176 lines
19 KiB
XML
176 lines
19 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6452741" ID-ISSN="2073-2333" ID-Zenodo-Dep="6452741" approvalRequired="12" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="8" approvalRequired_for_treatments="3" checkinTime="1649773237151" checkinUser="felipe" docAuthor="Korneyev, V. A." docDate="2015" docId="B83387CFE27D8168C9858A2BEEAE5C3E" docLanguage="en" docName="VestnikZoologii.49.6.497-518.pdf" docOrigin="Vestnik Zoologii 49 (6)" docSource="http://dx.doi.org/10.1515/vzoo-2015-0061" docStyle="DocumentStyle:9FE6554C034D0A4D8EA9CEAE255F4368.2:VestnikZoologii.2014-2019.journal_article" docStyleId="9FE6554C034D0A4D8EA9CEAE255F4368" docStyleName="VestnikZoologii.2014-2019.journal_article" docStyleVersion="2" docTitle="Geloemyia namibica V. Korneyev 2015, sp. n." docType="treatment" docVersion="2" lastPageId="13" lastPageNumber="510" masterDocId="440AFFB7E2768165C92B8E64EA645A74" masterDocTitle="Review Of The Genus Geloemyia (Diptera, Pyrgotidae), With Discussion Of Its Taxonomic Position" masterLastPageNumber="518" masterPageNumber="497" pageId="11" pageNumber="508" updateTime="1649773747346" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review Of The Genus Geloemyia (Diptera, Pyrgotidae), With Discussion Of Its Taxonomic Position</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Korneyev, V. A.</mods:namePart>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Vestnik Zoologii</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-12-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>49</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>6</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>497</mods:start>
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<mods:end>518</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.1515/vzoo-2015-0061</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1515/vzoo-2015-0061</mods:identifier>
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<mods:identifier type="ISSN">2073-2333</mods:identifier>
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<mods:identifier type="Zenodo-Dep">6452699</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6452741" ID-Zenodo-Dep="6452741" LSID="urn:lsid:plazi:treatment:B83387CFE27D8168C9858A2BEEAE5C3E" httpUri="http://treatment.plazi.org/id/B83387CFE27D8168C9858A2BEEAE5C3E" lastPageId="13" lastPageNumber="510" pageId="11" pageNumber="508">
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<subSubSection pageId="11" pageNumber="508" type="nomenclature">
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<paragraph blockId="11.[174,904,1103,1130]" box="[174,904,1103,1130]" pageId="11" pageNumber="508">
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<heading box="[174,904,1103,1130]" fontSize="36" level="2" pageId="11" pageNumber="508" reason="3">
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<taxonomicName authority="V. Korneyev" authorityName="V. Korneyev" authorityYear="2015" box="[174,579,1103,1130]" class="Insecta" family="Pyrgotidae" genus="Geloemyia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="508" phylum="Arthropoda" rank="species" species="namibica" status="sp. n.">
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<emphasis bold="true" box="[174,423,1103,1130]" italics="true" pageId="11" pageNumber="508">Geloemyia namibica</emphasis>
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V. Korneyev
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</taxonomicName>
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,
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<taxonomicNameLabel box="[590,652,1104,1130]" pageId="11" pageNumber="508" rank="species">sp. n.</taxonomicNameLabel>
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(fig. 1, 11, 17, 55–66)
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</heading>
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</paragraph>
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<paragraph blockId="11.[174,1232,1149,1225]" pageId="11" pageNumber="508">
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<materialsCitation collectingDate="1998-01-22" collectingDateMax="1998-01-23" collectingDateMin="1998-01-22" collectionCode="NICW" collectorName="Kirk-Spriggs & Marais" country="Namibia" county="Rundu Dist." latitude="-18.893888" location="River" municipality="Kotara" pageId="11" pageNumber="508" specimenCount="1" stateProvince="Kavango West" typeStatus="holotype">
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M a t e r i a l.
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<emphasis bold="true" box="[350,406,1150,1171]" pageId="11" pageNumber="508">Type.</emphasis>
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<typeStatus box="[412,502,1149,1171]" pageId="11" pageNumber="508">Holotype</typeStatus>
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ǫ:
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<emphasis bold="true" box="[535,628,1149,1171]" pageId="11" pageNumber="508">
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<collectingCountry box="[535,624,1149,1171]" name="Namibia" pageId="11" pageNumber="508">Namibia</collectingCountry>
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:
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</emphasis>
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<collectingCounty box="[634,754,1149,1171]" pageId="11" pageNumber="508">Rundu Dist.</collectingCounty>
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,
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<collectingMunicipality box="[763,829,1149,1171]" pageId="11" pageNumber="508">Kotara</collectingMunicipality>
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<collectingRegion box="[834,933,1149,1171]" country="Namibia" name="Kavango West" pageId="11" pageNumber="508">Okavango</collectingRegion>
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<location LSID="urn:lsid:plazi:treatment:B83387CFE27D8168C9858A2BEEAE5C3E:35456002E27D816ECA808A19E9855EE7" box="[939,993,1149,1171]" country="Namibia" county="Rundu Dist." latitude="-18.893888" municipality="Kotara" name="River" pageId="11" pageNumber="508" stateProvince="Kavango West">River</location>
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, 17̊48'56"
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<geoCoordinate box="[1099,1207,1149,1172]" degrees="18" direction="south" minutes="53" orientation="latitude" pageId="11" pageNumber="508" precision="15" seconds="38" value="-18.893888">S 18º53'38"</geoCoordinate>
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E,
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<collectingMethod box="[174,262,1176,1198]" pageId="11" pageNumber="508">light trap</collectingMethod>
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sample,
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<date box="[347,469,1176,1198]" pageId="11" pageNumber="508" value="1998-01-22" valueMax="1998-01-23" valueMin="1998-01-22">
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<collectingDate box="[347,469,1176,1198]" pageId="11" pageNumber="508" value="1998-01-22" valueMax="1998-01-23" valueMin="1998-01-22">22–23.i.1998</collectingDate>
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</date>
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(
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<collectorName box="[481,606,1176,1198]" pageId="11" pageNumber="508">Kirk-Spriggs</collectorName>
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&
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<collectorName box="[633,701,1176,1198]" pageId="11" pageNumber="508">Marais</collectorName>
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) (
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<collectionCode box="[720,791,1176,1198]" pageId="11" pageNumber="508">NICW</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation collectingDate="1993-10-01" collectingDateMax="1993-10-03" collectingDateMin="1993-10-01" collectionCode="NICW, SIZK, MNKB" collectorName="Koch" country="Namibia" location="Mahango Game Reserve" pageId="11" pageNumber="508" specimenCount="2" typeStatus="paratype">
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<typeStatus box="[806,899,1176,1198]" pageId="11" pageNumber="508">Paratypes</typeStatus>
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3 ǫ:
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<emphasis bold="true" box="[948,1035,1176,1198]" pageId="11" pageNumber="508">
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<collectingCountry box="[948,1035,1176,1198]" name="Namibia" pageId="11" pageNumber="508">Namibia</collectingCountry>
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</emphasis>
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: same label as in the
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<typeStatus box="[174,258,1203,1225]" pageId="11" pageNumber="508">holotype</typeStatus>
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(
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<collectionCode box="[270,339,1203,1225]" pageId="11" pageNumber="508">NICW</collectionCode>
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;
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<collectionCode box="[348,402,1203,1225]" country="Ukraine" httpUri="http://grbio.org/cool/c442-9tdv" name="Schmaulhausen Institute of Zoology" pageId="11" pageNumber="508">SIZK</collectionCode>
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); 1 Ơ:
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<location LSID="urn:lsid:plazi:treatment:B83387CFE27D8168C9858A2BEEAE5C3E:35456002E27D816EC8E68AD7E8DD5EBD" box="[461,697,1203,1225]" country="Namibia" name="Mahango Game Reserve" pageId="11" pageNumber="508">Mahango Game Reserve</location>
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,
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<date box="[706,810,1203,1225]" pageId="11" pageNumber="508" value="1993-10-01" valueMax="1993-10-03" valueMin="1993-10-01">
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<collectingDate box="[706,810,1203,1225]" pageId="11" pageNumber="508" value="1993-10-01" valueMax="1993-10-03" valueMin="1993-10-01">1–3.x.1993</collectingDate>
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</date>
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(
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<collectorName box="[822,877,1203,1225]" pageId="11" pageNumber="508">Koch</collectorName>
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) (
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<collectionCode box="[896,971,1203,1225]" pageId="11" pageNumber="508">MNKB</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="13" lastPageNumber="510" pageId="11" pageNumber="508" type="description">
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<paragraph blockId="11.[174,1233,1258,1828]" pageId="11" pageNumber="508">
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D i a g n o s i s. This species can be recognized from combination of the head triangular in profile, with the parafacial dorsally 1.5–2× as long as flagellomere 1, face strongly receding, and peristomal cavity very short (as long as or slightly shorter than parafacial), ocellar setae long, one pair of orbital setae; wing with mottled pattern covering whole disk, including anal cell and lobe; vein R2+3 with stump vein; vein R4+5 setulose dorsally; crossvein dm-cu forming almost right angle (80–85°) with vein CuA
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<subScript attach="left" box="[1003,1011,1437,1451]" fontSize="6" pageId="11" pageNumber="508">1</subScript>
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; mesonotum with 1 presutural and 3 postsutural dorsocentral setae; scutellum with 2 pairs of setae but without setulae; female mid femur without femoral organ; oviscape slightly longer than tergites 3–6 combined.
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</paragraph>
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<paragraph blockId="11.[174,1233,1258,1828]" lastBlockId="12.[166,1225,1247,1850]" lastPageId="12" lastPageNumber="509" pageId="11" pageNumber="508">D e s c r i p t i o n. Head (fig. 55–58) yellow to brownish yellow, triangular in profile, height: length: width ratio = 1: 1: 1.1; with face strongly receding, and peristomal cavity very short; frontal vitta yellow, matt, microtrichose, black setulose anterolaterally, medially bare, 1.3–1.4× as long as wide in posterior half; parafacial 1.5–2× as wide as flagellomere 1; eye 1.4× as high as wide, 1.4–1.5× as wide as parafacial and 2–2.5× as high as gena. Face concave, with faint medial suture, in ventral one-fourth with short carina. Epistome acute triangular, very long, slightly longer than antennal grooves. Gena with short and low, sometimes brownish, subocular sclerite; postgena anteriorly with 1–2 subequal setae and 7–7 shorter setulae. Occiput entirely yellow. First flagellomere entirely reddish yellow, 1.3× as long as wide, 1.3× as long as pedicel and 2.3× as long as scape. Arista reddish yellow in basal half, 2-segmented, very short pubescent. Scape as long as wide. Palp yellow, oval, 2.5× as long as wide, and as long as first flagellomere. Labellum small, almost as long as palp. Lateral vertical setae (vte) 0.2–0.3× as long as medial vertical setae (vti); vti 3× as long as pedicel, 1.5× as long as ocellar and 3× as long as postocellar and orbital setae and 3–4× as long as setulae (above lunule).</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.6452721" ID-Zenodo-Dep="6452721" httpUri="https://zenodo.org/record/6452721/files/figure.png" pageId="12" pageNumber="509" startId="12.[170,203,1146,1168]" targetBox="[211,1191,183,1124]" targetPageId="12">
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<paragraph blockId="12.[170,1228,1145,1194]" pageId="12" pageNumber="509">
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Fig. 54–61.
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<taxonomicName authority="Korneyev, 2015" authorityName="V. Korneyev" authorityYear="2015" box="[283,480,1146,1167]" class="Insecta" family="Pyrgotidae" genus="Geloemyia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="509" phylum="Arthropoda" rank="species" species="namibica" status="sp. n.">
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<emphasis box="[283,480,1146,1167]" italics="true" pageId="12" pageNumber="509">Geloemyia namibica</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[492,546,1146,1168]" pageId="12" pageNumber="509" rank="species">sp. n.</taxonomicNameLabel>
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, paratype ǫ (SIZK): 54 — habitus; 55–58 — head (55 — lateral, 56 — anterolateral, 57 — dorsal, 58 — anteroventral view); 59 — wing; 60 — mesonotum, dorsal view; 61 — labels.
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</paragraph>
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</caption>
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<paragraph blockId="12.[166,1225,1247,1850]" pageId="12" pageNumber="509">Thorax yellow to brownish yellow, without distinct dark pattern (fig. 60). Prosternum saddle-like, short setulose. Postpronotal lobe with 4–6 setulae and one long seta; proepisternum with 3–5 setae. Scutum as long as wide, subshining yellow with faint brown areas, sparsely black setulose, with white microtrichia not obscuring underlying cuticle; 2 notopleural, 1 presutural and 1 postsutural supra-alar, 1 intra-alar, 1 postalar, 1 short presutural and 2–3 postsutural dorsocentral, and 0–1 acrostichal setae (if present, slightly longer than setulae). Scutellum yellow, whitish microtrichose, with 2 pairs of setae: apical 1.5× as long as scutellum and basal as long as scutellum, but without setulae (fig. 1, 5). Anepisternum yellow, with reddish or brownish anterior part covered with 10–12 short setulae; posterior part with 1–2 long setae and 6–7 irregular long setulae; anepimeron with 1 long seta and 1–2 short setulae; katepisternum with 1–3 long setae and 1–2 thinner and slightly shorter setulae dorsally and 2–3 setae medioventrally, anterior of mid coxa. Katatergite, anatergite and mediotergite matt yellow.</paragraph>
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<paragraph blockId="13.[174,1233,175,1616]" pageId="13" pageNumber="510">
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Wing (fig. 1, 59) 2.5–2.6× as long as wide, yellowish or hyaline with mottled pattern of moderately small grey spots partly fused apically and from pterostigma to posterior margin through crossvein r-m, and dark spots in anterior cubital and anal cells, anal lobe and alula; veins brown or yellowish. Costa with conspicuous humeral and subcostal break, subcostal vein broken at apex and not joined to costa. Pterostigma half as long as costal cell. Vein R
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<subScript attach="left" box="[192,219,354,368]" fontSize="6" pageId="13" pageNumber="510">2+3</subScript>
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sinuate, with stump vein. Vein R
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<subScript attach="left" box="[603,629,354,368]" fontSize="6" pageId="13" pageNumber="510">4+5</subScript>
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dorsally setulose over almost entire length. Cell cup closed by slightly sinuate crossvein and very short lobe. Wing length 6.0–
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<quantity box="[1051,1146,367,394]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="13" pageNumber="510" unit="mm" value="6.4">6.4 mm</quantity>
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. Lower calypter narrow, without cilia; upper calypter with long and dense setulae forming black fringe. Halter creamy white.
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</paragraph>
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<paragraph blockId="13.[174,1233,175,1616]" pageId="13" pageNumber="510">Legs (fig. 54) entirely yellow, black setulose and setose. Coxae moderately long and sparsely setose and setulose, unmodified. Fore femur moderately narrow, with basiventral seta and row of 6–7 strong black posteroventral setae as long as femur width and 4–5 shorter and thinner postero-dorsal setae; long setulose on posterior surface. Fore tibia widened apically, with 3–4 short preapical setae on dorsal surface. Fore tarsus unmodified in both sexes, yellow, tarsomere 1 twice as long as tarsomere 2; tarsomere 5 orange, narrow, as wide as tarsomere 3; tarsomere 4 unmodified, as wide as and 0.7× as long as tarsomere 5 (fig. 11). Mid femur unmodified, without long setae, uniformly setulose on anterior surface setose without femoral organ. Hind femur narrow, short setulose, without 2 longer subapical dorsal setae.</paragraph>
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<paragraph blockId="13.[174,1233,175,1616]" pageId="13" pageNumber="510">Abdomen (fig. 54) moderately wide, as long as thorax, with syntergite 1+2 transverse, 0.6× as long as wide and 0.7× as long as tergites 3–6 combined. Sternites of female wide,with short setulae and long marginal and submarginal setae on posterior margins, synsternite 1+2 trapezoid, sternites 3–5 subrectangular, wider than long; sternite 6 wide trapezoid. Synsternite 1+2 of male triangular, sternites 3–4 very narrow, somewhat widened at anterior margin and with oval posterior portion bearing 6–8 setae and 5-6 short setulae; sternite 5 with narrow anterior half and equilaterally triangular posterior half (fig. 62).</paragraph>
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<paragraph blockId="13.[174,1233,175,1616]" pageId="13" pageNumber="510">
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Female genitalia. Oviscape entirely orange (fig. 54),
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<quantity box="[840,980,1007,1034]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="13" pageNumber="510" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
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long, uncompressed, cylindrical, twice as long as wide basally, sparsely black setulose, and as long as tergites 3–6 combined, with apex short and inconspicuously bent dorsally, laterally bearing long setulae 1.2–2× as long as oviscape width (fig. 17). Dorsal (dorso-medial) lobes short, latero-ventral lobes very short, medial lobe long, twice as long as width of apex. Eversible membrane not examined. Aculeus (fig. 65) basally widened, without preapical constriction, 7× as long as wide at middle. Ventral receptacle as in other Pyrgotinae. Spermathecae oval (fig. 66).
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</paragraph>
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<paragraph blockId="13.[174,1233,175,1616]" pageId="13" pageNumber="510">
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Male genitalia as on figs 63, 64; epandrium and surstyli covered by short setulae inserted into big sockets; dorsal portion of epandrium bearing no long setae, only 4–5 short setae at level of cerci and fusion of epandrium with surstyli; surstyli directed posteriorly, as long as epandrium, robust; inner surstylus without prensisetae; phallus bare, with glans as on fig. 64, very similar to that of
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<taxonomicName authorityName="Korneyev" authorityYear="2015" baseAuthorityName="Enderlein" baseAuthorityYear="1942" box="[555,706,1359,1386]" class="Insecta" family="Pyrgotidae" genus="Geloemyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="510" phylum="Arthropoda" rank="species" species="trifasciata">
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<emphasis box="[555,706,1359,1386]" italics="true" pageId="13" pageNumber="510">G. trifasciata</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph blockId="13.[174,1233,175,1616]" pageId="13" pageNumber="510">
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Remarks. The main difference of
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<taxonomicName authority="Korneyev, 2015" authorityName="V. Korneyev" authorityYear="2015" box="[644,788,1391,1418]" class="Insecta" family="Pyrgotidae" genus="Geloemyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="510" phylum="Arthropoda" rank="species" species="namibica" status="sp. n.">
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<emphasis box="[644,788,1391,1418]" italics="true" pageId="13" pageNumber="510">G. namibica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[796,860,1392,1418]" pageId="13" pageNumber="510" rank="species">sp. n.</taxonomicNameLabel>
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from most other species of the genus (at least from those known from both sexes) is that it shows no sexual dimorphism in the wing pattern or structure of fore and mid legs; it seems to be related to the Madagascan
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<taxonomicName authorityName="Korneyev" authorityYear="2015" baseAuthorityName="Enderlein" baseAuthorityYear="1942" box="[174,326,1487,1514]" class="Insecta" family="Pyrgotidae" genus="Geloemyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="510" phylum="Arthropoda" rank="species" species="trifasciata">
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<emphasis box="[174,326,1487,1514]" italics="true" pageId="13" pageNumber="510">G. trifasciata</emphasis>
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</taxonomicName>
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as they share the high epistome, developed presutural dorsocentral seta and conspicuously narrowed abdominal sternites of males (possible synapomorphies, not occurring in most other Pyrgotinae except
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<taxonomicName authorityName="Hendel" authorityYear="1934" box="[657,804,1552,1577]" class="Insecta" family="Pyrgotidae" genus="Pyrgotomyia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="510" phylum="Arthropoda" rank="genus">
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<emphasis box="[657,804,1552,1577]" italics="true" pageId="13" pageNumber="510">Pyrgotomyia</emphasis>
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</taxonomicName>
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), in combination with well developed spurious vein on R
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<subScript attach="left" box="[395,421,1602,1616]" fontSize="6" pageId="13" pageNumber="510">2+3</subScript>
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and unmodified fore tarsus of female (possible symplesiomorphies).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |