425 lines
57 KiB
XML
425 lines
57 KiB
XML
<document id="646D77315DEA235706EFFC7680EB1AF9" ID-DOI="10.1080/00222930802109140" ID-ISSN="1464-5262" ID-Zenodo-Dep="5219374" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629340522496" checkinUser="felipe" docAuthor="Berning, Björn, Tilbrook, Kevin J. & Rosso, Antonietta" docDate="2008" docId="0395A723FFBEFFE3FE35FC61FC4FEB31" docLanguage="en" docName="JNatHist.42.21-22.1509-1547.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 42 (21 - 22)" docSource="http://dx.doi.org/10.1080/00222930802109140" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Therenia cryptooecium Berning & Tilbrook & Rosso 2008, new species" docType="treatment" docUuid="165C1F52-B31E-45A3-8CEC-37D3F4AF06D8" docUuidSource="ZooBank" docVersion="3" lastPageNumber="1533" masterDocId="FFACDF5BFFABFFF9FF99FFD2FF8CEA13" masterDocTitle="Revision of the north-eastern Atlantic and Mediterranean species of the genera Herentia and Therenia (Bryozoa: Cheilostomata)" masterLastPageNumber="1547" masterPageNumber="1509" pageNumber="1528" updateTime="1699068555203" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A912BF0562F118D8B70161A9932DE1D1">Revision of the north-eastern Atlantic and Mediterranean species of the genera Herentia and Therenia (Bryozoa: Cheilostomata)</mods:title>
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<mods:namePart id="697CB0BB5E1CA62B2F4305A2279728FC">Berning, Björn</mods:namePart>
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<mods:namePart id="1FF026DA5C1175CD8862011DCD95DEA5">Tilbrook, Kevin J.</mods:namePart>
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<mods:roleTerm id="A01111B1067056C307F8315840EA179D">Author</mods:roleTerm>
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<mods:namePart id="3C7D2C7A589476EBB0FE55F6F8CAF01C">Rosso, Antonietta</mods:namePart>
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<mods:title id="66F648483C4AA949059D3A1F9EAE1F36">Journal of Natural History</mods:title>
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<mods:part id="6437C0503F994797C8D9F1AF3EF0448A">
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<mods:date id="6A70FD5D14930076B5C9E4AC1FF99867">2008</mods:date>
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<mods:title id="E0B99B8CDC95E914C511E4C51A09793E">J. Nat. Hist.</mods:title>
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<mods:number id="9A87E198E9471473A869FA232FD0C93C">2008-06-30</mods:number>
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<mods:number id="9378AA27F06A0BB201E99474536B2EE6">42</mods:number>
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<mods:number id="85008A4A86991E15D436E1ABA628E4D1">21 - 22</mods:number>
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<mods:start id="4D054B977A2ACD9990D34E34AEAA4D37">1509</mods:start>
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<treatment id="0395A723FFBEFFE3FE35FC61FC4FEB31" LSID="urn:lsid:zoobank.org:act:165C1F52-B31E-45A3-8CEC-37D3F4AF06D8" httpUri="http://treatment.plazi.org/id/0395A723FFBEFFE3FE35FC61FC4FEB31" lastPageId="26" lastPageNumber="1533" pageId="21" pageNumber="1528">
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<subSubSection id="C32645BEFFBEFFECFE35FC61FCB4E9D8" box="[428,824,947,971]" pageId="21" pageNumber="1528" type="nomenclature">
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<paragraph id="8B831635FFBEFFECFE35FC61FCB4E9D8" blockId="21.[146,831,947,1037]" box="[428,824,947,971]" pageId="21" pageNumber="1528">
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<heading id="D0CBA159FFBEFFECFE35FC61FCB4E9D8" box="[428,824,947,971]" centered="true" fontSize="36" level="2" pageId="21" pageNumber="1528" reason="7">
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<taxonomicName id="4C3C6DB6FFBEFFECFE35FC61FD27E9D8" ID-CoL="56CWS" authority="Berning & Tilbrook & Rosso, 2008" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[428,683,947,971]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1528" phylum="Bryozoa" rank="species" species="cryptooecium" status="sp. nov.">
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<emphasis id="B948CA27FFBEFFECFE35FC61FD27E9D8" bold="true" box="[428,683,947,971]" italics="true" pageId="21" pageNumber="1528">Therenia cryptooecium</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27B775CFFBEFFECFD2DFC61FCB4E9D8" box="[692,824,947,971]" pageId="21" pageNumber="1528" rank="species">new species</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32645BEFFBEFFECFDA0FC06FD2BE9FF" box="[569,679,980,1004]" pageId="21" pageNumber="1528" type="description">
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<paragraph id="8B831635FFBEFFECFDA0FC06FD2BE9FF" blockId="21.[146,831,947,1037]" box="[569,679,980,1004]" pageId="21" pageNumber="1528">
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(
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<figureCitation id="13070AB0FFBEFFECFDD9FC06FD13E9FF" box="[576,671,980,1004]" captionStart="Figure 5" captionStartId="22.[213,280,918,940]" captionTargetBox="[213,1170,165,882]" captionTargetId="figure-150@22.[213,1170,165,882]" captionTargetPageId="22" captionText="Figure 5. Therenia cryptooecium n. sp. Holotype, NHM 1972.1.2.16A, Ghana, off Tema. (A) General aspect of colony; note the presence of three ovicellate zooecia (arrows); (B) autozooecia at the colony margin; note differences in condyle length between orifices; (C) close-up of autozooecium orifice and avicularium; (D) close-up of ovicellate zooecium, which is characterized by a slightly larger aperture and shorter condyles. Scale bars: 200 mm (A); 100 mm (B, D); 50 mm (C)." figureDoi="http://doi.org/10.5281/zenodo.5219384" httpUri="https://zenodo.org/record/5219384/files/figure.png" pageId="21" pageNumber="1528">Figure 5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32645BEFFBEFFECFF0BFC27FCB3EE1E" box="[146,831,1013,1037]" pageId="21" pageNumber="1528" type="reference_group">
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<paragraph id="8B831635FFBEFFECFF0BFC27FCB3EE1E" blockId="21.[146,831,947,1037]" box="[146,831,1013,1037]" pageId="21" pageNumber="1528">
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<treatmentCitationGroup id="AB2C311BFFBEFFECFF0BFC27FCB3EE1E" box="[146,831,1013,1037]" pageId="21" pageNumber="1528">
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<treatmentCitation id="0A9D3024FFBEFFECFF0BFC27FDBAEE1E" author="Cook PL" box="[146,566,1013,1037]" page="165" pageId="21" pageNumber="1528" year="1985">
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<taxonomicName id="4C3C6DB6FFBEFFECFF0BFC27FDBAEE1E" ID-CoL="3BGNV" authority=": Cook 1985, p. 165" authorityName=": Cook" authorityPageNumber="165" authorityYear="1985" box="[146,566,1013,1037]" class="Gymnolaemata" family="Escharinidae" genus="Escharina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1528" phylum="Bryozoa" rank="species" species="porosa">
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<emphasis id="B948CA27FFBEFFECFF0BFC27FEDFEE1E" box="[146,339,1013,1037]" italics="true" pageId="21" pageNumber="1528">Escharina porosa</emphasis>
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:
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<bibRefCitation id="EFAD6BC4FFBEFFECFEFAFC27FE6FEE1E" author="Cook PL" box="[355,483,1013,1037]" pageId="21" pageNumber="1528" pagination="1 - 315" refId="ref19142" refString="Cook PL. 1985. Bryozoa from Ghana, a preliminary survey. Koninklijk Museum voor Midden-Afrika (Tervuren, Belgie). Zoolog Wetenschappen - Ann. 238: 1 - 315." type="journal article" year="1985">Cook 1985</bibRefCitation>
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, p. 165
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</taxonomicName>
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</treatmentCitation>
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, Plate 16, Figures E, F.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32645BEFFBEFFEEFF0BFB94FD25EF54" lastPageId="23" lastPageNumber="1530" pageId="21" pageNumber="1528" type="diagnosis">
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<paragraph id="8B831635FFBEFFECFF0BFB94FE0DEE4D" blockId="21.[146,385,1094,1118]" box="[146,385,1094,1118]" pageId="21" pageNumber="1528">
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<emphasis id="B948CA27FFBEFFECFF0BFB94FE0DEE4D" box="[146,385,1094,1118]" italics="true" pageId="21" pageNumber="1528">Differential diagnosis</emphasis>
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</paragraph>
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<paragraph id="8B831635FFBEFFECFF0BFBBDFC42EEB4" blockId="21.[146,1103,1134,1581]" pageId="21" pageNumber="1528">
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<taxonomicName id="4C3C6DB6FFBEFFECFF0BFBBDFEC9EE94" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[146,325,1135,1159]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1528" phylum="Bryozoa" rank="species" species="cryptooecium">
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<emphasis id="B948CA27FFBEFFECFF0BFBBDFEC9EE94" box="[146,325,1135,1159]" italics="true" pageId="21" pageNumber="1528">T. cryptooecium</emphasis>
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</taxonomicName>
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differs from all other species in the extremely dense perforation of its frontal shield and in its ooecium, which is almost completely immersed.
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</paragraph>
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<paragraph id="8B831635FFBEFFECFF24FB62FF61EF3A" blockId="21.[146,1103,1134,1581]" pageId="21" pageNumber="1528">
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It is furthermore distinguished from
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<taxonomicName id="4C3C6DB6FFBEFFECFDE5FB62FD61EEDB" authorityName=": Cook" authorityYear="1985" box="[636,749,1200,1224]" class="Gymnolaemata" family="Escharinidae" genus="Escharina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1528" phylum="Bryozoa" rank="species" species="porosa">
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<emphasis id="B948CA27FFBEFFECFDE5FB62FD61EEDB" box="[636,749,1200,1224]" italics="true" pageId="21" pageNumber="1528">T. porosa</emphasis>
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</taxonomicName>
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in its wider sinus and in the thickened rim framing only the distolateral orifice margin in autozooecia (
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<taxonomicName id="4C3C6DB6FFBEFFECFBACFB02FF52EF1A" authorityName=": Cook" authorityYear="1985" class="Gymnolaemata" family="Escharinidae" genus="Escharina" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1528" phylum="Bryozoa" rank="species" species="porosa">
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<emphasis id="B948CA27FFBEFFECFBACFB02FF52EF1A" italics="true" pageId="21" pageNumber="1528">T. porosa</emphasis>
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</taxonomicName>
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has a narrower and distally raised peristome that completely encircles the orifice).
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</paragraph>
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<paragraph id="8B831635FFBEFFECFF24FAE0FDFFEF98" blockId="21.[146,1103,1134,1581]" pageId="21" pageNumber="1528">
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Besides the differences given above,
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<taxonomicName id="4C3C6DB6FFBEFFECFDF6FAE0FCABEF59" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[623,807,1330,1354]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1528" phylum="Bryozoa" rank="species" species="cryptooecium">
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<emphasis id="B948CA27FFBEFFECFDF6FAE0FCABEF59" box="[623,807,1330,1354]" italics="true" pageId="21" pageNumber="1528">T. cryptooecium</emphasis>
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</taxonomicName>
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is distinguished from
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<taxonomicName id="4C3C6DB6FFBEFFECFBAFFAE0FF4BEF78" authority="Berning & Tilbrook & Rosso, 2008" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1528" phylum="Bryozoa" rank="species" species="rosei" status="sp. nov.">
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<emphasis id="B948CA27FFBEFFECFBAFFAE0FF4BEF78" italics="true" pageId="21" pageNumber="1528">T. rosei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27B775CFFBEFFECFF4BFA80FE9FEF79" box="[210,275,1362,1386]" pageId="21" pageNumber="1528" rank="species">n. sp.</taxonomicNameLabel>
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due to its wider sinus in ovicellate zooecia, and its narrower band of calcification framing the ooecial aperture.
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</paragraph>
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<paragraph id="8B831635FFBEFFECFF24FA41FC87EC3E" blockId="21.[146,1103,1134,1581]" pageId="21" pageNumber="1528">
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In contrast to
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<taxonomicName id="4C3C6DB6FFBEFFECFEFAFA46FD84EFBF" authority="Berning & Tilbrook & Rosso, 2008" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[355,520,1428,1452]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="21" pageNumber="1528" phylum="Bryozoa" rank="species" species="peristomata" status="sp. nov.">
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<emphasis id="B948CA27FFBEFFECFEFAFA46FD84EFBF" box="[355,520,1428,1452]" italics="true" pageId="21" pageNumber="1528">T. peristomata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27B775CFFBEFFECFD88FA41FDC3EFB8" box="[529,591,1427,1451]" pageId="21" pageNumber="1528" rank="species">n. sp.</taxonomicNameLabel>
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it has a flat frontal surface, a thickened rim merely framing the distolateral orifice margin, condyles that project medially beyond the shoulders of the sinus, a very narrow band of calcification framing the distolateral ooecial aperture, an avicularium with a much broader and kidneyshaped rostrum, and a crossbar that lacks a columella.
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</paragraph>
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<paragraph id="8B831635FFBEFFECFF0BF9B4FE87EC6D" blockId="21.[146,267,1638,1662]" box="[146,267,1638,1662]" pageId="21" pageNumber="1528">
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<emphasis id="B948CA27FFBEFFECFF0BF9B4FE87EC6D" box="[146,267,1638,1662]" italics="true" pageId="21" pageNumber="1528">Etymology</emphasis>
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</paragraph>
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<paragraph id="8B831635FFBEFFECFF0BF95DFDE0ECB4" blockId="21.[146,620,1679,1703]" box="[146,620,1679,1703]" pageId="21" pageNumber="1528">
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For its hidden (Greek:
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<emphasis id="B948CA27FFBEFFECFE07F95DFE7DECB4" box="[414,497,1679,1703]" italics="true" pageId="21" pageNumber="1528">kryptos</emphasis>
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) ooecium.
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</paragraph>
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<caption id="DF4346BDFFBDFFEFFF4CFC44FD98EE2C" ID-DOI="http://doi.org/10.5281/zenodo.5219384" ID-Zenodo-Dep="5219384" httpUri="https://zenodo.org/record/5219384/files/figure.png" pageId="22" pageNumber="1529" startId="22.[213,280,918,940]" targetBox="[213,1170,165,882]" targetPageId="22">
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<paragraph id="8B831635FFBDFFEFFF4CFC44FD98EE2C" blockId="22.[213,1170,918,1087]" pageId="22" pageNumber="1529">
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Figure 5.
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<taxonomicName id="4C3C6DB6FFBDFFEFFED8FC45FDA6E9BF" authority="Berning & Tilbrook & Rosso, 2008" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[321,554,919,940]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="22" pageNumber="1529" phylum="Bryozoa" rank="species" species="cryptooecium" status="sp. nov.">
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||
<emphasis id="B948CA27FFBDFFEFFED8FC45FDA6E9BF" box="[321,554,919,940]" italics="true" pageId="22" pageNumber="1529">Therenia cryptooecium</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A27B775CFFBDFFEFFDAFFC44FDFEE9BF" box="[566,626,918,940]" pageId="22" pageNumber="1529" rank="species">n. sp.</taxonomicNameLabel>
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||
Holotype, NHM 1972.1.2.16A, Ghana, off Tema. (A) General aspect of colony; note the presence of three ovicellate zooecia (arrows); (B) autozooecia at the colony margin; note differences in condyle length between orifices; (C) close-up of autozooecium orifice and avicularium; (D) close-up of ovicellate zooecium, which is characterized by a slightly larger aperture and shorter condyles. Scale bars: 200 mm (A); 100 mm (B, D); 50 mm (C).
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</paragraph>
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</caption>
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<paragraph id="8B831635FFBDFFEFFF4CFBA2FE25EE9B" blockId="22.[213,425,1136,1160]" box="[213,425,1136,1160]" pageId="22" pageNumber="1529">
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||
<emphasis id="B948CA27FFBDFFEFFF4CFBA2FE25EE9B" box="[213,425,1136,1160]" italics="true" pageId="22" pageNumber="1529">Material examined</emphasis>
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||
</paragraph>
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<paragraph id="8B831635FFBDFFEFFF4CFB4AFCF9EEE2" blockId="22.[213,1169,1176,1265]" pageId="22" pageNumber="1529">
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||
<materialsCitation id="3B541C68FFBDFFEFFF4CFB4AFEE0EEC2" collectorName="Plate & Figures E" country="Ghana" location="Ghana" pageId="22" pageNumber="1529" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5487A897FFBDFFEFFF4CFB4AFEC8EEA3" box="[213,324,1176,1200]" pageId="22" pageNumber="1529" type="holotype">Holotype</typeStatus>
|
||
: NHM 1972.1.2.16A, off Tema,
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<collectingCountry id="F32B56A5FFBDFFEFFD27FB4AFC83EEA3" box="[702,783,1176,1200]" name="Ghana" pageId="22" pageNumber="1529">Ghana</collectingCountry>
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, figured by
|
||
<bibRefCitation id="EFAD6BC4FFBDFFEFFC02FB4AFBA8EEA3" author="Cook PL" box="[923,1060,1176,1200]" pageId="22" pageNumber="1529" pagination="1 - 315" refId="ref19142" refString="Cook PL. 1985. Bryozoa from Ghana, a preliminary survey. Koninklijk Museum voor Midden-Afrika (Tervuren, Belgie). Zoolog Wetenschappen - Ann. 238: 1 - 315." type="journal article" year="1985">Cook (1985</bibRefCitation>
|
||
,
|
||
<collectorName id="26C973E3FFBDFFEFFBABFB4AFBE7EEA3" box="[1074,1131,1176,1200]" pageId="22" pageNumber="1529">Plate</collectorName>
|
||
16,
|
||
<collectorName id="26C973E3FFBDFFEFFF4CFB6BFECEEEC2" box="[213,322,1209,1233]" pageId="22" pageNumber="1529">Figures E</collectorName>
|
||
, F)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B541C68FFBDFFEFFEEEFB6BFB83EEC2" box="[375,1039,1209,1233]" country="Ghana" elevation="35" location="Tema" pageId="22" pageNumber="1529" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="5487A897FFBDFFEFFEEEFB6BFE6CEEC2" box="[375,480,1209,1233]" pageId="22" pageNumber="1529" type="paratype">Paratype</typeStatus>
|
||
: NHM 1972.1.2.16B, off
|
||
<location id="8EE340EEFFBDFFEFFC99FB6BFCC8EEC2" LSID="urn:lsid:plazi:treatment:0395A723FFBEFFE3FE35FC61FC4FEB31:8EE340EEFFBDFFEFFC99FB6BFCC8EEC2" box="[768,836,1209,1233]" country="Ghana" name="Tema" pageId="22" pageNumber="1529">Tema</location>
|
||
,
|
||
<collectingCountry id="F32B56A5FFBDFFEFFCD7FB6BFC13EEC2" box="[846,927,1209,1233]" name="Ghana" pageId="22" pageNumber="1529">Ghana</collectingCountry>
|
||
,
|
||
<quantity id="4CC4BBD0FFBDFFEFFC32FB6BFB87EEC2" box="[939,1035,1209,1233]" metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="22" pageNumber="1529" unit="m" value="35.0" valueMax="40.0" valueMin="30.0">
|
||
<elevation id="0011F106FFBDFFEFFC32FB6BFB87EEC2" box="[939,1035,1209,1233]" metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="22" pageNumber="1529" unit="m" value="35.0" valueMax="40.0" valueMin="30.0">30–40 m</elevation>
|
||
</quantity>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="3B541C68FFBDFFEFFB8FFB6BFCF9EEE2" country="Ghana" elevation="35" location="Tema" pageId="22" pageNumber="1529" specimenCount="1">
|
||
Additional material: NHM 1972.1.2.16pt, off
|
||
<location id="8EE340EEFFBDFFEFFDFBFB0BFD2AEEE2" LSID="urn:lsid:plazi:treatment:0395A723FFBEFFE3FE35FC61FC4FEB31:8EE340EEFFBDFFEFFDFBFB0BFD2AEEE2" box="[610,678,1241,1265]" country="Ghana" name="Tema" pageId="22" pageNumber="1529">Tema</location>
|
||
,
|
||
<collectingCountry id="F32B56A5FFBDFFEFFD2AFB0BFC8FEEE2" box="[691,771,1241,1265]" name="Ghana" pageId="22" pageNumber="1529">Ghana</collectingCountry>
|
||
,
|
||
<quantity id="4CC4BBD0FFBDFFEFFC88FB0BFCFDEEE2" box="[785,881,1241,1265]" metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="22" pageNumber="1529" unit="m" value="35.0" valueMax="40.0" valueMin="30.0">
|
||
<elevation id="0011F106FFBDFFEFFC88FB0BFCFDEEE2" box="[785,881,1241,1265]" metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="22" pageNumber="1529" unit="m" value="35.0" valueMax="40.0" valueMin="30.0">30–40 m</elevation>
|
||
</quantity>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B831635FFBDFFEFFF4CFAF8FEFAEF51" blockId="22.[213,374,1322,1346]" box="[213,374,1322,1346]" pageId="22" pageNumber="1529">
|
||
<emphasis id="B948CA27FFBDFFEFFF4CFAF8FEFAEF51" box="[213,374,1322,1346]" italics="true" pageId="22" pageNumber="1529">Measurements</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B831635FFBDFFEFFF4CFA81FB1DEFB8" blockId="22.[213,1169,1362,1452]" pageId="22" pageNumber="1529">ZL 620¡63, 548–796 (1, 14); ZW 480¡65, 378–624 (1, 14); OL 136¡9, 119–149 (1, 17); OW 140¡7, 127–152 (1, 17); ApL 157, 155–159 (1, 3); ApW 159, 153–163 (1, 3); AL 142¡13, 115–167 (1, 16); AW 122¡14, 106–147 (1, 16); PD 20¡2, 16–23 (1, 12).</paragraph>
|
||
<paragraph id="8B831635FFBDFFEFFF4CFA37FED8EFEE" blockId="22.[213,340,1509,1533]" box="[213,340,1509,1533]" pageId="22" pageNumber="1529">
|
||
<emphasis id="B948CA27FFBDFFEFFF4CFA37FED8EFEE" box="[213,340,1509,1533]" italics="true" pageId="22" pageNumber="1529">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B831635FFBDFFEEFF4CF9DFFCCBEBF7" blockId="22.[213,1169,1549,1703]" lastBlockId="23.[146,1103,168,810]" lastPageId="23" lastPageNumber="1530" pageId="22" pageNumber="1529">Colony encrusting unilaminar, multiserial. Zooecia rhomboid, hexagonal or polygonal, separated by shallow grooves or a slightly raised suture; frontal wall flat, closely and regularly perforated by numerous small pseudopores, and with few (two to five) large marginal areolar pores, one of these situated directly proximolateral to orifice margin opposite to avicularium. Orifice dimorphic with a deeply immersed shelf along the distolateral margin, proximolaterally framed by a smooth narrow band of gymnocystal calcification; primary orifice about as wide as long and widest just distal to mid-distance, anter three-quarters of a transverse ellipse, proximal border straight, sinus U-shaped with rather square shoulders, occupying about half of proximal margin, two slightly immersed, thick, straight and tapering condyles pointing medially, projecting beyond the shoulders of the sinus and occasionally approaching each other, distolateral orifice border bounded by a raised and curved rim of thick calcification sloping towards orifice; aperture in ovicellate zooecia of similar shape but larger, condyles shorter, not projecting beyond sinus, distolateral margin relatively narrow and flat.</paragraph>
|
||
<paragraph id="8B831635FFBCFFEEFF24FE3FFBC3E816" blockId="23.[146,1103,168,810]" box="[189,1103,493,517]" pageId="23" pageNumber="1530">Ooecium completely immersed by distal zooecium, not visible at colony surface.</paragraph>
|
||
<paragraph id="8B831635FFBCFFEEFF24FDDCFCCEE91A" blockId="23.[146,1103,168,810]" pageId="23" pageNumber="1530">Avicularium situated directly proximolateral to orifice on either left or right side, in distal pseudoporous chamber, which is level with, and often not clearly demarcated from, frontal shield of zooecium apart from one or two areolar pores at lateral and/or proximolateral margin; rostrum a suborbicular to kidney-shaped, broad, sloping band, laterally or distolaterally oriented; crossbar complete, without columella, thick, extending fairly deep into cystid, square-edged and twisted along its axis; proximal opesium bounded by an immersed convex shelf of calcification extending towards crossbar from proximal rostrum margin.</paragraph>
|
||
<paragraph id="8B831635FFBCFFEEFF24FCC0FBC8E939" blockId="23.[146,1103,168,810]" box="[189,1092,786,810]" pageId="23" pageNumber="1530">An ancestrula or early astogenetic stages of colony growth were not observed.</paragraph>
|
||
<paragraph id="8B831635FFBCFFEEFF0BFCA2FF79E99B" blockId="23.[146,245,880,904]" box="[146,245,880,904]" pageId="23" pageNumber="1530">
|
||
<emphasis id="B948CA27FFBCFFEEFF0BFCA2FF79E99B" box="[146,245,880,904]" italics="true" pageId="23" pageNumber="1530">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B831635FFBCFFEEFF0BFC4BFD3CEE40" blockId="23.[146,1103,921,1237]" pageId="23" pageNumber="1530">
|
||
The type material of
|
||
<taxonomicName id="4C3C6DB6FFBCFFEEFE1CFC4BFDB6E9A2" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[389,570,921,945]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="1530" phylum="Bryozoa" rank="species" species="cryptooecium">
|
||
<emphasis id="B948CA27FFBCFFEEFE1CFC4BFDB6E9A2" box="[389,570,921,945]" italics="true" pageId="23" pageNumber="1530">T. cryptooecium</emphasis>
|
||
</taxonomicName>
|
||
is from
|
||
<collectingCountry id="F32B56A5FFBCFFEEFD05FC4BFD66E9A2" box="[668,746,921,945]" name="Ghana" pageId="23" pageNumber="1530">Ghana</collectingCountry>
|
||
and was described and figured by
|
||
<bibRefCitation id="EFAD6BC4FFBCFFEEFF21FC6BFECFE9C2" author="Cook PL" box="[184,323,953,977]" pageId="23" pageNumber="1530" pagination="1 - 315" refId="ref19142" refString="Cook PL. 1985. Bryozoa from Ghana, a preliminary survey. Koninklijk Museum voor Midden-Afrika (Tervuren, Belgie). Zoolog Wetenschappen - Ann. 238: 1 - 315." type="journal article" year="1985">Cook (1985</bibRefCitation>
|
||
, p. 165, Plate 16, Figures E, F). Whether any or all of the material reported as
|
||
<taxonomicName id="4C3C6DB6FFBCFFEEFE80FC08FE0FE9E1" authorityName=": Cook" authorityYear="1985" box="[281,387,986,1010]" class="Gymnolaemata" family="Escharinidae" genus="Escharina" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="1530" phylum="Bryozoa" rank="species" species="porosa">
|
||
<emphasis id="B948CA27FFBCFFEEFE80FC08FE0FE9E1" box="[281,387,986,1010]" italics="true" pageId="23" pageNumber="1530">T. porosa</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFAD6BC4FFBCFFEEFE37FC0BFDB9E9E2" author="Cook PL" box="[430,565,985,1009]" pageId="23" pageNumber="1530" pagination="115 - 262" refId="ref19079" refString="Cook PL. 1968. Bryozoa (Polyzoa) from the coast of tropical West Africa. Atlantide Report 10: 115 - 262." type="book chapter" year="1968">Cook (1968</bibRefCitation>
|
||
, p. 195) from
|
||
<collectingCountry id="F32B56A5FFBCFFEEFD4BFC0BFCA9E9E2" box="[722,805,985,1009]" name="Guinea-Bissau" pageId="23" pageNumber="1530">Guinea</collectingCountry>
|
||
and the Canary Islands is conspecific with
|
||
<taxonomicName id="4C3C6DB6FFBCFFEEFEC0FC28FD9EEE01" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[345,530,1018,1042]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="1530" phylum="Bryozoa" rank="species" species="cryptooecium">
|
||
<emphasis id="B948CA27FFBCFFEEFEC0FC28FD9EEE01" box="[345,530,1018,1042]" italics="true" pageId="23" pageNumber="1530">T. cryptooecium</emphasis>
|
||
</taxonomicName>
|
||
or the other species introduced here cannot be proven at present because the respective material is housed in different collections, which could not be studied during this project.
|
||
</paragraph>
|
||
<paragraph id="8B831635FFBCFFEEFF24FB8EFE6DEEC6" blockId="23.[146,1103,921,1237]" pageId="23" pageNumber="1530">
|
||
<bibRefCitation id="EFAD6BC4FFBCFFEEFF24FB8EFEDCEE67" author="Cook PL" box="[189,336,1116,1140]" pageId="23" pageNumber="1530" pagination="1 - 315" refId="ref19142" refString="Cook PL. 1985. Bryozoa from Ghana, a preliminary survey. Koninklijk Museum voor Midden-Afrika (Tervuren, Belgie). Zoolog Wetenschappen - Ann. 238: 1 - 315." type="journal article" year="1985">Cook (1985</bibRefCitation>
|
||
, p. 166) frequently observed
|
||
<taxonomicName id="4C3C6DB6FFBCFFEEFD5EFB8EFC0BEE67" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[711,903,1116,1140]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="1530" phylum="Bryozoa" rank="species" species="cryptooecium">
|
||
<emphasis id="B948CA27FFBCFFEEFD5EFB8EFC0BEE67" box="[711,903,1116,1140]" italics="true" pageId="23" pageNumber="1530">T. cryptooecium</emphasis>
|
||
</taxonomicName>
|
||
to encrust the agglutinating larger foraminifer
|
||
<taxonomicName id="4C3C6DB6FFBCFFEEFDBEFBAEFC27EE87" authority="Schlumberger" authorityName="Schlumberger" authorityYear="1890" box="[551,939,1148,1172]" class="Monothalamea" family="Schizamminidae" genus="Jullienella" kingdom="Chromista" order="Astrorhizida" pageId="23" pageNumber="1530" phylum="Foraminifera" rank="species" species="foetida">
|
||
<emphasis id="B948CA27FFBCFFEEFDBEFBAEFD7BEE87" box="[551,759,1148,1172]" italics="true" pageId="23" pageNumber="1530">Jullienella foetida</emphasis>
|
||
Schlumberger
|
||
</taxonomicName>
|
||
, occasionally reaching a colony size of over 1000 zooids. The depth of occurrence off
|
||
<collectingCountry id="F32B56A5FFBCFFEEFB98FB4EFBC3EEA7" box="[1025,1103,1180,1204]" name="Ghana" pageId="23" pageNumber="1530">Ghana</collectingCountry>
|
||
ranged between 10 and
|
||
<specimenCount id="9D3ADDBCFFBCFFEEFE3CFB6FFE6DEEC6" box="[421,481,1213,1237]" pageId="23" pageNumber="1530" type="generic">
|
||
<quantity id="4CC4BBD0FFBCFFEEFE3CFB6FFE51EEC6" box="[421,477,1213,1237]" metricMagnitude="1" metricUnit="m" metricValue="5.1" pageId="23" pageNumber="1530" unit="m" value="51.0">51 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
<paragraph id="8B831635FFBCFFEEFE57FADCFD76EF35" blockId="23.[146,1103,1294,1416]" box="[462,762,1294,1318]" pageId="23" pageNumber="1530">
|
||
<taxonomicName id="4C3C6DB6FFBCFFEEFE57FADCFDE2EF35" authority="Berning & Tilbrook & Rosso, 2008" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[462,622,1294,1318]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="1530" phylum="Bryozoa" rank="species" species="rosei" status="sp. nov.">
|
||
<emphasis id="B948CA27FFBCFFEEFE57FADCFDE2EF35" bold="true" box="[462,622,1294,1318]" italics="true" pageId="23" pageNumber="1530">Therenia rosei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27B775CFFBCFFEEFDEEFADCFD76EF35" box="[631,762,1294,1318]" pageId="23" pageNumber="1530" rank="species">new species</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8B831635FFBCFFEEFDA2FAFDFD25EF54" blockId="23.[146,1103,1294,1416]" box="[571,681,1327,1351]" pageId="23" pageNumber="1530">
|
||
(
|
||
<figureCitation id="13070AB0FFBCFFEEFDDAFAFDFD2DEF54" box="[579,673,1327,1351]" captionStart="Figure 6" captionStartId="24.[213,280,918,940]" captionTargetBox="[213,1170,165,882]" captionTargetId="figure-145@24.[213,1170,165,882]" captionTargetPageId="24" captionText="Figure 6. Therenia rosei n. sp. Holotype, NHM 1975.1.12.17A, Aegean Sea. (A) Autozooecia; (B) ovicellate zooecia; (C) close-up of autozooecium orifice and avicularium; (D) close-up of ovicellate zooecium; note difference in condyle morphology compared to orifice of autozooecium. Scale bars: 200 mm (A, B); 50 mm (C, D)." figureDoi="http://doi.org/10.5281/zenodo.5219386" httpUri="https://zenodo.org/record/5219386/files/figure.png" pageId="23" pageNumber="1530">Figure 6</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32645BEFFBCFFEEFF0BFA82FF7EEF9B" pageId="23" pageNumber="1530" type="reference_group">
|
||
<paragraph id="8B831635FFBCFFEEFF0BFA82FF7EEF9B" blockId="23.[146,1103,1294,1416]" pageId="23" pageNumber="1530">
|
||
<treatmentCitationGroup id="AB2C311BFFBCFFEEFF0BFA82FF7EEF9B" pageId="23" pageNumber="1530">
|
||
<treatmentCitation id="0A9D3024FFBCFFEEFF0BFA82FD0EEF74" author="Hayward PJ" box="[146,642,1359,1384]" page="377" pageId="23" pageNumber="1530" year="1974">
|
||
<taxonomicName id="4C3C6DB6FFBCFFEEFF0BFA82FD0EEF74" ID-CoL="3BGNV" authority=": Hayward 1974, p. 377" authorityName=": Hayward" authorityPageNumber="377" authorityYear="1974" box="[146,642,1359,1384]" class="Gymnolaemata" family="Escharinidae" genus="Escharina" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="1530" phylum="Bryozoa" rank="species" species="porosa">
|
||
<emphasis id="B948CA27FFBCFFEEFF0BFA82FED7EF7B" box="[146,347,1360,1384]" italics="true" pageId="23" pageNumber="1530">Escharina porosa</emphasis>
|
||
:
|
||
<bibRefCitation id="EFAD6BC4FFBCFFEEFEEDFA9DFDABEF74" author="Hayward PJ" box="[372,551,1359,1383]" pageId="23" pageNumber="1530" pagination="369 - 402" refId="ref19936" refString="Hayward PJ. 1974. Studies on the cheilostome bryozoan fauna of the Aegean island of Chios. J Nat Hist. 8: 369 - 402." type="journal article" year="1974">Hayward 1974</bibRefCitation>
|
||
, p. 377
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
,
|
||
<figureCitation id="13070AB0FFBCFFEEFD0EFA9DFCB8EF74" box="[663,820,1359,1383]" captionStart="Figure 4" captionStartId="17.[146,213,1596,1618]" captionTargetBox="[146,1103,842,1559]" captionTargetId="figure-144@17.[146,1103,842,1559]" captionTargetPageId="17" captionText="Figure 4. Herentia thalassae David and Pouyet, 1978. MNHN 7619, off NW Spain; note that the vertical component in these perpendicular views is greatly understated, the distal parts of the zooecia and ooecia are distinctly raised. (A) General aspect of colony; (B) close-up of ovicellate zooecium; (C) close-up of autozooecium. Scale bars: 400 mm (A); 100 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.5219382" httpUri="https://zenodo.org/record/5219382/files/figure.png" pageId="23" pageNumber="1530">Figure 4A, B</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFAD6BC4FFBCFFEEFCD3FA9DFC6BEF74" author="Rosso A" box="[842,999,1359,1383]" pageId="23" pageNumber="1530" pagination="189 - 225" refId="ref21479" refString="Rosso A. 1996 a. Popolamenti e tanatocenosi a briozoi di fondi mobili circalitorali del Golfo di Noto (Sicilia SE). Naturalista Siciliano (ser. 4) 20: 189 - 225." type="journal article" year="1996">Rosso 1996a</bibRefCitation>
|
||
, Plate 4, Figure f.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32645BEFFBCFFE0FF0BFA13FEB9ECB6" lastPageId="25" lastPageNumber="1532" pageId="23" pageNumber="1530" type="diagnosis">
|
||
<paragraph id="8B831635FFBCFFEEFF0BFA13FE0DEFCA" blockId="23.[146,385,1473,1497]" box="[146,385,1473,1497]" pageId="23" pageNumber="1530">
|
||
<emphasis id="B948CA27FFBCFFEEFF0BFA13FE0DEFCA" box="[146,385,1473,1497]" italics="true" pageId="23" pageNumber="1530">Differential diagnosis</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B831635FFBCFFEEFF0BFA38FC30EC51" blockId="23.[146,1103,1514,1700]" pageId="23" pageNumber="1530">
|
||
<taxonomicName id="4C3C6DB6FFBCFFEEFF0BFA38FF6BEC11" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[146,231,1514,1538]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="1530" phylum="Bryozoa" rank="species" species="rosei">
|
||
<emphasis id="B948CA27FFBCFFEEFF0BFA38FF6BEC11" box="[146,231,1514,1538]" italics="true" pageId="23" pageNumber="1530">T. rosei</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C3C6DB6FFBCFFEEFE79FA38FDC6EC11" authorityName=": Hayward" authorityYear="1974" box="[480,586,1514,1538]" class="Gymnolaemata" family="Escharinidae" genus="Escharina" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="1530" phylum="Bryozoa" rank="species" species="porosa">
|
||
<emphasis id="B948CA27FFBCFFEEFE79FA38FDC6EC11" box="[480,586,1514,1538]" italics="true" pageId="23" pageNumber="1530">T. porosa</emphasis>
|
||
</taxonomicName>
|
||
owing to its larger pores in the frontal wall, a wider sinus with rounded shoulders, the presence of a thickened rim framing the distolateral orifice margin, and in a broader area of exposed ooecium.
|
||
</paragraph>
|
||
<paragraph id="8B831635FFBCFFEEFF24F999FD88ECB7" blockId="23.[146,1103,1514,1700]" pageId="23" pageNumber="1530">
|
||
Differences to
|
||
<taxonomicName id="4C3C6DB6FFBCFFEEFEF0F99EFD93EC77" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[361,543,1612,1636]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="23" pageNumber="1530" phylum="Bryozoa" rank="species" species="cryptooecium">
|
||
<emphasis id="B948CA27FFBCFFEEFEF0F99EFD93EC77" box="[361,543,1612,1636]" italics="true" pageId="23" pageNumber="1530">T. cryptooecium</emphasis>
|
||
</taxonomicName>
|
||
are found in the less densely perforated frontal wall, in a narrower sinus with rounded shoulders, in crenellated condyles, and in the broad area of exposed ooecium.
|
||
</paragraph>
|
||
<caption id="DF4346BDFFB3FFE1FF4CFC44FC95EE17" ID-DOI="http://doi.org/10.5281/zenodo.5219386" ID-Zenodo-Dep="5219386" httpUri="https://zenodo.org/record/5219386/files/figure.png" pageId="24" pageNumber="1531" startId="24.[213,280,918,940]" targetBox="[213,1170,165,882]" targetPageId="24">
|
||
<paragraph id="8B831635FFB3FFE1FF4CFC44FC95EE17" blockId="24.[213,1169,918,1028]" pageId="24" pageNumber="1531">
|
||
Figure 6.
|
||
<taxonomicName id="4C3C6DB6FFB3FFE1FEA5FC45FE46E9BF" authority="Berning & Tilbrook & Rosso, 2008" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[316,458,919,940]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="24" pageNumber="1531" phylum="Bryozoa" rank="species" species="rosei" status="sp. nov.">
|
||
<emphasis id="B948CA27FFB3FFE1FEA5FC45FE46E9BF" box="[316,458,919,940]" italics="true" pageId="24" pageNumber="1531">Therenia rosei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27B775CFFB3FFE1FE48FC44FD84E9BF" box="[465,520,918,940]" pageId="24" pageNumber="1531" rank="species">n. sp.</taxonomicNameLabel>
|
||
Holotype, NHM 1975.1.12.17A, Aegean Sea. (A) Autozooecia; (B) ovicellate zooecia; (C) close-up of autozooecium orifice and avicularium; (D) close-up of ovicellate zooecium; note difference in condyle morphology compared to orifice of autozooecium. Scale bars: 200 mm (A, B); 50 mm (C, D).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B831635FFB3FFE1FF66FBEFFDB5EEC5" blockId="24.[213,1169,1085,1238]" pageId="24" pageNumber="1531">
|
||
In contrast to
|
||
<taxonomicName id="4C3C6DB6FFB3FFE1FE39FBEFFDCFEE46" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[416,579,1085,1109]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="24" pageNumber="1531" phylum="Bryozoa" rank="species" species="peristomata">
|
||
<emphasis id="B948CA27FFB3FFE1FE39FBEFFDCFEE46" box="[416,579,1085,1109]" italics="true" pageId="24" pageNumber="1531">T. peristomata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3C6DB6FFB3FFE1FDC8FBEFFD2AEE46" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[593,678,1085,1109]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="24" pageNumber="1531" phylum="Bryozoa" rank="species" species="rosei">
|
||
<emphasis id="B948CA27FFB3FFE1FDC8FBEFFD2AEE46" box="[593,678,1085,1109]" italics="true" pageId="24" pageNumber="1531">T. rosei</emphasis>
|
||
</taxonomicName>
|
||
has a more densely perforated frontal wall, a thick rim framing the distolateral orifice margin, a distinctly larger aperture and primary orifice with condyles extending beyond the shoulders of the sinus, as well as larger avicularia with a more extensive immersed shelf in the proximal opesium and a crossbar lacking the columella.
|
||
</paragraph>
|
||
<paragraph id="8B831635FFB3FFE1FF4CFAC2FEC2EF3B" blockId="24.[213,334,1296,1320]" box="[213,334,1296,1320]" pageId="24" pageNumber="1531">
|
||
<emphasis id="B948CA27FFB3FFE1FF4CFAC2FEC2EF3B" box="[213,334,1296,1320]" italics="true" pageId="24" pageNumber="1531">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B831635FFB3FFE1FF4CFAEAFBD8EF43" blockId="24.[213,1108,1336,1360]" box="[213,1108,1336,1360]" pageId="24" pageNumber="1531">Named in honour of Doris and Hans Rose, the first author’s aunt and uncle.</paragraph>
|
||
<paragraph id="8B831635FFB3FFE1FF4CFA5BFE25EFB2" blockId="24.[213,425,1417,1441]" box="[213,425,1417,1441]" pageId="24" pageNumber="1531">
|
||
<emphasis id="B948CA27FFB3FFE1FF4CFA5BFE25EFB2" box="[213,425,1417,1441]" italics="true" pageId="24" pageNumber="1531">Material examined</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B831635FFB3FFE1FF4CFA60FD45EC18" blockId="24.[213,1169,1458,1547]" pageId="24" pageNumber="1531">
|
||
<typeStatus id="5487A897FFB3FFE1FF4CFA60FEC8EFD9" box="[213,324,1458,1482]" pageId="24" pageNumber="1531" type="holotype">Holotype</typeStatus>
|
||
: NHM 1975.1.12.17A, Chios, Aegean Sea, figured by
|
||
<bibRefCitation id="EFAD6BC4FFB3FFE1FC4CFA60FB01EFD9" author="Hayward PJ" box="[981,1165,1458,1482]" pageId="24" pageNumber="1531" pagination="369 - 402" refId="ref19936" refString="Hayward PJ. 1974. Studies on the cheilostome bryozoan fauna of the Aegean island of Chios. J Nat Hist. 8: 369 - 402." type="journal article" year="1974">Hayward (1974</bibRefCitation>
|
||
,
|
||
<figureCitation id="13070AB0FFB3FFE1FF4CFA00FEE7EFF9" box="[213,363,1490,1514]" captionStart="Figure 4" captionStartId="17.[146,213,1596,1618]" captionTargetBox="[146,1103,842,1559]" captionTargetId="figure-144@17.[146,1103,842,1559]" captionTargetPageId="17" captionText="Figure 4. Herentia thalassae David and Pouyet, 1978. MNHN 7619, off NW Spain; note that the vertical component in these perpendicular views is greatly understated, the distal parts of the zooecia and ooecia are distinctly raised. (A) General aspect of colony; (B) close-up of ovicellate zooecium; (C) close-up of autozooecium. Scale bars: 400 mm (A); 100 mm (B, C)." figureDoi="http://doi.org/10.5281/zenodo.5219382" httpUri="https://zenodo.org/record/5219382/files/figure.png" pageId="24" pageNumber="1531">Figure 4A, B</figureCitation>
|
||
).
|
||
<typeStatus id="5487A897FFB3FFE1FE1BFA00FE67EFF9" box="[386,491,1490,1514]" pageId="24" pageNumber="1531" type="paratype">Paratype</typeStatus>
|
||
: NHM 1975.1.12.17B, Chios, Aegean Sea.
|
||
<materialsCitation id="3B541C68FFB3FFE1FC78FA00FD48EC18" collectionCode="PMC, R" location="Gulf of Noto" municipality="Ionian Sea" pageId="24" pageNumber="1531" specimenCount="1">
|
||
Other material:
|
||
<collectionCode id="ED2D8EF0FFB3FFE1FF4CFA21FE97EC18" box="[213,283,1523,1547]" pageId="24" pageNumber="1531">PMC</collectionCode>
|
||
.
|
||
<collectionCode id="ED2D8EF0FFB3FFE1FE87FA21FEB8EC18" box="[286,308,1523,1547]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="24" pageNumber="1531">R</collectionCode>
|
||
.I.H.B-4,
|
||
<location id="8EE340EEFFB3FFE1FE05FA21FDB5EC18" LSID="urn:lsid:plazi:treatment:0395A723FFBEFFE3FE35FC61FC4FEB31:8EE340EEFFB3FFE1FE05FA21FDB5EC18" box="[412,569,1523,1547]" municipality="Ionian Sea" name="Gulf of Noto" pageId="24" pageNumber="1531">Gulf of Noto</location>
|
||
,
|
||
<collectingMunicipality id="6BE78C4FFFB3FFE1FDDFFA21FD48EC18" box="[582,708,1523,1547]" pageId="24" pageNumber="1531">Ionian Sea</collectingMunicipality>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B831635FFB3FFE1FF4CF996FEFAEC4F" blockId="24.[213,374,1604,1628]" box="[213,374,1604,1628]" pageId="24" pageNumber="1531">
|
||
<emphasis id="B948CA27FFB3FFE1FF4CF996FEFAEC4F" box="[213,374,1604,1628]" italics="true" pageId="24" pageNumber="1531">Measurements</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B831635FFB3FFE1FF4CF9BEFB1DECB6" blockId="24.[213,1169,1644,1701]" pageId="24" pageNumber="1531">ZL 681¡47, 588–716 (2, 6); ZW 520¡75, 404–641 (2, 6); OL 154¡10, 133–167 (2, 9); OW 163¡10, 145–175 (2, 9); ApL 157¡5, 151–164 (1, 4); ApW 182¡9, 173–193</paragraph>
|
||
<paragraph id="8B831635FFB2FFE0FF0BFF7AFC1EEAF2" blockId="25.[146,1103,168,225]" pageId="25" pageNumber="1532">(1, 4); OvL 197¡17, 171–208 (1, 4); OvW 326¡40, 282–374 (1, 4); AL 175¡15, 141– 192 (2, 12); AW 136¡11, 122–157 (2, 12); PD 10¡1, 9–13 (2, 12).</paragraph>
|
||
<paragraph id="8B831635FFB2FFE0FF0BFEC8FE9DEB21" blockId="25.[146,273,282,306]" box="[146,273,282,306]" pageId="25" pageNumber="1532">
|
||
<emphasis id="B948CA27FFB2FFE0FF0BFEC8FE9DEB21" box="[146,273,282,306]" italics="true" pageId="25" pageNumber="1532">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B831635FFB2FFE0FF0BFE90FDCCE972" blockId="25.[146,1103,322,1255]" pageId="25" pageNumber="1532">Colony encrusting unilaminar, multiserial. Zooecia rhomboid, hexagonal or polygonal, separated by distinct grooves; frontal wall flat, closely and regularly perforated by numerous small pseudopores, and with few, relatively large, marginal areolar pores (usually two or three, rarely more than five); lateral walls with some 10 relatively small, distolateral communication pores. Orifice very slightly dimorphic with a deeply immersed shelf along the distolateral margin, proximolaterally framed by a smooth narrow band of gymnocystal calcification; primary orifice in autozooecia about as broad as long and widest in distal third, anter three-quarters of a transverse ellipse, proximal border straight, sinus U-shaped with rounded shoulders, occupying about one-third of proximal margin, condyles slightly immersed, strong, crenellated, more or less straight with rounded ends pointing medially and projecting above shoulders of sinus, the flattened frontal faces sloping proximally towards a distinct edge parallel to proximal orifice margin, distolateral orifice border bounded by a slightly raised and curved rim of thick calcification sloping towards orifice; aperture in ovicellate zooecia of similar shape and length but slightly wider; condyles slightly ribbed, not as broad as in autozooecia and ends not projecting beyond shoulders of sinus.</paragraph>
|
||
<paragraph id="8B831635FFB2FFE0FF24FCB8FECCE9D0" blockId="25.[146,1103,322,1255]" pageId="25" pageNumber="1532">Ooecium initially hemispherical, later for the most part immersed, exposed frontal area a crescentic, broad, flat band of smooth calcification slightly oblique to colony surface.</paragraph>
|
||
<paragraph id="8B831635FFB2FFE0FF24FC1EFD47EED5" blockId="25.[146,1103,322,1255]" pageId="25" pageNumber="1532">Avicularium situated directly proximolateral to orifice on either left or right side, in distal pseudoporous cystid, which is level with and not clearly demarcated from frontal shield of zooecium, one or two areolar pores at lateral or proximolateral margin; rostrum oval to kidney-shaped, broad, sloping band, laterally or distolaterally oriented; crossbar complete, without columella, thick and extending fairly deep into cystid, square-edged and twisted along its axis, proximal opesium bounded by an immersed, broad and slightly convex shelf of calcification extending towards crossbar from proximal rostrum margin.</paragraph>
|
||
<paragraph id="8B831635FFB2FFE0FF24FB1DFBC8EEF4" blockId="25.[146,1103,322,1255]" box="[189,1092,1231,1255]" pageId="25" pageNumber="1532">An ancestrula or early astogenetic stages of colony growth were not observed.</paragraph>
|
||
<paragraph id="8B831635FFB2FFE0FF0BFAF3FF79EF2A" blockId="25.[146,245,1313,1337]" box="[146,245,1313,1337]" pageId="25" pageNumber="1532">
|
||
<emphasis id="B948CA27FFB2FFE0FF0BFAF3FF79EF2A" box="[146,245,1313,1337]" italics="true" pageId="25" pageNumber="1532">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B831635FFB2FFE0FF0BFA9BFEB9ECB6" blockId="25.[146,1103,1353,1701]" pageId="25" pageNumber="1532">
|
||
This species was reported as
|
||
<taxonomicName id="4C3C6DB6FFB2FFE0FE47FA9BFDC6EF72" authorityName=": Hayward" authorityYear="1974" box="[478,586,1353,1377]" class="Gymnolaemata" family="Escharinidae" genus="Escharina" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="1532" phylum="Bryozoa" rank="species" species="porosa">
|
||
<emphasis id="B948CA27FFB2FFE0FE47FA9BFDC6EF72" box="[478,586,1353,1377]" italics="true" pageId="25" pageNumber="1532">E. porosa</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFAD6BC4FFB2FFE0FDEFFA9BFCBCEF72" author="Hayward PJ" box="[630,816,1353,1377]" pageId="25" pageNumber="1532" pagination="369 - 402" refId="ref19936" refString="Hayward PJ. 1974. Studies on the cheilostome bryozoan fauna of the Aegean island of Chios. J Nat Hist. 8: 369 - 402." type="journal article" year="1974">Hayward (1974)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFAD6BC4FFB2FFE0FCF2FA9BFB83EF72" author="Rosso A" box="[875,1039,1353,1377]" pageId="25" pageNumber="1532" pagination="189 - 225" refId="ref21479" refString="Rosso A. 1996 a. Popolamenti e tanatocenosi a briozoi di fondi mobili circalitorali del Golfo di Noto (Sicilia SE). Naturalista Siciliano (ser. 4) 20: 189 - 225." type="journal article" year="1996">Rosso (1996a)</bibRefCitation>
|
||
from the eastern and central Mediterranean Sea, respectively. It is possible that
|
||
<taxonomicName id="4C3C6DB6FFB2FFE0FC6EFAB8FBC3EF91" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[1015,1103,1386,1410]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="1532" phylum="Bryozoa" rank="species" species="rosei">
|
||
<emphasis id="B948CA27FFB2FFE0FC6EFAB8FBC3EF91" box="[1015,1103,1386,1410]" italics="true" pageId="25" pageNumber="1532">T. rosei</emphasis>
|
||
</taxonomicName>
|
||
also occurs in the western Mediterranean and the
|
||
<collectingRegion id="49F8D8D7FFB2FFE0FD51FA58FCF4EFB1" box="[712,888,1418,1442]" country="Spain" name="Canarias" pageId="25" pageNumber="1532">Canary Islands</collectingRegion>
|
||
. While Zabala and Maluquer (1988, p. 129, Figure 291; Plate 15, Figure D) record this morphotype from off
|
||
<collectingCountry id="F32B56A5FFB2FFE0FF22FA19FF72EFF0" box="[187,254,1483,1507]" name="Spain" pageId="25" pageNumber="1532">Spain</collectingCountry>
|
||
,
|
||
<bibRefCitation id="EFAD6BC4FFB2FFE0FE92FA19FE18EFF0" author="Cook PL" box="[267,404,1483,1507]" pageId="25" pageNumber="1532" pagination="115 - 262" refId="ref19079" refString="Cook PL. 1968. Bryozoa (Polyzoa) from the coast of tropical West Africa. Atlantide Report 10: 115 - 262." type="book chapter" year="1968">Cook (1968</bibRefCitation>
|
||
, p. 195),
|
||
<bibRefCitation id="EFAD6BC4FFB2FFE0FD99FA19FD7CEFF0" author="Aristegui Ruiz J" box="[512,752,1483,1507]" pageId="25" pageNumber="1532" refId="ref18269" refString="Aristegui Ruiz J. 1984. Briozoos queilostomados (Ectoprocta, Cheilostomata) de Canarias: estudio sistematico, faunistico y bioegeografico [dissertation]. La Laguna (Tenerife): Universidad de La Laguna. 524 p." type="book" year="1984">Arístegui Ruiz (1984</bibRefCitation>
|
||
, p. 294, Figure 62a–c; Plate 20,
|
||
<figureCitation id="13070AB0FFB2FFE0FF0BFA39FE9BEC10" box="[146,279,1515,1539]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStart-2="Figure 8" captionStartId-0="24.[213,280,918,940]" captionStartId-1="26.[213,280,1538,1560]" captionStartId-2="34.[213,280,1625,1647]" captionTargetBox-0="[213,1170,165,882]" captionTargetBox-1="[213,1170,784,1501]" captionTargetBox-2="[371,1010,1031,1579]" captionTargetId-0="figure-145@24.[213,1170,165,882]" captionTargetId-1="figure-165@26.[213,1170,784,1501]" captionTargetId-2="figure-308@34.[371,1011,1031,1589]" captionTargetPageId-0="24" captionTargetPageId-1="26" captionTargetPageId-2="34" captionText-0="Figure 6. Therenia rosei n. sp. Holotype, NHM 1975.1.12.17A, Aegean Sea. (A) Autozooecia; (B) ovicellate zooecia; (C) close-up of autozooecium orifice and avicularium; (D) close-up of ovicellate zooecium; note difference in condyle morphology compared to orifice of autozooecium. Scale bars: 200 mm (A, B); 50 mm (C, D)." captionText-1="Figure 7. Therenia peristomata n. sp. Holotype, NHM 2007.10.9.3, off Madeira. (A) General aspect of zooecia at the colony growth margin; (B) early astogenetic zooecia showing a rather flared peristome; note that the ancestrula has been covered by the lowermost zooecia; (C) close-up of autozooecium orifice and avicularium; (D) ovicellate zooecium; note slight differences in condyle morphology to orifice of autozooecium. Scale bars 200 mm (A, B); 50 mm (C, D)." captionText-2="Figure 8. Geographic distribution of Herentia and Therenia species in the North Atlantic and Mediterranean Sea. Note the co-occurrence of three species (H. hyndmanni, H. andreasi n. sp. and T. peristomata n. sp.) at Madeira, and the wide range of H. hyndmanni." figureDoi-0="http://doi.org/10.5281/zenodo.5219386" figureDoi-1="http://doi.org/10.5281/zenodo.5219389" figureDoi-2="http://doi.org/10.5281/zenodo.5219391" httpUri-0="https://zenodo.org/record/5219386/files/figure.png" httpUri-1="https://zenodo.org/record/5219389/files/figure.png" httpUri-2="https://zenodo.org/record/5219391/files/figure.png" pageId="25" pageNumber="1532">Figures 6–8</figureCitation>
|
||
, 1987), and Arístegui and Cruz (1986, p. 166) report
|
||
<taxonomicName id="4C3C6DB6FFB2FFE0FCE0FA3EFC68EC17" authorityName=": Hayward" authorityYear="1974" box="[889,996,1516,1540]" class="Gymnolaemata" family="Escharinidae" genus="Escharina" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="1532" phylum="Bryozoa" rank="species" species="porosa">
|
||
<emphasis id="B948CA27FFB2FFE0FCE0FA3EFC68EC17" box="[889,996,1516,1540]" italics="true" pageId="25" pageNumber="1532">E. porosa</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<collectingRegion id="49F8D8D7FFB2FFE0FF0BF9DEFF76EC37" box="[146,250,1548,1572]" country="Spain" name="Canarias" pageId="25" pageNumber="1532">Canaries</collectingRegion>
|
||
. Especially Arístegui Ruiz’ (1984) figured specimens are very closely related, if not identical, to
|
||
<taxonomicName id="4C3C6DB6FFB2FFE0FE5BF9FEFD94EC57" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[450,536,1580,1604]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="1532" phylum="Bryozoa" rank="species" species="rosei">
|
||
<emphasis id="B948CA27FFB2FFE0FE5BF9FEFD94EC57" box="[450,536,1580,1604]" italics="true" pageId="25" pageNumber="1532">T. rosei</emphasis>
|
||
</taxonomicName>
|
||
. However, more SEM work is needed to enable a precise determination of these specimens. The same applies to the Mediterranean fossil
|
||
<taxonomicName id="4C3C6DB6FFB2FFE0FF4AF9BFFD8EEC96" authority="Pouyet, 1976" authorityName="Pouyet" authorityYear="1976" box="[211,514,1645,1669]" class="Gymnolaemata" family="Hippothoidae" genus="Hippothoa" kingdom="Animalia" order="Cheilostomatida" pageId="25" pageNumber="1532" phylum="Bryozoa" rank="species" species="montenati">
|
||
<emphasis id="B948CA27FFB2FFE0FF4AF9BFFEE9EC96" box="[211,357,1645,1669]" italics="true" pageId="25" pageNumber="1532">H. montenati</emphasis>
|
||
<bibRefCitation id="EFAD6BC4FFB2FFE0FEF4F9BFFD8EEC96" author="Pouyet S" box="[365,514,1645,1669]" pageId="25" pageNumber="1532" pagination="53 - 82" refId="ref21412" refString="Pouyet S. 1976. Bryozoaires cheilostomes de Pliocene d'Aguilas (Espagne meridionale). Nouv Arch Mus Hist Nat Lyon. 14: 53 - 82." type="journal article" year="1976">Pouyet, 1976</bibRefCitation>
|
||
</taxonomicName>
|
||
, which is also morphologically closely related to the recent species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32645BEFFB1FFE3FF66FF7BFC4FEB31" pageId="26" pageNumber="1533" type="reference_group">
|
||
<paragraph id="8B831635FFB1FFE3FF66FF7BFC4FEB31" blockId="26.[213,1169,168,290]" pageId="26" pageNumber="1533">
|
||
<taxonomicName id="4C3C6DB6FFB1FFE3FF66FF7BFED4EAD2" ID-CoL="56CWV" authorityName="Berning & Tilbrook & Rosso" authorityYear="2008" box="[255,344,169,193]" class="Gymnolaemata" family="Escharinidae" genus="Therenia" kingdom="Animalia" order="Cheilostomatida" pageId="26" pageNumber="1533" phylum="Bryozoa" rank="species" species="rosei">
|
||
<emphasis id="B948CA27FFB1FFE3FF66FF7BFED4EAD2" box="[255,344,169,193]" italics="true" pageId="26" pageNumber="1533">T. rosei</emphasis>
|
||
</taxonomicName>
|
||
was reported off the Greek island of Chios from
|
||
<quantity id="4CC4BBD0FFB1FFE3FC36FF7AFB93EAD3" box="[943,1055,168,192]" metricMagnitude="1" metricUnit="m" metricValue="3.05" metricValueMax="6.0" metricValueMin="0.1" pageId="26" pageNumber="1533" unit="m" value="30.5" valueMax="60.0" valueMin="1.0">1 to 60 m</quantity>
|
||
depth by
|
||
<bibRefCitation id="EFAD6BC4FFB1FFE3FF4CFF1BFE1CEAF2" author="Hayward PJ" box="[213,400,201,225]" pageId="26" pageNumber="1533" pagination="369 - 402" refId="ref19936" refString="Hayward PJ. 1974. Studies on the cheilostome bryozoan fauna of the Aegean island of Chios. J Nat Hist. 8: 369 - 402." type="journal article" year="1974">Hayward (1974)</bibRefCitation>
|
||
.
|
||
<bibRefCitation id="EFAD6BC4FFB1FFE3FE02FF1BFDB1EAF2" author="Rosso A" box="[411,573,201,225]" pageId="26" pageNumber="1533" pagination="189 - 225" refId="ref21479" refString="Rosso A. 1996 a. Popolamenti e tanatocenosi a briozoi di fondi mobili circalitorali del Golfo di Noto (Sicilia SE). Naturalista Siciliano (ser. 4) 20: 189 - 225." type="journal article" year="1996">Rosso (1996a)</bibRefCitation>
|
||
found a single living specimen at
|
||
<quantity id="4CC4BBD0FFB1FFE3FC27FF1BFC7FEAF2" box="[958,1011,201,225]" metricMagnitude="1" metricUnit="m" metricValue="3.3" pageId="26" pageNumber="1533" unit="m" value="33.0">33 m</quantity>
|
||
and few dead colony fragments between
|
||
<quantity id="4CC4BBD0FFB1FFE3FD88FF3BFD11EB12" box="[529,669,233,257]" metricMagnitude="1" metricUnit="m" metricValue="8.05" metricValueMax="13.0" metricValueMin="3.1" pageId="26" pageNumber="1533" unit="m" value="80.5" valueMax="130.0" valueMin="31.0">31 to 130 m</quantity>
|
||
depth off south-eastern
|
||
<collectingRegion id="49F8D8D7FFB1FFE3FC5FFF3BFB8BEB12" box="[966,1031,233,257]" country="Italy" name="Sicilia" pageId="26" pageNumber="1533">Sicily</collectingRegion>
|
||
. It encrusts rocks and calcareous substrata such as shells and coralline algae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |