498 lines
72 KiB
XML
498 lines
72 KiB
XML
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<mods:title id="26F07D54424AF0BA34BFCE71F4A7C69F">Cryptic diversity in the North American Dromochorus tiger beetles (Coleoptera: Carabidae: Cicindelinae): a congruence-based method for species discovery</mods:title>
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<mods:namePart id="DB4F8B45C1E0D8FF7965591085EF0DA9">Duran, Daniel P.</mods:namePart>
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<mods:namePart id="588057A1D78BF2BF4CBBAAD08C5BAC59">Herrmann, David P.</mods:namePart>
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<mods:namePart id="76E4D791C7AEEBBC5B3C18FE6282A488">Gwiazdowski, Rodger A.</mods:namePart>
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<mods:namePart id="7665B029EFAA693E51A48850D9D691C4">Drummond, Jennifer A.</mods:namePart>
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<mods:namePart id="0C25337877B58AAB17496712BFB62E46">Hood, Glen R.</mods:namePart>
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<mods:namePart id="316DA3E3053257E4048BEC5EAE513FCD">Egan, Scott P.</mods:namePart>
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<mods:title id="950536E62BEEF9E7D31669C06B732E13">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="3BDC66F354EFCE023991DD7EC7B109A3">2019</mods:date>
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<mods:number id="F9204DB51716493FB0642BF0E72AC661">2018-09-12</mods:number>
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<mods:identifier id="8B3308FC75307D7BDA821D2078489258" type="DOI">10.1093/zoolinnean/zly035</mods:identifier>
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<treatment id="B83387B9FF89F133FF20799CFAA5EBEC" ID-DOI="http://doi.org/10.5281/zenodo.5942797" ID-GBIF-Taxon="157232839" ID-Zenodo-Dep="5942797" LSID="urn:lsid:plazi:treatment:B83387B9FF89F133FF20799CFAA5EBEC" httpUri="http://treatment.plazi.org/id/B83387B9FF89F133FF20799CFAA5EBEC" lastPageId="15" lastPageNumber="265" pageId="13" pageNumber="263">
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<subSubSection id="78806524FF89F131FF20799CFEDDE94F" pageId="13" pageNumber="263" type="nomenclature">
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<paragraph id="302536AFFF89F131FF20799CFD3BEAF8" blockId="13.[166,739,1482,1506]" box="[166,739,1482,1506]" pageId="13" pageNumber="263">
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<heading id="6B6D81C3FF89F131FF20799CFD3BEAF8" allCaps="true" box="[166,739,1482,1506]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="263" reason="5">
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GENUS
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<taxonomicName id="F79A4D2CFF89F131FF7B799DFE77EAFB" ID-CoL="8KW25" box="[253,431,1483,1506]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="02EEEABDFF89F131FF7B799DFE77EAFB" box="[253,431,1483,1506]" italics="true" pageId="13" pageNumber="263">DROMOCHORUS</emphasis>
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GUÉRIN- MÉNEVILLE, 1845
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<paragraph id="302536AFFF89F131FF177A54FEFDE903" blockId="13.[145,760,1538,1621]" box="[145,293,1538,1561]" pageId="13" pageNumber="263">
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<heading id="6B6D81C3FF89F131FF177A54FEFDE903" box="[145,293,1538,1561]" fontSize="10" level="4" pageId="13" pageNumber="263" reason="1">
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<emphasis id="02EEEABDFF89F131FF177A54FEFDE903" box="[145,293,1538,1561]" italics="true" pageId="13" pageNumber="263">Type species</emphasis>
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</heading>
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</paragraph>
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<paragraph id="302536AFFF89F131FF177A77FEDDE94F" blockId="13.[145,760,1538,1621]" pageId="13" pageNumber="263">
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<taxonomicName id="F79A4D2CFF89F131FF177A77FD12E92D" authority="Guerin-Meneville, 1845" authorityName="Guerin-Meneville" authorityYear="1845" box="[145,714,1569,1591]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="species" species="pilatei">
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<emphasis id="02EEEABDFF89F131FF177A77FE4BE92C" box="[145,403,1569,1590]" italics="true" pageId="13" pageNumber="263">Dromochorus pilatei</emphasis>
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<bibRefCitation id="540B4B5EFF89F131FE1B7A77FD12E92D" author="Guerin-Meneville FE" box="[413,714,1569,1591]" journalOrPublisher="Annales de la Societe Entomologique de France (Bulletin Entomologique) Ser 2" pageId="13" pageNumber="263" pagination="1 - 4" part="7" refId="ref26455" refString="Guerin-Meneville FE. 1845. Sur le Dromochorus, nouveau genre de Cicindeletes. Annales de la Societe Entomologique de France (Bulletin Entomologique) Ser 2 7: 1 - 4." title="Sur le Dromochorus, nouveau genre de Cicindeletes" type="journal article" year="1845">Guérin-Méneville, 1845</bibRefCitation>
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</taxonomicName>
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. By monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection id="78806524FF89F132FF177A38FD36EE03" lastPageId="14" lastPageNumber="264" pageId="13" pageNumber="263" type="discussion">
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<paragraph id="302536AFFF89F131FF177A38FEA0E99F" blockId="13.[145,761,1645,1914]" box="[145,376,1645,1669]" pageId="13" pageNumber="263">
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<heading id="6B6D81C3FF89F131FF177A38FEA0E99F" box="[145,376,1645,1669]" fontSize="10" level="4" pageId="13" pageNumber="263" reason="1">
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<emphasis id="02EEEABDFF89F131FF177A38FEA0E99F" box="[145,376,1645,1669]" italics="true" pageId="13" pageNumber="263">Taxonomic history</emphasis>
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</heading>
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</paragraph>
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<paragraph id="302536AFFF89F131FF177ADBFAA4EB80" blockId="13.[145,761,1645,1914]" lastBlockId="13.[809,1427,727,1914]" pageId="13" pageNumber="263">
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<taxonomicName id="F79A4D2CFF89F131FF177ADBFEEEE9B8" box="[145,310,1677,1698]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="02EEEABDFF89F131FF177ADBFEEEE9B8" box="[145,310,1677,1698]" italics="true" pageId="13" pageNumber="263">Dromochorus</emphasis>
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</taxonomicName>
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was described by
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<bibRefCitation id="540B4B5EFF89F131FD987ADBFF05E9D8" author="Guerin-Meneville FE" journalOrPublisher="Annales de la Societe Entomologique de France (Bulletin Entomologique) Ser 2" pageId="13" pageNumber="263" pagination="1 - 4" part="7" refId="ref26455" refString="Guerin-Meneville FE. 1845. Sur le Dromochorus, nouveau genre de Cicindeletes. Annales de la Societe Entomologique de France (Bulletin Entomologique) Ser 2 7: 1 - 4." title="Sur le Dromochorus, nouveau genre de Cicindeletes" type="journal article" year="1845">Guérin-Méneville (1845)</bibRefCitation>
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as a new genus, believed to be most closely related to ‘
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<taxonomicName id="F79A4D2CFF89F131FE977A9DFE49E9FA" authorityName="Hope" authorityYear="1838" box="[273,401,1739,1760]" class="Insecta" family="Carabidae" genus="Apteroessa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="02EEEABDFF89F131FE977A9DFE49E9FA" box="[273,401,1739,1760]" italics="true" pageId="13" pageNumber="263">Apteroessa</emphasis>
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</taxonomicName>
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and especially
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<taxonomicName id="F79A4D2CFF89F131FDCD7A9DFD69E9FA" authorityName="Dejean" authorityYear="1826" box="[587,689,1739,1760]" class="Insecta" family="Carabidae" genus="Dromica" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="02EEEABDFF89F131FDCD7A9DFD69E9FA" box="[587,689,1739,1760]" italics="true" pageId="13" pageNumber="263">Dromica</emphasis>
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</taxonomicName>
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’. This perceived relatedness was due to the remarkable similarity in
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<emphasis id="02EEEABDFF89F131FEAD7B5EFEA1E807" box="[299,377,1800,1821]" italics="true" pageId="13" pageNumber="263">gestalt</emphasis>
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, although several morphological differences were identified, especially with respect to the mouthparts, such as the maxillary and labial palpi, and labrum. Subsequent authors have not shared his view about the systematic placement of the genus within the larger tiger beetle clade. In his revision of the
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<taxonomicName id="F79A4D2CFF89F131FBC07F42FA4BEC30" authority="Horn (1908)" authorityName="Horn" authorityYear="1908" box="[1094,1427,788,810]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="subFamily" subFamily="Cicindelinae">
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Cicindelinae,
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<bibRefCitation id="540B4B5EFF89F131FB717F42FA4BEC30" author="Horn W." box="[1271,1427,788,810]" editor="Wytsman P" journalOrPublisher="82 a" pageId="13" pageNumber="263" pagination="1 - 104" refId="ref26813" refString="Horn W. 1908. Cicindelinae. In: Wytsman P, ed. Genera insectorum. 82 a: 1 - 104." title="Cicindelinae" type="book chapter" volumeTitle="Genera insectorum" year="1908">Horn (1908)</bibRefCitation>
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</taxonomicName>
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placed
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<taxonomicName id="F79A4D2CFF89F131FC057F64FBF1EC5D" box="[899,1065,818,839]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="02EEEABDFF89F131FC057F64FBF1EC5D" box="[899,1065,818,839]" italics="true" pageId="13" pageNumber="263">Dromochorus</emphasis>
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</taxonomicName>
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in the subtribe Cicindelina, and
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<taxonomicName id="F79A4D2CFF89F131FCE77F04FC13EC7D" authorityName="Dejean" authorityYear="1826" box="[865,971,850,871]" class="Insecta" family="Carabidae" genus="Dromica" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="02EEEABDFF89F131FCE77F04FC13EC7D" box="[865,971,850,871]" italics="true" pageId="13" pageNumber="263">Dromica</emphasis>
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</taxonomicName>
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in a separate subtribe, Prothymina (
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<bibRefCitation id="540B4B5EFF89F131FCB47F26FC60EC9F" author="Horn W." box="[818,952,880,901]" editor="Wytsman P" journalOrPublisher="Genera insectorum." pageId="13" pageNumber="263" pagination="1 - 15" part="82" refId="ref26837" refString="Horn W. 1910. Cicindelinae. In: Wytsman P, ed. Genera insectorum. 82 b: 105 - 208, plates 1 - 15." title="Cicindelinae" type="journal article" year="1910">Horn 1910</bibRefCitation>
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). More recent molecular phylogenies have supported
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<bibRefCitation id="540B4B5EFF89F131FC797FD9FB6AECBF" author="Horn W." box="[1023,1202,911,933]" editor="Wytsman P" journalOrPublisher="82 a" pageId="13" pageNumber="263" pagination="1 - 104" refId="ref26813" refString="Horn W. 1908. Cicindelinae. In: Wytsman P, ed. Genera insectorum. 82 a: 1 - 104." title="Cicindelinae" type="book chapter" volumeTitle="Genera insectorum" year="1908">Horn’s (1908)</bibRefCitation>
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treatment, finding
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<taxonomicName id="F79A4D2CFF89F131FCDA7FFBFBDCECD8" box="[860,1028,941,962]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
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||
<emphasis id="02EEEABDFF89F131FCDA7FFBFBDCECD8" box="[860,1028,941,962]" italics="true" pageId="13" pageNumber="263">Dromochorus</emphasis>
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</taxonomicName>
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to be nested within a clade of other Nearctic Cicindelina (
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<bibRefCitation id="540B4B5EFF89F131FB047F9AFA53ECFB" author="Vogler AP & Welsh A." box="[1154,1419,972,994]" journalOrPublisher="Molecular phylogenetics and evolution" pageId="13" pageNumber="263" pagination="225 - 235" part="8" refId="ref28298" refString="Vogler AP, Welsh A. 1997. Phylogeny of North American Cicindela tiger beetles inferred from multiple mitochondrial DNA sequences. Molecular phylogenetics and evolution 8: 225 - 235." title="Phylogeny of North American Cicindela tiger beetles inferred from multiple mitochondrial DNA sequences" type="journal article" year="1997">Vogler & Welsh, 1997</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FCAF7FBCFBB4EB1A" author="Barraclough TG & Vogler AP" box="[809,1132,1002,1024]" journalOrPublisher="Molecular Biology and Evolution" pageId="13" pageNumber="263" pagination="1706 - 1716" part="19" refId="ref25096" refString="Barraclough TG, Vogler AP. 2002. Recent diversification rates in North American tiger beetles estimated from a dated mtDNA phylogenetic tree. Molecular Biology and Evolution 19: 1706 - 1716." title="Recent diversification rates in North American tiger beetles estimated from a dated mtDNA phylogenetic tree" type="journal article" year="2002">Barraclough & Vogler, 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FBFC7FBDFAE1EB1A" author="Pons J & Barraclough T & Theodorides K & Cardoso A & Vogler A." box="[1146,1337,1002,1024]" journalOrPublisher="Systematic Biology" pageId="13" pageNumber="263" pagination="554 - 570" part="53" refId="ref27711" refString="Pons J, Barraclough T, Theodorides K, Cardoso A, Vogler A. 2004. Using exon and intron sequences of the gene Mp 20 to resolve basal relationships in Cicindela (Coleoptera: Cicindelidae). Systematic Biology 53: 554 - 570." title="Using exon and intron sequences of the gene Mp 20 to resolve basal relationships in Cicindela (Coleoptera: Cicindelidae)" type="journal article" year="2004">
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||
Pons
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||
<emphasis id="02EEEABDFF89F131FB3F7FBDFB2BECE5" box="[1209,1267,1002,1024]" italics="true" pageId="13" pageNumber="263">et al.</emphasis>
|
||
, 2004
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FAC07FBCFC61EB05" author="Vogler AP & Cardoso A & Barraclough TG" journalOrPublisher="Systematic Biology" pageId="13" pageNumber="263" pagination="4 - 20" part="54" refId="ref28330" refString="Vogler AP, Cardoso A, Barraclough TG. 2005. Exploring rate variation among and within sites in a densely sampled tree: Species level phylogenetics of North American tiger beetles (genus Cicindela). Systematic Biology 54: 4 - 20." title="Exploring rate variation among and within sites in a densely sampled tree: Species level phylogenetics of North American tiger beetles (genus Cicindela)" type="journal article" year="2005">
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||
Vogler
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<emphasis id="02EEEABDFF89F131FCAF785CFCBAEB04" box="[809,866,1033,1055]" italics="true" pageId="13" pageNumber="263">et al</emphasis>
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||
., 2005
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||
</bibRefCitation>
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||
), and
|
||
<taxonomicName id="F79A4D2CFF89F131FB8A785CFBA0EB05" authorityName="Dejean" authorityYear="1826" box="[1036,1144,1034,1055]" class="Insecta" family="Carabidae" genus="Dromica" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF89F131FB8A785CFBA0EB05" box="[1036,1144,1034,1055]" italics="true" pageId="13" pageNumber="263">Dromica</emphasis>
|
||
</taxonomicName>
|
||
to be more related to
|
||
<taxonomicName id="F79A4D2CFF89F131FCAF787EFACCEB27" authority="(Galian, Hogan & Vogler, 2002)" authorityName="Galian, Hogan & Vogler" authorityYear="2002" baseAuthorityName="Galian, Hogan & Vogler" baseAuthorityYear="2002" box="[809,1300,1064,1086]" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF89F131FCAF787EFC45EB27" box="[809,925,1064,1085]" italics="true" pageId="13" pageNumber="263">Prothyma</emphasis>
|
||
(
|
||
<bibRefCitation id="540B4B5EFF89F131FC2B787EFAD2EB27" author="Galian J & Hogan JE & Vogler AP" box="[941,1290,1064,1086]" journalOrPublisher="Molecular Biology and Evolution" pageId="13" pageNumber="263" pagination="1792 - 1796" part="19" refId="ref26086" refString="Galian J, Hogan JE, Vogler AP. 2002. The origin of multiple sex chromosomes in tiger beetles. Molecular Biology and Evolution 19: 1792 - 1796." title="The origin of multiple sex chromosomes in tiger beetles" type="journal article" year="2002">Galián, Hogan & Vogler, 2002</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Any similarities between
|
||
<taxonomicName id="F79A4D2CFF89F131FC7F7810FB44EB41" box="[1017,1180,1094,1115]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF89F131FC7F7810FB44EB41" box="[1017,1180,1094,1115]" italics="true" pageId="13" pageNumber="263">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F79A4D2CFF89F131FB5D7811FA9CEB46" authorityName="Dejean" authorityYear="1826" box="[1243,1348,1095,1116]" class="Insecta" family="Carabidae" genus="Dromica" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF89F131FB5D7811FA9CEB46" box="[1243,1348,1095,1116]" italics="true" pageId="13" pageNumber="263">Dromica</emphasis>
|
||
</taxonomicName>
|
||
must, therefore, be the result of convergent evolution owing to their similar habitats and microhabitats.
|
||
</paragraph>
|
||
<paragraph id="302536AFFF89F131FCC778F4FA8EE932" blockId="13.[809,1427,727,1914]" pageId="13" pageNumber="263">
|
||
The
|
||
<taxonomicName id="F79A4D2CFF89F131FCFC78F4FBC5EBAD" box="[890,1053,1186,1207]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF89F131FCFC78F4FBC5EBAD" box="[890,1053,1186,1207]" italics="true" pageId="13" pageNumber="263">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
species have sometimes been treated as a distinct genus (
|
||
<bibRefCitation id="540B4B5EFF89F131FBF37897FA52EBCD" author="Guerin-Meneville FE" box="[1141,1418,1217,1239]" journalOrPublisher="Annales de la Societe Entomologique de France (Bulletin Entomologique) Ser 2" pageId="13" pageNumber="263" pagination="1 - 4" part="7" refId="ref26455" refString="Guerin-Meneville FE. 1845. Sur le Dromochorus, nouveau genre de Cicindeletes. Annales de la Societe Entomologique de France (Bulletin Entomologique) Ser 2 7: 1 - 4." title="Sur le Dromochorus, nouveau genre de Cicindeletes" type="journal article" year="1845">Guérin-Méneville, 1845</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FCAF78B6FC77EBEF" author="Salle A." box="[809,943,1248,1270]" journalOrPublisher="Annales de la Societe Entomologique de France (Bulletin des Seances) Ser 5" pageId="13" pageNumber="263" pagination="7 - 8" part="7" refId="ref28062" refString="Salle A. 1877. Note sur le genre Dromochorus. Annales de la Societe Entomologique de France (Bulletin des Seances) Ser 5 7: 7 - 8." title="Note sur le genre Dromochorus" type="journal article" year="1877">Sallé, 1877</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FC3978B6FBA6EBEF" author="Fleutiaux E." box="[959,1150,1248,1270]" journalOrPublisher="Vaillant-Carmanne: Imprimerie H" pageId="13" pageNumber="263" refId="ref26002" refString="Fleutiaux E. 1892. Catalogue systematique des Cicindelidae, decrits depuis Linne. Vaillant-Carmanne: Imprimerie H." title="Catalogue systematique des Cicindelidae, decrits depuis Linne" type="book" year="1892">Fleutiaux, 1892</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FB0878B6FAC4EBEF" author="Casey TL" box="[1166,1308,1248,1269]" journalOrPublisher="Annals of the New York Academy of Sciences" pageId="13" pageNumber="263" pagination="285 - 684" part="9" refId="ref25396" refString="Casey TL. 1897. Coleopterological notices VII. Annals of the New York Academy of Sciences 9: 285 - 684." title="Coleopterological notices VII" type="journal article" year="1897">Casey, 1897</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FAAA78B6FCB8EA0E" author="Rivalier E." journalOrPublisher="Revue Francaise d'Entomologie" pageId="13" pageNumber="263" pagination="249 - 268" part="21" refId="ref27888" refString="Rivalier E. 1954. Demembrement du genre Cicindela Linne. II. Faune americaine. Revue Francaise d'Entomologie 21: 249 - 268." title="Demembrement du genre Cicindela Linne. II. Faune americaine" type="journal article" year="1954">Rivalier, 1954</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="540B4B5EFF89F131FCEA78A8FC7BEA0E" author="Rivalier E." box="[876,931,1278,1300]" journalOrPublisher="Revue Francaise d'Entomologie" pageId="13" pageNumber="263" pagination="127 - 164" part="31" refId="ref27913" refString="Rivalier E. 1963. Demembrement du genre Cicindela L. (fin). V. Faune australienne (et liste recapitulative des genres et sous-genres proposes pour la faune mondiale). Revue Francaise d'Entomologie 31: 127 - 164." title="Demembrement du genre Cicindela L. (fin). V. Faune australienne (et liste recapitulative des genres et sous-genres proposes pour la faune mondiale)" type="journal article" year="1963">1963</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="540B4B5EFF89F131FC2978A9FC3EEA0E" author="Rivalier E." box="[943,998,1279,1300]" journalOrPublisher="Nouvelle revue d'Entomologie" pageId="13" pageNumber="263" pagination="135 - 143" part="1" refId="ref27956" refString="Rivalier E. 1971. Remarques sur la Tribu des Cicindelini (Col. Cicindelidae) et sa subdivision en sous-tribus. Nouvelle revue d'Entomologie 1: 135 - 143." title="Remarques sur la Tribu des Cicindelini (Col. Cicindelidae) et sa subdivision en sous-tribus" type="journal article" year="1971">1971</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FC7478A8FB4CEA0E" author="Johnson WN" box="[1010,1172,1278,1300]" journalOrPublisher="Cicindela" pageId="13" pageNumber="263" pagination="49 - 54" part="23" refId="ref26976" refString="Johnson WN. 1991. A new species of Dromochorus from southern Texas (Coleoptera: Cicindelidae). Cicindela 23: 49 - 54." title="A new species of Dromochorus from southern Texas (Coleoptera: Cicindelidae)" type="journal article" year="1991">Johnson, 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FB1978A8FAE6EA0E" author="Wiesner J." box="[1183,1342,1278,1300]" journalOrPublisher="Keltern: Erna Bauer" pageId="13" pageNumber="263" refId="ref28375" refString="Wiesner J. 1992. Verzeichnis der Sandlaufkafer der Welt (Coleoptera, Cicindelidae). Keltern: Erna Bauer." title="Verzeichnis der Sandlaufkafer der Welt (Coleoptera, Cicindelidae)" type="book" year="1992">Wiesner, 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FACC78A8FC39EA28" author="Erwin TL & Pearson DL" journalOrPublisher="Sofia-Moscow: Pensoft" pageId="13" pageNumber="263" refId="ref25900" refString="Erwin TL, Pearson DL. 2008. A treatise on the Western Hemisphere Caraboidea (Coleoptera). Their classification, distributions, and ways of life. Volume II (Carabidae- Nebriiformes 2 - Cicindelitae). Sofia-Moscow: Pensoft." title="A treatise on the Western Hemisphere Caraboidea (Coleoptera). Their classification, distributions, and ways of life. Volume II (Carabidae- Nebriiformes 2 - Cicindelitae)" type="book" year="2008">Erwin & Pearson, 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FC6B7948FB44EA28" author="Bousquet Y." box="[1005,1180,1309,1331]" journalOrPublisher="ZooKeys" pageId="13" pageNumber="263" pagination="1 - 1630" part="245" refId="ref25254" refString="Bousquet Y. 2012. Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico. ZooKeys 245: 1 - 1630." title="Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico" type="journal article" year="2012">Bousquet, 2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FB217948FA5BEA29" author="Pearson DL & Knisley CB & Duran DP & Kazilek CJ" box="[1191,1411,1309,1331]" journalOrPublisher="New York: Oxford University Press" pageId="13" pageNumber="263" refId="ref27666" refString="Pearson DL, Knisley CB, Duran DP, Kazilek CJ. 2015. A field guide to tiger beetles of the United States and Canada: identification, natural history, and distribution of the Cicindelinae. New York: Oxford University Press." title="A field guide to tiger beetles of the United States and Canada: identification, natural history, and distribution of the Cicindelinae" type="book" year="2015">
|
||
Pearson
|
||
<emphasis id="02EEEABDFF89F131FA8F7948FA98EA28" box="[1289,1344,1309,1331]" italics="true" pageId="13" pageNumber="263">et al.</emphasis>
|
||
, 2015
|
||
</bibRefCitation>
|
||
); or viewed to be a synonym of
|
||
<taxonomicName id="F79A4D2CFF89F131FBF9796AFB35EA4B" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[1151,1261,1340,1361]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF89F131FBF9796AFB35EA4B" box="[1151,1261,1340,1361]" italics="true" pageId="13" pageNumber="263">Cicindela</emphasis>
|
||
</taxonomicName>
|
||
(e.g.
|
||
<bibRefCitation id="540B4B5EFF89F131FAAC796AFCB9EA6A" author="LeConte JL" journalOrPublisher="Transactions of the American Entomological Society" pageId="13" pageNumber="263" pagination="157 - 162" part="5" refId="ref27170" refString="LeConte JL. 1875. Notes on Cicindelidae of the United States. Transactions of the American Entomological Society 5: 157 - 162." title="Notes on Cicindelidae of the United States" type="journal article" year="1875">LeConte, 1875</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FCE8790DFC35EA6A" author="Leng CW" box="[878,1005,1371,1392]" journalOrPublisher="Transactions of the American Entomological Society" pageId="13" pageNumber="263" pagination="93 - 186" part="28" refId="ref27195" refString="Leng CW. 1902. Revision of the Cicindelidae of boreal America. Transactions of the American Entomological Society 28: 93 - 186." title="Revision of the Cicindelidae of boreal America" type="journal article" year="1902">Leng, 1902</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FC7D790CFB54EA6A" author="Harris ED" box="[1019,1164,1370,1392]" journalOrPublisher="Yonkers: Truan Press" pageId="13" pageNumber="263" refId="ref26532" refString="Harris ED. 1911. List of North American Cicindelidae in the Harris Collection. Yonkers: Truan Press." title="List of North American Cicindelidae in the Harris Collection" type="book" year="1911">Harris, 1911</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FB1C790DFAC3EA6A" author="Horn W." box="[1178,1307,1370,1392]" editor="Wytsman P" journalOrPublisher="Genera insectorum." pageId="13" pageNumber="263" pagination="16 - 23" part="82" refId="ref26866" refString="Horn W. 1915. Cicindelinae. In: Wytsman P, ed. Genera insectorum. 82 c: 209 - 486, plates 16 - 23." title="Cicindelinae" type="journal article" year="1915">Horn, 1915</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FAAE790CFC73EA95" author="Harris ED & Leng C." journalOrPublisher="New York: The American Museum of Natural History Press" pageId="13" pageNumber="263" refId="ref26552" refString="Harris ED, Leng C. 1916. The Cicindelinae of North America as arranged by Dr. Walther Horn in Genera Insectorum. New York: The American Museum of Natural History Press." title="The Cicindelinae of North America as arranged by Dr. Walther Horn in Genera Insectorum" type="book" year="1916">Harris & Leng, 1916</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FC3C792FFB95EA95" author="Cazier MA" box="[954,1101,1401,1423]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="13" pageNumber="263" pagination="231 - 309" part="103" refId="ref25454" refString="Cazier MA. 1954. A review of the Mexican tiger beetles of the genus Cicindela (Coleoptera, Cicindelidae). Bulletin of the American Museum of Natural History 103: 231 - 309." title="A review of the Mexican tiger beetles of the genus Cicindela (Coleoptera, Cicindelidae)" type="journal article" year="1954">Cazier, 1954</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FBDC792FFB29EA95" author="Arnett RH Jr." box="[1114,1265,1401,1423]" journalOrPublisher="Washington DC: Catholic University of America Press" pageId="13" pageNumber="263" refId="ref25035" refString="Arnett RH Jr. 1963. The beetles of the United States (a manual for identification). Washington DC: Catholic University of America Press." title="The beetles of the United States (a manual for identification)" type="book" year="1963">Arnett, 1963</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FB78792FFA53EA95" author="Willis HL" box="[1278,1419,1401,1423]" journalOrPublisher="Journal of the Kansas Entomological Society" pageId="13" pageNumber="263" pagination="303 - 317" part="41" refId="ref28396" refString="Willis HL. 1968. Artificial key to the species of Cicindela o f N o r t h A m e r i c a n o r t h o f M e x i c o (C o l e o p t e r a: Cicindelidae). Journal of the Kansas Entomological Society 41: 303 - 317." title="Artificial key to the species of Cicindela o f N o r t h A m e r i c a n o r t h o f M e x i c o (C o l e o p t e r a: Cicindelidae)" type="journal article" year="1968">Willis, 1968</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FCAF79CEFB43EAB7" author="Bousquet Y & Larochelle A." box="[809,1179,1432,1454]" journalOrPublisher="Memoirs of the Entomological Society of Canada" pageId="13" pageNumber="263" pagination="1 - 197" part="67" refId="ref25281" refString="Bousquet Y, Larochelle A. 1993. Catalogue of the Geadephaga (Coleoptera: Trachypachidae, Rhysodidae, Carabidae including Cicindelini) of America north of Mexico. Memoirs of the Entomological Society of Canada 67: 1 - 197." title="Catalogue of the Geadephaga (Coleoptera: Trachypachidae, Rhysodidae, Carabidae including Cicindelini) of America north of Mexico" type="journal article" year="1993">Bousquet & Larochelle, 1993</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FB2A79CEFCB8EAD6" author="Arnett RH Jr & Thomas MC" journalOrPublisher="Boca Raton: CRC Press" pageId="13" pageNumber="263" refId="ref25063" refString="Arnett RH Jr, Thomas MC, eds. 2000. American beetles, Volume I: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. Boca Raton: CRC Press." title="American beetles, Volume I: Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia" type="book" year="2000">Arnett & Thomas, 2000</bibRefCitation>
|
||
), with some North American workers recognizing the taxon as a subgenus (
|
||
<bibRefCitation id="540B4B5EFF89F131FBD17983FB00EAF0" author="Boyd HP" box="[1111,1240,1493,1515]" journalOrPublisher="Marlton, NJ: Plexus" pageId="13" pageNumber="263" refId="ref25323" refString="Boyd HP. 1982. Checklist of Cicindelidae: the tiger beetles. Marlton, NJ: Plexus." title="Checklist of Cicindelidae: the tiger beetles" type="book" year="1982">Boyd, 1982</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FB607983FA5AEAF0" author="Freitag R." box="[1254,1410,1493,1515]" journalOrPublisher="Ottawa: NRC Research Press" pageId="13" pageNumber="263" refId="ref26021" refString="Freitag R. 1999. Catalogue of the tiger beetles of Canada and the United States. Ottawa: NRC Research Press." title="Catalogue of the tiger beetles of Canada and the United States" type="book" year="1999">Freitag, 1999</bibRefCitation>
|
||
). Despite the variety of treatments, justifications for different taxonomic ranks were almost never given.
|
||
</paragraph>
|
||
<paragraph id="302536AFFF89F132FCC77A64FD36EE03" blockId="13.[809,1427,727,1914]" lastBlockId="14.[163,779,198,282]" lastPageId="14" lastPageNumber="264" pageId="13" pageNumber="263">
|
||
Recent phylogenies have given support to many of the named clades within the
|
||
<emphasis id="02EEEABDFF89F131FBEA7A06FA94E97F" box="[1132,1356,1616,1637]" italics="true" pageId="13" pageNumber="263">
|
||
<taxonomicName id="F79A4D2CFF89F131FBEA7A06FB0EE97F" authority="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan, 2019" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[1132,1238,1616,1637]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus" sensu="lato">Cicindela</taxonomicName>
|
||
<taxonomicNameLabel id="19DD57C6FF89F131FB5B7A06FA94E97F" box="[1245,1356,1616,1637]" pageId="13" pageNumber="263" sensu="lato">sensu lato</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation id="540B4B5EFF89F131FADD7A06FC7BE99E" author="Pons J & Barraclough T & Theodorides K & Cardoso A & Vogler A." journalOrPublisher="Systematic Biology" pageId="13" pageNumber="263" pagination="554 - 570" part="53" refId="ref27711" refString="Pons J, Barraclough T, Theodorides K, Cardoso A, Vogler A. 2004. Using exon and intron sequences of the gene Mp 20 to resolve basal relationships in Cicindela (Coleoptera: Cicindelidae). Systematic Biology 53: 554 - 570." title="Using exon and intron sequences of the gene Mp 20 to resolve basal relationships in Cicindela (Coleoptera: Cicindelidae)" type="journal article" year="2004">
|
||
Pons
|
||
<emphasis id="02EEEABDFF89F131FCAF7A39FCB8E999" box="[809,864,1646,1668]" italics="true" pageId="13" pageNumber="263">et al.</emphasis>
|
||
, 2004
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="540B4B5EFF89F131FC297A38FBA0E99E" author="Vogler AP & Cardoso A & Barraclough TG" box="[943,1144,1646,1668]" journalOrPublisher="Systematic Biology" pageId="13" pageNumber="263" pagination="4 - 20" part="54" refId="ref28330" refString="Vogler AP, Cardoso A, Barraclough TG. 2005. Exploring rate variation among and within sites in a densely sampled tree: Species level phylogenetics of North American tiger beetles (genus Cicindela). Systematic Biology 54: 4 - 20." title="Exploring rate variation among and within sites in a densely sampled tree: Species level phylogenetics of North American tiger beetles (genus Cicindela)" type="journal article" year="2005">
|
||
Vogler
|
||
<emphasis id="02EEEABDFF89F131FC787A39FBF6E999" box="[1022,1070,1646,1668]" italics="true" pageId="13" pageNumber="263">et al</emphasis>
|
||
., 2005
|
||
</bibRefCitation>
|
||
; Gough
|
||
<emphasis id="02EEEABDFF89F131FB507A39FADFE999" box="[1238,1287,1646,1668]" italics="true" pageId="13" pageNumber="263">et al</emphasis>
|
||
., in review), and these generally correspond to groups recognized in Rivalier’s revision (1954, 1963, 1971), including the
|
||
<taxonomicName id="F79A4D2CFF89F131FCAF7A9CFC19E9C5" box="[809,961,1738,1759]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF89F131FCAF7A9CFC19E9C5" box="[809,961,1738,1759]" italics="true" pageId="13" pageNumber="263">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
. Based on the aforementioned research, and our own mtDNA tree (
|
||
<figureCitation id="A8A12A2AFF89F131FBE17ABFFB71E9E4" box="[1127,1193,1769,1791]" captionStart="Figure 2" captionStartId="6.[162,242,1619,1641]" captionTargetBox="[266,1340,196,1575]" captionTargetId="figure@6.[265,1340,195,1577]" captionTargetPageId="6" captionText="Figure 2A. Mitochondrial DNA (mtDNA) cytochrome oxidase b (Cytb) Bayesian genealogy of all members of the genus Dromochorus. Outgroups include North American representatives of Cylindera and Cicindelidia. On the left is a reduced version of the full tree indicating clades shown, in detail, in each figure. Colours are arbitrarily chosen to show distinctions among the taxa, and directly correspond to populations shown in Figure 3. Only posterior support values above 60 are displayed, and the scale bar at bottom shows a branch length representing nucleotide substitutions per site. Taxon names include the unique specimen number (see also Supporting Information, Table S1), along with the state and collection locality; genomic material from specimens marked with a ~ were also used in the multilocus analysis (Figure 4), and holotypes of species described in this work are marked with an ‘H’. The longitudinal bars on the right identify specimens and clades historically considered to be a single species." httpUri="https://zenodo.org/record/3089247/files/figure.png" pageId="13" pageNumber="263">Fig. 2</figureCitation>
|
||
),
|
||
<taxonomicName id="F79A4D2CFF89F131FB477ABFFA84E9E4" box="[1217,1372,1769,1790]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF89F131FB477ABFFA84E9E4" box="[1217,1372,1769,1790]" italics="true" pageId="13" pageNumber="263">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
was recovered as a well-supported monophyletic lineage, and sister to another clade of Nearctic tiger beetles. As such, we recognize the group as a distinct genus. Although
|
||
<taxonomicName id="F79A4D2CFF89F131FCAF7B32FC63E863" box="[809,955,1892,1913]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF89F131FCAF7B32FC63E863" box="[809,955,1892,1913]" italics="true" pageId="13" pageNumber="263">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
may appear nested within
|
||
<taxonomicName id="F79A4D2CFF89F131FB597B32FA93E863" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[1247,1355,1892,1913]" class="Insecta" family="Carabidae" genus="Cylindera" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="263" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF89F131FB597B32FA93E863" box="[1247,1355,1892,1913]" italics="true" pageId="13" pageNumber="263">Cylindera</emphasis>
|
||
</taxonomicName>
|
||
in our tree, this latter group is polyphyletic (
|
||
<bibRefCitation id="540B4B5EFF8AF132FDBC7C90FD25EFC6" author="Gough HM & Duran DP & Kawahara AY & Toussiant EF" box="[570,765,198,220]" journalOrPublisher="Systematic Entomology" pageId="14" pageNumber="264" refId="ref26379" refString="Gough HM, Duran DP, Kawahara AY, Toussiant EF. 2018. A comprehensive molecular phylogeny of tiger beetles (Coleoptera, Carabidae, Cicindelinae) challenges their current classification. Systematic Entomology (in press)." title="A comprehensive molecular phylogeny of tiger beetles (Coleoptera, Carabidae, Cicindelinae) challenges their current classification" type="book" year="2018">
|
||
Gough
|
||
<emphasis id="02EEEABDFF8AF132FD0F7C91FD6EEFC1" box="[649,694,198,220]" italics="true" pageId="14" pageNumber="264">et al</emphasis>
|
||
., 2018
|
||
</bibRefCitation>
|
||
), as previously believed by many workers, and the sister clade to
|
||
<taxonomicName id="F79A4D2CFF8AF132FF7B7D52FE57EE03" box="[253,399,260,281]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF8AF132FF7B7D52FE57EE03" box="[253,399,260,281]" italics="true" pageId="14" pageNumber="264">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
is being named as a new genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78806524FF8AF132FF257D17FD08EDFD" pageId="14" pageNumber="264" type="diagnosis">
|
||
<paragraph id="302536AFFF8AF132FF257D17FEFAEE42" blockId="14.[163,779,321,743]" box="[163,290,321,344]" pageId="14" pageNumber="264">
|
||
<heading id="6B6D81C3FF8AF132FF257D17FEFAEE42" box="[163,290,321,344]" fontSize="10" level="4" pageId="14" pageNumber="264" reason="1">
|
||
<emphasis id="02EEEABDFF8AF132FF257D17FEFAEE42" box="[163,290,321,344]" italics="true" pageId="14" pageNumber="264">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8AF132FF257D37FE5FED76" blockId="14.[163,779,321,743]" pageId="14" pageNumber="264">
|
||
<taxonomicName id="F79A4D2CFF8AF132FF257D37FE90EE6C" box="[163,328,353,374]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF8AF132FF257D37FE90EE6C" box="[163,328,353,374]" italics="true" pageId="14" pageNumber="264">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
are separable from all other North American tiger beetle genera by the following combination of characters, present in the adult stage. Beetles are flightless and lack flight wings, but the elytra are not fused. The elytra are oval shaped, and completely lacking pale maculations. The dorsum is dark, usually black or brown, but may also have a frosted blue, violet, grey or green sheen. The legs and tarsi are clothed in decumbent setae.
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8AF132FF3D7E23FD08EDFD" blockId="14.[163,779,321,743]" pageId="14" pageNumber="264">In subsequent species descriptions, the most salient or diagnostic characters are indicated in bold. Rarely are any of these characters diagnostic by themselves, but the combination of these characters may be.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78806524FF8AF132FF257F58FC4DEFE0" pageId="14" pageNumber="264" type="distribution">
|
||
<paragraph id="302536AFFF8AF132FF257F58FEE2EC3C" blockId="14.[163,780,782,1235]" box="[163,314,782,806]" pageId="14" pageNumber="264">
|
||
<heading id="6B6D81C3FF8AF132FF257F58FEE2EC3C" box="[163,314,782,806]" fontSize="10" level="4" pageId="14" pageNumber="264" reason="1">
|
||
<emphasis id="02EEEABDFF8AF132FF257F58FEE2EC3C" box="[163,314,782,806]" italics="true" pageId="14" pageNumber="264">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8AF132FF257F78FC4DEFE0" blockId="14.[163,780,782,1235]" lastBlockId="14.[827,1443,198,250]" pageId="14" pageNumber="264">
|
||
<taxonomicName id="F79A4D2CFF8AF132FF257F78FE9EEC59" box="[163,326,814,835]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF8AF132FF257F78FE9EEC59" box="[163,326,814,835]" italics="true" pageId="14" pageNumber="264">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
are geographically restricted to the south-central United States and adjacent Mexico (
|
||
<figureCitation id="A8A12A2AFF8AF132FF2D7F3AFF34EC9B" box="[171,236,876,898]" captionStart="Figure 3" captionStartId="8.[163,241,1807,1829]" captionTargetBox="[163,1443,581,1767]" captionTargetId="figure@8.[163,1443,581,1767]" captionTargetPageId="8" captionText="Figure 3. Distributionof sampled Dromochorus populations.Colours correspondto those used in Figure 1. Green = D.pilatei, pink = D. welderensis sp. nov., sky blue = D. velutinigrens, red = D. chaparralensis sp. nov., blue = D. pruininus, lime green = D. knisleyi sp. nov., orange = D. minimus sp. nov., black = D. belfragei." httpUri="https://zenodo.org/record/3089261/files/figure.png" pageId="14" pageNumber="264">Fig. 3</figureCitation>
|
||
). Texas is the centre of diversity, and all eight species are found there, at least in part of their ranges. Records from Mexico are few and imprecise. Historically,
|
||
<taxonomicName id="F79A4D2CFF8AF132FEB47F9EFE6CECC7" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[306,436,968,989]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="species" species="belfragei">
|
||
<emphasis id="02EEEABDFF8AF132FEB47F9EFE6CECC7" box="[306,436,968,989]" italics="true" pageId="14" pageNumber="264">D. belfragei</emphasis>
|
||
</taxonomicName>
|
||
was the only species recorded from Mexico, although it would appear as if these populations belong to
|
||
<taxonomicName id="F79A4D2CFF8AF132FEE07850FDF5EB00" authority="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan, 2019" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[358,557,1029,1051]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="species" species="chaparralensis" status="sp. nov.">
|
||
<emphasis id="02EEEABDFF8AF132FEE07850FDF5EB00" box="[358,557,1029,1051]" italics="true" pageId="14" pageNumber="264">D. chaparralensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19DD57C6FF8AF132FDB27853FD4AEB00" box="[564,658,1029,1050]" pageId="14" pageNumber="264" rank="species">
|
||
<emphasis id="02EEEABDFF8AF132FDB27853FD4AEB00" bold="true" box="[564,658,1029,1050]" pageId="14" pageNumber="264">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, described in this treatment. In general,
|
||
<taxonomicName id="F79A4D2CFF8AF132FE7A7872FD4DEB23" box="[508,661,1060,1081]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF8AF132FE7A7872FD4DEB23" box="[508,661,1060,1081]" italics="true" pageId="14" pageNumber="264">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
are found at low elevations; apparently, they do not occur in montane environments, such as the Ouachita Mountains of Arkansas and Oklahoma, which border a section of the eastern range of
|
||
<taxonomicName id="F79A4D2CFF8AF132FE1478C9FDFCEBAE" authorityName="Casey" authorityYear="1897" box="[402,548,1183,1204]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="species" species="pruininus">
|
||
<emphasis id="02EEEABDFF8AF132FE1478C9FDFCEBAE" box="[402,548,1183,1204]" italics="true" pageId="14" pageNumber="264">D. pruininus</emphasis>
|
||
</taxonomicName>
|
||
. Most
|
||
<taxonomicName id="F79A4D2CFF8AF132FDF478C8FCD3EBA9" box="[626,779,1182,1203]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF8AF132FDF478C8FCD3EBA9" box="[626,779,1182,1203]" italics="true" pageId="14" pageNumber="264">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
are found below 500 m, with only the westernmost populations of
|
||
<taxonomicName id="F79A4D2CFF8AF132FC597C91FB87EFC1" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[991,1119,198,220]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="species" species="belfragei">
|
||
<emphasis id="02EEEABDFF8AF132FC597C91FB87EFC1" box="[991,1119,198,220]" italics="true" pageId="14" pageNumber="264">D. belfragei</emphasis>
|
||
</taxonomicName>
|
||
occurring at elevations up to 1000 m.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="78806524FF8AF132FCBD7D74FAAAEBCB" pageId="14" pageNumber="264" type="biology_ecology">
|
||
<paragraph id="302536AFFF8AF132FCBD7D74FBB1EE20" blockId="14.[827,1443,290,1234]" box="[827,1129,290,314]" pageId="14" pageNumber="264">
|
||
<heading id="6B6D81C3FF8AF132FCBD7D74FBB1EE20" box="[827,1129,290,314]" fontSize="10" level="4" pageId="14" pageNumber="264" reason="1">
|
||
<emphasis id="02EEEABDFF8AF132FCBD7D74FBB1EE20" box="[827,1129,290,314]" italics="true" pageId="14" pageNumber="264">Ecology/natural history</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8AF132FCBD7D14FBE7EDD2" blockId="14.[827,1443,290,1234]" pageId="14" pageNumber="264">
|
||
<taxonomicName id="F79A4D2CFF8AF132FCBD7D14FC0BEE4D" box="[827,979,322,343]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF8AF132FCBD7D14FC0BEE4D" box="[827,979,322,343]" italics="true" pageId="14" pageNumber="264">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
appear to have a 2-year life cycle, based on observations of
|
||
<taxonomicName id="F79A4D2CFF8AF132FB957D34FB7DEE6D" authorityName="Casey" authorityYear="1897" box="[1043,1189,354,375]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="species" species="pruininus">
|
||
<emphasis id="02EEEABDFF8AF132FB957D34FB7DEE6D" box="[1043,1189,354,375]" italics="true" pageId="14" pageNumber="264">D. pruininus</emphasis>
|
||
</taxonomicName>
|
||
(Herrmann & Duran, unpublished). Adult beetles are terrestrial predators of small invertebrates, and are generally believed to be crepuscular or active in the late afternoon, but we have found them at all hours of the day in more shaded microhabitats or when significant cloud cover is present.
|
||
<taxonomicName id="F79A4D2CFF8AF132FCFB7E4FFBC5ED34" box="[893,1053,537,558]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF8AF132FCFB7E4FFBC5ED34" box="[893,1053,537,558]" italics="true" pageId="14" pageNumber="264">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
are extremely fast runners and may evade capture by darting into dense grasses, or in some species, hiding in cracks in the earth, especially
|
||
<taxonomicName id="F79A4D2CFF8AF132FCF97E20FBD9ED90" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[895,1025,629,651]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="species" species="belfragei">
|
||
<emphasis id="02EEEABDFF8AF132FCF97E20FBD9ED90" box="[895,1025,629,651]" italics="true" pageId="14" pageNumber="264">D. belfragei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F79A4D2CFF8AF132FBBD7E20FB13ED91" authorityName="Casey" authorityYear="1897" box="[1083,1227,630,651]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="species" species="pruininus">
|
||
<emphasis id="02EEEABDFF8AF132FBBD7E20FB13ED91" box="[1083,1227,630,651]" italics="true" pageId="14" pageNumber="264">D. pruininus</emphasis>
|
||
</taxonomicName>
|
||
. Larvae are poorly known, and were only recently described (
|
||
<bibRefCitation id="540B4B5EFF8AF132FAC57EC2FBF7EDDD" author="Spomer SM & Nabity PD & Brust ML" journalOrPublisher="The Coleopterist's Bulletin" pageId="14" pageNumber="264" pagination="37 - 41" part="62" refId="ref28197" refString="Spomer SM, Nabity PD, Brust ML. 2008. Larval description of Cicindela (Dromochorus) pruinina (Casey) (Coleoptera: Carabidae: Cicindelinae) with notes on habitat and adult behavior. The Coleopterist's Bulletin 62: 37 - 41." title="Larval description of Cicindela (Dromochorus) pruinina (Casey) (Coleoptera: Carabidae: Cicindelinae) with notes on habitat and adult behavior" type="journal article" year="2008">Spomer, Nabity & Brust, 2008</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8AF132FCD57E87FAAAEBCB" blockId="14.[827,1443,290,1234]" pageId="14" pageNumber="264">
|
||
There are remarkably few specimens of these beetles in major museum collections, even though several species occur near major cities and universities in Texas and Oklahoma. Many species records are based on one or a few specimens, and as such,
|
||
<taxonomicName id="F79A4D2CFF8AF132FB437F1AFABAEC7B" box="[1221,1378,844,865]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF8AF132FB437F1AFABAEC7B" box="[1221,1378,844,865]" italics="true" pageId="14" pageNumber="264">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
were reported to be rare, or at low densities. This is likely a consequence of their atypical natural history compared to other diurnal North American tiger beetles, and the fact that their habitats are not as commonly visited by collectors. We have found that
|
||
<taxonomicName id="F79A4D2CFF8AF132FB3C7FB3FA8AECE0" box="[1210,1362,997,1018]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF8AF132FB3C7FB3FA8AECE0" box="[1210,1362,997,1018]" italics="true" pageId="14" pageNumber="264">Dromochorus</emphasis>
|
||
</taxonomicName>
|
||
can be remarkably abundant in the appropriate habitat during the ideal time of year, rivalling or exceeding densities observed in some common riparian tiger beetle species. These observations are unlikely to represent unusual population explosions, as we have witnessed similarly large numbers of beetles every year in areas that we visited each year between 2012 and 2016.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="64E56627FF8AF132FF257B13FB38E862" httpUri="https://zenodo.org/record/3089275/files/figure.png" pageId="14" pageNumber="264" startId="14.[163,243,1861,1883]" targetBox="[166,1441,1284,1809]" targetPageId="14">
|
||
<paragraph id="302536AFFF8AF132FF257B13FB38E862" blockId="14.[163,1443,1861,1912]" pageId="14" pageNumber="264">
|
||
<emphasis id="02EEEABDFF8AF132FF257B13FED5E840" bold="true" box="[163,269,1861,1883]" pageId="14" pageNumber="264">Figure 7.</emphasis>
|
||
Labial palps of: A,
|
||
<taxonomicName id="F79A4D2CFF8AF132FE667B10FDBEE841" authorityName="Casey" authorityYear="1897" box="[480,614,1862,1883]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="species" species="pruininus">
|
||
<emphasis id="02EEEABDFF8AF132FE667B10FDBEE841" box="[480,614,1862,1883]" italics="true" pageId="14" pageNumber="264">D. pruininus</emphasis>
|
||
</taxonomicName>
|
||
; B,
|
||
<taxonomicName id="F79A4D2CFF8AF132FD097B10FCD1E840" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[655,777,1861,1883]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="species" species="belfragei">
|
||
<emphasis id="02EEEABDFF8AF132FD097B10FCD1E840" box="[655,777,1861,1883]" italics="true" pageId="14" pageNumber="264">D. belfragei</emphasis>
|
||
</taxonomicName>
|
||
; C,
|
||
<emphasis id="02EEEABDFF8AF132FCB47B13FBD6E840" bold="true" box="[818,1038,1861,1883]" pageId="14" pageNumber="264">
|
||
<taxonomicName id="F79A4D2CFF8AF132FCB47B13FC76E841" authority="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan, 2019" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[818,942,1861,1883]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="264" phylum="Arthropoda" rank="species" species="knisleyi" status="sp. nov.">
|
||
<emphasis id="02EEEABDFF8AF132FCB47B13FC76E841" bold="true" box="[818,942,1861,1883]" italics="true" pageId="14" pageNumber="264">D. knisleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19DD57C6FF8AF132FC337B13FBD6E840" box="[949,1038,1861,1882]" pageId="14" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A and B illustrate labial palps with a contrasting darker apical segment, and C illustrates labial palps that are consistently dark throughout.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="78806524FF8BF133FDCA7C88FAA5EBEC" pageId="15" pageNumber="265" type="key">
|
||
<paragraph id="302536AFFF8BF133FDCA7C88FC0FEFEB" blockId="15.[167,1405,222,1271]" box="[588,983,222,243]" pageId="15" pageNumber="265">
|
||
<heading id="6B6D81C3FF8BF133FDCA7C88FC0FEFEB" allCaps="true" box="[588,983,222,243]" fontSize="8" level="5" pageId="15" pageNumber="265" reason="3">
|
||
KEY TO THE GENUS
|
||
<taxonomicName id="F79A4D2CFF8BF133FCAC7C88FC0FEFEB" ID-CoL="8KW25" box="[810,983,222,243]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="265" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="02EEEABDFF8BF133FCAC7C88FC0FEFEB" box="[810,983,222,243]" italics="true" pageId="15" pageNumber="265">DROMOCHORUS</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF217D5AFAA4EE5A" blockId="15.[167,1405,222,1271]" pageId="15" pageNumber="265">
|
||
1a. Labial palps at least partly yellow to dark amber, with contrasting darker apical segment (
|
||
<figureCitation id="A8A12A2AFF8BF133FF6B7D7CFE85EE5A" box="[237,349,298,320]" captionStart="Figure 7" captionStartId="14.[163,243,1861,1883]" captionTargetBox="[166,1441,1284,1809]" captionTargetId="figure@14.[163,1443,1268,1820]" captionTargetPageId="14" captionText="Figure 7. Labial palps of: A, D. pruininus; B, D. belfragei; C, D. knisleyi sp. nov. A and B illustrate labial palps with a contrasting darker apical segment, and C illustrates labial palps that are consistently dark throughout." httpUri="https://zenodo.org/record/3089275/files/figure.png" pageId="15" pageNumber="265">Fig. 7A, B</figureCitation>
|
||
) ......................................................................................................................................................2
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF217D1FFAA4EE44" blockId="15.[167,1405,222,1271]" box="[167,1404,329,351]" pageId="15" pageNumber="265">
|
||
1b. Labial palps consistently dark brown to black throughout (
|
||
<figureCitation id="A8A12A2AFF8BF133FC047D1FFC0BEE44" box="[898,979,329,351]" captionStart="Figure 7" captionStartId="14.[163,243,1861,1883]" captionTargetBox="[166,1441,1284,1809]" captionTargetId="figure@14.[163,1443,1268,1820]" captionTargetPageId="14" captionText="Figure 7. Labial palps of: A, D. pruininus; B, D. belfragei; C, D. knisleyi sp. nov. A and B illustrate labial palps with a contrasting darker apical segment, and C illustrates labial palps that are consistently dark throughout." httpUri="https://zenodo.org/record/3089275/files/figure.png" pageId="15" pageNumber="265">Fig. 7C</figureCitation>
|
||
) ..........................................................4
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF217D3EFAA4EEA1" blockId="15.[167,1405,222,1271]" pageId="15" pageNumber="265">
|
||
2a. Elytral surface smooth and finely frosted in texture, without any pitting or subsutural foveae. Dorsum with strong blue to violet reflections throughout. Labial palps yellow, with contrasting darker apical seg- ment. Kansas and western Missouri, south to Central Texas ..................................................
|
||
<taxonomicName id="F79A4D2CFF8BF133FB6A7DF0FAA4EEA1" ID-CoL="6DKDD" authorityName="Casey" authorityYear="1897" box="[1260,1404,422,443]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="265" phylum="Arthropoda" rank="species" species="pruininus">
|
||
<emphasis id="02EEEABDFF8BF133FB6A7DF0FAA4EEA1" box="[1260,1404,422,443]" italics="true" pageId="15" pageNumber="265">D. pruininus</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF217D92FAA4EEED" blockId="15.[167,1405,222,1271]" pageId="15" pageNumber="265">
|
||
2b. Elytra surface dull, textured with fine to deep pitting throughout. Distinct shallow pits running along elytra suture (subsutural foveae,
|
||
<figureCitation id="A8A12A2AFF8BF133FDE67DB4FD69EEED" box="[608,689,482,504]" captionStart="Figure 8" captionStartId="16.[163,241,1096,1118]" captionTargetBox="[330,1279,195,1055]" captionTargetId="figure@16.[323,1283,195,1055]" captionTargetPageId="16" captionText="Figure 8. A, Elytra showing distinct punctures (foveae) running parallel to the suture of the elytra (D. knisleyi sp. nov.). B, lack of distinct subsutural foveae (D. welderensis sp. nov.)." httpUri="https://zenodo.org/record/3089283/files/figure.png" pageId="15" pageNumber="265">Fig. 8A</figureCitation>
|
||
) may be present ..........................................................................3
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF217E57FAA4ED68" blockId="15.[167,1405,222,1271]" pageId="15" pageNumber="265">
|
||
3a. Dorsum dark brown with prominent shallow subsutural foveae and irregular pitting throughout ely- tral surface. Foveae and smaller pits with metallic green reflections. Irregular green marbling may be present on elytra, head and pronotum. Labial palps yellow, with contrasting darker apical segment. Louisiana to East Texas....................................................................................................................
|
||
<taxonomicName id="F79A4D2CFF8BF133FA927E08FAA4ED68" ID-CoL="6DKDB" box="[1300,1404,605,627]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="265" phylum="Arthropoda" rank="species" species="pilatei">
|
||
<emphasis id="02EEEABDFF8BF133FA927E08FAA4ED68" box="[1300,1404,605,627]" italics="true" pageId="15" pageNumber="265">D. pilatei</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF217E2AFAA4EDF7" blockId="15.[167,1405,222,1271]" pageId="15" pageNumber="265">
|
||
3b. Dorsum black with shallow to deep pitting on elytral surface. Some metallic blue reflections may be pre- sent, especially on the supraorbital region of the head and humeral area of the elytra. Subsutural foveae may be present. Labial palps yellow to dark amber, with apical segment darkest. Oklahoma and Texas panhandle, south to East Central Texas......................................................................................
|
||
<taxonomicName id="F79A4D2CFF8BF133FB7C7E8EFAA4EDF7" ID-CoL="37QDY" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[1274,1404,728,749]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="265" phylum="Arthropoda" rank="species" species="belfragei">
|
||
<emphasis id="02EEEABDFF8BF133FB7C7E8EFAA4EDF7" box="[1274,1404,728,749]" italics="true" pageId="15" pageNumber="265">D. belfragei</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF217EA0FAA5EC30" blockId="15.[167,1405,222,1271]" pageId="15" pageNumber="265">
|
||
4a. Elytral surface rough, with shallow to deep pitting. Subsutural foveae present (
|
||
<figureCitation id="A8A12A2AFF8BF133FB107EA0FB34EC11" box="[1174,1260,758,780]" captionStart="Figure 8" captionStartId="16.[163,241,1096,1118]" captionTargetBox="[330,1279,195,1055]" captionTargetId="figure@16.[323,1283,195,1055]" captionTargetPageId="16" captionText="Figure 8. A, Elytra showing distinct punctures (foveae) running parallel to the suture of the elytra (D. knisleyi sp. nov.). B, lack of distinct subsutural foveae (D. welderensis sp. nov.)." httpUri="https://zenodo.org/record/3089283/files/figure.png" pageId="15" pageNumber="265">Fig. 8A</figureCitation>
|
||
), often with metallic green or blue reflections in pits. ‘Hill Country’ region of Central Texas...........
|
||
<taxonomicName id="F79A4D2CFF8BF133FB2B7F40FAFBEC30" ID-CoL="8LVBX" authority="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan, 2019" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[1197,1315,789,811]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="265" phylum="Arthropoda" rank="species" species="knisleyi" status="sp. nov.">
|
||
<emphasis id="02EEEABDFF8BF133FB2B7F40FAFBEC30" box="[1197,1315,789,811]" italics="true" pageId="15" pageNumber="265">D. knisleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19DD57C6FF8BF133FAAC7F43FAA5EC30" box="[1322,1405,789,810]" pageId="15" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF217F62FAA4EC72" blockId="15.[167,1405,222,1271]" pageId="15" pageNumber="265">
|
||
4b. Elytral surface smooth, often with velvety or frosted texture. No subsutural foveae present (
|
||
<figureCitation id="A8A12A2AFF8BF133FF6B7F04FEE6EC7D" box="[237,318,850,872]" captionStart="Figure 8" captionStartId="16.[163,241,1096,1118]" captionTargetBox="[330,1279,195,1055]" captionTargetId="figure@16.[323,1283,195,1055]" captionTargetPageId="16" captionText="Figure 8. A, Elytra showing distinct punctures (foveae) running parallel to the suture of the elytra (D. knisleyi sp. nov.). B, lack of distinct subsutural foveae (D. welderensis sp. nov.)." httpUri="https://zenodo.org/record/3089283/files/figure.png" pageId="15" pageNumber="265">Fig. 8B</figureCitation>
|
||
) ..........................................................................................................................................................5
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF217F27FAA4ECBC" blockId="15.[167,1405,222,1271]" pageId="15" pageNumber="265">5a. Pronotum glabrous or with few scattered long thin erect setae irregularly placed, rarely concentrated along margins.................................................................................................................................................6</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF217FF8FAA4ECDE" blockId="15.[167,1405,222,1271]" box="[167,1404,942,964]" pageId="15" pageNumber="265">5b. Pronotum with sparse to regular white decumbent setae, mostly in lateral third ....................................7</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF217F9BFAA3EB05" blockId="15.[167,1405,222,1271]" pageId="15" pageNumber="265">
|
||
6a. Dorsum finely velvety black, with strong violet, blue or green reflections, especially along margins. Male labrum entirely dark or very nearly so. Body form gracile. South Texas, coastal areas south of Corpus Christi and inland to vicinity of Dimmit County .................................................................
|
||
<taxonomicName id="F79A4D2CFF8BF133FB4E785DFAA3EB05" ID-CoL="6DKDF" authorityName="Johnson" authorityYear="1992" box="[1224,1403,1034,1056]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="265" phylum="Arthropoda" rank="species" species="velutinigrens">
|
||
<emphasis id="02EEEABDFF8BF133FB4E785DFAA3EB05" box="[1224,1403,1034,1056]" italics="true" pageId="15" pageNumber="265">D. velutinigrens</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF21787FFAA4EB47" blockId="15.[167,1405,222,1271]" pageId="15" pageNumber="265">
|
||
6b. Elytra dark ash-grey, sometimes with frosty blue reflections. South Texas in mesquite chaparral forest. Known only from Bexar, Frio and Atascosa Counties ....................................................
|
||
<taxonomicName id="F79A4D2CFF8BF133FB19781EFAFBEB47" ID-CoL="8LVBY" authority="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan, 2019" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[1183,1315,1096,1117]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="265" phylum="Arthropoda" rank="species" species="minimus" status="sp. nov.">
|
||
<emphasis id="02EEEABDFF8BF133FB19781EFAFBEB47" box="[1183,1315,1096,1117]" italics="true" pageId="15" pageNumber="265">D. minimus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19DD57C6FF8BF133FAAC781EFAA4EB47" box="[1322,1404,1096,1117]" pageId="15" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF217831FAA5EBA3" blockId="15.[167,1405,222,1271]" pageId="15" pageNumber="265">
|
||
7a. Elytra dull black, may have bluish reflections especially near margins. South Texas to Mexico, in mes- quite chaparral. Known from Dimmit, LaSalle, and Webb Counties in Texas, and the state of Tamaulipas, Mexico.....................................................................................................................
|
||
<taxonomicName id="F79A4D2CFF8BF133FBDA78F2FAFBEBA2" ID-CoL="8LVBW" authority="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan, 2019" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[1116,1315,1187,1209]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="265" phylum="Arthropoda" rank="species" species="chaparralensis" status="sp. nov.">
|
||
<emphasis id="02EEEABDFF8BF133FBDA78F2FAFBEBA2" box="[1116,1315,1187,1209]" italics="true" pageId="15" pageNumber="265">D. chaparralensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19DD57C6FF8BF133FAAC78F2FAA5EBA3" box="[1322,1405,1188,1209]" pageId="15" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="302536AFFF8BF133FF217894FAA5EBEC" blockId="15.[167,1405,222,1271]" pageId="15" pageNumber="265">
|
||
7b. Elytra finely velvety black, may have a faint dark blue sheen. Found in coastal prairie habitat near Gulf of Mexico ......................................................................................................................
|
||
<taxonomicName id="F79A4D2CFF8BF133FB0778B4FAFBEBEC" ID-CoL="8LVBZ" authority="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan, 2019" authorityName="Duran & Herrmann & Roman & Gwiazdowski & Drummond & Hood & Egan" authorityYear="2019" box="[1153,1315,1249,1271]" class="Insecta" family="Carabidae" genus="Dromochorus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="265" phylum="Arthropoda" rank="species" species="welderensis" status="sp. nov.">
|
||
<emphasis id="02EEEABDFF8BF133FB0778B4FAFBEBEC" box="[1153,1315,1249,1271]" italics="true" pageId="15" pageNumber="265">D. welderensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="19DD57C6FF8BF133FAAC78B7FAA5EBEC" box="[1322,1405,1249,1270]" pageId="15" pageNumber="265" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |