treatments-xml/data/03/A7/87/03A7873CFFE34532FF72236BFF681545.xml

<document id="3A79D9F5272454471ED7D164246DA76F" ID-DOI="10.11646/phytotaxa.645.1.8" ID-ISSN="1179-3163" ID-Zenodo-Dep="13214900" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="jonas" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1722596870361" checkinUser="felipe" docAuthor="Nguyen, Cuong Huu, Le, Anh Tuan, Ha, Van Bac &amp; Lin, Che Wei" docDate="2024" docId="03A7873CFFE34532FF72236BFF681545" docLanguage="en" docName="phytotaxa.645.1.8.pdf" docOrigin="Phytotaxa 645 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.645.1.8" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Begonia dakrongensis C. H. Nguyen, T. A. Le &amp; C. W. Lin, sp. nov." docType="treatment" docVersion="1" lastPageNumber="86" masterDocId="FF9EFF44FFE24533FFFA225FFF8A121F" masterDocTitle="Begonia dakrongensis (Begoniaceae), a new species from Dakrong Nature Reserve, Quang Tri province in Central Vietnam" masterLastPageNumber="90" masterPageNumber="85" pageNumber="86" updateTime="1724788287754" updateUser="jonas" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="730F17F75D55F947EBA22977816C76CF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4D68A8A02BAC9B8100BDFE6E15DA160B">
<mods:title id="0617CE739930F284F4EFE174EC12110C">Begonia dakrongensis (Begoniaceae), a new species from Dakrong Nature Reserve, Quang Tri province in Central Vietnam</mods:title>
</mods:titleInfo>
<mods:name id="DFC3AD2ABD0164E09E4AE770E101F28C" type="personal">
<mods:role id="A277D068B437BBCE73F043EE1D1B1BDF">
<mods:roleTerm id="6C4DED87CB9278A38D519EFADFF661C9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C5512F3918791790AD8A096B90232CFD">Nguyen, Cuong Huu</mods:namePart>
<mods:nameIdentifier id="8683C10AFF984DA8B7F35C50EF90B3F0" type="ORCID">0000-0003-2281-741X</mods:nameIdentifier>
<mods:affiliation id="702B480DED94399FBB3EFB5D3F202103">Vietnam National University of Forestry, 21 A Street, Xuan Mai, Chuong My, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="DFF8953ED4A04FCC589D45960E551D37" type="email">nguyenhuucuong.tvr@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="F3942563D7E98357ACDF7A866E3B44F4" type="personal">
<mods:role id="88A2F172EC2BCF9909817DCF40E18182">
<mods:roleTerm id="F758A9D31AE4CE1BBC0AADD469151F4E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1929B5F583ED76C5DA4EF5D4B03DBE1F">Le, Anh Tuan</mods:namePart>
<mods:nameIdentifier id="292D390F2F7881178A5B5EB1526BE5B4" type="ORCID">0000-0002-5917-3243</mods:nameIdentifier>
<mods:affiliation id="503BEC1140B61E88852BA53C43181E9C">Mien Trung Institute for Scientific Research, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 321 Huynh Thuc Khang street, Thua Thien Hue, Vietnam</mods:affiliation>
<mods:nameIdentifier id="3AE8D40D1FAFEF65A36FC7C4671EEC25" type="email">tasa207@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="EBE0BBADBEFA24C71BFA103A32DDE9AD" type="personal">
<mods:role id="193CDD6A386DBB7DBBA9DB2D55A09ACF">
<mods:roleTerm id="C7DD5BAD99FD9D0AC6D93F6F5DB5331F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="83F0FB01E24C3CFD64B4C52B8AC6B773">Ha, Van Bac</mods:namePart>
<mods:nameIdentifier id="8A0D1202DAACFA566CEF8EAD566B5FF0" type="ORCID">0009-0003-0024-2608</mods:nameIdentifier>
<mods:affiliation id="70A7C9B5B2A025C230CEB03E94FF0F5B">Dakrong Natural Reserve, Ta Long, Dakrong, Quang Tri, Vietnam</mods:affiliation>
<mods:nameIdentifier id="65C5986CFF0C0ECB6074F7B473310966" type="email">havanbacklqt@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="E123586E05A3E3597DDC0FBBFEE7CEE8" type="personal">
<mods:role id="BF417AF60B577E1D6706BDDC5C4D2C4F">
<mods:roleTerm id="160C9F50B2BC79BF6135F5D2424B13B0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B809451F55BAFFE7B73A9FB6DBA6D143">Lin, Che Wei</mods:namePart>
<mods:nameIdentifier id="47436819ABF2D47D6D3D3A08B4868632" type="ORCID">0000-0002-8564-5793</mods:nameIdentifier>
<mods:affiliation id="ED6592BBFDB70EC3C476F3A6EE379B06">Herbarium of Taiwan Forestry Research Institute, Taiwan Forestry Research Institute, No. 53, Nan-Hai Road, Taipei 100, Taiwan</mods:affiliation>
<mods:nameIdentifier id="4AB15CC69980CB0C49C1752C8D47081E" type="email">varalba@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="F1CE598A402CE146F20E54665869E7F8">text</mods:typeOfResource>
<mods:relatedItem id="D2983156C28C559A580F17B268BBC434" type="host">
<mods:titleInfo id="4A66007D18FAC9C786A4C2883E4C739E">
<mods:title id="2C761DD6D403AF17DBE21F90E6172CAD">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="F97719B2E35759F878CC5FE780B6A535">
<mods:date id="A95B1D91569A61A261FF3289438EB545">2024</mods:date>
<mods:detail id="DEECD428E9BDE75DEDF52A07E044F868" type="pubDate">
<mods:number id="18CDA87DE4F37A405DA621EA18A255D0">2024-04-18</mods:number>
</mods:detail>
<mods:detail id="77DB7A2216814D8A8499587727482D0C" type="volume">
<mods:number id="FF2DDBDA72E260E935BF1CBF8FDD2265">645</mods:number>
</mods:detail>
<mods:detail id="C609913A360E0B44C2A54417DB8B2AF8" type="issue">
<mods:number id="70A2470136DB11F03ACA1579C0F38623">1</mods:number>
</mods:detail>
<mods:extent id="687471387BC217644136FC5C3902C6EF" unit="page">
<mods:start id="B985D16D0683B5B73E9A200B0C3BED7C">85</mods:start>
<mods:end id="2DE966863A710AA16E8FEF877661D59D">90</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="3D546EF73763D3B9C6CCCD79989DED25">
<mods:url id="AA92B56F018A7B20FF544ACA948F39A8">http://dx.doi.org/10.11646/phytotaxa.645.1.8</mods:url>
</mods:location>
<mods:classification id="3527973E54BD9DE91306C9D954D4328F">journal article</mods:classification>
<mods:identifier id="998C13008FE1A6DC680BF76B3A02D9DA" type="DOI">10.11646/phytotaxa.645.1.8</mods:identifier>
<mods:identifier id="36A8B5523B8F11686FFD41CE1D1ECAF3" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="138A68A84D1FC9E7F7E259CF5B01AC54" type="Zenodo-Dep">13214900</mods:identifier>
</mods:mods>
<treatment id="03A7873CFFE34532FF72236BFF681545" LSID="urn:lsid:plazi:treatment:03A7873CFFE34532FF72236BFF681545" httpUri="http://treatment.plazi.org/id/03A7873CFFE34532FF72236BFF681545" lastPageNumber="86" pageId="1" pageNumber="86">
<subSubSection id="C31465A1FFE34532FF72236BFC721351" box="[136,1016,308,335]" pageId="1" pageNumber="86" type="nomenclature">
<paragraph id="8BB1362AFFE34532FF72236BFC721351" blockId="1.[136,1016,308,335]" box="[136,1016,308,335]" pageId="1" pageNumber="86">
<heading id="D0F98146FFE34532FF72236BFC721351" box="[136,1016,308,335]" fontSize="11" level="2" pageId="1" pageNumber="86" reason="3">
<taxonomicName id="4C0E4DA9FFE34532FF72236BFC8B1351" ID-CoL="CFGSN" authority="C.H.Nguyen, T.A.Le &amp; C.W.Lin" authorityName="C. H. Nguyen, T. A. Le &amp; C. W. Lin" box="[136,769,308,335]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="dakrongensis" status="sp. nov.">
<emphasis id="B97AEA38FFE34532FF72236BFE001350" bold="true" box="[136,394,308,335]" italics="true" pageId="1" pageNumber="86">Begonia dakrongensis</emphasis>
C.H.Nguyen, T.A.Le &amp; C.W.Lin
</taxonomicName>
, 
<emphasis id="B97AEA38FFE34532FCF6236AFCEB1351" box="[780,865,309,334]" italics="true" pageId="1" pageNumber="86">
<taxonomicNameLabel id="A2495743FFE34532FCF6236AFCEB1351" box="[780,865,309,334]" pageId="1" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13352AAFFFE34532FC95236BFC361351" box="[879,956,308,334]" captionStart="FIGURE 1" captionStartId="2.[136,234,1990,2012]" captionTargetBox="[200,1416,200,1948]" captionTargetId="figure-19@2.[163,1423,187,1968]" captionTargetPageId="2" captionText="FIGURE 1. Begonia dakrongensis C.H.Nguyen, T.A.Le &amp; C.W.Lin. A. Habit; B, B'. Portion of leaf, upper and lower surfaces; C. Stipule; D, D', D'', D'''. Bracts; E, E'. Staminate flower, face and side views; F, F'. Stamens, ventral and dorsal views; G, G'. Immature pistillate flower, face and side views; H, H'. Style and stigmatic band, ventral and dorsal views; I. Cross section of ovary; J. Capsule." figureDoi="http://doi.org/10.5281/zenodo.13214902" httpUri="https://zenodo.org/record/13214902/files/figure.png" pageId="1" pageNumber="86">Figs. 1</figureCitation>
&amp; 
<figureCitation id="13352AAFFFE34532FC1A236BFC651351" box="[992,1007,308,334]" captionStart="FIGURE 2" captionStartId="3.[136,234,1935,1957]" captionTargetBox="[151,1435,190,1911]" captionTargetId="figure-19@3.[151,1435,190,1911]" captionTargetPageId="3" captionText="FIGURE 2. Begonia dakrongensis C.H.Nguyen, T.A.Le &amp; C.W.Lin.A. Habit and habitat; B. Habitats; C. Inflorescence, showing bracts, immature staminate flowers and capsule; D. Portion of leaf, upper surface, E. same, lower surface; F. Staminate flowers; G. Immature staminate flower; H. Staminate flower, face view; I. Immature pistillate flower; J. Cross section of ovary; K. Capsule." figureDoi="http://doi.org/10.5281/zenodo.13214904" httpUri="https://zenodo.org/record/13214904/files/figure.png" pageId="1" pageNumber="86">2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31465A1FFE34532FF7223DFFBA517ED" pageId="1" pageNumber="86" type="materials_examined">
<paragraph id="8BB1362AFFE34532FF7223DFFDC1101D" blockId="1.[136,1452,384,514]" pageId="1" pageNumber="86">
<emphasis id="B97AEA38FFE34532FF7223DFFF50138A" bold="true" box="[136,218,384,405]" pageId="1" pageNumber="86">Type:—</emphasis>
<materialsCitation id="3B663C77FFE34532FF2023DFFEBA101D" collectingDate="2023-05-26" collectionCode="VNMN" collectorName="Tuan Anh Le &amp; Ha Van Bac" country="Vietnam" county="Dakrong district" elevation="800" latitude="16.463028" location="Dakrong Nature Reserve" longLatPrecision="1" longitude="107.09234" municipality="Huc Nghi" pageId="1" pageNumber="86" specimenCode="B0000608" specimenCount="1" stateProvince="Quang Tri province" typeStatus="holotype">
<collectingCountry id="F31976BAFFE34532FF2023DFFEC6138A" box="[218,332,384,405]" name="Vietnam" pageId="1" pageNumber="86">VIETNAM</collectingCountry>
. 
<collectingRegion id="49CAF8C8FFE34532FEAC23DFFD9C1389" box="[342,534,384,406]" country="Vietnam" name="Quang Tri" pageId="1" pageNumber="86">Quang Tri province</collectingRegion>
, 
<collectingCounty id="62D04EA6FFE34532FDDB23DFFD4B1389" box="[545,705,384,406]" pageId="1" pageNumber="86">Dakrong district</collectingCounty>
, 
<collectingMunicipality id="6BD5AC50FFE34532FD3623DFFCA11389" box="[716,811,384,406]" pageId="1" pageNumber="86">Huc Nghi</collectingMunicipality>
commune, 
<location id="8ED160F1FFE34532FC6423DFFB1B138A" LSID="urn:lsid:plazi:treatment:03A7873CFFE34532FF72236BFF681545:8ED160F1FFE34532FC6423DFFB1B138A" box="[926,1169,384,406]" country="Vietnam" county="Dakrong district" latitude="16.463028" longLatPrecision="1" longitude="107.09234" municipality="Huc Nghi" name="Dakrong Nature Reserve" pageId="1" pageNumber="86" stateProvince="Quang Tri province">Dakrong Nature Reserve</location>
, around point 
<geoCoordinate id="EE3A50EDFFE34532FAE423DFFA2D1389" box="[1310,1447,384,406]" degrees="16" direction="north" minutes="27" orientation="latitude" pageId="1" pageNumber="86" precision="1" seconds="46.9" value="16.463028">16°27’46.9”N</geoCoordinate>
, 
<geoCoordinate id="EE3A50EDFFE34532FF4723FBFEDA13A5" box="[189,336,420,442]" degrees="107" direction="east" minutes="05" orientation="longitude" pageId="1" pageNumber="86" precision="1" seconds="32.4" value="107.09234">107°05’32.4”E</geoCoordinate>
, common in the moist shady areas under evergreen primary evergreen forest on the rocky crevices in association with of waterfall Do Quyen at elevations of about 
<elevation id="0023D119FFE34532FD8B2397FD6813C1" box="[625,738,456,478]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="86" unit="m" value="800.0">
<quantity id="4CF69BCFFFE34532FD8B2397FD2413C2" box="[625,686,456,477]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="86" unit="m" value="800.0">800 m</quantity>
a.s.l.
</elevation>
, 
<date id="FFB010EAFFE34532FD172397FCE513C2" box="[749,879,456,478]" pageId="1" pageNumber="86" value="2023-05-26">
<collectingDate id="EFF4E902FFE34532FD172397FCE513C2" box="[749,879,456,478]" pageId="1" pageNumber="86" value="2023-05-26">26 May 2023</collectingDate>
</date>
, 
<emphasis id="B97AEA38FFE34532FC832397FB7013C2" box="[889,1274,456,477]" italics="true" pageId="1" pageNumber="86">
<collectorName id="26FB53FCFFE34532FC832397FC7C13C2" box="[889,1014,456,477]" pageId="1" pageNumber="86">Tuan Anh Le</collectorName>
, 
<collectorName id="26FB53FCFFE34532FBFA2397FBFC13C2" box="[1024,1142,456,477]" pageId="1" pageNumber="86">Ha Van Bac</collectorName>
, LTA.DKR.09
</emphasis>
(
<typeStatus id="54B58888FFE34532FAFD2397FAD713C1" box="[1287,1373,456,478]" pageId="1" pageNumber="86" type="holotype">holotype</typeStatus>
<collectionCode id="ED1FAEEFFFE34532FA982397FA2113C2" box="[1378,1451,456,477]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="1" pageNumber="86">VNMN</collectionCode>
<specimenCode id="DBA89E51FFE34532FF4723B3FEA3101D" box="[189,297,492,514]" collectionCode="B" country="Germany" lsid="urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="1" pageNumber="86" type="Herbarium">B0000608</specimenCode>
!;
</materialsCitation>
<materialsCitation id="3B663C77FFE34532FECC23B3FDC1101D" box="[310,587,492,514]" collectingDate="2023-05-26" collectionCode="VNF" collectorName="Tuan Anh Le &amp; Ha Van Bac" country="Vietnam" county="Dakrong district" elevation="800" latitude="16.463028" location="Dakrong Nature Reserve" longLatPrecision="1" longitude="107.09234" municipality="Huc Nghi" pageId="1" pageNumber="86" specimenCode="VNF 20230526010" specimenCount="1" stateProvince="Quang Tri province" typeStatus="isotype">
<typeStatus id="54B58888FFE34532FECC23B3FEF1101D" box="[310,379,492,514]" pageId="1" pageNumber="86" type="isotype">isotype</typeStatus>
<specimenCode id="DBA89E51FFE34532FE7B23B3FDC8101D" box="[385,578,492,514]" collectionCode="VNF" country="Viet Nam" httpUri="http://grbio.org/cool/merb-e6y6" name="Vietnam Forestry Herbarium" pageId="1" pageNumber="86" type="Herbarium">VNF 20230526010</specimenCode>
!)
</materialsCitation>
</paragraph>
<paragraph id="8BB1362AFFE34532FF72206EFBA517ED" blockId="1.[136,1452,560,1882]" pageId="1" pageNumber="86">
Rosette, rhizomatous perennial herbs, monoecious. 
<emphasis id="B97AEA38FFE34532FD2C206FFCC11055" bold="true" box="[726,843,560,586]" pageId="1" pageNumber="86">Rhizomes</emphasis>
dusky magenta to reddish-brown or brownish-green, terete, to 
<quantity id="4CF69BCFFFE34532FF15200BFEAF1070" box="[239,293,596,623]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="86" unit="cm" value="6.0">6 cm</quantity>
or longer, 
<quantity id="4CF69BCFFFE34532FE67200BFE731070" box="[413,505,596,623]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="1" pageNumber="86" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 mm</quantity>
across, creeping, sparsely villous-hirsute, internodes 
<quantity id="4CF69BCFFFE34532FBB1200BFB361070" box="[1099,1212,596,623]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="9.0" metricValueMin="1.5" pageId="1" pageNumber="86" unit="mm" value="5.25" valueMax="9.0" valueMin="1.5">1.5–9 mm</quantity>
long. 
<emphasis id="B97AEA38FFE34532FAFA200BFAD51071" bold="true" box="[1280,1375,596,622]" pageId="1" pageNumber="86">Stipules</emphasis>
ovate-triangular, slightly oblique, 5–8 × 
<quantity id="4CF69BCFFFE34532FDEB2027FD16108C" box="[529,668,632,659]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="1" pageNumber="86" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 mm</quantity>
, glabrous or very sparsely hirsute, apex attenuate and aristate, arista 
<quantity id="4CF69BCFFFE34532FF7220C3FF7310A8" box="[136,249,668,695]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="3.0" metricValueMin="0.8" pageId="1" pageNumber="86" unit="mm" value="1.9" valueMax="3.0" valueMin="0.8">0.8–3 mm</quantity>
long, margin entire or very sparsely denticulate. 
<emphasis id="B97AEA38FFE34532FCE020C3FCE610A9" bold="true" box="[794,876,668,694]" pageId="1" pageNumber="86">Leaves</emphasis>
alternate, nearly pressed to substrate; petiole dusky red, 
<quantity id="4CF69BCFFFE34532FF43209FFEBC10C4" box="[185,310,704,731]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="3.5" metricValueMin="1.3" pageId="1" pageNumber="86" unit="cm" value="2.4" valueMax="3.5" valueMin="1.3">1.3–3.5 cm</quantity>
long, 
<quantity id="4CF69BCFFFE34532FE86209FFE6710C4" box="[380,493,704,731]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.0" metricValueMin="2.5" pageId="1" pageNumber="86" unit="mm" value="3.25" valueMax="4.0" valueMin="2.5">2.5–4 mm</quantity>
across, grooved above, sparsely to densely villous-hirsute, leaf blade asymmetric, not oblique or nearly so, obtriangular-spathulate to oblong-obovate or widely obovate, 4.7–17 × 
<quantity id="4CF69BCFFFE34532FB6320BBFB7710E0" box="[1177,1277,740,767]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="12.0" metricValueMin="3.0" pageId="1" pageNumber="86" unit="cm" value="7.5" valueMax="12.0" valueMin="3.0">3–12 cm</quantity>
, base obliquely cordate, lobes 
<quantity id="4CF69BCFFFE34532FED62157FE1D113C" box="[300,407,776,803]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="12.0" metricValueMin="6.0" pageId="1" pageNumber="86" unit="mm" value="9.0" valueMax="12.0" valueMin="6.0">6–12 mm</quantity>
long, broad side 
<quantity id="4CF69BCFFFE34532FDA02157FD49113C" box="[602,707,776,803]" metricMagnitude="-2" metricUnit="m" metricValue="4.65" metricValueMax="7.3" metricValueMin="2.0" pageId="1" pageNumber="86" unit="cm" value="4.65" valueMax="7.3" valueMin="2.0">2–7.3 cm</quantity>
wide, margin denticulate on the upper two-thirds and subentire to repand on the lower one-third, slightly revolute and undulate with rows of sparsely white villous and red scabrous, sometimes with coarsely dentate and irregularly angular, apex acuminate to acute; leaf subcoriaceous, succulent, adaxially emerald green, glabrous, waxy, abaxially pale green, veins pink to red, sparsely pinkish-white villous-hirsute and red scabrous; venation palmate-pinnate, midrib distinct, 2 or 3 secondary veins on broad side and 0 or 1 on narrow side, lateral through connection between with other basal veins, minor veins inconspicuous. 
<emphasis id="B97AEA38FFE34532FB5421E3FAC611C9" bold="true" box="[1198,1356,956,982]" pageId="1" pageNumber="86">Inflorescence</emphasis>
axillary and terminal, unisexual or bisexual, cymose-paniculate or racemose, 
<quantity id="4CF69BCFFFE34532FC6E21BFFC7711E4" box="[916,1021,992,1019]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="1" pageNumber="86" unit="cm" value="2.25" valueMax="3.0" valueMin="1.5">1.5–3 cm</quantity>
long, rose pink to red, erect, glabrous or subglabrous; Bracts creamy pink to creamy white or greenish-white, hyaline, persistent, ovate-triangular to widely triangular or narrowly triangular, the first pair up to 7 × 
<quantity id="4CF69BCFFFE34532FD122677FCB4165C" box="[744,830,1064,1091]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="86" unit="mm" value="4.5">4.5 mm</quantity>
, decreasing in size towards the more distal inflorescence nodes, attenuate and aristate at apex, arista 
<quantity id="4CF69BCFFFE34532FD982613FD621678" box="[610,744,1100,1127]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.5" metricValueMin="0.5" pageId="1" pageNumber="86" unit="mm" value="1.5" valueMax="2.5" valueMin="0.5">0.5–2.5 mm</quantity>
long, margin denticulate; Pistillate flower produced from a very short peduncle at the basal node of the inflorescence, usually solitary; staminate flowers produced in distal inflorescence part, in several 1–3-flowered compressed monochasial partial inflorescences borne on lateral branches (
<quantity id="4CF69BCFFFE34532FAD726CBFA1516B1" box="[1325,1439,1172,1198]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="12.0" metricValueMin="2.0" pageId="1" pageNumber="86" unit="mm" value="7.0" valueMax="12.0" valueMin="2.0">2–12 mm</quantity>
). 
<emphasis id="B97AEA38FFE34532FF7226E7FEEA16CD" bold="true" box="[136,352,1208,1234]" pageId="1" pageNumber="86">Staminate flower:</emphasis>
pedicel 
<quantity id="4CF69BCFFFE34532FE3926E7FDB416CC" box="[451,574,1208,1235]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="1" pageNumber="86" unit="mm" value="12.0" valueMax="14.0" valueMin="10.0">10–14 mm</quantity>
long, tepals 4, outer 2 white to pinkish-white, sometimes with pale greyishpink veins, orbicular to obovate, 7–12 × 
<quantity id="4CF69BCFFFE34532FDAE2683FD5C16E8" box="[596,726,1244,1271]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="10.0" metricValueMin="8.5" pageId="1" pageNumber="86" unit="mm" value="9.25" valueMax="10.0" valueMin="8.5">8.5–10 mm</quantity>
, margin entire, abaxially red scabrous; Inner 2 white or pinkish-white, oblanceolate, apex round, base cuneate, 7.5–9 × 
<quantity id="4CF69BCFFFE34532FCF9275FFCEE1704" box="[771,868,1280,1307]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="86" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
, glabrous; Androecium actinomorphic, stamens 
<emphasis id="B97AEA38FFE34532FA70275EFA2E1705" box="[1418,1444,1281,1306]" italics="true" pageId="1" pageNumber="86">ca</emphasis>
. 25, yellow, filaments free, 
<emphasis id="B97AEA38FFE34532FE4B277AFE591721" box="[433,467,1317,1342]" italics="true" pageId="1" pageNumber="86">ca.</emphasis>
<quantity id="4CF69BCFFFE34532FE23277BFDA41720" box="[473,558,1316,1343]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="86" unit="mm" value="0.4">0.4 mm</quantity>
long, anthers widely obovate, 
<emphasis id="B97AEA38FFE34532FC7E277AFC141721" box="[900,926,1317,1342]" italics="true" pageId="1" pageNumber="86">ca</emphasis>
. 
<quantity id="4CF69BCFFFE34532FC51277BFB8B1720" box="[939,1025,1316,1343]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="86" unit="mm" value="0.7">0.7 mm</quantity>
long, 2-locular, apex retuse. 
<emphasis id="B97AEA38FFE34532FAB9277BFEE6177D" bold="true" pageId="1" pageNumber="86">Pistillate flower (immature):</emphasis>
subsessile, tepals 5, white or pinkish-white, outer 4 tepals widely ovate to oblong, margin entire, apex rounded to obtuse, abaxially red scabrous; inner 1 obovate to oblanceolate, glabrous; Ovary trigonous-globose, densely red scabrous, 3-locular, 3-winged, wings subequal, trapeziform, truncate to broadly acute proximally, pointed distally, placentation axle, bilamellate; styles 3, Y-shaped, divided to base, stigmatic band spiralled. 
<emphasis id="B97AEA38FFE34532FB0827EBFAD817D1" bold="true" box="[1266,1362,1460,1486]" pageId="1" pageNumber="86">Capsule</emphasis>
pedicel 
<emphasis id="B97AEA38FFE34532FF722786FF2817ED" box="[136,162,1497,1522]" italics="true" pageId="1" pageNumber="86">ca</emphasis>
. 
<quantity id="4CF69BCFFFE34532FF4A2787FF7B17EC" box="[176,241,1496,1523]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="86" unit="mm" value="1.0">1 mm</quantity>
long, body trigonous-globose, 10–11 × 
<quantity id="4CF69BCFFFE34532FD482787FCA117EC" box="[690,811,1496,1523]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="1" pageNumber="86" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">12–14 mm</quantity>
long (wings included).
</paragraph>
</subSubSection>
<subSubSection id="C31465A1FFE34532FF4727A3FA161441" pageId="1" pageNumber="86" type="distribution">
<paragraph id="8BB1362AFFE34532FF4727A3FA161441" blockId="1.[136,1452,560,1882]" pageId="1" pageNumber="86">
<emphasis id="B97AEA38FFE34532FF4727A3FE721409" bold="true" box="[189,504,1532,1558]" pageId="1" pageNumber="86">Distribution and ecology:</emphasis>
—The new species is only known from Dakrong Nature Reserve, in 
<collectingRegion id="49CAF8C8FFE34532FACE27A3FF671425" country="Vietnam" name="Quang Tri" pageId="1" pageNumber="86">Quang Tri province</collectingRegion>
, Central 
<collectingCountry id="F31976BAFFE34532FEA8247FFE3F1425" box="[338,437,1568,1594]" name="Vietnam" pageId="1" pageNumber="86">Vietnam</collectingCountry>
. 
<taxonomicName id="4C0E4DA9FFE34532FE45247EFD3D1425" box="[447,695,1568,1594]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="dakrongensis">
<emphasis id="B97AEA38FFE34532FE45247EFD3D1425" box="[447,695,1568,1594]" italics="true" pageId="1" pageNumber="86">Begonia dakrongensis</emphasis>
</taxonomicName>
usually grows in moist shady places near waterfalls, along streams and occasionally on the rocky crevices covered by evergreen broad-leaved forests at elevations of about 
<quantity id="4CF69BCFFFE34532FAE0241BFAEB1440" box="[1306,1377,1604,1631]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="86" unit="m" value="800.0">800 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="C31465A1FFE34532FF472437FF681545" pageId="1" pageNumber="86" type="etymology">
<paragraph id="8BB1362AFFE34532FF472437FAEE149D" blockId="1.[136,1452,560,1882]" box="[189,1380,1640,1667]" pageId="1" pageNumber="86">
<emphasis id="B97AEA38FFE34532FF472437FEC2149D" bold="true" box="[189,328,1640,1666]" pageId="1" pageNumber="86">Etymology:</emphasis>
—The specific epithet refers to the 
<typeStatus id="54B58888FFE34532FD2E2436FC8E149C" box="[724,772,1641,1667]" pageId="1" pageNumber="86">type</typeStatus>
locality (Dakrong Nature Reserve, Central 
<collectingCountry id="F31976BAFFE34532FB172437FADE149D" box="[1261,1364,1640,1666]" name="Vietnam" pageId="1" pageNumber="86">Vietnam</collectingCountry>
.).
</paragraph>
<paragraph id="8BB1362AFFE34532FF4724D3FB7814D5" blockId="1.[136,1452,560,1882]" pageId="1" pageNumber="86">
<emphasis id="B97AEA38FFE34532FF4724D3FE3914B9" bold="true" box="[189,435,1676,1702]" pageId="1" pageNumber="86">Conservation status:</emphasis>
—The new species is known from a single collection, and we do not have information about its abundance at 
<typeStatus id="54B58888FFE34532FEB924EEFEF914D4" box="[323,371,1713,1739]" pageId="1" pageNumber="86">type</typeStatus>
locality its conservation status is assessed as Data Deficient (
<collectionCode id="ED1FAEEFFFE34532FBDF24EFFBD814D5" box="[1061,1106,1712,1738]" country="India" lsid="urn:lsid:biocol.org:col:13033" name="Forest Research Institute, Indian Council of Forestry Research and Education" pageId="1" pageNumber="86" type="Herbarium">DD</collectionCode>
; 
<specimenCode id="DBA89E51FFE34532FBA724EFFB6C14D5" box="[1117,1254,1712,1738]" collectionCode="IUCN" pageId="1" pageNumber="86">IUCN 2022</specimenCode>
).
</paragraph>
<paragraph id="8BB1362AFFE34532FF47248BFF681545" blockId="1.[136,1452,560,1882]" pageId="1" pageNumber="86">
<emphasis id="B97AEA38FFE34532FF47248BFE8314F1" bold="true" box="[189,265,1748,1774]" pageId="1" pageNumber="86">Notes:</emphasis>
—Among the rhizomatous begonias in 
<collectingCountry id="F31976BAFFE34532FD2B248BFCB914F1" box="[721,819,1748,1774]" name="Vietnam" pageId="1" pageNumber="86">Vietnam</collectingCountry>
, 
<taxonomicName id="4C0E4DA9FFE34532FCBA248AFBB614F1" box="[832,1084,1748,1774]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="dakrongensis">
<emphasis id="B97AEA38FFE34532FCBA248AFBB614F1" box="[832,1084,1748,1774]" italics="true" pageId="1" pageNumber="86">Begonia dakrongensis</emphasis>
</taxonomicName>
closely resembles 
<taxonomicName id="4C0E4DA9FFE34532FAE3248AFA2614F1" box="[1305,1452,1748,1774]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="abbrevata">
<emphasis id="B97AEA38FFE34532FAE3248AFA2614F1" box="[1305,1452,1748,1774]" italics="true" pageId="1" pageNumber="86">B. abbrevata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C0E4DA9FFE34532FF4224A6FDEB150D" authority="Gagnepain (1919: 198)" authorityName="Gagnepain" authorityPageNumber="198" authorityYear="1919" box="[184,609,1784,1810]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="86" phylum="Tracheophyta" rank="species" species="eberhardtii">
<emphasis id="B97AEA38FFE34532FF4224A6FEDE150D" box="[184,340,1784,1810]" italics="true" pageId="1" pageNumber="86">B. eberhardtii</emphasis>
<bibRefCitation id="EF9F4BDBFFE34532FEA624A7FDEB150D" author="Gagnepain, M. F." box="[348,609,1784,1810]" pageId="1" pageNumber="86" pagination="194 - 201" refId="ref2390" refString="Gagnepain, M. F. (1919) Nouveaux Begonia d'Asie. Quelques synonymes. Bulletin du Museum National d'Histoire Naturelle 25: 194 - 201." type="journal article" year="1919">Gagnepain (1919: 198)</bibRefCitation>
</taxonomicName>
in their axillary and terminal inflorescence with densely bracts and flower structure, but these three species differ in many morphological characteristics. A detailed comparison is provided in 
<tableCitation id="C68C0391FFE34532FF72251FFF561545" box="[136,220,1856,1882]" captionStart="TABLE 1" captionStartId="4.[136,224,159,183]" captionTargetBox="[144,1439,240,1156]" captionTargetPageId="4" captionText="TABLE 1. Comparison of Begonia dakrongensis C.H.Nguyen, T.A.Le &amp; C.W.Lin, B. abbrevata C.I Peng and B. eberhardtii Gagnepain." pageId="1" pageNumber="86">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>