320 lines
46 KiB
XML
320 lines
46 KiB
XML
<document id="5FAFA26F8CF8F020D543D12654FFF045" ID-CLB-Dataset="222446" ID-DOI="10.11646/zootaxa.5227.4.6" ID-GBIF-Dataset="d12208f6-3ae6-40e6-8e09-9cab4b7be698" ID-ISSN="1175-5326" ID-Zenodo-Dep="7518988" ID-ZooBank="28D21568-70C8-4A85-9B82-943B336A234E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1673312589758" checkinUser="plazi" docAuthor="Tchemeris, Aleksey N." docDate="2023" docId="03EC8786FFDDFFB2EB99F9B891424460" docLanguage="en" docName="zootaxa.5227.4.6.pdf" docOrigin="Zootaxa 5227 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kovalius logunovi Tchemeris 2023, sp. nov." docType="treatment" docVersion="7" lastPageNumber="494" masterDocId="FFD5FFFEFFDCFFBAEB0EFFF3925C457D" masterDocTitle="Kovalius-a new genus of cave-dwelling harvestmen from the Caucasus (Opiliones Sclerosomatidae: Leiobuninae)" masterLastPageNumber="494" masterPageNumber="486" pageNumber="487" updateTime="1699473677005" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="52F7A72683E1AE0EBB27A363FD75BCA2">Kovalius-a new genus of cave-dwelling harvestmen from the Caucasus (Opiliones Sclerosomatidae: Leiobuninae)</mods:title>
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<mods:name id="9FC0D6B8045F0015E60DD2ABDAAC4712" type="personal">
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<mods:namePart id="1AD6F684A95ED910949DB0712E49DF5C">Tchemeris, Aleksey N.</mods:namePart>
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<mods:title id="2453ADDB91DCB64FAB40E79596752A11">Zootaxa</mods:title>
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<mods:part id="1F7F230A989D5C9DA30B16898B4B4143">
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<mods:date id="0360F9E42DC06228DF24B15054383F65">2023</mods:date>
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<mods:number id="B952C9CCE4961BEAF19CAD02C9772AC2">2023-01-09</mods:number>
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<treatment id="03EC8786FFDDFFB2EB99F9B891424460" ID-DOI="http://doi.org/10.5281/zenodo.7525195" ID-GBIF-Taxon="208788521" ID-Zenodo-Dep="7525195" LSID="urn:lsid:plazi:treatment:03EC8786FFDDFFB2EB99F9B891424460" httpUri="http://treatment.plazi.org/id/03EC8786FFDDFFB2EB99F9B891424460" lastPageId="8" lastPageNumber="494" pageId="1" pageNumber="487">
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<taxonomicName id="4C454D13FFDDFFBBEB99F9B8932E431B" ID-CoL="B2YKG" authority="Tchemeris, 2023" authorityName="Tchemeris" authorityYear="2023" box="[151,370,1611,1638]" class="Arachnida" family="Sclerosomatidae" genus="Kovalius" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="487" phylum="Arthropoda" rank="species" species="logunovi" status="sp. nov.">
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<emphasis id="B931EA82FFDDFFBBEB99F9B8932E431B" bold="true" box="[151,370,1611,1638]" italics="true" pageId="1" pageNumber="487">Kovalius logunovi</emphasis>
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<taxonomicNameLabel id="A20257F9FFDDFFBBEA74F9BF938B431B" box="[378,471,1612,1638]" pageId="1" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C35F651BFFDDFFBBEB99F983932F4225" pageId="1" pageNumber="487" type="description">
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<paragraph id="8BFA3690FFDDFFBBEB99F983935A43F7" blockId="1.[151,471,1611,1674]" box="[151,262,1648,1674]" pageId="1" pageNumber="487">
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<figureCitation id="137E2A15FFDDFFBBEB99F983935A43F7" box="[151,262,1648,1674]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–7" captionStart-2="FIGURES 8–10" captionStartId-0="2.[151,264,1943,1967]" captionStartId-1="3.[151,264,1943,1967]" captionStartId-2="4.[151,264,1943,1967]" captionTargetBox-0="[206,1383,192,1903]" captionTargetBox-1="[201,1386,192,1920]" captionTargetBox-2="[395,1191,194,1909]" captionTargetId-0="figure-15@2.[204,1383,181,1920]" captionTargetId-1="figure-22@3.[201,1386,181,1920]" captionTargetId-2="figure-15@4.[380,1208,181,1920]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1–2. Kovalius logunovi sp. nov., male holotype: 1—body dorsal view, 2—body lateral view. Scale bar 1,0 mm." captionText-1="FIGURES 3–7. Kovalius logunovi sp. nov., male holotype: 3—right pedipalp inner view, 4—tarsal claw of palp, 5—second and third segments of left chelicerae frontal view, 6—left chelicerae lateral view, 7—patellae and tibiae of left pedipalp dorsal view. 3, 5-7—scale bar 0,5 mm, 4—scale bar 0,25 mm." captionText-2="FIGURES 8–10. Kovalius logunovi sp. nov., penis of male holotype: 8—lateral view, 9—ventral view, 10—dorsal view. Scale bar 0,5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7518990" figureDoi-1="http://doi.org/10.5281/zenodo.7518992" figureDoi-2="http://doi.org/10.5281/zenodo.7518994" httpUri-0="https://zenodo.org/record/7518990/files/figure.png" httpUri-1="https://zenodo.org/record/7518992/files/figure.png" httpUri-2="https://zenodo.org/record/7518994/files/figure.png" pageId="1" pageNumber="487">Figs 1–10</figureCitation>
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</paragraph>
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http://zoobank.org/
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urn:lsid:zoobank.org:act:
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<paragraph id="8BFA3690FFDDFFBBEB99F906932F4225" blockId="1.[151,1437,1781,1988]" pageId="1" pageNumber="487">
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<emphasis id="B931EA82FFDDFFBBEB99F90692B84272" bold="true" box="[151,228,1781,1807]" pageId="1" pageNumber="487">Types.</emphasis>
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<materialsCitation id="3B2D3CCDFFDDFFBBEBE8F90691C3424E" ID-GBIF-Occurrence="4012651303" collectingDate="2015-09-17" collectionCode="ZIN" collectorName="NW Caucasus & A. G. Kovalleg" country="Russia" county="SokolovaCave" elevation="300" latitude="43.688614" location="Sochi" longLatPrecision="19" longitude="39.858887" municipality="River" pageId="1" pageNumber="487" specimenCode="ZIN5201" specimenCount="1" typeStatus="holotype">
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male(
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<specimenCode id="DBE39EEBFFDDFFBBEA9EF90693A0426D" box="[400,508,1781,1808]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="487" type="Museum">ZIN5201</specimenCode>
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)
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,
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<collectorName id="26B05346FFDDFFBBE954F90590A44272" box="[602,760,1781,1808]" pageId="1" pageNumber="487">NW Caucasus</collectorName>
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,region of
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,
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Alek
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</location>
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,gorge ofAts
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<collectingMunicipality id="6B9EACEAFFDDFFBBEFD9F906974B4272" box="[1239,1303,1781,1807]" pageId="1" pageNumber="487">River</collectingMunicipality>
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,
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a.s.l.
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</elevation>
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,
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<collectingCounty id="629B4E1CFFDDFFBBEB99F8EA9361424E" box="[151,317,1817,1843]" pageId="1" pageNumber="487">SokolovaCave</collectingCounty>
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,[
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/
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],
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<date id="FFFB1050FFDDFFBBE979F8EA90A74249" box="[631,763,1817,1844]" pageId="1" pageNumber="487" value="2015-09-17">
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<collectingDate id="EFBFE9B8FFDDFFBBE979F8EA90A74249" box="[631,763,1817,1844]" pageId="1" pageNumber="487" value="2015-09-17">17.IX.2015</collectingDate>
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</date>
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.
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<collectorName id="26B05346FFDDFFBBE9F0F8EA91C3424E" box="[766,927,1817,1844]" pageId="1" pageNumber="487">A.G.Kovalleg</collectorName>
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</materialsCitation>
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.
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<materialsCitation id="3B2D3CCDFFDDFFBBE8ADF8E99760424E" ID-GBIF-Occurrence="4012651301" box="[931,1340,1817,1844]" collectionCode="ISEA" country="Russia" pageId="1" pageNumber="487" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<typeStatus id="54FE8832FFDDFFBBE8ADF8E9964F4249" box="[931,1043,1818,1844]" pageId="1" pageNumber="487" type="paratype">Paratypes</typeStatus>
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:
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,(
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<collectionCode id="ED54AE55FFDDFFBBEF63F8EA96F7424E" box="[1133,1195,1817,1843]" pageId="1" pageNumber="487">ISEA</collectionCode>
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O.001.0291)
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</materialsCitation>
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,
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<materialsCitation id="3B2D3CCDFFDDFFBBEE4FF8EA93334225" ID-GBIF-Occurrence="4012651302" collectionCode="ZIN" country="Russia" pageId="1" pageNumber="487" specimenCode="ZIN 5202" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<specimenCount id="9D43FD19FFDDFFBBEE4FF8EA97C0424E" box="[1345,1436,1817,1843]" pageId="1" pageNumber="487" type="female">1female</specimenCount>
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(
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<specimenCode id="DBE39EEBFFDDFFBBEB91F8CE93534225" box="[159,271,1853,1880]" collectionCode="ZIN" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="1" pageNumber="487" type="Museum">ZIN 5202</specimenCode>
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||
)—idem
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F651BFFDDFFBBEBC9F892963E42BE" pageId="1" pageNumber="487" type="etymology">
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<paragraph id="8BFA3690FFDDFFBBEBC9F892963E42BE" blockId="1.[151,1437,1781,1988]" pageId="1" pageNumber="487">
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||
<emphasis id="B931EA82FFDDFFBBEBC9F89293124206" bold="true" box="[199,334,1889,1915]" pageId="1" pageNumber="487">Etymology.</emphasis>
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The new species is named in honour of my mentor Dr. Dmitri V. Logunov, the Russian and English arachnologist (
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<collectingRegion id="4981F872FFDDFFBBEA30F876939842E2" box="[318,452,1925,1951]" country="United Kingdom" name="Manchester" pageId="1" pageNumber="487">Manchester</collectingRegion>
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,
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<collectingCountry id="F3527600FFDDFFBBEADEF87693A042E2" box="[464,508,1925,1951]" name="United Kingdom" pageId="1" pageNumber="487">UK</collectingCountry>
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), professional collector and author of a number of publications on world fauna of spiders, in particular the family
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<taxonomicName id="4C454D13FFDDFFBBEAF3F85A903742BE" authorityName="Blackwall" authorityYear="1841" box="[509,619,1961,1987]" class="Arachnida" family="Salticidae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="487" phylum="Arthropoda" rank="family">Salticidae</taxonomicName>
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and some papers about
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<taxonomicName id="4C454D13FFDDFFBBE877F85A91BA42BE" authorityName="Sundevall" authorityYear="1833" box="[889,998,1961,1987]" class="Arachnida" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="487" phylum="Arthropoda" rank="order">Opiliones</taxonomicName>
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of Siberia.
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</paragraph>
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</subSubSection>
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||
<caption id="DF3A6618FFDEFFB8EB99F864972D42D2" ID-DOI="http://doi.org/10.5281/zenodo.7518990" ID-Zenodo-Dep="7518990" box="[151,1393,1943,1968]" httpUri="https://zenodo.org/record/7518990/files/figure.png" pageId="2" pageNumber="488" startId="2.[151,264,1943,1967]" targetBox="[206,1383,192,1903]" targetPageId="2">
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<paragraph id="8BFA3690FFDEFFB8EB99F864972D42D2" blockId="2.[151,1393,1943,1968]" box="[151,1393,1943,1968]" pageId="2" pageNumber="488">
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||
<emphasis id="B931EA82FFDEFFB8EB99F864936642D2" bold="true" box="[151,314,1943,1967]" pageId="2" pageNumber="488">FIGURES 1–2.</emphasis>
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||
<taxonomicName id="4C454D13FFDEFFB8EA4FF86B93A642CD" authority="Tchemeris, 2023" authorityName="Tchemeris" authorityYear="2023" box="[321,506,1944,1968]" class="Arachnida" family="Sclerosomatidae" genus="Kovalius" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="488" phylum="Arthropoda" rank="species" species="logunovi" status="sp. nov.">
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<emphasis id="B931EA82FFDEFFB8EA4FF86B93A642CD" box="[321,506,1944,1968]" italics="true" pageId="2" pageNumber="488">Kovalius logunovi</emphasis>
|
||
</taxonomicName>
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||
<taxonomicNameLabel id="A20257F9FFDEFFB8E90EF86B900D42D2" box="[512,593,1944,1967]" pageId="2" pageNumber="488" rank="species">sp. nov.</taxonomicNameLabel>
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||
, male holotype: 1—body dorsal view, 2—body lateral view. Scale bar 1,0 mm.
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||
</paragraph>
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||
</caption>
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||
<caption id="DF3A6618FFDFFFB9EB99F86490D0428A" ID-DOI="http://doi.org/10.5281/zenodo.7518992" ID-Zenodo-Dep="7518992" httpUri="https://zenodo.org/record/7518992/files/figure.png" pageId="3" pageNumber="489" startId="3.[151,264,1943,1967]" targetBox="[201,1386,192,1920]" targetPageId="3">
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<paragraph id="8BFA3690FFDFFFB9EB99F86490D0428A" blockId="3.[151,1437,1943,2040]" pageId="3" pageNumber="489">
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||
<emphasis id="B931EA82FFDFFFB9EB99F864936642D2" bold="true" box="[151,314,1943,1967]" pageId="3" pageNumber="489">FIGURES 3–7.</emphasis>
|
||
<taxonomicName id="4C454D13FFDFFFB9EA31F86B93AB42CD" authority="Tchemeris, 2023" authorityName="Tchemeris" authorityYear="2023" box="[319,503,1944,1968]" class="Arachnida" family="Sclerosomatidae" genus="Kovalius" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="489" phylum="Arthropoda" rank="species" species="logunovi" status="sp. nov.">
|
||
<emphasis id="B931EA82FFDFFFB9EA31F86B93AB42CD" box="[319,503,1944,1968]" italics="true" pageId="3" pageNumber="489">Kovalius logunovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20257F9FFDFFFB9EAF2F86B901042D2" box="[508,588,1944,1967]" pageId="3" pageNumber="489" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male holotype: 3—right pedipalp inner view, 4—tarsal claw of palp, 5—second and third segments of left chelicerae frontal view, 6—left chelicerae lateral view, 7—patellae and tibiae of left pedipalp dorsal view. 3, 5-7—scale bar 0,5 mm, 4—scale bar 0,25 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3A6618FFD8FFBEEB99F864934F42AE" ID-DOI="http://doi.org/10.5281/zenodo.7518994" ID-Zenodo-Dep="7518994" httpUri="https://zenodo.org/record/7518994/files/figure.png" pageId="4" pageNumber="490" startId="4.[151,264,1943,1967]" targetBox="[395,1191,194,1909]" targetPageId="4">
|
||
<paragraph id="8BFA3690FFD8FFBEEB99F864934F42AE" blockId="4.[151,1436,1943,2004]" pageId="4" pageNumber="490">
|
||
<emphasis id="B931EA82FFD8FFBEEB99F864931B42D2" bold="true" box="[151,327,1943,1967]" pageId="4" pageNumber="490">FIGURES 8–10.</emphasis>
|
||
<taxonomicName id="4C454D13FFD8FFBEEA43F86B905B42CD" authority="Tchemeris, 2023" authorityName="Tchemeris" authorityYear="2023" box="[333,519,1944,1968]" class="Arachnida" family="Sclerosomatidae" genus="Kovalius" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="490" phylum="Arthropoda" rank="species" species="logunovi" status="sp. nov.">
|
||
<emphasis id="B931EA82FFD8FFBEEA43F86B905B42CD" box="[333,519,1944,1968]" italics="true" pageId="4" pageNumber="490">Kovalius logunovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20257F9FFD8FFBEE903F86B900242D2" box="[525,606,1944,1967]" pageId="4" pageNumber="490" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, penis of male holotype: 8—lateral view, 9—ventral view, 10—dorsal view. Scale bar 0,5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3A6618FFD9FFBFEB99F98890B643CA" ID-DOI="http://doi.org/10.5281/zenodo.7518996" ID-Zenodo-Dep="7518996" httpUri="https://zenodo.org/record/7518996/files/figure.png" pageId="5" pageNumber="491" startId="5.[151,264,1659,1683]" targetBox="[158,1430,181,1635]" targetPageId="5">
|
||
<paragraph id="8BFA3690FFD9FFBFEB99F98890B643CA" blockId="5.[151,1436,1659,1719]" pageId="5" pageNumber="491">
|
||
<emphasis id="B931EA82FFD9FFBFEB99F988930F43EE" bold="true" box="[151,339,1659,1683]" pageId="5" pageNumber="491">FIGURES 11–12.</emphasis>
|
||
<taxonomicName id="4C454D13FFD9FFBFEA57F988904E43EE" authority="Tchemeris, 2023" authorityName="Tchemeris" authorityYear="2023" box="[345,530,1659,1683]" class="Arachnida" family="Sclerosomatidae" genus="Kovalius" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="491" phylum="Arthropoda" rank="species" species="logunovi" status="sp. nov.">
|
||
<emphasis id="B931EA82FFD9FFBFEA57F988904E43EE" box="[345,530,1659,1683]" italics="true" pageId="5" pageNumber="491">Kovalius logunovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20257F9FFD9FFBFE916F98F903543EE" box="[536,617,1660,1683]" pageId="5" pageNumber="491" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, female paratype: 11—body dorsal view; 12—patellae and tibiae of left pedipalp dorsal view. 11—scale bar 1.0 mm, 12—scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35F651BFFD9FFBCEBC9F913964F4098" lastPageId="6" lastPageNumber="492" pageId="5" pageNumber="491" type="description">
|
||
<paragraph id="8BFA3690FFD9FFBFEBC9F9139354428B" blockId="5.[151,1437,1760,2038]" pageId="5" pageNumber="491">
|
||
<emphasis id="B931EA82FFD9FFBFEBC9F91393054387" bold="true" box="[199,345,1760,1786]" pageId="5" pageNumber="491">Description.</emphasis>
|
||
MALE.
|
||
<emphasis id="B931EA82FFD9FFBFEAB3F91290034387" box="[445,607,1761,1786]" italics="true" pageId="5" pageNumber="491">Measurements</emphasis>
|
||
. Body: length 3.52; abdomen maximal width 2.60. Carapace length: 1.12; maximal width 1.92. Clypeus length (distance between anterior margin of carapace and ocularium): 0.58. Eye tubercle width: 0.54. Chelicera: basal segment 1.07 long; distal segment 1.64 long; chela maximal length 0.63. Length of pedipalpal trochanter, femur, patella, tibia and tarsus: 0.26 + 1.38 + 0.56 (apophysis 0.30 not included in total length) + 0.90 + 1.71 = 4.81. Length of leg segments (femur, patella, tibia, metatarsus, tarsus): I: 4.83 + 1.37 + 5.48 + 6.04 + 10.17 = 27.89; II: 8.87 + 1.80 + 9.72 + 8.92 + 23.47 = 52.78; III: 4.91 + 1.52 + 5.23 + 6.93 + 12.14 = 30.73; IV: 6.35 + 1.49 + 7.38 + 9.43 + 16.72 = 41.37. Penis 1.72 long, base width 0.225, glans 0.21 long, stylus 0.14 long.
|
||
</paragraph>
|
||
<paragraph id="8BFA3690FFDAFFBCEBC9FF64928B4460" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="492">
|
||
Body medium sized with soft integument (
|
||
<figureCitation id="137E2A15FFDAFFBCE992FF64915C45CC" box="[668,768,151,177]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[206,1383,192,1903]" captionTargetId="figure-15@2.[204,1383,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–2. Kovalius logunovi sp. nov., male holotype: 1—body dorsal view, 2—body lateral view. Scale bar 1,0 mm." figureDoi="http://doi.org/10.5281/zenodo.7518990" httpUri="https://zenodo.org/record/7518990/files/figure.png" pageId="6" pageNumber="492">Figs 1–2</figureCitation>
|
||
). Carapace with scant short setae, carapace clearly separated from abdomen. Ozophores distinct. Ocularium (
|
||
<figureCitation id="137E2A15FFDAFFBCE9A6FF4890B545A8" box="[680,745,186,213]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[206,1383,192,1903]" captionTargetId="figure-15@2.[204,1383,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–2. Kovalius logunovi sp. nov., male holotype: 1—body dorsal view, 2—body lateral view. Scale bar 1,0 mm." figureDoi="http://doi.org/10.5281/zenodo.7518990" httpUri="https://zenodo.org/record/7518990/files/figure.png" pageId="6" pageNumber="492">Figs 1</figureCitation>
|
||
-25) stocky, bearing scant short setae. Supra-cheliceral lamella small, unarmed. Meta- and mesopeltidium and all abdominal tergites and sternites with transverse rows of short setae.
|
||
</paragraph>
|
||
<paragraph id="8BFA3690FFDAFFBCEBC9FED496E04418" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="492">
|
||
Chelicerae of typical size (
|
||
<figureCitation id="137E2A15FFDAFFBCEAE2FED49017443C" box="[492,587,294,321]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[201,1386,192,1920]" captionTargetId="figure-22@3.[201,1386,181,1920]" captionTargetPageId="3" captionText="FIGURES 3–7. Kovalius logunovi sp. nov., male holotype: 3—right pedipalp inner view, 4—tarsal claw of palp, 5—second and third segments of left chelicerae frontal view, 6—left chelicerae lateral view, 7—patellae and tibiae of left pedipalp dorsal view. 3, 5-7—scale bar 0,5 mm, 4—scale bar 0,25 mm." figureDoi="http://doi.org/10.5281/zenodo.7518992" httpUri="https://zenodo.org/record/7518992/files/figure.png" pageId="6" pageNumber="492">Figs 5-6</figureCitation>
|
||
). First segment with ventral spine, dorsally with group of short setae. Second segment anteriorly with short and long setae, ectal surface with short setae grouped near chela.
|
||
</paragraph>
|
||
<paragraph id="8BFA3690FFDAFFBCEBC9FE9C92E94488" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="492">
|
||
Pedipalps not elongate (
|
||
<figureCitation id="137E2A15FFDAFFBCEAD7FE9C907D44F4" box="[473,545,367,393]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[206,1383,192,1903]" captionTargetId="figure-15@2.[204,1383,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–2. Kovalius logunovi sp. nov., male holotype: 1—body dorsal view, 2—body lateral view. Scale bar 1,0 mm." figureDoi="http://doi.org/10.5281/zenodo.7518990" httpUri="https://zenodo.org/record/7518990/files/figure.png" pageId="6" pageNumber="492">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="137E2A15FFDAFFBCE93FFE9C900344F4" box="[561,607,367,393]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[201,1386,192,1920]" captionTargetId="figure-22@3.[201,1386,181,1920]" captionTargetPageId="3" captionText="FIGURES 3–7. Kovalius logunovi sp. nov., male holotype: 3—right pedipalp inner view, 4—tarsal claw of palp, 5—second and third segments of left chelicerae frontal view, 6—left chelicerae lateral view, 7—patellae and tibiae of left pedipalp dorsal view. 3, 5-7—scale bar 0,5 mm, 4—scale bar 0,25 mm." figureDoi="http://doi.org/10.5281/zenodo.7518992" httpUri="https://zenodo.org/record/7518992/files/figure.png" pageId="6" pageNumber="492">3, 7</figureCitation>
|
||
). Patellae with distinctly pronounced inner apophysis exceeding half the length of patellae (
|
||
<figureCitation id="137E2A15FFDAFFBCEA65FE6093EC44D0" box="[363,432,403,429]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[201,1386,192,1920]" captionTargetId="figure-22@3.[201,1386,181,1920]" captionTargetPageId="3" captionText="FIGURES 3–7. Kovalius logunovi sp. nov., male holotype: 3—right pedipalp inner view, 4—tarsal claw of palp, 5—second and third segments of left chelicerae frontal view, 6—left chelicerae lateral view, 7—patellae and tibiae of left pedipalp dorsal view. 3, 5-7—scale bar 0,5 mm, 4—scale bar 0,25 mm." figureDoi="http://doi.org/10.5281/zenodo.7518992" httpUri="https://zenodo.org/record/7518992/files/figure.png" pageId="6" pageNumber="492">Fig. 7</figureCitation>
|
||
). Tibiae with small inner distal apophysis. Patellae, tibiae and tarsi densely covered with microsetae. All segments with variable length setae, longer on ventral surface of femur. Tarsal claw toothed (
|
||
<figureCitation id="137E2A15FFDAFFBCEE61FE4492F44488" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[201,1386,192,1920]" captionTargetId="figure-22@3.[201,1386,181,1920]" captionTargetPageId="3" captionText="FIGURES 3–7. Kovalius logunovi sp. nov., male holotype: 3—right pedipalp inner view, 4—tarsal claw of palp, 5—second and third segments of left chelicerae frontal view, 6—left chelicerae lateral view, 7—patellae and tibiae of left pedipalp dorsal view. 3, 5-7—scale bar 0,5 mm, 4—scale bar 0,25 mm." figureDoi="http://doi.org/10.5281/zenodo.7518992" httpUri="https://zenodo.org/record/7518992/files/figure.png" pageId="6" pageNumber="492">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFA3690FFDAFFBCEBC9FE0C93DB47D4" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="492">
|
||
Legs not long, thin, cylindrical in cross section (
|
||
<figureCitation id="137E2A15FFDAFFBCE80CFE0C911B4764" box="[770,839,511,537]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[206,1383,192,1903]" captionTargetId="figure-15@2.[204,1383,181,1920]" captionTargetPageId="2" captionText="FIGURES 1–2. Kovalius logunovi sp. nov., male holotype: 1—body dorsal view, 2—body lateral view. Scale bar 1,0 mm." figureDoi="http://doi.org/10.5281/zenodo.7518990" httpUri="https://zenodo.org/record/7518990/files/figure.png" pageId="6" pageNumber="492">Fig. 1</figureCitation>
|
||
). Femora I and III thickened, noticeably thickened femora I and provided with more developed armament than on other legs. All segments densely covered with trichomes and each accompanied by microsetae - a ‘sensillum chaeticum’. Coxae covered with long setae. Femora, patellae and tibiae with rows of short setae and acute inclined tubercles, more prominent on femora I. Tibia II with 4–5 pseudosegments.
|
||
</paragraph>
|
||
<paragraph id="8BFA3690FFDAFFBCEBC9FD4093B846B4" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="492">
|
||
Penis weakly sclerotized (
|
||
<figureCitation id="137E2A15FFDAFFBCEAE1FD40903E47B0" box="[495,610,691,717]" captionStart="FIGURES 8–10" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[395,1191,194,1909]" captionTargetId="figure-15@4.[380,1208,181,1920]" captionTargetPageId="4" captionText="FIGURES 8–10. Kovalius logunovi sp. nov., penis of male holotype: 8—lateral view, 9—ventral view, 10—dorsal view. Scale bar 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.7518994" httpUri="https://zenodo.org/record/7518994/files/figure.png" pageId="6" pageNumber="492">Figs 8–10</figureCitation>
|
||
) with extended base. Corpus, glans and stylus fused. Corpus moderately expanded from the glans towards the middle and narrowed to the base, ventrally with microdenticles from the center to the base (
|
||
<figureCitation id="137E2A15FFDAFFBCEA13FD08933E4668" box="[285,354,763,789]" captionStart="FIGURES 8–10" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[395,1191,194,1909]" captionTargetId="figure-15@4.[380,1208,181,1920]" captionTargetPageId="4" captionText="FIGURES 8–10. Kovalius logunovi sp. nov., penis of male holotype: 8—lateral view, 9—ventral view, 10—dorsal view. Scale bar 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.7518994" httpUri="https://zenodo.org/record/7518994/files/figure.png" pageId="6" pageNumber="492">Fig. 9</figureCitation>
|
||
). Wing-like oval structures directed upwards are provided below the middle of the corpus. These 'wings' are weakly developed and slightly protruding over sides of the corpus forming membrane-like cover. Glans in lateral view strongly expanded and forms a ‘window’ tightened by a transparent membrane in the center (
|
||
<figureCitation id="137E2A15FFDAFFBCEE44FCB097D24620" box="[1354,1422,835,861]" captionStart="FIGURES 8–10" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[395,1191,194,1909]" captionTargetId="figure-15@4.[380,1208,181,1920]" captionTargetPageId="4" captionText="FIGURES 8–10. Kovalius logunovi sp. nov., penis of male holotype: 8—lateral view, 9—ventral view, 10—dorsal view. Scale bar 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.7518994" httpUri="https://zenodo.org/record/7518994/files/figure.png" pageId="6" pageNumber="492">Fig. 8</figureCitation>
|
||
). Due to the thin cuticle protruding beyond the edge, the stylus in lateral view looks like harpoon. Apex of the stylus appearing flat in dorsal and ventral view, with widened base with lateral micropores. Base of penis sclerotized (
|
||
<figureCitation id="137E2A15FFDAFFBCEE61FC7892E946B4" captionStart="FIGURES 8–10" captionStartId="4.[151,264,1943,1967]" captionTargetBox="[395,1191,194,1909]" captionTargetId="figure-15@4.[380,1208,181,1920]" captionTargetPageId="4" captionText="FIGURES 8–10. Kovalius logunovi sp. nov., penis of male holotype: 8—lateral view, 9—ventral view, 10—dorsal view. Scale bar 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.7518994" httpUri="https://zenodo.org/record/7518994/files/figure.png" pageId="6" pageNumber="492">Fig. 10</figureCitation>
|
||
), wide and with two lobes.
|
||
</paragraph>
|
||
<paragraph id="8BFA3690FFDAFFBCEBC9FC2092BA41B8" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="492">Body darker dorsally, carapace with brownish patterns and spots, laterally with dark brown and whitish spots. Abdomen dark ochre in center and with large dark brown patterns and small round and oval light ochre spots on sides, anteriorly with light ochre band, last tergites entirely dark brown. Ocularium dark brown. Venter light ochre to ochre on coxae and ochre-brown on abdomen. Chelicerae light ochre with ochre spots. Pedipalps ventrally light ochre; femora, patellae, tibiae dorsally brown with small rounded ochre spots, tarsi completely ochre. Legs ochre, with small brown and light ochre spots on trochanters. Penis light ochre, with stylus and ventral border of glans brown.</paragraph>
|
||
<paragraph id="8BFA3690FFDAFFBCEBC9FB3C975B40E0" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="492">
|
||
FEMALE.
|
||
<emphasis id="B931EA82FFDAFFBCEA49FB3C93B54195" box="[327,489,1231,1256]" italics="true" pageId="6" pageNumber="492">Measurements</emphasis>
|
||
. Body: length 4.36; abdomen maximal width 2.88. Carapace length: 1.59; width in maximal 2.61. Clypeus length: 0.40. Eye tubercle width: 0.55. Chelicera: basal segment 1.01 long; distal segment 1.13 long; chela maximal long 0.54. Length of pedipalpal segments (trochanter, femur, patella, tibia, tarsus): 0.29 + 1.23 + 0.52 (apophysis 0.47 not included in total length) + 1.10 + 1.62 = 4.76. Length of leg segments (femur, patella, tibia, metatarsus, tarsus): I: 3.45 + 1.16 + 3.46 + 3.72 + 7.32 = 19.11; II: 6.22 + 1.68 + 6.63 + 5.82 + 18.82 = 39.17; III: 3.64 + 1.31 + 3.90 + 4.08 + 7.62 = 20.55; IV: 4.78 + 1.48 + 5.22 + 5.92 + 16.83 = 34.23.
|
||
</paragraph>
|
||
<paragraph id="8BFA3690FFDAFFBCEBC9FA54964F4098" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="492">General appearance as in male, but with longer, wider body. Apophysis of pedipalpal patellae and tibiae noticeably larger than in male. Tibia II same as male with 4–5 pseudosegments.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F651BFFDAFFBCEBC9FA1C917942A0" pageId="6" pageNumber="492" type="biology_ecology">
|
||
<paragraph id="8BFA3690FFDAFFBCEBC9FA1C972143E4" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="492">
|
||
<emphasis id="B931EA82FFDAFFBCEBC9FA1C937A4374" bold="true" box="[199,294,1519,1545]" pageId="6" pageNumber="492">Ecology</emphasis>
|
||
.
|
||
<taxonomicName id="4C454D13FFDAFFBCEA3AFA1C905D4374" authority="Tchemeris, 2023" authorityName="Tchemeris" authorityYear="2023" box="[308,513,1519,1545]" class="Arachnida" family="Sclerosomatidae" genus="Kovalius" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="492" phylum="Arthropoda" rank="species" species="logunovi" status="sp. nov.">
|
||
<emphasis id="B931EA82FFDAFFBCEA3AFA1C905D4374" box="[308,513,1519,1545]" italics="true" pageId="6" pageNumber="492">Kovalius logunovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20257F9FFDAFFBCE906FA1C90034374" box="[520,607,1519,1545]" pageId="6" pageNumber="492" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is a troglophilic species. The specimens were collected in Sokolova Cave about
|
||
<quantity id="4CBD9B75FFDAFFBCEBD2F9E193624350" box="[220,318,1554,1581]" metricMagnitude="1" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="6" pageNumber="492" unit="m" value="20.0" valueMax="25.0" valueMin="15.0">15–25 m</quantity>
|
||
from its entrance, on the wall of the cave gallery in the subhorizontal part of the cave, slightly above the cave lake located at the entrance to the cave and on the wall of this gallery above the puddles located behind the lake (
|
||
<figureCitation id="137E2A15FFDAFFBCEBDDF9A893084308" box="[211,340,1627,1653]" captionStart="FIGURES 13–18" captionStartId="7.[151,264,1908,1932]" captionTargetBox="[164,1419,181,1882]" captionTargetId="figure-22@7.[164,1423,181,1884]" captionTargetPageId="7" captionText="FIGURES 13–18. Type locality of Kovalius logunovi sp. nov. and specimen in situ: 13—entrance to the Sokolova Cave; 14- 16 habitat in type locality: 14—on the wall at the entrance nearby the lake; 15—16 wall with seeping water and stalagmite; 17—cave river (13—15, photo: R. Vargovitsh; 16—17, photo: A. Koval); 18—waterfall and river within the cave (photo: R. Vargovitsh)." figureDoi="http://doi.org/10.5281/zenodo.7518998" httpUri="https://zenodo.org/record/7518998/files/figure.png" pageId="6" pageNumber="492">Figs 13–16</figureCitation>
|
||
). Light does not penetrate into the cave; there is always darkness and a cool cave microclimate. The
|
||
<geoCoordinate id="EE715057FFDAFFBCEB99F98C928C43F0" box="[151,208,1663,1689]" degrees="10.2" direction="west" orientation="longitude" pageId="6" pageNumber="492" precision="5555" value="-10.2">
|
||
10.2
|
||
<superScript id="7C309BD8FFDAFFBCEBC6F98C928C43F0" attach="left" box="[200,208,1663,1677]" fontSize="6" pageId="6" pageNumber="492">о</superScript>
|
||
</geoCoordinate>
|
||
С air temperature and almost 100% absolute air humidity were registered when specimens were collected.
|
||
</paragraph>
|
||
<paragraph id="8BFA3690FFDAFFBCEBC9F950917942A0" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="492">
|
||
Brief description of Sokolova (Atsinskaya) Cave Based on
|
||
<bibRefCitation id="EFD44B61FFDAFFBCE879F950964E43C0" author="Caves" box="[887,1042,1699,1725]" pageId="6" pageNumber="492" refId="ref3803" refString="Caves (2017) Information and Search Engine. Available from: https: // speleoatlas. ru / caves / sokolova- 12229 / (accesses 13 May 2022)" type="url" year="2017">Caves (2017)</bibRefCitation>
|
||
, Sokolova (Atsinskaya) Cave was formed in subhorizontal Upper Jurassic limestones, and presents the lower part of the hydrological system of the Alek Mt Range divided into small blocks. The cave's entrance gallery first drops sharply, forming a small siphon lake, then the cave passage turns into a series of ledges leading to an underground river. Forty meters from the fork upstream is situated 8-meter waterfall, behind which appear two parallel courses located at different levels. An underground river flows along the right (lower) course in 0.8–2.5-m depth; the left (upper) course is dry, with accumulations of clay, pebbles and numerous leaks. After
|
||
<quantity id="4CBD9B75FFDAFFBCE812F889910A42E8" box="[796,854,1914,1941]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="492" unit="m" value="50.0">50 m</quantity>
|
||
, the passages merge again, forming one high gallery about
|
||
<quantity id="4CBD9B75FFDAFFBCEBD2F86C937E42C4" box="[220,290,1951,1977]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="492" unit="m" value="100.0">100 m</quantity>
|
||
in length. A lake with "upper" siphons is formed in a distant of the gallery. Extent of the cave is
|
||
<quantity id="4CBD9B75FFDAFFBCEE41F86C97CB42C4" box="[1359,1431,1951,1977]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="492" unit="m" value="600.0">600 m</quantity>
|
||
, depth
|
||
<quantity id="4CBD9B75FFDAFFBCEBD2F830934B42A0" box="[220,279,1987,2013]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="492" unit="m" value="20.0">20 m</quantity>
|
||
, area
|
||
<quantity id="4CBD9B75FFDAFFBCEA57F83093F242A0" box="[345,430,1987,2013]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="492" unit="m" value="2000.0">2000 m</quantity>
|
||
<superScript id="7C309BD8FFDAFFBCEAA0F83193EA42AD" attach="left" box="[430,438,1986,2000]" fontSize="6" pageId="6" pageNumber="492">2</superScript>
|
||
, volume
|
||
<quantity id="4CBD9B75FFDAFFBCE911F830902842A0" box="[543,628,1987,2013]" metricMagnitude="3" metricUnit="m" metricValue="8.4" pageId="6" pageNumber="492" unit="m" value="8400.0">8400 m</quantity>
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||
<superScript id="7C309BD8FFDAFFBCE97DF831902742AD" attach="left" box="[627,635,1986,2000]" fontSize="6" pageId="6" pageNumber="492">3</superScript>
|
||
, altitude
|
||
<specimenCount id="9D43FD19FFDAFFBCE9EBF831917942A0" box="[741,805,1986,2013]" pageId="6" pageNumber="492" type="generic">
|
||
<quantity id="4CBD9B75FFDAFFBCE9EBF831917D42A0" box="[741,801,1986,2013]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="492" unit="m" value="65.0">65 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3A6618FFDBFFBDEB99F88793484285" ID-DOI="http://doi.org/10.5281/zenodo.7518998" ID-Zenodo-Dep="7518998" httpUri="https://zenodo.org/record/7518998/files/figure.png" pageId="7" pageNumber="493" startId="7.[151,264,1908,1932]" targetBox="[164,1419,181,1882]" targetPageId="7">
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||
<paragraph id="8BFA3690FFDBFFBDEB99F88793484285" blockId="7.[151,1437,1908,2040]" pageId="7" pageNumber="493">
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||
<emphasis id="B931EA82FFDBFFBDEB99F887930942F1" bold="true" box="[151,341,1908,1932]" pageId="7" pageNumber="493">FIGURES 13–18.</emphasis>
|
||
Type locality of
|
||
<taxonomicName id="4C454D13FFDBFFBDE908F887909D42F1" authority="Tchemeris, 2023" authorityName="Tchemeris" authorityYear="2023" box="[518,705,1908,1932]" class="Arachnida" family="Sclerosomatidae" genus="Kovalius" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="493" phylum="Arthropoda" rank="species" species="logunovi" status="sp. nov.">
|
||
<emphasis id="B931EA82FFDBFFBDE908F887909D42F1" box="[518,705,1908,1932]" italics="true" pageId="7" pageNumber="493">Kovalius logunovi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20257F9FFDBFFBDE9C6F886914442F1" box="[712,792,1909,1932]" pageId="7" pageNumber="493" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and specimen in situ: 13—entrance to the Sokolova Cave; 14- 16 habitat in type locality: 14—on the wall at the entrance nearby the lake; 15—16 wall with seeping water and stalagmite; 17—cave river (13—15, photo: R. Vargovitsh; 16—17, photo: A. Koval); 18—waterfall and river within the cave (photo: R. Vargovitsh).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35F651BFFD4FFB2EBC9FF6491424460" pageId="8" pageNumber="494" type="discussion">
|
||
<paragraph id="8BFA3690FFD4FFB2EBC9FF6493C24584" blockId="8.[151,1436,151,285]" pageId="8" pageNumber="494">
|
||
<emphasis id="B931EA82FFD4FFB2EBC9FF64936745CC" bold="true" box="[199,315,151,177]" pageId="8" pageNumber="494">Remarks.</emphasis>
|
||
The Sokolova Cave fauna is unique and is the
|
||
<typeStatus id="54FE8832FFD4FFB2E852FF6491D045CC" box="[860,908,151,177]" pageId="8" pageNumber="494">type</typeStatus>
|
||
locality for two troglobiont species of ground beetles (
|
||
<taxonomicName id="4C454D13FFD4FFB2EBF6FF48931145A8" authorityName="Linnaeus" authorityYear="1758" box="[248,333,187,213]" class="Insecta" kingdom="Animalia" pageId="8" pageNumber="494" phylum="Arthropoda" rank="class">Insecta</taxonomicName>
|
||
:
|
||
<taxonomicName id="4C454D13FFD4FFB2EA55FF48938D45A8" authorityName="Latreille" authorityYear="1802" box="[347,465,187,213]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
|
||
),
|
||
<taxonomicName id="4C454D13FFD4FFB2EAE6FF48965C45A8" authority="Belousov, 1999" authorityName="Belousov" authorityYear="1999" box="[488,1024,187,213]" class="Insecta" family="Carabidae" genus="Caucasaphaenops" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="molchanovi">
|
||
<emphasis id="B931EA82FFD4FFB2EAE6FF48911845A8" box="[488,836,187,213]" italics="true" pageId="8" pageNumber="494">Caucasaphaenops molchanovi</emphasis>
|
||
<bibRefCitation id="EFD44B61FFD4FFB2E840FF48965C45A8" author="Belousov, I. A." box="[846,1024,187,213]" pageId="8" pageNumber="494" pagination="165 - 182" refId="ref3238" refString="Belousov, I. A. (1999) A new genus and species of cave dwelling trechine beetles from the West Caucasus (Coleoptera Carabidae Trechini). In: Zamotajlov, A. & Sciaky, R., (Eds.), Advances in Carabidology. Papers dedicated to the memory of Prof. Dr. Oleg L. Kryzhanovskij. MUISO Publishers, Krasnodar, pp. 165 - 182." type="book chapter" year="1999">Belousov, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C454D13FFD4FFB2EF31FF4893C64584" authority="Belousov, 1999" authorityName="Belousov" authorityYear="1999" class="Insecta" family="Carabidae" genus="Caucasorites" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="494" phylum="Arthropoda" rank="species" species="kovali">
|
||
<emphasis id="B931EA82FFD4FFB2EF31FF48977E45A8" box="[1087,1314,187,213]" italics="true" pageId="8" pageNumber="494">Caucasorites kovali</emphasis>
|
||
<bibRefCitation id="EFD44B61FFD4FFB2EE23FF4892934584" author="Belousov, I. A." pageId="8" pageNumber="494" pagination="165 - 182" refId="ref3238" refString="Belousov, I. A. (1999) A new genus and species of cave dwelling trechine beetles from the West Caucasus (Coleoptera Carabidae Trechini). In: Zamotajlov, A. & Sciaky, R., (Eds.), Advances in Carabidology. Papers dedicated to the memory of Prof. Dr. Oleg L. Kryzhanovskij. MUISO Publishers, Krasnodar, pp. 165 - 182." type="book chapter" year="1999">Belousov, 1999</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="EFD44B61FFD4FFB2EBD0FF2C93CD4584" author="Belousov, I. A." box="[222,401,223,249]" pageId="8" pageNumber="494" pagination="165 - 182" refId="ref3238" refString="Belousov, I. A. (1999) A new genus and species of cave dwelling trechine beetles from the West Caucasus (Coleoptera Carabidae Trechini). In: Zamotajlov, A. & Sciaky, R., (Eds.), Advances in Carabidology. Papers dedicated to the memory of Prof. Dr. Oleg L. Kryzhanovskij. MUISO Publishers, Krasnodar, pp. 165 - 182." type="book chapter" year="1999">Belousov, 1999</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFA3690FFD4FFB2EBC9FEF191424460" blockId="8.[151,1436,151,285]" box="[199,798,258,285]" pageId="8" pageNumber="494">
|
||
<emphasis id="B931EA82FFD4FFB2EBC9FEF193054461" bold="true" box="[199,345,258,284]" pageId="8" pageNumber="494">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F3527600FFD4FFB2EA69FEF093EA4460" box="[359,438,259,285]" name="Russia" pageId="8" pageNumber="494">Russia</collectingCountry>
|
||
: the
|
||
<typeStatus id="54FE8832FFD4FFB2EAE4FEF090464460" box="[490,538,259,285]" pageId="8" pageNumber="494">type</typeStatus>
|
||
locality only (
|
||
<figureCitation id="137E2A15FFD4FFB2E9B3FEF0914D4460" box="[701,785,259,285]" captionStart="FIGURES 13–18" captionStartId="7.[151,264,1908,1932]" captionTargetBox="[164,1419,181,1882]" captionTargetId="figure-22@7.[164,1423,181,1884]" captionTargetPageId="7" captionText="FIGURES 13–18. Type locality of Kovalius logunovi sp. nov. and specimen in situ: 13—entrance to the Sokolova Cave; 14- 16 habitat in type locality: 14—on the wall at the entrance nearby the lake; 15—16 wall with seeping water and stalagmite; 17—cave river (13—15, photo: R. Vargovitsh; 16—17, photo: A. Koval); 18—waterfall and river within the cave (photo: R. Vargovitsh)." figureDoi="http://doi.org/10.5281/zenodo.7518998" httpUri="https://zenodo.org/record/7518998/files/figure.png" pageId="8" pageNumber="494">Fig. 17</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |