559 lines
76 KiB
XML
559 lines
76 KiB
XML
<document id="FA62FBB098E347522AAB643B261107F6" ID-CLB-Dataset="32021" ID-DOI="10.11646/zootaxa.4323.3.1" ID-GBIF-Dataset="f1be4566-adeb-4995-83c7-2aebab3c7458" ID-ISSN="1175-5326" ID-Zenodo-Dep="919606" ID-ZooBank="3E58F99B-2A30-437A-95Ae-D68A1D3Ecc45" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1506057765384" checkinUser="plazi" docAuthor="Schmidt, Joachim" docDate="2017" docId="03956A03A3757F66FF67FF368E3FDD4C" docLanguage="en" docName="zootaxa.4323.3.1.pdf" docOrigin="Zootaxa 4323 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichotichnus (Bottchrus) pusillus Schmidt 2017, sp. n." docType="treatment" docVersion="7" lastPageNumber="328" masterDocId="FFAC127BA36D7F7DFFF0FFAE8B5CD930" masterDocTitle="Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species" masterLastPageNumber="358" masterPageNumber="301" pageNumber="325" updateTime="1698468866275" updateUser="ExternalLinkService">
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<mods:title id="4C1D5F55C50FB5B0AB2B4717ED24EFE0">Brachypterous ground beetles of the Trichotichnus subgenus Bottchrus Jedlička (Coleoptera, Carabidae) from the Himalaya, with description of fifteen new species</mods:title>
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<mods:namePart id="4BAA22B2E3656751A57EA3918ADE4905">Schmidt, Joachim</mods:namePart>
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<mods:typeOfResource id="086F37491797B7024A0F8FF1D0E0D177">text</mods:typeOfResource>
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<mods:title id="7BFF04380AF62B5BA5D326CE3EE52A63">Zootaxa</mods:title>
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<mods:part id="BE3A504A6F77F4AD6AA3B26F1FC7D7E7">
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<mods:date id="846946264C6061B33090E3A8D4316407">2017</mods:date>
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<mods:number id="E564A2A17CD13BB2D43207166C5E6A1F">2017-09-22</mods:number>
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<mods:number id="2D1F721E96B43C0124A8694A41252803">4323</mods:number>
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<mods:classification id="8A2C65178EE46571B4DBA363DFCB9951">journal article</mods:classification>
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<mods:identifier id="E895E7AF50E16AFE1CE68A004A68BC02" type="DOI">10.11646/zootaxa.4323.3.1</mods:identifier>
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<treatment id="03956A03A3757F66FF67FF368E3FDD4C" ID-DOI="http://doi.org/10.5281/zenodo.6015755" ID-GBIF-Taxon="134839416" ID-Zenodo-Dep="6015755" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C" httpUri="http://treatment.plazi.org/id/03956A03A3757F66FF67FF368E3FDD4C" lastPageId="27" lastPageNumber="328" pageId="24" pageNumber="325">
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<subSubSection id="C326889EA3757F65FF67FF368A92D9E3" pageId="24" pageNumber="325" type="nomenclature">
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<paragraph id="8B83DB15A3757F65FF67FF3689DED982" blockId="24.[151,642,152,211]" box="[151,642,152,178]" pageId="24" pageNumber="325">
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<heading id="D0CB6C79A3757F65FF67FF3689DED982" bold="true" box="[151,642,152,178]" fontSize="11" level="1" pageId="24" pageNumber="325" reason="1">
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<emphasis id="B9480707A3757F65FF67FF3689DED982" bold="true" box="[151,642,152,178]" pageId="24" pageNumber="325">
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<taxonomicName id="4C3CA096A3757F65FF67FF368964D982" ID-CoL="58LHT" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[151,568,152,178]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="325" phylum="Arthropoda" rank="species" species="pusillus" status="sp. nov." subGenus="Bottchrus">
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<emphasis id="B9480707A3757F65FF67FF368A1DD982" bold="true" box="[151,321,152,178]" italics="true" pageId="24" pageNumber="325">Trichotichnus</emphasis>
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||
(
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<emphasis id="B9480707A3757F65FEA2FF368A95D982" bold="true" box="[338,457,152,178]" italics="true" pageId="24" pageNumber="325">Bottchrus</emphasis>
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||
)
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||
<emphasis id="B9480707A3757F65FE2AFF368964D982" bold="true" box="[474,568,152,178]" italics="true" pageId="24" pageNumber="325">pusillus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27BBA7CA3757F65FDCFFF3689DED982" box="[575,642,152,178]" pageId="24" pageNumber="325" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8B83DB15A3757F65FF67FF148A92D9E3" blockId="24.[151,642,152,211]" box="[151,462,186,211]" pageId="24" pageNumber="325">
|
||
(
|
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<figureCitation id="1307C790A3757F65FF6FFF148A7DD9E3" box="[159,289,186,211]" captionStart="FIGURES 67 – 70" captionStartId="25.[151,264,1947,1970]" captionTargetBox="[311,1277,200,1923]" captionTargetId="figure@25.[305,1282,193,1927]" captionTargetPageId="25" captionText="FIGURES 67 – 70. Himalayan species of Trichotichnus subgenus Bottchrus, photographs of habitus. 67, T. pusillus sp. n., paratype, male, from the Bara Pokhari Lekh mountain range; 68, T. pusillus sp. n., paratype, male, from the Dudh Pokhari Lekh mountain range; 69, T. ganeshensis sp. n., paratype, male; 70, T. minutus sp. n., paratype, male." httpUri="https://zenodo.org/record/919638/files/figure.png" pageId="24" pageNumber="325">Figs 67–68</figureCitation>
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,
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<figureCitation id="1307C790A3757F65FEDBFF158A28D9E3" box="[299,372,187,211]" captionStart="FIGURES 71 – 79" captionStartId="26.[151,264,1133,1156]" captionTargetBox="[180,1420,209,1090]" captionTargetId="figure@26.[151,1436,193,1113]" captionTargetPageId="26" captionText="FIGURES 71 – 79. Trichotichnus (Bottchrus) pusillus sp. n. (71, 78, 79, paratypes from Dudh Pokhari Lekh; 72, specimen from Annapurna; 73, 74, paratype from Bara Pokhari Lekh; 75 – 77, holotype). 71, left metepisternum; 72 – 73, 75, 77, median lobe of aedeagus; 74, 76, apical capitulum; 78, apex of last (VII) abdominal sternite, male; 79, same, female. 71, 78 – 79, ventral view; 72 – 73, 75, lateral view; 74, 76, caudal view; 77, dorsal view. Scale bars = 0.25 mm (A: Figs 71 – 73, 75, 77 – 79; B: Figs 74, 76)." httpUri="https://zenodo.org/record/919642/files/figure.png" pageId="24" pageNumber="325">71–79</figureCitation>
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,
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<figureCitation id="1307C790A3757F65FE8FFF148AF7D9E3" box="[383,427,186,211]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="24" pageNumber="325">151</figureCitation>
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,
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<figureCitation id="1307C790A3757F65FE47FF158A99D9E3" box="[439,453,187,211]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="24" pageNumber="325">
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||
<emphasis id="B9480707A3757F65FE47FF158A99D9E3" box="[439,453,187,211]" italics="true" pageId="24" pageNumber="325">9</emphasis>
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</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C326889EA3757F65FF67FEAC8B84DA23" pageId="24" pageNumber="325" type="materials_examined">
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<paragraph id="8B83DB15A3757F65FF67FEAC8AA8D80F" blockId="24.[151,1437,258,2011]" pageId="24" pageNumber="325">
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<materialsCitation id="3B54D148A3757F65FF67FEAC8AA8D80F" ID-GBIF-Occurrence="1655986824" collectingDate="2003-04-22" collectorName="J. Schmidt" country="Nepal" elevation="2000" location="Manaslu Mts." pageId="24" pageNumber="325" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||
<emphasis id="B9480707A3757F65FF67FEAC8A1DD82B" bold="true" box="[151,321,258,283]" pageId="24" pageNumber="325">
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<typeStatus id="548765B7A3757F65FF67FEAC8B8ED82B" box="[151,210,258,283]" pageId="24" pageNumber="325">Type</typeStatus>
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||
material
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||
</emphasis>
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||
.
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||
<typeStatus id="548765B7A3757F65FEA2FEAC8AE3D82B" box="[338,447,258,283]" pageId="24" pageNumber="325" type="holotype">Holotype</typeStatus>
|
||
:
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||
<specimenCount id="9D3A109CA3757F65FE3CFEAD8ABFD82A" box="[460,483,259,282]" pageId="24" pageNumber="325" type="male">♂</specimenCount>
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||
, “
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||
<collectingCountry id="F32B9B85A3757F65FE0CFEAD890AD82B" box="[508,598,259,283]" name="Nepal" pageId="24" pageNumber="325">NEPAL</collectingCountry>
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<location id="8EE38DCEA3757F65FD90FEAC89A2D82A" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FD90FEAC89A2D82A" box="[608,766,258,283]" country="Nepal" name="Manaslu Mts." pageId="24" pageNumber="325">Manaslu Mts.</location>
|
||
<collectingDate id="EFC6043DA3757F65FCF8FEAD88C4D82B" box="[776,920,259,283]" pageId="24" pageNumber="325" value="2003-04-22">
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||
<date id="FF82FDD5A3757F65FCF8FEAD88C4D82B" box="[776,920,259,283]" pageId="24" pageNumber="325" value="2003-04-22">2003, 22.IV.</date>
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||
</collectingDate>
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||
, Dudh Pokhari Lekh, below Helam Pokhari,
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||
<elevation id="00113C26A3757F65FF67FE898BB2D80E" box="[151,238,295,319]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="325" unit="m" value="2000.0">
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||
<quantity id="4CC476F0A3757F65FF67FE898BB2D80E" box="[151,238,295,319]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="325" unit="m" value="2000.0">2000 m</quantity>
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||
</elevation>
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||
, leg.
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||
<collectorName id="26C9BEC3A3757F65FEDAFE898AFFD80F" box="[298,419,294,319]" pageId="24" pageNumber="325">J. Schmidt</collectorName>
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” (cJS).
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||
</materialsCitation>
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</paragraph>
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||
<paragraph id="8B83DB15A3757F65FF37FEE58BB6DBFB" blockId="24.[151,1437,258,2011]" pageId="24" pageNumber="325">
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||
<materialsCitation id="3B54D148A3757F65FF37FEE588E4DB96" ID-GBIF-Occurrence="1655986827" collectingDate="1995-09-15" collectingDateMax="2005-04-29" collectingDateMin="1995-09-15" collectionCode="ZIN" collectorName="J. Schmidt & Manaslu Mts, S & Manaslu Mts., S" country="Nepal" county="Barapokhari Lekh" elevation="2350" latitude="28.25" location="Manaslu Mts." longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" pageId="24" pageNumber="325" specimenCount="164" specimenCount-female="64" specimenCount-male="100" typeStatus="paratype">
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||
<typeStatus id="548765B7A3757F65FF37FEE58A65D853" box="[199,313,331,355]" pageId="24" pageNumber="325" type="paratype">Paratypes</typeStatus>
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||
:
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||
<specimenCount id="9D3A109CA3757F65FEB4FEE58ADCD852" box="[324,384,331,355]" pageId="24" pageNumber="325" type="male">21 ♂</specimenCount>
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,
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||
<specimenCount id="9D3A109CA3757F65FE7EFEE58AE6D852" box="[398,442,331,355]" pageId="24" pageNumber="325" type="female">8 ♀</specimenCount>
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||
, same data as holotype (cJS,
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||
<collectionCode id="ED2D43D0A3757F65FCFFFEE5881ED853" box="[783,834,331,355]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="24" pageNumber="325">ZIN</collectionCode>
|
||
);
|
||
<specimenCount id="9D3A109CA3757F65FCA7FEE588D9D852" box="[855,901,331,355]" pageId="24" pageNumber="325" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3A109CA3757F65FC63FEE588E3D852" box="[915,959,331,355]" pageId="24" pageNumber="325" type="female">1 ♀</specimenCount>
|
||
,
|
||
<location id="8EE38DCEA3757F65FC3FFEE48F31D852" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FC3FFEE48F31D852" box="[975,1133,330,355]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Manaslu Mts." pageId="24" pageNumber="325">Manaslu Mts.</location>
|
||
,
|
||
<location id="8EE38DCEA3757F65FB8BFEE48E02D853" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FB8BFEE48E02D853" box="[1147,1374,330,355]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Dudh Pokhari Lekh" pageId="24" pageNumber="325">Dudh Pokhari Lekh</location>
|
||
, upp.
|
||
<location id="8EE38DCEA3757F65FF67FEC18BBED8B7" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FF67FEC18BBED8B7" box="[151,226,367,391]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Jorney" pageId="24" pageNumber="325">Jorney</location>
|
||
vill.,
|
||
<elevation id="00113C26A3757F65FED9FEC08A95D8B6" box="[297,457,366,391]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.6" metricValueMin="2.1" pageId="24" pageNumber="325" unit="m" value="2350.0" valueMax="2600.0" valueMin="2100.0">
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<quantity id="4CC476F0A3757F65FED9FEC08A95D8B6" box="[297,457,366,391]" metricMagnitude="3" metricUnit="m" metricValue="2.35" metricValueMax="2.6" metricValueMin="2.1" pageId="24" pageNumber="325" unit="m" value="2350.0" valueMax="2600.0" valueMin="2100.0">2100–2600 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFC6043DA3757F65FE28FEC08906D8B7" box="[472,602,366,391]" pageId="24" pageNumber="325" value="1995-09-15">
|
||
<date id="FF82FDD5A3757F65FE28FEC08906D8B7" box="[472,602,366,391]" pageId="24" pageNumber="325" value="1995-09-15">15.IX.1995</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C9BEC3A3757F65FD99FEC189BED8B7" box="[617,738,366,391]" pageId="24" pageNumber="325">J. Schmidt</collectorName>
|
||
leg. (cJS);
|
||
<specimenCount id="9D3A109CA3757F65FC9CFEC188F4D8B6" box="[876,936,367,391]" pageId="24" pageNumber="325" type="male">11 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3A109CA3757F65FC47FEC088AED8B6" box="[951,1010,366,391]" pageId="24" pageNumber="325" type="female">15 ♀</specimenCount>
|
||
,
|
||
<location id="8EE38DCEA3757F65FBF3FEC08FFFD8B6" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FBF3FEC08FFFD8B6" box="[1027,1187,366,391]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Manaslu Mts." pageId="24" pageNumber="325">Manaslu Mts.</location>
|
||
,
|
||
<location id="8EE38DCEA3757F65FB42FEC08EC4D8B7" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FB42FEC08EC4D8B7" box="[1202,1432,366,391]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Dudh Pokhari Lekh" pageId="24" pageNumber="325">Dudh Pokhari Lekh</location>
|
||
,
|
||
<location id="8EE38DCEA3757F65FF67FE3D8AF9D89B" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FF67FE3D8AF9D89B" box="[151,421,402,427]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="upper Phulinggiri Madi" pageId="24" pageNumber="325">upper Phulinggiri Madi</location>
|
||
,
|
||
<elevation id="00113C26A3757F65FE42FE3C890ED89A" box="[434,594,402,427]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="24" pageNumber="325" unit="m" value="2300.0" valueMax="2500.0" valueMin="2100.0">
|
||
<quantity id="4CC476F0A3757F65FE42FE3C890ED89A" box="[434,594,402,427]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="24" pageNumber="325" unit="m" value="2300.0" valueMax="2500.0" valueMin="2100.0">2100–2500 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFC6043DA3757F65FD90FE3D8854D89B" box="[608,776,403,427]" pageId="24" pageNumber="325" value="2003-04-19" valueMax="2003-04-21" valueMin="2003-04-19">
|
||
<date id="FF82FDD5A3757F65FD90FE3D8854D89B" box="[608,776,403,427]" pageId="24" pageNumber="325" value="2003-04-19" valueMax="2003-04-21" valueMin="2003-04-19">19–21.IV.2003</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C9BEC3A3757F65FCE6FE3D88D2D89B" box="[790,910,402,427]" pageId="24" pageNumber="325">J. Schmidt</collectorName>
|
||
leg. (cJS);
|
||
<specimenCount id="9D3A109CA3757F65FBE5FE3D8F1DD89A" box="[1045,1089,403,427]" pageId="24" pageNumber="325" type="female">3 ♀</specimenCount>
|
||
,
|
||
<location id="8EE38DCEA3757F65FBA1FE3C8FACD89A" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FBA1FE3C8FACD89A" box="[1105,1264,402,427]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Manaslu Mts." pageId="24" pageNumber="325">Manaslu Mts.</location>
|
||
,
|
||
<location id="8EE38DCEA3757F65FB0EFE3C8B88D8FF" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FB0EFE3C8B88D8FF" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Dudh Pokhari Lekh" pageId="24" pageNumber="325">Dudh Pokhari Lekh</location>
|
||
,
|
||
<location id="8EE38DCEA3757F65FF11FE188A98D8FF" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FF11FE188A98D8FF" box="[225,452,438,463]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Dordi Khola Valley" pageId="24" pageNumber="325">Dordi Khola Valley</location>
|
||
,
|
||
<elevation id="00113C26A3757F65FE21FE19892DD8FE" box="[465,625,439,463]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="24" pageNumber="325" unit="m" value="2150.0" valueMax="2300.0" valueMin="2000.0">
|
||
<quantity id="4CC476F0A3757F65FE21FE19892DD8FE" box="[465,625,439,463]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="24" pageNumber="325" unit="m" value="2150.0" valueMax="2300.0" valueMin="2000.0">2000–2300 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFC6043DA3757F65FD70FE188874D8FF" box="[640,808,438,463]" pageId="24" pageNumber="325" value="2003-04-14" valueMax="2003-04-15" valueMin="2003-04-14">
|
||
<date id="FF82FDD5A3757F65FD70FE188874D8FF" box="[640,808,438,463]" pageId="24" pageNumber="325" value="2003-04-14" valueMax="2003-04-15" valueMin="2003-04-14">14–15.IV.2003</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C9BEC3A3757F65FCC7FE1988F3D8FF" box="[823,943,438,463]" pageId="24" pageNumber="325">J. Schmidt</collectorName>
|
||
leg. (cJS);
|
||
<specimenCount id="9D3A109CA3757F65FBC8FE198F28D8FE" box="[1080,1140,439,463]" pageId="24" pageNumber="325" type="male">11 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3A109CA3757F65FB73FE198FECD8FE" box="[1155,1200,439,463]" pageId="24" pageNumber="325" type="female">8 ♀</specimenCount>
|
||
,
|
||
<location id="8EE38DCEA3757F65FB31FE188E05D8FE" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FB31FE188E05D8FE" box="[1217,1369,438,463]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Manaslu Mts" pageId="24" pageNumber="325">Manaslu Mts</location>
|
||
,
|
||
<location id="8EE38DCEA3757F65FA97FE198A6ED8C3" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FA97FE198A6ED8C3" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Bara Pokhari Lekh" pageId="24" pageNumber="325">Bara Pokhari Lekh</location>
|
||
, Chhandi Khola Vall.,
|
||
<elevation id="00113C26A3757F65FDDEFE758997D8C2" box="[558,715,475,499]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="24" pageNumber="325" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">
|
||
<quantity id="4CC476F0A3757F65FDDEFE758997D8C2" box="[558,715,475,499]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="24" pageNumber="325" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000–2200 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFC6043DA3757F65FD27FE758822D8C3" box="[727,894,475,499]" pageId="24" pageNumber="325" value="2003-04-11" valueMax="2003-04-12" valueMin="2003-04-11">
|
||
<date id="FF82FDD5A3757F65FD27FE758822D8C3" box="[727,894,475,499]" pageId="24" pageNumber="325" value="2003-04-11" valueMax="2003-04-12" valueMin="2003-04-11">11–12.IV.2003</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C9BEC3A3757F65FC79FE7588A2D8C3" box="[905,1022,474,499]" pageId="24" pageNumber="325">J. Schmidt</collectorName>
|
||
leg. (cJS);
|
||
<specimenCount id="9D3A109CA3757F65FB8EFE758FF5D8C2" box="[1150,1193,475,499]" pageId="24" pageNumber="325" type="male">4 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3A109CA3757F65FB45FE758F83D8C2" box="[1205,1247,475,498]" pageId="24" pageNumber="325" type="female">7 ♀</specimenCount>
|
||
,
|
||
<collectorName id="26C9BEC3A3757F65FB1CFE748EC0D8C3" box="[1260,1436,474,499]" pageId="24" pageNumber="325">Manaslu Mts, S</collectorName>
|
||
of
|
||
<location id="8EE38DCEA3757F65FF47FE518A13DB27" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FF47FE518A13DB27" box="[183,335,510,535]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Bara Pokhari" pageId="24" pageNumber="325">Bara Pokhari</location>
|
||
,
|
||
<elevation id="00113C26A3757F65FEADFE518AEADB26" box="[349,438,511,535]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="325" unit="m" value="2000.0">
|
||
<quantity id="4CC476F0A3757F65FEADFE518AEADB26" box="[349,438,511,535]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="325" unit="m" value="2000.0">2000 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFC6043DA3757F65FE34FE518969DB27" box="[452,565,511,535]" pageId="24" pageNumber="325" value="2003-04-08">
|
||
<date id="FF82FDD5A3757F65FE34FE518969DB27" box="[452,565,511,535]" pageId="24" pageNumber="325" value="2003-04-08">8.IV.2003</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C9BEC3A3757F65FDB2FE5189E6DB27" box="[578,698,510,535]" pageId="24" pageNumber="325">J. Schmidt</collectorName>
|
||
leg. (cJS);
|
||
<specimenCount id="9D3A109CA3757F65FCB1FE518833DB26" box="[833,879,511,535]" pageId="24" pageNumber="325" type="male">3 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3A109CA3757F65FC8DFE5188F6DB26" box="[893,938,511,535]" pageId="24" pageNumber="325" type="female">3 ♀</specimenCount>
|
||
,
|
||
<collectorName id="26C9BEC3A3757F65FC4AFE508F33DB27" box="[954,1135,510,535]" pageId="24" pageNumber="325">Manaslu Mts, S</collectorName>
|
||
of
|
||
<location id="8EE38DCEA3757F65FB69FE518E6DDB27" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FB69FE518E6DDB27" box="[1177,1329,510,535]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Bara Pokhari" pageId="24" pageNumber="325">Bara Pokhari</location>
|
||
,
|
||
<elevation id="00113C26A3757F65FACEFE518ECBDB26" box="[1342,1431,511,535]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="325" unit="m" value="2000.0">
|
||
<quantity id="4CC476F0A3757F65FACEFE518ECBDB26" box="[1342,1431,511,535]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="325" unit="m" value="2000.0">2000 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFC6043DA3757F65FF67FD8D8A5BDB0B" box="[151,263,547,571]" pageId="24" pageNumber="325" value="1999-04-01">
|
||
<date id="FF82FDD5A3757F65FF67FD8D8A5BDB0B" box="[151,263,547,571]" pageId="24" pageNumber="325" value="1999-04-01">1.IV.1999</date>
|
||
</collectingDate>
|
||
,
|
||
<location id="8EE38DCEA3757F65FEE3FD8D8A63DB0A" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FEE3FD8D8A63DB0A" box="[275,319,547,570]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Lau" pageId="24" pageNumber="325">Lau</location>
|
||
&
|
||
<collectorName id="26C9BEC3A3757F65FE94FD8D8A86DB0B" box="[356,474,546,571]" pageId="24" pageNumber="325">J. Schmidt</collectorName>
|
||
leg. (cJS,
|
||
<collectionCode id="ED2D43D0A3757F65FDA2FD8D89D9DB0B" box="[594,645,547,571]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="24" pageNumber="325">ZIN</collectionCode>
|
||
);
|
||
<specimenCount id="9D3A109CA3757F65FD69FD8D899FDB0A" box="[665,707,547,571]" pageId="24" pageNumber="325" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3A109CA3757F65FD22FD8D89A1DB0A" box="[722,765,547,571]" pageId="24" pageNumber="325" type="female">1 ♀</specimenCount>
|
||
,
|
||
<location id="8EE38DCEA3757F65FCFBFD8C88FDDB0A" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FCFBFD8C88FDDB0A" box="[779,929,546,571]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Manaslu Mts" pageId="24" pageNumber="325">Manaslu Mts</location>
|
||
,
|
||
<location id="8EE38DCEA3757F65FC5DFD8D8FD9DB0B" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FC5DFD8D8FD9DB0B" box="[941,1157,546,571]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Bara Pokhari Lekh" pageId="24" pageNumber="325">Bara Pokhari Lekh</location>
|
||
, upp.
|
||
<location id="8EE38DCEA3757F65FB39FD8C8E4FDB0B" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FB39FD8C8E4FDB0B" box="[1225,1299,546,571]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Taksar" pageId="24" pageNumber="325">Taksar</location>
|
||
vill.,
|
||
<elevation id="00113C26A3757F65FAA6FD8D8BAEDB6E" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="24" pageNumber="325" unit="m" value="2050.0" valueMax="2100.0" valueMin="2000.0">
|
||
<quantity id="4CC476F0A3757F65FAA6FD8D8BAEDB6E" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="24" pageNumber="325" unit="m" value="2050.0" valueMax="2100.0" valueMin="2000.0">2000– 2100 m</quantity>
|
||
</elevation>
|
||
, 1+
|
||
<collectingDate id="EFC6043DA3757F65FEECFDE98AC7DB6F" box="[284,411,583,607]" pageId="24" pageNumber="325" value="1999-04-11">
|
||
<date id="FF82FDD5A3757F65FEECFDE98AC7DB6F" box="[284,411,583,607]" pageId="24" pageNumber="325" value="1999-04-11">11.IV.1999</date>
|
||
</collectingDate>
|
||
(cJS);
|
||
<specimenCount id="9D3A109CA3757F65FE03FDE8897FDB6E" box="[499,547,582,607]" pageId="24" pageNumber="325" type="male">6 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3A109CA3757F65FDC3FDE9893DDB6E" box="[563,609,583,607]" pageId="24" pageNumber="325" type="female">4 ♀</specimenCount>
|
||
, same as preceding but
|
||
<collectingDate id="EFC6043DA3757F65FC73FDE988A8DB6F" box="[899,1012,583,607]" pageId="24" pageNumber="325" value="2003-04-07">
|
||
<date id="FF82FDD5A3757F65FC73FDE988A8DB6F" box="[899,1012,583,607]" pageId="24" pageNumber="325" value="2003-04-07">7.IV.2003</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C9BEC3A3757F65FBF4FDE98F22DB6F" box="[1028,1150,582,607]" pageId="24" pageNumber="325">J. Schmidt</collectorName>
|
||
leg. (cJS);
|
||
<specimenCount id="9D3A109CA3757F65FAFAFDE98E15DB6E" box="[1290,1353,583,607]" pageId="24" pageNumber="325" type="male">42 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3A109CA3757F65FAA9FDE98EC9DB6E" box="[1369,1429,583,607]" pageId="24" pageNumber="325" type="female">13 ♀</specimenCount>
|
||
,
|
||
<collectorName id="26C9BEC3A3757F65FF67FDC48A11DBB3" box="[151,333,618,643]" pageId="24" pageNumber="325">
|
||
<location id="8EE38DCEA3757F65FF67FDC48A6FDBB2" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FF67FDC48A6FDBB2" box="[151,307,618,643]" county="Barapokhari Lekh" latitude="28.25" longLatPrecision="1233" longitude="84.416664" municipality="Manaslu" name="Manaslu Mts." pageId="24" pageNumber="325">Manaslu Mts.</location>
|
||
, S
|
||
</collectorName>
|
||
Bara Pokhari, ca.
|
||
<geoCoordinate id="EE08BDD2A3757F65FDE9FDC48926DBB3" box="[537,634,618,643]" direction="north" orientation="latitude" pageId="24" pageNumber="325" precision="925" value="28.25">28°15’N</geoCoordinate>
|
||
<geoCoordinate id="EE08BDD2A3757F65FD70FDC589BCDBB3" box="[640,736,618,643]" direction="east" orientation="longitude" pageId="24" pageNumber="325" precision="925" value="84.416664">84°25’E</geoCoordinate>
|
||
,
|
||
<elevation id="00113C26A3757F65FD1BFDC588D4DBB2" box="[747,904,619,643]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="24" pageNumber="325" unit="m" value="2050.0" valueMax="2100.0" valueMin="2000.0">
|
||
<quantity id="4CC476F0A3757F65FD1BFDC588D4DBB2" box="[747,904,619,643]" metricMagnitude="3" metricUnit="m" metricValue="2.05" metricValueMax="2.1" metricValueMin="2.0" pageId="24" pageNumber="325" unit="m" value="2050.0" valueMax="2100.0" valueMin="2000.0">2000–2100 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFC6043DA3757F65FC63FDC48F52DBB3" box="[915,1038,618,643]" pageId="24" pageNumber="325" value="2005-04-29">
|
||
<date id="FF82FDD5A3757F65FC63FDC48F52DBB3" box="[915,1038,618,643]" pageId="24" pageNumber="325" value="2005-04-29">29.IV.2005</date>
|
||
</collectingDate>
|
||
(cJS,
|
||
<collectionCode id="ED2D43D0A3757F65FBA3FDC58FDADBB3" box="[1107,1158,619,643]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="24" pageNumber="325">ZIN</collectionCode>
|
||
);
|
||
<specimenCount id="9D3A109CA3757F65FB69FDC58F9EDBB2" box="[1177,1218,619,643]" pageId="24" pageNumber="325" type="female">1 ♀</specimenCount>
|
||
, C-Nepal,
|
||
<collectingMunicipality id="6BE7416FA3757F65FACCFDC48EC0DBB3" box="[1340,1436,618,643]" pageId="24" pageNumber="325">Manaslu</collectingMunicipality>
|
||
massif,
|
||
<collectingCounty id="62E2A399A3757F65FF03FD208A9ADB97" box="[243,454,654,679]" pageId="24" pageNumber="325">Barapokhari Lekh</collectingCounty>
|
||
,
|
||
<locationDeviation id="9A31C237A3757F65FE25FD218997DB97" box="[469,715,654,679]" pageId="24" pageNumber="325">
|
||
<quantity id="4CC476F0A3757F65FE25FD21897DDB97" box="[469,545,654,679]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="24" pageNumber="325" unit="km" value="23.0">23 km</quantity>
|
||
NE Besisahar
|
||
</locationDeviation>
|
||
vill.,
|
||
<elevation id="00113C26A3757F65FCE6FD2188E4DB96" box="[790,952,655,679]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="24" pageNumber="325" unit="m" value="2500.0" valueMax="3000.0" valueMin="2000.0">
|
||
<quantity id="4CC476F0A3757F65FCE6FD2188E4DB96" box="[790,952,655,679]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="24" pageNumber="325" unit="m" value="2500.0" valueMax="3000.0" valueMin="2000.0">2000–3000 m</quantity>
|
||
</elevation>
|
||
</materialsCitation>
|
||
, 28°21'N, 84°33'E,
|
||
<date id="FF82FDD5A3757F65FB54FD218E50DB97" box="[1188,1292,655,679]" pageId="24" pageNumber="325" value="2010-07">VII.2010</date>
|
||
, Santos leg. (cWR).
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3757F65FF37FD788B84DA23" blockId="24.[151,1437,258,2011]" pageId="24" pageNumber="325">
|
||
<materialsCitation id="3B54D148A3757F65FF37FD788B84DA23" ID-GBIF-Occurrence="1655986832" collectingDate="1997-06-15" collectorName="J. Schmidt" country="Nepal" elevation="2000" location="Annapurna" municipality="Telbrung Danda" pageId="24" pageNumber="325" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<emphasis id="B9480707A3757F65FF37FD788AEADBDF" bold="true" box="[199,438,726,751]" pageId="24" pageNumber="325">Additional material</emphasis>
|
||
.
|
||
<specimenCount id="9D3A109CA3757F65FE38FD798AABDBDE" box="[456,503,727,751]" pageId="24" pageNumber="325" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D3A109CA3757F65FDF7FD798969DBDE" box="[519,565,727,751]" pageId="24" pageNumber="325" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingCountry id="F32B9B85A3757F65FDB6FD7889D0DBDF" box="[582,652,726,751]" name="Nepal" pageId="24" pageNumber="325">Nepal</collectingCountry>
|
||
,
|
||
<location id="8EE38DCEA3757F65FD6BFD798846DBDF" LSID="urn:lsid:plazi:treatment:03956A03A3757F66FF67FF368E3FDD4C:8EE38DCEA3757F65FD6BFD798846DBDF" box="[667,794,727,751]" country="Nepal" municipality="Telbrung Danda" name="Annapurna" pageId="24" pageNumber="325">Annapurna</location>
|
||
,
|
||
<collectingMunicipality id="6BE7416FA3757F65FCD9FD7888B8DBDF" box="[809,996,726,751]" pageId="24" pageNumber="325">Telbrung Danda</collectingMunicipality>
|
||
,
|
||
<elevation id="00113C26A3757F65FC03FD798F12DBDE" box="[1011,1102,727,751]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="325" unit="m" value="2000.0">
|
||
<quantity id="4CC476F0A3757F65FC03FD798F12DBDE" box="[1011,1102,727,751]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="325" unit="m" value="2000.0">2000 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingDate id="EFC6043DA3757F65FBADFD788F83DBDF" box="[1117,1247,726,751]" pageId="24" pageNumber="325" value="1997-06-15">
|
||
<date id="FF82FDD5A3757F65FBADFD788F83DBDF" box="[1117,1247,726,751]" pageId="24" pageNumber="325" value="1997-06-15">15.VI.1997</date>
|
||
</collectingDate>
|
||
,
|
||
<collectorName id="26C9BEC3A3757F65FB1EFD798E34DBDF" box="[1262,1384,726,751]" pageId="24" pageNumber="325">J. Schmidt</collectorName>
|
||
leg. (cJS).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA3757F66FF37FCB08F52D8E0" lastPageId="27" lastPageNumber="328" pageId="24" pageNumber="325" type="description">
|
||
<paragraph id="8B83DB15A3757F65FF37FCB08F24DA06" blockId="24.[151,1437,258,2011]" box="[199,1144,798,823]" pageId="24" pageNumber="325">
|
||
<emphasis id="B9480707A3757F65FF37FCB08A0DDA07" bold="true" box="[199,337,798,823]" pageId="24" pageNumber="325">Description</emphasis>
|
||
. Habitus as in figs 67–68. Body length 5.3–6.0 mm, width
|
||
<quantity id="4CC476F0A3757F65FC19FCB08F28DA06" box="[1001,1140,798,823]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.6" metricValueMin="2.4" pageId="24" pageNumber="325" unit="mm" value="3.0" valueMax="3.6" valueMin="2.4">2.4–3.6 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3757F65FF37FCEC8F26DA93" blockId="24.[151,1437,258,2011]" pageId="24" pageNumber="325">Colour: Body piceous, shiny on dorsum, with light, but distinct bluish or greenish tinge on elytra and often also on pronotum; labrum, in most specimens also base of mandibles, lateral gutters of pronotum and elytra reddish brown; palpi, antennae and legs brownish yellow, occasionally femora slightly infuscate.</paragraph>
|
||
<paragraph id="8B83DB15A3757F65FF37FC008A66DCD2" blockId="24.[151,1437,258,2011]" pageId="24" pageNumber="325">
|
||
Head: Comparatively large, with wide neck (in males, HWmax/PWmax = 0.68–0.71, HWmin/PWmax = 0.54– 0.60; in females, these indices 0.69–0.73 and 0.57–0.60, respectively). Eyes moderately convex (in males, HWmax/HWmin = 1.21–1.30, and in females, 1.18–1.23), in lateral view somewhat wide oval. Tempora about one third as long as eye, slightly convex, sloped somewhat abruptly to neck. Genae wide, about 1.75 as wide as width of antennomere 1, with few short setae. Lateral margin of head between antenna and eye, like in all the preceding species, slightly protruded laterally, forming an obtuse angle near eyes, distinctly bordered. Area between supraorbital furrow and upper margin of eye approximately as wide as or slightly shorter than width of antennomere 3 basally. Supraorbital setigerous pore located just behind level of posterior margin of eye, separated from supraorbital furrow approximately by width of antennomere 3 apically. Labrum slightly or moderately concave anteriorly. Clypeus slightly emarginate along apical margin, slightly convex basally, flatly sloped to apex, with one setigerous pore at each apical angle. Frons and vertex convex.
|
||
<collectingRegion id="49F815F7A3757F65FC48FAB98F4BDC1E" box="[952,1047,1303,1326]" country="Myanmar" name="Chin" pageId="24" pageNumber="325">Mentum</collectingRegion>
|
||
and submentum completely fused, submentum on each side with one long seta and additionally with one short seta laterally of long seta. Left mandible not truncate, slightly blunted at apex. Dorsal microsculpture visible laterally behind eyes, consisting of a mixture of transverse and almost isodiametric meshes. Antennae in male almost extending to the basal edge of pronotum, with antennomeres 4–8 markedly longer than wide, in female antennae slightly shorter, with shorter antennomeres.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3757F65FF37FA418F92DEEB" blockId="24.[151,1437,258,2011]" pageId="24" pageNumber="325">Pronotum: Moderately wide (PWmax/PL = 1.33–1.44), widest at the end of anterior third, slightly to moderately narrowed posteriad (PWmax/PWmin = 1.14–1.28), with one lateral seta inserted slightly before widest point. Sides rounded anteriorly, almost straight or more widely rounded in basal third, in some specimens slightly sinuate in short distance just before basal angles. Apical margin slightly emarginate, finely bordered laterally. Apical angles slightly, but clearly protruded ahead, comparatively narrowly rounded at tip. Basal margin nearly straight or slightly oblique laterally, vaguely bordered laterally, markedly longer than apical margin and slightly shorter than base of elytra between humeral angles. Basal angles more or less obtuse, more or less clearly denticulate at tip, often with an acute apex laterally projected. Pronotal disc moderately convex, strongly sloped to apical angles and moderately so to basal ones, not depressed basally, but slightly flattened in comparatively wide latero-basal area. Lateral gutter very narrow, slightly widened in apical third. Median line very fine, not deepened, rather long, ended slightly before apical and basal margins. Basal foveae small, oval or somewhat elongate, very shallow, usually poorly delimited. Surface sparsely, irregularly punctate in latero-basal areas before lateral seta, with few indistinct punctures or almost impunctate at anterior margin and in middle basal portion. Microsculpture very obliterate, recognizable usually only along sides, consisting of very fine transverse meshes.</paragraph>
|
||
<caption id="DF438B9DA3747F64FF67F8358F3DDEDF" httpUri="https://zenodo.org/record/919638/files/figure.png" pageId="25" pageNumber="326" targetBox="[311,1277,200,1923]" targetPageId="25">
|
||
<paragraph id="8B83DB15A3747F64FF67F8358F3DDEDF" blockId="25.[151,1436,1947,2031]" pageId="25" pageNumber="326">
|
||
<emphasis id="B9480707A3747F64FF67F8358A0BDE81" bold="true" box="[151,343,1947,1970]" pageId="25" pageNumber="326">FIGURES 67–70.</emphasis>
|
||
Himalayan species of
|
||
<taxonomicName id="4C3CA096A3747F64FDBCF832898BDE82" authorityName="Morawitz" authorityYear="1863" box="[588,727,1948,1970]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9480707A3747F64FDBCF832898BDE82" box="[588,727,1948,1970]" italics="true" pageId="25" pageNumber="326">Trichotichnus</emphasis>
|
||
</taxonomicName>
|
||
subgenus
|
||
<taxonomicName id="4C3CA096A3747F64FCB9F83288F7DE82" authorityName="Jedlicka" authorityYear="1935" box="[841,939,1948,1970]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9480707A3747F64FCB9F83288F7DE82" box="[841,939,1948,1970]" italics="true" pageId="25" pageNumber="326">Bottchrus</emphasis>
|
||
</taxonomicName>
|
||
, photographs of habitus. 67,
|
||
<taxonomicName id="4C3CA096A3747F64FB14F8338E13DE82" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1252,1359,1948,1970]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="pusillus" status="sp. nov.">
|
||
<emphasis id="B9480707A3747F64FB14F8338E13DE82" box="[1252,1359,1948,1970]" italics="true" pageId="25" pageNumber="326">T. pusillus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9480707A3747F64FAA9F8328EC9DE81" bold="true" box="[1369,1429,1948,1969]" pageId="25" pageNumber="326">
|
||
<taxonomicNameLabel id="A27BBA7CA3747F64FAA9F8328EC9DE81" box="[1369,1429,1948,1969]" pageId="25" pageNumber="326" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, male, from the Bara Pokhari Lekh mountain range; 68,
|
||
<taxonomicName id="4C3CA096A3747F64FCD7F81288D3DEE1" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[807,911,1979,2001]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="pusillus" status="sp. nov.">
|
||
<emphasis id="B9480707A3747F64FCD7F81288D3DEE1" box="[807,911,1979,2001]" italics="true" pageId="25" pageNumber="326">T. pusillus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3747F64FC64F8158891DEE0" box="[916,973,1979,2000]" pageId="25" pageNumber="326" rank="species">
|
||
<emphasis id="B9480707A3747F64FC64F8158891DEE0" bold="true" box="[916,973,1979,2000]" pageId="25" pageNumber="326">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype, male, from the Dudh Pokhari Lekh mountain range; 69,
|
||
<taxonomicName id="4C3CA096A3747F64FE98F8748AA7DEDF" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[360,507,2009,2031]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="ganeshensis" status="sp. nov.">
|
||
<emphasis id="B9480707A3747F64FE98F8748AA7DEDF" box="[360,507,2009,2031]" italics="true" pageId="25" pageNumber="326">T. ganeshensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9480707A3747F64FDF2F8748966DEDF" bold="true" box="[514,570,2010,2031]" pageId="25" pageNumber="326">
|
||
<taxonomicNameLabel id="A27BBA7CA3747F64FDF2F8748966DEDF" box="[514,570,2010,2031]" pageId="25" pageNumber="326" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, male; 70,
|
||
<taxonomicName id="4C3CA096A3747F64FCE0F8748825DEDF" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[784,889,2010,2031]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov.">
|
||
<emphasis id="B9480707A3747F64FCE0F8748825DEDF" box="[784,889,2010,2031]" italics="true" pageId="25" pageNumber="326">T. minutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3747F64FC8FF87488E4DEDF" box="[895,952,2010,2031]" pageId="25" pageNumber="326" rank="species">
|
||
<emphasis id="B9480707A3747F64FC8FF87488E4DEDF" bold="true" box="[895,952,2010,2031]" pageId="25" pageNumber="326">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype, male.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF438B9DA3777F67FF67FBC38EC0DDD0" httpUri="https://zenodo.org/record/919642/files/figure.png" pageId="26" pageNumber="327" targetBox="[180,1420,209,1090]" targetPageId="26">
|
||
<paragraph id="8B83DB15A3777F67FF67FBC38EC0DDD0" blockId="26.[151,1436,1133,1248]" pageId="26" pageNumber="327">
|
||
<emphasis id="B9480707A3777F67FF67FBC38A0FDDB3" bold="true" box="[151,339,1133,1156]" pageId="26" pageNumber="327">FIGURES 71–79.</emphasis>
|
||
<taxonomicName id="4C3CA096A3777F67FEA9FBC089EEDDB4" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[345,690,1134,1156]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="327" phylum="Arthropoda" rank="species" species="pusillus" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3777F67FEA9FBC08AB8DDB4" box="[345,484,1134,1156]" italics="true" pageId="26" pageNumber="327">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3777F67FE02FBC08909DDB4" box="[498,597,1134,1156]" italics="true" pageId="26" pageNumber="327">Bottchrus</emphasis>
|
||
)
|
||
<emphasis id="B9480707A3777F67FD93FBC089EEDDB4" box="[611,690,1134,1156]" italics="true" pageId="26" pageNumber="327">pusillus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3777F67FD47FBC189ACDDB4" box="[695,752,1135,1156]" pageId="26" pageNumber="327" rank="species">
|
||
<emphasis id="B9480707A3777F67FD47FBC189ACDDB4" bold="true" box="[695,752,1135,1156]" pageId="26" pageNumber="327">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(71, 78, 79, paratypes from Dudh Pokhari Lekh; 72, specimen from Annapurna; 73, 74, paratype from Bara Pokhari Lekh; 75–77, holotype). 71, left metepisternum; 72–73, 75, 77, median lobe of aedeagus; 74, 76, apical capitulum; 78, apex of last (VII) abdominal sternite, male; 79, same, female. 71, 78–79, ventral view; 72–73, 75, lateral view; 74, 76, caudal view; 77, dorsal view. Scale bars = 0.25 mm (A: Figs 71–73, 75, 77–79; B: Figs 74, 76).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B83DB15A3777F67FF37FAA98981DF9C" blockId="26.[151,1437,1287,2032]" pageId="26" pageNumber="327">Elytra: In lateral and caudal view convex, in dorsal view oval, moderately wide (in males, EL/EW = 1.31–1.37; EL/PL = 2.20–2.39; EW/PWmax = 1.24–1.29; in females, these indices 1.28–1.38, 2.19–2.44, 1.21–1.30, respectively), widest just behind middle, with somewhat acute apex. Humeri prominent, angularly rounded at tip, with a small, but distinct acute denticle visible from behind. Subapical sinuation very shallow. Sutural angle acutangular in general, sharp or blunted at tip. Basal edge evenly curved ahead laterally, forming a sharp, obtuse angle with lateral margin. Elytral striae thin, usually slightly impressed; intervals flat or subconvex throughout, weakly narrowed posteriad. Parascutellar setigerous pore present, large. Parascutellar striole short, at most as long as distance from parascutellar pore to suture, with apex free or connected with stria 1. Interval 3 with a distinct discal setigerous pore at stria 2 behind middle. In many specimens elytra near apex with a short additional stria 9 separated from lateral gutter by distinct convexity. Marginal umbilicate series widely interrupted at middle, with 7– 8 pores in anterior group and 8–10 pores in posterior group. Microsculpture strongly reduced, very fine transverse meshes evident in very narrow areas along margins.</paragraph>
|
||
<paragraph id="8B83DB15A3777F67FF37F919890FDFE0" blockId="26.[151,1437,1287,2032]" box="[199,595,1719,1744]" pageId="26" pageNumber="327">Hindwings: Reduced to tiny scales.</paragraph>
|
||
<paragraph id="8B83DB15A3777F67FF37F97589C2DE0C" blockId="26.[151,1437,1287,2032]" pageId="26" pageNumber="327">
|
||
Ventral surface: Prosternum and metasternum finely pubescent. Metepisternum (
|
||
<figureCitation id="1307C790A3777F67FB90F9758FE8DFC4" box="[1120,1204,1755,1780]" captionStart="FIGURES 71 – 79" captionStartId="26.[151,264,1133,1156]" captionTargetBox="[180,1420,209,1090]" captionTargetId="figure@26.[151,1436,193,1113]" captionTargetPageId="26" captionText="FIGURES 71 – 79. Trichotichnus (Bottchrus) pusillus sp. n. (71, 78, 79, paratypes from Dudh Pokhari Lekh; 72, specimen from Annapurna; 73, 74, paratype from Bara Pokhari Lekh; 75 – 77, holotype). 71, left metepisternum; 72 – 73, 75, 77, median lobe of aedeagus; 74, 76, apical capitulum; 78, apex of last (VII) abdominal sternite, male; 79, same, female. 71, 78 – 79, ventral view; 72 – 73, 75, lateral view; 74, 76, caudal view; 77, dorsal view. Scale bars = 0.25 mm (A: Figs 71 – 73, 75, 77 – 79; B: Figs 74, 76)." httpUri="https://zenodo.org/record/919642/files/figure.png" pageId="26" pageNumber="327">Fig. 71</figureCitation>
|
||
) slightly wider than long, narrowed posteriad. Apex of last visible (VII) abdominal sternite in both sexes rounded at apex, in female (
|
||
<figureCitation id="1307C790A3777F67FF6FF88D8BB5DE0C" box="[159,233,1827,1852]" captionStart="FIGURES 71 – 79" captionStartId="26.[151,264,1133,1156]" captionTargetBox="[180,1420,209,1090]" captionTargetId="figure@26.[151,1436,193,1113]" captionTargetPageId="26" captionText="FIGURES 71 – 79. Trichotichnus (Bottchrus) pusillus sp. n. (71, 78, 79, paratypes from Dudh Pokhari Lekh; 72, specimen from Annapurna; 73, 74, paratype from Bara Pokhari Lekh; 75 – 77, holotype). 71, left metepisternum; 72 – 73, 75, 77, median lobe of aedeagus; 74, 76, apical capitulum; 78, apex of last (VII) abdominal sternite, male; 79, same, female. 71, 78 – 79, ventral view; 72 – 73, 75, lateral view; 74, 76, caudal view; 77, dorsal view. Scale bars = 0.25 mm (A: Figs 71 – 73, 75, 77 – 79; B: Figs 74, 76)." httpUri="https://zenodo.org/record/919642/files/figure.png" pageId="26" pageNumber="327">Fig 79</figureCitation>
|
||
) more angularly than in male (
|
||
<figureCitation id="1307C790A3777F67FDB3F88D89CDDE0C" box="[579,657,1827,1852]" captionStart="FIGURES 71 – 79" captionStartId="26.[151,264,1133,1156]" captionTargetBox="[180,1420,209,1090]" captionTargetId="figure@26.[151,1436,193,1113]" captionTargetPageId="26" captionText="FIGURES 71 – 79. Trichotichnus (Bottchrus) pusillus sp. n. (71, 78, 79, paratypes from Dudh Pokhari Lekh; 72, specimen from Annapurna; 73, 74, paratype from Bara Pokhari Lekh; 75 – 77, holotype). 71, left metepisternum; 72 – 73, 75, 77, median lobe of aedeagus; 74, 76, apical capitulum; 78, apex of last (VII) abdominal sternite, male; 79, same, female. 71, 78 – 79, ventral view; 72 – 73, 75, lateral view; 74, 76, caudal view; 77, dorsal view. Scale bars = 0.25 mm (A: Figs 71 – 73, 75, 77 – 79; B: Figs 74, 76)." httpUri="https://zenodo.org/record/919642/files/figure.png" pageId="26" pageNumber="327">Fig 78</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3777F67FF37F8E689DBDEC0" blockId="26.[151,1437,1287,2032]" pageId="26" pageNumber="327">Legs: Protibia very finely longitudinally sulcate on dorsal side in its basal two-third. Profemur usually with four, more rarely with five or three setae on anterio-ventral margin. Tarsi short, metatarsus noticeably shorter than HWmin, with metatarsomeres (particularly 2–4) moderately widened posteriad, metatarsomere 1 approximately as long as metatarsomeres 2+3. Tarsomere 5 with two, occasionally three pairs of ventro-lateral setae. In male, pro- and mesotarsi moderately strongly widened.</paragraph>
|
||
<paragraph id="8B83DB15A3767F66FF37FF398952D9C8" blockId="27.[151,1437,151,1149]" pageId="27" pageNumber="328">Female genitalia: Laterotergite (hemisternite) symmetrical, with two thick setae apically. Basal stylomere with one preapical spine on external margin. Apical stylomere moderately curved, with a peg-like spine at both ventral and dorsal edges of outer margin.</paragraph>
|
||
<paragraph id="8B83DB15A3767F66FF37FEAA8F52D8E0" blockId="27.[151,1437,151,1149]" pageId="27" pageNumber="328">
|
||
Male genitalia: Median lobe of aedeagus (
|
||
<figureCitation id="1307C790A3767F66FD58FEAA8876D82C" box="[680,810,260,285]" captionStart="FIGURES 71 – 79" captionStartId="26.[151,264,1133,1156]" captionTargetBox="[180,1420,209,1090]" captionTargetId="figure@26.[151,1436,193,1113]" captionTargetPageId="26" captionText="FIGURES 71 – 79. Trichotichnus (Bottchrus) pusillus sp. n. (71, 78, 79, paratypes from Dudh Pokhari Lekh; 72, specimen from Annapurna; 73, 74, paratype from Bara Pokhari Lekh; 75 – 77, holotype). 71, left metepisternum; 72 – 73, 75, 77, median lobe of aedeagus; 74, 76, apical capitulum; 78, apex of last (VII) abdominal sternite, male; 79, same, female. 71, 78 – 79, ventral view; 72 – 73, 75, lateral view; 74, 76, caudal view; 77, dorsal view. Scale bars = 0.25 mm (A: Figs 71 – 73, 75, 77 – 79; B: Figs 74, 76)." httpUri="https://zenodo.org/record/919642/files/figure.png" pageId="27" pageNumber="328">Figs 72–77</figureCitation>
|
||
) in lateral view evenly arcuate, C-shaped, with ventral margin more or less roundly emarginate throughout, with apical third strongly bent ventrad; in dorsal view median lobe almost straight, markedly narrowed to apex in distal half. Terminal lamella in dorsal view moderately short, markedly narrowed posteriad and more or less clearly sinuate at sides just at apex. Apical capitulum in lateral view (
|
||
<figureCitation id="1307C790A3767F66FF6FFE3A8A17D89D" box="[159,331,404,429]" captionStart="FIGURES 71 – 79" captionStartId="26.[151,264,1133,1156]" captionTargetBox="[180,1420,209,1090]" captionTargetId="figure@26.[151,1436,193,1113]" captionTargetPageId="26" captionText="FIGURES 71 – 79. Trichotichnus (Bottchrus) pusillus sp. n. (71, 78, 79, paratypes from Dudh Pokhari Lekh; 72, specimen from Annapurna; 73, 74, paratype from Bara Pokhari Lekh; 75 – 77, holotype). 71, left metepisternum; 72 – 73, 75, 77, median lobe of aedeagus; 74, 76, apical capitulum; 78, apex of last (VII) abdominal sternite, male; 79, same, female. 71, 78 – 79, ventral view; 72 – 73, 75, lateral view; 74, 76, caudal view; 77, dorsal view. Scale bars = 0.25 mm (A: Figs 71 – 73, 75, 77 – 79; B: Figs 74, 76)." httpUri="https://zenodo.org/record/919642/files/figure.png" pageId="27" pageNumber="328">Figs 72–73, 75</figureCitation>
|
||
) almost transverse, more strongly prominent dorsally than ventrally, in caudal view (
|
||
<figureCitation id="1307C790A3767F66FAE2FE3A8EC8D89D" box="[1298,1428,404,429]" captionStart="FIGURES 71 – 79" captionStartId="26.[151,264,1133,1156]" captionTargetBox="[180,1420,209,1090]" captionTargetId="figure@26.[151,1436,193,1113]" captionTargetPageId="26" captionText="FIGURES 71 – 79. Trichotichnus (Bottchrus) pusillus sp. n. (71, 78, 79, paratypes from Dudh Pokhari Lekh; 72, specimen from Annapurna; 73, 74, paratype from Bara Pokhari Lekh; 75 – 77, holotype). 71, left metepisternum; 72 – 73, 75, 77, median lobe of aedeagus; 74, 76, apical capitulum; 78, apex of last (VII) abdominal sternite, male; 79, same, female. 71, 78 – 79, ventral view; 72 – 73, 75, lateral view; 74, 76, caudal view; 77, dorsal view. Scale bars = 0.25 mm (A: Figs 71 – 73, 75, 77 – 79; B: Figs 74, 76)." httpUri="https://zenodo.org/record/919642/files/figure.png" pageId="27" pageNumber="328">Figs 74, 76</figureCitation>
|
||
) discoidal, rounded at tip. Internal sac without spines and distinct spiny patches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA3767F66FF37FE728A14DADC" pageId="27" pageNumber="328" type="etymology">
|
||
<paragraph id="8B83DB15A3767F66FF37FE728A00DB28" blockId="27.[151,1437,151,1149]" pageId="27" pageNumber="328">
|
||
<emphasis id="B9480707A3767F66FF37FE728A1AD8C5" bold="true" box="[199,326,476,501]" pageId="27" pageNumber="328">Etymology</emphasis>
|
||
. The specific name is derived from the Latin adjective meaning “tiny” and refers to the small body size of the beetle.
|
||
</paragraph>
|
||
<paragraph id="8B83DB15A3767F66FF37FD8A8A14DADC" blockId="27.[151,1437,151,1149]" pageId="27" pageNumber="328">
|
||
<emphasis id="B9480707A3767F66FF37FD8A8A06DB0D" bold="true" box="[199,346,548,573]" pageId="27" pageNumber="328">Comparison</emphasis>
|
||
. In size and general habitus,
|
||
<taxonomicName id="4C3CA096A3767F66FD5BFD8B880EDB0D" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[683,850,548,573]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="pusillus" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3767F66FD5BFD8B899CDB0C" box="[683,704,549,572]" italics="true" pageId="27" pageNumber="328">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3767F66FD21FD8B89BEDB0C" box="[721,738,549,572]" italics="true" pageId="27" pageNumber="328">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3767F66FD0BFD8A880EDB0D" box="[763,850,548,573]" italics="true" pageId="27" pageNumber="328">pusillus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3767F66FCABFD8A88C0DB0D" box="[859,924,548,573]" pageId="27" pageNumber="328" rank="species">
|
||
<emphasis id="B9480707A3767F66FCABFD8A88C0DB0D" bold="true" box="[859,924,548,573]" pageId="27" pageNumber="328">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is very similar to
|
||
<taxonomicName id="4C3CA096A3767F66FB9EFD8B8E0FDB0D" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1134,1363,548,573]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="baglungensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3767F66FB9EFD8B8FD8DB0C" box="[1134,1156,549,572]" italics="true" pageId="27" pageNumber="328">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3767F66FB65FD8B8FFADB0C" box="[1173,1190,549,572]" italics="true" pageId="27" pageNumber="328">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3767F66FB4FFD8A8E0FDB0D" box="[1215,1363,548,573]" italics="true" pageId="27" pageNumber="328">baglungensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3767F66FAABFD8A8EC0DB0D" box="[1371,1436,548,573]" pageId="27" pageNumber="328" rank="species">
|
||
<emphasis id="B9480707A3767F66FAABFD8A8EC0DB0D" bold="true" box="[1371,1436,548,573]" pageId="27" pageNumber="328">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and slightly more robust than
|
||
<taxonomicName id="4C3CA096A3767F66FE16FDE789B8DB50" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[486,740,583,608]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="panchhaseensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3767F66FE16FDE78AAFDB50" box="[486,499,585,608]" italics="true" pageId="27" pageNumber="328">T</emphasis>
|
||
. (
|
||
<emphasis id="B9480707A3767F66FDFCFDE78941DB50" box="[524,541,585,608]" italics="true" pageId="27" pageNumber="328">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3767F66FDC4FDE989B8DB50" box="[564,740,583,608]" italics="true" pageId="27" pageNumber="328">panchhaseensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3767F66FD1BFDE98876DB50" box="[747,810,583,608]" pageId="27" pageNumber="328" rank="species">
|
||
<emphasis id="B9480707A3767F66FD1BFDE98876DB50" bold="true" box="[747,810,583,608]" pageId="27" pageNumber="328">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C3CA096A3767F66FC91FDE78F78DB50" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[865,1060,583,608]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="siklesensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3767F66FC91FDE7882ADB50" box="[865,886,585,608]" italics="true" pageId="27" pageNumber="328">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3767F66FC75FDE788CADB50" box="[901,918,585,608]" italics="true" pageId="27" pageNumber="328">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3767F66FC5EFDE98F78DB50" box="[942,1060,583,608]" italics="true" pageId="27" pageNumber="328">siklesensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3767F66FBDCFDE98F37DB50" box="[1068,1131,583,608]" pageId="27" pageNumber="328" rank="species">
|
||
<emphasis id="B9480707A3767F66FBDCFDE98F37DB50" bold="true" box="[1068,1131,583,608]" pageId="27" pageNumber="328">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<taxonomicName id="4C3CA096A3767F66FB82FDE98EC0DB50" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[1138,1436,583,608]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="pusillus" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3767F66FB82FDE98E50DB50" box="[1138,1292,583,608]" italics="true" pageId="27" pageNumber="328">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3767F66FAECFDE78E71DB50" box="[1308,1325,585,608]" italics="true" pageId="27" pageNumber="328">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3767F66FAB5FDE98EC0DB50" box="[1349,1436,583,608]" italics="true" pageId="27" pageNumber="328">pusillus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3767F66FF67FDC28B82DBB5" box="[151,222,620,645]" pageId="27" pageNumber="328" rank="species">
|
||
<emphasis id="B9480707A3767F66FF67FDC28B82DBB5" bold="true" box="[151,222,620,645]" pageId="27" pageNumber="328">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is also characterized by metepisternum slightly wider than long (
|
||
<figureCitation id="1307C790A3767F66FBF2FDC28F00DBB4" box="[1026,1116,620,645]" captionStart="FIGURES 71 – 79" captionStartId="26.[151,264,1133,1156]" captionTargetBox="[180,1420,209,1090]" captionTargetId="figure@26.[151,1436,193,1113]" captionTargetPageId="26" captionText="FIGURES 71 – 79. Trichotichnus (Bottchrus) pusillus sp. n. (71, 78, 79, paratypes from Dudh Pokhari Lekh; 72, specimen from Annapurna; 73, 74, paratype from Bara Pokhari Lekh; 75 – 77, holotype). 71, left metepisternum; 72 – 73, 75, 77, median lobe of aedeagus; 74, 76, apical capitulum; 78, apex of last (VII) abdominal sternite, male; 79, same, female. 71, 78 – 79, ventral view; 72 – 73, 75, lateral view; 74, 76, caudal view; 77, dorsal view. Scale bars = 0.25 mm (A: Figs 71 – 73, 75, 77 – 79; B: Figs 74, 76)." httpUri="https://zenodo.org/record/919642/files/figure.png" pageId="27" pageNumber="328">Fig. 71</figureCitation>
|
||
), mentum and submentum completely fused, pronotal lateral gutter slightly widened in apical third, humerus with a distinct acute denticle at apex, protibia very finely longitudinally sulcate on dorsal side, and microsculpture indistinct on pronotal and elytral discs. It is distinctly distinguished from all other species of
|
||
<taxonomicName id="4C3CA096A3767F66FC40FD798F41DBC0" authorityName="Jedlicka" authorityYear="1935" box="[944,1053,727,752]" class="Insecta" family="Carabidae" genus="Bottchrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9480707A3767F66FC40FD798F41DBC0" box="[944,1053,727,752]" italics="true" pageId="27" pageNumber="328">Bottchrus</emphasis>
|
||
</taxonomicName>
|
||
of the Himalayan region by the peculiar male genitalia with C-shaped aedeagal median lobe (
|
||
<figureCitation id="1307C790A3767F66FC91FD5288B9DA24" box="[865,997,764,789]" captionStart="FIGURES 71 – 79" captionStartId="26.[151,264,1133,1156]" captionTargetBox="[180,1420,209,1090]" captionTargetId="figure@26.[151,1436,193,1113]" captionTargetPageId="26" captionText="FIGURES 71 – 79. Trichotichnus (Bottchrus) pusillus sp. n. (71, 78, 79, paratypes from Dudh Pokhari Lekh; 72, specimen from Annapurna; 73, 74, paratype from Bara Pokhari Lekh; 75 – 77, holotype). 71, left metepisternum; 72 – 73, 75, 77, median lobe of aedeagus; 74, 76, apical capitulum; 78, apex of last (VII) abdominal sternite, male; 79, same, female. 71, 78 – 79, ventral view; 72 – 73, 75, lateral view; 74, 76, caudal view; 77, dorsal view. Scale bars = 0.25 mm (A: Figs 71 – 73, 75, 77 – 79; B: Figs 74, 76)." httpUri="https://zenodo.org/record/919642/files/figure.png" pageId="27" pageNumber="328">Figs 72–77</figureCitation>
|
||
) which is bent ventrad approximately equally basally and apically; its terminal lamella is moderately long, slightly widened apically and supplied with a small transverse apical capitulum; the latter is discoidal in caudal view (
|
||
<figureCitation id="1307C790A3767F66FBEFFCEA8FF1DA6D" box="[1055,1197,836,861]" captionStart="FIGURES 71 – 79" captionStartId="26.[151,264,1133,1156]" captionTargetBox="[180,1420,209,1090]" captionTargetId="figure@26.[151,1436,193,1113]" captionTargetPageId="26" captionText="FIGURES 71 – 79. Trichotichnus (Bottchrus) pusillus sp. n. (71, 78, 79, paratypes from Dudh Pokhari Lekh; 72, specimen from Annapurna; 73, 74, paratype from Bara Pokhari Lekh; 75 – 77, holotype). 71, left metepisternum; 72 – 73, 75, 77, median lobe of aedeagus; 74, 76, apical capitulum; 78, apex of last (VII) abdominal sternite, male; 79, same, female. 71, 78 – 79, ventral view; 72 – 73, 75, lateral view; 74, 76, caudal view; 77, dorsal view. Scale bars = 0.25 mm (A: Figs 71 – 73, 75, 77 – 79; B: Figs 74, 76)." httpUri="https://zenodo.org/record/919642/files/figure.png" pageId="27" pageNumber="328">Figs 74–76</figureCitation>
|
||
).
|
||
<taxonomicName id="4C3CA096A3767F66FB37FCEA8A72DAB0" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="manasluensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3767F66FB37FCEA8E3DDA6D" box="[1223,1377,836,861]" italics="true" pageId="27" pageNumber="328">Trichotichnus</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3767F66FA8AFCEB8ED7DA6C" box="[1402,1419,837,860]" italics="true" pageId="27" pageNumber="328">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3767F66FF67FCC98A72DAB0" box="[151,302,871,896]" italics="true" pageId="27" pageNumber="328">manasluensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3767F66FEC7FCC98A25DAB0" box="[311,377,871,896]" pageId="27" pageNumber="328" rank="species">
|
||
<emphasis id="B9480707A3767F66FEC7FCC98A25DAB0" bold="true" box="[311,377,871,896]" pageId="27" pageNumber="328">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, occurring sympatricly with
|
||
<taxonomicName id="4C3CA096A3767F66FD37FCC7882CDAB0" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[711,880,871,896]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="pusillus" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3767F66FD37FCC78986DAB0" box="[711,730,873,896]" italics="true" pageId="27" pageNumber="328">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3767F66FD1EFCC789A3DAB0" box="[750,767,873,896]" italics="true" pageId="27" pageNumber="328">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3767F66FCE9FCC9882CDAB0" box="[793,880,871,896]" italics="true" pageId="27" pageNumber="328">pusillus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3767F66FC8AFCC988E7DAB0" box="[890,955,871,896]" pageId="27" pageNumber="328" rank="species">
|
||
<emphasis id="B9480707A3767F66FC8AFCC988E7DAB0" bold="true" box="[890,955,871,896]" pageId="27" pageNumber="328">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, is easily distinguished from the latter by having larger size (
|
||
<quantity id="4CC476F0A3767F66FE8AFC228957DA94" box="[378,523,908,933]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" metricValueMax="6.3" metricValueMin="5.9" pageId="27" pageNumber="328" unit="mm" value="6.1" valueMax="6.3" valueMin="5.9">5.9–6.3 mm</quantity>
|
||
), humerus without denticle, last (VII) abdominal sternite in female somewhat angulate at apex (
|
||
<figureCitation id="1307C790A3767F66FEAEFC018AE7DAF8" box="[350,443,943,968]" captionStart="FIGURES 109 – 114" captionStartId="38.[151,264,1913,1936]" captionTargetBox="[212,1384,835,1874]" captionTargetId="figure@38.[187,1400,823,1891]" captionTargetPageId="38" captionText="FIGURES 109 – 114. Trichotichnus (Bottchrus) manasluensis sp. n. (109, 111, paratypes; 112 – 114, holotype). 109, left metepisternum; 110, apex of last (VII) abdominal sternite, male; 111, same, female; 112, 114, median lobe of aedeagus; 113, apical capitulum. 109 – 111, ventral view; 112, lateral view; 113, caudal view; 114, dorsal view. Scale bars = 0.25 mm (A: Figs 109 – 112, 114; B: Fig. 113)." httpUri="https://zenodo.org/record/919664/files/figure.png" pageId="27" pageNumber="328">Fig. 111</figureCitation>
|
||
) and median lobe of the aedeagus in lateral view almost straight, only slightly S-shaped (
|
||
<figureCitation id="1307C790A3767F66FF6FFC7A8A67DADC" box="[159,315,980,1005]" captionStart="FIGURES 109 – 114" captionStartId="38.[151,264,1913,1936]" captionTargetBox="[212,1384,835,1874]" captionTargetId="figure@38.[187,1400,823,1891]" captionTargetPageId="38" captionText="FIGURES 109 – 114. Trichotichnus (Bottchrus) manasluensis sp. n. (109, 111, paratypes; 112 – 114, holotype). 109, left metepisternum; 110, apex of last (VII) abdominal sternite, male; 111, same, female; 112, 114, median lobe of aedeagus; 113, apical capitulum. 109 – 111, ventral view; 112, lateral view; 113, caudal view; 114, dorsal view. Scale bars = 0.25 mm (A: Figs 109 – 112, 114; B: Fig. 113)." httpUri="https://zenodo.org/record/919664/files/figure.png" pageId="27" pageNumber="328">Figs 112, 114</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C326889EA3767F66FF37FC598E3FDD4C" pageId="27" pageNumber="328" type="distribution">
|
||
<paragraph id="8B83DB15A3767F66FF37FC598E3FDD4C" blockId="27.[151,1437,151,1149]" pageId="27" pageNumber="328">
|
||
<emphasis id="B9480707A3767F66FF37FC598A05DD20" bold="true" box="[199,345,1015,1040]" pageId="27" pageNumber="328">Distribution</emphasis>
|
||
(
|
||
<figureCitation id="1307C790A3767F66FE98FC598A94DD20" box="[360,456,1015,1040]" captionStart="FIGURE 151" captionStartId="53.[151,250,1080,1102]" captionTargetBox="[151,1436,339,1059]" captionTargetId="figure@53.[151,1436,339,1059]" captionTargetPageId="53" captionText="FIGURE 151. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. holzschuhi Kirschenhofer, 1992; 2, T. baglungensis sp. n.; 3, T. parvulus sp. n.; 4, T. bubsaensis sp. n.; 5, T. schawalleri sp. n.; 6, T. obliquebasalis sp. n.; 7, T. panchhaseensis sp. n.; 8, T. siklesensis sp. n.; 9, T. pusillus sp. n.; 10, T. ganeshensis sp. n.; 11, T. minutus sp. n.; 12, T. cyanescens Ito, 1998; 13, T. martensi sp. n.; 14, T. opacus Ito, 1998." httpUri="https://zenodo.org/record/919694/files/figure.png" pageId="27" pageNumber="328">Fig. 151</figureCitation>
|
||
,
|
||
<figureCitation id="1307C790A3767F66FE24FC568AB8DD20" box="[468,484,1016,1040]" captionStart="FIGURES 5 – 13" captionStartId="6.[151,264,1196,1219]" captionTargetBox="[152,1425,180,1152]" captionTargetId="figure@6.[151,1436,180,1193]" captionTargetPageId="6" captionText="FIGURES 5 – 13. Trichotichnus (Bottchrus) holzschuhi Kirschenhofer, 1992 (5 – 7, paratype; 8 – 13, specimens from Sete). 5, terminal lamella of aedeagus; 6 – 8, 11, median lobe of aedeagus; 9, apical capitulum; 10, left metepisternum; 12, apex of last (VII) abdominal sternite, male; 13, same, female. 5, 7, 8, dorsal view; 6, 11, lateral view; 9, caudal view; 10, 12, 13, ventral view. Scale bars = 0.25 mm (A: Figs 5 – 8, 10 – 13; B: Fig. 9)." httpUri="https://zenodo.org/record/919610/files/figure.png" pageId="27" pageNumber="328">9</figureCitation>
|
||
). Dudh Pokhari Lekh and Bara Pokhari Lekh mountain ranges on the southern slopes of Manaslu Himal, western
|
||
<collectingRegion id="49F815F7A3767F66FE11FBB28968DD05" box="[481,564,1052,1077]" country="Sri Lanka" name="Central" pageId="27" pageNumber="328">Central</collectingRegion>
|
||
<collectingCountry id="F32B9B85A3767F66FDCFFBB289D9DD05" box="[575,645,1052,1077]" name="Nepal" pageId="27" pageNumber="328">Nepal</collectingCountry>
|
||
, Lamjung District, at altitudes of
|
||
<quantity id="4CC476F0A3767F66FBE9FBB28FE0DD04" box="[1049,1212,1052,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.6" metricValueMin="2.0" pageId="27" pageNumber="328" unit="m" value="2300.0" valueMax="2600.0" valueMin="2000.0">2000–2600 m</quantity>
|
||
. Also found in the Telbrung Danda mountain range on southern slope of Lamjung Himal (south-eastern slope of Annapurna Himal). In the Manaslu Himal,
|
||
<taxonomicName id="4C3CA096A3767F66FE6AFBCB8962DD4D" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[410,574,1124,1149]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="pusillus" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3767F66FE6AFBCB8AF2DD4C" box="[410,430,1125,1148]" italics="true" pageId="27" pageNumber="328">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3767F66FE4EFBCB8A93DD4C" box="[446,463,1125,1148]" italics="true" pageId="27" pageNumber="328">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3767F66FE17FBCA8962DD4D" box="[487,574,1124,1149]" italics="true" pageId="27" pageNumber="328">pusillus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3767F66FDB5FBCA89D9DD4D" box="[581,645,1124,1149]" pageId="27" pageNumber="328" rank="species">
|
||
<emphasis id="B9480707A3767F66FDB5FBCA89D9DD4D" bold="true" box="[581,645,1124,1149]" pageId="27" pageNumber="328">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
occurs sympatricly with
|
||
<taxonomicName id="4C3CA096A3767F66FC6FFBCB8FDFDD4D" authority="Schmidt, 2017" authorityName="Schmidt" authorityYear="2017" box="[927,1155,1124,1149]" class="Insecta" family="Carabidae" genus="Trichotichnus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="328" phylum="Arthropoda" rank="species" species="manasluensis" status="sp. nov." subGenus="Bottchrus">
|
||
<emphasis id="B9480707A3767F66FC6FFBCB88EFDD4C" box="[927,947,1125,1148]" italics="true" pageId="27" pageNumber="328">T.</emphasis>
|
||
(
|
||
<emphasis id="B9480707A3767F66FC34FBCB8889DD4C" box="[964,981,1125,1148]" italics="true" pageId="27" pageNumber="328">B</emphasis>
|
||
.)
|
||
<emphasis id="B9480707A3767F66FC1CFBCA8FDFDD4D" box="[1004,1155,1124,1149]" italics="true" pageId="27" pageNumber="328">manasluensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27BBA7CA3767F66FB7AFBCA8F96DD4D" box="[1162,1226,1124,1149]" pageId="27" pageNumber="328" rank="species">
|
||
<emphasis id="B9480707A3767F66FB7AFBCA8F96DD4D" bold="true" box="[1162,1226,1124,1149]" pageId="27" pageNumber="328">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="1307C790A3767F66FB29FBCA8E66DD4D" box="[1241,1338,1124,1149]" captionStart="FIGURE 152" captionStartId="53.[151,250,1975,1997]" captionTargetBox="[151,1436,1234,1954]" captionTargetId="figure@53.[151,1436,1234,1954]" captionTargetPageId="53" captionText="FIGURE 152. Distribution of the Himalayan species of Trichotichnus subgenus Bottchrus. 1, T. gupchiensis sp. n.; 2, T. brancuccii sp. n.; 3, T. manasluensis sp. n.; 4, T. loebli Ito, 1998; 5, T. sikkimensis sp. n.; 6, T. hingstoni Andrewes, 1930." httpUri="https://zenodo.org/record/919698/files/figure.png" pageId="27" pageNumber="328">Fig. 152</figureCitation>
|
||
,
|
||
<emphasis id="B9480707A3767F66FAB5FBCA8E0FDD4C" box="[1349,1363,1124,1148]" italics="true" pageId="27" pageNumber="328">3</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |