treatments-xml/data/1D/57/B2/1D57B212FFAFFFFC7EB9FE8A1014FE27.xml
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<document id="A1EF4EFA79B4F166C022F291705C8A8F" ID-CLB-Dataset="9025" ID-DOI="10.11646/zootaxa.4861.4.6" ID-GBIF-Dataset="59dc47c8-01b5-4f14-baa6-95b57158ca96" ID-ISSN="1175-5326" ID-Zenodo-Dep="4416789" ID-ZooBank="1A9329DD-74BA-4B44-9008-383D4C88C9D1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609801562384" checkinUser="plazi" docAuthor="Ma, Zijun, Huang, Sunbin &amp; Tian, Mingyi" docDate="2020" docId="1D57B212FFAFFFFC7EB9FE8A1014FE27" docLanguage="en" docName="zootaxa.4861.4.6.pdf" docOrigin="Zootaxa 4861 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sinaphaenops (Sinaphaenops) chengguangyuani Ma &amp; Huang &amp; Tian 2020, n. sp." docType="treatment" docVersion="9" lastPageNumber="587" masterDocId="E16ECA6AFFAEFFFA7E2EFFAD151DFF41" masterDocTitle="Two new species of cavernicolous trechines from central Guizhou and northwestern Guangxi, China (Coleoptera: Carabidae: Trechinae)" masterLastPageNumber="593" masterPageNumber="581" pageNumber="582" updateTime="1698852245223" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="026EC6A7FC121C5744F0DEF4F6A7211A">Two new species of cavernicolous trechines from central Guizhou and northwestern Guangxi, China (Coleoptera: Carabidae: Trechinae)</mods:title>
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<mods:namePart id="D2FEDCBC63DA1EAAD46946FF95A07AD2">Ma, Zijun</mods:namePart>
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<mods:affiliation id="E3808A1EC8417978568D6414204D3732">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, CHINA &amp; 3275124967 @ qq. com; https: // orcid. org / 0000 - 0001 - 8884 - 3376</mods:affiliation>
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<mods:namePart id="E3DA438141772A1CA60F7A703C8F247D">Huang, Sunbin</mods:namePart>
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<mods:affiliation id="BD851B2A0B645197C5453441A7281299">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, CHINA &amp; Mécanismes adaptatifs et évolution (MECADEV), UMR 7179 CNRS-MNHN, Muséum National dHistoire Naturelle, CP 50, 57 rue Cuvier, 75005 Paris, France. huangsunbin @ 163. com; https: // orcid. org / 0000 - 0001 - 8357 - 6651</mods:affiliation>
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<mods:namePart id="7AFF93E5953C04D9A9CF22093C9206AF">Tian, Mingyi</mods:namePart>
<mods:affiliation id="AF1EAEC09ED7436634856DF82E06E070">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, CHINA</mods:affiliation>
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<taxonomicName id="52FE7887FFAFFFFB7EB9FE8A17F7FE03" ID-CoL="8MGFB" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[151,746,295,322]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="chengguangyuani" status="sp. nov." subGenus="Sinaphaenops">
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(
<emphasis id="A78ADF16FFAFFFFB7F7DFE8A14E3FE03" bold="true" box="[339,510,295,322]" italics="true" pageId="1" pageNumber="582">Sinaphaenops</emphasis>
)
<emphasis id="A78ADF16FFAFFFFB7C3EFE8A17F7FE03" bold="true" box="[528,746,295,322]" italics="true" pageId="1" pageNumber="582">chengguangyuani</emphasis>
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(
<figureCitation id="0DC51F81FFAFFFFB7EB1FEE61414FE27" box="[159,265,331,358]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,250,1341,1365]" captionStartId-1="2.[151,250,1967,1991]" captionStartId-2="3.[151,250,1636,1660]" captionStartId-3="4.[151,250,1203,1227]" captionStartId-4="4.[151,250,2007,2031]" captionStartId-5="5.[151,250,1943,1967]" captionTargetBox-1="[152,1433,1396,1936]" captionTargetBox-2="[155,1430,181,1604]" captionTargetBox-3="[321,1274,208,1155]" captionTargetBox-5="[179,1408,181,1920]" captionTargetId-0="figure-23@2.[509,1078,181,1317]" captionTargetId-1="figure-39@2.[151,1436,1390,1944]" captionTargetId-2="figure-20@3.[151,1436,181,1613]" captionTargetId-3="figure-23@4.[295,1292,181,1178]" captionTargetId-4="figure-61@4.[302,1285,1287,1983]" captionTargetId-5="figure-20@5.[179,1408,181,1920]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="4" captionTargetPageId-5="5" captionText-0="FIGURE 1. Habitus of Sinaphaenops (s. str.) chengguangyuani n. sp.." captionText-1="FIGURE 2. Pronotum of Sinaphaenops species (A. S. (s. str.) chengguangyuani n. sp., holotype, male; B. S. (s. str.) lipoi, holotype, male; C. S. (s. str.) banshanicus, holotype, male)." captionText-2="FIGURE 3. Elytra chaetotaxy of Sinaphaenops species (A. S. (s. str.) chengguangyuani n. sp., holotype, male; B. S. (s. str.) lipoi, holotype, male; C. S. (s. str.) banshanicus, holotype, male)." captionText-3="FIGURE 4. Male genitalia of Sinaphaenops (s. str.) chengguangyuani n. sp., holotype (A. median lobe and parameres, lateral view; B. apex of median lobe, dorsal view; C. genital ring, ventral view)." captionText-4="FIGURE 5. Distribution map of the subgenus Sinaphaenops." captionText-5="FIGURE 6. Cave Shuijing Dong, type locality of Sinaphaenops (s. str.) chengguangyuani n. sp. (A. entrance; B, C. cave environment; D. Sinaphaenops (s. str.) chengguangyuani n. sp.)." figureDoi-0="http://doi.org/10.5281/zenodo.4426432" figureDoi-1="http://doi.org/10.5281/zenodo.4416791" figureDoi-2="http://doi.org/10.5281/zenodo.4416793" figureDoi-3="http://doi.org/10.5281/zenodo.4416795" figureDoi-4="http://doi.org/10.5281/zenodo.4426434" figureDoi-5="http://doi.org/10.5281/zenodo.4416797" httpUri-0="https://zenodo.org/record/4426432/files/figure.png" httpUri-1="https://zenodo.org/record/4416791/files/figure.png" httpUri-2="https://zenodo.org/record/4416793/files/figure.png" httpUri-3="https://zenodo.org/record/4416795/files/figure.png" httpUri-4="https://zenodo.org/record/4426434/files/figure.png" httpUri-5="https://zenodo.org/record/4416797/files/figure.png" pageId="1" pageNumber="582">Figs. 16</figureCitation>
)
</paragraph>
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<emphasis id="A78ADF16FFAFFFFB7EB9FE3E141BFEEC" bold="true" box="[151,262,403,429]" pageId="1" pageNumber="582">Material.</emphasis>
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: male,
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,
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,
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,
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,
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&amp;
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,
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,
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,
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in altitude,
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, leg.
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&amp;
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.
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,
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;
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, leg.
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,
<collectorName id="380B66D2FFAFFFFB7A74FE5211A5FD58" box="[1114,1208,511,537]" pageId="1" pageNumber="582">Yi Zhao</collectorName>
,
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,
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&amp;
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, deposited in
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.
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<subSubSection id="DDE4508FFFAFFFF97EE9FDEA11A5F8A4" lastPageId="3" lastPageNumber="584" pageId="1" pageNumber="582" type="description">
<paragraph id="95410304FFAFFFFB7EE9FDEA1728FDB0" blockId="1.[151,1437,403,2014]" pageId="1" pageNumber="582">
<emphasis id="A78ADF16FFAFFFFB7EE9FDEA1444FD20" bold="true" box="[199,345,583,609]" pageId="1" pageNumber="582">Description.</emphasis>
Length:
<quantity id="5206AEE1FFAFFFFB7FEDFDEA1751FD23" box="[451,588,583,610]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="7.1" metricValueMin="6.5" pageId="1" pageNumber="582" unit="mm" value="6.8" valueMax="7.1" valueMin="6.5">6.57.1 mm</quantity>
from apex of right mandible to elytral apex,
<quantity id="5206AEE1FFAFFFFB7A75FDEA11F9FD23" box="[1115,1252,583,610]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.4" metricValueMin="6.1" pageId="1" pageNumber="582" unit="mm" value="6.25" valueMax="6.4" valueMin="6.1">6.16.4 mm</quantity>
from labrum to apex of elytra; width:
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. Habitus as in
<figureCitation id="0DC51F81FFAFFFFB7CDCFDC61625FDC4" box="[754,824,619,645]" captionStart="FIGURE 1" captionStartId="2.[151,250,1341,1365]" captionTargetId="figure-23@2.[509,1078,181,1317]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Sinaphaenops (s. str.) chengguangyuani n. sp.." figureDoi="http://doi.org/10.5281/zenodo.4426432" httpUri="https://zenodo.org/record/4426432/files/figure.png" pageId="1" pageNumber="582">Fig. 1</figureCitation>
. Fore body (including mandibles) longer than elytra, (HLm+PrL)/EL=1.021.03. Overall yellowish brown, with paler tarsi and palps; body moderately shiny, impunctate and glabrous; microsculpture meshes transversely striate on vertex of head and pronotum, moderately transverse on clypeus and frons of head and elytra.
</paragraph>
<paragraph id="95410304FFAFFFFB7EE9FD561495FBA8" blockId="1.[151,1437,403,2014]" pageId="1" pageNumber="582">
Head elongate subtriangular, much longer than wide (HLm/HW=2.842.90, HLl/HW=2.242.26), widest at about 3/7 from base (including mandibles); neck constriction wide, about 0.480.49 times as wide as head; frons depressed, frontal furrows well-marked, not parallel-sided, strongly divergent posteriorly; clypeus transverse, 4-setose; labrum transverse, 6-setose; mandibles stout and long, with obvious bidentate; two pairs of frontal setiferous pores present, the posterior more narrowly spaced than the anterior, at about the 1/3 level between neck constriction and anterior frontal pores; labial suture clearly visible, submentum 10-setose; mentum with two setae near base of median tooth which is thin and bifid; basal foveae concave and separated each other; ligula adnate to paraglossae, 10-setose; palps long, slender and glabrous except for the 2
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labial palpomere which possesses two setae on inner margin and another seta on outer margin; the 2
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labial palpomere 1.2 times longer than the 3
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; the 3
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maxillary palpomere 1.3 times longer than the 4
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. Ventral head with sparse and long setae; suborbital setae present, closer to neck of head than to submentum. Antennae slender, both the 10
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and 11
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antennomeres extending beyond elytral apices; the 1
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antennomere stouter and shorter than others, the 4
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longest, the comparative length ratio of antennomere as follows: 1
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(1.00), 2
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(1.33), 3
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(1.40), 4
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(2.07), 5
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(1.87), 6
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(1.80), 7
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(1.50), 8
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(1.37), 9
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(1.33), 10
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(1.10) and 11
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(1.20).
</paragraph>
<paragraph id="95410304FFAFFFFB7EE9FB5E149FFAA4" blockId="1.[151,1437,403,2014]" pageId="1" pageNumber="582">
Prothorax stouter than in
<taxonomicName id="52FE7887FFAFFFFB7FCEFB5E165FFA4C" authority="Tian et al., 2017" authorityName="Tian" authorityYear="2017" box="[480,834,1267,1294]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="banshanicus">
<emphasis id="A78ADF16FFAFFFFB7FCEFB5E14F3FA4C" box="[480,494,1267,1293]" italics="true" pageId="1" pageNumber="582">S</emphasis>
.
<emphasis id="A78ADF16FFAFFFFB7FD2FB5E1795FA4C" box="[508,648,1267,1293]" italics="true" pageId="1" pageNumber="582">banshanicus</emphasis>
Tian
<emphasis id="A78ADF16FFAFFFFB7CE8FB5917E0FA4C" box="[710,765,1267,1293]" italics="true" pageId="1" pageNumber="582">et al.</emphasis>
, 2017
</taxonomicName>
and
<taxonomicName id="52FE7887FFAFFFFB7D59FB5E11E5FA4C" authority="Chen, Huang &amp; Tian, 2020" authorityName="Chen, Huang &amp; Tian" authorityYear="2020" box="[887,1272,1267,1294]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="lipoi">
<emphasis id="A78ADF16FFAFFFFB7D59FB5E1698FA4C" box="[887,901,1267,1293]" italics="true" pageId="1" pageNumber="582">S</emphasis>
.
<emphasis id="A78ADF16FFAFFFFB7DBCFB5E16D8FA4C" box="[914,965,1267,1293]" italics="true" pageId="1" pageNumber="582">lipoi</emphasis>
<bibRefCitation id="F16F7EF5FFAFFFFB7DE2FB5E11E5FA4C" author="Chen, M. Z. &amp; Huang, S. B. &amp; Tian, M. Y." box="[972,1272,1267,1294]" pageId="1" pageNumber="582" pagination="1 - 8" refId="ref4718" refString="Chen, M. Z., Huang, S. B. &amp; Tian, M. Y. (2020) A new species of the aphaenopsian genus Sinaphaenops Ueno and Wang, with notes on male genitalia of S. orthogenys Ueno (Coleoptera: Carabidae: Trechini). The Coleopterists Bulletin, 74 (2), 1 - 8. https: // doi. org / 10.1649 / 0010 - 065 X- 74.2.343" type="journal article" year="2020">Chen, Huang &amp; Tian, 2020</bibRefCitation>
</taxonomicName>
(
<figureCitation id="0DC51F81FFAFFFFB7B28FB5E1057FA4C" box="[1286,1354,1267,1293]" captionStart="FIGURE 2" captionStartId="2.[151,250,1967,1991]" captionTargetBox="[152,1433,1396,1936]" captionTargetId="figure-39@2.[151,1436,1390,1944]" captionTargetPageId="2" captionText="FIGURE 2. Pronotum of Sinaphaenops species (A. S. (s. str.) chengguangyuani n. sp., holotype, male; B. S. (s. str.) lipoi, holotype, male; C. S. (s. str.) banshanicus, holotype, male)." figureDoi="http://doi.org/10.5281/zenodo.4416791" httpUri="https://zenodo.org/record/4416791/files/figure.png" pageId="1" pageNumber="582">Fig. 2</figureCitation>
), much shorter than head (PrL/HLm=0.470.61, PrL/HLl=0.750.80); distinctly wider than head (PrW/HW=1.241.33); widest at about 2/5 from base; anterior lateral-marginal setae present at about 1/3 distance from front; propleura distinctly expanded and visible from above (PrW/PnW=1.111.14); pronotum faintly wider than head (PnW/ HW=1.091.20), widest at about 1/2 from base; base distinctly wider than front (PbW/PfW=1.241.27); front not stoutly emarginate; lateral margins feebly sinuate, posterior angles acuminate, front angles obtuse and rounded; median line obvious.
</paragraph>
<paragraph id="95410304FFAFFFFB7EE9FA421452F830" blockId="1.[151,1437,403,2014]" pageId="1" pageNumber="582">
Elytra oval-like, much longer than pronotum (EL/PnL=2.552.68), moderately elongate for
<taxonomicName id="52FE7887FFAFFFFB7AE4FA42107AF948" authorityName="Ueno &amp; Wang" authorityYear="1991" box="[1226,1383,1519,1545]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="genus">
<emphasis id="A78ADF16FFAFFFFB7AE4FA42107AF948" box="[1226,1383,1519,1545]" italics="true" pageId="1" pageNumber="582">Sinaphaenops</emphasis>
</taxonomicName>
species, about twice as long as wide (EL/EW=1.942.02), widest at about 3/5 from base, basally narrow and depressed, middle strongly convex; shoulders distinctly oblique, not serrulate, humeral angles broad; lateral margins finely margined and visible throughout, not parallel-sided; elytral striae generally invisible except punctures of the 1
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stria more or less clear at basal part. Chaetotaxy similar in
<taxonomicName id="52FE7887FFAFFFFB7CDDF9D216EEF9D8" box="[755,1011,1663,1690]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="banshanicus">
<emphasis id="A78ADF16FFAFFFFB7CDDF9D2161CF9D8" box="[755,769,1663,1689]" italics="true" pageId="1" pageNumber="582">S</emphasis>
. (
<emphasis id="A78ADF16FFAFFFFB7D37F92D1639F9D8" box="[793,804,1664,1689]" italics="true" pageId="1" pageNumber="582">s</emphasis>
.
<emphasis id="A78ADF16FFAFFFFB7D1CF92D164DF9D8" box="[818,848,1664,1689]" italics="true" pageId="1" pageNumber="582">str</emphasis>
.)
<emphasis id="A78ADF16FFAFFFFB7D49F9D216EEF9D8" box="[871,1011,1663,1689]" italics="true" pageId="1" pageNumber="582">banshanicus</emphasis>
</taxonomicName>
and
<taxonomicName id="52FE7887FFAFFFFB7A07F9D211CDF9D8" authorityName="Chen, Huang &amp; Tian" authorityYear="2020" box="[1065,1232,1663,1690]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="582" phylum="Arthropoda" rank="species" species="lipoi">
<emphasis id="A78ADF16FFAFFFFB7A07F9D2112AF9D8" box="[1065,1079,1663,1689]" italics="true" pageId="1" pageNumber="582">S</emphasis>
. (
<emphasis id="A78ADF16FFAFFFFB7A61F92D1147F9D8" box="[1103,1114,1664,1689]" italics="true" pageId="1" pageNumber="582">s</emphasis>
.
<emphasis id="A78ADF16FFAFFFFB7A46F92D119BF9D8" box="[1128,1158,1664,1689]" italics="true" pageId="1" pageNumber="582">str</emphasis>
.)
<emphasis id="A78ADF16FFAFFFFB7AB3F9D211CDF9D8" box="[1181,1232,1663,1689]" italics="true" pageId="1" pageNumber="582">lipoi</emphasis>
</taxonomicName>
(
<figureCitation id="0DC51F81FFAFFFFB7ACEF9D21039F9D8" box="[1248,1316,1663,1689]" captionStart="FIGURE 3" captionStartId="3.[151,250,1636,1660]" captionTargetBox="[155,1430,181,1604]" captionTargetId="figure-20@3.[151,1436,181,1613]" captionTargetPageId="3" captionText="FIGURE 3. Elytra chaetotaxy of Sinaphaenops species (A. S. (s. str.) chengguangyuani n. sp., holotype, male; B. S. (s. str.) lipoi, holotype, male; C. S. (s. str.) banshanicus, holotype, male)." figureDoi="http://doi.org/10.5281/zenodo.4416793" httpUri="https://zenodo.org/record/4416793/files/figure.png" pageId="1" pageNumber="582">Fig. 3</figureCitation>
): presence of two dorsal pores on approximate position of 3
<superScript id="628BAE4CFFAFFFFB7C94F90E17D5F9F0" attach="left" box="[698,712,1699,1713]" fontSize="6" pageId="1" pageNumber="582">rd</superScript>
stria, at about 3/10 and 1/2 from elytral base, respectively; presence of preapical pore, which is well-marked, closer to suture than to apical margin of elytra; 1
<superScript id="628BAE4CFFAFFFFB7AF6F96A11FEF994" attach="left" box="[1240,1251,1735,1749]" fontSize="6" pageId="1" pageNumber="582">st</superScript>
umbilicate pore of the humeral group roughly shifted to the position of 5
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interval and away from marginal gutter; 2
<superScript id="628BAE4CFFAFFFFB7AD8F946101BF9B8" attach="left" box="[1270,1286,1771,1785]" fontSize="6" pageId="1" pageNumber="582">nd</superScript>
and 3
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pores close to each other, the former near the marginal gutter; 4
<superScript id="628BAE4CFFAFFFFB7D38F8A2163EF85C" attach="left" box="[790,803,1807,1821]" fontSize="6" pageId="1" pageNumber="582">th</superScript>
pore more or less located on the site of 5
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stria, closer to 5
<superScript id="628BAE4CFFAFFFFB7E8BF89E15AFF800" attach="left" box="[165,178,1843,1857]" fontSize="6" pageId="1" pageNumber="582">th</superScript>
pore than to 3
<superScript id="628BAE4CFFAFFFFB7F7AF89E147FF800" attach="none" box="[340,354,1843,1857]" fontSize="6" pageId="1" pageNumber="582">rd</superScript>
; 5
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and 6
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pores of the middle group close to each other, both also widely distant from marginal gutter (
<figureCitation id="0DC51F81FFAFFFFB7EC6F8FA145FF830" box="[232,322,1879,1905]" captionStart="FIGURE 3" captionStartId="3.[151,250,1636,1660]" captionTargetBox="[155,1430,181,1604]" captionTargetId="figure-20@3.[151,1436,181,1613]" captionTargetPageId="3" captionText="FIGURE 3. Elytra chaetotaxy of Sinaphaenops species (A. S. (s. str.) chengguangyuani n. sp., holotype, male; B. S. (s. str.) lipoi, holotype, male; C. S. (s. str.) banshanicus, holotype, male)." figureDoi="http://doi.org/10.5281/zenodo.4416793" httpUri="https://zenodo.org/record/4416793/files/figure.png" pageId="1" pageNumber="582">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="95410304FFAFFFFB7EE9F8D117BBF89C" blockId="1.[151,1437,403,2014]" pageId="1" pageNumber="582">
Legs slender, the 1
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tarsomere shorter than 2
<superScript id="628BAE4CFFAFFFFB7CE1F8D617F3F8D4" attach="none" box="[719,750,1915,1941]" fontSize="6" pageId="1" pageNumber="582">nd</superScript>
4
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tarsomeres together in fore legs, while slightly longer in middle and as long as in hind legs, respectively. The 1
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and 2
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protarsomeres modified in male, each denticulate on inner side of apex, with spongy setae ventrally.
</paragraph>
<caption id="C181538CFFACFFF87EB9FA901679FA14" ID-DOI="http://doi.org/10.5281/zenodo.4426432" ID-Zenodo-Dep="4426432" box="[151,868,1341,1366]" httpUri="https://zenodo.org/record/4426432/files/figure.png" pageId="2" pageNumber="583" targetBox="[518,1062,181,1306]" targetPageId="2">
<paragraph id="95410304FFACFFF87EB9FA901679FA14" blockId="2.[151,868,1341,1366]" box="[151,868,1341,1366]" pageId="2" pageNumber="583">
<emphasis id="A78ADF16FFACFFF87EB9FA90140EFA14" bold="true" box="[151,275,1341,1365]" pageId="2" pageNumber="583">FIGURE 1.</emphasis>
Habitus of
<taxonomicName id="52FE7887FFACFFF87FA4FA931603FA17" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[394,798,1342,1366]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="chengguangyuani" status="sp. nov.">
<emphasis id="A78ADF16FFACFFF87FA4FA931705FA17" box="[394,536,1342,1366]" italics="true" pageId="2" pageNumber="583">Sinaphaenops</emphasis>
(
<emphasis id="A78ADF16FFACFFF87C09FA931740FA14" box="[551,605,1342,1365]" italics="true" pageId="2" pageNumber="583">s. str.</emphasis>
)
<emphasis id="A78ADF16FFACFFF87C44FA931603FA17" box="[618,798,1342,1366]" italics="true" pageId="2" pageNumber="583">chengguangyuani</emphasis>
</taxonomicName>
<emphasis id="A78ADF16FFACFFF87D0BFA931643FA17" bold="true" box="[805,862,1342,1366]" pageId="2" pageNumber="583">
<taxonomicNameLabel id="BCB9626DFFACFFF87D0BFA931643FA17" box="[805,862,1342,1366]" pageId="2" pageNumber="583" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</caption>
<caption id="C181538CFFACFFF87EB9F80217C4F8AA" ID-DOI="http://doi.org/10.5281/zenodo.4416791" ID-Zenodo-Dep="4416791" httpUri="https://zenodo.org/record/4416791/files/figure.png" pageId="2" pageNumber="583" startId="2.[151,250,1967,1991]" targetBox="[152,1433,1396,1936]" targetPageId="2">
<paragraph id="95410304FFACFFF87EB9F80217C4F8AA" blockId="2.[151,1436,1967,2028]" pageId="2" pageNumber="583">
<emphasis id="A78ADF16FFACFFF87EB9F802140EF886" bold="true" box="[151,275,1967,1991]" pageId="2" pageNumber="583">FIGURE 2.</emphasis>
Pronotum of
<taxonomicName id="52FE7887FFACFFF87F8EF8021733F886" authorityName="Ueno &amp; Wang" authorityYear="1991" box="[416,558,1967,1991]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="genus">
<emphasis id="A78ADF16FFACFFF87F8EF8021733F886" box="[416,558,1967,1991]" italics="true" pageId="2" pageNumber="583">Sinaphaenops</emphasis>
</taxonomicName>
species (A.
<taxonomicName id="52FE7887FFACFFF87C85F81D16D5F886" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[683,968,1967,1992]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="chengguangyuani" status="sp. nov.">
<emphasis id="A78ADF16FFACFFF87C85F81D17A5F886" box="[683,696,1968,1991]" italics="true" pageId="2" pageNumber="583">S</emphasis>
. (
<emphasis id="A78ADF16FFACFFF87CE3F81D17CAF886" box="[717,727,1968,1991]" italics="true" pageId="2" pageNumber="583">s</emphasis>
.
<emphasis id="A78ADF16FFACFFF87CCAF81D17E2F886" box="[740,767,1968,1991]" italics="true" pageId="2" pageNumber="583">str</emphasis>
.)
<emphasis id="A78ADF16FFACFFF87D3AF80216D5F886" box="[788,968,1967,1991]" italics="true" pageId="2" pageNumber="583">chengguangyuani</emphasis>
</taxonomicName>
<emphasis id="A78ADF16FFACFFF87DE1F81D1115F889" bold="true" box="[975,1032,1968,1992]" pageId="2" pageNumber="583">
<taxonomicNameLabel id="BCB9626DFFACFFF87DE1F81D1115F889" box="[975,1032,1968,1992]" pageId="2" pageNumber="583" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male; B.
<taxonomicName id="52FE7887FFACFFF87AF9F81D1070F886" authorityName="Chen, Huang &amp; Tian" authorityYear="2020" box="[1239,1389,1967,1992]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="lipoi">
<emphasis id="A78ADF16FFACFFF87AF9F81D11F9F886" box="[1239,1252,1968,1991]" italics="true" pageId="2" pageNumber="583">S</emphasis>
. (
<emphasis id="A78ADF16FFACFFF87AD7F81D101EF886" box="[1273,1283,1968,1991]" italics="true" pageId="2" pageNumber="583">s</emphasis>
.
<emphasis id="A78ADF16FFACFFF87B21F81D1037F886" box="[1295,1322,1968,1991]" italics="true" pageId="2" pageNumber="583">str</emphasis>
.)
<emphasis id="A78ADF16FFACFFF87B11F8021070F886" box="[1343,1389,1967,1991]" italics="true" pageId="2" pageNumber="583">lipoi</emphasis>
</taxonomicName>
, holotype, male; C.
<taxonomicName id="52FE7887FFACFFF87F11F879173BF8AA" box="[319,550,2003,2028]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="583" phylum="Arthropoda" rank="species" species="banshanicus">
<emphasis id="A78ADF16FFACFFF87F11F8791451F8AA" box="[319,332,2004,2027]" italics="true" pageId="2" pageNumber="583">S</emphasis>
. (
<emphasis id="A78ADF16FFACFFF87F4FF8791476F8AA" box="[353,363,2004,2027]" italics="true" pageId="2" pageNumber="583">s</emphasis>
.
<emphasis id="A78ADF16FFACFFF87F56F879148EF8AA" box="[376,403,2004,2027]" italics="true" pageId="2" pageNumber="583">str</emphasis>
.)
<emphasis id="A78ADF16FFACFFF87F89F87E173BF8AA" box="[423,550,2003,2027]" italics="true" pageId="2" pageNumber="583">banshanicus</emphasis>
</taxonomicName>
, holotype, male).
</paragraph>
</caption>
<caption id="C181538CFFADFFF97EB9F9C91630F9E0" ID-DOI="http://doi.org/10.5281/zenodo.4416793" ID-Zenodo-Dep="4416793" httpUri="https://zenodo.org/record/4416793/files/figure.png" pageId="3" pageNumber="584" startId="3.[151,250,1636,1660]" targetBox="[155,1430,181,1604]" targetPageId="3">
<paragraph id="95410304FFADFFF97EB9F9C91630F9E0" blockId="3.[151,1436,1636,1697]" pageId="3" pageNumber="584">
<emphasis id="A78ADF16FFADFFF97EB9F9C91409F93D" bold="true" box="[151,276,1636,1660]" pageId="3" pageNumber="584">FIGURE 3.</emphasis>
Elytra chaetotaxy of
<taxonomicName id="52FE7887FFADFFF97FDDF9C8179CF93C" authorityName="Ueno &amp; Wang" authorityYear="1991" box="[499,641,1637,1661]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="genus">
<emphasis id="A78ADF16FFADFFF97FDDF9C8179CF93C" box="[499,641,1637,1661]" italics="true" pageId="3" pageNumber="584">Sinaphaenops</emphasis>
</taxonomicName>
species (A.
<taxonomicName id="52FE7887FFADFFF97D2DF9C8113EF93C" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[771,1059,1637,1661]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="species" species="chengguangyuani" status="sp. nov.">
<emphasis id="A78ADF16FFADFFF97D2DF9C8160BF93D" box="[771,790,1637,1660]" italics="true" pageId="3" pageNumber="584">S.</emphasis>
(
<emphasis id="A78ADF16FFADFFF97D08F9C8162DF93D" box="[806,816,1637,1660]" italics="true" pageId="3" pageNumber="584">s</emphasis>
.
<emphasis id="A78ADF16FFADFFF97D13F9C81645F93D" box="[829,856,1637,1660]" italics="true" pageId="3" pageNumber="584">str</emphasis>
.)
<emphasis id="A78ADF16FFADFFF97D41F9C8113EF93C" box="[879,1059,1637,1661]" italics="true" pageId="3" pageNumber="584">chengguangyuani</emphasis>
</taxonomicName>
<emphasis id="A78ADF16FFADFFF97A04F9C81178F93C" bold="true" box="[1066,1125,1637,1661]" pageId="3" pageNumber="584">
<taxonomicNameLabel id="BCB9626DFFADFFF97A04F9C81178F93C" box="[1066,1125,1637,1661]" pageId="3" pageNumber="584" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, male; B.
<taxonomicName id="52FE7887FFADFFF97B16F9C815D8F9E0" authorityName="Chen, Huang &amp; Tian" authorityYear="2020" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="species" species="lipoi">
<emphasis id="A78ADF16FFADFFF97B16F9C81058F93D" box="[1336,1349,1637,1660]" italics="true" pageId="3" pageNumber="584">S</emphasis>
. (
<emphasis id="A78ADF16FFADFFF97B75F9C81078F93D" box="[1371,1381,1637,1660]" italics="true" pageId="3" pageNumber="584">s</emphasis>
.
<emphasis id="A78ADF16FFADFFF97B5DF9C81093F93D" box="[1395,1422,1637,1660]" italics="true" pageId="3" pageNumber="584">str</emphasis>
.)
<emphasis id="A78ADF16FFADFFF97EB9F92415D8F9E0" box="[151,197,1673,1697]" italics="true" pageId="3" pageNumber="584">lipoi</emphasis>
</taxonomicName>
, holotype, male; C.
<taxonomicName id="52FE7887FFADFFF97FBAF9241766F9E0" box="[404,635,1673,1697]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="584" phylum="Arthropoda" rank="species" species="banshanicus">
<emphasis id="A78ADF16FFADFFF97FBAF92414BCF9E1" box="[404,417,1673,1696]" italics="true" pageId="3" pageNumber="584">S</emphasis>
. (
<emphasis id="A78ADF16FFADFFF97F98F92414DDF9E1" box="[438,448,1673,1696]" italics="true" pageId="3" pageNumber="584">s</emphasis>
.
<emphasis id="A78ADF16FFADFFF97FE2F92414FAF9E1" box="[460,487,1673,1696]" italics="true" pageId="3" pageNumber="584">str</emphasis>
.)
<emphasis id="A78ADF16FFADFFF97FD2F9241766F9E0" box="[508,635,1673,1697]" italics="true" pageId="3" pageNumber="584">banshanicus</emphasis>
</taxonomicName>
, holotype, male).
</paragraph>
</caption>
<paragraph id="95410304FFADFFF97EE9F96211EFF870" blockId="3.[151,1437,1743,2021]" pageId="3" pageNumber="584">Prosternum with several setae near base and apex, propleura smooth and glabrous; fore, middle and hind coxae with several setae. All ventrites sparsely setose; ventrite IVVI each with 4 paramedial setae, and a few additional setae which are much shorter, ventrite VII with two pairs of setae in male, but three pairs in female.</paragraph>
<paragraph id="95410304FFADFFF97EE9F89611A5F8A4" blockId="3.[151,1437,1743,2021]" pageId="3" pageNumber="584">
Male genitalia (
<figureCitation id="0DC51F81FFADFFF97F59F89614A6F814" box="[375,443,1851,1877]" captionStart="FIGURE 4" captionStartId="4.[151,250,1203,1227]" captionTargetBox="[321,1274,208,1155]" captionTargetId="figure-23@4.[295,1292,181,1178]" captionTargetPageId="4" captionText="FIGURE 4. Male genitalia of Sinaphaenops (s. str.) chengguangyuani n. sp., holotype (A. median lobe and parameres, lateral view; B. apex of median lobe, dorsal view; C. genital ring, ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4416795" httpUri="https://zenodo.org/record/4416795/files/figure.png" pageId="3" pageNumber="584">Fig. 4</figureCitation>
) moderately sclerotized, distinctly arcuate at middle in lateral view, widely membranous on its dorsal part; base large, sagittal aileron large and almost rounded; apex almost straight, and blunt at tip; inner sac armed with a wide copulatory piece, which is about 1/5 as long as median lobe; apical lobe wide, although longer than wide, slightly sinuate on the right side, broadly rounded at apex (
<figureCitation id="0DC51F81FFADFFF97DC4F80A115FF880" box="[1002,1090,1959,1985]" captionStart="FIGURE 4" captionStartId="4.[151,250,1203,1227]" captionTargetBox="[321,1274,208,1155]" captionTargetId="figure-23@4.[295,1292,181,1178]" captionTargetPageId="4" captionText="FIGURE 4. Male genitalia of Sinaphaenops (s. str.) chengguangyuani n. sp., holotype (A. median lobe and parameres, lateral view; B. apex of median lobe, dorsal view; C. genital ring, ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4416795" httpUri="https://zenodo.org/record/4416795/files/figure.png" pageId="3" pageNumber="584">Fig. 4B</figureCitation>
); parameres elongate and welldeveloped, each armed with two long setae at apex (
<figureCitation id="0DC51F81FFADFFF97CCEF8661626F8A4" box="[736,827,1995,2021]" captionStart="FIGURE 4" captionStartId="4.[151,250,1203,1227]" captionTargetBox="[321,1274,208,1155]" captionTargetId="figure-23@4.[295,1292,181,1178]" captionTargetPageId="4" captionText="FIGURE 4. Male genitalia of Sinaphaenops (s. str.) chengguangyuani n. sp., holotype (A. median lobe and parameres, lateral view; B. apex of median lobe, dorsal view; C. genital ring, ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4416795" httpUri="https://zenodo.org/record/4416795/files/figure.png" pageId="3" pageNumber="584">Fig. 4A</figureCitation>
); genital ring triangular (
<figureCitation id="0DC51F81FFADFFF97A7DF86611B1F8A4" box="[1107,1196,1995,2021]" captionStart="FIGURE 4" captionStartId="4.[151,250,1203,1227]" captionTargetBox="[321,1274,208,1155]" captionTargetId="figure-23@4.[295,1292,181,1178]" captionTargetPageId="4" captionText="FIGURE 4. Male genitalia of Sinaphaenops (s. str.) chengguangyuani n. sp., holotype (A. median lobe and parameres, lateral view; B. apex of median lobe, dorsal view; C. genital ring, ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4416795" httpUri="https://zenodo.org/record/4416795/files/figure.png" pageId="3" pageNumber="584">Fig. 4C</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="C181538CFFAAFFFE7EB9FB1E1660FBAE" ID-DOI="http://doi.org/10.5281/zenodo.4416795" ID-Zenodo-Dep="4416795" httpUri="https://zenodo.org/record/4416795/files/figure.png" pageId="4" pageNumber="585" startId="4.[151,250,1203,1227]" targetBox="[321,1274,208,1155]" targetPageId="4">
<paragraph id="95410304FFAAFFFE7EB9FB1E1660FBAE" blockId="4.[151,1436,1203,1263]" pageId="4" pageNumber="585">
<emphasis id="A78ADF16FFAAFFFE7EB9FB1E140EFB8A" bold="true" box="[151,275,1203,1227]" pageId="4" pageNumber="585">FIGURE 4.</emphasis>
Male genitalia of
<taxonomicName id="52FE7887FFAAFFFE7FE0FB1E167BFB8A" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[462,870,1203,1227]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="species" species="chengguangyuani" status="sp. nov.">
<emphasis id="A78ADF16FFAAFFFE7FE0FB1E1741FB8A" box="[462,604,1203,1227]" italics="true" pageId="4" pageNumber="585">Sinaphaenops</emphasis>
(
<emphasis id="A78ADF16FFAAFFFE7C45FB191768FB8A" box="[619,629,1204,1227]" italics="true" pageId="4" pageNumber="585">s</emphasis>
.
<emphasis id="A78ADF16FFAAFFFE7CACFB191780FB8A" box="[642,669,1204,1227]" italics="true" pageId="4" pageNumber="585">str</emphasis>
.)
<emphasis id="A78ADF16FFAAFFFE7C9CFB1E167BFB8A" box="[690,870,1203,1227]" italics="true" pageId="4" pageNumber="585">chengguangyuani</emphasis>
</taxonomicName>
<emphasis id="A78ADF16FFAAFFFE7D43FB1E16BBFB8A" bold="true" box="[877,934,1203,1227]" pageId="4" pageNumber="585">
<taxonomicNameLabel id="BCB9626DFFAAFFFE7D43FB1E16BBFB8A" box="[877,934,1203,1227]" pageId="4" pageNumber="585" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype (A. median lobe and parameres, lateral view; B. apex of median lobe, dorsal view; C. genital ring, ventral view).
</paragraph>
</caption>
<caption id="C181538CFFAAFFFE7EB9F87A161BF8AF" ID-DOI="http://doi.org/10.5281/zenodo.4426434" ID-Zenodo-Dep="4426434" box="[151,774,2007,2031]" httpUri="https://zenodo.org/record/4426434/files/figure.png" pageId="4" pageNumber="585" targetBox="[305,1279,1288,1982]" targetPageId="4">
<paragraph id="95410304FFAAFFFE7EB9F87A161BF8AF" blockId="4.[151,774,2007,2031]" box="[151,774,2007,2031]" pageId="4" pageNumber="585">
<emphasis id="A78ADF16FFAAFFFE7EB9F87A140EF8AE" bold="true" box="[151,275,2007,2031]" pageId="4" pageNumber="585">FIGURE 5.</emphasis>
Distribution map of the subgenus
<taxonomicName id="52FE7887FFAAFFFE7C5FF87A17E2F8AE" authorityName="Ueno &amp; Wang" authorityYear="1991" box="[625,767,2007,2031]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="585" phylum="Arthropoda" rank="genus">
<emphasis id="A78ADF16FFAAFFFE7C5FF87A17E2F8AE" box="[625,767,2007,2031]" italics="true" pageId="4" pageNumber="585">Sinaphaenops</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="C181538CFFABFFFF7EB9F83A17E0F895" ID-DOI="http://doi.org/10.5281/zenodo.4416797" ID-Zenodo-Dep="4416797" httpUri="https://zenodo.org/record/4416797/files/figure.png" pageId="5" pageNumber="586" startId="5.[151,250,1943,1967]" targetBox="[179,1408,181,1920]" targetPageId="5">
<paragraph id="95410304FFABFFFF7EB9F83A17E0F895" blockId="5.[151,1436,1943,2004]" pageId="5" pageNumber="586">
<emphasis id="A78ADF16FFABFFFF7EB9F83A140EF8EE" bold="true" box="[151,275,1943,1967]" pageId="5" pageNumber="586">FIGURE 6.</emphasis>
Cave Shuijing Dong, type locality of
<taxonomicName id="52FE7887FFABFFFF7CBDF8351137F8F1" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[659,1066,1944,1968]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="chengguangyuani" status="sp. nov.">
<emphasis id="A78ADF16FFABFFFF7CBDF835163CF8F1" box="[659,801,1944,1968]" italics="true" pageId="5" pageNumber="586">Sinaphaenops</emphasis>
(
<emphasis id="A78ADF16FFABFFFF7D1EF8351627F8EE" box="[816,826,1944,1967]" italics="true" pageId="5" pageNumber="586">s</emphasis>
.
<emphasis id="A78ADF16FFABFFFF7D68F835167CF8EE" box="[838,865,1944,1967]" italics="true" pageId="5" pageNumber="586">str</emphasis>
.)
<emphasis id="A78ADF16FFABFFFF7D58F8351137F8F1" box="[886,1066,1944,1968]" italics="true" pageId="5" pageNumber="586">chengguangyuani</emphasis>
</taxonomicName>
<emphasis id="A78ADF16FFABFFFF7A1EF8351174F8F1" bold="true" box="[1072,1129,1944,1968]" pageId="5" pageNumber="586">
<taxonomicNameLabel id="BCB9626DFFABFFFF7A1EF8351174F8F1" box="[1072,1129,1944,1968]" pageId="5" pageNumber="586" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(A. entrance; B, C. cave environment; D.
<taxonomicName id="52FE7887FFABFFFF7F36F81117B2F895" authority="Ma &amp; Huang &amp; Tian, 2020" authorityName="Ma &amp; Huang &amp; Tian" authorityYear="2020" box="[280,687,1980,2004]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="586" phylum="Arthropoda" rank="species" species="chengguangyuani" status="sp. nov.">
<emphasis id="A78ADF16FFABFFFF7F36F81114BBF895" box="[280,422,1980,2004]" italics="true" pageId="5" pageNumber="586">Sinaphaenops</emphasis>
(
<emphasis id="A78ADF16FFABFFFF7F9BF81114A2F892" box="[437,447,1980,2003]" italics="true" pageId="5" pageNumber="586">s</emphasis>
.
<emphasis id="A78ADF16FFABFFFF7FE5F81114FBF892" box="[459,486,1980,2003]" italics="true" pageId="5" pageNumber="586">str</emphasis>
.)
<emphasis id="A78ADF16FFABFFFF7FD5F81117B2F895" box="[507,687,1980,2004]" italics="true" pageId="5" pageNumber="586">chengguangyuani</emphasis>
</taxonomicName>
<emphasis id="A78ADF16FFABFFFF7C98F81117F2F895" bold="true" box="[694,751,1980,2004]" pageId="5" pageNumber="586">
<taxonomicNameLabel id="BCB9626DFFABFFFF7C98F81117F2F895" box="[694,751,1980,2004]" pageId="5" pageNumber="586" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</caption>
<subSubSection id="DDE4508FFFA8FFFC7EE9FF3A1077FFB8" pageId="6" pageNumber="587" type="discussion">
<paragraph id="95410304FFA8FFFC7EE9FF3A1077FFB8" blockId="6.[151,1436,151,358]" pageId="6" pageNumber="587">
<emphasis id="A78ADF16FFA8FFFC7EE9FF3A1426FFF0" bold="true" box="[199,315,151,177]" pageId="6" pageNumber="587">Remarks.</emphasis>
This new species belongs to the same lineage as
<taxonomicName id="52FE7887FFA8FFFC7D42FF3A11EBFFF0" box="[876,1270,151,177]" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="banshanicus">
<emphasis id="A78ADF16FFA8FFFC7D42FF3A1114FFF0" box="[876,1033,151,177]" italics="true" pageId="6" pageNumber="587">Sinaphaenops</emphasis>
(
<emphasis id="A78ADF16FFA8FFFC7A34FF3A1138FFF1" box="[1050,1061,151,176]" italics="true" pageId="6" pageNumber="587">s</emphasis>
.
<emphasis id="A78ADF16FFA8FFFC7A1AFF3A114FFFF1" box="[1076,1106,151,176]" italics="true" pageId="6" pageNumber="587">str</emphasis>
.)
<emphasis id="A78ADF16FFA8FFFC7A44FF3A11EBFFF0" box="[1130,1270,151,177]" italics="true" pageId="6" pageNumber="587">banshanicus</emphasis>
</taxonomicName>
and
<taxonomicName id="52FE7887FFA8FFFC7B00FF3A15D7FF94" authorityName="Chen, Huang &amp; Tian" authorityYear="2020" class="Insecta" family="Carabidae" genus="Sinaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="587" phylum="Arthropoda" rank="species" species="lipoi">
<emphasis id="A78ADF16FFA8FFFC7B00FF3A1021FFF0" box="[1326,1340,151,177]" italics="true" pageId="6" pageNumber="587">S</emphasis>
. (
<emphasis id="A78ADF16FFA8FFFC7B7BFF3A107DFFF1" box="[1365,1376,151,176]" italics="true" pageId="6" pageNumber="587">s</emphasis>
.
<emphasis id="A78ADF16FFA8FFFC7B40FF3A1091FFF1" box="[1390,1420,151,176]" italics="true" pageId="6" pageNumber="587">str</emphasis>
.)
<emphasis id="A78ADF16FFA8FFFC7EB9FF1615D7FF94" box="[151,202,187,213]" italics="true" pageId="6" pageNumber="587">lipoi</emphasis>
</taxonomicName>
, in possessing anterior lateral-marginal setae on pronotum and preapical dorsal pores on elytra. It differs from the above two species by the shorter pronotum and presence of a distinct sagittal aileron on the male genitalia.
</paragraph>
</subSubSection>
<subSubSection id="DDE4508FFFA8FFFC7EE9FEAE11A8FE5C" box="[199,1205,259,285]" pageId="6" pageNumber="587" type="etymology">
<paragraph id="95410304FFA8FFFC7EE9FEAE11A8FE5C" blockId="6.[151,1436,151,358]" box="[199,1205,259,285]" pageId="6" pageNumber="587">
<emphasis id="A78ADF16FFA8FFFC7EE9FEAE1453FE5C" bold="true" box="[199,334,259,285]" pageId="6" pageNumber="587">Etymology.</emphasis>
Dedicated to Mr. Guangyuan Cheng, who discovered this interesting species.
</paragraph>
</subSubSection>
<subSubSection id="DDE4508FFFA8FFFC7EE9FE8B1014FE27" pageId="6" pageNumber="587" type="distribution">
<paragraph id="95410304FFA8FFFC7EE9FE8B1014FE27" blockId="6.[151,1436,151,358]" pageId="6" pageNumber="587">
<emphasis id="A78ADF16FFA8FFFC7EE9FE8B147CFE01" bold="true" box="[199,353,294,320]" pageId="6" pageNumber="587">Distribution.</emphasis>
<collectingRegion id="573ACDE6FFA8FFFC7F48FE8A14DBFE00" box="[358,454,295,321]" country="China" name="Guizhou" pageId="6" pageNumber="587">Guizhou</collectingRegion>
(Guiyang). Known only from a limestone cave called Shuijing Dong (
<figureCitation id="0DC51F81FFA8FFFC7AFFFE8A1009FE00" box="[1233,1300,294,321]" captionStart="FIGURE 5" captionStartId="4.[151,250,2007,2031]" captionTargetId="figure-61@4.[302,1285,1287,1983]" captionTargetPageId="4" captionText="FIGURE 5. Distribution map of the subgenus Sinaphaenops." figureDoi="http://doi.org/10.5281/zenodo.4426434" httpUri="https://zenodo.org/record/4426434/files/figure.png" pageId="6" pageNumber="587">Fig. 5</figureCitation>
). The single beetle was collected on wet floor inside the cave, a place about 250 meters from the entrance (
<figureCitation id="0DC51F81FFA8FFFC7A98FEE611E7FE27" box="[1206,1274,331,358]" captionStart="FIGURE 6" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-20@5.[179,1408,181,1920]" captionTargetPageId="5" captionText="FIGURE 6. Cave Shuijing Dong, type locality of Sinaphaenops (s. str.) chengguangyuani n. sp. (A. entrance; B, C. cave environment; D. Sinaphaenops (s. str.) chengguangyuani n. sp.)." figureDoi="http://doi.org/10.5281/zenodo.4416797" httpUri="https://zenodo.org/record/4416797/files/figure.png" pageId="6" pageNumber="587">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>