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<document id="92EEDAFABC1C8C1CF9763B0BEA88DEE2" ID-DOI="10.5281/zenodo.178242" ID-GBIF-Dataset="ad386ac9-d6e0-4ab3-a5c5-b417d35e2b44" ID-ISSN="1175-5326" ID-Zenodo-Dep="178242" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459886466042" checkinUser="plazi" docAuthor="Saito, Fuki &amp; Kojima, Jun-Ichi" docDate="2007" docId="4E55E763FFC2FFC7FF53FA8CFCCA034C" docLanguage="en" docName="zt01556p030.pdf" docOrigin="Zootaxa 1556" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Eustenogaster fumipennis Saito, sp. nov." docType="treatment" docVersion="10" lastPageNumber="22" masterDocId="B26C9F1BFFD6FFD2FFC4FF33FFF50016" masterDocTitle="A taxonomic revision of the hover wasp genus Eustenogaster van der Vecht (Insecta: Hymenoptera; Vespidae, Stenogastrinae)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="21" updateTime="1698227849508" updateUser="plazi">
<mods:mods id="8FDB87F6F87B52DC66E832D3D2221F39" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="715609DDE1685CB2C7386D616E97D4FE">
<mods:title id="395FB15AD1EF85E2ACDF407C2029B808">A taxonomic revision of the hover wasp genus Eustenogaster van der Vecht (Insecta: Hymenoptera; Vespidae, Stenogastrinae)</mods:title>
</mods:titleInfo>
<mods:name id="69963724DFA3F05A3280EECFC37701C5" type="personal">
<mods:role id="654D1E4A27B2BEEE14FDD32BCE08C5FA">
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</mods:role>
<mods:namePart id="643FA2E153BFC04A89FA4A7CFA5185F7">Saito, Fuki</mods:namePart>
</mods:name>
<mods:name id="EA9CEE803D3A8987969F71C749A98F88" type="personal">
<mods:role id="B4CF56E7F395F155F2B255CD6A752890">
<mods:roleTerm id="285E50D0B45BB0F9490758B8D568A09F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="70927A0CA5A72F527C5ED973683DC2F4">Kojima, Jun-Ichi</mods:namePart>
</mods:name>
<mods:typeOfResource id="3B3670AC181ED13C4BD28E345F053EBF">text</mods:typeOfResource>
<mods:relatedItem id="3AF617614ECDB9232FE8C36F327BA505" type="host">
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<mods:title id="CEE2C11CCC98B6E3BF4EB8A435E7A597">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="812B66280502F9880A08F36F7AA62A68">
<mods:date id="8837FD9A39AC1E64B821C398833D35D0">2007</mods:date>
<mods:detail id="7D2C71F28BA7B8CAE6FF07E5AC5CC3D7" type="volume">
<mods:number id="86223EAA2900F594880D6DB3377F4CAA">1556</mods:number>
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<mods:classification id="C08DCA7BC037C2C74655489C7F72BD68">journal article</mods:classification>
<mods:identifier id="F2A762F539F0D3D164247351F9F3270D" type="DOI">10.5281/zenodo.178242</mods:identifier>
<mods:identifier id="A04887502278611D5AB4B1335812D007" type="GBIF-Dataset">ad386ac9-d6e0-4ab3-a5c5-b417d35e2b44</mods:identifier>
<mods:identifier id="CD88C82E73F63CA7EEE8E48FD833FB59" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CD7AF34F82E965CCA9D854A4617C2AC2" type="Zenodo-Dep">178242</mods:identifier>
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<treatment id="4E55E763FFC2FFC7FF53FA8CFCCA034C" ID-DOI="http://doi.org/10.5281/zenodo.3507583" ID-GBIF-Taxon="119371594" ID-Zenodo-Dep="3507583" LSID="urn:lsid:plazi:treatment:4E55E763FFC2FFC7FF53FA8CFCCA034C" httpUri="http://treatment.plazi.org/id/4E55E763FFC2FFC7FF53FA8CFCCA034C" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
<subSubSection id="8EE605FEFFC2FFC6FF53FA8CFF0506FA" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="C6435675FFC2FFC6FF53FA8CFD7205CF" blockId="20.[151,647,1471,1497]" box="[151,647,1471,1497]" pageId="20" pageNumber="21">
<heading id="9D0BE119FFC2FFC6FF53FA8CFD7205CF" bold="true" box="[151,647,1471,1497]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="F4888A67FFC2FFC6FF53FA8CFD7205CF" bold="true" box="[151,647,1471,1497]" pageId="20" pageNumber="21">
<taxonomicName id="01FC2DF6FFC2FFC6FF53FA8CFDE905CF" authority="Saito" authorityName="Saito" box="[151,540,1471,1497]" class="Insecta" family="Vespidae" genus="Eustenogaster" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fumipennis" status="sp. nov.">
<emphasis id="F4888A67FFC2FFC6FF53FA8CFE2605CF" bold="true" box="[151,467,1471,1497]" italics="true" pageId="20" pageNumber="21">Eustenogaster fumipennis</emphasis>
Saito
</taxonomicName>
,
<taxonomicNameLabel id="EFBB371CFFC2FFC6FDEDFA8CFD7205CF" box="[553,647,1471,1497]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C6435675FFC2FFC6FF53F939FF0506FA" blockId="20.[151,1437,1546,1972]" pageId="20" pageNumber="21">
This species is known only from the female and is similar to
<taxonomicName id="01FC2DF6FFC2FFC6FCA5F938FBEC0632" box="[865,1049,1547,1572]" class="Insecta" family="Vespidae" genus="Stenogaster" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="calyptodoma">
<emphasis id="F4888A67FFC2FFC6FCA5F938FBEC0632" box="[865,1049,1547,1572]" italics="true" pageId="20" pageNumber="21">E. calyptodoma</emphasis>
</taxonomicName>
, but it can be distinguished from
<taxonomicName id="01FC2DF6FFC2FFC6FF53F900FEA0065A" box="[151,341,1587,1612]" class="Insecta" family="Vespidae" genus="Stenogaster" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="calyptodoma">
<emphasis id="F4888A67FFC2FFC6FF53F900FEA0065A" box="[151,341,1587,1612]" italics="true" pageId="20" pageNumber="21">E. calyptodoma</emphasis>
</taxonomicName>
by the combination of the following characters: body size larger than
<taxonomicName id="01FC2DF6FFC2FFC6FB12F900FA61065A" box="[1238,1428,1587,1612]" class="Insecta" family="Vespidae" genus="Stenogaster" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="calyptodoma">
<emphasis id="F4888A67FFC2FFC6FB12F900FA61065A" box="[1238,1428,1587,1612]" italics="true" pageId="20" pageNumber="21">E. calyptodoma</emphasis>
</taxonomicName>
; extreme apex of clypeus pointed (
<figureCitation id="5EC74AF0FFC2FFC6FDF1F969FD790662" box="[565,652,1626,1652]" captionStart="FIGURES 26 31" captionStartId="9.[151,269,1734,1758]" captionTargetBox="[183,1431,191,1688]" captionTargetId="figure@9.[155,1432,170,1730]" captionTargetPageId="9" captionText="FIGURES 26 31. Eustenogaster fumipennis Saito sp. nov., holotype female; 26, head in frontal view. 27, mandible. 28, flat area of sixth metasomal tergum. 29, ocelli. 30, second metasomal tergum in dorsal view. 31, propodeum in dorsal view. Scale bars show 1 mm." httpUri="https://zenodo.org/record/178243/files/figure.png" pageId="20" pageNumber="21">Fig. 26</figureCitation>
<emphasis id="F4888A67FFC2FFC6FD53F968FD5A0662" box="[663,687,1627,1652]" italics="true" pageId="20" pageNumber="21">vs</emphasis>
. truncate in
<taxonomicName id="01FC2DF6FFC2FFC6FC88F968FBFD0662" box="[844,1032,1627,1652]" class="Insecta" family="Vespidae" genus="Stenogaster" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="calyptodoma">
<emphasis id="F4888A67FFC2FFC6FC88F968FBFD0662" box="[844,1032,1627,1652]" italics="true" pageId="20" pageNumber="21">E. calyptodoma</emphasis>
</taxonomicName>
, Fig. 21); mid-tooth of mandible rounded apically (
<figureCitation id="5EC74AF0FFC2FFC6FEA9F9B1FE34068A" box="[365,449,1666,1692]" captionStart="FIGURES 26 31" captionStartId="9.[151,269,1734,1758]" captionTargetBox="[183,1431,191,1688]" captionTargetId="figure@9.[155,1432,170,1730]" captionTargetPageId="9" captionText="FIGURES 26 31. Eustenogaster fumipennis Saito sp. nov., holotype female; 26, head in frontal view. 27, mandible. 28, flat area of sixth metasomal tergum. 29, ocelli. 30, second metasomal tergum in dorsal view. 31, propodeum in dorsal view. Scale bars show 1 mm." httpUri="https://zenodo.org/record/178243/files/figure.png" pageId="20" pageNumber="21">Fig. 27</figureCitation>
<emphasis id="F4888A67FFC2FFC6FE0EF9B0FE17068A" box="[458,482,1667,1692]" italics="true" pageId="20" pageNumber="21">vs</emphasis>
. Fig.
<quantity id="0104FB90FFC2FFC6FDEDF9B1FD90068A" box="[553,613,1666,1692]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="20" pageNumber="21" unit="in" value="22.0">22 in</quantity>
<taxonomicName id="01FC2DF6FFC2FFC6FDA9F9B0FCD3068A" box="[621,806,1667,1692]" class="Insecta" family="Vespidae" genus="Stenogaster" kingdom="Animalia" order="Hymenoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="calyptodoma">
<emphasis id="F4888A67FFC2FFC6FDA9F9B0FCD3068A" box="[621,806,1667,1692]" italics="true" pageId="20" pageNumber="21">E. calyptodoma</emphasis>
</taxonomicName>
); frons covered with dense, long hairs; sixth metasomal tergum with sharp spine and wide flat area in front of the spine (
<figureCitation id="5EC74AF0FFC2FFC6FC16F999FBDE06D2" box="[978,1067,1706,1732]" captionStart="FIGURES 26 31" captionStartId="9.[151,269,1734,1758]" captionTargetBox="[183,1431,191,1688]" captionTargetId="figure@9.[155,1432,170,1730]" captionTargetPageId="9" captionText="FIGURES 26 31. Eustenogaster fumipennis Saito sp. nov., holotype female; 26, head in frontal view. 27, mandible. 28, flat area of sixth metasomal tergum. 29, ocelli. 30, second metasomal tergum in dorsal view. 31, propodeum in dorsal view. Scale bars show 1 mm." httpUri="https://zenodo.org/record/178243/files/figure.png" pageId="20" pageNumber="21">Fig. 28</figureCitation>
); bright markings reduced (see below).
</paragraph>
</subSubSection>
<subSubSection id="8EE605FEFFC2FFC7FF03F9C8FE78020C" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="C6435675FFC2FFC6FF03F9C8FCB9072A" blockId="20.[151,1437,1546,1972]" pageId="20" pageNumber="21">
<emphasis id="F4888A67FFC2FFC6FF03F9C8FEEA0702" box="[199,287,1787,1812]" italics="true" pageId="20" pageNumber="21">Female</emphasis>
. Body length (head + mesosoma + first two terga)
<quantity id="0104FB90FFC2FFC6FC62F9C9FBCA0702" box="[934,1087,1786,1812]" metricMagnitude="-2" metricUnit="m" metricValue="1.925" metricValueMax="2.0" metricValueMin="1.85" pageId="20" pageNumber="21" unit="mm" value="19.25" valueMax="20.0" valueMin="18.5">18.520 mm</quantity>
(
<typeStatus id="1947E8D7FFC2FFC6FB97F9C9FB4A0702" box="[1107,1215,1786,1812]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
: about
<quantity id="0104FB90FFC2FFC6FAD9F9C9FA7A0702" box="[1309,1423,1786,1812]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="20" pageNumber="21" unit="mm" value="19.5">19.5 mm</quantity>
); forewing length
<quantity id="0104FB90FFC2FFC6FE9CF811FDF6072A" box="[344,515,1826,1852]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.55" metricValueMin="1.45" pageId="20" pageNumber="21" unit="mm" value="15.0" valueMax="15.5" valueMin="14.5">14.515.5 mm</quantity>
(
<typeStatus id="1947E8D7FFC2FFC6FDD6F811FD8A072A" box="[530,639,1826,1852]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
: about 15.0 mm).
</paragraph>
<paragraph id="C6435675FFC2FFC6FF03F879FEB407A2" blockId="20.[151,1437,1546,1972]" pageId="20" pageNumber="21">
Anterior ocellus about 1.2 times larger in width than posterior ocellus (
<figureCitation id="5EC74AF0FFC2FFC6FBC0F879FBAE0772" box="[1028,1115,1866,1892]" captionStart="FIGURES 26 31" captionStartId="9.[151,269,1734,1758]" captionTargetBox="[183,1431,191,1688]" captionTargetId="figure@9.[155,1432,170,1730]" captionTargetPageId="9" captionText="FIGURES 26 31. Eustenogaster fumipennis Saito sp. nov., holotype female; 26, head in frontal view. 27, mandible. 28, flat area of sixth metasomal tergum. 29, ocelli. 30, second metasomal tergum in dorsal view. 31, propodeum in dorsal view. Scale bars show 1 mm." httpUri="https://zenodo.org/record/178243/files/figure.png" pageId="20" pageNumber="21">Fig. 29</figureCitation>
); distance between anterior and posterior ocelli shorter than diameter of posterior ocellus; distance between posterior ocelli about equal to their diameter.
</paragraph>
<paragraph id="C6435675FFC3FFC7FF03FFABFEF00114" blockId="21.[151,1436,152,858]" pageId="21" pageNumber="22">
First metasomal tergum narrow and long (
<quantity id="0104FB90FFC3FFC7FD06FFABFCA100A4" box="[706,852,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="21" pageNumber="22" unit="mm" value="8.0" valueMax="8.5" valueMin="7.5">7.58.5 mm</quantity>
long), about 6.3 times longer than its maximum height and about 4.8 times longer than the maximum width; second tergum in dorsal view longer than wide (
<figureCitation id="5EC74AF0FFC3FFC7FF5BFFDBFF020114" box="[159,247,232,258]" captionStart="FIGURES 26 31" captionStartId="9.[151,269,1734,1758]" captionTargetBox="[183,1431,191,1688]" captionTargetId="figure@9.[155,1432,170,1730]" captionTargetPageId="9" captionText="FIGURES 26 31. Eustenogaster fumipennis Saito sp. nov., holotype female; 26, head in frontal view. 27, mandible. 28, flat area of sixth metasomal tergum. 29, ocelli. 30, second metasomal tergum in dorsal view. 31, propodeum in dorsal view. Scale bars show 1 mm." httpUri="https://zenodo.org/record/178243/files/figure.png" pageId="21" pageNumber="22">Fig. 30</figureCitation>
).
</paragraph>
<paragraph id="C6435675FFC3FFC7FF03FE23FD7B01E4" blockId="21.[151,1436,152,858]" pageId="21" pageNumber="22">
Color: Body shiny black; bright markings reduced and restricted to the following parts: spot below antennal socket (
<figureCitation id="5EC74AF0FFC3FFC7FEDAFE0BFE830144" box="[286,374,312,338]" captionStart="FIGURE 57" captionStartId="15.[151,255,1665,1689]" captionTargetBox="[186,1381,184,1648]" captionTargetId="figure@15.[183,1402,178,1648]" captionTargetPageId="15" captionText="FIGURE 57. Facial marking pattern of Eustenogaster species." httpUri="https://zenodo.org/record/178247/files/figure.png" pageId="21" pageNumber="22">Fig. 57</figureCitation>
), paired small lateral spot on clypeus (sometimes absent), small spot at posterodorsal corner of pronotum, paired lateral spots on metanotum (sometimes absent), paired posterior spots on propodeum (
<figureCitation id="5EC74AF0FFC3FFC7FF5BFEBBFF0301B4" box="[159,246,392,418]" captionStart="FIGURES 26 31" captionStartId="9.[151,269,1734,1758]" captionTargetBox="[183,1431,191,1688]" captionTargetId="figure@9.[155,1432,170,1730]" captionTargetPageId="9" captionText="FIGURES 26 31. Eustenogaster fumipennis Saito sp. nov., holotype female; 26, head in frontal view. 27, mandible. 28, flat area of sixth metasomal tergum. 29, ocelli. 30, second metasomal tergum in dorsal view. 31, propodeum in dorsal view. Scale bars show 1 mm." httpUri="https://zenodo.org/record/178243/files/figure.png" pageId="21" pageNumber="22">Fig. 31</figureCitation>
), all yellow; second metasomal tergum with very small, yellow or pale-brown lateral spots. Legs dark brown; fore tibia with variable-sized lateral yellow spot. Wings semihyaline, smoky brown, darker along anterior margin of forewing; veins dark brown.
</paragraph>
<paragraph id="C6435675FFC3FFC7FF03FD33FE78020C" blockId="21.[151,1436,152,858]" box="[199,397,512,538]" pageId="21" pageNumber="22">
<emphasis id="F4888A67FFC3FFC7FF03FD33FEF6020F" box="[199,259,512,537]" italics="true" pageId="21" pageNumber="22">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="8EE605FEFFC3FFC7FF03FD1BFBE602AC" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="C6435675FFC3FFC7FF03FD1BFD38027C" blockId="21.[151,1436,152,858]" pageId="21" pageNumber="22">
<emphasis id="F4888A67FFC3FFC7FF03FD1BFE9C0257" box="[199,361,552,577]" italics="true" pageId="21" pageNumber="22">
<typeStatus id="1947E8D7FFC3FFC7FF03FD1BFF0B0257" box="[199,254,552,577]" pageId="21" pageNumber="22">Type</typeStatus>
material
</emphasis>
:
<typeStatus id="1947E8D7FFC3FFC7FEBEFD1BFE120254" box="[378,487,552,578]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
female (deposited in
<collectionCode id="A0EDCEB0FFC3FFC7FD2DFD1BFCB00254" LSID="urn:lsid:biocol.org:col:34925" box="[745,837,552,578]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="21" pageNumber="22">AMNH</collectionCode>
) labeled, “Antique: Alojipan, Culasi,
<date id="B24270B5FFC3FFC7FAC3FD1BFA620254" box="[1287,1431,552,578]" pageId="21" pageNumber="22" value="1982-12-20">20xii 1982</date>
, Starr &amp; Godoy, Nest series No. 591” and “GM”.
</paragraph>
<paragraph id="C6435675FFC3FFC7FF03FD4BFBE602AC" blockId="21.[151,1436,152,858]" pageId="21" pageNumber="22">
<typeStatus id="1947E8D7FFC3FFC7FF03FD4BFECB0284" box="[199,318,632,658]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
:
<emphasis id="F4888A67FFC3FFC7FE88FD44FD2D0287" bold="true" box="[332,728,631,657]" pageId="21" pageNumber="22">
The
<collectingCountry id="BEEB16E5FFC3FFC7FE43FD44FDED0287" box="[391,536,631,657]" name="Philippines" pageId="21" pageNumber="22">Philippines</collectingCountry>
: Panay Island:
</emphasis>
4Ψ (
<collectionCode id="A0EDCEB0FFC3FFC7FCDCFD4BFC920284" box="[792,871,632,658]" pageId="21" pageNumber="22">IUNH</collectionCode>
), Alojipan, Culasi, Antique,
<date id="B24270B5FFC3FFC7FB00FD4BFABE0284" box="[1220,1355,632,658]" pageId="21" pageNumber="22" value="1982-12-16">16.xii.1982</date>
, CKS, [1Ψ, Nest series No. 553; 1Ψ, Nest series No. 555; 2Ψ, Nest series No. 556].
</paragraph>
</subSubSection>
<subSubSection id="8EE605FEFFC3FFC7FF03FDFBFDB202F4" box="[199,583,712,738]" pageId="21" pageNumber="22" type="distribution">
<paragraph id="C6435675FFC3FFC7FF03FDFBFDB202F4" blockId="21.[151,1436,152,858]" box="[199,583,712,738]" pageId="21" pageNumber="22">
<emphasis id="F4888A67FFC3FFC7FF03FDFBFEA302F7" box="[199,342,712,737]" italics="true" pageId="21" pageNumber="22">Distribution</emphasis>
:
<collectingCountry id="BEEB16E5FFC3FFC7FEA1FDFBFE1B02F4" box="[357,494,712,738]" name="Philippines" pageId="21" pageNumber="22">Philippines</collectingCountry>
: Panay.
</paragraph>
</subSubSection>
<subSubSection id="8EE605FEFFC3FFC7FF03FDC3FCCA034C" pageId="21" pageNumber="22" type="etymology">
<paragraph id="C6435675FFC3FFC7FF03FDC3FCCA034C" blockId="21.[151,1436,152,858]" pageId="21" pageNumber="22">
<emphasis id="F4888A67FFC3FFC7FF03FDC3FEB6031F" box="[199,323,752,777]" italics="true" pageId="21" pageNumber="22">Etymology</emphasis>
. The specific name
<taxonomicName id="01FC2DF6FFC3FFC7FDF3FDC3FD4D031F" box="[567,696,752,777]" class="Insecta" family="Vespidae" genus="Eustenogaster" kingdom="Animalia" order="Hymenoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fumipennis">
<emphasis id="F4888A67FFC3FFC7FDF3FDC3FD4D031F" box="[567,696,752,777]" italics="true" pageId="21" pageNumber="22">fumipennis</emphasis>
</taxonomicName>
is based on van der Vechts manuscript name. This is a compound name formed with a masculine Latin noun,
<emphasis id="F4888A67FFC3FFC7FCCDFC2BFCC80327" box="[777,829,792,817]" italics="true" pageId="21" pageNumber="22">fumi</emphasis>
(= smoke) and a feminine Latin noun,
<emphasis id="F4888A67FFC3FFC7FAEDFC2BFA800327" box="[1321,1397,792,817]" italics="true" pageId="21" pageNumber="22">pennis</emphasis>
(= feather), to refer to the dark-colored wings of this species.
</paragraph>
</subSubSection>
</treatment>
</document>