356 lines
47 KiB
XML
356 lines
47 KiB
XML
<document id="AC30AFCE35A5696E909E434FA2418041" ENCODING="UTF-8" ID-GBIF-Dataset="d1cd8ac5-9818-47fb-a2ad-bee8b13694a1" ModsDocID="z01297p001" checkinTime="1247147669357" checkinUser="thomas" docAuthor="Lucila C. Protogino, Amalia M. Miquelarena & Hugo L. López" docDate="2006" docId="B833298EA9604FB6D606A2D66B4D6D55" docLanguage="en" docName="2006_Protogino_Miquelarena_Lopez_gg1.xml" docOrigin="Zootaxa 1297" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:8B4411D9-087E-4182-81EF-23D764E0FE27" docTitle="Astyanax aramburui Protogino, Miquelarena & López, 2006, new species" docType="treatment" docVersion="11" lastPageNumber="12" masterDocId="FFDA194F4ED8F70E1084B316265B7B54" masterDocTitle="A new species of Astyanax (Teleostei, Characiformes, Characidae), with breeding tubercles, from the Paraná and Uruguay river basins." masterLastPageNumber="16" masterPageNumber="1" pageNumber="2" updateTime="1701314452692" updateUser="plazi">
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<mods:title id="CB25163A31B7AD221EE65CD9935DC8E7">A new species of Astyanax (Teleostei, Characiformes, Characidae), with breeding tubercles, from the Paraná and Uruguay river basins.</mods:title>
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<mods:namePart id="18E487DDECCEB75EC8E31B4AE9749AD5">Lucila C. Protogino</mods:namePart>
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<mods:namePart id="3C7F9E6F4ABD841FD5EFF8E97DCD4987">Amalia M. Miquelarena</mods:namePart>
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<mods:roleTerm id="8DF6AD06879A0FFC6AC10A3D860F79C1">Author</mods:roleTerm>
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<mods:namePart id="2F0C0D7372724C66A0D36C1635A982FA">Hugo L. López</mods:namePart>
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<mods:date id="8369B0D7A042F0769FA3FDC4698718C8">2006</mods:date>
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<mods:classification id="A1D2E63B645571B105B14BB5EA5B9C1C">journal article</mods:classification>
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<treatment id="B833298EA9604FB6D606A2D66B4D6D55" ID-DOI="http://doi.org/10.5281/zenodo.6258680" ID-GBIF-Taxon="114923930" ID-Zenodo-Dep="6258680" LSID="urn:lsid:plazi:treatment:B833298EA9604FB6D606A2D66B4D6D55" httpUri="http://treatment.plazi.org/id/B833298EA9604FB6D606A2D66B4D6D55" lastPageNumber="12" pageNumber="2">
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<subSubSection id="60A507873B4E5789C50967580A6E9B48" type="nomenclature">
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<paragraph id="6A3FE6C7646A666BD85959218C3EE762" pageNumber="2">
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<taxonomicName id="DCB31A3FEBD5F6174AB2C0ED060CFC0A" ID-CoL="J5Q5" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="2" phylum="Chordata" rank="species" species="aramburui">Astyanax aramburui</taxonomicName>
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<taxonomicNameLabel id="FA5EAF3300A701FCB07739E349E081B3" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="ADE21D30D39F1AB9BE9E3830DCA09113" type="description">
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<paragraph id="C296158345FC8D59D4F79B9801DF6DA8" pageNumber="2">Fig. 1</paragraph>
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</subSubSection>
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<subSubSection id="68E85B2E3E771759ECD657188E45217A" type="nomenclature">
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<paragraph id="C797CC69756AE06CCD2A767F71A4D327" pageNumber="2">
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<materialsCitation id="9BC85C95B76EBAE214F0F87119C87850" ID-GBIF-Occurrence="1226907498" country="Argentina" stateProvince="Entre Rios Province">
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<typeStatus id="ED74CC3FB078E6E2D5E989D2DACCE576">Holotype</typeStatus>
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.
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<collectionCode id="FEF8417DF90CECE38A1ABC0630CF70D5" collectionName="ILPLA">ILPLA</collectionCode>
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1712, male, 61.1 mm SL; Villaguay Creek at Balneario (
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<geoCoordinate id="567C8CBF2DAF13D311679D1CB058EB6D" direction="south" orientation="latitude" value="-31.916666">31º55’S</geoCoordinate>
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<geoCoordinate id="50A5045F511FF6C104B8F73F967E36E6" direction="west" orientation="longitude" value="-59.05">59º03’W</geoCoordinate>
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), Gualeguay River basin,
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<collectingRegion id="BFED7C9863258DD6268C5D7ECA94A5F7">
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Entre
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<normalizedToken id="9AD906A1C909B307A4EE0B1729AEEF2A" originalValue="Ríos">Rios</normalizedToken>
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Province
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</collectingRegion>
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,
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<collectingCountry id="5BBA785B8E55A96065CA37E0E14D7D5D">Argentina</collectingCountry>
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; coll.: A. Miquelarena, et al.,
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<date id="541C6FF191CD8BF867B74BDA6649E268" month="11" value="2004-11" year="2004">November 2004</date>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="225E4E016AD5ACAF6D01C8498E507D72" pageNumber="3">
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<materialsCitation id="9F494A754BE190B8CE317E0DF123C417" ID-GBIF-Occurrence="1226907490" country="Argentina" stateProvince="Entre Rios Province">
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<typeStatus id="D5C867228AB90A81AF0DCA60CAE48BD0">Paratypes</typeStatus>
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.
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<collectionCode id="B252D37087ECC95BF1B7AF6A28FBA2A7" collectionName="ILPLA">ILPLA</collectionCode>
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569, (13), 7 males, 6 females, 59.2-78.9 mm SL; Brazo Chico Creek, 10 km from Uruguay River (
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<geoCoordinate id="8A678B0FB402F2E787FF8A2D42991838" direction="south" orientation="latitude" value="-33.75">33°45’S</geoCoordinate>
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<geoCoordinate id="5492DEA9B6FA1AE4B71B9F2FC58FF817" direction="west" orientation="longitude" value="-58.533333">58°32’W</geoCoordinate>
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),
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<collectingRegion id="116B5E83CAB3C8CAE5DBCDDE6F931AD8">
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Entre
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<normalizedToken id="DF41A1787722DF0BB266594BA8AB3C30" originalValue="Ríos">Rios</normalizedToken>
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Province
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</collectingRegion>
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,
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<collectingCountry id="AC45E5EDD457D778A5182B4AD1047F9F">Argentina</collectingCountry>
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, coll: N. Landoni,
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<date id="A890A487E1B25BF3E74F01327E197B55" month="01" value="1985-01" year="1985">January 1985</date>
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</materialsCitation>
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;
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<materialsCitation id="30F64258EF7F003E9C74EE6E338F4A5F" ID-GBIF-Occurrence="1226907500" country="Argentina" stateProvince="Entre Rios Province">
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<collectionCode id="5D247BBD1DACFF4526F8259EB3BCD1F9" collectionName="ILPLA">ILPLA</collectionCode>
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1709, (4) 2 males, 2 females (c&s), 60.2-80.9 mm SL; same locality as previous specimen
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</materialsCitation>
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;
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<materialsCitation id="C34AD22AB6892ED85F05073861259E19" ID-GBIF-Occurrence="1226907496" country="Argentina" stateProvince="Entre Rios Province">
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<collectionCode id="AA512EBDADFBC4BF18E993E9DB7D68ED" collectionName="ILPLA">ILPLA</collectionCode>
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1710, (7) males, 55.0-60.6 mm SL
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</materialsCitation>
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;
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<materialsCitation id="71C393A67D90C3DE63BC74422421B8EC" ID-GBIF-Occurrence="1226907486" country="Argentina" stateProvince="Entre Rios Province">
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<collectionCode id="0467396982963E71F53592C60EFC6275" collectionName="ILPLA">ILPLA</collectionCode>
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1711, (5) 4 males, 1 juv. (c&s), 41.0-74.2 mm SL
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</materialsCitation>
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;
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<materialsCitation id="6A92B4E8431EDA5877B3853F04EFFA0C" ID-GBIF-Occurrence="1226907489" country="Argentina" stateProvince="Entre Rios Province">
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<collectionCode id="D129034E3EF6447AAF7C42705C9D8425" collectionName="ILPLA">ILPLA</collectionCode>
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1713, (4) males, 57.7-66.0 mm SL
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</materialsCitation>
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,
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<materialsCitation id="9A5C98C231968039AC30E115CEB7EF80" ID-GBIF-Occurrence="1226907485" country="Argentina" stateProvince="Entre Rios Province">
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<collectionCode id="4D11CEF9EA013FFD7B705D3FCAB52721" collectionName="Argentina, La Plata, Universidad Nacional de La Plata, Museo de la Plata">MLP</collectionCode>
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9672, male, 55.0 mm SL, collected with holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="E2D3244AC7C89ABCA9E6B8CD85BF58F7" type="diagnosis">
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<paragraph id="AF4F461E12F65CF5AE0F13AF35A9E111" lastPageNumber="4" pageNumber="3">
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Diagnosis:
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<taxonomicName id="EB715294A72F98DE88A6FFB6688F4225" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="3" phylum="Chordata" rank="species" species="aramburui">Astyanax aramburui</taxonomicName>
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is distinguished from all congeners by the following unique combination of characters: round or trapezoidal black humeral spot, dorsal to the lateral line; a branch of the infraorbital sensory canal directed ventrally, continued as a series of pores totally or partially traversing the third infraorbital; body relatively slender (31.8-37.4 % SL); eye large (41.1-45.8 % HL); snout short (15.5-20.9 % HL); caudal peduncle relatively slender (10.3-12.0 % SL); one pentacuspid maxillary tooth; 38-42 perforated scales in lateral line; and v -vi, 23-29 anal-fin rays. In addition, the species is characterized by the presence, in males, of breeding tubercles on the head and scales, and bony hooks on all fins.
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</paragraph>
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</subSubSection>
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<subSubSection id="69F05165555FD5B2286B0F4C5A337419" type="description">
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<paragraph id="45E15FF55A21DAA8A4500B3BFD1DA53A" pageNumber="4">Description: Morphometric data appear in table 1. Body elongate, with maximum body depth at dorsal-fin origin (31.8-37.4% SL). Dorsal body profile convex, slightly concave between eye and supraoccipital process, relatively straight from that point to dorsal-fin origin and descending slightly to adipose-fin origin. Dorsal and ventral margins of caudal peduncle slightly concave. Caudal peduncle relatively slender (10.3-12.0% SL). Ventral body profile convex between tip of snout and pelvic-fin origin, nearly straight between that point and anal-fin origin. Body compressed posteriorly from anal-fin origin. Head short (22.6-25.2% SL). Snout short (15.5-20.9% HL). Eye large (41.1-45.8% HL). Interorbital less than eye diameter (30.3-35.3% HL). Mouth terminal or slightly superior, lower jaw included. Maxilla relatively long, extending slightly beyond anterior edge of orbit.</paragraph>
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<paragraph id="14F5ED7F5720E19CAD0D5B71967781AD" pageNumber="4">Dorsal-fin origin almost equidistant between tip of snout and base of caudal-fin. Tip of pectoral fin surpassing pelvic-fin origin in both males and females. Tip of pelvic fin almost never reaching anal-fin origin in most females (except in one specimen). Tip of pelvic fin of males either falling short of, reaching or surpassing anal-fin origin. Bony hooks present on rays of all fins in matures males. Hooks mostly present on posterior branches of rays. Usually one pair of bony hooks per ray segment.</paragraph>
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<paragraph id="70A382B6AD9F0981CE1767CC3B866F19" pageNumber="4">Dorsal-fin rays ii,9 (6*); iii,9 (4); posterior margin of dorsal fin typically straight, the last unbranched ray and first two branched rays longest. Bony hooks of males small, scattered on branches of first to eighth fin rays.</paragraph>
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<paragraph id="5733FDEECB26818D1EFFF9DEE1C655A1" pageNumber="4">Pectoral-fin rays i,12 (3); i,13 (6); i,14 (1*). Bony hooks of large specimens small and scarce, 4-6 hooks on a few branched rays.</paragraph>
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<paragraph id="25AB6CCB395426374633E8C5F2AECF19" pageNumber="5">Pelvic-fin rays i,7 (10*). Bony hooks numerous, more developed on first to fourth branched rays. Origin of pelvic fin anterior to dorsal-fin origin. Pelvic axillary scale without hooks. Posterior margin of pectoral and pelvic fins rounded.</paragraph>
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<paragraph id="2BA5EC7381CD88EFEE2297F99416D584" pageNumber="5">Anal-fin rays v,23 (1); v,24 (1); vi,25 (1*); v,25 (3); v,26 (2); v,27 (1); vi,29 (1). Bony hooks conspicuous, curved, distributed from last unbranched ray to, usually, 15th branched ray. Minute hooks distributed along distal third of some remaining rays. Origin of anal fin anterior to posteriormost dorsal fin ray insertion. Posterior margin of anal fin nearly straight in males, and with anterior one-third slightly falcate in females. Caudal-fin forked, with lobes of similar size. Principal caudal-fin rays i,17,i. Dorsal procurrent rays 11 (1) or 12 (6). Ventral procurrent rays 8 (1), 10 (5), 11(1). Caudal fin with only few bony hooks on principal rays.</paragraph>
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<paragraph id="1A043942375A9B970DC61223163F49D6" pageNumber="5">Cycloid scales large, deeply striated, regularly distributed on body. Lateral line complete, perforated scales 38 (1), 39 (5), 40 (13), 41 (1*), 42 (1). Scale rows between dorsal-fin origin and lateral line 7 (19*); 8 (2). Scale rows between lateral line and anal-fin origin 6 (5), 7 (16*). Predorsal scales 11 (6), 12 (13*), 13 (2), usually in a regular series. Single row of scales at base of anal fin, 12 (1), 14 (4), 15 (6), 16 (3*), 17 (2), 18 (2), 19 (1), 20 (1), 22 (1). Preventral scales large. Pelvic-fin axillary scale with evident grooves. Males with small regularly-arranged tubercles on posterior edge of scales (see remarks below).</paragraph>
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<paragraph id="CD90D704B51E78D0A590C8FF3885EBF7" pageNumber="5">Premaxilla with short ascending process and relatively elongated lateral process, with two series of teeth (Fig. 2a -b). Outer row with 4 to 6 teeth, somewhat irregularly arranged, with 3 to 5 (typically 3) cusps. Inner row with 5 broader teeth, a symphysial tooth, narrowest and highest, with 4 or 5 (typically 5) cusps; the second and third with 6 or 7 cusps; the fourth with 4 to 7 (typically 6) cusps. Fifth tooth smallest and situated internally with respect to fourth, with 4 to 6 cusps (single specimen with only 3). Maxilla (Fig. 2c) with one pentacuspid tooth on upper third (one tricuspid tooth in only two individuals). Dentary with 3 or 4 large teeth with 5-7 cusps, followed by a middle-sized tooth with 4 or 5 cusps, and 3-6 smaller teeth, monocuspid or tricuspid (Fig. 2d).</paragraph>
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<paragraph id="3675FAB464FB01F7D1BB433335AC9D09" pageNumber="5">Vertebrae 36 (6), 37 (2). Supraneurals 5 or 6, typically 5. Infraorbitals 6. A branch of infraorbital sensory canal directed ventrally, continued as a series of pores totally or partially traversing third infraorbital (Fig. 3). Upper gill rakers 9(1), 10(2), 11(3), 12 (2); lower gill rakers 13 (1), 14 (1), 15 (4), 16 (2).</paragraph>
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<paragraph id="5DA49EBB9E134BF652FE4139C8D904FB" pageNumber="5">Color in alcohol: Background light yellowish. Dorsum of head and body darker. Bones in infraorbital series and operculum silvery. Small dark humeral spot, round or trapezoidal, often with diffuse margins, centered on third and fourth scales of scale row just dorsal to lateral line. Lateral body stripe broad and dark, extending from behind humeral spot, and ending as a black rhombic caudal spot on middle caudal-fin rays. Lateral body stripe silvery in some specimens. Ventrolateral region of body uniformly colored. Dorsal, anal and caudal fins with scattered dark chromatophores. Pectoral and pelvic fins hyaline.</paragraph>
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<paragraph id="C91D6AB96C7213CDEEA822439C2FF453" pageNumber="7">Color in life: Body silvery white, iridescent with yellowish hues. Dorsum of head and body with dark chromatophores. Lateral surface of head silvery. Lateral stripe silvery, terminating in black caudal spot continued on to middle caudal-fin rays. Dorsal and anal fins covered with scattered dark chromatophores. Pectoral fin hyaline, except for dark first unbranched ray. Pelvic fin hyaline. Caudal fin grayish, with black distal margin (Fig. 4).</paragraph>
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</subSubSection>
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<subSubSection id="C01A57CBD2DAB3EB471D922AD2061B47" type="etymology">
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<paragraph id="301C3A5A57B0533E0DB4C95899D3EF13" pageNumber="7">
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Etymology: Named after Prof.
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<normalizedToken id="33DF9A6DEE154714AF00E7C301ABD698" originalValue="Raúl">Raul</normalizedToken>
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H.
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<normalizedToken id="7D3183808C1AE0B4DA7533DD756FF738" originalValue="Arámburu">Aramburu</normalizedToken>
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(1924-2004), researcher and professor of the Museo de La Plata, Buenos Aires, founder of the first chair of Ichthyology in Argentina.
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</paragraph>
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</subSubSection>
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<subSubSection id="22AD34DAFF1D1FC3867846959C983B01" type="distribution">
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<paragraph id="925BDB6CCF4C026B51D16D5587AA6910" pageNumber="7">
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Distribution and habitat:
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<taxonomicName id="AA31BE3443456C44637CCB045E4C6941" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="7" phylum="Chordata" rank="species" species="aramburui">Astyanax aramburui</taxonomicName>
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is known from Villaguay Creek and Brazo Chico Creek, which flow into the
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<normalizedToken id="377E8621D4DFDD0629E5010FFEFF0C1D" originalValue="Paraná">Parana</normalizedToken>
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and Uruguay Rivers, respectively, in Entre
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<normalizedToken id="5E2E8A7EA8196FF79A856975417CF012" originalValue="Ríos">Rios</normalizedToken>
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Province, Argentina (Fig. 5).
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</paragraph>
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</subSubSection>
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<subSubSection id="A9034ADDD526DD23D2E32AD6CCBB5CE7" type="biology_ecology">
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<paragraph id="C39D7F8D96A375593BF7AF85B1922E7C" pageNumber="7">Villaguay creek is a relatively extensive watercourse, with sections about 50 m wide and pools over 2 m deep. The banks are vegetated with grasses and other plants, and the bottom consists of mud and clay (Fig. 6). Brazo Chico creek is a watercourse situated in the deltaic area known as Islas del Ibicuy.</paragraph>
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</subSubSection>
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<subSubSection id="1F7B6988ACF151E890F548A43B0327CB" type="discussion">
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<paragraph id="04CF4AF787AC311E6A7B0639A52D4728" lastPageNumber="10" pageNumber="8">
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Remarks: Of all the families of Neotropical Characiformes, only the Characidae, Parodontidae and Lebiasinidae are known to develop true nuptial tubercles (Wiley & Collette, 1970:164-167; Collette, 1977: 236-241). Those authors state that breeding tubercles are probably present in all species of Parodontidae and at least five species of Lebiasinidae. Within the Characidae, similar structures have been reported in
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<taxonomicName id="D04131260652B8972F1F0D53ABD6F4D0" family="Characidae" genus="Bryconamericus" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="8" phylum="Chordata" rank="species" species="emperador">Bryconamericus emperador (Eigenmann & Ogle)</taxonomicName>
|
||
, by Meek & Hildebrand (1916:284); for an unidentified
|
||
<taxonomicName id="5E90790BE0FE163E8A67F52D21A924B0" LSID-ZBK="urn:lsid:zoobank.org:act:A694B2D7-A3FB-417F-A952-1A84FFB5C7B2" family="Characidae" genus="Bryconamericus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Bryconamericus Eigenmann 1907:139" order="Characiformes" pageNumber="8" phylum="Chordata" rank="genus">Bryconamericus</taxonomicName>
|
||
species from Panama by Fink (1976:342); in two species of
|
||
<taxonomicName id="E71BB7490A5B3923CB22F452010AC479" LSID-ZBK="urn:lsid:zoobank.org:act:871425B1-C450-4BB3-BF06-5F3ACA538A84" family="Characidae" genus="Monotocheirodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Monotocheirodon Eigenmann & Pearson 1924:34" order="Characiformes" pageNumber="8" phylum="Chordata" rank="genus">Monotocheirodon</taxonomicName>
|
||
by Collette (1977:238); and in two Brazilian species,
|
||
<taxonomicName id="C649ECBB9BA51B08CA20E6CCA79A0EF3" LSID-ZBK="urn:lsid:zoobank.org:act:687C0FBB-5487-488E-B2E3-1652A3464838" family="Characidae" genus="Myxiops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Myxiops aphos Zanata & Akama 2004:47" order="Characiformes" pageNumber="8" phylum="Chordata" rank="species" species="aphos">Myxiops aphos Zanata & Akama</taxonomicName>
|
||
, and an undescribed species of
|
||
<taxonomicName id="F8DD30821BE2D3BD3A5C76522D539D95" LSID-ZBK="urn:lsid:zoobank.org:act:4DB39814-5BBA-4D7B-8C5D-F830AB26824D" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax Baird & Girard 1854:26" order="Characiformes" pageNumber="8" phylum="Chordata" rank="genus">Astyanax</taxonomicName>
|
||
by Zanata & Akama (2004:50-51). We were able to observe small tubercles on the dorsal part of the head, as well as on the snout, infraorbital, and opercular region in males of
|
||
<taxonomicName id="4922389533326F368378ED7D08545164" LSID-ZBK="urn:lsid:zoobank.org:act:E5E50087-8076-460E-AE11-71F2D06827ED" family="Parodontidae" genus="Parodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Parodon carrikeri Fowler 1940:47" order="Characiformes" pageNumber="8" phylum="Chordata" rank="species" species="carrikeri">Parodon carrikeri Fowler</taxonomicName>
|
||
. These structures are irregularly distributed on the scales, particularly in the predorsal area (Figs. 7a -d). In mature males of
|
||
<taxonomicName id="6F0A69D71B46BF6B87AC7EFD6E456C12" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="8" phylum="Chordata" rank="species" species="aramburui">A. aramburui</taxonomicName>
|
||
, the tubercles are arranged as in
|
||
<taxonomicName id="A42358B25DFEA2FCF61109D724697261" LSID-ZBK="urn:lsid:zoobank.org:act:E5E50087-8076-460E-AE11-71F2D06827ED" family="Parodontidae" genus="Parodon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Parodon carrikeri Fowler 1940:47" order="Characiformes" pageNumber="8" phylum="Chordata" rank="species" species="carrikeri">P. carrikeri</taxonomicName>
|
||
in the cephalic region, but they are distributed regularly on the posterior edge of the scales over most of the body (Figs. 8a -d). Mature male specimens of
|
||
<taxonomicName id="6C78E331B32A79950A53A3A4AF50B0CF" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="8" phylum="Chordata" rank="species" species="aeneus">
|
||
Astyanax aeneus (
|
||
<normalizedToken id="95554DA70E6ABF1CDAC471D3132D7B80" originalValue="Günther">Guenther</normalizedToken>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="AAEF0ACF2E3C069C95D69AF495CC826E" LSID-ZBK="urn:lsid:zoobank.org:act:9EA48613-2CCE-422A-93F1-B0466DB136A3" family="Characidae" genus="Bryconamericus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Bryconamericus thomasi Fowler 1940:49" order="Characiformes" pageNumber="8" phylum="Chordata" rank="species" species="thomasi">Bryconamericus thomasi Fowler</taxonomicName>
|
||
also show breeding tubercles arranged as in
|
||
<taxonomicName id="65EA18793D22CDF84FFB58FE3336D2E2" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="8" phylum="Chordata" rank="species" species="aramburui">A. aramburui</taxonomicName>
|
||
. Histological examination confirmed that they are of epitelial origin, and formed by cells distributed in a concentric manner. In all the above species the breeding tubercles were more noticeable in the specimens collected during October, November and March. Zanata & Akama (2004), in their discussion of characters that distinguish the genus
|
||
<taxonomicName id="D1CD19B855F7E36742E075A6B5F15F27" LSID-ZBK="urn:lsid:zoobank.org:act:87E87117-8304-4D4E-B12E-C79CB5EACC7E" family="Characidae" genus="Myxiops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Myxiops Zanata & Akama 2004:46" order="Characiformes" pageNumber="8" phylum="Chordata" rank="genus">Myxiops</taxonomicName>
|
||
, state that these structures are located along the posterior border of the scales, thus delineating the scale outline; this agrees with our observations of
|
||
<taxonomicName id="2D56978FFC5F55A59F19B9AAA3E32589" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="8" phylum="Chordata" rank="species" species="aeneus">A. aeneus</taxonomicName>
|
||
,
|
||
<taxonomicName id="C5AA112DDA6327F95125183270DF4F97" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="8" phylum="Chordata" rank="species" species="aramburui">A. aramburui</taxonomicName>
|
||
and
|
||
<taxonomicName id="4D78C4A2542942F75ADC9F22A7D5E2A5" LSID-ZBK="urn:lsid:zoobank.org:act:9EA48613-2CCE-422A-93F1-B0466DB136A3" family="Characidae" genus="Bryconamericus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Bryconamericus thomasi Fowler 1940:49" order="Characiformes" pageNumber="8" phylum="Chordata" rank="species" species="thomasi">B. thomasi</taxonomicName>
|
||
. However, in contrast with the condition in the above-mentioned species, the accumulation of epithelial cells of
|
||
<taxonomicName id="70F4591EC2D85B02A2417581FB9FDA96" LSID-ZBK="urn:lsid:zoobank.org:act:87E87117-8304-4D4E-B12E-C79CB5EACC7E" family="Characidae" genus="Myxiops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Myxiops Zanata & Akama 2004:46" order="Characiformes" pageNumber="8" phylum="Chordata" rank="genus">Myxiops</taxonomicName>
|
||
is more conspicuous in juveniles, and occurs both in males and females (Zanata & Akama, 2004).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A7BAD3B26799C59C56B970D9707BC7E8" type="discussion">
|
||
<paragraph id="85977F9667C0766F4D9F0A9F0F234A06" pageNumber="10">Discussion</paragraph>
|
||
<paragraph id="0B32B2F463199D27F746BE9FD28DFED9" pageNumber="10">
|
||
Nineteen species of the genus
|
||
<taxonomicName id="B79C993275589CFFCD561B6CC14BC0A6" LSID-ZBK="urn:lsid:zoobank.org:act:4DB39814-5BBA-4D7B-8C5D-F830AB26824D" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax Baird & Girard 1854:26" order="Characiformes" pageNumber="10" phylum="Chordata" rank="genus">Astyanax</taxonomicName>
|
||
have thus far been reported from Argentina (Miquelarena & Menni, 2005). Although
|
||
<taxonomicName id="5BC46BC7B3AC57255B950DD263914B25" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="10" phylum="Chordata" rank="species" species="fasciatus">A. fasciatus Cuvier</taxonomicName>
|
||
is indicated, in some references, to be widely distributed in Argentina, Melo & Buckup (2006) concluded that this species is restricted to the
|
||
<normalizedToken id="F0CBC290777600D3440E304F06E3A0E5" originalValue="São">Sao</normalizedToken>
|
||
Francisco River drainage, in Brazil.
|
||
<taxonomicName id="F245E4A70728FE31DF9C278F805DC9EF" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="10" phylum="Chordata" rank="species" species="aramburui">A. aramburui</taxonomicName>
|
||
resembles
|
||
<taxonomicName id="FD5042C36E437AFF5BE154E95EABC4B1" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="10" phylum="Chordata" rank="species" species="fasciatus">A. fasciatus</taxonomicName>
|
||
in possessing a single humeral spot, a relatively high number of anal-fin rays, and one tooth on the anterior margin of the maxilla. It differs from the
|
||
<normalizedToken id="B59D841B957DB9FF779367E5BC98C227" originalValue="São">Sao</normalizedToken>
|
||
Francisco populations by usually having 5 (vs. 4) teeth in the outer row of the premaxilla [
|
||
<taxonomicName id="3156CEC6EC0654203E3C97C32551064E" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="10" phylum="Chordata" rank="species" species="aramburui">A. aramburui</taxonomicName>
|
||
4(3), 5(17), 6(1);
|
||
<taxonomicName id="6CE934AA32F7E07564568E063B3C05D8" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="10" phylum="Chordata" rank="species" species="fasciatus">A. fasciatus</taxonomicName>
|
||
3(1), 4(38), 5(4), 6(1)]. Moreover,
|
||
<taxonomicName id="87B36DE93BE0BDEA24C64B05127C9A22" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="10" phylum="Chordata" rank="species" species="fasciatus">A. fasciatus</taxonomicName>
|
||
is diagnosed by the presence of an elongated dorsal fin in mature males (Melo & Buckup, 2006), a character not shared by
|
||
<taxonomicName id="CA9B49CDAB7EF7BCC0C98D7C70186478" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="10" phylum="Chordata" rank="species" species="aramburui">A. aramburui</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B9827F4711917DBC033986265EC5B98C" pageNumber="10">
|
||
<taxonomicName id="B0A0C5E0E6D9294D683AE057CA53AE42" LSID-ZBK="urn:lsid:zoobank.org:act:2F695358-258B-4815-B9B8-C106DE72B515" family="Characidae" genus="Tetragonopterus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Tetragonopterus rutilus Jenyns 1842:125" order="Characiformes" pageNumber="10" phylum="Chordata" rank="species" species="rutilus">Tetragonopterus rutilus Jenyns</taxonomicName>
|
||
, originally described from the
|
||
<normalizedToken id="CEC7603B42364831CA4C689D12C11EB3" originalValue="Paraná">Parana</normalizedToken>
|
||
River, was considered by Melo & Buckup (2006:49, 50; table2) as a probable valid species of
|
||
<taxonomicName id="2D7F811455EFA60AB5C1DA61C7EFC880" LSID-ZBK="urn:lsid:zoobank.org:act:4DB39814-5BBA-4D7B-8C5D-F830AB26824D" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax Baird & Girard 1854:26" order="Characiformes" pageNumber="10" phylum="Chordata" rank="genus">Astyanax</taxonomicName>
|
||
. An analysis of morphometric and meristic data of the
|
||
<taxonomicName id="A81233C806134DB3C3929D92FEFDD15A" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="10" phylum="Chordata" rank="species" species="rutilus">A. rutilus</taxonomicName>
|
||
holotype provided by those authors shows that
|
||
<taxonomicName id="B279B160290C7B4A9CA37D4FF24652DF" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="10" phylum="Chordata" rank="species" species="aramburui">A. aramburui</taxonomicName>
|
||
(21 specimens analyzed unless otherwise indicated) differs from
|
||
<taxonomicName id="01234B18434B780D865DA096DD04C4AF" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="10" phylum="Chordata" rank="species" species="rutilus">A. rutilus</taxonomicName>
|
||
in several characters: lesser body depth (31.8-37.4 [mean=33.9] vs. 40% SL), shorter preanal distance (54.5-61.8 [mean=59.6] vs. 66.4% SL), shorter snout length (15.5-20.9 [mean=17.8] vs. 23.0% HL), lower vertebrae count (36-37 [mean=36.2] [8 specimens] vs. 37), lesser number of scale rows between lateral line and dorsal-fin origin (7-8 [mean=7.1] vs. 8) and lesser number of scale rows between lateral line and anal-fin origin (6-7 [mean=6.8] vs. 8). In addition, according to the original description, the fins of
|
||
<taxonomicName id="02B05376CF1F58EF99B64DF643925DC6" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="10" phylum="Chordata" rank="species" species="rutilus">A. rutilus</taxonomicName>
|
||
are “dirty orange or bright red in color” (vs. paired fins hyaline and unpaired fins grayish).
|
||
</paragraph>
|
||
<paragraph id="6C86AEE161AAD8FB059C434C195DC622" pageNumber="11">
|
||
<taxonomicName id="5B48F4B43DFEC875FDBFAA79756CB1A1" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="aramburui">A. aramburui</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="C83B49E22761C9C036C3F423C6933D31" LSID-ZBK="urn:lsid:zoobank.org:act:B24C46C3-E368-49E4-B408-4876F70121D1" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax biotae Castro & Vari 2004:331" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="biotae">A. biotae Castro & Vari</taxonomicName>
|
||
, described from the upper Rio
|
||
<normalizedToken id="54855E1383AA550810D7217CB84D2736" originalValue="Paraná">Parana</normalizedToken>
|
||
system in southeastern Brazil, by having a greater number of scales in the lateral line (38-42 vs. 32-35), a greater number of scales in transverse series from origin of anal fin to lateral line (6-7 vs. 4-5), and a rounded humeral spot (vs. vertically elongated spot).
|
||
</paragraph>
|
||
<paragraph id="AB55344D1ECF3BB3D1AA460439313C3F" pageNumber="11">
|
||
<taxonomicName id="FBF0A01C208C24918ECC4A5A541390F8" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="aramburui">A. aramburui</taxonomicName>
|
||
exhibits bony hooks on all fins of mature males. This dimorphic character has also been reported in several species that have been recently described from the Paranoplatensean basin:
|
||
<taxonomicName id="4C9DA03F053AB9D4A96D3905CCC66263" LSID-ZBK="urn:lsid:zoobank.org:act:277C9FEE-EFEC-4675-8415-DC69F6205FE7" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax chico Casciotta & Almirón 2004:12" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="chico">
|
||
A. chico Casciotta &
|
||
<normalizedToken id="A77AFFDAAABC81CBFC839E50B181B557" originalValue="Almirón">Almiron</normalizedToken>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="6233E72DC29B9B5E1E52C41951E90E3D" LSID-ZBK="urn:lsid:zoobank.org:act:15E68D20-6FA1-43D1-A098-7843F661D1CD" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax hermosus Miquelarena, Protogino & López 2005:14" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="hermosus">
|
||
A. hermosus Miquelarena, Protogino &
|
||
<normalizedToken id="019C44CB3A40D755565AD2B54CC0DA15" originalValue="López">Lopez</normalizedToken>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="7A31385A6A56E75A28A91FC6A53761C3" LSID-ZBK="urn:lsid:zoobank.org:act:AE44CFDC-63B3-46DF-A875-9C84D52198F0" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax ojiara Azpelicueta & Garcia 2000:246" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="ojiara">
|
||
A. ojiara Azpelicueta &
|
||
<normalizedToken id="CE79E7D21E4E9BC5978C342910E56094" originalValue="García">Garcia</normalizedToken>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C4C6846C505692D7CAE51D4EF5B9717" LSID-ZBK="urn:lsid:zoobank.org:act:96A907F5-8583-4DFB-8209-1B4852986965" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax pynandi Casciotta, Almirón, Bechara, Roux & Ruiz Diaz 2003:808" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="pynandi">
|
||
A. pynandi Casciotta,
|
||
<normalizedToken id="EA01A32A5B91AEB69A8F82C99C6DF32B" originalValue="Almirón">Almiron</normalizedToken>
|
||
, Bechara, Roux & Ruiz Diaz
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="84796122E333762D476A7A000E0A265A" LSID-ZBK="urn:lsid:zoobank.org:act:290BA648-48A7-485B-AFDF-9A33A7AE0F21" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax troya Azpelicueta, Casciotta & Almirón 2002:249" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="troya">
|
||
A. troya Azpelicueta, Casciotta &
|
||
<normalizedToken id="FF14E3557368A6DED278CF5EEEC7C377" originalValue="Almirón">Almiron</normalizedToken>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="8689B4413923E43404995FACA6073C7A" LSID-ZBK="urn:lsid:zoobank.org:act:E5863476-57AD-4DFB-9869-C74D9004AFD1" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax leonidas Azpelicueta, Casciotta & Almirón 2002:245" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="leonidas">
|
||
A. leonidas Azpelicueta, Casciotta &
|
||
<normalizedToken id="77A09913F428911E5D5FD991FECC59AC" originalValue="Almirón">Almiron</normalizedToken>
|
||
</taxonomicName>
|
||
(without bony hooks on dorsal fin); and
|
||
<taxonomicName id="AAD1CC8AC38DB130ED85C0945E260307" LSID-ZBK="urn:lsid:zoobank.org:act:E47C3BA8-9E72-4C96-B1B3-9167C828A693" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax tumbayaensis Miquelarena & Menni 2005" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="tumbayaensis">A. tumbayaensis Miquelarena & Menni</taxonomicName>
|
||
(without bony hooks on pectoral fin). The new species differs from
|
||
<taxonomicName id="25C8EC51525FA094EFBE6F1EBA4B74E9" LSID-ZBK="urn:lsid:zoobank.org:act:277C9FEE-EFEC-4675-8415-DC69F6205FE7" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax chico Casciotta & Almirón 2004:12" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="chico">A.chico</taxonomicName>
|
||
,
|
||
<taxonomicName id="73CA41EBB567DBB480F0E081C34F3902" LSID-ZBK="urn:lsid:zoobank.org:act:E5863476-57AD-4DFB-9869-C74D9004AFD1" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax leonidas Azpelicueta, Casciotta & Almirón 2002:245" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="leonidas">A. leonidas</taxonomicName>
|
||
,
|
||
<taxonomicName id="260350198B174FAE18D34C05EE847C89" LSID-ZBK="urn:lsid:zoobank.org:act:AE44CFDC-63B3-46DF-A875-9C84D52198F0" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax ojiara Azpelicueta & Garcia 2000:246" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="ojiara">A. ojiara</taxonomicName>
|
||
,
|
||
<taxonomicName id="3CB1CDC92E9FEC8CA3EFD9F6557B9CB3" LSID-ZBK="urn:lsid:zoobank.org:act:96A907F5-8583-4DFB-8209-1B4852986965" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax pynandi Casciotta, Almirón, Bechara, Roux & Ruiz Diaz 2003:808" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="pynandi">A. pynandi</taxonomicName>
|
||
,
|
||
<taxonomicName id="7A3D77880F17723D6005FBB999AC0274" LSID-ZBK="urn:lsid:zoobank.org:act:290BA648-48A7-485B-AFDF-9A33A7AE0F21" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax troya Azpelicueta, Casciotta & Almirón 2002:249" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="troya">A. troya</taxonomicName>
|
||
, and
|
||
<taxonomicName id="E12C60757BE3B151DBD32FE43DBDA67A" LSID-ZBK="urn:lsid:zoobank.org:act:E47C3BA8-9E72-4C96-B1B3-9167C828A693" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax tumbayaensis Miquelarena & Menni 2005" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="tumbayaensis">A. tumbayaensis</taxonomicName>
|
||
by having a coloration pattern that includes a single humeral spot (vs. two humeral spots).
|
||
<taxonomicName id="DDFDABBE7724B883BB443B572571D675" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="aramburui">A. aramburui</taxonomicName>
|
||
shares with
|
||
<taxonomicName id="DDEAD79B5F61A7F62102205F2D3D149D" LSID-ZBK="urn:lsid:zoobank.org:act:15E68D20-6FA1-43D1-A098-7843F661D1CD" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax hermosus Miquelarena, Protogino & López 2005:14" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="hermosus">A. hermosus</taxonomicName>
|
||
the presence of a single humeral spot, which, however, differs in shape (rounded vs. Y-shaped).
|
||
</paragraph>
|
||
<paragraph id="538402D081442348DE6A5CD70A7A2A84" pageNumber="11">
|
||
Other characters that distinguish
|
||
<taxonomicName id="6A45D5B5E2879B5DCAE3384EE541C820" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="aramburui">A. aramburui</taxonomicName>
|
||
from
|
||
<taxonomicName id="E9EFF7BCD6FFE8EEE39F2B395CE94822" LSID-ZBK="urn:lsid:zoobank.org:act:15E68D20-6FA1-43D1-A098-7843F661D1CD" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax hermosus Miquelarena, Protogino & López 2005:14" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="hermosus">A.hermosus</taxonomicName>
|
||
,
|
||
<taxonomicName id="0AE63A7F30CAF0E814B9A81BE8297291" LSID-ZBK="urn:lsid:zoobank.org:act:AE44CFDC-63B3-46DF-A875-9C84D52198F0" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax ojiara Azpelicueta & Garcia 2000:246" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="ojiara">A. ojiara</taxonomicName>
|
||
,
|
||
<taxonomicName id="3C5AE3561CA44B7ECCBACFBEC1DD56A4" LSID-ZBK="urn:lsid:zoobank.org:act:96A907F5-8583-4DFB-8209-1B4852986965" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax pynandi Casciotta, Almirón, Bechara, Roux & Ruiz Diaz 2003:808" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="pynandi">A. pynandi</taxonomicName>
|
||
, and
|
||
<taxonomicName id="1D65BC66FB62270DBB86D967D37294E0" LSID-ZBK="urn:lsid:zoobank.org:act:290BA648-48A7-485B-AFDF-9A33A7AE0F21" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax troya Azpelicueta, Casciotta & Almirón 2002:249" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="troya">A. troya</taxonomicName>
|
||
are: greater number of scales in lateral line series (38-42 vs. 35-38; 36-38; 34-37 and 34-37, respectively); greater number of branched rays in anal fin (23-29 vs. 17-22; 20-23; 21-26 and 18-21, respectively), and shorter snout length (15.5-20.9% HL vs. 18.3-26.6; 24.5-30.9; 22.7-26.9 and 22.9-28.9% HL, respectively). It also differs from
|
||
<taxonomicName id="2A01D640C988DFBD5DA338154DF4B285" LSID-ZBK="urn:lsid:zoobank.org:act:15E68D20-6FA1-43D1-A098-7843F661D1CD" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax hermosus Miquelarena, Protogino & López 2005:14" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="hermosus">A. hermosus</taxonomicName>
|
||
by the greater eye diameter (41.1-45.8% HL vs. 29.7-34.7% HL) and lesser depth of the caudal peduncle (10.3-12.0% SL vs. 12.6-14.7% SL).
|
||
</paragraph>
|
||
<paragraph id="DCEB22FBE61004E634F0C5BDE2BE4F2C" pageNumber="11">
|
||
<taxonomicName id="476353D84948C53DCFB67D90CCD055BD" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="aramburui">A. aramburui</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="5FD6A2657CDFFAC2FF218E421BCD8198" LSID-ZBK="urn:lsid:zoobank.org:act:AE44CFDC-63B3-46DF-A875-9C84D52198F0" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax ojiara Azpelicueta & Garcia 2000:246" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="ojiara">A. ojiara</taxonomicName>
|
||
in having a greater eye diameter (41.1-45.8% HL vs. 29.1-37.2% HL) and by the presence of one pentacuspid (vs. heptacuspid) maxillary tooth; from
|
||
<taxonomicName id="88656E2BE60D4F71DE21CD54F194D10F" LSID-ZBK="urn:lsid:zoobank.org:act:96A907F5-8583-4DFB-8209-1B4852986965" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax pynandi Casciotta, Almirón, Bechara, Roux & Ruiz Diaz 2003:808" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="pynandi">A. pynandi</taxonomicName>
|
||
by a lesser body depth (31.8-37.4% SL vs. 35.4-42.9% SL); from
|
||
<taxonomicName id="0E59D7D532E633BB89A8D467A386BE1C" LSID-ZBK="urn:lsid:zoobank.org:act:E47C3BA8-9E72-4C96-B1B3-9167C828A693" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax tumbayaensis Miquelarena & Menni 2005" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="tumbayaensis">A. tumbayaensis</taxonomicName>
|
||
by having a greater eye diameter (41.1-45.8% HL vs. 28.9-35.0% HL), lesser body depth (31.8-37.4% SL vs. 39.2-45.3% SL) and lesser depth of the caudal peduncle (10.3-12.0% SL vs. 13.6-15.0% SL); and from
|
||
<taxonomicName id="A371C2E9FBCC9FBDAB4E518DF069565C" LSID-ZBK="urn:lsid:zoobank.org:act:277C9FEE-EFEC-4675-8415-DC69F6205FE7" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax chico Casciotta & Almirón 2004:12" order="Characiformes" pageNumber="11" phylum="Chordata" rank="species" species="chico">A. chico</taxonomicName>
|
||
by a greater number of branched anal-fin-rays (23-29 vs. 19-24).
|
||
</paragraph>
|
||
<paragraph id="121EE6F5EA0AD844375D539EBBF48B55" pageNumber="12">
|
||
Males of
|
||
<taxonomicName id="AD9AD953F48AB088FF3C9B9C00FF53F5" LSID-ZBK="urn:lsid:zoobank.org:act:E1334B2A-947E-4C19-BA1F-7745DCDAC7AA" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Astyanax elachylepis Bertaco & Lucinda 2005" order="Characiformes" pageNumber="12" phylum="Chordata" rank="species" species="elachylepis">A. elachylepis Bertaco & Lucinda</taxonomicName>
|
||
, described from the Tocantins River drainage, also have bony hooks on the dorsal, anal, pectoral, and pelvic-fin rays, but that species is easily distinguished from
|
||
<taxonomicName id="3EEA385C83BE01BCF9423C08EB4E785D" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="12" phylum="Chordata" rank="species" species="aramburui">A. aramburui</taxonomicName>
|
||
by the greater number of perforated scales in the lateral line (48-53 vs. 38-42).
|
||
</paragraph>
|
||
<paragraph id="6FBE61318D9C514F5EE80F4E4A0F20ED" pageNumber="12">
|
||
<taxonomicName id="F43C58FDFACADF020265CE28CBF18195" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="12" phylum="Chordata" rank="species" species="aeneus">A. aeneus</taxonomicName>
|
||
, from Arroyo Las Flores,
|
||
<normalizedToken id="F4A8D2C1E5EA93B67CAACF0BB5E03A6A" originalValue="México">Mexico</normalizedToken>
|
||
, shares with
|
||
<taxonomicName id="77F27A9E86088A0514C2B798056A467D" family="Characidae" genus="Astyanax" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageNumber="12" phylum="Chordata" rank="species" species="aramburui">A aramburui</taxonomicName>
|
||
the presence of breeding tubercles and bony hooks on all fins of mature males, but differs from the latter species by the greater preanal distance (62.1-64.8 vs. 54.5-61.8) and smaller eye diameter (38.8-40.5 vs. 41.1-45.8).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |