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<mods:title id="A95986A7057C38F87EA77FF013C1938A">Cryptic diversity in the North American Dromochorus tiger beetles (Coleoptera: Carabidae: Cicindelinae): a congruence-based method for species discovery</mods:title>
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<taxonomicName id="F79A4D2CFF91F129FCCB7C90FBA4EFC1" ID-CoL="8LVBX" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[845,1148,198,221]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="281" phylum="Arthropoda" rank="species" species="knisleyi">
<emphasis id="02EEEABDFF91F129FCCB7C90FBA4EFC1" bold="true" box="[845,1148,198,221]" italics="true" pageId="21" pageNumber="271">DROMOCHORUS KNISLEYI</emphasis>
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DURAN, HERRMANN,
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ROMAN &amp; EGAN
<emphasis id="02EEEABDFF91F129FB0D7CB1FB36EFE6" bold="true" box="[1163,1262,228,252]" pageId="21" pageNumber="271">SP. NOV.</emphasis>
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<paragraph id="302536AFFF91F129FC7F7D5DFB18EE39" blockId="21.[845,1388,198,291]" box="[1017,1216,267,291]" pageId="21" pageNumber="271">
<heading id="6B6D81C3FF91F129FC7F7D5DFB18EE39" allCaps="true" box="[1017,1216,267,291]" centered="true" fontSize="10" level="1" pageId="21" pageNumber="271" reason="1">
(
<figureCitation id="A8A12A2AFF91F129FB877D5DFB85EE39" box="[1025,1117,267,291]" captionStart="Figure 7" captionStartId="14.[163,243,1861,1883]" captionTargetBox="[166,1441,1284,1809]" captionTargetId="figure@14.[163,1443,1268,1820]" captionTargetPageId="14" captionText="Figure 7. Labial palps of: A, D. pruininus; B, D. belfragei; C, D. knisleyi sp. nov. A and B illustrate labial palps with a contrasting darker apical segment, and C illustrates labial palps that are consistently dark throughout." httpUri="https://zenodo.org/record/3089275/files/figure.png" pageId="21" pageNumber="271">FIGS 7C</figureCitation>
,
<figureCitation id="A8A12A2AFF91F129FBEC7D5DFB53EE39" box="[1130,1163,267,291]" captionStart="Figure 8" captionStartId="16.[163,241,1096,1118]" captionTargetBox="[330,1279,195,1055]" captionTargetId="figure@16.[323,1283,195,1055]" captionTargetPageId="16" captionText="Figure 8. A, Elytra showing distinct punctures (foveae) running parallel to the suture of the elytra (D. knisleyi sp. nov.). B, lack of distinct subsutural foveae (D. welderensis sp. nov.)." httpUri="https://zenodo.org/record/3089283/files/figure.png" pageId="21" pageNumber="271">8A</figureCitation>
,
<figureCitation id="A8A12A2AFF91F129FB1E7D5DFB60EE39" box="[1176,1208,267,291]" captionStart="Figure 9" captionStartId="17.[145,223,1847,1869]" captionTargetBox="[178,1403,196,1806]" captionTargetId="figure@17.[164,1406,196,1806]" captionTargetPageId="17" captionText="Figure 9. Dorsal habitus for species of Dromochorus. A, D.pruininus; B, D.belfragei; C, D. knisleyi sp. nov.; D, D. minimus sp. nov.; E, D. chaparralensis sp. nov.; F, D. welderensis sp. nov.; G, D. velutinigrens; H, D. pilatei." httpUri="https://zenodo.org/record/3089287/files/figure.png" pageId="21" pageNumber="271">9C</figureCitation>
)
</heading>
</paragraph>
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<subSubSection id="78806524FF91F129FCAF7D14FB8DEE62" pageId="21" pageNumber="271" type="vernacular_names">
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<heading id="6B6D81C3FF91F129FCAF7D14FC39EE43" box="[809,993,322,345]" fontSize="10" level="4" pageId="21" pageNumber="271" reason="1">
<emphasis id="02EEEABDFF91F129FCAF7D14FC39EE43" box="[809,993,322,345]" italics="true" pageId="21" pageNumber="271">Common name</emphasis>
</heading>
</paragraph>
<paragraph id="302536AFFF91F129FCAF7D34FB8DEE62" blockId="21.[809,1109,322,376]" box="[809,1109,354,376]" pageId="21" pageNumber="271">Juniper grove tiger beetle.</paragraph>
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<subSubSection id="78806524FF91F129FCAF7DD9FADEEC0C" pageId="21" pageNumber="271" type="materials_examined">
<paragraph id="302536AFFF91F129FCAF7DD9FC1CEEBC" blockId="21.[809,1425,398,790]" box="[809,964,398,422]" pageId="21" pageNumber="271">
<heading id="6B6D81C3FF91F129FCAF7DD9FC1CEEBC" box="[809,964,398,422]" fontSize="10" level="4" pageId="21" pageNumber="271" reason="1">
<emphasis id="02EEEABDFF91F129FCAF7DD9FC1CEEBC" box="[809,964,398,422]" italics="true" pageId="21" pageNumber="271">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="302536AFFF91F129FCAF7DF9FADEEC0C" blockId="21.[809,1425,398,790]" pageId="21" pageNumber="271">
<materialsCitation id="80F23CF2FF91F129FCAF7DF9FAD9ED18" ID-GBIF-Occurrence="2243053341" collectingDate="2013-06-19" collectionCode="USNM" collectorName="D. Duran" country="United States of America" county="Blanco Co." location="Vi c i n i t y o f Pe d e r n a l e s Fa l l s, Te x a s" pageId="21" pageNumber="271" specimenCount="1" specimenCount-male="1" stateProvince="Texas" typeStatus="holotype">
<location id="35456074FF91F129FCAF7DF9FAD1EEDE" LSID="urn:lsid:plazi:treatment:B83387B9FF91F12AFCCB7C90FC20EAB5:35456074FF91F129FCAF7DF9FAD1EEDE" box="[809,1289,431,453]" country="United States of America" county="Blanco Co." name="Vi c i n i t y o f Pe d e r n a l e s Fa l l s, Te x a s" pageId="21" pageNumber="271" stateProvince="Texas">Vi c i n i t y o f Pe d e r n a l e s Fa l l s, Te x a s</location>
.
<typeStatus id="EF21880DFF91F129FA9D7DF9FA48EEDE" box="[1307,1424,431,453]" pageId="21" pageNumber="271">H o l o t y p e</typeStatus>
(
<collectionCode id="568BAE6AFF91F129FCB77D98FC53EEFE" box="[817,907,462,484]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="21" pageNumber="271">USNM</collectionCode>
):
<specimenCount id="269CFD26FF91F129FC267D98FC13EEFE" box="[928,971,462,484]" pageId="21" pageNumber="271" type="male">1 ♂</specimenCount>
,
<collectingCountry id="488D763FFF91F129FC587D98FBC3EEF9" box="[990,1051,462,483]" name="United States of America" pageId="21" pageNumber="271">USA</collectingCountry>
:
<collectingRegion id="F25EF84DFF91F129FBA17D98FBAAEEF9" box="[1063,1138,462,483]" country="United States of America" name="Texas" pageId="21" pageNumber="271">Texas</collectingRegion>
:
<collectingCounty id="D9444E23FF91F129FBF97D98FADCEEFE" box="[1151,1284,462,484]" pageId="21" pageNumber="271">Blanco Co.</collectingCounty>
/
<location id="35456074FF91F129FA8B7D98FC6BED18" LSID="urn:lsid:plazi:treatment:B83387B9FF91F12AFCCB7C90FC20EAB5:35456074FF91F129FA8B7D98FC6BED18" country="United States of America" county="Blanco Co." name="Vicinity of Pedernales" pageId="21" pageNumber="271" stateProvince="Texas">Vicinity of Pedernales</location>
/
<collectingDate id="5460E987FF91F129FC3B7DBAFB91ED18" box="[957,1097,492,514]" pageId="21" pageNumber="271" value="2013-06-19">
<date id="4424106FFF91F129FC3B7DBAFB91ED18" box="[957,1097,492,514]" pageId="21" pageNumber="271" value="2013-06-19">19-VI-2013</date>
</collectingDate>
/leg
<collectorName id="9D6F5379FF91F129FB047DBBFB25ED18" box="[1154,1277,493,514]" pageId="21" pageNumber="271">D. Duran</collectorName>
.
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Paratypes: 14 ♂♂, 19 ♀♀, USA: Texas: Blanco Co./Vicinity of Pedernales Falls St. Pk./19-VI-2013/leg D. Duran. 6 ♂♂, 7 ♀♀, USA: Texas: Blanco Co./Vicinity of Pedernales Falls St. Pk./08-VI-2015/leg S.J Roman. 5 ♂♂, 3 ♀♀, USA: Texas: Blanco Co./Vicinity of Pedernales Falls St. Pk./01-VI-2014/leg D. Brzoska. 2 ♂, USA: Texas: Blanco Co./Park E. of Pedernales Falls S.P./20-VI-2014/ leg D. Duran. 2 ♀, USA: TEXAS: Bandera Co./3 miles W of Pipe Creek/22-VI-2013/leg D. Duran.
</paragraph>
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<emphasis id="02EEEABDFF91F129FCAF7F6CFC18EC48" box="[809,960,826,850]" italics="true" pageId="21" pageNumber="271">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="302536AFFF91F129FCAF7F0DFC23EB7C" blockId="21.[809,1426,826,1126]" pageId="21" pageNumber="271">
This species is only known from the Edwards Plateau of central Texas, locally known as the Texas Hill Country. It comes in close geographic proximity to
<taxonomicName id="F79A4D2CFF91F129FA8B7FCFFA49ECB7" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[1293,1425,920,942]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="271" phylum="Arthropoda" rank="species" species="belfragei">
<emphasis id="02EEEABDFF91F129FA8B7FCFFA49ECB7" box="[1293,1425,920,942]" italics="true" pageId="21" pageNumber="271">D. belfragei</emphasis>
</taxonomicName>
and
<taxonomicName id="F79A4D2CFF91F129FCDD7FEEFC39ECD6" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[859,993,951,973]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="271" phylum="Arthropoda" rank="species" species="minimus">
<emphasis id="02EEEABDFF91F129FCDD7FEEFC39ECD6" box="[859,993,951,973]" italics="true" pageId="21" pageNumber="271">D. minimus</emphasis>
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, where all three species distributions converge at the edge of the Balcones Escarpment in Bexar County. Analyses of mtDNA data indicate that hybridization occurs in this contact zone, and apparent hybrid
<taxonomicName id="F79A4D2CFF91F129FC227864FBC1EB5C" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[932,1049,1073,1095]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="271" phylum="Arthropoda" rank="species" species="knisleyi">
<emphasis id="02EEEABDFF91F129FC227864FBC1EB5C" box="[932,1049,1073,1095]" italics="true" pageId="21" pageNumber="271">D. knisleyi</emphasis>
</taxonomicName>
x
<taxonomicName id="F79A4D2CFF91F129FBB47867FB4DEB5C" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[1074,1173,1073,1094]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="271" phylum="Arthropoda" rank="species" species="belfragei">
<emphasis id="02EEEABDFF91F129FBB47867FB4DEB5C" box="[1074,1173,1073,1094]" italics="true" pageId="21" pageNumber="271">belfragei</emphasis>
</taxonomicName>
individuals have been found in this area.
</paragraph>
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<emphasis id="02EEEABDFF91F129FCAF78D5FC7AEB80" box="[809,930,1155,1178]" italics="true" pageId="21" pageNumber="271">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="302536AFFF91F129FCAF78F5FBBFEBEC" blockId="21.[809,1425,1155,1270]" pageId="21" pageNumber="271">
<taxonomicName id="F79A4D2CFF91F129FCAF78F5FBC7EBA2" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[809,1055,1187,1208]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="271" phylum="Arthropoda" rank="species" species="knisleyi">
<emphasis id="02EEEABDFF91F129FCAF78F5FBC7EBA2" box="[809,1055,1187,1208]" italics="true" pageId="21" pageNumber="271">Dromochorus knisleyi</emphasis>
</taxonomicName>
is most easily confused with the sister taxa
<taxonomicName id="F79A4D2CFF91F129FC2D7894FBF6EBCC" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[939,1070,1217,1239]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="271" phylum="Arthropoda" rank="species" species="belfragei">
<emphasis id="02EEEABDFF91F129FC2D7894FBF6EBCC" box="[939,1070,1217,1239]" italics="true" pageId="21" pageNumber="271">D. belfragei</emphasis>
</taxonomicName>
. For differential diagnosis, see
<taxonomicName id="F79A4D2CFF91F129FCAF78B7FC72EBEF" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[809,938,1248,1270]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="271" phylum="Arthropoda" rank="species" species="belfragei">
<emphasis id="02EEEABDFF91F129FCAF78B7FC72EBEF" box="[809,938,1248,1270]" italics="true" pageId="21" pageNumber="271">D. belfragei</emphasis>
</taxonomicName>
species account.
</paragraph>
</subSubSection>
<subSubSection id="78806524FF91F12AFCAF7958FC47EEA3" lastPageId="22" lastPageNumber="272" pageId="21" pageNumber="271" type="description">
<paragraph id="302536AFFF91F129FCAF7958FC6DEA3F" blockId="21.[809,1427,1294,1869]" box="[809,949,1294,1317]" pageId="21" pageNumber="271">
<heading id="6B6D81C3FF91F129FCAF7958FC6DEA3F" box="[809,949,1294,1317]" fontSize="10" level="4" pageId="21" pageNumber="271" reason="1">
<emphasis id="02EEEABDFF91F129FCAF7958FC6DEA3F" box="[809,949,1294,1317]" italics="true" pageId="21" pageNumber="271">Description</emphasis>
</heading>
</paragraph>
<paragraph id="302536AFFF91F12AFCAF7978FD68EDFC" blockId="21.[809,1427,1294,1869]" lastBlockId="22.[163,780,198,742]" lastPageId="22" lastPageNumber="272" pageId="21" pageNumber="271">
Medium-sized
<taxonomicName id="F79A4D2CFF91F129FC617978FB48EA59" box="[999,1168,1326,1347]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="271" phylum="Arthropoda" rank="genus">
<emphasis id="02EEEABDFF91F129FC617978FB48EA59" box="[999,1168,1326,1347]" italics="true" pageId="21" pageNumber="271">Dromochorus</emphasis>
</taxonomicName>
. Body length 10.9 14.4 mm, mean ♀ 13.0 mm, mean ♂ 12.2 mm. Head slightly wider than pronotum.
<emphasis id="02EEEABDFF91F129FB01793DFBADEAC7" bold="true" pageId="21" pageNumber="271">Head predominantly charcoal black with blue reflections mostly concentrated near the anterior margin and edges of the supraorbital region</emphasis>
. Fine rugosity often present on the frons and vertex. All head portions glabrous except for two supraorbital setae next to each eye. Frons concave in median area, especially in male, bulging towards slightly convex near anterior margin, clearly delimited from clypeus, gradually blending into vertex. Genae black or bright polished metallic violet to blue, with shallow longitudinal striae gradually ending at border of vertex. Clypeus bronze with green to blue reflections throughout. Male labrum tridentate with 68 setae, central area pale ochre-testaceous, with a thin dark-brown to black border posteriorly and sometimes anteriorly, dark-brown to black laterally; the pale central area of the labrum may exist as a small spot, up to one-third of the total labrum surface; female labrum tridentate with 68 setae, entirely dark-brown to black with polished metallic cupreous to green reflections.
<emphasis id="02EEEABDFF92F12AFEAA7D17FD82EE8E" bold="true" pageId="22" pageNumber="272">All segments of maxillary and labial palpi consistently dark-brown; apical segment is not darker than other segments</emphasis>
. Antennae normal length, reaching back to humerus and basal third of elytron, slightly longer in male than female; scape dark testaceous to black with metallic reflections of violet, cupreous and green, with 23 apical setae; pedicel dark testaceous with metallic reflections of violet, cupreous and green, lacking any setae; flagellum dark testaceous, antennomeres 34 with metallic violet and green reflections, densely clothed in short white setae, antennomeres 511 dull-textured without metallic reflections and possessing erect setae in apical rings only, covered with fine pubescence throughout.
</paragraph>
<paragraph id="302536AFFF92F12AFF257EA8FE29EAEE" blockId="22.[163,780,766,1524]" pageId="22" pageNumber="272">
<emphasis id="02EEEABDFF92F12AFF257EA8FF23EC09" box="[163,251,766,787]" italics="true" pageId="22" pageNumber="272">Thorax</emphasis>
: Pronotum 2.43.2 mm in length, mean ♀ 2.9 mm, mean ♂ 2.8 mm; width 2.53.2 mm, mean ♀ 3.0 mm, mean ♂ 2.9 mm. Pronotum charcoal black, with green to blue or violet reflections, especially along lateral margins, slightly wider than long, widest near anterior margin, width to length ratio 0.9 to 1.1, setae sparse to regularly spaced, mostly present along lateral third of dorsal surface; disc finely rugose, with thin but distinct median line, with well-defined shallow sulci present anteriorly and posteriorly; notopleural sutures clearly defined, not visible from dorsal view; proepisternum black with weak to strong iridescent blue to violet reflections, glabrous. Elytra elongate, 6.88.7 mm length, mean ♀ 7.9 mm, mean ♂ 7.7 mm, shape similar in both sexes, but slightly wider in female, especially toward apical third; sutural spine absent, microserrations not present on elytral apices; elytral dorsal surface convex;
<emphasis id="02EEEABDFF92F12AFE847970FEADEA82" bold="true" pageId="22" pageNumber="272">elytral texture dull, with regular small pits present throughout disk, elytral coloration charcoal black, often with blue reflections near humeral region</emphasis>
; elytral maculations absent;
<emphasis id="02EEEABDFF92F12AFD5C79D4FE32EAEE" bold="true" pageId="22" pageNumber="272">two dark oblique infuscations present; subsutural foveae prominent, typically with metallic blue, green, or gold reflections</emphasis>
.
</paragraph>
<paragraph id="302536AFFF92F12AFF257A46FC29EFE1" blockId="22.[163,779,1551,1880]" lastBlockId="22.[827,1443,198,251]" pageId="22" pageNumber="272">
<emphasis id="02EEEABDFF92F12AFF257A46FF03E93F" box="[163,219,1552,1573]" italics="true" pageId="22" pageNumber="272">Legs</emphasis>
: Pro-, meso-, and metacoxae dark testaceous to black with iridescent blue to violet and cupreous reflections, sparse setae on pro- and mesocoxae, fewer on metacoxae; pro- and mesotrochanters with a single erect seta, metatrochanter glabrous, trochanters dark brown-testaceous; femora black with metallic violet and green reflections, densely clothed in decumbent white setae; tibiae testaceous brown, clothed with setae of two types: sparser brown-testaceous long setae and dense short decumbent white setae; two tibial spines present; tarsi brown-testaceous, first three dilated protarsomeres in male with dense greyishwhite setal pad.
</paragraph>
<paragraph id="302536AFFF92F12AFCBD7D7FFC47EEA3" blockId="22.[827,1443,297,441]" pageId="22" pageNumber="272">
<emphasis id="02EEEABDFF92F12AFCBD7D7FFC7EEE24" box="[827,934,297,318]" italics="true" pageId="22" pageNumber="272">Abdomen</emphasis>
: Venter mostly black with occasional metallic green to violet reflections. Decumbent white setae present on ventrite 1. Ventrites 26 have sparse short brown erect setae present throughout, but often abraded.
</paragraph>
</subSubSection>
<subSubSection id="78806524FF92F12AFCBD7D87FC6FED98" pageId="22" pageNumber="272" type="etymology">
<paragraph id="302536AFFF92F12AFCBD7D87FC64EEF3" blockId="22.[827,1443,465,642]" box="[827,956,465,489]" pageId="22" pageNumber="272">
<heading id="6B6D81C3FF92F12AFCBD7D87FC64EEF3" box="[827,956,465,489]" fontSize="10" level="4" pageId="22" pageNumber="272" reason="1">
<emphasis id="02EEEABDFF92F12AFCBD7D87FC64EEF3" box="[827,956,465,489]" italics="true" pageId="22" pageNumber="272">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="302536AFFF92F12AFCBD7DA7FC6FED98" blockId="22.[827,1443,465,642]" pageId="22" pageNumber="272">Named for Dr C. Barry Knisley, one of the leading authorities on North American tiger beetle conservation and ecology. D.P. Duran and R.A. Gwiazdowski are greatly indebted to Barry for his mentorship and friendship.</paragraph>
</subSubSection>
<subSubSection id="78806524FF92F12AFCBD7ECFFC20EAB5" pageId="22" pageNumber="272" type="biology_ecology">
<paragraph id="302536AFFF92F12AFCBD7ECFFBBBEDAB" blockId="22.[827,1444,665,1455]" box="[827,1123,665,689]" pageId="22" pageNumber="272">
<heading id="6B6D81C3FF92F12AFCBD7ECFFBBBEDAB" box="[827,1123,665,689]" fontSize="10" level="4" pageId="22" pageNumber="272" reason="1">
<emphasis id="02EEEABDFF92F12AFCBD7ECFFBBBEDAB" box="[827,1123,665,689]" italics="true" pageId="22" pageNumber="272">Ecology/natural history</emphasis>
</heading>
</paragraph>
<paragraph id="302536AFFF92F12AFCBD7EEFFB51EC16" blockId="22.[827,1444,665,1455]" pageId="22" pageNumber="272">
<taxonomicName id="F79A4D2CFF92F12AFCBD7EEFFBE7EDD4" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[827,1087,697,718]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="272" phylum="Arthropoda" rank="species" species="knisleyi">
<emphasis id="02EEEABDFF92F12AFCBD7EEFFBE7EDD4" box="[827,1087,697,718]" italics="true" pageId="22" pageNumber="272">Dromochorus knisleyi</emphasis>
</taxonomicName>
adults have been found from mid-May to late June, but it is likely that they could be active outside of this window.
</paragraph>
<paragraph id="302536AFFF92F12AFCD57F43FB00EA48" blockId="22.[827,1444,665,1455]" pageId="22" pageNumber="272">
<taxonomicName id="F79A4D2CFF92F12AFCD57F43FB8EEC30" authorityName="Duran &amp; Herrmann &amp; Roman &amp; Gwiazdowski &amp; Drummond &amp; Hood &amp; Egan" authorityYear="2019" box="[851,1110,789,810]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="272" phylum="Arthropoda" rank="species" species="knisleyi">
<emphasis id="02EEEABDFF92F12AFCD57F43FB8EEC30" box="[851,1110,789,810]" italics="true" pageId="22" pageNumber="272">Dromochorus knisleyi</emphasis>
</taxonomicName>
is found in upland juniperoak woodlands in the Edwards Plateau, and does not appear to be strongly associated with riparian areas. The preferred habitat is late succession stands of juniper and, as such, it can be difficult for a collector to easily walk through these areas. Adult beetles are active throughout the day and are present in semiopen grassy areas under the cover of juniper trees. The first author observed dozens of beetles over a span of two days, and all adult activity was restricted to these forested areas. Beetles foraged and mated exclusively near or under juniper boughs. Moreover, even during cloudy periods and late in the afternoon, none were observed moving into more open grassy areas outside of the juniper stands. Moreover, beetles were not present in woodlands dominated by oaks. In mixed juni-peroak woodlands, beetles were found exclusively near junipers. This species may be the most ecologically specialized of all
<taxonomicName id="F79A4D2CFF92F12AFBBC796BFB0AEA48" box="[1082,1234,1341,1362]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="02EEEABDFF92F12AFBBC796BFB0AEA48" box="[1082,1234,1341,1362]" italics="true" pageId="22" pageNumber="272">Dromochorus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="302536AFFF92F12AFCD5790AFC20EAB5" blockId="22.[827,1444,665,1455]" pageId="22" pageNumber="272">More observations are needed for this rarely collected species. Many aspects of the biology are currently unknown.</paragraph>
</subSubSection>
</treatment>
</document>