269 lines
32 KiB
XML
269 lines
32 KiB
XML
<document id="FB24D2BDEEE282A9EDE355A55EEAC876" ID-DOI="10.5281/zenodo.182839" ID-GBIF-Dataset="beac9aa2-83cc-4dd0-bd88-9dfc232b4830" ID-ISSN="1175-5326" ID-Zenodo-Dep="182839" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459878942779" checkinUser="plazi" docAuthor="García-Alzate, Carlos A., Román-Valencia, César & Taphorn, Donald C." docDate="2008" docId="03ED879D542EFFECFF45FACAFE6CFC05" docLanguage="en" docName="zt01813p050.pdf" docOrigin="Zootaxa 1813" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hyphessobrycon oritoensis García-Alzate, Román-Valencia & Taphorn, 2008, new species" docType="treatment" docVersion="7" lastPageNumber="47" masterDocId="FFD4FFE5542FFFE9FFD2FF88FFB4FFFF" masterDocTitle="Hyphessobrycon oritoensis (Characiformes: Characidae), a new species from the Putumayo River drainage, Colombian Amazon" masterLastPageNumber="50" masterPageNumber="42" pageNumber="43" updateTime="1698227283718" updateUser="plazi">
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<mods:mods id="71CD02A843F202254065E62D79584880" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="960ABF98B5482D3C291867D403022E17">
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<mods:title id="A768D142710D4E2CE5A961E91A3047FB">Hyphessobrycon oritoensis (Characiformes: Characidae), a new species from the Putumayo River drainage, Colombian Amazon</mods:title>
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<mods:name id="D7EC37573BFB85DFAB77483FAC6713EB" type="personal">
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<mods:role id="E8B7F31C6D42475DAE55BEFA86818388">
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<mods:roleTerm id="BD034CA89BCB9474E43E5EA4B4C1B28D">Author</mods:roleTerm>
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<mods:namePart id="33D6A60375D2B34441D49DE71054DE04">García-Alzate, Carlos A.</mods:namePart>
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<mods:roleTerm id="3BE22936581DAC0F1D9CF6B824845556">Author</mods:roleTerm>
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<mods:namePart id="16E1CFDDF6899DD9B0531403A062F5EA">Román-Valencia, César</mods:namePart>
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</mods:name>
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<mods:name id="FD847B63DE67B85B8432834CCFEC4D2F" type="personal">
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<mods:roleTerm id="02758A0B8351677EFA515BDC807F2317">Author</mods:roleTerm>
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<mods:namePart id="B4E116F3C0DC48F3AE189A7FFA75770F">Taphorn, Donald C.</mods:namePart>
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<mods:typeOfResource id="61865A7FB79E29ECB81BB636A53C2DBC">text</mods:typeOfResource>
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<mods:title id="E432828DD075D791035D3427A1D77967">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="49840D777BEE268781B2C916BD9400DB">
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<mods:date id="995A58418C089EF1E72AA2101D396B56">2008</mods:date>
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<mods:detail id="32E418170B4575773C5B02970CBADC81" type="volume">
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<mods:number id="49C222AE4E27B5850FE6A103F4C89279">1813</mods:number>
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<mods:classification id="0AB70AA16C14115CBC33300C614BCB64">journal article</mods:classification>
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<mods:identifier id="602F1406CAFB7A20310D2C49600A6D1C" type="DOI">10.5281/zenodo.182839</mods:identifier>
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<mods:identifier id="1288F826906C2D17C05B0EB788F0AB2D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="52FFFC8F9EDEEF3C8FBF51D8C6ECBFDF" type="Zenodo-Dep">182839</mods:identifier>
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<treatment id="03ED879D542EFFECFF45FACAFE6CFC05" ID-DOI="http://doi.org/10.5281/zenodo.6232853" ID-GBIF-Taxon="119370550" ID-Zenodo-Dep="6232853" LSID="urn:lsid:plazi:treatment:03ED879D542EFFECFF45FACAFE6CFC05" httpUri="http://treatment.plazi.org/id/03ED879D542EFFECFF45FACAFE6CFC05" lastPageId="5" lastPageNumber="47" pageId="1" pageNumber="43">
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<subSubSection id="C35E6500542EFFE8FF45FACAFEDEFA80" pageId="1" pageNumber="43" type="nomenclature">
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<paragraph id="8BFB368B542EFFE8FF45FACAFDCEFAA3" blockId="1.[151,634,1346,1407]" box="[151,634,1346,1372]" pageId="1" pageNumber="43">
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<heading id="D0B381E7542EFFE8FF45FACAFDCEFAA3" bold="true" box="[151,634,1346,1372]" fontSize="11" level="1" pageId="1" pageNumber="43" reason="1">
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<emphasis id="B930EA99542EFFE8FF45FACAFDCEFAA3" bold="true" box="[151,634,1346,1372]" pageId="1" pageNumber="43">
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<emphasis id="B930EA99542EFFE8FF45FACAFE57FAA3" bold="true" box="[151,483,1346,1372]" italics="true" pageId="1" pageNumber="43">
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<taxonomicName id="4C444D08542EFFE8FF45FACAFE68FAA3" ID-CoL="3NRZ7" box="[151,476,1346,1372]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="43" phylum="Chordata" rank="species" species="oritoensis" status="sp. nov.">Hyphessobrycon oritoensis</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A20357E2542EFFE8FE38FACAFDCEFAA3" box="[490,634,1346,1372]" pageId="1" pageNumber="43" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BFB368B542EFFE8FF45FAEDFEDEFA80" blockId="1.[151,634,1346,1407]" box="[151,362,1381,1407]" pageId="1" pageNumber="43">
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<tableCitation id="C6C60330542EFFE8FF45FAEDFF5AFA80" box="[151,238,1381,1407]" captionStart="TABLE 1" captionStartId="2.[151,239,151,175]" captionText="TABLE 1. Morphometric and meristic data for Hyphessobrycon oritoensis n. sp. Standard and total length in mm. Means given in parentheses. Standard deviation = SD." pageId="1" pageNumber="43">Table 1</tableCitation>
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,
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<figureCitation id="137F2A0E542EFFE8FF2EFAEDFEDEFA80" box="[252,362,1381,1407]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,255,847,871]" captionStartId-1="4.[151,255,1855,1879]" captionStartId-2="5.[151,255,1735,1759]" captionTargetBox-0="[263,1322,357,820]" captionTargetBox-1="[159,1432,204,1812]" captionTargetBox-2="[231,1355,1065,1711]" captionTargetId-0="figure@3.[263,1323,357,820]" captionTargetId-1="figure@4.[151,1436,194,1831]" captionTargetId-2="figure@5.[225,1358,1061,1719]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Hyphessobrycon oritoensis. Holotype; 34.63 mm SL; Colombia, Putumayo, Rio Orito, Amazon basin." captionText-1="FIGURE 2. Upper (A) and lower jaws (B) of Hyphessobrycon oritoensis, IUQ 1575, paratype." captionText-2="FIGURE 3. Principal components analyses (PCA) (component 1 in the axis X, component 2 in the axis Y) of morphometics data in Hyphessobrycon oritoensis n. sp (+), H. poeciloides (Π), H. heterorhabdus (), H. vilmae (), H. melanostichos (), H. diancistrus (*) and H. notidanos (Ϛ)" httpUri-0="https://zenodo.org/record/182840/files/figure.png" httpUri-1="https://zenodo.org/record/182841/files/figure.png" httpUri-2="https://zenodo.org/record/182842/files/figure.png" pageId="1" pageNumber="43">Figs. 1–3</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C35E6500542EFFE8FF45FA3DFB4DF940" pageId="1" pageNumber="43" type="materials_examined">
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<paragraph id="8BFB368B542EFFE8FF45FA3DFF5FF9E0" blockId="1.[151,1436,1461,2007]" pageId="1" pageNumber="43">
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<emphasis id="B930EA99542EFFE8FF45FA3DFEA6FA30" bold="true" box="[151,274,1461,1487]" pageId="1" pageNumber="43">
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<typeStatus id="54FF8829542EFFE8FF45FA3DFEBAFA30" box="[151,270,1461,1487]" pageId="1" pageNumber="43" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<collectionCode id="ED55AE4E542EFFE8FECDFA3DFEE7FA30" box="[287,339,1461,1487]" httpUri="http://grbio.org/cool/8xqv-ibyz" name="Laboratorio de Ictiologia" pageId="1" pageNumber="43">IUQ</collectionCode>
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1574,
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<quantity id="4CBC9B6E542EFFE8FE7CFA3DFD9FFA30" box="[430,555,1461,1487]" metricMagnitude="-2" metricUnit="m" metricValue="2.463" pageId="1" pageNumber="43" unit="mm" value="24.63">24.63 mm</quantity>
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SL;
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<collectingCountry id="F353761B542EFFE8FDBCFA3DFD51FA30" box="[622,741,1461,1487]" name="Colombia" pageId="1" pageNumber="43">Colombia</collectingCountry>
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, Putumayo, Orito, Quebrada La Palma, finca La Palma, Vereda Calimonte,
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<date id="FFFA104B542EFFE8FEAEFA55FDADFA08" box="[380,537,1501,1527]" pageId="1" pageNumber="43" value="1998-06-29">29 Jun. 1998</date>
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. Approximately
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<geoCoordinate id="EE70504C542EFFE8FD33FA55FCF3FA08" box="[737,839,1501,1527]" direction="north" orientation="latitude" pageId="1" pageNumber="43" precision="925" value="0.68333334">0º 41’ N</geoCoordinate>
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&
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<geoCoordinate id="EE70504C542EFFE8FCA3FA55FC59FA08" box="[881,1005,1501,1527]" direction="west" orientation="longitude" pageId="1" pageNumber="43" precision="925" value="-75.86667">75º 52’ W</geoCoordinate>
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. C. Román–Valencia, J. Tovar & H. Hoyos.
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</paragraph>
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<paragraph id="8BFB368B542EFFE8FF17F9A5FB4DF940" blockId="1.[151,1436,1461,2007]" pageId="1" pageNumber="43">
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<emphasis id="B930EA99542EFFE8FF17F9A5FEF8F9B8" bold="true" box="[197,332,1581,1607]" pageId="1" pageNumber="43">
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<typeStatus id="54FF8829542EFFE8FF17F9A5FEF2F9B8" box="[197,326,1581,1607]" pageId="1" pageNumber="43" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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<collectionCode id="ED55AE4E542EFFE8FE87F9A5FE3DF9B8" box="[341,393,1581,1607]" httpUri="http://grbio.org/cool/8xqv-ibyz" name="Laboratorio de Ictiologia" pageId="1" pageNumber="43">IUQ</collectionCode>
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139, (6 ex.) collected with the
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<typeStatus id="54FF8829542EFFE8FD2DF9A5FCDEF9B8" box="[767,874,1581,1607]" pageId="1" pageNumber="43" type="holotype">holotype</typeStatus>
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.
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<collectionCode id="ED55AE4E542EFFE8FCA7F9A5FC1DF9B8" box="[885,937,1581,1607]" httpUri="http://grbio.org/cool/8xqv-ibyz" name="Laboratorio de Ictiologia" pageId="1" pageNumber="43">IUQ</collectionCode>
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1575, (2 ex.) (C. & S.) collected with the
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<typeStatus id="54FF8829542EFFE8FF45F9DDFEB5F990" box="[151,257,1621,1647]" pageId="1" pageNumber="43" type="holotype">holotype</typeStatus>
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;
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<collectionCode id="ED55AE4E542EFFE8FEDDF9DDFECFF990" box="[271,379,1621,1647]" httpUri="http://grbio.org/cool/7nxg-etxt" name="Museo de Biologia de la Universidad Central de Venezuela" pageId="1" pageNumber="43">MBUCV</collectionCode>
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–
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<accessionNumber id="9417AB68542EFFE8FE58F9DDFE40F990" box="[394,500,1621,1647]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V33737" pageId="1" pageNumber="43" type="EnaNcbi">V 33737</accessionNumber>
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, (2 ex.) collected with the
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<typeStatus id="54FF8829542EFFE8FCE8F9DDFC10F990" box="[826,932,1621,1647]" pageId="1" pageNumber="43" type="holotype">holotype</typeStatus>
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;
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<collectionCode id="ED55AE4E542EFFE8FC60F9DDFBBEF990" box="[946,1034,1621,1647]" httpUri="http://grbio.org/cool/tyz9-grz8" name="Museo de Ciencias Naturales de la UNELLEZ en Guanare" pageId="1" pageNumber="43">MCNG</collectionCode>
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55844, (2 ex.) collected with the
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<typeStatus id="54FF8829542EFFE8FF45F9F5FEB5F968" box="[151,257,1661,1687]" pageId="1" pageNumber="43" type="holotype">holotype</typeStatus>
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;
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<collectingCountry id="F353761B542EFFE8FEDFF9F5FE30F968" box="[269,388,1661,1687]" name="Colombia" pageId="1" pageNumber="43">Colombia</collectingCountry>
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, Putumayo, Orito, Quebrada La Palma, finca La Palma, Vereda Calimonte,
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<date id="FFFA104B542EFFE8FB2CF9F5FA2CF968" box="[1278,1432,1661,1687]" pageId="1" pageNumber="43" value="1998-06-29">29 Jun. 1998</date>
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. Approx.
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<geoCoordinate id="EE70504C542EFFE8FF2DF92DFED5F940" box="[255,353,1701,1727]" direction="north" orientation="latitude" pageId="1" pageNumber="43" precision="925" value="0.68333334">0º 41’ N</geoCoordinate>
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&
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<geoCoordinate id="EE70504C542EFFE8FE55F92DFDB4F940" box="[391,512,1701,1727]" direction="west" orientation="longitude" pageId="1" pageNumber="43" precision="925" value="-75.86667">75º 52’ W</geoCoordinate>
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. C. Román–Valencia, J. Tovar & H. Hoyos.
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<quantity id="4CBC9B6E542EFFE8FBD4F92DFB7DF940" box="[1030,1225,1701,1727]" metricMagnitude="-2" metricUnit="m" metricValue="3.448" metricValueMax="4.277" metricValueMin="2.619" pageId="1" pageNumber="43" unit="mm" value="34.48" valueMax="42.77" valueMin="26.19">26.19-42.77 mm</quantity>
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SL.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35E6500542EFFE8FF17F945FBA1F828" pageId="1" pageNumber="43" type="diagnosis">
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<paragraph id="8BFB368B542EFFE8FF17F945FBA1F828" blockId="1.[151,1436,1461,2007]" pageId="1" pageNumber="43">
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<emphasis id="B930EA99542EFFE8FF17F945FEF3F918" bold="true" box="[197,327,1741,1767]" pageId="1" pageNumber="43">Diagnosis:</emphasis>
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<taxonomicName id="4C444D08542EFFE8FE81F945FD21F919" box="[339,661,1741,1766]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="43" phylum="Chordata" rank="species" species="oritoensis">
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<emphasis id="B930EA99542EFFE8FE81F945FD21F919" box="[339,661,1741,1766]" italics="true" pageId="1" pageNumber="43">Hyphessobrycon oritoensis</emphasis>
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</taxonomicName>
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is distinguished from all congeners by having three simple and eight branched rays in the dorsal fin (vs. two simple and nine branched rays, except
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<taxonomicName id="4C444D08542EFFE8FB51F97DFAA8F8F1" box="[1155,1308,1781,1806]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="43" phylum="Chordata" rank="species" species="notidanos">
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<emphasis id="B930EA99542EFFE8FB51F97DFAA8F8F1" box="[1155,1308,1781,1806]" italics="true" pageId="1" pageNumber="43">H. notidanos</emphasis>
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</taxonomicName>
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which has iii, 8); four simple and 26–27 branched anal–fin rays (vs. three simple and 17–24 branched rays); 19 teeth on the dentary (vs. 6–12); 35 lateral line scales (vs. 31–34, except for
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<taxonomicName id="4C444D08542EFFE8FC16F8CDFB27F8A1" box="[964,1171,1861,1886]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="43" phylum="Chordata" rank="species" species="heterorhabdus">
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<emphasis id="B930EA99542EFFE8FC16F8CDFB27F8A1" box="[964,1171,1861,1886]" italics="true" pageId="1" pageNumber="43">H. heterorhabdus</emphasis>
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</taxonomicName>
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which has 37); 10-11 pored lateral–line scales (vs. 8–9); seven longitudinal scale rows from the lateral line to the dorsal fin (vs. five, except in
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<taxonomicName id="4C444D08542EFFE8FEDBF81DFE62F851" box="[265,470,1941,1966]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="43" phylum="Chordata" rank="species" species="heterorhabdus">
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<emphasis id="B930EA99542EFFE8FEDBF81DFE62F851" box="[265,470,1941,1966]" italics="true" pageId="1" pageNumber="43">H. heterorhabdus</emphasis>
|
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</taxonomicName>
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with seven) and fourteen predorsal scales (vs. 10–11).
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<taxonomicName id="4C444D08542EFFE8FB8FF81DFA28F851" box="[1117,1436,1941,1966]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="43" phylum="Chordata" rank="species" species="oritoensis">
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<emphasis id="B930EA99542EFFE8FB8FF81DFA28F851" box="[1117,1436,1941,1966]" italics="true" pageId="1" pageNumber="43">Hyphessobrycon oritoensis</emphasis>
|
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</taxonomicName>
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is distinguished in detail from
|
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<taxonomicName id="4C444D08542EFFE8FE2EF835FD27F829" box="[508,659,1981,2006]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="1" pageNumber="43" phylum="Chordata" rank="species" species="notidanos">
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<emphasis id="B930EA99542EFFE8FE2EF835FD27F829" box="[508,659,1981,2006]" italics="true" pageId="1" pageNumber="43">H. notidanos</emphasis>
|
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</taxonomicName>
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in the “Remarks” section below.
|
||
</paragraph>
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||
</subSubSection>
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||
<caption id="DF3B6603542DFFEBFF45FF1FFD6DFF2E" pageId="2" pageNumber="44">
|
||
<paragraph id="8BFB368B542DFFEBFF45FF1FFD6DFF2E" blockId="2.[151,1437,151,209]" pageId="2" pageNumber="44">
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||
<emphasis id="B930EA99542DFFEBFF45FF1FFEB2FF50" bold="true" box="[151,262,151,175]" pageId="2" pageNumber="44">TABLE 1</emphasis>
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||
. Morphometric and meristic data for
|
||
<taxonomicName id="4C444D08542DFFEBFD62FF10FC61FF50" box="[688,981,152,175]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="2" pageNumber="44" phylum="Chordata" rank="species" species="oritoensis" status="sp. nov.">
|
||
<emphasis id="B930EA99542DFFEBFD62FF10FC61FF50" box="[688,981,152,175]" italics="true" pageId="2" pageNumber="44">Hyphessobrycon oritoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20357E2542DFFEBFC32FF1FFB94FF50" box="[992,1056,151,175]" pageId="2" pageNumber="44" rank="species">
|
||
<emphasis id="B930EA99542DFFEBFC32FF1FFB94FF50" bold="true" box="[992,1056,151,175]" pageId="2" pageNumber="44">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Standard and total length in mm. Means given in parentheses. Standard deviation = SD.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35E6500542DFFECFF17F839FEA5FF25" lastPageId="5" lastPageNumber="47" pageId="2" pageNumber="44" type="description">
|
||
<paragraph id="8BFB368B542DFFEAFF17F839FDDEFED5" blockId="2.[151,1436,1969,2036]" lastBlockId="3.[151,1436,152,298]" lastPageId="3" lastPageNumber="45" pageId="2" pageNumber="44">
|
||
<emphasis id="B930EA99542DFFEBFF17F839FED4F834" bold="true" box="[197,352,1969,1995]" pageId="2" pageNumber="44">Description:</emphasis>
|
||
Morfometric and meristic data are given in
|
||
<tableCitation id="C6C60330542DFFEBFCA2F83AFC7DF833" box="[880,969,1970,1996]" captionStart="TABLE 1" captionStartId="2.[151,239,151,175]" captionText="TABLE 1. Morphometric and meristic data for Hyphessobrycon oritoensis n. sp. Standard and total length in mm. Means given in parentheses. Standard deviation = SD." pageId="2" pageNumber="44">Table 1</tableCitation>
|
||
. Body short and deep, dorsal profile of head and body oblique from snout to supraoccipital, convex from there to dorsal fin origin, and from the last dorsal–fin ray to the base of the caudal fin. Ventral profile of body convex from snout to base of anal fin, more pronounced behind pectorals. Head and snout short; mandibles equal; mouth subterminal; lips soft and flexible, not covering the external row of premaxillary teeth; ventral part of the upper jaw flat; posterior end of maxilla passing anterior border of orbit.
|
||
</paragraph>
|
||
<caption id="DF3B6603542CFFEAFF45FCC7FAD7FC99" box="[151,1379,847,871]" httpUri="https://zenodo.org/record/182840/files/figure.png" pageId="3" pageNumber="45" targetBox="[263,1322,357,820]" targetPageId="3">
|
||
<paragraph id="8BFB368B542CFFEAFF45FCC7FAD7FC99" blockId="3.[151,1379,847,871]" box="[151,1379,847,871]" pageId="3" pageNumber="45">
|
||
<emphasis id="B930EA99542CFFEAFF45FCC7FEADFC98" bold="true" box="[151,281,847,871]" pageId="3" pageNumber="45">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C444D08542CFFEAFEF2FCC7FDF5FC99" box="[288,577,847,870]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="3" pageNumber="45" phylum="Chordata" rank="species" species="oritoensis">
|
||
<emphasis id="B930EA99542CFFEAFEF2FCC7FDF5FC99" box="[288,577,847,870]" italics="true" pageId="3" pageNumber="45">Hyphessobrycon oritoensis</emphasis>
|
||
</taxonomicName>
|
||
. Holotype; 34.63 mm SL; Colombia, Putumayo, Rio Orito, Amazon basin.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFB368B542CFFEAFF17FC12FE3EFB83" blockId="3.[151,1436,922,2028]" pageId="3" pageNumber="45">
|
||
Premaxillary with a long lateral process, pointed and with two rows of teeth (
|
||
<figureCitation id="137F2A0E542CFFEAFB95FC12FB3AFC4B" box="[1095,1166,922,948]" captionStart="FIGURE 2" captionStartId="4.[151,255,1855,1879]" captionTargetBox="[159,1432,204,1812]" captionTargetId="figure@4.[151,1436,194,1831]" captionTargetPageId="4" captionText="FIGURE 2. Upper (A) and lower jaws (B) of Hyphessobrycon oritoensis, IUQ 1575, paratype." httpUri="https://zenodo.org/record/182841/files/figure.png" pageId="3" pageNumber="45">Fig. 2</figureCitation>
|
||
); the external row with 3–4 tricuspid teeth arranged in zig zag; the internal row with 5–6 tricuspid teeth, diminishing gradually in size from inner to outer laterally. Maxilla long with 5–6 tricuspid teeth, and with posterior end reaching the posterior edge of second infraorbital; maxilla with two small foramina near upper margin of the tooth series. Two pentacuspid teeth in front of dentary, followed by three tricuspid lateral teeth larger than the rest, followed by 14 small conic teeth.
|
||
</paragraph>
|
||
<paragraph id="8BFB368B542CFFEAFF17FB02FA8BFB33" blockId="3.[151,1436,922,2028]" pageId="3" pageNumber="45">Six infraorbitals, the third longest in width, wide but not in contact with the sensory canal of preopercle; fourth infraorbital the smallest of series. Antorbital with anterior border concave. Supraorbital absent.</paragraph>
|
||
<paragraph id="8BFB368B542CFFEAFF17FB52FE7AF9CB" blockId="3.[151,1436,922,2028]" pageId="3" pageNumber="45">Rhinosphenoid heavily ossified, united with orbitosphenoid by a band of cartilage that extends above its posterior border; cartilage band extends to the dorso–anterior end and is in contact with the postero–dorsal margin of the prevomer. Orbitosphenoid heavily ossified with a round apophysis on anterior part. Paraesphenoid not divided, united to ventral surface of vomer; posterior end of parasphenoid in contact with prootic and basioccipital through a band of cartilage. Metapterygoid with upper margin wide and two crests, a small foramen in postero–medial region, with a band of cartilage along anterior and posterior edges. Ectopterygoid elongate and wide, in contact via cartilage with the quadrate. Nasal bone present. Basihyal cartilaginous and divided in two. Pharyngeal plate elongate and curved, having cartilage on the dorsal and antero–ventral edges. 16 gill rakers on first arch.</paragraph>
|
||
<paragraph id="8BFB368B542CFFEAFF17F9CAFC89F8DB" blockId="3.[151,1436,922,2028]" pageId="3" pageNumber="45">Total number of vertebrae 34–35. Proximal dorsal pterygiophores inserted between neural spines 10 and 17; 28 radial pterygiophores in anal fin. Four to five supraneurals present with cartilage on their dorsal and ventral ends. Pectoral girdle with a pointed dorsal elongation. Cleithrum elongate, widening near posterior edge and with the margin wavy, located below the ventral edge of the opercle. Scapula united to internal surface of cleithrum by cartilage. Postcleithrum 3 elongate and curved with a rounded laminar bony prolongation of its postero–medial edge. Four proximal radials present.</paragraph>
|
||
<paragraph id="8BFB368B542CFFEAFF17F8BAFDACF83B" blockId="3.[151,1436,922,2028]" pageId="3" pageNumber="45">Edge of dorsal fin oblique. Pelvic fin short, tip not reaching anal-fin origin. Pelvic bone elongate and straight, parallel to medial area of body with cartilage on anterior and postero-lateral edges; ischial process short, curved, with pointed apophysis cartilaginous. Caudal fin forked, with long pointed lobes, scaleless. Principal caudal rays 9/8 (n=14).</paragraph>
|
||
<paragraph id="8BFB368B542CFFEAFF17F859FC59F813" blockId="3.[151,1436,922,2028]" box="[197,1005,2001,2028]" pageId="3" pageNumber="45">
|
||
<emphasis id="B930EA99542CFFEAFF17F859FE74F814" bold="true" box="[197,448,2001,2027]" pageId="3" pageNumber="45">Sexual dimorphism.</emphasis>
|
||
No apparent sexual dimorphism was observed.
|
||
</paragraph>
|
||
<caption id="DF3B6603542BFFEDFF45F8B7FB3BF8A9" box="[151,1167,1855,1879]" httpUri="https://zenodo.org/record/182841/files/figure.png" pageId="4" pageNumber="46" targetBox="[159,1432,204,1812]" targetPageId="4">
|
||
<paragraph id="8BFB368B542BFFEDFF45F8B7FB3BF8A9" blockId="4.[151,1167,1855,1879]" box="[151,1167,1855,1879]" pageId="4" pageNumber="46">
|
||
<emphasis id="B930EA99542BFFEDFF45F8B7FEADF8A8" bold="true" box="[151,281,1855,1879]" pageId="4" pageNumber="46">FIGURE 2.</emphasis>
|
||
Upper (A) and lower jaws (B) of
|
||
<taxonomicName id="4C444D08542BFFEDFD56F8C8FC11F8A8" box="[644,933,1856,1879]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="4" pageNumber="46" phylum="Chordata" rank="species" species="oritoensis">
|
||
<emphasis id="B930EA99542BFFEDFD56F8C8FC11F8A8" box="[644,933,1856,1879]" italics="true" pageId="4" pageNumber="46">Hyphessobrycon oritoensis</emphasis>
|
||
</taxonomicName>
|
||
, IUQ 1575, paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFB368B542BFFECFF17F80FFEA5FF25" blockId="4.[151,1436,1927,2034]" lastBlockId="5.[151,1437,152,1018]" lastPageId="5" lastPageNumber="47" pageId="4" pageNumber="46">
|
||
<emphasis id="B930EA99542BFFEDFF17F80FFE23F85E" bold="true" box="[197,407,1927,1953]" pageId="4" pageNumber="46">Color in alcohol:</emphasis>
|
||
Body light yellow, lateral portion of body with dark band from behind humeral spot to tips of central caudal fin rays, wider beneath dorsal fin origin. Humeral spot dark. Guanine present dorsally and laterally in many specimens. Posterior edge of scales on dorsum and upper side black. Tip of caudal lobes dark. Dorsal, anal, pectoral and pelvic fins hyaline. Anal fin lightly pigmented at tips of rays. Head darker than body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35E6500542AFFECFF17FF6FFCB8FEFD" box="[197,780,231,258]" pageId="5" pageNumber="47" type="distribution">
|
||
<paragraph id="8BFB368B542AFFECFF17FF6FFCB8FEFD" blockId="5.[151,1437,152,1018]" box="[197,780,231,258]" pageId="5" pageNumber="47">
|
||
<emphasis id="B930EA99542AFFECFF17FF6FFEDCFEFE" bold="true" box="[197,360,231,257]" pageId="5" pageNumber="47">Distribution:</emphasis>
|
||
Only known from the
|
||
<typeStatus id="54FF8829542AFFECFDA6FF60FD13FEFD" box="[628,679,232,258]" pageId="5" pageNumber="47">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35E6500542AFFECFF17FE87FF76FEAD" pageId="5" pageNumber="47" type="etymology">
|
||
<paragraph id="8BFB368B542AFFECFF17FE87FF76FEAD" blockId="5.[151,1437,152,1018]" pageId="5" pageNumber="47">
|
||
<emphasis id="B930EA99542AFFECFF17FE87FEE2FED6" bold="true" box="[197,342,271,297]" pageId="5" pageNumber="47">Etymology:</emphasis>
|
||
Named for the collection site, the Orito River in the Putumayo drainage, Amazonía,
|
||
<collectingCountry id="F353761B542AFFECFA90FE98FF0BFEAD" name="Colombia" pageId="5" pageNumber="47">Colombia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35E6500542AFFECFF17FED7FE6CFC05" pageId="5" pageNumber="47" type="discussion">
|
||
<paragraph id="8BFB368B542AFFECFF17FED7FBAFFDBD" blockId="5.[151,1437,152,1018]" pageId="5" pageNumber="47">
|
||
<emphasis id="B930EA99542AFFECFF17FED7FEF5FE86" bold="true" box="[197,321,351,377]" pageId="5" pageNumber="47">Remarks:</emphasis>
|
||
The new species is very similar to
|
||
<taxonomicName id="4C444D08542AFFECFD3DFEE8FC3DFE86" box="[751,905,352,377]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="47" phylum="Chordata" rank="species" species="notidanos">
|
||
<emphasis id="B930EA99542AFFECFD3DFEE8FC3DFE86" box="[751,905,352,377]" italics="true" pageId="5" pageNumber="47">H. notidanos</emphasis>
|
||
</taxonomicName>
|
||
, however it differs from it in the number of lateral scales (35 vs.
|
||
<quantity id="4CBC9B6E542AFFECFE58FE00FE46FE5D" box="[394,498,392,418]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" metricValueMax="8.636" metricValueMin="8.128" pageId="5" pageNumber="47" unit="in" value="33.0" valueMax="34.0" valueMin="32.0">32–34 in</quantity>
|
||
<taxonomicName id="4C444D08542AFFECFE2AFE00FD3BFE5E" box="[504,655,392,417]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="47" phylum="Chordata" rank="species" species="notidanos">
|
||
<emphasis id="B930EA99542AFFECFE2AFE00FD3BFE5E" box="[504,655,392,417]" italics="true" pageId="5" pageNumber="47">H. notidanos</emphasis>
|
||
</taxonomicName>
|
||
), in the number of pored scales (12–14 vs. 9–11), in the number of scale rows between dorsal–fin origin and lateral line (7 vs. 6), in the number of predorsal median scales (12– 14 vs. 9–11), in the number of teeth in the dentary (19 vs. 7), in the number of teeth in the maxillary (5–6 vs. 2–4) in the dorsal–fin rays (iv, 26–27 vs. iii, 17–20), in body depth (37.84–5.59% LS vs. 36.84–37.79% LS) and in the snout–anal fin distance (56.71–65.05% LS vs. 62.52–80.39% LS).
|
||
</paragraph>
|
||
<paragraph id="8BFB368B542AFFECFF17FDD8FCC3FCF5" blockId="5.[151,1437,152,1018]" pageId="5" pageNumber="47">
|
||
Principal components analysis (fig. 3) indicates that
|
||
<taxonomicName id="4C444D08542AFFECFC95FDD8FB3DFD96" box="[839,1161,592,617]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="47" phylum="Chordata" rank="species" species="oritoensis" status="sp. nov.">
|
||
<emphasis id="B930EA99542AFFECFC95FDD8FB3DFD96" box="[839,1161,592,617]" italics="true" pageId="5" pageNumber="47">Hyphessobrycon oritoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20357E2542AFFECFB46FDD8FB62FD95" box="[1172,1238,592,618]" pageId="5" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
|
||
is distinguished from
|
||
<emphasis id="B930EA99542AFFECFF09FDF0FA8BFD6E" box="[219,1343,632,657]" italics="true" pageId="5" pageNumber="47">
|
||
<taxonomicName id="4C444D08542AFFECFF09FDF0FD8EFD6E" box="[219,570,632,657]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="47" phylum="Chordata" rank="species" species="poecilioides">Hyphessobrycon poecilioides</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C444D08542AFFECFD9AFDF0FCA8FD6E" box="[584,796,632,657]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="47" phylum="Chordata" rank="species" species="heterorhabdus">H. heterorhabdus</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C444D08542AFFECFCFEFDF0FC10FD6E" box="[812,932,632,657]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="47" phylum="Chordata" rank="species" species="vilmae">H. vilmae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C444D08542AFFECFC61FDF0FB31FD6E" box="[947,1157,632,657]" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="47" phylum="Chordata" rank="species" species="melanostichos">H. melanostichos</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C444D08542AFFECFB47FDF0FA8BFD6E" box="[1173,1343,632,657]" class="Actinopterygii" family="Characidae" genus="Hemigrammus" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="47" phylum="Chordata" rank="species" species="diancistrus">H. diancistrus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C444D08542AFFECFAADFDF0FEBFFD46" class="Actinopterygii" family="Characidae" genus="Hyphessobrycon" kingdom="Animalia" order="Characiformes" pageId="5" pageNumber="47" phylum="Chordata" rank="species" species="notidanos">
|
||
<emphasis id="B930EA99542AFFECFAADFDF0FEBFFD46" italics="true" pageId="5" pageNumber="47">H. notidanos</emphasis>
|
||
</taxonomicName>
|
||
in axis 1 by caudal peduncle depth in relationship to upper jaw length; and in axis 2, by body depth vs. dorsal fin–anal fin length. The first component explains 89.60% of the total variance, and combined with the second component they explain 92.74% of total variability.
|
||
</paragraph>
|
||
<paragraph id="8BFB368B542AFFECFF17FC9FFE6CFC05" blockId="5.[151,1437,152,1018]" pageId="5" pageNumber="47">
|
||
<emphasis id="B930EA99542AFFECFF17FC9FFE2DFCCE" bold="true" box="[197,409,791,817]" pageId="5" pageNumber="47">Ecological notes:</emphasis>
|
||
Data obtained from two stomachs of cleared and stained specimens indicates that this species is probably mostly insectivorous, feeding on allocthonous material, with unidentified insect parts being the most important element (56% by Number of prey items and 31% by Volume), followed by odonate larvae (8% N and 10% V), vegetable matter (16% N and 11% V) and unidentifiable material (20% N and 48% V). Their habitat had clear water over a substrate of sand and stone, with low dissolved oxygen (
|
||
<quantity id="4CBC9B6E542AFFECFAF7FC30FAC9FC2D" box="[1317,1405,952,978]" metricMagnitude="-6" metricUnit="kg" metricValue="5.2" pageId="5" pageNumber="47" unit="mg" value="5.2">5.2 mg</quantity>
|
||
/l), 67% saturation and
|
||
<collectingCountry id="F353761B542AFFECFE52FC68FE10FC05" box="[384,420,992,1018]" name="Philippines" pageId="5" pageNumber="47">pH</collectingCountry>
|
||
6.0.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |