179 lines
22 KiB
XML
179 lines
22 KiB
XML
<document id="9CC6927323B50888EBBD8C91E6505B51" ID-CLB-Dataset="116165" ID-DOI="10.11646/zootaxa.5171.1.1" ID-GBIF-Dataset="9923ae54-fea7-4394-9c56-f5f4db376e0d" ID-ISSN="1175-5326" ID-Zenodo-Dep="6966207" ID-ZooBank="5B00C374-33B0-4433-95A0-DC9B5FFC5B0C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1659689746521" checkinUser="plazi" docAuthor="Borowiec, Lech & Świętojańska, Jolanta" docDate="2022" docId="E417E526B17BA90FFF6CFAE4080F93CE" docLanguage="en" docName="zootaxa.5171.1.1.pdf" docOrigin="Zootaxa 5171 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5171.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cassida blanda Spaeth 1933" docType="treatment" docVersion="7" lastPageNumber="37" masterDocId="182E9D5EB15FA92BFFFBFA730C63955A" masterDocTitle="A monograph of the Afrotropical Cassidinae (Coleoptera: Chrysomelidae). Part 6. Revision of the tribe Cassidini 3, the genus Cassida L." masterLastPageNumber="250" masterPageNumber="1" pageNumber="37" updateTime="1699364714896" updateUser="ExternalLinkService">
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<mods:titleInfo id="5D7D3C60EE4A02CD0C5BC689D13A184F">
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<mods:title id="A78B1A22F92C6879ECC02E3F600C1511">A monograph of the Afrotropical Cassidinae (Coleoptera: Chrysomelidae). Part 6. Revision of the tribe Cassidini 3, the genus Cassida L.</mods:title>
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<mods:namePart id="AF798D3E5092E04E3E0FE7C4AC44D408">Borowiec, Lech</mods:namePart>
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<mods:nameIdentifier id="946327D3D59927FF546F00D1D64A1CD9" type="email">lech.borowiec@uwr.edu.pl</mods:nameIdentifier>
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<mods:namePart id="97345649532DEC77D6A310DDC00ED2D0">Świętojańska, Jolanta</mods:namePart>
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<mods:nameIdentifier id="66A5065C1ECE32626A1DBFFEE835712B" type="email">jolanta.swietojanska@uwr.edu.pl</mods:nameIdentifier>
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<mods:title id="39B41C45599486C558D9096E34A7FAC6">Zootaxa</mods:title>
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<mods:part id="53D2208D18040A77C25D3F80E0387186">
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<mods:date id="7FED8DE5D6A65AED06DEA692A17E7F84">2022</mods:date>
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<mods:number id="59943F1E0FCDA22CC41FA92174ECF48A">2022-08-04</mods:number>
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<mods:number id="1B3F5F7DCED20BB1ABDAD55937964151">5171</mods:number>
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<treatment id="E417E526B17BA90FFF6CFAE4080F93CE" ID-DOI="http://doi.org/10.5281/zenodo.6966236" ID-GBIF-Taxon="198839401" ID-Zenodo-Dep="6966236" LSID="urn:lsid:plazi:treatment:E417E526B17BA90FFF6CFAE4080F93CE" httpUri="http://treatment.plazi.org/id/E417E526B17BA90FFF6CFAE4080F93CE" lastPageNumber="37" pageId="36" pageNumber="37">
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<subSubSection id="24A407BBB17BA90FFF6CFAE40D9B95EB" box="[151,504,151,178]" pageId="36" pageNumber="37" type="nomenclature">
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<paragraph id="6C015430B17BA90FFF6CFAE40D9B95EB" blockId="36.[151,504,151,213]" box="[151,504,151,178]" pageId="36" pageNumber="37">
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<heading id="3749E35CB17BA90FFF6CFAE40D9B95EB" bold="true" box="[151,504,151,178]" fontSize="11" level="2" pageId="36" pageNumber="37" reason="4">
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<taxonomicName id="ABBE2FB3B17BA90FFF6CFAE40D9B95EB" ID-CoL="RMC6" authority="Spaeth, 1933" authorityName="Spaeth" authorityYear="1933" box="[151,504,151,178]" class="Insecta" family="Chrysomelidae" genus="Cassida" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="blanda">
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<emphasis id="5ECA8822B17BA90FFF6CFAE40D9B95EB" bold="true" box="[151,504,151,178]" pageId="36" pageNumber="37">
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<emphasis id="5ECA8822B17BA90FFF6CFAE40D3395E8" bold="true" box="[151,336,151,178]" italics="true" pageId="36" pageNumber="37">Cassida blanda</emphasis>
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Spaeth, 1933
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="24A407BBB17BA90FFF6CFAC80D27958F" box="[151,324,187,213]" pageId="36" pageNumber="37" type="description">
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<paragraph id="6C015430B17BA90FFF6CFAC80D27958F" blockId="36.[151,504,151,213]" box="[151,324,187,213]" pageId="36" pageNumber="37">
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(
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<figureCitation id="F48548B5B17BA90FFF64FAC80C8A958F" box="[159,233,187,213]" captionStart="FIGURE 2" captionStartId="24.[151,250,1967,1991]" captionTargetBox="[159,1426,753,1933]" captionTargetId="figure-292@24.[151,1436,741,1944]" captionTargetPageId="24" captionText="FIGURE 2. Distribution of Cassida altiuscula (black circles), Cassida blanda (black square) and Cassida procurva (black diamond)." figureDoi="http://doi.org/10.5281/zenodo.6966250" httpUri="https://zenodo.org/record/6966250/files/figure.png" pageId="36" pageNumber="37">figs. 2</figureCitation>
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, 67–68)
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</paragraph>
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</subSubSection>
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<subSubSection id="24A407BBB17BA90FFF6CFB730E8C9442" box="[151,751,255,280]" pageId="36" pageNumber="37" type="reference_group">
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<paragraph id="6C015430B17BA90FFF6CFB730E8C9442" blockId="36.[151,751,255,280]" box="[151,751,255,280]" pageId="36" pageNumber="37">
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<treatmentCitationGroup id="4CAE731EB17BA90FFF6CFB730E8C9442" box="[151,751,255,280]" pageId="36" pageNumber="37">
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<treatmentCitation id="ED1F7221B17BA90FFF6CFB730E6B944D" author="Spaeth, F." box="[151,520,255,280]" page="350" pageId="36" pageNumber="37" year="1933">
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<taxonomicName id="ABBE2FB3B17BA90FFF6CFB730E6B944D" ID-CoL="RMC6" authority="Spaeth, 1933 a: 350" authorityName="Spaeth" authorityPageNumber="350" authorityYear="1933" box="[151,520,255,280]" class="Insecta" family="Chrysomelidae" genus="Cassida" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="blanda">
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<emphasis id="5ECA8822B17BA90FFF6CFB730D569442" box="[151,309,256,280]" italics="true" pageId="36" pageNumber="37">Cassida blanda</emphasis>
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<bibRefCitation id="082F29C1B17BA90FFEC7FB730E6B944D" author="Spaeth, F." box="[316,520,255,280]" pageId="36" pageNumber="37" pagination="345 - 359" refId="ref188213" refString="Spaeth, F. (1933 a) Neue Beitrage zur Kenntnis der Afrikanischen Cassidinen (Col. Chrys.). Revue de Zoologie et de Botanique Africaines, 22, 345 - 359." type="journal article" year="1933">Spaeth, 1933 a: 350</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
;
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<treatmentCitation id="ED1F7221B17BA90FFDE9FB730E889442" author="Borowiec, L." box="[530,747,256,280]" page="241" pageId="36" pageNumber="37" year="1999">
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<bibRefCitation id="082F29C1B17BA90FFDE9FB730E889442" author="Borowiec, L." box="[530,747,256,280]" pageId="36" pageNumber="37" refId="ref184613" refString="Borowiec, L. (1999) A world catalogue of the Cassidinae (Coleoptera: Chrysomelidae). Biologica Silesiae, Wroclaw, 476 pp." type="book" year="1999">Borowiec, 1999: 241</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="24A407BBB17BA90FFF6CFB360D059076" pageId="36" pageNumber="37" type="description">
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<paragraph id="6C015430B17BA90FFF6CFB360C9994DE" blockId="36.[151,1437,325,1684]" pageId="36" pageNumber="37">
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<emphasis id="5ECA8822B17BA90FFF6CFB360D4A9405" bold="true" box="[151,297,325,351]" pageId="36" pageNumber="37">Description.</emphasis>
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L: 8.65 mm, W: 6.80 mm, Lp: 2.80 mm, Wp: 5.40 mm, L/W: 1.27, Wp/Lp: 1.93. Body short–oval (fig. 67).
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</paragraph>
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<paragraph id="6C015430B17BA90FFF3CFBFD0DB8974E" blockId="36.[151,1437,325,1684]" pageId="36" pageNumber="37">Pronotum and scutellum ochraceous, disc rusty yellow, all punctures with darker centre and some with ochraceous areola, especially in postscutellar impression, half length of disc and on slope (figs. 67, 68). Explanate margin rusty yellow. Head, ventrites and legs yellow. Two basal antennal segments yellow, rest of antennae broken in the only known specimen.</paragraph>
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<paragraph id="6C015430B17BA90FFF3CF86D09F597FE" blockId="36.[151,1437,325,1684]" pageId="36" pageNumber="37">Pronotum transverse, with maximum width behind the middle, anterior margin regularly convex, sides narrowly angulate, no basal corners but apices of angulate sides placed close to humeral angle. Disc indistinctly bordered from explanate margin except small lateral impressions, lateral lobes absent. Surface of disc shiny, impunctate, only with sparse pricks. Explanate margin broad, impunctate, shiny, transparent with well visible honeycomb structure.</paragraph>
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<paragraph id="6C015430B17BA90FFF3CF8DD0E2F9156" blockId="36.[151,1437,325,1684]" pageId="36" pageNumber="37">Base of elytra not wider than base of pronotum, humeral angles moderately protruding anterad, subangulate. Disc strongly convex in profile, with postscutellar impressions and high, tuberculate X–shaped postscutellar elevation and some relief (fig. 68). Punctation fine, arranged in regular rows but interrupted by elytral relief, distance between punctures from as wide as to thrice wider than puncture diameter. Marginal row distinct, its punctures only slightly coarser than punctures in central rows. Intervals on slope slightly convex, on sides flat, in sutural half of disc thrice wider, on sides twice wider than rows, marginal interval broad, as wide as submarginal interval and submarginal row combined, with broad and only slightly convex humeral and lateral folds. Explanate margin moderately narrow, moderately declivous with external part tend to be subhorizontal, in the widest part six times narrower than disc, surface shiny with very shallow and sparse punctation and transverse grooves, appears regular, semitransparent but with well visible honeycomb structure.</paragraph>
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<paragraph id="6C015430B17BA90FFF3CFE6508E991C6" blockId="36.[151,1437,325,1684]" pageId="36" pageNumber="37">Eyes very large, gena obsolete. Clypeus very broad, approximately 1.8 times as wide as long. Clypeal grooves very fine, in basal part hardly marked, run close to margin of eye, on apex converging in broad arch, surface of clypeus convex before antennal insertions, shiny, impunctate, clypeal grooves with row of small, setose punctures. Labrum narrowly emarginate to 1/5 length. Antennae broken in the only known specimen.</paragraph>
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<paragraph id="6C015430B17BA90FFF3CFED508689052" blockId="36.[151,1437,325,1684]" pageId="36" pageNumber="37">Prosternum narrow in the middle, strongly expanded apically, between coxae shallowly impressed, shiny, without special sculpture but with very small sparse, punctures armed with long hair, expanded apex slightly convex centrally, with several punctures armed with hair and slightly irregular surface.</paragraph>
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<paragraph id="6C015430B17BA90FFF3CFF610D059076" blockId="36.[151,1437,325,1684]" box="[199,358,1298,1324]" pageId="36" pageNumber="37">Claws simple.</paragraph>
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</subSubSection>
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<subSubSection id="24A407BBB17BA90FFF3CFF460E44900A" box="[199,551,1333,1360]" pageId="36" pageNumber="37" type="distribution">
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<paragraph id="6C015430B17BA90FFF3CFF460E44900A" blockId="36.[151,1437,325,1684]" box="[199,551,1333,1360]" pageId="36" pageNumber="37">
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<emphasis id="5ECA8822B17BA90FFF3CFF460D029015" bold="true" box="[199,353,1333,1359]" pageId="36" pageNumber="37">Distribution.</emphasis>
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<collectingCountry id="14A914A0B17BA90FFE9CFF450DAF900A" box="[359,460,1334,1360]" name="Tanzania" pageId="36" pageNumber="37">Tanzania</collectingCountry>
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(
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<figureCitation id="F48548B5B17BA90FFE20FF450E79900A" box="[475,538,1334,1360]" captionStart="FIGURE 2" captionStartId="24.[151,250,1967,1991]" captionTargetBox="[159,1426,753,1933]" captionTargetId="figure-292@24.[151,1436,741,1944]" captionTargetPageId="24" captionText="FIGURE 2. Distribution of Cassida altiuscula (black circles), Cassida blanda (black square) and Cassida procurva (black diamond)." figureDoi="http://doi.org/10.5281/zenodo.6966250" httpUri="https://zenodo.org/record/6966250/files/figure.png" pageId="36" pageNumber="37">fig. 2</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="24A407BBB17BA90FFF3CFF290FD7932A" pageId="36" pageNumber="37" type="discussion">
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<paragraph id="6C015430B17BA90FFF3CFF290FD7932A" blockId="36.[151,1437,325,1684]" pageId="36" pageNumber="37">
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<emphasis id="5ECA8822B17BA90FFF3CFF290D58902E" bold="true" box="[199,315,1370,1396]" pageId="36" pageNumber="37">Remarks.</emphasis>
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A very distinct species. The large size, narrowly angulate pronotal sides, base of elytra not wider than pronotum, and strongly convex elytral disc with a distinct postscutellar hump (figs. 67, 68) distinguish this species from all African members of the genus
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<taxonomicName id="ABBE2FB3B17BA90FFD53FFD10F6190E6" authorityName="Linnaeus" authorityYear="1758" box="[680,770,1442,1468]" class="Insecta" family="Chrysomelidae" genus="Cassida" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
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<emphasis id="5ECA8822B17BA90FFD53FFD10F6190E6" box="[680,770,1442,1468]" italics="true" pageId="36" pageNumber="37">Cassida</emphasis>
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</taxonomicName>
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. A similar size and more or less distinct postscutellar hump are known only in species of the
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<taxonomicName id="ABBE2FB3B17BA90FFE04FFB50EF690BA" authorityName="Klug" authorityYear="1835" box="[511,661,1478,1504]" class="Insecta" family="Chrysomelidae" genus="Cassida" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="tosta">
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<emphasis id="5ECA8822B17BA90FFE04FFB50EF690BA" box="[511,661,1478,1504]" italics="true" pageId="36" pageNumber="37">Cassida tosta</emphasis>
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</taxonomicName>
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group (
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<taxonomicName id="ABBE2FB3B17BA90FFD17FFB50F1A90BA" authorityName="Spaeth" authorityYear="1916" box="[748,889,1478,1504]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="altiuscula">
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<emphasis id="5ECA8822B17BA90FFD17FFB50F1A90BA" box="[748,889,1478,1504]" italics="true" pageId="36" pageNumber="37">C. altiuscula</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="ABBE2FB3B17BA90FFC7DFFB5087890BA" authorityName="Thomson" authorityYear="1858" box="[902,1051,1478,1504]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="inaequalis">
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<emphasis id="5ECA8822B17BA90FFC7DFFB5087890BA" box="[902,1051,1478,1504]" italics="true" pageId="36" pageNumber="37">C. inaequalis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="ABBE2FB3B17BA90FFBDCFFB508C990BA" authorityName="Borowiec & Świętojańska" authorityYear="2022" box="[1063,1194,1478,1504]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="overlaeti">
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<emphasis id="5ECA8822B17BA90FFBDCFFB508C990BA" box="[1063,1194,1478,1504]" italics="true" pageId="36" pageNumber="37">C. overlaeti</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="ABBE2FB3B17BA90FFB25FFB5095090BA" authorityName="Klug" authorityYear="1835" box="[1246,1331,1478,1504]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="tosta">
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<emphasis id="5ECA8822B17BA90FFB25FFB5095090BA" box="[1246,1331,1478,1504]" italics="true" pageId="36" pageNumber="37">C. tosta</emphasis>
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</taxonomicName>
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) but they differ in pronotum broadly rounded and base of elytra distinctly wider than pronotum. Individuals are usually dark coloured, black, reddish brown, or mixed reddish brown and black. Only
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<taxonomicName id="ABBE2FB3B17BA90FFC3DFC7D08389372" authorityName="Thomson" authorityYear="1858" box="[966,1115,1550,1576]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="inaequalis">
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<emphasis id="5ECA8822B17BA90FFC3DFC7D08389372" box="[966,1115,1550,1576]" italics="true" pageId="36" pageNumber="37">C. inaequalis</emphasis>
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</taxonomicName>
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occasionally has the dorsum rusty reddish while in
|
||
<taxonomicName id="ABBE2FB3B17BA90FFE69FC410E629316" authorityName="Spaeth" authorityYear="1933" box="[402,513,1586,1612]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="blanda">
|
||
<emphasis id="5ECA8822B17BA90FFE69FC410E629316" box="[402,513,1586,1612]" italics="true" pageId="36" pageNumber="37">C. blanda</emphasis>
|
||
</taxonomicName>
|
||
the dorsum is ochraceous to rusty yellow (
|
||
<figureCitation id="F48548B5B17BA90FFC18FC4108599316" box="[995,1082,1586,1612]" captionStart="FIGURES 63–66" captionStartId="187.[151,264,1634,1658]" captionTargetBox="[173,1420,198,1584]" captionTargetId="figure-19@187.[151,1435,181,1610]" captionTargetPageId="187" captionText="FIGURES 63–66. Cassida inaequalis. FIGURES 67–68. Cassida blanda. 63–64, 66–67. Dorsal; 65, 68. Lateral." figureDoi="http://doi.org/10.5281/zenodo.6966400" httpUri="https://zenodo.org/record/6966400/files/figure.png" pageId="36" pageNumber="37">figs. 63</figureCitation>
|
||
, 67–68). No members of the
|
||
<taxonomicName id="ABBE2FB3B17BA90FFA78FC410CAE932A" authorityName="Klug" authorityYear="1835" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="tosta">
|
||
<emphasis id="5ECA8822B17BA90FFA78FC410CAE932A" italics="true" pageId="36" pageNumber="37">C. tosta</emphasis>
|
||
</taxonomicName>
|
||
species–group has the postscutellar hump as high as in
|
||
<taxonomicName id="ABBE2FB3B17BA90FFCC4FC250FCD932A" authorityName="Spaeth" authorityYear="1933" box="[831,942,1622,1648]" class="Magnoliopsida" family="Lamiaceae" genus="Cassida" kingdom="Plantae" order="Lamiales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="blanda">
|
||
<emphasis id="5ECA8822B17BA90FFCC4FC250FCD932A" box="[831,942,1622,1648]" italics="true" pageId="36" pageNumber="37">C. blanda</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="24A407BBB17BA90FFF3CFC09080F93CE" box="[199,1132,1657,1684]" pageId="36" pageNumber="37" type="materials_examined">
|
||
<paragraph id="6C015430B17BA90FFF3CFC09080F93CE" blockId="36.[151,1437,325,1684]" box="[199,1132,1657,1684]" pageId="36" pageNumber="37">
|
||
<emphasis id="5ECA8822B17BA90FFF3CFC090DE093C9" bold="true" box="[199,387,1657,1684]" pageId="36" pageNumber="37">Type examined.</emphasis>
|
||
<materialsCitation id="DCD65E6DB17BA90FFE71FC09080F93CE" ID-GBIF-Occurrence="3881324818" box="[394,1132,1658,1684]" collectionCode="ZSM" collectorName="D. O. Afrika & Ertl" country="Tanzania" location="TANZANIA" pageId="36" pageNumber="37" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="B305EA92B17BA90FFE71FC090D9493CE" box="[394,503,1658,1684]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
|
||
: [
|
||
<collectingCountry id="14A914A0B17BA90FFDFCFC090EC393CE" box="[519,672,1658,1684]" name="Tanzania" pageId="36" pageNumber="37">TANZANIA</collectingCountry>
|
||
]:
|
||
<collectorName id="C14B31E6B17BA90FFD4AFC090F2393CE" box="[689,832,1658,1684]" pageId="36" pageNumber="37">D. O. Afrika</collectorName>
|
||
, Ndanda, coll.
|
||
<collectorName id="C14B31E6B17BA90FFC13FC09087193CE" box="[1000,1042,1658,1684]" pageId="36" pageNumber="37">Ertl</collectorName>
|
||
(
|
||
<collectionCode id="0AAFCCF5B17BA90FFBDAFC09080393CE" box="[1057,1120,1658,1684]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="36" pageNumber="37" type="Museum">ZSM</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |