org/record/13245534/files/figure.png" pageId="11" pageNumber="256">Fig. 5D</figureCitation>
) 3.7 times as long as wide.
</paragraph>
<paragraph id="8BFA369A550FCD572A746C5FFBBD2151" blockId="11.[812,1452,1898,2081]" pageId="11" pageNumber="256">
Mouthparts similar to that of
<taxonomicName id="4C454D19550FCD572D0E6C5EFAB02EF2" authorityName="Shokita" authorityYear="2006" box="[1110,1310,1899,1922]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="256" phylum="Arthropoda" rank="species" species="cebuensis" status="sp. nov.">
<emphasis id="B931EA88550FCD572D0E6C5EFAB02EF2" box="[1110,1310,1899,1922]" italics="true" pageId="11" pageNumber="256">Caridina cebuensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20257F3550FCD572C716C5FFA062EF1" box="[1321,1448,1898,1921]" pageId="11" pageNumber="256" rank="species">new species</taxonomicNameLabel>
. Palp of first maxilliped broadly triangular, ending in a fingerlike projection. Second maxilliped typical of the genus. Third maxilliped reaching beyond end of second segment of antennular peduncle, with ultimate segment longer than penultimate segment.
</paragraph>
<caption id="DF3A66125508CD5029D06CE3FAB12131" ID-DOI="http://doi.org/10.5281/zenodo.13245534" ID-Zenodo-Dep="13245534" httpUri="https://zenodo.org/record/13245534/files/figure.png" pageId="12" pageNumber="257" startId="12.[136,171,2006,2027]" targetBox="[244,1371,222,1931]" targetPageId="12" targetType="figure">
<paragraph id="8BFA369A5508CD5029D06CE3FAB12131" blockId="12.[136,1474,2006,2113]" pageId="12" pageNumber="257">
Fig. 5.
<taxonomicName id="4C454D195508CD5029966CE3FE012E9B" authorityName="Shokita" authorityYear="2006" box="[206,431,2006,2027]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="257" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B931EA885508CD5029966CE3FE012E9B" box="[206,431,2006,2027]" italics="true" pageId="12" pageNumber="257">Caridina palawanensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20257F35508CD5028E46CE3FD9D2E9B" box="[444,563,2006,2027]" pageId="12" pageNumber="257" rank="species">new species</taxonomicNameLabel>
, ovig. females, cl 4.1 mm, paratype, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island Philippines:A, cephalothorax and cephalic appendages, lateral view; B, telson; C, distal portion of telson; D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scale bars: A, B, D = 1 mm; E-G, I, K = 0.5 mm; C = 0.2 mm; H, J = 0.1 mm.
</paragraph>
</caption>
<caption id="DF3A66125509CD5129296CABFD352133" ID-DOI="http://doi.org/10.5281/zenodo.13245536" ID-Zenodo-Dep="13245536" httpUri="https://zenodo.org/record/13245536/files/figure.png" pageId="13" pageNumber="258" startId="13.[113,148,1950,1971]" targetBox="[244,1310,216,1897]" targetPageId="13" targetType="figure">
<paragraph id="8BFA369A5509CD5129296CABFD352133" blockId="13.[113,1451,1950,2115]" pageId="13" pageNumber="258">
Fig. 6.
<taxonomicName id="4C454D195509CD5129E16CABFE352EC3" authorityName="Shokita" authorityYear="2006" box="[185,411,1950,1971]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="258" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B931EA885509CD5129E16CABFE352EC3" box="[185,411,1950,1971]" italics="true" pageId="13" pageNumber="258">Caridina palawanensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20257F35509CD5128F06CABFD8F2EC3" box="[424,545,1950,1971]" pageId="13" pageNumber="258" rank="species">new species</taxonomicNameLabel>
, male, cl 2.8 mm, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island, Philippines: A, male second pleopod; B, male first pleopod.
<taxonomicName id="4C454D195509CD512BA16C8EFC662EA0" authorityName="De Man" authorityYear="1892" box="[761,968,1979,2000]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="258" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="B931EA885509CD512BA16C8EFC662EA0" box="[761,968,1979,2000]" italics="true" pageId="13" pageNumber="258">Caridina sumatrensis</emphasis>
</taxonomicName>
, male, cl 3.2 mm, ZRC, Santacruz River, Palawan Island, Philippines: C, cephalothorax and cephalic appendages, lateral view; D, first pereiopod; E, second pereiopod; F, third pereiopod; G, preanal carina.
<taxonomicName id="4C454D195509CD5128596CC0FE61217A" authorityName="Yeo &amp; Ng" authorityYear="1997" box="[257,463,2037,2058]" class="Malacostraca" family="Alpheidae" genus="Potamalpheops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="258" phylum="Arthropoda" rank="species" species="miyai">
<emphasis id="B931EA885509CD5128596CC0FE61217A" box="[257,463,2037,2058]" italics="true" pageId="13" pageNumber="258">Potamalpheops miyai</emphasis>
</taxonomicName>
, male, cl 3.0 mm, ZRC, Surigao mangrove, Mindanao Island, Philippines: H, cephalothorax and cephalic appendages, lateral view; I, second pereiopod; J, propodus and dactylus of third pereiopod; K, fifth pereiopod; L, uropodal diaeresis. Scale bars: A, B = 0.2 mm, C, H = 1 mm, D-G, I-L = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BFA369A550ACD5229D06BF4FDC52A68" blockId="14.[136,775,193,792]" pageId="14" pageNumber="259">
Epipods on first four pereiopods. First pereiopod (
<figureCitation id="137E2A1F550ACD522BF46BF4FD5029A8" box="[684,766,193,216]" captionStart="Fig" captionStartId="12.[136,171,2006,2027]" captionTargetBox="[244,1371,222,1931]" captionTargetId="figure-103@12.[211,1422,191,1960]" captionTargetPageId="12" captionText="Fig. 5. Caridina palawanensis, new species, ovig. females, cl 4.1 mm, paratype, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island Philippines:A, cephalothorax and cephalic appendages, lateral view; B, telson;C, distal portion of telson;D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scale bars: A, B, D = 1 mm; E-G, I, K = 0.5 mm; C = 0.2 mm; H, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13245534" httpUri="https://zenodo.org/record/13245534/files/figure.png" pageId="14" pageNumber="259">Fig. 5E</figureCitation>
) reaching to anterior margin of the eye stalk; ischium shorter than merus; merus 2.7 times as long as broad, slightly shorter than carpus; carpus excavated anteriorly, shorter than chela, 1.8 times as long as high; chela 2.0 times as long as broad; fingers distinctly longer than palm. Second pereiopod (
<figureCitation id="137E2A1F550ACD522B836A54FF0A28E8" captionStart="Fig" captionStartId="12.[136,171,2006,2027]" captionTargetBox="[244,1371,222,1931]" captionTargetId="figure-103@12.[211,1422,191,1960]" captionTargetPageId="12" captionText="Fig. 5. Caridina palawanensis, new species, ovig. females, cl 4.1 mm, paratype, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island Philippines:A, cephalothorax and cephalic appendages, lateral view; B, telson;C, distal portion of telson;D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scale bars: A, B, D = 1 mm; E-G, I, K = 0.5 mm; C = 0.2 mm; H, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13245534" httpUri="https://zenodo.org/record/13245534/files/figure.png" pageId="14" pageNumber="259">Fig. 5F</figureCitation>
) reaching beyond second segment of antennular peduncle; ischium shorter than merus; merus shorter than carpus, 5.3 times as long as broad; carpus 1.3 times as long as chela, 5.5 times as long as high; chela 2.6 times as long as broad; fingers 1.6 times as long as palm. Third pereiopod (
<figureCitation id="137E2A1F550ACD522BDA6934FD512B68" box="[642,767,513,536]" captionStart="Fig" captionStartId="12.[136,171,2006,2027]" captionTargetBox="[244,1371,222,1931]" captionTargetId="figure-103@12.[211,1422,191,1960]" captionTargetPageId="12" captionText="Fig. 5. Caridina palawanensis, new species, ovig. females, cl 4.1 mm, paratype, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island Philippines:A, cephalothorax and cephalic appendages, lateral view; B, telson;C, distal portion of telson;D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scale bars: A, B, D = 1 mm; E-G, I, K = 0.5 mm; C = 0.2 mm; H, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13245534" httpUri="https://zenodo.org/record/13245534/files/figure.png" pageId="14" pageNumber="259">Fig. 5G, H</figureCitation>
) reaching to end of antennular peduncle, propodus distinctly shorter than merus, 13 times as long as broad, 4.2 times as long as dactylus; dactylus 3.0 times as long as wide (spines included), with 6 accessory spines on its flexor margin. Fifth pereiopod (
<figureCitation id="137E2A1F550ACD52285A6994FECA2BC8" box="[258,356,673,696]" captionStart="Fig" captionStartId="12.[136,171,2006,2027]" captionTargetBox="[244,1371,222,1931]" captionTargetId="figure-103@12.[211,1422,191,1960]" captionTargetPageId="12" captionText="Fig. 5. Caridina palawanensis, new species, ovig. females, cl 4.1 mm, paratype, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island Philippines:A, cephalothorax and cephalic appendages, lateral view; B, telson;C, distal portion of telson;D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scale bars: A, B, D = 1 mm; E-G, I, K = 0.5 mm; C = 0.2 mm; H, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13245534" httpUri="https://zenodo.org/record/13245534/files/figure.png" pageId="14" pageNumber="259">Fig. 5I, J</figureCitation>
) reaching to end of second segment of antennular peduncle, propodus 14 times as long as broad, 3.0- 3.6 times as long as dactylus; dactylus 3.7 times as long as broad, with 49 spinules on its flexor margin.
</paragraph>
<paragraph id="8BFA369A550ACD5229D06874FF7C2AA8" blockId="14.[136,774,833,984]" pageId="14" pageNumber="259">
Endopod of male first pleopod (
<figureCitation id="137E2A1F550ACD5228976874FDB02A28" box="[463,542,833,856]" captionStart="Fig" captionStartId="13.[113,148,1950,1971]" captionTargetBox="[244,1310,216,1897]" captionTargetId="figure-129@13.[192,1396,180,1922]" captionTargetPageId="13" captionText="Fig. 6. Caridina palawanensis, new species, male, cl 2.8 mm, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island, Philippines: A, male second pleopod; B, male first pleopod. Caridina sumatrensis, male, cl 3.2 mm, ZRC, Santacruz River, Palawan Island, Philippines: C, cephalothorax and cephalic appendages, lateral view; D, first pereiopod; E, second pereiopod; F, third pereiopod; G, preanal carina. Potamalpheops miyai, male, cl 3.0 mm, ZRC, Surigao mangrove, Mindanao Island, Philippines:H, cephalothorax and cephalic appendages, lateral view; I, second pereiopod; J, propodus and dactylus of third pereiopod; K, fifth pereiopod; L, uropodal diaeresis. Scale bars: A, B = 0.2 mm, C, H = 1 mm, D-G, I-L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13245536" httpUri="https://zenodo.org/record/13245536/files/figure.png" pageId="14" pageNumber="259">Fig. 6B</figureCitation>
) triangular, one fourth length of exopod, no appendix interna. Appendix masculina of male second pleopod (
<figureCitation id="137E2A1F550ACD5228C768B4FE5B2AE8" box="[415,501,897,920]" captionStart="Fig" captionStartId="13.[113,148,1950,1971]" captionTargetBox="[244,1310,216,1897]" captionTargetId="figure-129@13.[192,1396,180,1922]" captionTargetPageId="13" captionText="Fig. 6. Caridina palawanensis, new species, male, cl 2.8 mm, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island, Philippines: A, male second pleopod; B, male first pleopod. Caridina sumatrensis, male, cl 3.2 mm, ZRC, Santacruz River, Palawan Island, Philippines: C, cephalothorax and cephalic appendages, lateral view; D, first pereiopod; E, second pereiopod; F, third pereiopod; G, preanal carina. Potamalpheops miyai, male, cl 3.0 mm, ZRC, Surigao mangrove, Mindanao Island, Philippines:H, cephalothorax and cephalic appendages, lateral view; I, second pereiopod; J, propodus and dactylus of third pereiopod; K, fifth pereiopod; L, uropodal diaeresis. Scale bars: A, B = 0.2 mm, C, H = 1 mm, D-G, I-L = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13245536" httpUri="https://zenodo.org/record/13245536/files/figure.png" pageId="14" pageNumber="259">Fig. 6A</figureCitation>
) half of endopod length, with appendix interna reaching half of appendix masculina length.
</paragraph>
<paragraph id="8BFA369A550ACD5229D06F34FD122D68" blockId="14.[136,700,1025,1048]" box="[136,700,1025,1048]" pageId="14" pageNumber="259">
Uropodal diaeresis (
<figureCitation id="137E2A1F550ACD52283A6F34FE1D2D68" box="[354,435,1025,1048]" captionStart="Fig" captionStartId="12.[136,171,2006,2027]" captionTargetBox="[244,1371,222,1931]" captionTargetId="figure-103@12.[211,1422,191,1960]" captionTargetPageId="12" captionText="Fig. 5. Caridina palawanensis, new species, ovig. females, cl 4.1 mm, paratype, ZRC, Panitian River, upper stream, ca 30 km from river mouth, Palawan Island Philippines:A, cephalothorax and cephalic appendages, lateral view; B, telson;C, distal portion of telson;D, scaphocerite; E, first pereiopod; F, second pereiopod; G, third pereiopod; H, dactylus of third pereiopod; I, fifth pereiopod; J, dactylus of fifth pereiopod; K, preanal carina; L, uropodal diaeresis. Scale bars: A, B, D = 1 mm; E-G, I, K = 0.5 mm; C = 0.2 mm; H, J = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13245534" httpUri="https://zenodo.org/record/13245534/files/figure.png" pageId="14" pageNumber="259">Fig. 5L</figureCitation>
) with 17 movable teeth.
</paragraph>
<paragraph id="8BFA369A550ACD5229D06F74FEA72D08" blockId="14.[136,774,1089,1144]" pageId="14" pageNumber="259">
Ovigerous females with egg sized 0.78-0.82x
<quantity id="4CBD9B7F550ACD522B376F74FCA82D28" box="[623,774,1089,1112]" metricMagnitude="-4" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="14" pageNumber="259" unit="mm" value="0.425" valueMax="0.45" valueMin="0.4">0.40-0.45 mm</quantity>
in diameter.
</paragraph>
<paragraph id="8BFA369A550ACD5229D06F94FE9D2D88" blockId="14.[136,775,1184,1272]" pageId="14" pageNumber="259">
<emphasis id="B931EA88550ACD5229D06F94FE462DC8" box="[136,488,1184,1209]" italics="true" pageId="14" pageNumber="259">
<emphasis id="B931EA88550ACD5229D06F94FF5F2DC8" bold="true" box="[136,241,1184,1209]" italics="true" pageId="14" pageNumber="259">Habitat. –</emphasis>
<taxonomicName id="4C454D19550ACD5229AE6F94FE462DC8" authorityName="Shokita" authorityYear="2006" box="[246,488,1185,1208]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="259" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">Caridina palawanensis</taxonomicName>
</emphasis>
,
<taxonomicNameLabel id="A20257F3550ACD5228AB6F94FDDC2DC8" box="[499,626,1185,1208]" pageId="14" pageNumber="259" rank="species">new species</taxonomicNameLabel>
, was collected from Panitian River, upper stream, ca
<quantity id="4CBD9B7F550ACD522B536FF4FDE22DA8" box="[523,588,1217,1240]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="259" unit="km" value="30.0">30 km</quantity>
from river mouth,
<collectingRegion id="4981F878550ACD5229D06FD4FF4C2D88" box="[136,226,1249,1272]" country="Philippines" name="Palawan" pageId="14" pageNumber="259">Palawan</collectingRegion>
Island.
</paragraph>
</subSubSection>
<subSubSection id="C35F6511550ACD5229D06E14FE612C28" pageId="14" pageNumber="259" type="etymology">
<paragraph id="8BFA369A550ACD5229D06E14FE612C28" blockId="14.[136,775,1312,1368]" pageId="14" pageNumber="259">
<emphasis id="B931EA88550ACD5229D06E14FEBB2C48" bold="true" box="[136,277,1312,1337]" pageId="14" pageNumber="259">
<emphasis id="B931EA88550ACD5229D06E14FF522C49" bold="true" box="[136,252,1313,1337]" italics="true" pageId="14" pageNumber="259">Etymology</emphasis>
. –
</emphasis>
The new species is named after its
<typeStatus id="54FE8838550ACD522BDC6E14FD1C2C48" box="[644,690,1313,1336]" pageId="14" pageNumber="259">type</typeStatus>
locality –
<collectingRegion id="4981F878550ACD5229C56E74FF592C28" box="[157,247,1345,1368]" country="Philippines" name="Palawan" pageId="14" pageNumber="259">Palawan</collectingRegion>
Island,
<collectingCountry id="F352760A550ACD5228176E74FE652C28" box="[335,459,1345,1368]" name="Philippines" pageId="14" pageNumber="259">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35F6511550ACD5229D06EB4FEB42FC8" pageId="14" pageNumber="259" type="discussion">
<paragraph id="8BFA369A550ACD5229D06EB4FEB42FC8" blockId="14.[136,775,1408,1720]" pageId="14" pageNumber="259">
<emphasis id="B931EA88550ACD5229D06EB4FEA92CE8" bold="true" box="[136,263,1408,1433]" pageId="14" pageNumber="259">
<emphasis id="B931EA88550ACD5229D06EB4FF5E2CE9" bold="true" box="[136,240,1409,1433]" italics="true" pageId="14" pageNumber="259">Remarks.</emphasis>
–
</emphasis>
With respect to the rostrum, the egg size, the pereiopods,
<taxonomicName id="4C454D19550ACD5228576E94FDA02CC8" authorityName="Shokita" authorityYear="2006" box="[271,526,1441,1464]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="259" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B931EA88550ACD5228576E94FDA02CC8" box="[271,526,1441,1464]" italics="true" pageId="14" pageNumber="259">Caridina palawanensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A20257F3550ACD522B476E94FD092CC8" box="[543,679,1441,1464]" pageId="14" pageNumber="259" rank="species">new species</taxonomicNameLabel>
, is most similar to
<taxonomicName id="4C454D19550ACD5228516EF4FDDF2CA8" authority="Cai &amp; Ng, 2000" authorityName="Cai &amp; Ng" authorityYear="2000" box="[265,625,1473,1496]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="259" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B931EA88550ACD5228516EF4FE352CA8" box="[265,411,1473,1496]" italics="true" pageId="14" pageNumber="259">C. williamsi</emphasis>
Cai &amp; Ng, 2000
</taxonomicName>
. It could be distinguished from
<taxonomicName id="4C454D19550ACD5228006ED4FE742C88" authorityName="Cai &amp; Ng" authorityYear="2000" box="[344,474,1505,1528]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="259" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B931EA88550ACD5228006ED4FE742C88" box="[344,474,1505,1528]" italics="true" pageId="14" pageNumber="259">C. williamsi</emphasis>
</taxonomicName>
by the endopod of the male first pleopod, which has no appendix interna (vs. has a distinct appendix interna at the distal end of the endopod in
<taxonomicName id="4C454D19550ACD522BB56D14FF442F28" authorityName="Cai &amp; Ng" authorityYear="2000" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="259" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B931EA88550ACD522BB56D14FF442F28" italics="true" pageId="14" pageNumber="259">C. williamsi</emphasis>
</taxonomicName>
), the telson which terminates in a small projection (vs. no such projection in the latter), and the shorter antennular peduncle (0.7 times as long as carapace vs. 0.85-1.0 times in
<taxonomicName id="4C454D19550ACD5229D06D94FEA42FC8" authorityName="Cai &amp; Ng" authorityYear="2000" box="[136,266,1697,1720]" class="Malacostraca" family="Atyidae" genus="Caridina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="259" phylum="Arthropoda" rank="species" species="williamsi">
<emphasis id="B931EA88550ACD5229D06D94FEA42FC8" box="[136,266,1697,1720]" italics="true" pageId="14" pageNumber="259">C. williamsi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F6511550ACD5229D06DD4FE1E2F88" box="[136,432,1761,1785]" pageId="14" pageNumber="259" type="distribution">
<paragraph id="8BFA369A550ACD5229D06DD4FE1E2F88" blockId="14.[136,432,1761,1785]" box="[136,432,1761,1785]" pageId="14" pageNumber="259">
<emphasis id="B931EA88550ACD5229D06DD4FE872F89" bold="true" box="[136,297,1761,1785]" italics="true" pageId="14" pageNumber="259">Distribution. –</emphasis>
<collectingCountry id="F352760A550ACD5228696DD4FE032F88" box="[305,429,1761,1784]" name="Philippines" pageId="14" pageNumber="259">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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