treatments-xml/data/B8/2C/44/B82C4464FFBCB830FF45F968F8C6FE0F.xml
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<document id="54AF8EB1A9299CA2B47AAA7C8E7EBE0F" ID="10.11646/zootaxa.4150.2.3" ID-DOI="10.11646/zootaxa.4150.2.3" ID-GBIF-Dataset="29be7477-05ac-4a36-a518-bddbf8d8464d" ID-ISSN="1175-5326" ID-Zenodo-Dep="256323" ID-ZooBank="8F929395-D423-4B3F-BB5F-C1D08B0FD8BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470896765710" checkinUser="plazi" docAuthor="Yang, Jian-Huan, Wang, Ying-Yong &amp; Chan, Bosco Pui-Lok" docDate="2016" docId="B82C4464FFBCB830FF45F968F8C6FE0F" docLanguage="en" docName="zootaxa.4150.2.3.pdf" docOrigin="Zootaxa 4150 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptobrachium tengchongense Yang, Wang &amp; Chan, 2016, sp. nov." docType="treatment" docVersion="7" lastPageNumber="145" masterDocId="44153C1CFFBFB83CFFD2FFDFFFF9F821" masterDocTitle="A new species of the genus Leptobrachium (Anura: Megophryidae) from the Gaoligongshan Mountain Range, China" masterLastPageNumber="148" masterPageNumber="133" pageNumber="136" updateTime="1698381807993" updateUser="plazi">
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<mods:title id="31331006CB02661487B74FF972CC3FC9">A new species of the genus Leptobrachium (Anura: Megophryidae) from the Gaoligongshan Mountain Range, China</mods:title>
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<mods:namePart id="94284FE08E5DF92D63EA54C35CBEA8C2">Yang, Jian-Huan</mods:namePart>
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<mods:namePart id="2CB30DD0ECF6D1D33586279B6A540E78">Wang, Ying-Yong</mods:namePart>
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<mods:namePart id="3E2FAA8B06D717D43A486F249D8F3ABD">Chan, Bosco Pui-Lok</mods:namePart>
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<mods:date id="1ACAD22EB4627D7DA553846BA1F1D48B">2016</mods:date>
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<treatment id="B82C4464FFBCB830FF45F968F8C6FE0F" ID-DOI="http://doi.org/10.5281/zenodo.5662298" ID-GBIF-Taxon="124329190" ID-Zenodo-Dep="5662298" LSID="urn:lsid:plazi:treatment:B82C4464FFBCB830FF45F968F8C6FE0F" httpUri="http://treatment.plazi.org/id/B82C4464FFBCB830FF45F968F8C6FE0F" lastPageId="12" lastPageNumber="145" pageId="3" pageNumber="136">
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<paragraph id="303AF572FFBCB83FFF45F968FD88FEF3" blockId="3.[151,625,1719,1746]" box="[151,625,1719,1746]" pageId="3" pageNumber="136">
<heading id="6B72421EFFBCB83FFF45F968FD88FEF3" bold="true" box="[151,625,1719,1746]" fontSize="11" level="1" pageId="3" pageNumber="136" reason="1">
<emphasis id="02F12960FFBCB83FFF45F968FD88FEF3" bold="true" box="[151,625,1719,1746]" pageId="3" pageNumber="136">
<taxonomicName id="F7858EF1FFBCB83FFF45F968FDF4FEF0" ID-CoL="3TKD2" box="[151,525,1719,1745]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="3" pageNumber="136" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFBCB83FFF45F968FDF4FEF0" bold="true" box="[151,525,1719,1745]" italics="true" pageId="3" pageNumber="136">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFBCB83FFDC6F967FD88FEF3" box="[532,625,1720,1746]" pageId="3" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="303AF572FFBCB83FFF45F921FB4FFFA2" blockId="3.[151,1437,1790,2031]" pageId="3" pageNumber="136">
<materialsCitation id="80EDFF2FFFBCB83FFF45F921FB4FFFA2" ID-GBIF-Occurrence="1291547720" collectingDate="2015-03-13" collectionCode="SYS" collectorName="J. H. Yang." country="China" county="Tengchong County" elevation="3" latitude="25.74062" location="The" longLatPrecision="1" longitude="98.69604" municipality="Gaoligongshan National Nature Reserve" pageId="3" pageNumber="136" specimenCount="1" stateProvince="Yunnan Province" typeStatus="holotype">
<emphasis id="02F12960FFBCB83FFF45F921FEF2FF36" bold="true" box="[151,267,1790,1815]" pageId="3" pageNumber="136">
<typeStatus id="EF3E4BD0FFBCB83FFF45F921FEFFFF36" box="[151,262,1790,1815]" pageId="3" pageNumber="136" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="56946DB7FFBCB83FFEC0F920FEBFFF36" box="[274,326,1791,1815]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="3" pageNumber="136">SYS</collectionCode>
a004603, adult male, calling on the floor in shrub about
<elevation id="BBA81241FFBCB83FFC15F920FC0BFF37" box="[967,1010,1791,1815]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="136" unit="m" value="3.0">
<quantity id="F77D5897FFBCB83FFC15F920FC0BFF37" box="[967,1010,1791,1815]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="136" unit="m" value="3.0">3 m</quantity>
</elevation>
from a rocky stream.
<location id="355AA3A9FFBCB83FFB39F921FAEEFF36" LSID="urn:lsid:plazi:treatment:B82C4464FFBCB830FF45F968F8C6FE0F:355AA3A9FFBCB83FFB39F921FAEEFF36" box="[1259,1303,1790,1815]" country="China" county="Tengchong County" latitude="25.74062" longLatPrecision="1" longitude="98.69604" municipality="Gaoligongshan National Nature Reserve" name="The" pageId="3" pageNumber="136" stateProvince="Yunnan Province">The</location>
stream was
<elevation id="BBA81241FFBCB83FFF45F8FDFF1BFF1B" box="[151,226,1826,1851]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="3" pageNumber="136" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">
<quantity id="F77D5897FFBCB83FFF45F8FDFF1BFF1B" box="[151,226,1826,1851]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="3" pageNumber="136" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">35 m</quantity>
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wide, flowing pass mature montane evergreen broadleaf forest at
<location id="355AA3A9FFBCB83FFC24F8FCFBBEFF1A" LSID="urn:lsid:plazi:treatment:B82C4464FFBCB830FF45F968F8C6FE0F:355AA3A9FFBCB83FFC24F8FCFBBEFF1A" box="[1014,1095,1827,1851]" country="China" county="Tengchong County" latitude="25.74062" longLatPrecision="1" longitude="98.69604" municipality="Gaoligongshan National Nature Reserve" name="Datang" pageId="3" pageNumber="136" stateProvince="Yunnan Province">Datang</location>
substation of the
<location id="355AA3A9FFBCB83FFAF2F8FDFF12FF7E" LSID="urn:lsid:plazi:treatment:B82C4464FFBCB830FF45F968F8C6FE0F:355AA3A9FFBCB83FFAF2F8FDFF12FF7E" country="China" county="Tengchong County" latitude="25.74062" longLatPrecision="1" longitude="98.69604" municipality="Gaoligongshan National Nature Reserve" name="Tengchong Section" pageId="3" pageNumber="136" stateProvince="Yunnan Province">Tengchong Section</location>
of
<collectingMunicipality id="D05E6F08FFBCB83FFEC2F899FD20FF7F" box="[272,729,1862,1887]" pageId="3" pageNumber="136">Gaoligongshan National Nature Reserve</collectingMunicipality>
,
<collectingCounty id="D95B8DFEFFBCB83FFD37F899FC45FF7E" box="[741,956,1862,1887]" pageId="3" pageNumber="136">Tengchong County</collectingCounty>
,
<collectingRegion id="F2413B90FFBCB83FFC15F898FB73FF7E" box="[967,1162,1862,1887]" country="China" name="Yunnan" pageId="3" pageNumber="136">Yunnan Province</collectingRegion>
,
<collectingCountry id="4892B5E2FFBCB83FFB47F899FB21FF7E" box="[1173,1240,1862,1887]" name="China" pageId="3" pageNumber="136">China</collectingCountry>
(
<geoCoordinate id="55B193B5FFBCB83FFB35F899FA61FF7F" box="[1255,1432,1862,1887]" direction="north" orientation="latitude" pageId="3" pageNumber="136" precision="1" value="25.74062">25°44'26.23'' N</geoCoordinate>
,
<geoCoordinate id="55B193B5FFBCB83FFF45F8B5FEBBFFA3" box="[151,322,1898,1923]" direction="east" orientation="longitude" pageId="3" pageNumber="136" precision="1" value="98.69604">98°41'45.74'' E</geoCoordinate>
,
<elevation id="BBA81241FFBCB83FFE9CF8B5FDA0FFA2" box="[334,601,1898,1923]" metricMagnitude="3" metricUnit="m" metricValue="2.06" pageId="3" pageNumber="136" unit="m" value="2060.0">
<quantity id="F77D5897FFBCB83FFE9CF8B5FE5AFFA3" box="[334,419,1898,1923]" metricMagnitude="3" metricUnit="m" metricValue="2.06" pageId="3" pageNumber="136" unit="m" value="2060.0">2060 m</quantity>
above sea level
</elevation>
), collected at 2100 h on
<collectingDate id="547F2A5AFFBCB83FFCB9F8B4FBEFFFA2" box="[875,1046,1898,1923]" pageId="3" pageNumber="136" value="2015-03-13">
<date id="443BD3B2FFBCB83FFCB9F8B4FBEFFFA2" box="[875,1046,1898,1923]" pageId="3" pageNumber="136" value="2015-03-13">13 March 2015</date>
</collectingDate>
by
<collectorName id="9D7090A4FFBCB83FFB92F8B4FB4FFFA2" box="[1088,1206,1899,1923]" pageId="3" pageNumber="136">J.H. Yang.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="303AF572FFBCB838FF15F851FEF4F8F4" blockId="3.[151,1437,1790,2031]" lastBlockId="4.[151,1436,151,213]" lastPageId="4" lastPageNumber="137" pageId="3" pageNumber="136">
<materialsCitation id="80EDFF2FFFBCB83FFF15F851FDA4FFEA" ID-GBIF-Occurrence="1291547718" collectingDate="2016-03-17" collectingDateMax="2016-03-18" collectingDateMin="2016-03-17" collectionCode="SYS" collectorName="J. H. Yang" country="China" county="Tengchong County" elevation="3" latitude="25.74062" location="The" longLatPrecision="1" longitude="98.69604" municipality="Gaoligongshan National Nature Reserve" pageId="3" pageNumber="136" specimenCount="1" stateProvince="Yunnan Province" typeStatus="paratype">
<emphasis id="02F12960FFBCB83FFF15F851FEBFFF86" bold="true" box="[199,326,1934,1959]" pageId="3" pageNumber="136">
<typeStatus id="EF3E4BD0FFBCB83FFF15F851FEB8FF86" box="[199,321,1934,1959]" pageId="3" pageNumber="136" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="56946DB7FFBCB83FFE9FF850FE78FF86" box="[333,385,1935,1959]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="3" pageNumber="136">SYS</collectionCode>
a0046234627, five adult males, from the same locality as holotype, collected at 20000130 h on
<collectingDate id="547F2A5AFFBCB83FFF05F86CFE4CFFEA" box="[215,437,1970,1995]" pageId="3" pageNumber="136" value="2016-03-17" valueMax="2016-03-18" valueMin="2016-03-17">
<date id="443BD3B2FFBCB83FFF05F86CFE4CFFEA" box="[215,437,1970,1995]" pageId="3" pageNumber="136" value="2016-03-17" valueMax="2016-03-18" valueMin="2016-03-17">1718 March 2016</date>
</collectingDate>
by
<collectorName id="9D7090A4FFBCB83FFE35F86CFDA4FFEA" box="[487,605,1971,1995]" pageId="3" pageNumber="136">J.H. Yang</collectorName>
</materialsCitation>
. SYS a0046284632, five adult males, from Cizhuhe substation of the Tengchong Section of Gaoligongshan National Nature Reserve (25°46'16.85'' N, 98°37'54.27'' E,
<quantity id="F77D5897FFBCB83FFB06F808FAD0FFCF" box="[1236,1321,2007,2031]" metricMagnitude="3" metricUnit="m" metricValue="2.2399999999999998" pageId="3" pageNumber="136" unit="m" value="2240.0">2240 m</quantity>
above sea level; 25°47'22.33'' N, 98°37'33.60'' E,
<quantity id="F77D5897FFBBB838FD99FF48FD66F88E" box="[587,671,151,176]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="4" pageNumber="137" unit="m" value="2510.0">2510 m</quantity>
above sea level), collected at 20000130 h on
<date id="443BD3B2FFBBB838FB74FF47FA83F891" box="[1190,1402,151,176]" pageId="4" pageNumber="137" value="2016-03-20" valueMax="2016-03-21" valueMin="2016-03-20">2021 March 2016</date>
by J.H. Yang.
</paragraph>
</subSubSection>
<caption id="64FAA5FAFFBBB838FF45F847FD6BFFCA" httpUri="https://zenodo.org/record/256325/files/figure.png" pageId="4" pageNumber="137" targetBox="[180,1393,274,1914]" targetPageId="4">
<paragraph id="303AF572FFBBB838FF45F847FD6BFFCA" blockId="4.[151,1436,1944,2027]" pageId="4" pageNumber="137">
<emphasis id="02F12960FFBBB838FF45F847FEEBFF8C" bold="true" box="[151,274,1944,1966]" pageId="4" pageNumber="137">FIGURE 2.</emphasis>
Bayesian inference (BI) tree derived from partial sequences of the mitochondrial 16S rRNA gene. Numbers above branches are bootstrap support values for maximum likelihood analyses (&gt;70 retained) and numbers below branches indicate Bayesian posterior probabilities (&gt; 70% retained).
</paragraph>
</caption>
<caption id="64FAA5FAFFBAB839FFAFFDE5FA06FA71" ID-Table-UUID="64FAA5FAFFBAB839FFAFFDE5FA06FA71" box="[125,1535,570,592]" httpUri="http://table.plazi.org/id/64FAA5FAFFBAB839FFAFFDE5FA06FA71" pageId="5" pageNumber="138" targetBox="[125,1862,616,1575]" targetIsTable="true" targetPageId="5">
<paragraph id="303AF572FFBAB839FFAFFDE5FA06FA71" blockId="5.[125,1535,570,592]" box="[125,1535,570,592]" pageId="5" pageNumber="138">
<emphasis id="02F12960FFBAB839FFAFFDE5FF11FA71" bold="true" box="[125,232,570,592]" pageId="5" pageNumber="138">TABLE 2.</emphasis>
Uncorrecteđ
<emphasis id="02F12960FFBAB839FEA3FDE4FE84FA71" box="[369,381,571,592]" italics="true" pageId="5" pageNumber="138">p</emphasis>
-đistances (%) among the
<taxonomicName id="F7858EF1FFBAB839FD51FDE5FCE4FA6E" box="[643,797,570,591]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="02F12960FFBAB839FD51FDE5FCE4FA6E" box="[643,797,570,591]" italics="true" pageId="5" pageNumber="138">Leptobrachium</emphasis>
</taxonomicName>
species anđ outgroups baseđ on a 16S rRNA fragment. (To be continueđ)
</paragraph>
</caption>
<paragraph id="303AF572FFBAB839FE3CFDB7F8F4FE06" pageId="5" pageNumber="138">
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<th id="CD649E4CFFBA47C3FE3CFDB7FDC0FA5F" box="[494,569,616,638]" gridcol="1" gridrow="0" pageId="5" pageNumber="138">(1!5)</th>
<th id="CD649E4CFFBA47C3FDA8FDB7FC19FA5F" box="[634,992,616,638]" gridcol="2" gridrow="0" pageId="5" pageNumber="138">(6!9) 10 11</th>
<th id="CD649E4CFFBA47C3FBF2FDB7FBB4FA5F" box="[1056,1101,616,638]" gridcol="3" gridrow="0" pageId="5" pageNumber="138">12</th>
<th id="CD649E4CFFBA47C3FB7FFDB7FB00FA5F" box="[1197,1273,616,638]" gridcol="4" gridrow="0" pageId="5" pageNumber="138">13</th>
<th id="CD649E4CFFBA47C3FAE8FDB7FA9FFA5F" box="[1338,1382,616,638]" gridcol="5" gridrow="0" pageId="5" pageNumber="138">14</th>
<th id="CD649E4CFFBA47C3FA15FDB7FA0AFA5F" box="[1479,1523,616,638]" gridcol="6" gridrow="0" pageId="5" pageNumber="138">15</th>
<th id="CD649E4CFFBA47C3F986FDB7F8BFFA5F" box="[1620,1862,616,638]" gridcol="7" gridrow="0" pageId="5" pageNumber="138">16 (17!21)</th>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFD50F8BFFA84" box="[125,1862,655,677]" gridrow="1" pageId="5" pageNumber="138" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="CD649E4CFFBA47C3FFAFFD50FE4AFA84" box="[125,435,655,677]" gridcol="0" gridrow="1" pageId="5" pageNumber="138">
(1!6)
<taxonomicName id="F7858EF1FFBAB839FF6BFD4FFEABFA84" box="[185,338,656,677]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="genus">
<emphasis id="02F12960FFBAB839FF6BFD4FFEABFA84" box="[185,338,656,677]" italics="true" pageId="5" pageNumber="138">Leptobrachium</emphasis>
</taxonomicName>
sp.
</th>
<td id="CD649E4CFFBA47C3FE3CFD50FDC0FA84" box="[494,569,655,677]" gridcol="1" gridrow="1" pageId="5" pageNumber="138">0.0</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFD6AF8BFFAEA" box="[125,1862,693,715]" gridrow="2" pageId="5" pageNumber="138" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="CD649E4CFFBA47C3FFAFFD6AFE4AFAEA" box="[125,435,693,715]" gridcol="0" gridrow="2" pageId="5" pageNumber="138">
(6!9)
<taxonomicName id="F7858EF1FFBAB839FF6BFD69FEBCFAEA" box="[185,325,694,715]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="ailaonicum">
<emphasis id="02F12960FFBAB839FF6BFD69FEBCFAEA" box="[185,325,694,715]" italics="true" pageId="5" pageNumber="138">L. ailaonicum</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFD6AFDC0FAEA" box="[494,569,693,715]" gridcol="1" gridrow="2" pageId="5" pageNumber="138">2.3!3.1</td>
<td id="CD649E4CFFBA47C3FDA8FD6AFC19FAEA" box="[634,992,693,715]" gridcol="2" gridrow="2" pageId="5" pageNumber="138">0.0!0.8</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFD03F8BFFAD2" box="[125,1862,732,755]" gridrow="3" pageId="5" pageNumber="138" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="CD649E4CFFBA47C3FFAFFD03FE4AFAD2" box="[125,435,732,755]" gridcol="0" gridrow="3" pageId="5" pageNumber="138">
(10)
<taxonomicName id="F7858EF1FFBAB839FF7FFD02FEFDFAD3" box="[173,260,733,754]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="banae">
<emphasis id="02F12960FFBAB839FF7FFD02FEFDFAD3" box="[173,260,733,754]" italics="true" pageId="5" pageNumber="138">L. banae</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFD03FDC0FAD2" box="[494,569,732,755]" gridcol="1" gridrow="3" pageId="5" pageNumber="138">1 2.8</td>
<td id="CD649E4CFFBA47C3FDA8FD03FC19FAD2" box="[634,992,732,755]" gridcol="2" gridrow="3" pageId="5" pageNumber="138">1 3.0!1 3.5!</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFCDCF8BFFB3B" box="[125,1862,771,794]" gridrow="4" pageId="5" pageNumber="138" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="CD649E4CFFBA47C3FFAFFCDCFE4AFB3B" box="[125,435,771,794]" gridcol="0" gridrow="4" pageId="5" pageNumber="138">
(11)
<taxonomicName id="F7858EF1FFBAB839FF7EFCDAFEEEFB38" box="[172,279,772,794]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="boringii">
<emphasis id="02F12960FFBAB839FF7EFCDAFEEEFB38" box="[172,279,772,794]" italics="true" pageId="5" pageNumber="138">L. boringii</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFCDCFDC0FB3B" box="[494,569,771,794]" gridcol="1" gridrow="4" pageId="5" pageNumber="138">4.8</td>
<td id="CD649E4CFFBA47C3FDA8FCDCFC19FB3B" box="[634,992,771,794]" gridcol="2" gridrow="4" pageId="5" pageNumber="138">4.6!4.8 1 3.0!</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFCF5F8BFFB60" box="[125,1862,810,833]" gridrow="5" pageId="5" pageNumber="138" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="CD649E4CFFBA47C3FFAFFCF5FE4AFB60" box="[125,435,810,833]" gridcol="0" gridrow="5" pageId="5" pageNumber="138">
(12)
<taxonomicName id="F7858EF1FFBAB839FF7FFCF3FED8FB61" box="[173,289,811,833]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="buchardi">
<emphasis id="02F12960FFBAB839FF7FFCF3FED8FB61" box="[173,289,811,833]" italics="true" pageId="5" pageNumber="138">L. buchardi</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFCF5FDC0FB60" box="[494,569,810,833]" gridcol="1" gridrow="5" pageId="5" pageNumber="138">1 0.6</td>
<td id="CD649E4CFFBA47C3FDA8FCF5FC19FB60" box="[634,992,810,833]" gridcol="2" gridrow="5" pageId="5" pageNumber="138">1 1.0!1 1.6 7.9 1 1.6</td>
<td id="CD649E4CFFBA47C3FBF2FCF5FBB4FB60" box="[1056,1101,810,833]" gridcol="3" gridrow="5" pageId="5" pageNumber="138">!</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFC8EF8BFFB49" box="[125,1862,849,872]" gridrow="6" pageId="5" pageNumber="138" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="CD649E4CFFBA47C3FFAFFC8EFE4AFB49" box="[125,435,849,872]" gridcol="0" gridrow="6" pageId="5" pageNumber="138">
(13)
<taxonomicName id="F7858EF1FFBAB839FF7FFC8CFECBFB46" box="[173,306,850,872]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="chapaense">
<emphasis id="02F12960FFBAB839FF7FFC8CFECBFB46" box="[173,306,850,872]" italics="true" pageId="5" pageNumber="138">L. chapaense</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFC8EFDC0FB49" box="[494,569,849,872]" gridcol="1" gridrow="6" pageId="5" pageNumber="138">5.0</td>
<td id="CD649E4CFFBA47C3FDA8FC8EFC19FB49" box="[634,992,849,872]" gridcol="2" gridrow="6" pageId="5" pageNumber="138">4.3!4.6 1 3.3 5.8</td>
<td id="CD649E4CFFBA47C3FBF2FC8EFBB4FB49" box="[1056,1101,849,872]" gridcol="3" gridrow="6" pageId="5" pageNumber="138">1 1.8</td>
<td id="CD649E4CFFBA47C3FB7FFC8EFB00FB49" box="[1197,1273,849,872]" gridcol="4" gridrow="6" pageId="5" pageNumber="138">!</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFCA7F8BFFBAE" box="[125,1862,888,911]" gridrow="7" pageId="5" pageNumber="138" rowspan-6="1" rowspan-7="1">
<th id="CD649E4CFFBA47C3FFAFFCA7FE4AFBAE" box="[125,435,888,911]" gridcol="0" gridrow="7" pageId="5" pageNumber="138">
(14)
<taxonomicName id="F7858EF1FFBAB839FF7FFCA5FEBCFBAE" box="[173,325,890,911]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="guangxiense">
<emphasis id="02F12960FFBAB839FF7FFCA5FEBCFBAE" box="[173,325,890,911]" italics="true" pageId="5" pageNumber="138">L. guangxiense</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFCA7FDC0FBAE" box="[494,569,888,911]" gridcol="1" gridrow="7" pageId="5" pageNumber="138">10.4</td>
<td id="CD649E4CFFBA47C3FDA8FCA7FC19FBAE" box="[634,992,888,911]" gridcol="2" gridrow="7" pageId="5" pageNumber="138">11.2!11.8 7.7 11.4</td>
<td id="CD649E4CFFBA47C3FBF2FCA7FBB4FBAE" box="[1056,1101,888,911]" gridcol="3" gridrow="7" pageId="5" pageNumber="138">4.8</td>
<td id="CD649E4CFFBA47C3FB7FFCA7FB00FBAE" box="[1197,1273,888,911]" gridcol="4" gridrow="7" pageId="5" pageNumber="138">11.6</td>
<td id="CD649E4CFFBA47C3FAE8FCA7FA9FFBAE" box="[1338,1382,888,911]" gridcol="5" gridrow="7" pageId="5" pageNumber="138">!</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFC7FF8BFFB97" box="[125,1862,928,950]" gridrow="8" pageId="5" pageNumber="138" rowspan-7="1">
<th id="CD649E4CFFBA47C3FFAFFC7FFE4AFB97" box="[125,435,928,950]" gridcol="0" gridrow="8" pageId="5" pageNumber="138">
(15)
<taxonomicName id="F7858EF1FFBAB839FF7FFC7EFEC0FB94" box="[173,313,928,950]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="hainanense">
<emphasis id="02F12960FFBAB839FF7FFC7EFEC0FB94" box="[173,313,928,950]" italics="true" pageId="5" pageNumber="138">L. hainanense</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFC7FFDC0FB97" box="[494,569,928,950]" gridcol="1" gridrow="8" pageId="5" pageNumber="138">10.8</td>
<td id="CD649E4CFFBA47C3FDA8FC7FFC19FB97" box="[634,992,928,950]" gridcol="2" gridrow="8" pageId="5" pageNumber="138">11.0!11.6 7.7 11.4</td>
<td id="CD649E4CFFBA47C3FBF2FC7FFBB4FB97" box="[1056,1101,928,950]" gridcol="3" gridrow="8" pageId="5" pageNumber="138">5.2</td>
<td id="CD649E4CFFBA47C3FB7FFC7FFB00FB97" box="[1197,1273,928,950]" gridcol="4" gridrow="8" pageId="5" pageNumber="138">12.0</td>
<td id="CD649E4CFFBA47C3FAE8FC7FFA9FFB97" box="[1338,1382,928,950]" gridcol="5" gridrow="8" pageId="5" pageNumber="138">1.2</td>
<td id="CD649E4CFFBA47C3FA15FC7FFA0AFB97" box="[1479,1523,928,950]" gridcol="6" gridrow="8" pageId="5" pageNumber="138">!</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFC18F8BFFBFC" box="[125,1862,967,989]" gridrow="9" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFC18FE4AFBFC" box="[125,435,967,989]" gridcol="0" gridrow="9" pageId="5" pageNumber="138">
(16)
<taxonomicName id="F7858EF1FFBAB839FF7FFC17FEE3FBFD" box="[173,282,967,989]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="hasseltii">
<emphasis id="02F12960FFBAB839FF7FFC17FEE3FBFD" box="[173,282,967,989]" italics="true" pageId="5" pageNumber="138">L. hasseltii</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFC18FDC0FBFC" box="[494,569,967,989]" gridcol="1" gridrow="9" pageId="5" pageNumber="138">11.4</td>
<td id="CD649E4CFFBA47C3FDA8FC18FC19FBFC" box="[634,992,967,989]" gridcol="2" gridrow="9" pageId="5" pageNumber="138">11.4!11.8 13.3 12.6</td>
<td id="CD649E4CFFBA47C3FBF2FC18FBB4FBFC" box="[1056,1101,967,989]" gridcol="3" gridrow="9" pageId="5" pageNumber="138">11.2</td>
<td id="CD649E4CFFBA47C3FB7FFC18FB00FBFC" box="[1197,1273,967,989]" gridcol="4" gridrow="9" pageId="5" pageNumber="138">11.6</td>
<td id="CD649E4CFFBA47C3FAE8FC18FA9FFBFC" box="[1338,1382,967,989]" gridcol="5" gridrow="9" pageId="5" pageNumber="138">12.0</td>
<td id="CD649E4CFFBA47C3FA15FC18FA0AFBFC" box="[1479,1523,967,989]" gridcol="6" gridrow="9" pageId="5" pageNumber="138">12.6</td>
<td id="CD649E4CFFBA47C3F986FC18F8BFFBFC" box="[1620,1862,967,989]" gridcol="7" gridrow="9" pageId="5" pageNumber="138">!</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFC31F8BFFC24" box="[125,1862,1006,1029]" gridrow="10" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFC31FE4AFC24" box="[125,435,1006,1029]" gridcol="0" gridrow="10" pageId="5" pageNumber="138">
(17!21)
<taxonomicName id="F7858EF1FFBAB839FF01FC30FEB8FC22" box="[211,321,1006,1028]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="huashen">
<emphasis id="02F12960FFBAB839FF01FC30FEB8FC22" box="[211,321,1006,1028]" italics="true" pageId="5" pageNumber="138">L. huashen</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFC31FDC0FC24" box="[494,569,1006,1029]" gridcol="1" gridrow="10" pageId="5" pageNumber="138">4.8!5.0</td>
<td id="CD649E4CFFBA47C3FDA8FC31FC19FC24" box="[634,992,1006,1029]" gridcol="2" gridrow="10" pageId="5" pageNumber="138">4.6!5.0 13.0!13.3 5.4!5.8</td>
<td id="CD649E4CFFBA47C3FBF2FC31FBB4FC24" box="[1056,1101,1006,1029]" gridcol="3" gridrow="10" pageId="5" pageNumber="138">12.0</td>
<td id="CD649E4CFFBA47C3FB7FFC31FB00FC24" box="[1197,1273,1006,1029]" gridcol="4" gridrow="10" pageId="5" pageNumber="138">2.5!2.7</td>
<td id="CD649E4CFFBA47C3FAE8FC31FA9FFC24" box="[1338,1382,1006,1029]" gridcol="5" gridrow="10" pageId="5" pageNumber="138">11.4</td>
<td id="CD649E4CFFBA47C3FA15FC31FA0AFC24" box="[1479,1523,1006,1029]" gridcol="6" gridrow="10" pageId="5" pageNumber="138">11.8</td>
<td id="CD649E4CFFBA47C3F986FC31F8BFFC24" box="[1620,1862,1006,1029]" gridcol="7" gridrow="10" pageId="5" pageNumber="138">12.0!12.2 0.0!0.6</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFBCAF8BFFC0D" box="[125,1862,1045,1068]" gridrow="11" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFBCAFE4AFC0D" box="[125,435,1045,1068]" gridcol="0" gridrow="11" pageId="5" pageNumber="138">
(22)
<taxonomicName id="F7858EF1FFBAB839FF7FFBC9FEFAFC0A" box="[173,259,1046,1067]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="ingeri">
<emphasis id="02F12960FFBAB839FF7FFBC9FEFAFC0A" box="[173,259,1046,1067]" italics="true" pageId="5" pageNumber="138">L. ingeri</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFBCAFDC0FC0D" box="[494,569,1045,1068]" gridcol="1" gridrow="11" pageId="5" pageNumber="138">13.7</td>
<td id="CD649E4CFFBA47C3FDA8FBCAFC19FC0D" box="[634,992,1045,1068]" gridcol="2" gridrow="11" pageId="5" pageNumber="138">13.5!13.7 13.5 15.3</td>
<td id="CD649E4CFFBA47C3FBF2FBCAFBB4FC0D" box="[1056,1101,1045,1068]" gridcol="3" gridrow="11" pageId="5" pageNumber="138">12.8</td>
<td id="CD649E4CFFBA47C3FB7FFBCAFB00FC0D" box="[1197,1273,1045,1068]" gridcol="4" gridrow="11" pageId="5" pageNumber="138">13.9</td>
<td id="CD649E4CFFBA47C3FAE8FBCAFA9FFC0D" box="[1338,1382,1045,1068]" gridcol="5" gridrow="11" pageId="5" pageNumber="138">13.9</td>
<td id="CD649E4CFFBA47C3FA15FBCAFA0AFC0D" box="[1479,1523,1045,1068]" gridcol="6" gridrow="11" pageId="5" pageNumber="138">14.5</td>
<td id="CD649E4CFFBA47C3F986FBCAF8BFFC0D" box="[1620,1862,1045,1068]" gridcol="7" gridrow="11" pageId="5" pageNumber="138">6.6 13.9</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFBE3F8BFFC72" box="[125,1862,1084,1107]" gridrow="12" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFBE3FE4AFC72" box="[125,435,1084,1107]" gridcol="0" gridrow="12" pageId="5" pageNumber="138">
(23)
<taxonomicName id="F7858EF1FFBAB839FF7FFBE2FEC5FC73" box="[173,316,1085,1106]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="leishanense">
<emphasis id="02F12960FFBAB839FF7FFBE2FEC5FC73" box="[173,316,1085,1106]" italics="true" pageId="5" pageNumber="138">L. leishanense</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFBE3FDC0FC72" box="[494,569,1084,1107]" gridcol="1" gridrow="12" pageId="5" pageNumber="138">2.5</td>
<td id="CD649E4CFFBA47C3FDA8FBE3FC19FC72" box="[634,992,1084,1107]" gridcol="2" gridrow="12" pageId="5" pageNumber="138">2.1!2.3 13.0 3.7</td>
<td id="CD649E4CFFBA47C3FBF2FBE3FBB4FC72" box="[1056,1101,1084,1107]" gridcol="3" gridrow="12" pageId="5" pageNumber="138">12.0</td>
<td id="CD649E4CFFBA47C3FB7FFBE3FB00FC72" box="[1197,1273,1084,1107]" gridcol="4" gridrow="12" pageId="5" pageNumber="138">3.7</td>
<td id="CD649E4CFFBA47C3FAE8FBE3FA9FFC72" box="[1338,1382,1084,1107]" gridcol="5" gridrow="12" pageId="5" pageNumber="138">11.8</td>
<td id="CD649E4CFFBA47C3FA15FBE3FA0AFC72" box="[1479,1523,1084,1107]" gridcol="6" gridrow="12" pageId="5" pageNumber="138">12.2</td>
<td id="CD649E4CFFBA47C3F986FBE3F8BFFC72" box="[1620,1862,1084,1107]" gridcol="7" gridrow="12" pageId="5" pageNumber="138">12.0 3.3!3.7</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFBBCF8BFFC5B" box="[125,1862,1123,1146]" gridrow="13" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFBBCFE4AFC5B" box="[125,435,1123,1146]" gridcol="0" gridrow="13" pageId="5" pageNumber="138">
(24)
<taxonomicName id="F7858EF1FFBAB839FF7FFBBBFEEDFC58" box="[173,276,1124,1145]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="leucops">
<emphasis id="02F12960FFBAB839FF7FFBBBFEEDFC58" box="[173,276,1124,1145]" italics="true" pageId="5" pageNumber="138">L. leucops</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFBBCFDC0FC5B" box="[494,569,1123,1146]" gridcol="1" gridrow="13" pageId="5" pageNumber="138">10.1</td>
<td id="CD649E4CFFBA47C3FDA8FBBCFC19FC5B" box="[634,992,1123,1146]" gridcol="2" gridrow="13" pageId="5" pageNumber="138">11.2!11.8 8.9 11.2</td>
<td id="CD649E4CFFBA47C3FBF2FBBCFBB4FC5B" box="[1056,1101,1123,1146]" gridcol="3" gridrow="13" pageId="5" pageNumber="138">4.6</td>
<td id="CD649E4CFFBA47C3FB7FFBBCFB00FC5B" box="[1197,1273,1123,1146]" gridcol="4" gridrow="13" pageId="5" pageNumber="138">10.8</td>
<td id="CD649E4CFFBA47C3FAE8FBBCFA9FFC5B" box="[1338,1382,1123,1146]" gridcol="5" gridrow="13" pageId="5" pageNumber="138">6.0</td>
<td id="CD649E4CFFBA47C3FA15FBBCFA0AFC5B" box="[1479,1523,1123,1146]" gridcol="6" gridrow="13" pageId="5" pageNumber="138">6.2</td>
<td id="CD649E4CFFBA47C3F986FBBCF8BFFC5B" box="[1620,1862,1123,1146]" gridcol="7" gridrow="13" pageId="5" pageNumber="138">11.6 11.2</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFB55F8BFFC80" box="[125,1862,1162,1185]" gridrow="14" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFB55FE4AFC80" box="[125,435,1162,1185]" gridcol="0" gridrow="14" pageId="5" pageNumber="138">
(25)
<taxonomicName id="F7858EF1FFBAB839FF7FFB53FF13FC81" box="[173,234,1163,1185]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="liui">
<emphasis id="02F12960FFBAB839FF7FFB53FF13FC81" box="[173,234,1163,1185]" italics="true" pageId="5" pageNumber="138">L. liui</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFB55FDC0FC80" box="[494,569,1162,1185]" gridcol="1" gridrow="14" pageId="5" pageNumber="138">3.5</td>
<td id="CD649E4CFFBA47C3FDA8FB55FC19FC80" box="[634,992,1162,1185]" gridcol="2" gridrow="14" pageId="5" pageNumber="138">3.3!3.7 14.1 4.1</td>
<td id="CD649E4CFFBA47C3FBF2FB55FBB4FC80" box="[1056,1101,1162,1185]" gridcol="3" gridrow="14" pageId="5" pageNumber="138">12.6</td>
<td id="CD649E4CFFBA47C3FB7FFB55FB00FC80" box="[1197,1273,1162,1185]" gridcol="4" gridrow="14" pageId="5" pageNumber="138">5.2</td>
<td id="CD649E4CFFBA47C3FAE8FB55FA9FFC80" box="[1338,1382,1162,1185]" gridcol="5" gridrow="14" pageId="5" pageNumber="138">12.4</td>
<td id="CD649E4CFFBA47C3FA15FB55FA0AFC80" box="[1479,1523,1162,1185]" gridcol="6" gridrow="14" pageId="5" pageNumber="138">12.8</td>
<td id="CD649E4CFFBA47C3F986FB55F8BFFC80" box="[1620,1862,1162,1185]" gridcol="7" gridrow="14" pageId="5" pageNumber="138">12.4 4.6!4.8</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFB6EF8BFFCE9" box="[125,1862,1201,1224]" gridrow="15" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFB6EFE4AFCE9" box="[125,435,1201,1224]" gridcol="0" gridrow="15" pageId="5" pageNumber="138">
(26)
<taxonomicName id="F7858EF1FFBAB839FF7FFB6CFEA1FCE6" box="[173,344,1202,1224]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="masatakasatoi">
<emphasis id="02F12960FFBAB839FF7FFB6CFEA1FCE6" box="[173,344,1202,1224]" italics="true" pageId="5" pageNumber="138">L. masatakasatoi</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFB6EFDC0FCE9" box="[494,569,1201,1224]" gridcol="1" gridrow="15" pageId="5" pageNumber="138">4.3</td>
<td id="CD649E4CFFBA47C3FDA8FB6EFC19FCE9" box="[634,992,1201,1224]" gridcol="2" gridrow="15" pageId="5" pageNumber="138">5.0!5.8 12.8 4.8</td>
<td id="CD649E4CFFBA47C3FBF2FB6EFBB4FCE9" box="[1056,1101,1201,1224]" gridcol="3" gridrow="15" pageId="5" pageNumber="138">11.6</td>
<td id="CD649E4CFFBA47C3FB7FFB6EFB00FCE9" box="[1197,1273,1201,1224]" gridcol="4" gridrow="15" pageId="5" pageNumber="138">5.4</td>
<td id="CD649E4CFFBA47C3FAE8FB6EFA9FFCE9" box="[1338,1382,1201,1224]" gridcol="5" gridrow="15" pageId="5" pageNumber="138">11.0</td>
<td id="CD649E4CFFBA47C3FA15FB6EFA0AFCE9" box="[1479,1523,1201,1224]" gridcol="6" gridrow="15" pageId="5" pageNumber="138">11.4</td>
<td id="CD649E4CFFBA47C3F986FB6EF8BFFCE9" box="[1620,1862,1201,1224]" gridcol="7" gridrow="15" pageId="5" pageNumber="138">12.0 5.0!5.2</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFB07F8BFFCCE" box="[125,1862,1240,1263]" gridrow="16" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFB07FE4AFCCE" box="[125,435,1240,1263]" gridcol="0" gridrow="16" pageId="5" pageNumber="138">
(27)
<taxonomicName id="F7858EF1FFBAB839FF7FFB05FEE0FCCF" box="[173,281,1241,1263]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="mouhoti">
<emphasis id="02F12960FFBAB839FF7FFB05FEE0FCCF" box="[173,281,1241,1263]" italics="true" pageId="5" pageNumber="138">L. mouhoti</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFB07FDC0FCCE" box="[494,569,1240,1263]" gridcol="1" gridrow="16" pageId="5" pageNumber="138">9.0</td>
<td id="CD649E4CFFBA47C3FDA8FB07FC19FCCE" box="[634,992,1240,1263]" gridcol="2" gridrow="16" pageId="5" pageNumber="138">10.8!11.0 7.7 10.6</td>
<td id="CD649E4CFFBA47C3FBF2FB07FBB4FCCE" box="[1056,1101,1240,1263]" gridcol="3" gridrow="16" pageId="5" pageNumber="138">4.1</td>
<td id="CD649E4CFFBA47C3FB7FFB07FB00FCCE" box="[1197,1273,1240,1263]" gridcol="4" gridrow="16" pageId="5" pageNumber="138">10.8</td>
<td id="CD649E4CFFBA47C3FAE8FB07FA9FFCCE" box="[1338,1382,1240,1263]" gridcol="5" gridrow="16" pageId="5" pageNumber="138">4.3</td>
<td id="CD649E4CFFBA47C3FA15FB07FA0AFCCE" box="[1479,1523,1240,1263]" gridcol="6" gridrow="16" pageId="5" pageNumber="138">4.6</td>
<td id="CD649E4CFFBA47C3F986FB07F8BFFCCE" box="[1620,1862,1240,1263]" gridcol="7" gridrow="16" pageId="5" pageNumber="138">12.0 11.2</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFADFF8BFFD37" box="[125,1862,1280,1302]" gridrow="17" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFADFFE4AFD37" box="[125,435,1280,1302]" gridcol="0" gridrow="17" pageId="5" pageNumber="138">
(28)
<taxonomicName id="F7858EF1FFBAB839FF7FFADEFEB2FD34" box="[173,331,1280,1302]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="ngoclinhense">
<emphasis id="02F12960FFBAB839FF7FFADEFEB2FD34" box="[173,331,1280,1302]" italics="true" pageId="5" pageNumber="138">L. ngoclinhense</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFADFFDC0FD37" box="[494,569,1280,1302]" gridcol="1" gridrow="17" pageId="5" pageNumber="138">10.1</td>
<td id="CD649E4CFFBA47C3FDA8FADFFC19FD37" box="[634,992,1280,1302]" gridcol="2" gridrow="17" pageId="5" pageNumber="138">10.6!11.2 6.0 10.8</td>
<td id="CD649E4CFFBA47C3FBF2FADFFBB4FD37" box="[1056,1101,1280,1302]" gridcol="3" gridrow="17" pageId="5" pageNumber="138">2.9</td>
<td id="CD649E4CFFBA47C3FB7FFADFFB00FD37" box="[1197,1273,1280,1302]" gridcol="4" gridrow="17" pageId="5" pageNumber="138">11.0</td>
<td id="CD649E4CFFBA47C3FAE8FADFFA9FFD37" box="[1338,1382,1280,1302]" gridcol="5" gridrow="17" pageId="5" pageNumber="138">3.7</td>
<td id="CD649E4CFFBA47C3FA15FADFFA0AFD37" box="[1479,1523,1280,1302]" gridcol="6" gridrow="17" pageId="5" pageNumber="138">4.1</td>
<td id="CD649E4CFFBA47C3F986FADFF8BFFD37" box="[1620,1862,1280,1302]" gridcol="7" gridrow="17" pageId="5" pageNumber="138">11.0 11.2!11.4</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFAF8F8BFFD1C" box="[125,1862,1319,1341]" gridrow="18" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFAF8FE4AFD1C" box="[125,435,1319,1341]" gridcol="0" gridrow="18" pageId="5" pageNumber="138">
(29)
<taxonomicName id="F7858EF1FFBAB839FF7FFAF7FEB1FD1D" box="[173,328,1319,1341]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="promustache">
<emphasis id="02F12960FFBAB839FF7FFAF7FEB1FD1D" box="[173,328,1319,1341]" italics="true" pageId="5" pageNumber="138">L. promustache</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFAF8FDC0FD1C" box="[494,569,1319,1341]" gridcol="1" gridrow="18" pageId="5" pageNumber="138">4.8</td>
<td id="CD649E4CFFBA47C3FDA8FAF8FC19FD1C" box="[634,992,1319,1341]" gridcol="2" gridrow="18" pageId="5" pageNumber="138">3.7!4.1 13.5 5.8</td>
<td id="CD649E4CFFBA47C3FBF2FAF8FBB4FD1C" box="[1056,1101,1319,1341]" gridcol="3" gridrow="18" pageId="5" pageNumber="138">11.6</td>
<td id="CD649E4CFFBA47C3FB7FFAF8FB00FD1C" box="[1197,1273,1319,1341]" gridcol="4" gridrow="18" pageId="5" pageNumber="138">6.8</td>
<td id="CD649E4CFFBA47C3FAE8FAF8FA9FFD1C" box="[1338,1382,1319,1341]" gridcol="5" gridrow="18" pageId="5" pageNumber="138">12.4</td>
<td id="CD649E4CFFBA47C3FA15FAF8FA0AFD1C" box="[1479,1523,1319,1341]" gridcol="6" gridrow="18" pageId="5" pageNumber="138">12.2</td>
<td id="CD649E4CFFBA47C3F986FAF8F8BFFD1C" box="[1620,1862,1319,1341]" gridcol="7" gridrow="18" pageId="5" pageNumber="138">13.3 7.2!7.5</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFA91F8BFFD44" box="[125,1862,1358,1381]" gridrow="19" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFA91FE4AFD44" box="[125,435,1358,1381]" gridcol="0" gridrow="19" pageId="5" pageNumber="138">
(30)
<taxonomicName id="F7858EF1FFBAB839FF7FFA90FEF4FD42" box="[173,269,1358,1380]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="pullum">
<emphasis id="02F12960FFBAB839FF7FFA90FEF4FD42" box="[173,269,1358,1380]" italics="true" pageId="5" pageNumber="138">L. pullum</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFA91FDC0FD44" box="[494,569,1358,1381]" gridcol="1" gridrow="19" pageId="5" pageNumber="138">10.4</td>
<td id="CD649E4CFFBA47C3FDA8FA91FC19FD44" box="[634,992,1358,1381]" gridcol="2" gridrow="19" pageId="5" pageNumber="138">11.0!11.4 7.9 11.2</td>
<td id="CD649E4CFFBA47C3FBF2FA91FBB4FD44" box="[1056,1101,1358,1381]" gridcol="3" gridrow="19" pageId="5" pageNumber="138">4.3</td>
<td id="CD649E4CFFBA47C3FB7FFA91FB00FD44" box="[1197,1273,1358,1381]" gridcol="4" gridrow="19" pageId="5" pageNumber="138">11.0</td>
<td id="CD649E4CFFBA47C3FAE8FA91FA9FFD44" box="[1338,1382,1358,1381]" gridcol="5" gridrow="19" pageId="5" pageNumber="138">5.4</td>
<td id="CD649E4CFFBA47C3FA15FA91FA0AFD44" box="[1479,1523,1358,1381]" gridcol="6" gridrow="19" pageId="5" pageNumber="138">5.2</td>
<td id="CD649E4CFFBA47C3F986FA91F8BFFD44" box="[1620,1862,1358,1381]" gridcol="7" gridrow="19" pageId="5" pageNumber="138">12.6 11.8</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFAAAF8BFFDAD" box="[125,1862,1397,1420]" gridrow="20" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFAAAFE4AFDAD" box="[125,435,1397,1420]" gridcol="0" gridrow="20" pageId="5" pageNumber="138">
(31)
<taxonomicName id="F7858EF1FFBAB839FF7FFAA9FEC2FDAA" box="[173,315,1398,1419]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="rakhinensis">
<emphasis id="02F12960FFBAB839FF7FFAA9FEC2FDAA" box="[173,315,1398,1419]" italics="true" pageId="5" pageNumber="138">L. rakhinensis</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFAAAFDC0FDAD" box="[494,569,1397,1420]" gridcol="1" gridrow="20" pageId="5" pageNumber="138">14.3</td>
<td id="CD649E4CFFBA47C3FDA8FAAAFC19FDAD" box="[634,992,1397,1420]" gridcol="2" gridrow="20" pageId="5" pageNumber="138">14.5!14.9 14.5 15.1</td>
<td id="CD649E4CFFBA47C3FBF2FAAAFBB4FDAD" box="[1056,1101,1397,1420]" gridcol="3" gridrow="20" pageId="5" pageNumber="138">14.7</td>
<td id="CD649E4CFFBA47C3FB7FFAAAFB00FDAD" box="[1197,1273,1397,1420]" gridcol="4" gridrow="20" pageId="5" pageNumber="138">13.5</td>
<td id="CD649E4CFFBA47C3FAE8FAAAFA9FFDAD" box="[1338,1382,1397,1420]" gridcol="5" gridrow="20" pageId="5" pageNumber="138">15.3</td>
<td id="CD649E4CFFBA47C3FA15FAAAFA0AFDAD" box="[1479,1523,1397,1420]" gridcol="6" gridrow="20" pageId="5" pageNumber="138">15.5</td>
<td id="CD649E4CFFBA47C3F986FAAAF8BFFDAD" box="[1620,1862,1397,1420]" gridcol="7" gridrow="20" pageId="5" pageNumber="138">11.2 12.8!13.3</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFA43F8BFFD92" box="[125,1862,1436,1459]" gridrow="21" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFA43FE4AFD92" box="[125,435,1436,1459]" gridcol="0" gridrow="21" pageId="5" pageNumber="138">
(32)
<taxonomicName id="F7858EF1FFBAB839FF7FFA42FED8FD93" box="[173,289,1437,1458]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="xanthops">
<emphasis id="02F12960FFBAB839FF7FFA42FED8FD93" box="[173,289,1437,1458]" italics="true" pageId="5" pageNumber="138">L. xanthops</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFA43FDC0FD92" box="[494,569,1436,1459]" gridcol="1" gridrow="21" pageId="5" pageNumber="138">9.9</td>
<td id="CD649E4CFFBA47C3FDA8FA43FC19FD92" box="[634,992,1436,1459]" gridcol="2" gridrow="21" pageId="5" pageNumber="138">10.1!10.8 7.0 10.6</td>
<td id="CD649E4CFFBA47C3FBF2FA43FBB4FD92" box="[1056,1101,1436,1459]" gridcol="3" gridrow="21" pageId="5" pageNumber="138">4.1</td>
<td id="CD649E4CFFBA47C3FB7FFA43FB00FD92" box="[1197,1273,1436,1459]" gridcol="4" gridrow="21" pageId="5" pageNumber="138">10.8</td>
<td id="CD649E4CFFBA47C3FAE8FA43FA9FFD92" box="[1338,1382,1436,1459]" gridcol="5" gridrow="21" pageId="5" pageNumber="138">4.8</td>
<td id="CD649E4CFFBA47C3FA15FA43FA0AFD92" box="[1479,1523,1436,1459]" gridcol="6" gridrow="21" pageId="5" pageNumber="138">4.6</td>
<td id="CD649E4CFFBA47C3F986FA43F8BFFD92" box="[1620,1862,1436,1459]" gridcol="7" gridrow="21" pageId="5" pageNumber="138">11.2 11.2</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFA1CF8BFFDFB" box="[125,1862,1475,1498]" gridrow="22" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFA1CFE4AFDFB" box="[125,435,1475,1498]" gridcol="0" gridrow="22" pageId="5" pageNumber="138">
(33)
<taxonomicName id="F7858EF1FFBAB839FF7FFA1BFEB7FDF8" box="[173,334,1476,1497]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="xanthospilum">
<emphasis id="02F12960FFBAB839FF7FFA1BFEB7FDF8" box="[173,334,1476,1497]" italics="true" pageId="5" pageNumber="138">L. xanthospilum</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFA1CFDC0FDFB" box="[494,569,1475,1498]" gridcol="1" gridrow="22" pageId="5" pageNumber="138">11.8</td>
<td id="CD649E4CFFBA47C3FDA8FA1CFC19FDFB" box="[634,992,1475,1498]" gridcol="2" gridrow="22" pageId="5" pageNumber="138">12.2!12.6 7.9 12.8</td>
<td id="CD649E4CFFBA47C3FBF2FA1CFBB4FDFB" box="[1056,1101,1475,1498]" gridcol="3" gridrow="22" pageId="5" pageNumber="138">6.6</td>
<td id="CD649E4CFFBA47C3FB7FFA1CFB00FDFB" box="[1197,1273,1475,1498]" gridcol="4" gridrow="22" pageId="5" pageNumber="138">12.4</td>
<td id="CD649E4CFFBA47C3FAE8FA1CFA9FFDFB" box="[1338,1382,1475,1498]" gridcol="5" gridrow="22" pageId="5" pageNumber="138">8.1</td>
<td id="CD649E4CFFBA47C3FA15FA1CFA0AFDFB" box="[1479,1523,1475,1498]" gridcol="6" gridrow="22" pageId="5" pageNumber="138">8.5</td>
<td id="CD649E4CFFBA47C3F986FA1CF8BFFDFB" box="[1620,1862,1475,1498]" gridcol="7" gridrow="22" pageId="5" pageNumber="138">12.2 13.3</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFFA35F8BFFE20" box="[125,1862,1514,1537]" gridrow="23" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFFA35FE4AFE20" box="[125,435,1514,1537]" gridcol="0" gridrow="23" pageId="5" pageNumber="138">
(34)
<taxonomicName id="F7858EF1FFBAB839FF7FFA34FE4AFE21" box="[173,435,1515,1536]" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="rhodostigmatus">
<emphasis id="02F12960FFBAB839FF7FFA34FE4AFE21" box="[173,435,1515,1536]" italics="true" pageId="5" pageNumber="138">Oreolalax rhodostigmatus</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CFA35FDC0FE20" box="[494,569,1514,1537]" gridcol="1" gridrow="23" pageId="5" pageNumber="138">11.4</td>
<td id="CD649E4CFFBA47C3FDA8FA35FC19FE20" box="[634,992,1514,1537]" gridcol="2" gridrow="23" pageId="5" pageNumber="138">11.4!11.6 13.3 12.4</td>
<td id="CD649E4CFFBA47C3FBF2FA35FBB4FE20" box="[1056,1101,1514,1537]" gridcol="3" gridrow="23" pageId="5" pageNumber="138">11.6</td>
<td id="CD649E4CFFBA47C3FB7FFA35FB00FE20" box="[1197,1273,1514,1537]" gridcol="4" gridrow="23" pageId="5" pageNumber="138">10.4</td>
<td id="CD649E4CFFBA47C3FAE8FA35FA9FFE20" box="[1338,1382,1514,1537]" gridcol="5" gridrow="23" pageId="5" pageNumber="138">11.8</td>
<td id="CD649E4CFFBA47C3FA15FA35FA0AFE20" box="[1479,1523,1514,1537]" gridcol="6" gridrow="23" pageId="5" pageNumber="138">11.8</td>
<td id="CD649E4CFFBA47C3F986FA35F8BFFE20" box="[1620,1862,1514,1537]" gridcol="7" gridrow="23" pageId="5" pageNumber="138">12.2 11.4</td>
</tr>
<tr id="8EB5F730FFBA47C3FFAFF9CEF8BFFE06" box="[125,1862,1553,1575]" gridrow="24" pageId="5" pageNumber="138">
<th id="CD649E4CFFBA47C3FFAFF9CEFE4AFE06" box="[125,435,1553,1575]" gridcol="0" gridrow="24" pageId="5" pageNumber="138">
(35)
<taxonomicName id="F7858EF1FFBAB839FF7FF9CDFE72FE06" box="[173,395,1554,1575]" class="Amphibia" family="Megophryidae" genus="Scutiger" kingdom="Animalia" order="Anura" pageId="5" pageNumber="138" phylum="Chordata" rank="species" species="chintingensis">
<emphasis id="02F12960FFBAB839FF7FF9CDFE72FE06" box="[173,395,1554,1575]" italics="true" pageId="5" pageNumber="138">Scutiger chintingensis</emphasis>
</taxonomicName>
</th>
<td id="CD649E4CFFBA47C3FE3CF9CEFDC0FE06" box="[494,569,1553,1575]" gridcol="1" gridrow="24" pageId="5" pageNumber="138">12.8</td>
<td id="CD649E4CFFBA47C3FDA8F9CEFC19FE06" box="[634,992,1553,1575]" gridcol="2" gridrow="24" pageId="5" pageNumber="138">12.0!12.2 14.5 12.6</td>
<td id="CD649E4CFFBA47C3FBF2F9CEFBB4FE06" box="[1056,1101,1553,1575]" gridcol="3" gridrow="24" pageId="5" pageNumber="138">12.4</td>
<td id="CD649E4CFFBA47C3FB7FF9CEFB00FE06" box="[1197,1273,1553,1575]" gridcol="4" gridrow="24" pageId="5" pageNumber="138">12.0</td>
<td id="CD649E4CFFBA47C3FAE8F9CEFA9FFE06" box="[1338,1382,1553,1575]" gridcol="5" gridrow="24" pageId="5" pageNumber="138">12.6</td>
<td id="CD649E4CFFBA47C3FA15F9CEFA0AFE06" box="[1479,1523,1553,1575]" gridcol="6" gridrow="24" pageId="5" pageNumber="138">12.2</td>
<td id="CD649E4CFFBA47C3F986F9CEF8BFFE06" box="[1620,1862,1553,1575]" gridcol="7" gridrow="24" pageId="5" pageNumber="138">12.6 12.2</td>
</tr>
</table>
</paragraph>
<subSubSection id="789FA6F9FFBAB83AFA29F9E5F8E9FE75" lastPageId="6" lastPageNumber="139" pageId="5" pageNumber="138" type="description">
<paragraph id="303AF572FFBAB83AFA29F9E5F8E5FA8A" blockId="5.[1531,1862,1594,1616]" lastBlockId="6.[119,1820,615,1621]" lastPageId="6" lastPageNumber="139" pageId="5" pageNumber="138">
<emphasis id="02F12960FFBAB839FA29F9E5F8BFFE71" box="[1531,1862,1594,1616]" italics="true" pageId="5" pageNumber="138">……continued on the next page</emphasis>
22 23 24 25 26 27 28 29 30 31 32 33 34 35
</paragraph>
<caption id="64FAA5FAFFB9B83AFFA5FDB8FE9AFA5F" box="[119,355,615,638]" pageId="6" pageNumber="139">
<paragraph id="303AF572FFB9B83AFFA5FDB8FE9AFA5F" blockId="6.[119,1820,615,1621]" box="[119,355,615,638]" pageId="6" pageNumber="139">
<emphasis id="02F12960FFB9B83AFFA5FDB8FF1AFA5C" bold="true" box="[119,227,615,637]" pageId="6" pageNumber="139">TABLE 2.</emphasis>
(Continueđ)
</paragraph>
</caption>
<paragraph id="303AF572FFB9B83AFF43FD61FE89FAF5" blockId="6.[119,1820,615,1621]" box="[145,368,702,724]" pageId="6" pageNumber="139">
6)
<taxonomicName id="F7858EF1FFB9B83AFF66FD61FEB4FAF2" box="[180,333,702,723]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="genus">
<emphasis id="02F12960FFB9B83AFF66FD61FEB4FAF2" box="[180,333,702,723]" italics="true" pageId="6" pageNumber="139">Leptobrachium</emphasis>
</taxonomicName>
sp.
</paragraph>
<paragraph id="303AF572FFB9B83AFF43FD3DFEC6FAD9" blockId="6.[119,1820,615,1621]" box="[145,319,738,761]" pageId="6" pageNumber="139">
!9)
<taxonomicName id="F7858EF1FFB9B83AFF66FD3BFEC6FAD9" box="[180,319,739,761]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="ailaonicum">
<emphasis id="02F12960FFB9B83AFF66FD3BFEC6FAD9" box="[180,319,739,761]" italics="true" pageId="6" pageNumber="139">L. ailaonicum</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="303AF572FFB9B83AFF4BFCD5FF06FB3E" blockId="6.[119,1820,615,1621]" box="[153,255,778,800]" pageId="6" pageNumber="139">
)
<taxonomicName id="F7858EF1FFB9B83AFF75FCD4FF06FB3E" box="[167,255,778,800]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="banae">
<emphasis id="02F12960FFB9B83AFF75FCD4FF06FB3E" box="[167,255,778,800]" italics="true" pageId="6" pageNumber="139">L. banae</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="303AF572FFB9B83AFF4AFCEEFEEBFB67" blockId="6.[119,1820,615,1621]" box="[152,274,817,839]" pageId="6" pageNumber="139">
)
<taxonomicName id="F7858EF1FFB9B83AFF75FCEDFEEBFB67" box="[167,274,817,839]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="boringii">
<emphasis id="02F12960FFB9B83AFF75FCEDFEEBFB67" box="[167,274,817,839]" italics="true" pageId="6" pageNumber="139">L. boringii</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="303AF572FFB9B83AFF4BFC87FEE5FB4C" blockId="6.[119,1820,615,1621]" box="[153,284,856,878]" pageId="6" pageNumber="139">
)
<taxonomicName id="F7858EF1FFB9B83AFF75FC86FEE5FB4C" box="[167,284,856,878]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="buchardi">
<emphasis id="02F12960FFB9B83AFF75FC86FEE5FB4C" box="[167,284,856,878]" italics="true" pageId="6" pageNumber="139">L. buchardi</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="303AF572FFB9B83AFF4BFC5FFED5FBB5" blockId="6.[119,1820,615,1621]" box="[153,300,895,918]" pageId="6" pageNumber="139">
)
<taxonomicName id="F7858EF1FFB9B83AFF75FC5FFED5FBB5" box="[167,300,895,917]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="chapaense">
<emphasis id="02F12960FFB9B83AFF75FC5FFED5FBB5" box="[167,300,895,917]" italics="true" pageId="6" pageNumber="139">L. chapaense</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="303AF572FFB9B83AFF4BFC78FEB9FB9D" blockId="6.[119,1820,615,1621]" box="[153,320,935,957]" pageId="6" pageNumber="139">
)
<taxonomicName id="F7858EF1FFB9B83AFF75FC78FEB9FB9D" box="[167,320,935,956]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="guangxiense">
<emphasis id="02F12960FFB9B83AFF75FC78FEB9FB9D" box="[167,320,935,956]" italics="true" pageId="6" pageNumber="139">L. guangxiense</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="303AF572FFB9B83AFF4BFC12FECDFBC2" blockId="6.[119,1820,615,1621]" box="[153,308,973,995]" pageId="6" pageNumber="139">
)
<taxonomicName id="F7858EF1FFB9B83AFF75FC11FECDFBC2" box="[167,308,974,995]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="hainanense">
<emphasis id="02F12960FFB9B83AFF75FC11FECDFBC2" box="[167,308,974,995]" italics="true" pageId="6" pageNumber="139">L. hainanense</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="303AF572FFB9B83AFF4BFC2BFEECFC2B" blockId="6.[119,1820,615,1621]" box="[153,277,1012,1034]" pageId="6" pageNumber="139">
)
<taxonomicName id="F7858EF1FFB9B83AFF75FC2AFEECFC2B" box="[167,277,1013,1034]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="hasseltii">
<emphasis id="02F12960FFB9B83AFF75FC2AFEECFC2B" box="[167,277,1013,1034]" italics="true" pageId="6" pageNumber="139">L. hasseltii</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="303AF572FFB9B83AFF4FFBC3FEC2FC10" blockId="6.[119,1820,615,1621]" box="[157,315,1052,1074]" pageId="6" pageNumber="139">
!21)
<taxonomicName id="F7858EF1FFB9B83AFF1FFBC2FEC2FC10" box="[205,315,1052,1074]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="huashen">
<emphasis id="02F12960FFB9B83AFF1FFBC2FEC2FC10" box="[205,315,1052,1074]" italics="true" pageId="6" pageNumber="139">L. huashen</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="303AF572FFB9B83AFF48FB9CFE0DFC79" blockId="6.[119,1820,615,1621]" box="[154,500,1091,1113]" pageId="6" pageNumber="139">
)
<taxonomicName id="F7858EF1FFB9B83AFF75FB9BFF07FC78" box="[167,254,1092,1113]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="ingeri">
<emphasis id="02F12960FFB9B83AFF75FB9BFF07FC78" box="[167,254,1092,1113]" italics="true" pageId="6" pageNumber="139">L. ingeri</emphasis>
</taxonomicName>
!
</paragraph>
<paragraph id="303AF572FFB9B83AFF48FBB5FDA0FC5E" blockId="6.[119,1820,615,1621]" box="[154,601,1130,1152]" pageId="6" pageNumber="139">
)
<taxonomicName id="F7858EF1FFB9B83AFF75FBB4FECEFC5E" box="[167,311,1130,1152]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="leishanense">
<emphasis id="02F12960FFB9B83AFF75FBB4FECEFC5E" box="[167,311,1130,1152]" italics="true" pageId="6" pageNumber="139">L. leishanense</emphasis>
</taxonomicName>
1 4.7!
</paragraph>
<paragraph id="303AF572FFB9B83AFF48FB4EFD47FC87" blockId="6.[119,1820,615,1621]" box="[154,702,1168,1191]" pageId="6" pageNumber="139">
)
<taxonomicName id="F7858EF1FFB9B83AFF75FB4DFEF6FC87" box="[167,271,1169,1191]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="leucops">
<emphasis id="02F12960FFB9B83AFF75FB4DFEF6FC87" box="[167,271,1169,1191]" italics="true" pageId="6" pageNumber="139">L. leucops</emphasis>
</taxonomicName>
1 2.2 1 1.6!
</paragraph>
<paragraph id="303AF572FFB9B83AFF48FB67FCDBFCEC" blockId="6.[119,1820,615,1621]" box="[154,802,1208,1230]" pageId="6" pageNumber="139">
)
<taxonomicName id="F7858EF1FFB9B83AFF75FB66FF1DFCEC" box="[167,228,1208,1230]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="liui">
<emphasis id="02F12960FFB9B83AFF75FB66FF1DFCEC" box="[167,228,1208,1230]" italics="true" pageId="6" pageNumber="139">L. liui</emphasis>
</taxonomicName>
1 5.3 2.3 1 2.2!
</paragraph>
<paragraph id="303AF572FFB9B83AFF48FB00F8E9FE75" blockId="6.[119,1820,615,1621]" pageId="6" pageNumber="139">
)
<taxonomicName id="F7858EF1FFB9B83AFF75FB3FFEAAFCD5" box="[167,339,1247,1269]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="masatakasatoi">
<emphasis id="02F12960FFB9B83AFF75FB3FFEAAFCD5" box="[167,339,1247,1269]" italics="true" pageId="6" pageNumber="139">L. masatakasatoi</emphasis>
</taxonomicName>
1 4.5 4.3 1 1.0 5.2!)
<taxonomicName id="F7858EF1FFB9B83AFF75FAD8FEEDFD3A" box="[167,276,1286,1308]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="mouhoti">
<emphasis id="02F12960FFB9B83AFF75FAD8FEEDFD3A" box="[167,276,1286,1308]" italics="true" pageId="6" pageNumber="139">L. mouhoti</emphasis>
</taxonomicName>
1 3.3 1 1.0 4.8 1 1.6 1 1.0!)
<taxonomicName id="F7858EF1FFB9B83AFF75FAF1FEBFFD62" box="[167,326,1326,1347]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="ngoclinhense">
<emphasis id="02F12960FFB9B83AFF75FAF1FEBFFD62" box="[167,326,1326,1347]" italics="true" pageId="6" pageNumber="139">L. ngoclinhense</emphasis>
</taxonomicName>
1 3.0 1 0.8 4.6 1 1.8 1 0.4 2.3!)
<taxonomicName id="F7858EF1FFB9B83AFF75FA8AFEBAFD4B" box="[167,323,1365,1386]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="promustache">
<emphasis id="02F12960FFB9B83AFF75FA8AFEBAFD4B" box="[167,323,1365,1386]" italics="true" pageId="6" pageNumber="139">L. promustache</emphasis>
</taxonomicName>
15.5 4.6 12.2 5.6 7.0 11.2 11.6!)
<taxonomicName id="F7858EF1FFB9B83AFF75FAA3FEF1FDB0" box="[167,264,1404,1425]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="pullum">
<emphasis id="02F12960FFB9B83AFF75FAA3FEF1FDB0" box="[167,264,1404,1425]" italics="true" pageId="6" pageNumber="139">L. pullum</emphasis>
</taxonomicName>
14.7 11.6 5.2 12.2 11.4 2.1 3.3 11.0!)
<taxonomicName id="F7858EF1FFB9B83AFF75FA7BFECCFD99" box="[167,309,1443,1465]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="rakhinensis">
<emphasis id="02F12960FFB9B83AFF75FA7BFECCFD99" box="[167,309,1443,1465]" italics="true" pageId="6" pageNumber="139">L. rakhinensis</emphasis>
</taxonomicName>
13.3 14.1 14.1 14.7 15.1 14.3 14.1 15.9 14.5!)
<taxonomicName id="F7858EF1FFB9B83AFF75FA14FEE5FDFE" box="[167,284,1482,1504]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="xanthops">
<emphasis id="02F12960FFB9B83AFF75FA14FEE5FDFE" box="[167,284,1482,1504]" italics="true" pageId="6" pageNumber="139">L. xanthops</emphasis>
</taxonomicName>
13.3 11.0 5.0 11.2 10.6 3.3 2.9 11.2 3.9 14.3!)
<taxonomicName id="F7858EF1FFB9B83AFF75FA2DFEB0FE27" box="[167,329,1521,1543]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="xanthospilum">
<emphasis id="02F12960FFB9B83AFF75FA2DFEB0FE27" box="[167,329,1521,1543]" italics="true" pageId="6" pageNumber="139">L. xanthospilum</emphasis>
</taxonomicName>
13.9 12.8 7.9 13.7 12.4 6.6 5.6 12.6 6.8 15.1 6.4!)
<taxonomicName id="F7858EF1FFB9B83AFF75F9C7FE54FE0C" box="[167,429,1560,1581]" class="Amphibia" family="Megophryidae" genus="Oreolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="rhodostigmatus">
<emphasis id="02F12960FFB9B83AFF75F9C7FE54FE0C" box="[167,429,1560,1581]" italics="true" pageId="6" pageNumber="139">Oreolalax rhodostigmatus</emphasis>
</taxonomicName>
13.3 11.6 11.6 12.6 11.4 11.8 11.0 11.0 11.4 14.9 11.8 13.9!)
<taxonomicName id="F7858EF1FFB9B83AFF75F99FFE7CFE75" box="[167,389,1599,1621]" class="Amphibia" family="Megophryidae" genus="Scutiger" kingdom="Animalia" order="Anura" pageId="6" pageNumber="139" phylum="Chordata" rank="species" species="chintingensis">
<emphasis id="02F12960FFB9B83AFF75F99FFE7CFE75" box="[167,389,1599,1621]" italics="true" pageId="6" pageNumber="139">Scutiger chintingensis</emphasis>
</taxonomicName>
14.5 12.6 12.2 13.7 12.8 12.0 11.6 13.3 12.0 13.9 11.8 14.3 9.9!
</paragraph>
</subSubSection>
<caption id="64FAA5FAFFB8B83BFF45FA21FB30FE70" httpUri="https://zenodo.org/record/256326/files/figure.png" pageId="7" pageNumber="140" targetBox="[151,1436,193,1512]" targetPageId="7">
<paragraph id="303AF572FFB8B83BFF45FA21FB30FE70" blockId="7.[151,1436,1534,1617]" pageId="7" pageNumber="140">
<emphasis id="02F12960FFB8B83BFF45FA21FEEAFE32" bold="true" box="[151,275,1534,1556]" pageId="7" pageNumber="140">FIGURE 3.</emphasis>
Holotype of
<taxonomicName id="F7858EF1FFB8B83BFE4EFA21FD2DFE35" box="[412,724,1534,1556]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="7" pageNumber="140" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB8B83BFE4EFA21FD2DFE35" box="[412,724,1534,1556]" italics="true" pageId="7" pageNumber="140">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB8B83BFD0EFA20FCD4FE35" box="[732,813,1535,1556]" pageId="7" pageNumber="140" rank="species">
<emphasis id="02F12960FFB8B83BFD0EFA20FCD4FE35" bold="true" box="[732,813,1535,1556]" pageId="7" pageNumber="140">sp. nov.</emphasis>
</taxonomicNameLabel>
(SYS a004603) in preservative: (A) dorsal view; (B) ventral view; (C) plantar view of the left foot; (E) posteroventral surface of thigh, note the imperceptible femoral gland; (D) volar view of the right hand; (F) ventral surface of right chest, note the indistinct axillary gland. Photos by J.H. Yang.
</paragraph>
</caption>
<subSubSection id="789FA6F9FFB8B83BFF15F9A6FEA6FE97" pageId="7" pageNumber="140" type="materials_examined">
<paragraph id="303AF572FFB8B83BFF15F9A6FEA6FE97" blockId="7.[151,1436,1657,2006]" pageId="7" pageNumber="140">
<emphasis id="02F12960FFB8B83BFF15F9A6FE42FEB3" bold="true" box="[199,443,1657,1682]" pageId="7" pageNumber="140">Referred specimens.</emphasis>
SYS a004604, seven tadpoles (a004604ag), collection data identical to holotype, not assigned as types.
</paragraph>
</subSubSection>
<subSubSection id="789FA6F9FFB8B83BFF15F91EFE56FEDF" pageId="7" pageNumber="140" type="etymology">
<paragraph id="303AF572FFB8B83BFF15F91EFE56FEDF" blockId="7.[151,1436,1657,2006]" pageId="7" pageNumber="140">
<emphasis id="02F12960FFB8B83BFF15F91EFEB7FEFB" bold="true" box="[199,334,1729,1754]" pageId="7" pageNumber="140">Etymology.</emphasis>
The specific epithet, “
<taxonomicName id="F7858EF1FFB8B83BFD81F91EFD00FEF8" box="[595,761,1729,1753]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="7" pageNumber="140" phylum="Chordata" rank="species" species="tengchongense">
<emphasis id="02F12960FFB8B83BFD81F91EFD00FEF8" box="[595,761,1729,1753]" italics="true" pageId="7" pageNumber="140">tengchongense</emphasis>
</taxonomicName>
”, is in reference to the
<typeStatus id="EF3E4BD0FFB8B83BFBD8F91DFBC3FEFB" box="[1034,1082,1730,1754]" pageId="7" pageNumber="140">type</typeStatus>
locality, Tengchong County of
<collectingRegion id="F2413B90FFB8B83BFF45F939FEA0FEDF" box="[151,345,1765,1790]" country="China" name="Yunnan" pageId="7" pageNumber="140">Yunnan Province</collectingRegion>
,
<collectingCountry id="4892B5E2FFB8B83BFEB7F93AFE55FEDF" box="[357,428,1765,1790]" name="China" pageId="7" pageNumber="140">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="789FA6F9FFB8B834FF15F8D6FF1FF93C" lastPageId="8" lastPageNumber="141" pageId="7" pageNumber="140" type="diagnosis">
<paragraph id="303AF572FFB8B834FF15F8D6FF1FF93C" blockId="7.[151,1436,1657,2006]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="141" pageId="7" pageNumber="140">
<emphasis id="02F12960FFB8B83BFF15F8D6FEB8FF03" bold="true" box="[199,321,1801,1826]" pageId="7" pageNumber="140">Diagnosis.</emphasis>
Assigned to the genus
<taxonomicName id="F7858EF1FFB8B83BFD9DF8D6FD03FF00" box="[591,762,1801,1825]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="7" pageNumber="140" phylum="Chordata" rank="genus">
<emphasis id="02F12960FFB8B83BFD9DF8D6FD03FF00" box="[591,762,1801,1825]" italics="true" pageId="7" pageNumber="140">Leptobrachium</emphasis>
</taxonomicName>
on the basis of having head width larger than tibia length; skin above with a network of ridges; oval and large axillary glands present; extremities of digits rounded; breeding males lacking spines on fingers and breast; and iris bicolored (
<bibRefCitation id="54148883FFB8B83BFC72F88EFB48FF48" author="Dubois" box="[928,1201,1873,1898]" pageId="7" pageNumber="140" refString="Dubois, A. &amp; Ohler, A. (1998) A new species of Leptobrachium (Vibrissaphora) from northern Vietnam, with a review of the taxonomy of the genus Leptobrachium (Pelobatidae, Megophyinae). Dumerilia, 4, 1 - 32." type="journal article" year="1998">Dubois &amp; Ohler 1998</bibRefCitation>
). The new species,
<taxonomicName id="F7858EF1FFB8B83BFF45F8AAFE16FFAC" box="[151,495,1909,1933]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="7" pageNumber="140" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB8B83BFF45F8AAFE16FFAC" box="[151,495,1909,1933]" italics="true" pageId="7" pageNumber="140">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB8B83BFE25F8AAFDA8FFAF" box="[503,593,1909,1934]" pageId="7" pageNumber="140" rank="species">
<emphasis id="02F12960FFB8B83BFE25F8AAFDA8FFAF" bold="true" box="[503,593,1909,1934]" pageId="7" pageNumber="140">sp. nov.</emphasis>
</taxonomicNameLabel>
, can be distinguished from other congeners by a combination of following characters: (1) relatively small size (adult males SVL
<quantity id="F77D5897FFB8B83BFCDFF846FC4EFF90" box="[781,951,1945,1970]" metricMagnitude="-2" metricUnit="m" metricValue="4.66" metricValueMax="5.15" metricValueMin="4.17" pageId="7" pageNumber="140" unit="mm" value="46.6" valueMax="51.5" valueMin="41.7">41.751.5 mm</quantity>
); (2) head width slightly larger than head length; (3) tympanum indistinct; (4) two palmar tubercles oval and distinct, inner one larger than outer one; (5) sexually active males without spines on the upper lip; (6) dorsal skin smooth with distinct network of ridges; (7) dorsum pinkish grey and scattered with irregular black markings; (8) venter dark purplish-gray with numerous small white spots on tubercles, solid white chest; (9) iris bicolored, upper one-third light blue, lower two-third dark brown.
</paragraph>
</subSubSection>
<subSubSection id="789FA6F9FFB7B835FF15FEF8FCDCF8D9" lastPageId="9" lastPageNumber="142" pageId="8" pageNumber="141" type="description">
<paragraph id="303AF572FFB7B834FF15FEF8FD49FAA4" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="141">
<emphasis id="02F12960FFB7B834FF15FEF8FE1DF961" bold="true" box="[199,484,295,320]" pageId="8" pageNumber="141">
Description of
<typeStatus id="EF3E4BD0FFB7B834FEA4FEF8FE26F961" box="[374,479,295,320]" pageId="8" pageNumber="141" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male, SVL
<quantity id="F77D5897FFB7B834FD62FEF8FCEEF91E" box="[688,791,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="4.529999999999999" pageId="8" pageNumber="141" unit="mm" value="45.3">45.3 mm</quantity>
. Habitus moderately stocky; body tapering to groin. Head broad and depressed; head length (HDL
<quantity id="F77D5897FFB7B834FDAEFE93FD1EF945" box="[636,743,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" pageId="8" pageNumber="141" unit="mm" value="18.4">18.4 mm</quantity>
) slightly shorter than head width (HDW
<quantity id="F77D5897FFB7B834FB04FE93FABAF945" box="[1238,1347,332,356]" metricMagnitude="-2" metricUnit="m" metricValue="1.9899999999999998" pageId="8" pageNumber="141" unit="mm" value="19.9">19.9 mm</quantity>
). Snout rounded in dorsal view, sharply sloping in profile, barely projecting beyond lower jaw in profile; nostril slightly closer to eye than to tip of snout, below canthus; internarial distance (IND
<quantity id="F77D5897FFB7B834FBC8FE4BFB8EF98D" box="[1050,1143,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="141" unit="mm" value="4.8">4.8 mm</quantity>
) shorter than interorbital distance (IOD 6.0 mm); canthus rostralis distinct; lores oblique, moderately concave; eye large, slightly projecting from side of head, diameter (EYE
<quantity id="F77D5897FFB7B834FDF2FE03FD80F9D5" box="[544,633,476,501]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="8" pageNumber="141" unit="mm" value="6.7">6.7 mm</quantity>
) shorter than snout length (SNT
<quantity id="F77D5897FFB7B834FC22FE03FBB2F9D5" box="[1008,1099,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="8" pageNumber="141" unit="mm" value="7.8">7.8 mm</quantity>
), interorbital distance slightly greater than upper eyelid width (UEW 5.0 mm); no pineal ocellus; tympanum round, nearly invisible, tympanum diameter (TYM
<quantity id="F77D5897FFB7B834FE9DFDFBFE5EFA1D" box="[335,423,548,573]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="141" unit="mm" value="2.5">2.5 mm</quantity>
) about 37.3% eye diameter and slightly greater than distance between tympanum and eye (TED 2.0 mm); tongue heart-shaped, notched posteriorly; large, slit-like vocal sac openings on floor of mouth near lateral margin of tongue; vomerine teeth absent.
</paragraph>
<paragraph id="303AF572FFB7B834FF15FD50FDC6FBE9" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="141">
Forelimb slender. Fingers moderately slender, without webbing. Tip of fingers blunt, that of fingers I slightly swollen; relative finger lengths II = IV &lt;I &lt;III; two oval palmar tubercles not in contact, inner larger than outer; inner palmar tubercle larger than tip of fingers I; low callous tissue on ventral surface of fingers; nuptial pad absent. Hindlimb slender and relatively short; heels not meeting when limbs are held at right angles to body; tibiotarsal articulation of adpressed limb reaching to the posterior margin of jaws. Toes moderately slender; webbing poorly developed, formula:
<date id="443BD3B2FFB7B834FEADFC9AFE43FB7C" box="[383,442,836,861]" pageId="8" pageNumber="141">I 22</date>
¼
<date id="443BD3B2FFB7B834FE07FC9AFE01FB7C" box="[469,504,836,861]" pageId="8" pageNumber="141">II 1</date>
½3
<date id="443BD3B2FFB7B834FDE6FC9AFD79FB7C" box="[564,640,836,861]" pageId="8" pageNumber="141">III 23</date>
½
<date id="443BD3B2FFB7B834FD4FFC9AFD12FB7D" box="[669,747,836,860]" pageId="8" pageNumber="141">IV 42</date>
V. Tips of toes similar to those of fingers; relative toe lengths I&lt;II&lt;V&lt;III&lt;IV; toes with lateral fringes; subarticular tubercles obscure, but elongate, replaced by low callous tissue; distinct, oval, inner metatarsal tubercle, length (IML 3.0 mm) about 76.9 % distance between tip of toe I and tubercle; no outer metatarsal tubercle.
</paragraph>
<paragraph id="303AF572FFB7B834FF15FC0BFE5EFCE5" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="141">
Skin above smooth with distinct network of ridges, lower flank scattered with small tubercles; no spines on upper lip; low supratympanic ridge from posterior edge of eye to axilla; temporal region granular; tubercles on sacrum absent; sparsely granulated around vent; chin, chest and belly densely covered with white small tubercles; limbs with distinct ridges dorsally and smooth ventrally; a large, round axillary gland (
<quantity id="F77D5897FFB7B834FB8DFBE0FB4FFC76" box="[1119,1206,1087,1112]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="141" unit="mm" value="2.5">2.5 mm</quantity>
in diameter) present on ventrolateral surface slightly posterior to insertion of forelimb with body, slightly visible (
<figureCitation id="A8BEE9F7FFB7B834FB72FBBBFB1BFC5D" box="[1184,1250,1124,1149]" captionStart="FIGURE 3" captionStartId="7.[151,250,1534,1556]" captionTargetBox="[151,1436,193,1512]" captionTargetId="figure@7.[151,1436,193,1513]" captionTargetPageId="7" captionText="FIGURE 3. Holotype of Leptobrachium tengchongense sp. nov. (SYS a 004603) in preservative: (A) dorsal view; (B) ventral view; (C) plantar view of the left foot; (E) posteroventral surface of thigh, note the imperceptible femoral gland; (D) volar view of the right hand; (F) ventral surface of right chest, note the indistinct axillary gland. Photos by J. H. Yang." httpUri="https://zenodo.org/record/256326/files/figure.png" pageId="8" pageNumber="141">Fig. 3</figureCitation>
F); a flat, oblong femoral gland (
<quantity id="F77D5897FFB7B834FE97FB58FE67FC81" box="[325,414,1159,1184]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="141" unit="mm" value="4.5">4.5 mm</quantity>
in length) present on distal half of posteroventral surface of thigh, not in white color and imperceptible (
<figureCitation id="A8BEE9F7FFB7B834FE92FB73FE7AFCE5" box="[320,387,1196,1221]" captionStart="FIGURE 3" captionStartId="7.[151,250,1534,1556]" captionTargetBox="[151,1436,193,1512]" captionTargetId="figure@7.[151,1436,193,1513]" captionTargetPageId="7" captionText="FIGURE 3. Holotype of Leptobrachium tengchongense sp. nov. (SYS a 004603) in preservative: (A) dorsal view; (B) ventral view; (C) plantar view of the left foot; (E) posteroventral surface of thigh, note the imperceptible femoral gland; (D) volar view of the right hand; (F) ventral surface of right chest, note the indistinct axillary gland. Photos by J. H. Yang." httpUri="https://zenodo.org/record/256326/files/figure.png" pageId="8" pageNumber="141">Fig. 3</figureCitation>
D).
</paragraph>
<paragraph id="303AF572FFB7B834FF15FB10FF4EFDE1" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="141">
<emphasis id="02F12960FFB7B834FF15FB10FDD6FCC9" bold="true" box="[199,559,1231,1256]" pageId="8" pageNumber="141">
Color of the
<typeStatus id="EF3E4BD0FFB7B834FEBEFB10FE2BFCC9" box="[364,466,1231,1256]" pageId="8" pageNumber="141" type="holotype">holotype</typeStatus>
in life.
</emphasis>
Dorsum pinkish grey, head slightly brighter, with distinct, black irregular markings; upper flank like dorsum, lower flank like belly, with small white spots on small tubercles; dorsal limbs with distinct wide black bandings; lower two-third of iris dark brown, upper one-third of iris light blue; black streak under canthus and supratympanic fold; a small black spot present on lower anterior corner of eye; ventral surface of body and limbs dark purplish-gray except the chest remarkably white; numerous minute white spots on tubercles on chin, chest, and belly; axillary gland and femoral gland not in white color and imperceptible (
<figureCitation id="A8BEE9F7FFB7B834FAECFA5BFA79FDBD" box="[1342,1408,1412,1437]" captionStart="FIGURE 3" captionStartId="7.[151,250,1534,1556]" captionTargetBox="[151,1436,193,1512]" captionTargetId="figure@7.[151,1436,193,1513]" captionTargetPageId="7" captionText="FIGURE 3. Holotype of Leptobrachium tengchongense sp. nov. (SYS a 004603) in preservative: (A) dorsal view; (B) ventral view; (C) plantar view of the left foot; (E) posteroventral surface of thigh, note the imperceptible femoral gland; (D) volar view of the right hand; (F) ventral surface of right chest, note the indistinct axillary gland. Photos by J. H. Yang." httpUri="https://zenodo.org/record/256326/files/figure.png" pageId="8" pageNumber="141">Fig. 3</figureCitation>
D, F).
</paragraph>
<paragraph id="303AF572FFB7B834FF15FA13FBF7FE29" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="141">
<emphasis id="02F12960FFB7B834FF15FA13FD87FDC4" bold="true" box="[199,638,1484,1509]" pageId="8" pageNumber="141">
Color of the
<typeStatus id="EF3E4BD0FFB7B834FE8CFA13FE3CFDC4" box="[350,453,1484,1509]" pageId="8" pageNumber="141" type="holotype">holotype</typeStatus>
in preservative
</emphasis>
. Color in preservative dark gray-brown, dark markings on dorsum and bandings on limbs still visible. Upper part of iris and scleral arc faded to white.
</paragraph>
<paragraph id="303AF572FFB7B834FF15F9CBFEC9FE9C" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="141">
<emphasis id="02F12960FFB7B834FF15F9CBFEC2FE0C" bold="true" box="[199,315,1556,1581]" pageId="8" pageNumber="141">Variation.</emphasis>
All ten paratypes match the overall characters of the holotype (for measurements of the type series see Table 3). The femoral gland imperceptible in the holotype and other nine paratypes, but is dull white and slightly visible in a single paratype SYS a004627; the axillary gland is distinctly visible in five paratypes (SYS a004623, a004624, a004627, a004630 and a004632), but barely visible or imperceptible in the other five paratypes and holotype.
</paragraph>
<paragraph id="303AF572FFB7B834FF15F918FE86FF09" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="141">
<emphasis id="02F12960FFB7B834FF15F918FE57FEC1" bold="true" box="[199,430,1735,1760]" pageId="8" pageNumber="141">Advertisement call.</emphasis>
A single, pulsed note of approximately
<geoCoordinate id="55B193B5FFB7B834FCBEF917FC56FEC1" box="[876,943,1736,1760]" direction="south" orientation="latitude" pageId="8" pageNumber="141" precision="555" value="-0.13">0.13 s</geoCoordinate>
duration similar to other
<taxonomicName id="F7858EF1FFB7B834FB18F917FA8CFEC1" box="[1226,1397,1736,1760]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="8" pageNumber="141" phylum="Chordata" rank="genus">
<emphasis id="02F12960FFB7B834FB18F917FA8CFEC1" box="[1226,1397,1736,1760]" italics="true" pageId="8" pageNumber="141">Leptobrachium</emphasis>
</taxonomicName>
for which the advertisement call is known. A full description of the advertisement call will be presented elsewhere (Yang
<emphasis id="02F12960FFB7B834FF33F8CEFEE0FF09" box="[225,281,1808,1832]" italics="true" pageId="8" pageNumber="141">et al.</emphasis>
in prep).
</paragraph>
<paragraph id="303AF572FFB7B835FF15F8EBFCDCF8D9" blockId="8.[151,1437,151,2013]" lastBlockId="9.[151,1436,151,248]" lastPageId="9" lastPageNumber="142" pageId="8" pageNumber="141">
<emphasis id="02F12960FFB7B834FF15F8EBFEC1FF6C" bold="true" box="[199,312,1844,1869]" pageId="8" pageNumber="141">Tadpoles.</emphasis>
The following description based on specimen SYS a004604a preserved in 96 % ethanol: stage 40, SVL
<quantity id="F77D5897FFB7B834FF05F888FEBBFF51" box="[215,322,1879,1904]" metricMagnitude="-2" metricUnit="m" metricValue="3.16" pageId="8" pageNumber="141" unit="mm" value="31.6">31.6 mm</quantity>
, total length
<quantity id="F77D5897FFB7B834FE32F887FDB3FF51" box="[480,586,1880,1904]" metricMagnitude="-2" metricUnit="m" metricValue="9.12" pageId="8" pageNumber="141" unit="mm" value="91.2">91.2 mm</quantity>
, tail height
<quantity id="F77D5897FFB7B834FD09F888FCBFFF51" box="[731,838,1879,1904]" metricMagnitude="-2" metricUnit="m" metricValue="1.86" pageId="8" pageNumber="141" unit="mm" value="18.6">18.6 mm</quantity>
; body elliptical in dorsal view, snout rounded; eye moderate, positioned dorsolaterally, not visible in ventral view; spiracle tube short with a free distal end, positioned laterally; tail musculature strong, relatively straight anteriorly and gradually tapering posteriorly, almost reaching the tip; the fin relatively high, tip of tail pointed; oral disk ventral, labial tooth row formula: 1:(5+5)/(4+4). Body and tail light brown, with a wide light V-shaped marking at posterior body, the base of the V slightly extending to the upper fin; dorsal and lateral body scattered with moderate dark brown spots; tail with large dark brown blotches in the proximal half and small dark spots in the distal half.
</paragraph>
</subSubSection>
<caption id="64FAA5FAFFB6B835FF45F88DFD45FF87" httpUri="https://zenodo.org/record/256327/files/figure.png" pageId="9" pageNumber="142" targetBox="[151,1436,301,1851]" targetPageId="9">
<paragraph id="303AF572FFB6B835FF45F88DFD45FF87" blockId="9.[151,1435,1874,1958]" pageId="9" pageNumber="142">
<emphasis id="02F12960FFB6B835FF45F88DFEEDFF46" bold="true" box="[151,276,1874,1896]" pageId="9" pageNumber="142">FIGURE 4.</emphasis>
(A) Holotype of
<taxonomicName id="F7858EF1FFB6B835FE1EF88DFCFCFF49" box="[460,773,1874,1896]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="9" pageNumber="142" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB6B835FE1EF88DFCFCFF49" box="[460,773,1874,1896]" italics="true" pageId="9" pageNumber="142">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB6B835FCDCF88CFC98FF49" box="[782,865,1875,1896]" pageId="9" pageNumber="142" rank="species">
<emphasis id="02F12960FFB6B835FCDCF88CFC98FF49" bold="true" box="[782,865,1875,1896]" pageId="9" pageNumber="142">sp. nov.</emphasis>
</taxonomicNameLabel>
(SYS a004603) in life, inserted picture depicts the iris coloration of holotype in life; (B)
<taxonomicName id="F7858EF1FFB6B835FE2AF8AEFD16FFA6" box="[504,751,1905,1927]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="9" pageNumber="142" phylum="Chordata" rank="species" species="huashen">
<emphasis id="02F12960FFB6B835FE2AF8AEFD16FFA6" box="[504,751,1905,1927]" italics="true" pageId="9" pageNumber="142">Leptobrachium huashen</emphasis>
</taxonomicName>
from Linjiapu substation of Gaoligongshan NNR, inserted picture depicts the iris coloration in life. Photos by J.H. Yang.
</paragraph>
</caption>
<caption id="64FAA5FAFFB5B836FF45F935FBBCFF1C" httpUri="https://zenodo.org/record/256328/files/figure.png" pageId="10" pageNumber="143" targetBox="[151,1436,193,1747]" targetPageId="10">
<paragraph id="303AF572FFB5B836FF45F935FBBCFF1C" blockId="10.[151,1436,1770,1853]" pageId="10" pageNumber="143">
<emphasis id="02F12960FFB5B836FF45F935FEEAFF21" bold="true" box="[151,275,1770,1792]" pageId="10" pageNumber="143">FIGURE 5.</emphasis>
(A) Paratype of
<taxonomicName id="F7858EF1FFB5B836FE6DF935FD0EFF21" box="[447,759,1770,1792]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="10" pageNumber="143" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB5B836FE6DF935FD0EFF21" box="[447,759,1770,1792]" italics="true" pageId="10" pageNumber="143">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB5B836FD2CF934FCB6FF21" box="[766,847,1771,1792]" pageId="10" pageNumber="143" rank="species">
<emphasis id="02F12960FFB5B836FD2CF934FCB6FF21" bold="true" box="[766,847,1771,1792]" pageId="10" pageNumber="143">sp. nov.</emphasis>
</taxonomicNameLabel>
(SYS a004625) in life from Datang Substation of GLGS, inserted picture depicts the microhabitat; (B) paratype of
<taxonomicName id="F7858EF1FFB5B836FD34F8D6FBE7FF3E" box="[742,1054,1801,1823]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="10" pageNumber="143" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB5B836FD34F8D6FBE7FF3E" box="[742,1054,1801,1823]" italics="true" pageId="10" pageNumber="143">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB5B836FBF4F8D6FB81FF3F" box="[1062,1144,1801,1822]" pageId="10" pageNumber="143" rank="species">
<emphasis id="02F12960FFB5B836FBF4F8D6FB81FF3F" bold="true" box="[1062,1144,1801,1822]" pageId="10" pageNumber="143">sp. nov.</emphasis>
</taxonomicNameLabel>
(SYS a004628) in life from Cizhuhe Substation of GLGS, inserted picture depicts the microhabitat. Photos by J.H. Yang.
</paragraph>
</caption>
<subSubSection id="789FA6F9FFB5B830FF15F8B5FA8AFAB0" lastPageId="12" lastPageNumber="145" pageId="10" pageNumber="143" type="distribution">
<paragraph id="303AF572FFB5B837FF15F8B5FD37F8D9" blockId="10.[151,1436,1898,2031]" lastBlockId="11.[151,1437,151,1977]" lastPageId="11" lastPageNumber="144" pageId="10" pageNumber="143">
<emphasis id="02F12960FFB5B836FF15F8B5FDA7FFA2" bold="true" box="[199,606,1898,1923]" pageId="10" pageNumber="143">Distribution and natural history.</emphasis>
<taxonomicName id="F7858EF1FFB5B836FDB8F8B5FC31FFA3" box="[618,968,1898,1922]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="10" pageNumber="143" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB5B836FDB8F8B5FC31FFA3" box="[618,968,1898,1922]" italics="true" pageId="10" pageNumber="143">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB5B836FC06F8B5FBCAFFA2" box="[980,1075,1898,1923]" pageId="10" pageNumber="143" rank="species">
<emphasis id="02F12960FFB5B836FC06F8B5FBCAFFA2" bold="true" box="[980,1075,1898,1923]" pageId="10" pageNumber="143">sp. nov.</emphasis>
</taxonomicNameLabel>
is so far known from Datang substation and Cizhuhe substation of the Tengchong Section of Gaoligongshan National Nature Reserve, Tengchong County,
<collectingRegion id="F2413B90FFB5B836FEAAF86CFDC1FFEA" box="[376,568,1970,1995]" country="China" name="Yunnan" pageId="10" pageNumber="143">Yunnan Province</collectingRegion>
of
<collectingCountry id="4892B5E2FFB5B836FD8EF86DFD5AFFEA" box="[604,675,1970,1995]" name="China" pageId="10" pageNumber="143">China</collectingCountry>
. The new species was found mainly on the riparian forest floor, near shallow, clear-water rocky streams in well-preserved montane evergreen broadleaf forest at elevations between
<quantity id="F77D5897FFB4B837FF45FF48FECCF88E" box="[151,309,151,176]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="11" pageNumber="144" unit="m" value="2250.0" valueMax="2500.0" valueMin="2000.0">20002500 m</quantity>
. Most adult males were found calling with their bodies buried in the moist topsoil and heads hidden under leaf litter, making them difficult to detect (
<figureCitation id="A8BEE9F7FFB4B837FD17FF63FCF1F8F4" box="[709,776,188,213]" captionStart="FIGURE 5" captionStartId="10.[151,250,1770,1792]" captionTargetBox="[151,1436,193,1747]" captionTargetId="figure@10.[151,1436,193,1749]" captionTargetPageId="10" captionText="FIGURE 5. (A) Paratype of Leptobrachium tengchongense sp. nov. (SYS a 004625) in life from Datang Substation of GLGS, inserted picture depicts the microhabitat; (B) paratype of Leptobrachium tengchongense sp. nov. (SYS a 004628) in life from Cizhuhe Substation of GLGS, inserted picture depicts the microhabitat. Photos by J. H. Yang." httpUri="https://zenodo.org/record/256328/files/figure.png" pageId="11" pageNumber="144">Fig. 5</figureCitation>
A). A few males were calling amongst rock crevices (
<figureCitation id="A8BEE9F7FFB4B837FABDFF63FF5FF8D9" captionStart="FIGURE 5" captionStartId="10.[151,250,1770,1792]" captionTargetBox="[151,1436,193,1747]" captionTargetId="figure@10.[151,1436,193,1749]" captionTargetPageId="10" captionText="FIGURE 5. (A) Paratype of Leptobrachium tengchongense sp. nov. (SYS a 004625) in life from Datang Substation of GLGS, inserted picture depicts the microhabitat; (B) paratype of Leptobrachium tengchongense sp. nov. (SYS a 004628) in life from Cizhuhe Substation of GLGS, inserted picture depicts the microhabitat. Photos by J. H. Yang." httpUri="https://zenodo.org/record/256328/files/figure.png" pageId="11" pageNumber="144">Fig. 5</figureCitation>
B). No females were detected during the surveys.
</paragraph>
<paragraph id="303AF572FFB4B837FF15FEDAFAE8FAEC" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="144">
We conducted field surveys in the type localities of the new species, namely Datang and Cizhuhe substations of GLGS, between
<date id="443BD3B2FFB4B837FEBDFEF8FE10F961" box="[367,489,295,320]" pageId="11" pageNumber="144" value="2014-04">April 2014</date>
and
<date id="443BD3B2FFB4B837FDCCFEF8FD53F961" box="[542,682,295,320]" pageId="11" pageNumber="144" value="2016-03">March 2016</date>
. A few males were heard calling during the survey on
<date id="443BD3B2FFB4B837FAD5FEF7FF36F944" pageId="11" pageNumber="144" value="2015-03-13" valueMax="2015-03-14" valueMin="2015-03-13">
1314 March
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</date>
in Datang, and only one calling adult male (holotype) was collected; but during the surveys on 1718/
<date id="443BD3B2FFB4B837FA87FE93FED9F9A9" pageId="11" pageNumber="144" value="2016-03-20" valueMax="2016-03-21" valueMin="2016-03-20">
2021 March
<quantity id="F77D5897FFB4B837FF3AFEB0FED9F9A9" box="[232,288,367,392]" metricMagnitude="1" metricUnit="m" metricValue="5.12064" pageId="11" pageNumber="144" unit="in" value="2016.0">2016</quantity>
</date>
in Datang and Cizhuhe respectively, calling males of the new species were ubiquitous along streams and riparian forests, and ten calling adult males were collected from both localities. In
<date id="443BD3B2FFB4B837FB4FFE4BFAD5F98C" box="[1181,1324,404,429]" pageId="11" pageNumber="144" value="2016-03">March 2016</date>
, montane forests between
<quantity id="F77D5897FFB4B837FE9FFE67FE10F9EE" box="[333,489,440,464]" metricMagnitude="3" metricUnit="m" metricValue="2.55" metricValueMax="2.9" metricValueMin="2.2" pageId="11" pageNumber="144" unit="m" value="2550.0" valueMax="2900.0" valueMin="2200.0">22002900 m</quantity>
a.
<materialsCitation id="80EDFF2FFFB4B837FDD7FE68FAE8FAEC" ID-GBIF-Occurrence="1291547719" collectingDate="2014-04-26" collectingDateMax="2015-05-19" collectingDateMin="2014-04-26" county="Datang" elevation="2355" location="Sympatric" municipality="Cizhuhe" pageId="11" pageNumber="144" specimenCount="1" stateProvince="Yunnan">
s.l. were surveyed in Cizhuhe, but mating calls and individuals of the new species were only detected from the lower elevation band (i.e.
<elevation id="BBA81241FFB4B837FCC1FE03FC09F9D4" box="[787,1008,476,501]" metricMagnitude="3" metricUnit="m" metricValue="2.355" metricValueMax="2.51" metricValueMin="2.2" pageId="11" pageNumber="144" unit="m" value="2355.0" valueMax="2510.0" valueMin="2200.0">
<quantity id="F77D5897FFB4B837FCC1FE03FC49F9D5" box="[787,944,476,501]" metricMagnitude="3" metricUnit="m" metricValue="2.355" metricValueMax="2.51" metricValueMin="2.2" pageId="11" pageNumber="144" unit="m" value="2355.0" valueMax="2510.0" valueMin="2200.0">22002510 m</quantity>
a.s.l.
</elevation>
). No calls and specimens of the new species were detected during the survey on
<collectingDate id="547F2A5AFFB4B837FD54FE20FCB6FA39" box="[646,847,511,536]" pageId="11" pageNumber="144" value="2014-04-26" valueMax="2014-04-27" valueMin="2014-04-26">
<date id="443BD3B2FFB4B837FD54FE20FCB6FA39" box="[646,847,511,536]" pageId="11" pageNumber="144" value="2014-04-26" valueMax="2014-04-27" valueMin="2014-04-26">
2627 April
<quantity id="F77D5897FFB4B837FCC5FDDFFCB6FA39" box="[791,847,512,536]" metricMagnitude="1" metricUnit="m" metricValue="5.11556" pageId="11" pageNumber="144" unit="in" value="2014.0">2014</quantity>
</date>
</collectingDate>
in
<collectingCounty id="D95B8DFEFFB4B837FCA4FDDFFC3EFA39" box="[886,967,512,536]" pageId="11" pageNumber="144">Datang</collectingCounty>
and
<collectingDate id="547F2A5AFFB4B837FC2DFDDFFB38FA39" box="[1023,1217,511,536]" pageId="11" pageNumber="144" value="2015-05-18" valueMax="2015-05-19" valueMin="2015-05-18">
<date id="443BD3B2FFB4B837FC2DFDDFFB38FA39" box="[1023,1217,511,536]" pageId="11" pageNumber="144" value="2015-05-18" valueMax="2015-05-19" valueMin="2015-05-18">
1819 May
<quantity id="F77D5897FFB4B837FB5BFE20FB38FA39" box="[1161,1217,511,536]" metricMagnitude="1" metricUnit="m" metricValue="5.1181" pageId="11" pageNumber="144" unit="in" value="2015.0">2015</quantity>
</date>
</collectingDate>
in
<collectingMunicipality id="D05E6F08FFB4B837FB35FE20FABEFA39" box="[1255,1351,511,536]" pageId="11" pageNumber="144">Cizhuhe</collectingMunicipality>
, except the tadpoles that were found in the stream in both localities in all surveys.
<location id="355AA3A9FFB4B837FC0DFDFBFBABFA1C" LSID="urn:lsid:plazi:treatment:B82C4464FFBCB830FF45F968F8C6FE0F:355AA3A9FFB4B837FC0DFDFBFBABFA1C" box="[991,1106,548,573]" county="Datang" municipality="Cizhuhe" name="Sympatric" pageId="11" pageNumber="144" stateProvince="Yunnan">Sympatric</location>
herpetofauna recorded in the locality during the survey included the
<location id="355AA3A9FFB4B837FD8FFD98FD66FA41" LSID="urn:lsid:plazi:treatment:B82C4464FFBCB830FF45F968F8C6FE0F:355AA3A9FFB4B837FD8FFD98FD66FA41" box="[605,671,583,608]" county="Datang" municipality="Cizhuhe" name="Spiny" pageId="11" pageNumber="144" stateProvince="Yunnan">Spiny</location>
stream toad
<taxonomicName id="F7858EF1FFB4B837FCE4FD97FB0BFA41" authority="Yang, Liu &amp; Rao" authorityName="Yang, Liu &amp; Rao" box="[822,1266,583,608]" class="Amphibia" family="Bufonidae" genus="Bufo" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="tuberospinius">
<emphasis id="02F12960FFB4B837FCE4FD97FBF2FA41" box="[822,1035,584,608]" italics="true" pageId="11" pageNumber="144">Bufo tuberospinius</emphasis>
(Yang, Liu &amp; Rao)
</taxonomicName>
,
<collectingRegion id="F2413B90FFB4B837FAD2FD97FAAFFA7E" box="[1280,1366,584,607]" country="China" name="Yunnan" pageId="11" pageNumber="144">Yunnan</collectingRegion>
spiny frog
<taxonomicName id="F7858EF1FFB4B837FF07FDB2FD8AFAA4" authority="Anderson" authorityName="Anderson" box="[213,627,620,645]" class="Amphibia" family="Dicroglossidae" genus="Nanorana" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="yunnanensis">
<emphasis id="02F12960FFB4B837FF07FDB2FE19FAA5" box="[213,480,621,644]" italics="true" pageId="11" pageNumber="144">Nanorana yunnanensis</emphasis>
(Anderson)
</taxonomicName>
,
<location id="355AA3A9FFB4B837FD54FDB3FD0FFAA4" LSID="urn:lsid:plazi:treatment:B82C4464FFBCB830FF45F968F8C6FE0F:355AA3A9FFB4B837FD54FDB3FD0FFAA4" box="[646,758,620,645]" county="Datang" municipality="Cizhuhe" name="Glandular" pageId="11" pageNumber="144" stateProvince="Yunnan">Glandular</location>
horned toad
<taxonomicName id="F7858EF1FFB4B837FC71FDB3FA61FAA4" authority="Fei, Ye &amp; Huang" authorityName="Fei, Ye &amp; Huang" box="[931,1432,620,645]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="glandulosa">
<emphasis id="02F12960FFB4B837FC71FDB3FB57FAA5" box="[931,1198,620,644]" italics="true" pageId="11" pageNumber="144">Megophrys glandulosa</emphasis>
Fei, Ye &amp; Huang
</taxonomicName>
,
<location id="355AA3A9FFB4B837FF45FD50FEEDFA89" LSID="urn:lsid:plazi:treatment:B82C4464FFBCB830FF45F968F8C6FE0F:355AA3A9FFB4B837FF45FD50FEEDFA89" box="[151,276,655,680]" county="Datang" municipality="Cizhuhe" name="Anderson's" pageId="11" pageNumber="144" stateProvince="Yunnan">Anderson's</location>
frog
<taxonomicName id="F7858EF1FFB4B837FE86FD4FFD21FA89" authority="Boulenger" authorityName="Boulenger" box="[340,728,655,680]" class="Amphibia" family="Ranidae" genus="Odorrana" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="andersonii">
<emphasis id="02F12960FFB4B837FE86FD4FFDBCFA89" box="[340,581,656,680]" italics="true" pageId="11" pageNumber="144">Odorrana andersonii</emphasis>
(Boulenger)
</taxonomicName>
,
<location id="355AA3A9FFB4B837FD36FD50FCA5FA89" LSID="urn:lsid:plazi:treatment:B82C4464FFBCB830FF45F968F8C6FE0F:355AA3A9FFB4B837FD36FD50FCA5FA89" box="[740,860,655,680]" county="Datang" municipality="Cizhuhe" name="Chaochiao" pageId="11" pageNumber="144" stateProvince="Yunnan">Chaochiao</location>
brown frog
<taxonomicName id="F7858EF1FFB4B837FC3FFD4EFAE9FA89" authority="Liu" authorityName="Liu" box="[1005,1296,655,680]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="chaochiaoensis">
<emphasis id="02F12960FFB4B837FC3FFD4EFB27FA89" box="[1005,1246,656,680]" italics="true" pageId="11" pageNumber="144">Rana chaochiaoensis</emphasis>
Liu
</taxonomicName>
,
<location id="355AA3A9FFB4B837FACFFD50FA65FA89" LSID="urn:lsid:plazi:treatment:B82C4464FFBCB830FF45F968F8C6FE0F:355AA3A9FFB4B837FACFFD50FA65FA89" box="[1309,1436,655,680]" county="Datang" municipality="Cizhuhe" name="Large-eyed" pageId="11" pageNumber="144" stateProvince="Yunnan">Large-eyed</location>
bamboo snake
<taxonomicName id="F7858EF1FFB4B837FEEDFD6BFD48FAEC" authority="Blyth" authorityName="Blyth" box="[319,689,692,717]" class="Reptilia" family="Colubridae" genus="Pseudoxenodon" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="macrops">
<emphasis id="02F12960FFB4B837FEEDFD6BFDAFFAED" box="[319,598,692,716]" italics="true" pageId="11" pageNumber="144">Pseudoxenodon macrops</emphasis>
(Blyth)
</taxonomicName>
, and
<location id="355AA3A9FFB4B837FD3EFD6BFC33FAEC" LSID="urn:lsid:plazi:treatment:B82C4464FFBCB830FF45F968F8C6FE0F:355AA3A9FFB4B837FD3EFD6BFC33FAEC" box="[748,970,692,717]" county="Datang" municipality="Cizhuhe" name="Leonard's Keelback" pageId="11" pageNumber="144" stateProvince="Yunnan">Leonard's Keelback</location>
<taxonomicName id="F7858EF1FFB4B837FC03FD6BFAF4FAEC" authority="Wall" authorityName="Wall" box="[977,1293,692,717]" class="Reptilia" family="Colubridae" genus="Rhabdophis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="leonardi">
<emphasis id="02F12960FFB4B837FC03FD6BFB42FAED" box="[977,1211,692,716]" italics="true" pageId="11" pageNumber="144">Rhabdophis leonardi</emphasis>
(Wall)
</taxonomicName>
.
</materialsCitation>
</paragraph>
<paragraph id="303AF572FFB4B837FF15FD08FAC9FBCC" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="144">
<emphasis id="02F12960FFB4B837FF15FD08FE9BFAD1" bold="true" box="[199,354,727,752]" pageId="11" pageNumber="144">Comparison.</emphasis>
We compared
<taxonomicName id="F7858EF1FFB4B837FDF0FD07FC7BFAD1" box="[546,898,728,752]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB4B837FDF0FD07FC7BFAD1" box="[546,898,728,752]" italics="true" pageId="11" pageNumber="144">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB4B837FC43FD08FC0AFAD1" box="[913,1011,727,752]" pageId="11" pageNumber="144" rank="species">
<emphasis id="02F12960FFB4B837FC43FD08FC0AFAD1" bold="true" box="[913,1011,727,752]" pageId="11" pageNumber="144">sp. nov.</emphasis>
</taxonomicNameLabel>
to all other recognized species of
<taxonomicName id="F7858EF1FFB4B837FF45FD23FEBAFB35" box="[151,323,764,788]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="genus">
<emphasis id="02F12960FFB4B837FF45FD23FEBAFB35" box="[151,323,764,788]" italics="true" pageId="11" pageNumber="144">Leptobrachium</emphasis>
</taxonomicName>
from
<collectingCountry id="4892B5E2FFB4B837FE55FD23FE33FB34" box="[391,458,764,789]" name="China" pageId="11" pageNumber="144">China</collectingCountry>
and Indochina, all of which formed a monophyletic clade in molecular studies (
<bibRefCitation id="54148883FFB4B837FA9CFD23FEEAFB19" author="Brown" pageId="11" pageNumber="144" refString="Brown, R. M., Siler, C. D., Diesmos, A. C. &amp; Alcala, A. C. (2009) Philippine frogs of the genus Leptobrachium (Anura; Megophryidae): phylogeny-based species delimitation, taxonomic review, and descriptions of three new species. Herpetological Monographs, 23 (1), 1 - 44. http: // dx. doi. org / 10.1655 / 09 - 037.1" type="journal article" year="2009">
Brown
<emphasis id="02F12960FFB4B837FF45FCFEFF29FB19" box="[151,208,800,824]" italics="true" pageId="11" pageNumber="144">et al.</emphasis>
2009
</bibRefCitation>
; Matsui
<emphasis id="02F12960FFB4B837FEA1FCFEFE55FB19" box="[371,428,800,824]" italics="true" pageId="11" pageNumber="144">et al.</emphasis>
2011; this study): i.e.,
<taxonomicName id="F7858EF1FFB4B837FD62FCFEFCB2FB19" box="[688,843,800,824]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="ailaonicum">
<emphasis id="02F12960FFB4B837FD62FCFEFCB2FB19" box="[688,843,800,824]" italics="true" pageId="11" pageNumber="144">L. ailaonicum</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FC88FCFEFC45FB19" box="[858,956,800,824]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="banae">
<emphasis id="02F12960FFB4B837FC88FCFEFC45FB19" box="[858,956,800,824]" italics="true" pageId="11" pageNumber="144">L. banae</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FC19FCFEFBBAFB19" box="[971,1091,800,824]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="boringii">
<emphasis id="02F12960FFB4B837FC19FCFEFBBAFB19" box="[971,1091,800,824]" italics="true" pageId="11" pageNumber="144">L. boringii</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FB80FCFEFB2DFB19" box="[1106,1236,800,824]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="buchardi">
<emphasis id="02F12960FFB4B837FB80FCFEFB2DFB19" box="[1106,1236,800,824]" italics="true" pageId="11" pageNumber="144">L. buchardi</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FB31FCFEFA8EFB19" box="[1251,1399,800,824]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="chapaense">
<emphasis id="02F12960FFB4B837FB31FCFEFA8EFB19" box="[1251,1399,800,824]" italics="true" pageId="11" pageNumber="144">L. chapaense</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FA57FCFEFEDAFB7D" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="guangxiense">
<emphasis id="02F12960FFB4B837FA57FCFEFEDAFB7D" italics="true" pageId="11" pageNumber="144">L. guangxiense</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FEE4FC9AFE21FB7D" box="[310,472,836,860]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="hainanense">
<emphasis id="02F12960FFB4B837FEE4FC9AFE21FB7D" box="[310,472,836,860]" italics="true" pageId="11" pageNumber="144">L. hainanense</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FE39FC9AFD93FB7D" box="[491,618,836,860]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="huashen">
<emphasis id="02F12960FFB4B837FE39FC9AFD93FB7D" box="[491,618,836,860]" italics="true" pageId="11" pageNumber="144">L. huashen</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FDACFC9AFCDDFB7D" box="[638,804,836,860]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="leishanense">
<emphasis id="02F12960FFB4B837FDACFC9AFCDDFB7D" box="[638,804,836,860]" italics="true" pageId="11" pageNumber="144">L. leishanense</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FCE4FC9AFC57FB7D" box="[822,942,836,860]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="leucops">
<emphasis id="02F12960FFB4B837FCE4FC9AFC57FB7D" box="[822,942,836,860]" italics="true" pageId="11" pageNumber="144">L. leucops</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FC13FC9AFBF0FB7D" box="[961,1033,836,860]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="liui">
<emphasis id="02F12960FFB4B837FC13FC9AFBF0FB7D" box="[961,1033,836,860]" italics="true" pageId="11" pageNumber="144">L. liui</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FBCEFC9AFB18FB7D" box="[1052,1249,836,860]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="masatakasatoi">
<emphasis id="02F12960FFB4B837FBCEFC9AFB18FB7D" box="[1052,1249,836,860]" italics="true" pageId="11" pageNumber="144">L. masatakasatoi</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FB26FC9AFA8BFB7D" box="[1268,1394,836,860]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="mouhoti">
<emphasis id="02F12960FFB4B837FB26FC9AFA8BFB7D" box="[1268,1394,836,860]" italics="true" pageId="11" pageNumber="144">L. mouhoti</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FA57FC9AFED2FBA1" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="ngoclinhense">
<emphasis id="02F12960FFB4B837FA57FC9AFED2FBA1" italics="true" pageId="11" pageNumber="144">L. ngoclinhense</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FEE8FCB6FE10FBA1" box="[314,489,872,896]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="promustache">
<emphasis id="02F12960FFB4B837FEE8FCB6FE10FBA1" box="[314,489,872,896]" italics="true" pageId="11" pageNumber="144">L. promustache</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FE2BFCB6FD9EFBA1" box="[505,615,872,896]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="pullum">
<emphasis id="02F12960FFB4B837FE2BFCB6FD9EFBA1" box="[505,615,872,896]" italics="true" pageId="11" pageNumber="144">L. pullum</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FDA4FCB6FD02FBA1" box="[630,763,872,896]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="xanthops">
<emphasis id="02F12960FFB4B837FDA4FCB6FD02FBA1" box="[630,763,872,896]" italics="true" pageId="11" pageNumber="144">L. xanthops</emphasis>
</taxonomicName>
and
<taxonomicName id="F7858EF1FFB4B837FCE7FCB6FC12FBA1" box="[821,1003,872,896]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="xanthospilum">
<emphasis id="02F12960FFB4B837FCE7FCB6FC12FBA1" box="[821,1003,872,896]" italics="true" pageId="11" pageNumber="144">L. xanthospilum</emphasis>
</taxonomicName>
. We also compared
<taxonomicName id="F7858EF1FFB4B837FB04FCB6FA62FBA1" box="[1238,1435,872,896]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB4B837FB04FCB6FA62FBA1" box="[1238,1435,872,896]" italics="true" pageId="11" pageNumber="144">L. tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB4B837FF45FC53FF0BFB84" box="[151,242,908,933]" pageId="11" pageNumber="144" rank="species">
<emphasis id="02F12960FFB4B837FF45FC53FF0BFB84" bold="true" box="[151,242,908,933]" pageId="11" pageNumber="144">sp. nov.</emphasis>
</taxonomicNameLabel>
with the newly described
<taxonomicName id="F7858EF1FFB4B837FDF0FC52FD74FB85" box="[546,653,908,932]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="bompu">
<emphasis id="02F12960FFB4B837FDF0FC52FD74FB85" box="[546,653,908,932]" italics="true" pageId="11" pageNumber="144">L. bompu</emphasis>
</taxonomicName>
from
<collectingCountry id="4892B5E2FFB4B837FD07FC53FCF6FB84" box="[725,783,908,933]" name="India" pageId="11" pageNumber="144">India</collectingCountry>
of which comparative 16S mtDNA data was unavailable from the GenBank. Unrelated Sunderland/
<collectingCountry id="4892B5E2FFB4B837FDABFC70FD25FBE9" box="[633,732,943,968]" name="Thailand" pageId="11" pageNumber="144">Thailand</collectingCountry>
clade taxa (occur in southern
<collectingCountry id="4892B5E2FFB4B837FBE5FC6FFB5FFBE9" box="[1079,1190,944,968]" name="Myanmar" pageId="11" pageNumber="144">Myanmar</collectingCountry>
,
<collectingCountry id="4892B5E2FFB4B837FB61FC70FAE0FBE9" box="[1203,1305,943,968]" name="Thailand" pageId="11" pageNumber="144">Thailand</collectingCountry>
, Peninsular
<collectingCountry id="4892B5E2FFB4B837FF45FC0BFEF8FBCC" box="[151,257,980,1005]" name="Malaysia" pageId="11" pageNumber="144">Malaysia</collectingCountry>
,
<collectingRegion id="F2413B90FFB4B837FEDEFC0BFE95FBCD" box="[268,364,980,1004]" country="Indonesia" name="Sumatera Utara" pageId="11" pageNumber="144">Sumatra</collectingRegion>
,
<collectingRegion id="F2413B90FFB4B837FEA5FC0AFE52FBCD" box="[375,427,981,1004]" country="Indonesia" name="Jawa Timur" pageId="11" pageNumber="144">Java</collectingRegion>
,
<collectingRegion id="F2413B90FFB4B837FE64FC0BFE1EFBCC" box="[438,487,980,1005]" country="Indonesia" name="Bali" pageId="11" pageNumber="144">Bali</collectingRegion>
, Borneo, and the
<collectingCountry id="4892B5E2FFB4B837FD7EFC0BFCD5FBCC" box="[684,812,980,1005]" name="Philippines" pageId="11" pageNumber="144">Philippines</collectingCountry>
) are omitted from comparisons for simplicity.
</paragraph>
<paragraph id="303AF572FFB4B837FF15FC27FF00FEB9" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="144">
By the absence of spines on the upper lip in sexually active males,
<taxonomicName id="F7858EF1FFB4B837FC08FC27FACFFC31" box="[986,1334,1016,1040]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB4B837FC08FC27FACFFC31" box="[986,1334,1016,1040]" italics="true" pageId="11" pageNumber="144">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB4B837FAEDFC28FA65FC31" box="[1343,1436,1015,1040]" pageId="11" pageNumber="144" rank="species">
<emphasis id="02F12960FFB4B837FAEDFC28FA65FC31" bold="true" box="[1343,1436,1015,1040]" pageId="11" pageNumber="144">sp. nov.</emphasis>
</taxonomicNameLabel>
distinctly differs from six other congeners, i.e.,
<emphasis id="02F12960FFB4B837FD60FBC2FB23FC15" box="[690,1242,1052,1076]" italics="true" pageId="11" pageNumber="144">
<taxonomicName id="F7858EF1FFB4B837FD60FBC2FCABFC15" box="[690,850,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="ailaonicum">L. ailaonicum</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FC8CFBC2FC23FC15" box="[862,986,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="boringii">L. boringii</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FC34FBC2FB73FC15" box="[998,1162,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="leishanense">L. leishanense</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FB44FBC2FB23FC15" box="[1174,1242,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="liui">L. liui</taxonomicName>
</emphasis>
,
<taxonomicName id="F7858EF1FFB4B837FB38FBC2FA65FC15" box="[1258,1436,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="ngoclinhense">
<emphasis id="02F12960FFB4B837FB38FBC2FA65FC15" box="[1258,1436,1052,1076]" italics="true" pageId="11" pageNumber="144">L. ngoclinhense</emphasis>
</taxonomicName>
and
<taxonomicName id="F7858EF1FFB4B837FF18FB9EFE85FC79" box="[202,380,1088,1112]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="promustache">
<emphasis id="02F12960FFB4B837FF18FB9EFE85FC79" box="[202,380,1088,1112]" italics="true" pageId="11" pageNumber="144">L. promustache</emphasis>
</taxonomicName>
, all of which process spines on the upper lip in sexually active males and were formerly referred to the genus
<taxonomicName id="F7858EF1FFB4B837FE55FBBBFDDCFC5D" box="[391,549,1124,1148]" class="Amphibia" family="Megophryidae" genus="Vibrissaphora" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="genus">
<emphasis id="02F12960FFB4B837FE55FBBBFDDCFC5D" box="[391,549,1124,1148]" italics="true" pageId="11" pageNumber="144">Vibrissaphora</emphasis>
</taxonomicName>
(
<bibRefCitation id="54148883FFB4B837FDE7FBBBFD11FC5C" author="Rao" box="[565,744,1124,1149]" pageId="11" pageNumber="144" refString="Rao, D. Q., Wilkinson, J. A. &amp; Zhang, M. W. (2006) A new species of the genus Vibrissaphora (Anura: Megophryidae) from Yunnan Province, China. Herpetologica, 62 (1), 90 - 95. http: // dx. doi. org / 10.1655 / 05 - 05.1" type="journal article" year="2006">
Rao
<emphasis id="02F12960FFB4B837FDB9FBBAFD5DFC5D" box="[619,676,1124,1148]" italics="true" pageId="11" pageNumber="144">et al.</emphasis>
2006
</bibRefCitation>
;
<bibRefCitation id="54148883FFB4B837FD21FBBBFC63FC5C" author="Fei" box="[755,922,1124,1149]" pageId="11" pageNumber="144" refString="Fei, L., Ye, C. Y., Huang, Y. Z., Jiang, J. P. &amp; Xie, F. (2005) An illustrated key to Chinese amphibians. Sichuan Publishing House of Science and Technology, Chongqing, 340 pp. [In Chinese]" type="book" year="2005">
Fei
<emphasis id="02F12960FFB4B837FCCCFBBAFCA1FC5D" box="[798,856,1124,1148]" italics="true" pageId="11" pageNumber="144">et al.</emphasis>
2005
</bibRefCitation>
,
<bibRefCitation id="54148883FFB4B837FC77FBBBFC1BFC5D" author="Fei" box="[933,994,1124,1148]" pageId="11" pageNumber="144" refString="Fei, L., Hu, S. Q., Ye, C. Y. &amp; Huang, Y. Z. (2009) Fauna Sinica. Amphibia Vol. 2 Anura. Science Press, Beijing, 957 pp. [In Chinese]" type="book" year="2009">2009</bibRefCitation>
). By having a bicolored iris with upper one-third light blue,
<taxonomicName id="F7858EF1FFB4B837FE56FB57FD26FC81" box="[388,735,1160,1184]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB4B837FE56FB57FD26FC81" box="[388,735,1160,1184]" italics="true" pageId="11" pageNumber="144">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB4B837FD3AFB58FCBDFC81" box="[744,836,1159,1184]" pageId="11" pageNumber="144" rank="species">
<emphasis id="02F12960FFB4B837FD3AFB58FCBDFC81" bold="true" box="[744,836,1159,1184]" pageId="11" pageNumber="144">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="F7858EF1FFB4B837FB52FB56FB15FC81" box="[1152,1260,1160,1184]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="bompu">
<emphasis id="02F12960FFB4B837FB52FB56FB15FC81" box="[1152,1260,1160,1184]" italics="true" pageId="11" pageNumber="144">L. bompu</emphasis>
</taxonomicName>
(iris uniformly grey-blue),
<taxonomicName id="F7858EF1FFB4B837FECCFB72FE4FFCE5" box="[286,438,1196,1220]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="chapaense">
<emphasis id="02F12960FFB4B837FECCFB72FE4FFCE5" box="[286,438,1196,1220]" italics="true" pageId="11" pageNumber="144">L. chapaense</emphasis>
</taxonomicName>
(iris uniformly dark brown),
<taxonomicName id="F7858EF1FFB4B837FCC5FB72FC6EFCE5" box="[791,919,1196,1220]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="huashen">
<emphasis id="02F12960FFB4B837FCC5FB72FC6EFCE5" box="[791,919,1196,1220]" italics="true" pageId="11" pageNumber="144">L. huashen</emphasis>
</taxonomicName>
(iris uniformly dark brown, [see discussion below]),
<taxonomicName id="F7858EF1FFB4B837FF29FB0EFE62FCC9" box="[251,411,1232,1256]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="leishanense">
<emphasis id="02F12960FFB4B837FF29FB0EFE62FCC9" box="[251,411,1232,1256]" italics="true" pageId="11" pageNumber="144">L. leishanense</emphasis>
</taxonomicName>
(upper part of iris light green),
<taxonomicName id="F7858EF1FFB4B837FD2BFB0EFC92FCC9" box="[761,875,1232,1256]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="leucops">
<emphasis id="02F12960FFB4B837FD2BFB0EFC92FCC9" box="[761,875,1232,1256]" italics="true" pageId="11" pageNumber="144">L. leucops</emphasis>
</taxonomicName>
(upper part of iris white),
<taxonomicName id="F7858EF1FFB4B837FB42FB0EFB2AFCC9" box="[1168,1235,1232,1256]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="liui">
<emphasis id="02F12960FFB4B837FB42FB0EFB2AFCC9" box="[1168,1235,1232,1256]" italics="true" pageId="11" pageNumber="144">L. liui</emphasis>
</taxonomicName>
(upper part of iris light green),
<taxonomicName id="F7858EF1FFB4B837FEE6FB2AFE4CFD2D" box="[308,437,1268,1292]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="mouhoti">
<emphasis id="02F12960FFB4B837FEE6FB2AFE4CFD2D" box="[308,437,1268,1292]" italics="true" pageId="11" pageNumber="144">L. mouhoti</emphasis>
</taxonomicName>
(iris black with an orange-yellow crescent on outer margin),
<taxonomicName id="F7858EF1FFB4B837FB7BFB2AFA98FD2D" box="[1193,1377,1268,1292]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="ngoclinhense">
<emphasis id="02F12960FFB4B837FB7BFB2AFA98FD2D" box="[1193,1377,1268,1292]" italics="true" pageId="11" pageNumber="144">L. ngoclinhense</emphasis>
</taxonomicName>
(iris uniformly dark brown),
<taxonomicName id="F7858EF1FFB4B837FE63FAC6FDD9FD11" box="[433,544,1304,1328]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="pullum">
<emphasis id="02F12960FFB4B837FE63FAC6FDD9FD11" box="[433,544,1304,1328]" italics="true" pageId="11" pageNumber="144">L. pullum</emphasis>
</taxonomicName>
(upper part of iris red),
<taxonomicName id="F7858EF1FFB4B837FC97FAC6FC32FD11" box="[837,971,1304,1328]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="xanthops">
<emphasis id="02F12960FFB4B837FC97FAC6FC32FD11" box="[837,971,1304,1328]" italics="true" pageId="11" pageNumber="144">L. xanthops</emphasis>
</taxonomicName>
(upper part of iris pale yellow), and
<taxonomicName id="F7858EF1FFB4B837FA57FAC6FED7FD75" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="xanthospilum">
<emphasis id="02F12960FFB4B837FA57FAC6FED7FD75" italics="true" pageId="11" pageNumber="144">L. xanthospilum</emphasis>
</taxonomicName>
(upper half of iris white). By having a relatively smaller body size (SVL
<quantity id="F77D5897FFB4B837FB46FAE3FAC3FD75" box="[1172,1338,1340,1365]" metricMagnitude="-2" metricUnit="m" metricValue="4.66" metricValueMax="5.15" metricValueMin="4.17" pageId="11" pageNumber="144" unit="mm" value="46.6" valueMax="51.5" valueMin="41.7">41.751.5 mm</quantity>
in adult males),
<taxonomicName id="F7858EF1FFB4B837FF22FABFFDB1FD59" box="[240,584,1376,1400]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB4B837FF22FABFFDB1FD59" box="[240,584,1376,1400]" italics="true" pageId="11" pageNumber="144">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB4B837FD82FA80FD53FD59" box="[592,682,1375,1400]" pageId="11" pageNumber="144" rank="species">
<emphasis id="02F12960FFB4B837FD82FA80FD53FD59" bold="true" box="[592,682,1375,1400]" pageId="11" pageNumber="144">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from the larger
<taxonomicName id="F7858EF1FFB4B837FB9EFABEFB11FD59" box="[1100,1256,1376,1400]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="ailaonicum">
<emphasis id="02F12960FFB4B837FB9EFABEFB11FD59" box="[1100,1256,1376,1400]" italics="true" pageId="11" pageNumber="144">L. ailaonicum</emphasis>
</taxonomicName>
(
<quantity id="F77D5897FFB4B837FB25FA80FA65FD56" box="[1271,1436,1375,1400]" metricMagnitude="-2" metricUnit="m" metricValue="7.805" metricValueMax="8.76" metricValueMin="6.85" pageId="11" pageNumber="144" unit="mm" value="78.05" valueMax="87.6" valueMin="68.5">68.587.6 mm</quantity>
in males),
<taxonomicName id="F7858EF1FFB4B837FEDEFA5AFE97FDBD" box="[268,366,1412,1436]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="banae">
<emphasis id="02F12960FFB4B837FEDEFA5AFE97FDBD" box="[268,366,1412,1436]" italics="true" pageId="11" pageNumber="144">L. banae</emphasis>
</taxonomicName>
(57.270.0 mm in males),
<taxonomicName id="F7858EF1FFB4B837FD49FA5AFCEBFDBD" box="[667,786,1412,1436]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="boringii">
<emphasis id="02F12960FFB4B837FD49FA5AFCEBFDBD" box="[667,786,1412,1436]" italics="true" pageId="11" pageNumber="144">L. boringii</emphasis>
</taxonomicName>
(69.889.0 mm in males),
<taxonomicName id="F7858EF1FFB4B837FB92FA5AFB13FDBD" box="[1088,1258,1413,1436]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="guangxiense">
<emphasis id="02F12960FFB4B837FB92FA5AFB13FDBD" box="[1088,1258,1413,1436]" italics="true" pageId="11" pageNumber="144">L. guangxiense</emphasis>
</taxonomicName>
(
<quantity id="F77D5897FFB4B837FB2AFA5BFA65FDBD" box="[1272,1436,1412,1437]" metricMagnitude="-2" metricUnit="m" metricValue="5.58" metricValueMax="5.81" metricValueMin="5.35" pageId="11" pageNumber="144" unit="mm" value="55.8" valueMax="58.1" valueMin="53.5">53.558.1 mm</quantity>
in males),
<taxonomicName id="F7858EF1FFB4B837FEDEFA76FE55FDE1" box="[268,428,1448,1472]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="leishanense">
<emphasis id="02F12960FFB4B837FEDEFA76FE55FDE1" box="[268,428,1448,1472]" italics="true" pageId="11" pageNumber="144">L. leishanense</emphasis>
</taxonomicName>
(69.0
<quantity id="F77D5897FFB4B837FE29FA78FDA6FD9E" box="[507,607,1447,1472]" metricMagnitude="-2" metricUnit="m" metricValue="9.57" pageId="11" pageNumber="144" unit="mm" value="95.7">95.7 mm</quantity>
in males),
<taxonomicName id="F7858EF1FFB4B837FD09FA76FCE4FDE1" box="[731,797,1448,1472]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="liui">
<emphasis id="02F12960FFB4B837FD09FA76FCE4FDE1" box="[731,797,1448,1472]" italics="true" pageId="11" pageNumber="144">L. liui</emphasis>
</taxonomicName>
(
<quantity id="F77D5897FFB4B837FCFEFA78FC28FD9E" box="[812,977,1447,1472]" metricMagnitude="-2" metricUnit="m" metricValue="7.915" metricValueMax="9.05" metricValueMin="6.779999999999999" pageId="11" pageNumber="144" unit="mm" value="79.15" valueMax="90.5" valueMin="67.8">67.890.5 mm</quantity>
in males),
<taxonomicName id="F7858EF1FFB4B837FB9EFA76FAF5FDE1" box="[1100,1292,1448,1472]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="masatakasatoi">
<emphasis id="02F12960FFB4B837FB9EFA76FAF5FDE1" box="[1100,1292,1448,1472]" italics="true" pageId="11" pageNumber="144">L. masatakasatoi</emphasis>
</taxonomicName>
(
<quantity id="F77D5897FFB4B837FAC8FA78FA79FD9E" box="[1306,1408,1447,1472]" metricMagnitude="-2" metricUnit="m" metricValue="5.52" pageId="11" pageNumber="144" unit="mm" value="55.2">55.2 mm</quantity>
in male),
<taxonomicName id="F7858EF1FFB4B837FF37FA12FE61FDC5" box="[229,408,1484,1508]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="ngoclinhense">
<emphasis id="02F12960FFB4B837FF37FA12FE61FDC5" box="[229,408,1484,1508]" italics="true" pageId="11" pageNumber="144">L. ngoclinhense</emphasis>
</taxonomicName>
(
<quantity id="F77D5897FFB4B837FE75FA13FDF4FDC5" box="[423,525,1484,1508]" metricMagnitude="-2" metricUnit="m" metricValue="7.709999999999999" pageId="11" pageNumber="144" unit="mm" value="77.1">77.1 mm</quantity>
in male) and
<taxonomicName id="F7858EF1FFB4B837FD7BFA12FCA7FDC5" box="[681,862,1484,1508]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="xanthospilum">
<emphasis id="02F12960FFB4B837FD7BFA12FCA7FDC5" box="[681,862,1484,1508]" italics="true" pageId="11" pageNumber="144">L. xanthospilum</emphasis>
</taxonomicName>
(
<quantity id="F77D5897FFB4B837FCBFFA13FBEAFDC5" box="[877,1043,1484,1509]" metricMagnitude="-2" metricUnit="m" metricValue="6.81" metricValueMax="7.34" metricValueMin="6.279999999999999" pageId="11" pageNumber="144" unit="mm" value="68.1" valueMax="73.4" valueMin="62.8">62.873.4 mm</quantity>
in males). By having inner palmar tubercle larger than outer one,
<taxonomicName id="F7858EF1FFB4B837FE2AFA2FFCAAFE29" box="[504,851,1520,1544]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB4B837FE2AFA2FFCAAFE29" box="[504,851,1520,1544]" italics="true" pageId="11" pageNumber="144">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB4B837FC8EFA30FC4EFE29" box="[860,951,1519,1544]" pageId="11" pageNumber="144" rank="species">
<emphasis id="02F12960FFB4B837FC8EFA30FC4EFE29" bold="true" box="[860,951,1519,1544]" pageId="11" pageNumber="144">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="F7858EF1FFB4B837FB20FA2EFA8CFE29" box="[1266,1397,1520,1544]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="buchardi">
<emphasis id="02F12960FFB4B837FB20FA2EFA8CFE29" box="[1266,1397,1520,1544]" italics="true" pageId="11" pageNumber="144">L. buchardi</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FA57FA2EFED2FE0D" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="ngoclinhense">
<emphasis id="02F12960FFB4B837FA57FA2EFED2FE0D" italics="true" pageId="11" pageNumber="144">L. ngoclinhense</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FEE8F9CAFE5FFE0D" box="[314,422,1556,1580]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="pullum">
<emphasis id="02F12960FFB4B837FEE8F9CAFE5FFE0D" box="[314,422,1556,1580]" italics="true" pageId="11" pageNumber="144">L. pullum</emphasis>
</taxonomicName>
and
<taxonomicName id="F7858EF1FFB4B837FE0DF9CAFD87FE0D" box="[479,638,1556,1580]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="hainanense">
<emphasis id="02F12960FFB4B837FE0DF9CAFD87FE0D" box="[479,638,1556,1580]" italics="true" pageId="11" pageNumber="144">L. hainanense</emphasis>
</taxonomicName>
(versus two palmar tubercles subequal in size in former three species, and outer one slightly larger or equal to the inner one in
<taxonomicName id="F7858EF1FFB4B837FCF7F9E6FC3AFE71" box="[805,963,1592,1616]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="hainanense">
<emphasis id="02F12960FFB4B837FCF7F9E6FC3AFE71" box="[805,963,1592,1616]" italics="true" pageId="11" pageNumber="144">L. hainanense</emphasis>
</taxonomicName>
).
<taxonomicName id="F7858EF1FFB4B837FC0FF9E7FACEFE71" box="[989,1335,1592,1616]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB4B837FC0FF9E7FACEFE71" box="[989,1335,1592,1616]" italics="true" pageId="11" pageNumber="144">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB4B837FA92F9E8FA65FE71" box="[1344,1436,1591,1616]" pageId="11" pageNumber="144" rank="species">
<emphasis id="02F12960FFB4B837FA92F9E8FA65FE71" bold="true" box="[1344,1436,1591,1616]" pageId="11" pageNumber="144">sp. nov.</emphasis>
</taxonomicNameLabel>
further differs from
<taxonomicName id="F7858EF1FFB4B837FEAEF982FDD1FE55" box="[380,552,1629,1652]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="guangxiense">
<emphasis id="02F12960FFB4B837FEAEF982FDD1FE55" box="[380,552,1629,1652]" italics="true" pageId="11" pageNumber="144">L. guangxiense</emphasis>
</taxonomicName>
and
<taxonomicName id="F7858EF1FFB4B837FDB3F982FCF9FE55" box="[609,768,1628,1652]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="hainanense">
<emphasis id="02F12960FFB4B837FDB3F982FCF9FE55" box="[609,768,1628,1652]" italics="true" pageId="11" pageNumber="144">L. hainanense</emphasis>
</taxonomicName>
by having an indistinct tympanum (versus distinct in both species).
</paragraph>
<paragraph id="303AF572FFB4B837FF15F97BFC16FF98" blockId="11.[151,1437,151,1977]" pageId="11" pageNumber="144">
Besides those differences mentioned above,
<taxonomicName id="F7858EF1FFB4B837FD14F97BFBD8FE9D" box="[710,1057,1700,1724]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB4B837FD14F97BFBD8FE9D" box="[710,1057,1700,1724]" italics="true" pageId="11" pageNumber="144">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB4B837FBF9F97BFB7EFE9C" box="[1067,1159,1700,1725]" pageId="11" pageNumber="144" rank="species">
<emphasis id="02F12960FFB4B837FBF9F97BFB7EFE9C" bold="true" box="[1067,1159,1700,1725]" pageId="11" pageNumber="144">sp. nov.</emphasis>
</taxonomicNameLabel>
further differs from the phylogenetically closely related congeners without spines on the upper lip in adult males, namely
<taxonomicName id="F7858EF1FFB4B837FB31F916FA8EFEC1" box="[1251,1399,1736,1760]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="chapaense">
<emphasis id="02F12960FFB4B837FB31F916FA8EFEC1" box="[1251,1399,1736,1760]" italics="true" pageId="11" pageNumber="144">L. chapaense</emphasis>
</taxonomicName>
,
<taxonomicName id="F7858EF1FFB4B837FA57F916FF0DFF25" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="huashen">
<emphasis id="02F12960FFB4B837FA57F916FF0DFF25" italics="true" pageId="11" pageNumber="144">L. huashen</emphasis>
</taxonomicName>
and
<taxonomicName id="F7858EF1FFB4B837FEF8F932FE10FF25" box="[298,489,1772,1796]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="masatakasatoi">
<emphasis id="02F12960FFB4B837FEF8F932FE10FF25" box="[298,489,1772,1796]" italics="true" pageId="11" pageNumber="144">L. masatakasatoi</emphasis>
</taxonomicName>
(
<figureCitation id="A8BEE9F7FFB4B837FE2AF933FDC5FF25" box="[504,572,1772,1797]" captionStart="FIGURE 2" captionStartId="4.[151,250,1944,1966]" captionTargetBox="[180,1393,274,1914]" captionTargetId="figure@4.[180,1406,265,1923]" captionTargetPageId="4" captionText="FIGURE 2. Bayesian inference (BI) tree derived from partial sequences of the mitochondrial 16 S rRNA gene. Numbers above branches are bootstrap support values for maximum likelihood analyses (&gt; 70 retained) and numbers below branches indicate Bayesian posterior probabilities (&gt; 70 % retained)." httpUri="https://zenodo.org/record/256325/files/figure.png" pageId="11" pageNumber="144">Fig. 2</figureCitation>
), by having a pinkish grey dorsal surface of body in life (versus brown in life in
<taxonomicName id="F7858EF1FFB4B837FF61F8CEFEBFFF09" box="[179,326,1808,1832]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="chapaense">
<emphasis id="02F12960FFB4B837FF61F8CEFEBFFF09" box="[179,326,1808,1832]" italics="true" pageId="11" pageNumber="144">L. chapaense</emphasis>
</taxonomicName>
and
<taxonomicName id="F7858EF1FFB4B837FEAEF8CEFE0CFF09" box="[380,501,1808,1832]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="huashen">
<emphasis id="02F12960FFB4B837FEAEF8CEFE0CFF09" box="[380,501,1808,1832]" italics="true" pageId="11" pageNumber="144">L. huashen</emphasis>
</taxonomicName>
,
<figureCitation id="A8BEE9F7FFB4B837FDD1F8D0FDB1FF09" box="[515,584,1807,1832]" captionStart="FIGURE 4" captionStartId="9.[151,250,1874,1896]" captionTargetBox="[151,1436,301,1851]" captionTargetId="figure@9.[151,1436,301,1853]" captionTargetPageId="9" captionText="FIGURE 4. (A) Holotype of Leptobrachium tengchongense sp. nov. (SYS a 004603) in life, inserted picture depicts the iris coloration of holotype in life; (B) Leptobrachium huashen from Linjiapu substation of Gaoligongshan NNR, inserted picture depicts the iris coloration in life. Photos by J. H. Yang." httpUri="https://zenodo.org/record/256327/files/figure.png" pageId="11" pageNumber="144">Fig. 4</figureCitation>
.), presence of a distinct network of ridges on the dorsum (versus absent in
<taxonomicName id="F7858EF1FFB4B837FA57F8CEFEC0FF6D" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="masatakasatoi">
<emphasis id="02F12960FFB4B837FA57F8CEFEC0FF6D" italics="true" pageId="11" pageNumber="144">L. masatakasatoi</emphasis>
</taxonomicName>
); inner metatarsal tubercle larger than first toe (versus smaller in
<taxonomicName id="F7858EF1FFB4B837FBF8F8EAFB12FF6D" box="[1066,1259,1844,1868]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="masatakasatoi">
<emphasis id="02F12960FFB4B837FBF8F8EAFB12FF6D" box="[1066,1259,1844,1868]" italics="true" pageId="11" pageNumber="144">L. masatakasatoi</emphasis>
</taxonomicName>
); presence of a dark black spot on lower anterior corner of eye (versus absent in
<taxonomicName id="F7858EF1FFB4B837FC58F886FBD9FF51" box="[906,1056,1880,1904]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="chapaense">
<emphasis id="02F12960FFB4B837FC58F886FBD9FF51" box="[906,1056,1880,1904]" italics="true" pageId="11" pageNumber="144">L. chapaense</emphasis>
</taxonomicName>
and
<taxonomicName id="F7858EF1FFB4B837FB8EF886FB20FF51" box="[1116,1241,1880,1904]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="11" pageNumber="144" phylum="Chordata" rank="species" species="huashen">
<emphasis id="02F12960FFB4B837FB8EF886FB20FF51" box="[1116,1241,1880,1904]" italics="true" pageId="11" pageNumber="144">L. huashen</emphasis>
</taxonomicName>
), relatively wide black bands on the dorsal surface of the limbs (versus relatively narrower bands in these species); and large and distinct cream spots on lower flanks absent (versus present in these species).
</paragraph>
<paragraph id="303AF572FFB3B830FFA5FDA5FA8AFAB0" blockId="12.[119,1869,634,1582]" box="[119,1395,634,657]" pageId="12" pageNumber="145">
<emphasis id="02F12960FFB3B830FFA5FDA5FF1CFAB0" bold="true" box="[119,229,634,657]" pageId="12" pageNumber="145">TABLE ³.</emphasis>
Measurements (mm) of type specimens of
<taxonomicName id="F7858EF1FFB3B830FD70FDA5FC24FAB0" box="[674,989,634,657]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="12" pageNumber="145" phylum="Chordata" rank="species" species="tengchongense" status="sp. nov.">
<emphasis id="02F12960FFB3B830FD70FDA5FC24FAB0" box="[674,989,634,657]" italics="true" pageId="12" pageNumber="145">Leptobrachium tengchongense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="19C2941BFFB3B830FC36FDA5FBCFFAB0" box="[996,1078,634,657]" pageId="12" pageNumber="145" rank="species">
<emphasis id="02F12960FFB3B830FC36FDA5FBCFFAB0" bold="true" box="[996,1078,634,657]" pageId="12" pageNumber="145">sp. nov.</emphasis>
</taxonomicNameLabel>
. Abbreviations đefineđ in text.
</paragraph>
</subSubSection>
<subSubSection id="789FA6F9FFB3B830FEDEFD76F8C6FE0F" pageId="12" pageNumber="145" type="description">
<paragraph id="303AF572FFB3B830FEDEFD76F8B4FAE1" blockId="12.[119,1869,634,1582]" box="[268,1869,681,704]" pageId="12" pageNumber="145">SYS SYS SYS SYS SYS SYS SYS SYS SYS SYS SYS Mean ± SD</paragraph>
<paragraph id="303AF572FFB3B830FEDEFD08F8DBFACF" blockId="12.[119,1869,634,1582]" box="[268,1826,727,750]" pageId="12" pageNumber="145">a004603 a004623 a004624 a004625 a004626 a004627 a004628 a004629 a004630 a004631 a004632 (N=11)</paragraph>
<paragraph id="303AF572FFB3B830FEDEFCD7F953FB3E" blockId="12.[119,1869,634,1582]" box="[268,1706,776,799]" pageId="12" pageNumber="145">Holotype Paratype Paratype Paratype Paratype Paratype Paratype Paratype Paratype Paratype Paratype</paragraph>
<paragraph id="303AF572FFB3B830FEDEFCE9F97CFB6C" blockId="12.[119,1869,634,1582]" box="[268,1669,822,845]" pageId="12" pageNumber="145">Male Male Male Male Male Male Male Male Male Male Male</paragraph>
<paragraph id="303AF572FFB3B830FEDEFCBBF8B5FBF6" blockId="12.[119,1869,634,1582]" pageId="12" pageNumber="145">45.3 51.5 48.0 45.9 47.9 50.5 48.2 48.9 41.7 48.6 50.3 47.89±2.76 HDL 18.4 20.7 19.4 18.7 19.4 20.2 19.7 20.8 17.5 18.9 20.2 19.45±1.02 HDW 19.9 21.7 20.4 20.0 20.2 21.2 20.9 22.3 18.2 20.5 21.9 20.65±1.14</paragraph>
<paragraph id="303AF572FFB3B830FEDEFC31F8C7FC43" blockId="12.[119,1869,634,1582]" pageId="12" pageNumber="145">7.8 8.4 8.1 7.9 7.7 8.1 8.1 7.9 7.0 8.2 8.5 7.97±0.40 EYE 6.7 7.4 6.8 6.2 6.5 6.9 6.7 6.9 5.4 6.5 6.4 6.58±0.50 UEW 5.0 5.6 5.1 5.0 5.0 5.3 5.2 5.8 4.8 4.8 4.6 5.11±0.35</paragraph>
<paragraph id="303AF572FFB3B830FEDEFBA6F8C6FCB1" blockId="12.[119,1869,634,1582]" box="[268,1855,1145,1168]" pageId="12" pageNumber="145">6.0 6.0 6.7 6.1 6.3 6.9 6.6 6.3 6.5 6.6 6.6 6.42±0.30</paragraph>
<paragraph id="303AF572FFB3B830FEDEFB78F8B5FD3B" blockId="12.[119,1869,634,1582]" pageId="12" pageNumber="145">4.8 5.1 5.1 4.7 4.6 4.9 5.1 5.2 4.7 5.2 5.1 4.95±0.22 LAL 14.1 14.4 14.8 13.7 13.6 14.4 13.7 14.1 12.6 14.3 14.0 13.97±0.58 HND 11.9 13.5 12.9 12.6 12.4 13.7 12.0 13.3 11.5 13.2 13.2 12.75±0.72</paragraph>
<paragraph id="303AF572FFB3B830FEDEFAEEF8B5FD57" blockId="12.[119,1869,634,1582]" pageId="12" pageNumber="145">16.9 19.0 17.5 17.2 17.6 18.8 17.6 18.4 15.9 17.9 18.0 17.71±0.88 TGH 20.6 22.2 20.9 20.1 20.6 22.0 20.4 21.6 16.9 20.3 21.6 20.65±1.44</paragraph>
<paragraph id="303AF572FFB3B830FEDEFA52F8C6FD85" blockId="12.[119,1869,634,1582]" box="[268,1855,1421,1444]" pageId="12" pageNumber="145">9.8 9.8 9.2 9.7 9.4 9.5 9.9 9.8 9.3 10.0 10.7 9.74±0.41</paragraph>
<paragraph id="303AF572FFB3B830FEDEFA64F8B5FDF3" blockId="12.[119,1869,634,1582]" box="[268,1868,1467,1490]" pageId="12" pageNumber="145">18.3 20.4 19.6 18.4 18.3 20.9 18.5 20.0 16.6 20.0 20.3 19.21±1.28</paragraph>
<paragraph id="303AF572FFB3B830FEDEFA36F8C6FE0F" blockId="12.[119,1869,634,1582]" pageId="12" pageNumber="145">3.0 3.4 3.4 2.7 2.9 3.7 3.0 3.3 2.6 3.6 3.6 3.20±0.38 IMW 1.5 1.8 1.5 1.3 1.5 1.6 1.5 1.8 1.3 1.6 1.7 1.55±0.17</paragraph>
</subSubSection>
</treatment>
</document>