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<document id="7E4DEA237454A40C4169F4FB87F1347D" ID-DOI="10.37828/em.2021.43.2" ID-ISSN="2336-9744" ID-Zenodo-Dep="13233553" ID-ZooBank="urn:lsid:zoobank.org:pub:72F0B1F1-0185-443A-8457-4874C439C83C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722914924524" checkinUser="felipe" docAuthor="Yakovlev, Roman V. &amp; Zolotuhin, Vadim V." docDate="2021" docId="B83687CEFF83FC20FF75F00D615BFC55" docLanguage="en" docName="EcolMontenegrina.43.16-29.pdf" docOrigin="Ecologica Montenegrina 43" docSource="http://dx.doi.org/10.37828/em.2021.43.2" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Stueningeria csovarii Yakovlev &amp; Zolotuhin, 2021, sp. nov." docType="treatment" docUuid="DE0C3CBF-2947-4B87-A7F8-BE88565B4049" docUuidSource="ZooBank" docVersion="3" lastPageNumber="23" masterDocId="440FFFB6FF86FC27FFE2F72E6404FFB3" masterDocTitle="Revision of the family Metarbelidae (Lepidoptera) of the Oriental Region. III. Genus Stueningeria Lehmann, 2019" masterLastPageNumber="29" masterPageNumber="16" pageNumber="21" updateTime="1722919289534" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="101C717A2F2D95EB11F82448C22B864C">Revision of the family Metarbelidae (Lepidoptera) of the Oriental Region. III. Genus Stueningeria Lehmann, 2019</mods:title>
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<mods:name id="3586FEC9980BEE2399C753DEAF749DAF" type="personal">
<mods:role id="160D8048FA93103496DFA5BF728FCA83">
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<mods:namePart id="C1528CC0DBD6787297AD162F8E85A87F">Yakovlev, Roman V.</mods:namePart>
<mods:affiliation id="6B6690D623A05B0E955E59CEC7D8CA83">Altai State University, Lenina pr. 61, Barnaul, 656049, Russia. E-mail: yakovlev _ asu @ mail. ru &amp; Tomsk State University, Laboratory of Biodiversity and Ecology, Lenina pr. 36, 634050 Tomsk, Russia. &amp; Paleo Data Lab., Institute of Archaeology and Ethnography SB RAS, Novosibirsk, Russia.</mods:affiliation>
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<mods:role id="BA58CC40343FE09599125AC653FFA8E0">
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</mods:role>
<mods:namePart id="1EBFE111D4D6F5801B66211396A6F6A9">Zolotuhin, Vadim V.</mods:namePart>
<mods:affiliation id="933D7EACA39712DD19DF53CD4E245E7E">Ulyanovsk State Pedagogical University, pl. 100 - letia Lenina 4, RUS- 432700, Ulyanovsk, Russia. E-mail: v. zolot @ mail. ru</mods:affiliation>
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<mods:title id="8AE4BD2816D2BDC698BE64007B0739AA">Ecologica Montenegrina</mods:title>
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<mods:part id="6F93CDE0AD32D95E1541E7DC927319CB">
<mods:date id="A82988922C4962D7B4E5AD01FE5452A9">2021</mods:date>
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<mods:number id="925B72D4F1F3A9BC436FD439C041C927">2021-06-16</mods:number>
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<mods:url id="2897A59FF7818DCB451DDDC42EF92CA8">http://dx.doi.org/10.37828/em.2021.43.2</mods:url>
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<mods:identifier id="4D60C96F6CE579141A653835C8CD31A3" type="DOI">10.37828/em.2021.43.2</mods:identifier>
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<treatment id="B83687CEFF83FC20FF75F00D615BFC55" ID-DOI="http://doi.org/10.5281/zenodo.13233781" ID-Zenodo-Dep="13233781" LSID="urn:lsid:zoobank.org:act:DE0C3CBF-2947-4B87-A7F8-BE88565B4049" httpUri="http://treatment.plazi.org/id/B83687CEFF83FC20FF75F00D615BFC55" lastPageId="7" lastPageNumber="23" pageId="5" pageNumber="21">
<subSubSection id="78856553FF83FC22FF75F00D6073F8E8" pageId="5" pageNumber="21" type="nomenclature">
<paragraph id="302036D8FF83FC22FF75F00D6604F88E" blockId="5.[151,1143,1827,1917]" box="[151,512,1827,1853]" pageId="5" pageNumber="21">
<heading id="6B6881B4FF83FC22FF75F00D6604F88E" box="[151,512,1827,1853]" fontSize="11" level="2" pageId="5" pageNumber="21" reason="2">
<emphasis id="02EBEACAFF83FC22FF75F00D6604F88E" bold="true" box="[151,512,1827,1853]" pageId="5" pageNumber="21">
<taxonomicName id="F79F4D5BFF83FC22FF75F00D659DF88E" authorityName="Yakovlev &amp; Zolotuhin" authorityYear="2021" box="[151,409,1827,1853]" class="Insecta" family="Metarbelidae" genus="Stueningeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="21" phylum="Arthropoda" rank="species" species="csovarii" status="sp. nov.">
<emphasis id="02EBEACAFF83FC22FF75F00D659DF88E" bold="true" box="[151,409,1827,1853]" italics="true" pageId="5" pageNumber="21">Stueningeria csovarii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19D857B1FF83FC22FE43F00D6604F88E" box="[417,512,1827,1853]" pageId="5" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="302036D8FF83FC22FF75F06D6073F8E8" blockId="5.[151,1143,1827,1917]" box="[151,1143,1859,1883]" pageId="5" pageNumber="21">
https://zoobank.org/
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urn:lsid:zoobank.org:act:
<uuid id="44390C0DFF83FC22FD97F06D6073F8E8" box="[629,1143,1859,1883]" pageId="5" pageNumber="21">DE0C3CBF-2947-4B87-A7F8-BE88565B4049</uuid>
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</paragraph>
</subSubSection>
<subSubSection id="78856553FF83FC22FF75F04C6587F8CE" box="[151,387,1890,1917]" pageId="5" pageNumber="21" type="description">
<paragraph id="302036D8FF83FC22FF75F04C6587F8CE" blockId="5.[151,1143,1827,1917]" box="[151,387,1890,1917]" pageId="5" pageNumber="21">
<figureCitation id="A8A42A5DFF83FC22FF75F04C6506F8CE" box="[151,258,1890,1917]" captionStart="Figures 1−15" captionStartId="4.[151,237,1832,1856]" captionTargetBox="[194,1394,201,1813]" captionTargetId="figure-15@4.[192,1395,189,1813]" captionTargetPageId="4" captionText="Figures 1−15. Adult specimens of Stueningeria: 1. S. nepalensis, male, Nepal, Annapura Himal (MWM); 2. S. nepalensis, male, Nepal, Annapura Himal (MWM); 3. S. nepalensis, male, India, Kumaon-Himalaya (MWM); 4. S. campbelli, female, holotype (NHMUK); 5. S. phaga, female, holotype (NHMUK); 6. S. phaga, male, India, Assam (NHMUK, individual number NHMUK: 012832501); 7. S. htetae, male, holotype (MWM); 8. S. csovarii, male, holotype (MWM); 9. S. csovarii, female, paratype (MWM); 10. S. loeffleri, male, holotype (MWM); 11. S. ihlei, male, holotype (ZISP); 12. S. murzini, male, holotype (MWM); 13. S. murzini, female, paratype (MWM); 14. S. pinratanai, male, holotype (MWM); 15. S. pinratanai, female, paratype (MWM)." figureDoi="http://doi.org/10.5281/zenodo.13233555" httpUri="https://zenodo.org/record/13233555/files/figure.png" pageId="5" pageNumber="21">Figs 8−9</figureCitation>
,
<figureCitation id="A8A42A5DFF83FC22FEECF04C652AF8CE" box="[270,302,1890,1917]" captionStart="Figures 16−19" captionStartId="5.[151,237,1666,1690]" captionTargetBox="[184,1362,218,1624]" captionTargetId="figure-22@5.[151,1390,189,1648]" captionTargetPageId="5" captionText="Figures 16−19. Male genitalia of Stueningeria: 16. S. nepalensis, India, Kumaon-Himalaya GenPr MWM: 32.909; 17. S. phaga, India, Assam (slide NHMUK: 010315532); 18. S. htetae, holotype (GenPr MWM: 28.019); 19. S. csovarii, holotype (GenPr MWM: 28.020)." figureDoi="http://doi.org/10.5281/zenodo.13233557" httpUri="https://zenodo.org/record/13233557/files/figure.png" pageId="5" pageNumber="21">19</figureCitation>
,
<figureCitation id="A8A42A5DFF83FC22FED8F04C655EF8CE" box="[314,346,1890,1917]" captionStart="Figures 24−26" captionStartId="8.[151,237,1058,1082]" captionTargetBox="[163,1434,189,1039]" captionTargetId="figure-15@8.[151,1436,189,1039]" captionTargetPageId="8" captionText="Figures 24−26. Female genitalia of Stueningeria: 24. S. campbelli, holotype (slide NHMUK: Coss-260♀); 25. S. csovarii, paratype (GenPr MWM: 32.907); 26. 23. S. pinratanai, paratype (GenPr MWM: 32.905)." figureDoi="http://doi.org/10.5281/zenodo.13233561" httpUri="https://zenodo.org/record/13233561/files/figure.png" pageId="5" pageNumber="21">25</figureCitation>
,
<figureCitation id="A8A42A5DFF83FC22FE84F04C6587F8CE" box="[358,387,1890,1917]" captionStart="Figures 27−32" captionStartId="10.[151,237,1997,2021]" captionTargetBox="[169,1411,774,1942]" captionTargetId="figure-193@10.[151,1437,762,1956]" captionTargetPageId="10" captionText="Figures 27−32. Forewings of males of Stueningeria: 27. S. nepalensis, India; 28. S. htetae, holotype; 29. S. csovarii, holotype; 30. S. ihlei, holotype; 31. S. murzini, holotype; 32. S. pinratanai, holotype." figureDoi="http://doi.org/10.5281/zenodo.13233563" httpUri="https://zenodo.org/record/13233563/files/figure.png" pageId="5" pageNumber="21">29</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="78856553FF83FC22FF75F088674BF7B7" pageId="5" pageNumber="21" type="materials_examined">
<paragraph id="302036D8FF83FC22FF75F088674BF7B7" blockId="5.[151,1436,1957,2052]" pageId="5" pageNumber="21">
<emphasis id="02EBEACAFF83FC22FF75F0886583F873" bold="true" box="[151,391,1958,1984]" pageId="5" pageNumber="21">Material examined</emphasis>
.
<materialsCitation id="80F73C85FF83FC22FE75F08B603EF851" collectingDate="1998-04-07" collectionCode="MWM" collectorName="Tibor Csovari &amp; Pal Steger" country="Thailand" elevation="1000" location="Male" pageId="5" pageNumber="21" specimenCount="1" stateProvince="Nan" typeStatus="holotype">
<typeStatus id="EF24887AFF83FC22FE75F08B660CF873" box="[407,520,1957,1984]" pageId="5" pageNumber="21" type="holotype">Holotype</typeStatus>
.
<location id="35406003FF83FC22FDF7F08B6651F873" LSID="urn:lsid:plazi:treatment:B83687CEFF83FC20FF75F00D615BFC55:35406003FF83FC22FDF7F08B6651F873" box="[533,597,1957,1984]" country="Thailand" name="Male" pageId="5" pageNumber="21" stateProvince="Nan">Male</location>
,
<collectingCountry id="48887648FF83FC22FD80F08B66CAF873" box="[610,718,1957,1984]" name="Thailand" pageId="5" pageNumber="21">Thailand</collectingCountry>
,
<collectingRegion id="F25BF83AFF83FC22FD39F08B678BF873" box="[731,911,1957,1984]" country="Thailand" name="Nan" pageId="5" pageNumber="21">Changwat Nan</collectingRegion>
,
<locationDeviation id="21922FFAFF83FC22FC7FF08B60A8F873" box="[925,1196,1957,1984]" location="Bo Luang" pageId="5" pageNumber="21">
<quantity id="F7679B3DFF83FC22FC7FF08B67EDF873" box="[925,1001,1957,1984]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="21" unit="km" value="15.0">15 km</quantity>
N of Bo Luang
</locationDeviation>
,
<quantity id="F7679B3DFF83FC22FB58F08B6113F873" box="[1210,1303,1957,1984]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="21" unit="m" value="1000.0">
<elevation id="BBB2D1EBFF83FC22FB58F08B6113F873" box="[1210,1303,1957,1984]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="21" unit="m" value="1000.0">1000 m</elevation>
</quantity>
,
<date id="44211018FF83FC22FAC7F08B6193F873" box="[1317,1431,1957,1984]" pageId="5" pageNumber="21" value="1998-04-07">
<collectingDate id="5465E9F0FF83FC22FAC7F08B6193F873" box="[1317,1431,1957,1984]" pageId="5" pageNumber="21" value="1998-04-07">7.iv.1998</collectingDate>
</date>
, leg.
<collectorName id="9D6A530EFF83FC22FF2EF0E96576F851" box="[204,370,1991,2018]" pageId="5" pageNumber="21">Tibor Csovari</collectorName>
&amp;
<collectorName id="9D6A530EFF83FC22FE7EF0E96612F851" box="[412,534,1991,2018]" pageId="5" pageNumber="21">Pal Steger</collectorName>
(
<collectionCode id="568EAE1DFF83FC22FDCAF0E96678F851" box="[552,636,1991,2018]" pageId="5" pageNumber="21">MWM</collectionCode>
, GenPr-Heterocera
<collectionCode id="568EAE1DFF83FC22FC88F0E667BEF851" box="[874,954,1992,2018]" pageId="5" pageNumber="21">MWM</collectionCode>
– 28.020)
</materialsCitation>
.
<materialsCitation id="80F73C85FF83FC22FBA5F0E6674FF7B7" collectingDate="1998-04-07" collectionCode="MWM" collectorName="Tibor Csovari &amp; Pal Steger" country="Thailand" elevation="1000" location="Male" pageId="5" pageNumber="21" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Nan" typeStatus="paratype">
<typeStatus id="EF24887AFF83FC22FBA5F0E660BAF851" box="[1095,1214,1992,2018]" pageId="5" pageNumber="21" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="2699FD51FF83FC22FB29F0E9611BF851" box="[1227,1311,1991,2018]" count="1" pageId="5" pageNumber="21" type="male">1 male</specimenCount>
,
<specimenCount id="2699FD51FF83FC22FACFF0E96193F851" box="[1325,1431,1991,2018]" count="1" pageId="5" pageNumber="21" type="female">1 female</specimenCount>
, same locality (
<collectionCode id="568EAE1DFF83FC22FEA0F0C76593F7B7" box="[322,407,2025,2052]" pageId="5" pageNumber="21">MWM</collectionCode>
, GenPr-Heterocera
<collectionCode id="568EAE1DFF83FC22FD9DF0C466CBF7B7" box="[639,719,2026,2052]" pageId="5" pageNumber="21">MWM</collectionCode>
– 32.907)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="78856553FF80FC20FF75F7CB679EFD04" lastPageId="7" lastPageNumber="23" pageId="6" pageNumber="22" type="description">
<paragraph id="302036D8FF80FC21FF75F7CB60DDFDBD" blockId="6.[151,1436,229,526]" pageId="6" pageNumber="22">
<emphasis id="02EBEACAFF80FC21FF75F7CB652CFF4C" bold="true" box="[151,296,229,255]" pageId="6" pageNumber="22">Description</emphasis>
. Length of fore wing
<quantity id="F7679B3DFF80FC21FDD4F7CB668BFEB3" box="[566,655,229,256]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="22" unit="mm" value="14.0">14 mm</quantity>
. Antenna short, bipectinate, setae in antenna medium third twice longer than antenna rod diameter, thorax and abdomen covered with long brown scales (except for medium third of abdomen which is covered with light-grey scales, abdomen apically with long modified scales. Fore wing very mottled, bright, with bright brown spots of various shape and size throughout all wing area, thin creamy lines between spots; oblique band submarginally expressed from R
<subScript id="AC1B349DFF80FC21FBF9F6586020FE34" attach="left" box="[1051,1060,374,391]" fontSize="7" pageId="6" pageNumber="22">2</subScript>
to CuA
<subScript id="AC1B349DFF80FC21FB64F658608BFE34" attach="left" box="[1158,1167,374,391]" fontSize="7" pageId="6" pageNumber="22">2</subScript>
; long narrow crescent spot of grey androconium scales cubitally (from base to tornal angle), two short thin portions of grey androconium scales between radial trunk veins; marginal area and fringe mottled (with alternating brown and creamy spots). Hind wing light-brown with poorly expressed blurred dark-brown pattern, anal edge creamy, without pattern, marginal area and fringe mottled (with alternating brown and creamy spots).
</paragraph>
<caption id="64E06650FF80FC21FF75F02C64F5F8E4" ID-DOI="http://doi.org/10.5281/zenodo.13233559" ID-Zenodo-Dep="13233559" httpUri="https://zenodo.org/record/13233559/files/figure.png" pageId="6" pageNumber="22" startId="6.[151,237,1794,1818]" targetBox="[188,1405,559,1776]" targetPageId="6" targetType="figure">
<paragraph id="302036D8FF80FC21FF75F02C64F5F8E4" blockId="6.[151,1437,1794,1879]" pageId="6" pageNumber="22">
<emphasis id="02EBEACAFF80FC21FF75F02C66A7F8A9" bold="true" box="[151,675,1794,1818]" pageId="6" pageNumber="22">
Figures 20−23. Male genitalia of
<taxonomicName id="F79F4D5BFF80FC21FDEEF02C669EF8A9" authorityName="Lehmann" authorityYear="2019" box="[524,666,1794,1818]" class="Insecta" family="Metarbelidae" genus="Stueningeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="02EBEACAFF80FC21FDEEF02C669EF8A9" bold="true" box="[524,666,1794,1818]" italics="true" pageId="6" pageNumber="22">Stueningeria</emphasis>
</taxonomicName>
:
</emphasis>
20.
<taxonomicName id="F79F4D5BFF80FC21FD30F02C6746F8A9" authorityName="Yakovlev &amp; Zolotuhin" authorityYear="2021" box="[722,834,1794,1818]" class="Insecta" family="Metarbelidae" genus="Stueningeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="22" phylum="Arthropoda" rank="species" species="loeffleri">
<emphasis id="02EBEACAFF80FC21FD30F02C6746F8A9" box="[722,834,1794,1818]" italics="true" pageId="6" pageNumber="22">S. loeffleri</emphasis>
</taxonomicName>
, holotype (GenPr MWM: 32.908); 21.
<taxonomicName id="F79F4D5BFF80FC21FB0AF02C6135F8A9" authorityName="Yakovlev &amp; Zolotuhin" authorityYear="2021" box="[1256,1329,1794,1818]" class="Insecta" family="Metarbelidae" genus="Stueningeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="22" phylum="Arthropoda" rank="species" species="ihlei">
<emphasis id="02EBEACAFF80FC21FB0AF02C6135F8A9" box="[1256,1329,1794,1818]" italics="true" pageId="6" pageNumber="22">S. ihlei</emphasis>
</taxonomicName>
, holotype (slyde RYB 2017/5); 22.
<taxonomicName id="F79F4D5BFF80FC21FE5AF00F6629F88A" authorityName="Yakovlev &amp; Zolotuhin" authorityYear="2021" box="[440,557,1825,1849]" class="Insecta" family="Metarbelidae" genus="Stueningeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="22" phylum="Arthropoda" rank="species" species="murzini">S. murzini</taxonomicName>
, holotype (GenPr MWM: 28.018); 23.
<taxonomicName id="F79F4D5BFF80FC21FC08F00F607EF88A" authorityName="Yakovlev &amp; Zolotuhin" authorityYear="2021" box="[1002,1146,1825,1849]" class="Insecta" family="Metarbelidae" genus="Stueningeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="22" phylum="Arthropoda" rank="species" species="pinratanai">
<emphasis id="02EBEACAFF80FC21FC08F00F607EF88A" box="[1002,1146,1825,1849]" italics="true" pageId="6" pageNumber="22">S. pinratanai</emphasis>
</taxonomicName>
, holotype (GenPr MWM: 32.906).
</paragraph>
</caption>
<paragraph id="302036D8FF80FC20FF75F0B1651AFEF0" blockId="6.[151,1436,1951,2046]" lastBlockId="7.[151,1436,195,323]" lastPageId="7" lastPageNumber="23" pageId="6" pageNumber="22">
Male genitalia (
<figureCitation id="A8A42A5DFF80FC21FEB6F0B165A8F809" box="[340,428,1951,1978]" captionStart="Figures 16−19" captionStartId="5.[151,237,1666,1690]" captionTargetBox="[184,1362,218,1624]" captionTargetId="figure-22@5.[151,1390,189,1648]" captionTargetPageId="5" captionText="Figures 16−19. Male genitalia of Stueningeria: 16. S. nepalensis, India, Kumaon-Himalaya GenPr MWM: 32.909; 17. S. phaga, India, Assam (slide NHMUK: 010315532); 18. S. htetae, holotype (GenPr MWM: 28.019); 19. S. csovarii, holotype (GenPr MWM: 28.020)." figureDoi="http://doi.org/10.5281/zenodo.13233557" httpUri="https://zenodo.org/record/13233557/files/figure.png" pageId="6" pageNumber="22">Fig. 19</figureCitation>
). Uncus with deep semicircular bifurcation, uncus halves slightly extended apically, relatively long; scaphium and subscaphium fused into wide short spindle-like tube; gnathos arms short, lamellar; gnathos lamellar with long thin lateral processes; valve short with semicircular costal edge, slightly curved saccular and outer edges; saccular edge (in basal half) strongly sclerotized, tubulate, distally sharply cut, short, with pyramidal twin-peaked harpe directed dorsally; juxta lamellar, robust; saccus almost reduced; phallus twice shorter than valve, poorly curved in medium third, relatively thin, with small spiky cornutus in distal third.
</paragraph>
<paragraph id="302036D8FF81FC20FF75F64260E4FD83" blockId="7.[151,1437,364,560]" pageId="7" pageNumber="23">
Female (
<figureCitation id="A8A42A5DFF81FC20FEE2F642654EFE34" box="[256,330,364,391]" captionStart="Figures 1−15" captionStartId="4.[151,237,1832,1856]" captionTargetBox="[194,1394,201,1813]" captionTargetId="figure-15@4.[192,1395,189,1813]" captionTargetPageId="4" captionText="Figures 1−15. Adult specimens of Stueningeria: 1. S. nepalensis, male, Nepal, Annapura Himal (MWM); 2. S. nepalensis, male, Nepal, Annapura Himal (MWM); 3. S. nepalensis, male, India, Kumaon-Himalaya (MWM); 4. S. campbelli, female, holotype (NHMUK); 5. S. phaga, female, holotype (NHMUK); 6. S. phaga, male, India, Assam (NHMUK, individual number NHMUK: 012832501); 7. S. htetae, male, holotype (MWM); 8. S. csovarii, male, holotype (MWM); 9. S. csovarii, female, paratype (MWM); 10. S. loeffleri, male, holotype (MWM); 11. S. ihlei, male, holotype (ZISP); 12. S. murzini, male, holotype (MWM); 13. S. murzini, female, paratype (MWM); 14. S. pinratanai, male, holotype (MWM); 15. S. pinratanai, female, paratype (MWM)." figureDoi="http://doi.org/10.5281/zenodo.13233555" httpUri="https://zenodo.org/record/13233555/files/figure.png" pageId="7" pageNumber="23">Fig. 9</figureCitation>
). Length of fore wing
<quantity id="F7679B3DFF81FC20FDBCF64266B2FE34" box="[606,694,364,391]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="23" unit="mm" value="12.0">12 mm</quantity>
. Antenna short, bipectinate, setae in medium third of antenna 1.5 times longer than antenna rod diameter, thorax and abdomen from above covered with light-grey scales, abdomen apically with long modified brown scales. Fore wing creamy with poorly expressed brown pattern of small spots along costal edge and submarginally, big brown spot in discal cell, slight sputtering of brown scales throughout all wing area. Hind wing light-brown with poorly expressed dim dark-brown pattern, anal angle creamy without pattern. Fringe on all wings mottled, light between veins, dark at veins.
</paragraph>
<paragraph id="302036D8FF81FC20FF75F576679EFD04" blockId="7.[151,1436,600,695]" pageId="7" pageNumber="23">
Female genitalia (
<figureCitation id="A8A42A5DFF81FC20FE91F57665CBFDC0" box="[371,463,600,627]" captionStart="Figures 24−26" captionStartId="8.[151,237,1058,1082]" captionTargetBox="[163,1434,189,1039]" captionTargetId="figure-15@8.[151,1436,189,1039]" captionTargetPageId="8" captionText="Figures 24−26. Female genitalia of Stueningeria: 24. S. campbelli, holotype (slide NHMUK: Coss-260♀); 25. S. csovarii, paratype (GenPr MWM: 32.907); 26. 23. S. pinratanai, paratype (GenPr MWM: 32.905)." figureDoi="http://doi.org/10.5281/zenodo.13233561" httpUri="https://zenodo.org/record/13233561/files/figure.png" pageId="7" pageNumber="23">Fig. 25</figureCitation>
). Ovipositor very short, papillae anales semicircular, anterior apophyses almost equal to posterior apophyses in length, short, antrum strongly sclerotized, elongated, funnel-like; ductus wide, smoothly passing into small bursa copulatrix without signa.
</paragraph>
</subSubSection>
<subSubSection id="78856553FF81FC20FF75F5F16721FCEC" pageId="7" pageNumber="23" type="diagnosis">
<paragraph id="302036D8FF81FC20FF75F5F16721FCEC" blockId="7.[151,1437,735,863]" pageId="7" pageNumber="23">
<emphasis id="02EBEACAFF81FC20FF75F5F16514FD4A" bold="true" box="[151,272,735,761]" pageId="7" pageNumber="23">Diagnosis</emphasis>
. Externally, the new species is similar to
<taxonomicName id="F79F4D5BFF81FC20FD1CF5CE674BFD49" authorityName="Yakovlev &amp; Zolotuhin" authorityYear="2021" box="[766,847,735,762]" class="Insecta" family="Metarbelidae" genus="Stueningeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="23" phylum="Arthropoda" rank="species" species="ihlei" status="sp. nov.">
<emphasis id="02EBEACAFF81FC20FD1CF5CE674BFD49" box="[766,847,735,762]" italics="true" pageId="7" pageNumber="23">S. ihlei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19D857B1FF81FC20FCB5F5CE67B7FD49" box="[855,947,736,762]" pageId="7" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="F79F4D5BFF81FC20FC0CF5CE608CFD4A" authorityName="Yakovlev &amp; Zolotuhin" authorityYear="2021" box="[1006,1160,736,762]" class="Insecta" family="Metarbelidae" genus="Stueningeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="23" phylum="Arthropoda" rank="species" species="pinratanai" status="sp. nov.">
<emphasis id="02EBEACAFF81FC20FC0CF5CE608CFD4A" box="[1006,1160,736,762]" italics="true" pageId="7" pageNumber="23">S. pinratanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19D857B1FF81FC20FB72F5CE60EBFD49" box="[1168,1263,736,762]" pageId="7" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
, from which it differs in very narrow androconium portions in the radial area of the fore wing, in the long distal halves of the uncus and in the twin-peaked harpe on the saccular edge of the valve. From
<taxonomicName id="F79F4D5BFF81FC20FBB3F40D60DEFC8E" authorityName="Yakovlev &amp; Zolotuhin" authorityYear="2021" box="[1105,1242,802,829]" class="Insecta" family="Metarbelidae" genus="Stueningeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="23" phylum="Arthropoda" rank="species" species="loeffeleri">
<emphasis id="02EBEACAFF81FC20FBB3F40D60DEFC8E" box="[1105,1242,802,829]" italics="true" pageId="7" pageNumber="23">S. loeffeleri</emphasis>
</taxonomicName>
the new species differs in the absence of small serrations on the phallus.
</paragraph>
</subSubSection>
<subSubSection id="78856553FF81FC20FF75F4A6666FFC10" box="[151,619,904,931]" pageId="7" pageNumber="23" type="distribution">
<paragraph id="302036D8FF81FC20FF75F4A6666FFC10" blockId="7.[151,619,904,931]" box="[151,619,904,931]" pageId="7" pageNumber="23">
<emphasis id="02EBEACAFF81FC20FF75F4A66535FC11" bold="true" box="[151,305,904,930]" pageId="7" pageNumber="23">Distribution</emphasis>
.
<collectingCountry id="48887648FF81FC20FEDDF4A665A3FC10" box="[319,423,904,931]" name="Thailand" pageId="7" pageNumber="23">Thailand</collectingCountry>
(
<collectingRegion id="F25BF83AFF81FC20FE55F4A6665BFC10" box="[439,607,904,931]" country="Thailand" name="Nan" pageId="7" pageNumber="23">Nan Province</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="78856553FF81FC20FF75F4E2615BFC55" box="[151,1375,971,998]" pageId="7" pageNumber="23" type="etymology">
<paragraph id="302036D8FF81FC20FF75F4E2615BFC55" blockId="7.[151,1375,971,998]" box="[151,1375,971,998]" pageId="7" pageNumber="23">
<emphasis id="02EBEACAFF81FC20FF75F4E2651BFC55" bold="true" box="[151,287,972,998]" pageId="7" pageNumber="23">Etymology</emphasis>
. The new species is named after the entomologist Tibor Csovari – the new species collector.
</paragraph>
</subSubSection>
</treatment>
</document>
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