242 lines
25 KiB
XML
242 lines
25 KiB
XML
<document ID-CLB-Dataset="129873" ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-PMC="PMC9848625" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-PubMed="36762355" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="1D5B716EC5CB5397A1659BC9C8255F0B" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupina dorri subsp. isanensis Jirapatrasilp 2022, ssp. nov." docType="treatment" docVersion="6" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1732923707409" updateUser="ExternalLinkService">
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<mods:mods id="21AEB1B926AEE9A012B42C94EB2A62C9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2BD923B9C04E4901BA237D1230F0FA45">
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<mods:title id="CE524A287A09EFF5821B415EBC06A8F4">Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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</mods:titleInfo>
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<mods:name id="15F0339A56DBD3581C1C92F3159B316C" type="personal">
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<mods:role id="75375726AE8735BBCF1462857345E225">
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<mods:roleTerm id="4929B3CEF3D156E8B3A4095B3B0FB3FD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E0105E3D2EA681DAAE99BD02526C126A">Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier id="E17D0B7AA381009873779A0BAD13B7B9" type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation id="D611B54532A655726C4F2CA675F2241A">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="14D08B21F318B9721F38984D79C69A66" type="personal">
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<mods:role id="8B0F014F6E4E48228D1FC1D32FF539D1">
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<mods:roleTerm id="AB47F541AB2270E710DF2984DBC4BFF6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4E4E8D37E7743E8E78B57527C509BAF8">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="79D1AF1E796AC5DC6F98042274E177A2">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="CE04A8DCD269942CCEFE49FC381EC3C0" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name id="D3E7EAA4071F47791D161602E66EB117" type="personal">
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<mods:roleTerm id="297E5A763861C6BADE5BB322541E8A9E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="247D279E3281C69C7111BAB4548B45CD">Panha, Somsak</mods:namePart>
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<mods:affiliation id="462C411D921471C584F02DC89E7B0150">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier id="17EFA95F0DBF265E228AAE3FFD82D4E9" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="E684EC5E0307B2081D9BF90361E64C46">text</mods:typeOfResource>
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<mods:titleInfo id="98A3EDAF8D790614697BB698073EABD0">
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<mods:title id="A88AC622D39DED568F1D4DCD65AD8CAB">ZooKeys</mods:title>
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<mods:part id="1CF3A0D48DE8504126A4D14D7305B5B5">
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<mods:date id="A531B1DBEA8E2472ED8D00C35467B747">2022</mods:date>
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<mods:number id="0D8BAADAE59E9CE31CF8B2EE81CA5796">2022-08-25</mods:number>
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</mods:detail>
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<mods:detail id="474A91DB44365468A59772BAB3648768" type="volume">
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<mods:number id="4C67C384086A8B54D0AE5D7FA2AA70E4">1119</mods:number>
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</mods:detail>
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<mods:extent id="B751904512532065DDF62A3154121E55" unit="page">
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<mods:start id="37D345FEBBD7B36A486A5E373CA5D6B2">1</mods:start>
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<mods:end id="A07D81C5B1DBE526821D3B9A57C8116F">115</mods:end>
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<mods:url id="EF376DBA1964A037BC9DC0F10D5826C8">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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</mods:location>
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<mods:classification id="41CAD6ECA17148E783CE20815678DA6C">journal article</mods:classification>
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<mods:identifier id="A307EE6D029716D005097735B484FCA1" type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier id="261D1516740515D7F8BA58CF93EA4D8A" type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<mods:identifier id="7CC1B6AC2A5291D0A0B9CB70EF31BCC0" type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
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<mods:identifier id="2C2BCBE9078731BCB6F4970ECB62A07F" type="Pensoft-UUID">F158A7C5261D52B69288A62F3C777CAF</mods:identifier>
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</mods:mods>
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<treatment id="1D5B716EC5CB5397A1659BC9C8255F0B" ID-GBIF-Taxon="199838999" LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B" httpUri="http://treatment.plazi.org/id/1D5B716EC5CB5397A1659BC9C8255F0B" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Pupinidae" scope_phylum="Mollusca" scope_subClass="Caenogastropoda" scope_subFamily="Pupininae" scope_superFamily="Cyclophoroidea">
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<subSubSection id="55D5EA3B516132F4E70FA38C71C045F2" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="A1A056BF3188300AA039B2198F9C1127" pageId="0" pageNumber="1">
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<taxonomicName id="0BEBBC48D2FB3CDA6FD9EFB3FD9216DD" LSID="1D5B716E-C5CB-5397-A165-9BC9C8255F0B" authority="Jirapatrasilp" authorityName="Jirapatrasilp" authorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina dorri subsp. isanensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dorri" status="ssp. nov." subSpecies="isanensis">Pupina dorri isanensis Jirapatrasilp</taxonomicName>
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<taxonomicNameLabel id="3A980362F3B7A52D23C759D4DCE31FB7" pageId="0" pageNumber="1">ssp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="4A7C35E8D0E91545B969755F97D7A7EC" pageId="0" pageNumber="1" type="description">
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<paragraph id="913B5A7C014AEE50C1E396F6D4D32756" pageId="0" pageNumber="1">
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<figureCitation id="29A3C425D2B92A23D0AF0C9950B15361" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">Figs 36K, L</figureCitation>
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<figureCitation id="72C22380BCEF8F1937DF9BC53BEE58DA" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. A, B Pupina paviei: specimens A NHMUK ex. Dautzenberg coll. from Kampot, Cambodge and B CUMZ 12134 from Lalu, Sa Kaeo C, D Pupina tchehelensis: specimens C SMF 109947 / 6 from Bukit Pondong, Perak and D CUMZ 12136 from limestone mountain, Phang Nga E, F Pupina dorri isanensis ssp. nov. E holotype CUMZ 12140 / 1 and F specimen CUMZ 12137 from Khao Wang Pha, Nong Bua Lam Phu." figureDoi="10.3897/zookeys.1119.85400.figure39" httpUri="https://binary.pensoft.net/fig/735400" pageId="0" pageNumber="1">, 39E, F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="963310801EC1D7BCE6EDD9E3BAF203EA" pageId="0" pageNumber="1" type="type material">
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<paragraph id="49F03F49734E810C49A0069985FB91FA" pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph id="FF703B08ACBA93B525AE2C04A13FE133" pageId="0" pageNumber="1">
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<emphasis id="B7E8BC4837E2DDAB42ECE978C1F0C9C5" bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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CUMZ 12140/1 (Figs
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<figureCitation id="3E8E901888D4D8F0F6CBD5BD9CDF70E7" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36K</figureCitation>
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,
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<figureCitation id="24B4A95DB4CEB3A18275A7728E8E1222" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. A, B Pupina paviei: specimens A NHMUK ex. Dautzenberg coll. from Kampot, Cambodge and B CUMZ 12134 from Lalu, Sa Kaeo C, D Pupina tchehelensis: specimens C SMF 109947 / 6 from Bukit Pondong, Perak and D CUMZ 12136 from limestone mountain, Phang Nga E, F Pupina dorri isanensis ssp. nov. E holotype CUMZ 12140 / 1 and F specimen CUMZ 12137 from Khao Wang Pha, Nong Bua Lam Phu." figureDoi="10.3897/zookeys.1119.85400.figure39" httpUri="https://binary.pensoft.net/fig/735400" pageId="0" pageNumber="1">39E</figureCitation>
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), 31 Aug. 2020, coll. C. Sutcharit, P. Jirapatrasilp, A. Pholyotha. Measurement: shell height 6.6 mm, shell width 4.6 mm and
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<normalizedToken id="30FAACCE7CF41008FF9C7C50C735EEA4" originalValue="5½">51/2</normalizedToken>
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whorls.
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<emphasis id="2007D0F84CDEEE9EAF7E03B949BC066B" bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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CUMZ 12140/2 (22 shells) and NHMUK 20210337 (3 shells), same data as holotype.
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</paragraph>
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</subSubSection>
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<subSubSection id="D48FEDD46D557C425BBD92259916D5CF" pageId="0" pageNumber="1" type="type locality">
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<paragraph id="FE8942E9BECC5F316A3CC474D654A4F1" pageId="0" pageNumber="1">Type locality.</paragraph>
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<paragraph id="DCE0495E30D51FDDCDCED4E4A8BA9FFD" pageId="0" pageNumber="1">
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Pa Pha Ya Temple, Suwannakhuha District, Nong Bua Lam Phu Province, Thailand,
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<geoCoordinate id="49C07EE578EA226B24B2FB28B0EC0323" degrees="17" direction="north" minutes="37" orientation="latitude" precision="1" seconds="38.8" value="17.627445">17°37'38.8"N</geoCoordinate>
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,
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<geoCoordinate id="36F826A395FAB25D37A89F1BE20CF0CC" degrees="102" direction="east" minutes="10" orientation="longitude" precision="1" seconds="13.7" value="102.17047">102°10'13.7"E</geoCoordinate>
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, 250 m amsl.
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</paragraph>
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</subSubSection>
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<subSubSection id="3E2B0127EBAF4C2A296B0C011057B024" pageId="0" pageNumber="1" type="materials_examined">
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<paragraph id="F11DE43798C6C471DA5623CD03D343AF" pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph id="34AEE9B2DB01C38CDD821BF3B00B1D78" pageId="0" pageNumber="1">
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<materialsCitation id="FA27B2B855BDFEEE162F3DB2FDD296C3" collectingDate="2007-10-15" collectorName="Figs" location="Na Wang District" municipality="Nong Bua Lam Phu Province" specimenCount="1">
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CUMZ 12137 (1 shell;
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<collectorName id="12CAE71B033FBD5B17A12948C065A562">Figs</collectorName>
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<figureCitation id="F13D698F212123FF8C19F8B858D89527" captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Shells of Pupina aureola species group from mainland Southeast Asia A-F Pupina aureola A possible syntype NHMUK 1988.12.4.101 and specimens B CUMZ 12124 C CUMZ 12126 D CUMZ 12130 E CUMZ 12133, and F CUMZ 12121 G, H Pupina stoliczkai sp. nov. G holotype CUMZ 12147 / 1 and H paratype NHMUK 20210336 I, J Pupina dorri dorri I paralectotype MNHN-IM- 2000 - 35836 and J specimen NHMUK ex. A. J. Piele Colln. Acc. No. 2242 K, L Pupina dorri isanensis spp. nov. K holotype CUMZ 12140 / 1 and L specimen CUMZ 12137 M Pupina brachysoma, lectotype MNHN-IM- 2000 - 9652. Photo: P. Maestrati, MNHN (I, M)" figureDoi="10.3897/zookeys.1119.85400.figure36" httpUri="https://binary.pensoft.net/fig/735397" pageId="0" pageNumber="1">36L</figureCitation>
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,
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<figureCitation id="E9D1089E3F89EAA5C6C4202985EA3AD5" captionStart="Figure 39" captionStartId="F39" captionText="Figure 39. A, B Pupina paviei: specimens A NHMUK ex. Dautzenberg coll. from Kampot, Cambodge and B CUMZ 12134 from Lalu, Sa Kaeo C, D Pupina tchehelensis: specimens C SMF 109947 / 6 from Bukit Pondong, Perak and D CUMZ 12136 from limestone mountain, Phang Nga E, F Pupina dorri isanensis ssp. nov. E holotype CUMZ 12140 / 1 and F specimen CUMZ 12137 from Khao Wang Pha, Nong Bua Lam Phu." figureDoi="10.3897/zookeys.1119.85400.figure39" httpUri="https://binary.pensoft.net/fig/735400" pageId="0" pageNumber="1">39F</figureCitation>
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) from Khao Wang Pha,
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<location id="E93DDB9D345C751B0BFC5A62F9144728" LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:E93DDB9D345C751B0BFC5A62F9144728" municipality="Nong Bua Lam Phu Province" name="Na Wang District">Na Wang District</location>
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,
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<collectingMunicipality id="5DCF3EF2B25C9CF4262A4E89F043866E">Nong Bua Lam Phu Province</collectingMunicipality>
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,
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<collectingDate id="1D53898D0F1AE88B1BA68E51C1F9DDD6" value="2007-10-15">15 Oct. 2007</collectingDate>
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</materialsCitation>
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.
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<materialsCitation id="28C97F0562629328F983F5048CABD5A4" collectingDate="2007-10-15" county="Nong Bua Lam Phu Province" location="Pa Jor Cave" municipality="Na Wang District" specimenCount="1">
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CUMZ 12138 (1 shell) from
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<location id="2D28654533D89CBDC4FAAEC7B05C40D0" LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:2D28654533D89CBDC4FAAEC7B05C40D0" county="Nong Bua Lam Phu Province" municipality="Na Wang District" name="Pa Jor Cave">Pa Jor Cave</location>
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,
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<collectingMunicipality id="57B752A818752A1515BFB731E7F84ACF">Na Wang District</collectingMunicipality>
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,
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<collectingCounty id="6B4814C7260793701FBEC09044610082">Nong Bua Lam Phu Province</collectingCounty>
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,
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<collectingDate id="525A6159332D9B794F964AC45B6590D1" value="2007-10-15">15 Oct. 2007</collectingDate>
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</materialsCitation>
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.
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<materialsCitation id="7798053D2ED441F1558FC2DE7056EB71" collectingDate="2020-08-31" collectorName="Tham Suwannakhuha Temple" location="Suwannakhuha District" municipality="Nong Bua Lam Phu Province" specimenCount="1">
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CUMZ 12139 (9 shells) from
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<collectorName id="745B11132EA7544B5A654C72AA90C142">Tham Suwannakhuha Temple</collectorName>
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,
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<location id="59A3B16627A6DACFFE6B6AC70D5E40B6" LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:59A3B16627A6DACFFE6B6AC70D5E40B6" municipality="Nong Bua Lam Phu Province" name="Suwannakhuha District">Suwannakhuha District</location>
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,
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<collectingMunicipality id="7977654470D36D526B27604A0AC7C1EA">Nong Bua Lam Phu Province</collectingMunicipality>
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,
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<collectingDate id="BEEBD9706A5CE6D587FDC033682DC5AC" value="2020-08-31">31 Aug. 2020</collectingDate>
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</materialsCitation>
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.
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<materialsCitation id="A3DE73B3446E7996160F6704B14E94FF" collectingDate="2020-08-31" county="Nong Bua Lam Phu Province" location="Namtok Thao To Forest" municipality="Mueang Nong Bua Lam Phu District" specimenCount="1">
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CUMZ 12141 (2 shells) from
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<location id="DE23C23CEA71AC910AC94D5A0D768342" LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:DE23C23CEA71AC910AC94D5A0D768342" county="Nong Bua Lam Phu Province" municipality="Mueang Nong Bua Lam Phu District" name="Namtok Thao To Forest">Namtok Thao To Forest</location>
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Park,
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<collectingMunicipality id="4356AA03FFC44590194235046AFADC7F">Mueang Nong Bua Lam Phu District</collectingMunicipality>
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,
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<collectingCounty id="742AC157FB3D047092FB2C60DA4BD221">Nong Bua Lam Phu Province</collectingCounty>
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,
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<collectingDate id="AA67A61491B84F524816A3C6183C105A" value="2020-08-31">31 Aug. 2020</collectingDate>
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</materialsCitation>
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.
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<materialsCitation id="1E6656804DBD743CDAEC6457E7B36862" collectingDate="2020-09-01" collectorName="Phu Pha Lom" location="Mueang Loei District" municipality="Loei Province" specimenCount="1">
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CUMZ 12142 (3 shells and
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<specimenCount id="A6582CD4260EDAE2BA510BF7DCEADE9E" type="generic">1 specimen</specimenCount>
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in ethanol) from
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<collectorName id="3E832EC8D07D9FB559B2AA83A9992BE0">Phu Pha Lom</collectorName>
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,
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<location id="55B18CDB07CD41BF4F16E9A44D7F6E0E" LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:55B18CDB07CD41BF4F16E9A44D7F6E0E" municipality="Loei Province" name="Mueang Loei District">Mueang Loei District</location>
|
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,
|
|
<collectingMunicipality id="03A037439DCB0C7EBE080A453F057006">Loei Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="7626AF02345170D8398F9F25DF708ECB" value="2020-09-01">1 Sept. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="38CB3FD4F11B355899BD7289DB16521C" collectingDate="2020-09-02" collectorName="Hin Pha Ngam Park" location="Nong Hin District" municipality="Loei Province" specimenCount="1">
|
|
CUMZ 12143 (
|
|
<specimenCount id="ADF0B2B97830D5316A5D16AB103DA086" type="generic">1 specimen</specimenCount>
|
|
in ethanol) from
|
|
<collectorName id="75039F1D7953919DEC82316CA12EAC11">Hin Pha Ngam Park</collectorName>
|
|
,
|
|
<location id="C2401E5B90FE7FA1199C2B3BD1DFCF5D" LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:C2401E5B90FE7FA1199C2B3BD1DFCF5D" municipality="Loei Province" name="Nong Hin District">Nong Hin District</location>
|
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,
|
|
<collectingMunicipality id="9407D6D7F8A992D27C33338EDD5F82E7">Loei Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="DE21931D7BBC27D104789E5AE3FC3B54" value="2020-09-02">2 Sept. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="5F9D0087CB380858D5FD8DC31C8E1B23" collectingDate="2020-07-21" county="Khon Kaen Province" location="Phraya Nakharaj Cave" municipality="Phu Pha Man District" specimenCount="1">
|
|
CUMZ 12144 (1 shell) from
|
|
<location id="67E285BDD6E12ED410106652C6D4E082" LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:67E285BDD6E12ED410106652C6D4E082" county="Khon Kaen Province" municipality="Phu Pha Man District" name="Phraya Nakharaj Cave">Phraya Nakharaj Cave</location>
|
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,
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|
<collectingMunicipality id="0623D847B13BE44C20B701C4BB608953">Phu Pha Man District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="AFFABAE695D98BFF8A2414D290D6DD67">Khon Kaen Province</collectingCounty>
|
|
,
|
|
<collectingDate id="1430D643EBFBA74983B23B3317ACA221" value="2020-07-21">21 July 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="1D6916566728D05EB190EECB874341A5" collectingDate="2020-08-30" collectorName="Phu Thong Thep Nimit Temple" location="Nong Saeng District" municipality="Udon Thani Province" specimenCount="1">
|
|
CUMZ 12145 (
|
|
<specimenCount id="E804B12E3B423C5B984E68BE7DB40156" type="generic">1 specimen</specimenCount>
|
|
in ethanol) from
|
|
<collectorName id="E9A9506867AD8F3F99196C44641F664F">Phu Thong Thep Nimit Temple</collectorName>
|
|
,
|
|
<location id="8A0BBAB5FA466B934DE29172C6CC20D1" LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:8A0BBAB5FA466B934DE29172C6CC20D1" municipality="Udon Thani Province" name="Nong Saeng District">Nong Saeng District</location>
|
|
,
|
|
<collectingMunicipality id="E5E8A321B92A7DF18591759CAE71F424">Udon Thani Province</collectingMunicipality>
|
|
,
|
|
<collectingDate id="C3F9E415049EF2DACDA72A04A8789ECF" value="2020-08-30">30 Aug. 2020</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="67DC718278B4EEF6F07836732EEC08AB" collectingDate="2009-09-15" county="Chanthaburi Province" location="Khao Wong Cave" municipality="Kaeng Hang Maeo District" specimenCount="1">
|
|
CUMZ 12170 (1 shell) from
|
|
<location id="82DE7C824E0DA7FE3720F53B43B74F47" LSID="urn:lsid:plazi:treatment:1D5B716EC5CB5397A1659BC9C8255F0B:82DE7C824E0DA7FE3720F53B43B74F47" county="Chanthaburi Province" municipality="Kaeng Hang Maeo District" name="Khao Wong Cave">Khao Wong Cave</location>
|
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,
|
|
<collectingMunicipality id="BAC1CD7D186E701C830EB6D3C2529766">Kaeng Hang Maeo District</collectingMunicipality>
|
|
,
|
|
<collectingCounty id="1DC56DB7505ED347CD4D0927D645CC5E">Chanthaburi Province</collectingCounty>
|
|
,
|
|
<collectingDate id="DB22F698900F51A9EA7973CED809898C" value="2009-09-15">15 Sept. 2009</collectingDate>
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="E32DA985BC15C76D4B529589E77A3312" pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph id="3AE515C20D2A23875809F0D22F767546" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph id="B02FF6E2855D1D948013BB9AA3189333" pageId="0" pageNumber="1">Shell ovate-fusiform; last whorl ca. 70% of shell height. Apertural lip thickened but not expanded. Parietal tooth triangular, thickened, covering posterior canal, approaching but not extending beyond the outer margin of apertural lip; columellar tooth fin-shaped, thickened, located next to slit-like anterior canal.</paragraph>
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|
</subSubSection>
|
|
<subSubSection id="DD67CC5AC421A28FE58BF4F2FC0D226E" pageId="0" pageNumber="1" type="differential diagnosis">
|
|
<paragraph id="0683E6B94B612872470E5E092B955AE1" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
|
<paragraph id="6DE9152B48F56DE46FF1D9B2B54CFFDA" pageId="0" pageNumber="1">This new subspecies is slightly different from the nominotypical subspecies in having the apertural lip, and parietal and columellar teeth more thickened.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="56719E6FBA5A085EA365990EA2883328" pageId="0" pageNumber="1" type="description">
|
|
<paragraph id="367E52BDD015789C8C3207FC82626436" pageId="0" pageNumber="1">Description.</paragraph>
|
|
<paragraph id="65690052DDACF35513DA6C1785F5274F" pageId="0" pageNumber="1">
|
|
Shell height 6.0-6.6 mm; shell width 4.2-4.6 mm. Shell ovate-fusiform, solid, semi-transparent, grey to pale brown, devoid of prominent sculpture on glazed smooth surface. Apex obtuse. Growth lines on shell surface inconspicuous. Whorls
|
|
<normalizedToken id="37B5247B848B40EAD72A06245F89CE2B" originalValue="5½">51/2</normalizedToken>
|
|
, last whorl large (ca. 70% of shell height). Spire angle ca. 75-80°, slightly extended. Sutures slightly impressed, but shallow. Aperture circular; lip thickened (ca. 0.2 mm wide and 0.3-0.4 mm thick) with paler colour, not expanded. Parietal callus not sharply defined and somewhat thickened with paler colour. Peristome interrupted by two canals; posterior canal slit-like ca. 0.7 mm long; anterior canal slit-like continuing horizontally ca. 0.8-0.9 mm. Parietal tooth triangular, thickened (ca. 0.7 mm long, 0.5 mm at its widest and 0.3 mm thick), covering posterior canal, approaching but not extending beyond the outer margin of apertural lip. Columellar tooth thickened (ca. 1.0 mm long, 0.3 mm at its widest and 0.3 mm thick), fin-shaped. Umbilicus closed. Operculum round, thin, flat, multispiral, whitish to pale yellow, opaque corneous with smooth edge.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B9C85886322B0A7A4546F1588B5A635C" pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph id="24315FABEC7AE97B084C6FA9D56AEEC4" pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph id="4F8ED6243E29E56114B38D0841586658" pageId="0" pageNumber="1">
|
|
The specific epithet refers to the Thai name
|
|
<normalizedToken id="285FAFEDB04DAF3729CB35D460EB05C5" originalValue="“Isan”">"Isan"</normalizedToken>
|
|
for the northeastern region of Thailand, where this new subspecies is mainly distributed.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="B52BA55B965EB7B0F48AFFFC069AC363" pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph id="5FF33F9BDE01D95C4EE89E7370CBF842" pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph id="CFB20DF345691953FD7A95678E67A52C" pageId="0" pageNumber="1">Northeastern and eastern Thailand.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="0A327B922318B853A06C9DC88294C902" pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph id="ED88EA5B0A490A257D93438A035F2C55" pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph id="43FCE0968F68DA6178E75BEC911926F7" pageId="0" pageNumber="1">Although the collecting localities of this new subspecies are ca. 600 km from the known occurrence of the nominotypical subspecies, DNA data and morphometric analyses are required to demonstrate whether these Thai specimens are distinct from the Vietnamese specimens and should be elevated to specific status.</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |