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<document id="C26388ABB49CCCBDBAD7F9C62EFF5AB3" ID-CLB-Dataset="26735" ID-DOI="10.1093/zoolinnean/zly035" ID-GBIF-Dataset="9d667866-47c7-4e0c-9fa9-22a0e202f36c" ID-Zenodo-Dep="3089237" ID-ZooBank="urn:lsid:zoobank.org:pub:FB357841" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1558456175796" checkinUser="plazi" docAuthor="Duran, Daniel P., Herrmann, David P., Roman, Stephen J., Gwiazdowski, Rodger A., Drummond, Jennifer A., Hood, Glen R. &amp; Egan, Scott P." docDate="2019" docId="B83387B9FF9CF125FF287852FD5BE96D" docLanguage="en" docName="zjls.186.250-285.pdf" docOrigin="Zoological Journal of the Linnean Society 186" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.1:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="1" docTitle="Dromochorus chaparralensis Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan 2019" docType="treatment" docVersion="7" lastPageNumber="275" masterDocId="440AFFC1FF84F13CFF867C56FFD8EF1A" masterDocTitle="Cryptic diversity in the North American Dromochorus tiger beetles (Coleoptera: Carabidae: Cicindelinae): a congruence-based method for species discovery" masterLastPageNumber="285" masterPageNumber="250" pageNumber="274" updateTime="1698731441728" updateUser="ExternalLinkService">
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<mods:title id="39ADD1D13580EC7449AEB1C860CCB04C">Cryptic diversity in the North American Dromochorus tiger beetles (Coleoptera: Carabidae: Cicindelinae): a congruence-based method for species discovery</mods:title>
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<treatment id="B83387B9FF9CF125FF287852FD5BE96D" ID-DOI="http://doi.org/10.5281/zenodo.5942806" ID-GBIF-Taxon="157232842" ID-Zenodo-Dep="5942806" LSID="urn:lsid:plazi:treatment:B83387B9FF9CF125FF287852FD5BE96D" httpUri="http://treatment.plazi.org/id/B83387B9FF9CF125FF287852FD5BE96D" lastPageId="25" lastPageNumber="275" pageId="24" pageNumber="274">
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<paragraph id="302536AFFF9CF124FF287852FDA7EB20" blockId="24.[174,767,1028,1121]" pageId="24" pageNumber="274">
<taxonomicName id="F79A4D2CFF9CF124FF287852FDE7EB00" ID-CoL="8LVBW" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[174,575,1028,1051]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="281" phylum="Arthropoda" rank="species" species="chaparralensis">
<emphasis id="02EEEABDFF9CF124FF287852FDE7EB00" bold="true" box="[174,575,1028,1051]" italics="true" pageId="24" pageNumber="274">DROMOCHORUS CHAPARRALENSIS</emphasis>
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DURAN, ROMAN, HERRMANN &amp; EGAN
<emphasis id="02EEEABDFF9CF124FD9B7873FDA7EB20" bold="true" box="[541,639,1058,1082]" pageId="24" pageNumber="274">SP. NOV.</emphasis>
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<subSubSection id="78806524FF9CF124FE25781FFDD2EB7B" box="[419,522,1097,1121]" pageId="24" pageNumber="274" type="description">
<paragraph id="302536AFFF9CF124FE25781FFDD2EB7B" blockId="24.[174,767,1028,1121]" box="[419,522,1097,1121]" pageId="24" pageNumber="274">
(
<figureCitation id="A8A12A2AFF9CF124FE2D781FFDDAEB7A" box="[427,514,1097,1121]" captionStart="Figure 9" captionStartId="17.[145,223,1847,1869]" captionTargetBox="[178,1403,196,1806]" captionTargetId="figure@17.[164,1406,196,1806]" captionTargetPageId="17" captionText="Figure 9. Dorsal habitus for species of Dromochorus. A, D.pruininus; B, D.belfragei; C, D. knisleyi sp. nov.; D, D. minimus sp. nov.; E, D. chaparralensis sp. nov.; F, D. welderensis sp. nov.; G, D. velutinigrens; H, D. pilatei." httpUri="https://zenodo.org/record/3089287/files/figure.png" pageId="24" pageNumber="274">FIG. 9E</figureCitation>
)
</paragraph>
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<subSubSection id="78806524FF9CF124FF25782CFEE4EBC5" pageId="24" pageNumber="274" type="vernacular_names">
<paragraph id="302536AFFF9CF124FF25782CFE83EB88" blockId="24.[163,423,1146,1200]" box="[163,347,1146,1170]" pageId="24" pageNumber="274">
<heading id="6B6D81C3FF9CF124FF25782CFE83EB88" box="[163,347,1146,1170]" fontSize="10" level="4" pageId="24" pageNumber="274" reason="1">
<emphasis id="02EEEABDFF9CF124FF25782CFE83EB88" box="[163,347,1146,1170]" italics="true" pageId="24" pageNumber="274">Common name</emphasis>
</heading>
</paragraph>
<paragraph id="302536AFFF9CF124FF2578CCFE7FEBAA" blockId="24.[163,423,1146,1200]" box="[163,423,1178,1200]" pageId="24" pageNumber="274">Chaparral tiger beetle.</paragraph>
<paragraph id="302536AFFF9CF124FF25789EFEE4EBC5" blockId="24.[163,780,1223,1492]" box="[163,316,1223,1247]" pageId="24" pageNumber="274">
<heading id="6B6D81C3FF9CF124FF25789EFEE4EBC5" box="[163,316,1223,1247]" fontSize="10" level="4" pageId="24" pageNumber="274" reason="1">
<emphasis id="02EEEABDFF9CF124FF25789EFEE4EBC5" box="[163,316,1223,1247]" italics="true" pageId="24" pageNumber="274">Type locality</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="78806524FF9CF124FF2578BEFDA1EAC9" pageId="24" pageNumber="274" type="materials_examined">
<paragraph id="302536AFFF9CF124FF2578BEFDA1EAC9" blockId="24.[163,780,1223,1492]" pageId="24" pageNumber="274">
<materialsCitation id="80F23CF2FF9CF124FF2578BEFDE1EA21" ID-GBIF-Occurrence="2243053344" collectingDate="1932-05-27" collectionCode="NMNH" collectorName="E. G. Lindsley Collector &amp; M. A. Cazier / Collection" location="Carrizo Springs, TX" pageId="24" pageNumber="274" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<location id="35456074FF9CF124FF2578BEFE55EBE7" LSID="urn:lsid:plazi:treatment:B83387B9FF9CF125FF287852FD5BE96D:35456074FF9CF124FF2578BEFE55EBE7" box="[163,397,1256,1278]" name="Carrizo Springs, TX" pageId="24" pageNumber="274">Carrizo Springs, TX</location>
.
<typeStatus id="EF21880DFF9CF124FE1F78BEFDDAEBE4" box="[409,514,1256,1278]" pageId="24" pageNumber="274" type="holotype">Holotype</typeStatus>
(deposited in
<collectionCode id="568BAE6AFF9CF124FD2178BEFCD8EBE7" box="[679,768,1256,1277]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="24" pageNumber="274">NMNH</collectionCode>
):
<specimenCount id="269CFD26FF9CF124FF257951FF16EA07" box="[163,206,1287,1309]" pageId="24" pageNumber="274" type="female">1 ♀</specimenCount>
,
<location id="35456074FF9CF124FF677950FE1AEA06" LSID="urn:lsid:plazi:treatment:B83387B9FF9CF125FF287852FD5BE96D:35456074FF9CF124FF677950FE1AEA06" box="[225,450,1286,1308]" name="Carrizo Spgs / Tex." pageId="24" pageNumber="274">Carrizo Spgs/Tex.</location>
<collectingDate id="5460E987FF9CF124FE4D7951FDE8EA06" box="[459,560,1287,1308]" pageId="24" pageNumber="274" value="1932-05-27">
<date id="4424106FFF9CF124FE4D7951FDE8EA06" box="[459,560,1287,1308]" pageId="24" pageNumber="274" value="1932-05-27">V-2732</date>
</collectingDate>
//
<collectorName id="9D6F5379FF9CF124FDD37951FED5EA21" pageId="24" pageNumber="274">E. G. Lindsley Collector</collectorName>
//
<collectorName id="9D6F5379FF9CF124FEAD7973FDE1EA21" box="[299,569,1317,1339]" pageId="24" pageNumber="274">M.A. Cazier/Collection</collectorName>
</materialsCitation>
. Paratypes: 3 ♂♂, 5 ♀♀, Carrizo Spgs./Tex VI-1232 // E.G. Lindsley Collector // A. Nicolay collection 1950. 1 ♀, Mexico, Tamaulipas/Nuevo Laredo/20-VI-2010/leg J. Stamatov. 1 ♀, USA: Texas: LaSalle Co./Chaparral W.M.A./19-VI- 2013/E. San Gregario.
<figureCitation id="A8A12A2AFF9CF124FE2579E8FE2CEAC9" box="[419,500,1470,1492]" captionStart="Figure 9" captionStartId="17.[145,223,1847,1869]" captionTargetBox="[178,1403,196,1806]" captionTargetId="figure@17.[164,1406,196,1806]" captionTargetPageId="17" captionText="Figure 9. Dorsal habitus for species of Dromochorus. A, D.pruininus; B, D.belfragei; C, D. knisleyi sp. nov.; D, D. minimus sp. nov.; E, D. chaparralensis sp. nov.; F, D. welderensis sp. nov.; G, D. velutinigrens; H, D. pilatei." httpUri="https://zenodo.org/record/3089287/files/figure.png" pageId="24" pageNumber="274">Fig. 9E</figureCitation>
, Map
<figureCitation id="A8A12A2AFF9CF124FDBD79E8FDA1EAC9" box="[571,633,1470,1492]" captionStart="Figure 3" captionStartId="8.[163,241,1807,1829]" captionTargetBox="[163,1443,581,1767]" captionTargetId="figure@8.[163,1443,581,1767]" captionTargetPageId="8" captionText="Figure 3. Distributionof sampled Dromochorus populations.Colours correspondto those used in Figure 1. Green = D.pilatei, pink = D. welderensis sp. nov., sky blue = D. velutinigrens, red = D. chaparralensis sp. nov., blue = D. pruininus, lime green = D. knisleyi sp. nov., orange = D. minimus sp. nov., black = D. belfragei." httpUri="https://zenodo.org/record/3089261/files/figure.png" pageId="24" pageNumber="274">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="78806524FF9CF124FF2579BDFDD3E9C0" pageId="24" pageNumber="274" type="distribution">
<paragraph id="302536AFFF9CF124FF2579BDFEE2E919" blockId="24.[163,780,1515,1754]" box="[163,314,1515,1539]" pageId="24" pageNumber="274">
<heading id="6B6D81C3FF9CF124FF2579BDFEE2E919" box="[163,314,1515,1539]" fontSize="10" level="4" pageId="24" pageNumber="274" reason="1">
<emphasis id="02EEEABDFF9CF124FF2579BDFEE2E919" box="[163,314,1515,1539]" italics="true" pageId="24" pageNumber="274">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="302536AFFF9CF124FF257A5AFDD3E9C0" blockId="24.[163,780,1515,1754]" pageId="24" pageNumber="274">
Inland South Texas (currently known from Dimmit, LaSalle, and Webb Counties) and Tamaulipas, Mexico. This species has not been well sampled and is likely present in adjacent areas within the Gulf Coastal Plain, especially in northern Mexico. It can occur with
<taxonomicName id="F79A4D2CFF9CF124FF257AF0FE81E9A0" authorityName="Johnson" authorityYear="1992" box="[163,345,1701,1723]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="274" phylum="Arthropoda" rank="species" species="velutinigrens">
<emphasis id="02EEEABDFF9CF124FF257AF0FE81E9A0" box="[163,345,1701,1723]" italics="true" pageId="24" pageNumber="274">D. velutinigrens</emphasis>
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in places where sandy soils mix with the dominant heavier red clays.
</paragraph>
</subSubSection>
<subSubSection id="78806524FF9CF124FF257AAFFB2CED70" pageId="24" pageNumber="274" type="diagnosis">
<paragraph id="302536AFFF9CF124FF257AAFFEC4E80A" blockId="24.[163,778,1785,1870]" box="[163,284,1785,1808]" pageId="24" pageNumber="274">
<heading id="6B6D81C3FF9CF124FF257AAFFEC4E80A" box="[163,284,1785,1808]" fontSize="10" level="4" pageId="24" pageNumber="274" reason="1">
<emphasis id="02EEEABDFF9CF124FF257AAFFEC4E80A" box="[163,284,1785,1808]" italics="true" pageId="24" pageNumber="274">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="302536AFFF9CF124FF257B4FFB17EE03" blockId="24.[163,778,1785,1870]" lastBlockId="24.[827,1443,198,619]" pageId="24" pageNumber="274">
<taxonomicName id="F79A4D2CFF9CF124FF257B4FFE29E834" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[163,497,1817,1838]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="274" phylum="Arthropoda" rank="species" species="chaparralensis">
<emphasis id="02EEEABDFF9CF124FF257B4FFE29E834" box="[163,497,1817,1838]" italics="true" pageId="24" pageNumber="274">Dromochorus chaparralensis</emphasis>
</taxonomicName>
is a robust, dark beetle, and most specimens are dull black with little colour on dorsal or ventral surfaces. Some specimens may have bluish reflections along the margins. It is restricted to mesquite chaparral in South Texas.
</paragraph>
<paragraph id="302536AFFF9CF124FCD57D74FBB6EE4D" blockId="24.[827,1443,198,619]" pageId="24" pageNumber="274">
This species is most likely to be confused with
<taxonomicName id="F79A4D2CFF9CF124FCBD7D14FC66EE4D" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[827,958,322,343]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="274" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="02EEEABDFF9CF124FCBD7D14FC66EE4D" box="[827,958,322,343]" italics="true" pageId="24" pageNumber="274">D. minimus</emphasis>
</taxonomicName>
or
<taxonomicName id="F79A4D2CFF9CF124FC627D17FBBFEE4C" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[996,1127,321,342]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="274" phylum="Arthropoda" rank="species" species="welderensis">
<emphasis id="02EEEABDFF9CF124FC627D17FBBFEE4C" box="[996,1127,321,342]" italics="true" pageId="24" pageNumber="274">welderensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="302536AFFF9CF124FCD57D36FB70EECB" blockId="24.[827,1443,198,619]" pageId="24" pageNumber="274">
<taxonomicName id="F79A4D2CFF9CF124FCD57D36FBB1EE6F" box="[851,1129,352,373]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="274" phylum="Arthropoda" rank="species" species="mimimus">
<emphasis id="02EEEABDFF9CF124FCD57D36FBB1EE6F" box="[851,1129,352,373]" italics="true" pageId="24" pageNumber="274">Dromochorus mimimus</emphasis>
</taxonomicName>
is usually smaller (
<figureCitation id="A8A12A2AFF9CF124FADF7D36FA43EE6C" box="[1369,1435,352,374]" captionStart="Figure 6" captionStartId="12.[163,242,1698,1720]" captionTargetBox="[291,1314,499,1657]" captionTargetId="figure@12.[291,1314,499,1657]" captionTargetPageId="12" captionText="Figure 6. Boxplots, by species, from morphometric data used in principal component analyses (PCA) for each of 411 individuals. PCA indicated three measurements: PC1, overall body length; PC2, shape of the pronotum; and PC3, elytral length to body length ratio, to explain 9295% of the total variation in the dataset. Shown are targeted comparisons among two main phylogenetic groups: 1, the D. minimusD. velutinigrensD. welderensischaparralensis group; and 2, the D. knisleyi D. belfrageiD. pruininus clade. Species labelled with the same letters did not differ significantly from each other, species labelled with ab did not differ between other species which were significantly different. Percentages are relative contributions per analyses, and group; outliers are shown as black dots." httpUri="https://zenodo.org/record/3089273/files/figure.png" pageId="24" pageNumber="274">Fig. 6</figureCitation>
;
<tableCitation id="7D180314FF9CF124FCBD7D28FC56EE89" box="[827,910,382,404]" captionStart="Table 2" captionStartId="11.[145,209,1268,1289]" captionText="Table 2. Mean ± SE for morphological measurements of body size and dimensions" pageId="24" pageNumber="274">Table 2</tableCitation>
), and possesses a prominent frosted ashy grey, beige, or blue sheen on the dorsal surface; the pronotum has sparse thin erect setae.
</paragraph>
<paragraph id="302536AFFF9CF124FCD57D8CFB2CED70" blockId="24.[827,1443,198,619]" pageId="24" pageNumber="274">
<taxonomicName id="F79A4D2CFF9CF124FCD57D8CFBA0EEF5" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[851,1144,474,495]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="274" phylum="Arthropoda" rank="species" species="welderensis">
<emphasis id="02EEEABDFF9CF124FCD57D8CFBA0EEF5" box="[851,1144,474,495]" italics="true" pageId="24" pageNumber="274">Dromochorus welderensis</emphasis>
</taxonomicName>
is black with a faint dark blue sheen dorsally, in many individuals. The habitat of
<taxonomicName id="F79A4D2CFF9CF124FCD17E4EFC25ED37" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[855,1021,536,557]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="274" phylum="Arthropoda" rank="species" species="welderensis">
<emphasis id="02EEEABDFF9CF124FCD17E4EFC25ED37" box="[855,1021,536,557]" italics="true" pageId="24" pageNumber="274">D. welderensis</emphasis>
</taxonomicName>
is Gulf Coast Prairie grasslands, as opposed to the more inland and forested mesquitechaparral habitat of
<taxonomicName id="F79A4D2CFF9CF124FBA07E00FB35ED70" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[1062,1261,597,619]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="274" phylum="Arthropoda" rank="species" species="chaparralensis">
<emphasis id="02EEEABDFF9CF124FBA07E00FB35ED70" box="[1062,1261,597,619]" italics="true" pageId="24" pageNumber="274">D. chaparralensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="78806524FF9CF125FCBD7EDDFF20EA05" lastPageId="25" lastPageNumber="275" pageId="24" pageNumber="274" type="description">
<paragraph id="302536AFFF9CF124FCBD7EDDFC1FEDB8" blockId="24.[827,1443,651,1777]" box="[827,967,651,674]" pageId="24" pageNumber="274">
<heading id="6B6D81C3FF9CF124FCBD7EDDFC1FEDB8" box="[827,967,651,674]" fontSize="10" level="4" pageId="24" pageNumber="274" reason="1">
<emphasis id="02EEEABDFF9CF124FCBD7EDDFC1FEDB8" box="[827,967,651,674]" italics="true" pageId="24" pageNumber="274">Description</emphasis>
</heading>
</paragraph>
<paragraph id="302536AFFF9CF124FCBD7EFCFA90E9EB" blockId="24.[827,1443,651,1777]" pageId="24" pageNumber="274">
Medium- to large-sized
<taxonomicName id="F79A4D2CFF9CF124FBD97EFCFAD8EDA5" box="[1119,1280,682,703]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="274" phylum="Arthropoda" rank="genus">
<emphasis id="02EEEABDFF9CF124FBD97EFCFAD8EDA5" box="[1119,1280,682,703]" italics="true" pageId="24" pageNumber="274">Dromochorus</emphasis>
</taxonomicName>
. Body length 12.514.5 mm, mean ♀ 13.7 mm, mean ♂ 13.0 mm. Head slightly wider than pronotum. Head black with metallic blue to green reflections mostly limited to the lateral ridge of the supraorbital region. Fine to marked rugosity often present on the frons and vertex. All head portions glabrous except for two supraorbital setae next to each eye. Frons concave in median area, especially in male, bulging towards slightly convex near anterior margin, clearly delimited from clypeus, gradually blending into vertex. Genae black often with weak metallic green to violet reflections, with shallow, longitudinal striae, gradually ending at border of vertex. Clypeus shining black, apparently lacking coloured reflections. Male labrum tridentate with 68 setae, central area pale ochre-testaceous, with a thin dark-brown to black border posteriorly and sometimes anteriorly, dark-brown to black laterally; in some populations, the pale central area of the labrum may exist as a small spot, up to one-quarter of the total labrum surface; female labrum tridentate with 68 setae, entirely dark-brown to black with polished metallic cupreous to green reflections.
<emphasis id="02EEEABDFF9CF124FBA5791BFAF2EA85" bold="true" pageId="24" pageNumber="274">All segments of maxillary and labial palpi consistently dark brown; apical segment is not darker than other segments</emphasis>
. Antennae normal length, reaching back to humerus and basal third of elytron, slightly longer in male than female; scape dark testaceous to black with metallic reflections of violet, cupreous and green, with 23 apical setae; pedicel dark testaceous with metallic reflections of violet, cupreous and green, lacking any setae; flagellum dark testaceous, antennomeres 34 with metallic violet and green reflections, densely clothed in short white setae, antennomeres 511 dull-textured without metallic reflections and possessing erect setae in apical rings only, covered with fine pubescence throughout.
</paragraph>
<paragraph id="302536AFFF9CF124FCBD7B41FA7AE857" blockId="24.[827,1442,1815,1869]" pageId="24" pageNumber="274">
<emphasis id="02EEEABDFF9CF124FCBD7B41FC4AE836" box="[827,914,1815,1836]" italics="true" pageId="24" pageNumber="274">Thorax</emphasis>
: Pronotum 2.63.4 mm in length, mean ♀ 3.1 mm, mean ♂ 3.1 mm; width 2.93.5 mm, mean ♀
</paragraph>
<paragraph id="302536AFFF9DF125FF177C91FF35EDFF" blockId="25.[145,762,198,741]" pageId="25" pageNumber="275">
3.4 mm, mean ♂ 3.1 mm. Pronotum dull black, slightly wider than long, widest near anterior margin, width to length ratio 1.0 to 1.1, setae sparse, mostly present along lateral third of dorsal surface; disc finely rugose, with thin but distinct median line, with well-defined shallow sulci present anteriorly and posteriorly; notopleural sutures clearly defined, not visible from dorsal view; proepisternum black, lacking prominent metallic coloured reflections, glabrous. Elytra elongate, dorsal surface convex, 7.28.6 mm length, mean ♀ 8.2 mm, mean ♂ 7.7 mm, shape similar in both sexes, but slightly wider in female, especially toward apical third; sutural spine absent, microserrations not present on elytral apices;
<emphasis id="02EEEABDFF9DF125FE147E03FF1FEDDD" bold="true" pageId="25" pageNumber="275">elytral texture dull, with no pitting present, elytral coloration black, may have weak blue reflections along lateral margins</emphasis>
; elytral maculations absent;
<emphasis id="02EEEABDFF9DF125FD957EE7FF3FEDFF" bold="true" pageId="25" pageNumber="275">subsutural foveae absent</emphasis>
.
</paragraph>
<paragraph id="302536AFFF9DF125FF177F5CFE9FEB6A" blockId="25.[145,762,777,1136]" pageId="25" pageNumber="275">
<emphasis id="02EEEABDFF9DF125FF177F5CFF1DEC05" box="[145,197,778,799]" italics="true" pageId="25" pageNumber="275">Legs</emphasis>
: Pro-, meso- and metacoxae black, without or with minimal metallic reflections, numerous setae on pro- and mesocoxae, sparse on metacoxae; pro- and mesotrochanters with a single erect seta, metatrochanter glabrous, trochanters dark brown-testaceous; femora black with metallic violet reflections, densely clothed in decumbent white setae; tibiae brown, clothed with setae of two types: sparser brown-testaceous long setae and dense short decumbent white setae; two tibial spines present; tarsi brown-testaceous, first three dilated protarsomeres in male with dense greyishwhite setal pad.
</paragraph>
<paragraph id="302536AFFF9DF125FF1778D8FF20EA05" blockId="25.[145,761,1166,1311]" pageId="25" pageNumber="275">
<emphasis id="02EEEABDFF9DF125FF1778D8FEDAEBB9" box="[145,258,1166,1187]" italics="true" pageId="25" pageNumber="275">Abdomen</emphasis>
: Venter mostly black with metallic olive green and violet reflections. Decumbent setae present on ventrite 1. Ventrites 26 have sparse short brown erect setae present throughout, but often abraded.
</paragraph>
</subSubSection>
<subSubSection id="78806524FF9DF125FF17796DFDEFEAD7" pageId="25" pageNumber="275" type="etymology">
<paragraph id="302536AFFF9DF125FF17796DFECAEA49" blockId="25.[145,761,1339,1486]" box="[145,274,1339,1363]" pageId="25" pageNumber="275">
<heading id="6B6D81C3FF9DF125FF17796DFECAEA49" box="[145,274,1339,1363]" fontSize="10" level="4" pageId="25" pageNumber="275" reason="1">
<emphasis id="02EEEABDFF9DF125FF17796DFECAEA49" box="[145,274,1339,1363]" italics="true" pageId="25" pageNumber="275">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="302536AFFF9DF125FF17790AFDEFEAD7" blockId="25.[145,761,1339,1486]" pageId="25" pageNumber="275">Named for the dominant mesquite-chaparral plant community found throughout the species range. Also, this species has been collected at Chaparral Wildlife Management Area, in Cotulla, Texas.</paragraph>
</subSubSection>
<subSubSection id="78806524FF9DF125FF1779B3FD5BE96D" pageId="25" pageNumber="275" type="biology_ecology">
<paragraph id="302536AFFF9DF125FF1779B3FE61EAE7" blockId="25.[145,761,1509,1655]" box="[145,441,1509,1533]" pageId="25" pageNumber="275">
<heading id="6B6D81C3FF9DF125FF1779B3FE61EAE7" box="[145,441,1509,1533]" fontSize="10" level="4" pageId="25" pageNumber="275" reason="1">
<emphasis id="02EEEABDFF9DF125FF1779B3FE61EAE7" box="[145,441,1509,1533]" italics="true" pageId="25" pageNumber="275">Ecology/natural history</emphasis>
</heading>
</paragraph>
<paragraph id="302536AFFF9DF125FF177A50FD5BE96D" blockId="25.[145,761,1509,1655]" pageId="25" pageNumber="275">
Very little is known about this species, as most museum specimens are older, and nothing has been recorded about its ecology. The authors of this study have not observed
<emphasis id="02EEEABDFF9DF125FEE37A34FDA4E96D" box="[357,636,1633,1655]" italics="true" pageId="25" pageNumber="275">
<taxonomicName id="F79A4D2CFF9DF125FEE37A34FDF4E96C" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[357,556,1633,1655]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="275" phylum="Arthropoda" rank="species" species="chaparralensis">D. chaparralensis</taxonomicName>
in situ
</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>