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<document id="C81A4E0344A5D0DF32BCC76137C2E141" ID-DOI="10.11646/phytotaxa.626.4.7" ID-ISSN="1179-3163" ID-Zenodo-Dep="13879337" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1727835996397" checkinUser="felipe" docAuthor="Al-Shehbaz, Ihsan A." docDate="2023" docId="BC0FC82EB35E277FFF73FB301291F9A8" docLanguage="en" docName="phytotaxa.626.4.7.pdf" docOrigin="Phytotaxa 626 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.626.4.7" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Lepidium weigendii Al-Shehbaz 2023, sp. nov." docType="treatment" docVersion="1" lastPageNumber="299" masterDocId="4036B056B35E277DFFFBFF89107EFFB5" masterDocTitle="Lepidium weigendii (Brassicaceae), a new species from southern Peru" masterLastPageNumber="300" masterPageNumber="297" pageNumber="297" updateTime="1729864979648" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="65FB186340E4C8ED6286D98A276D47CE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="633C49A199BE1318458EFB367C2CC970">
<mods:title id="9C984FC702941C3D4450F294DB95D149">Lepidium weigendii (Brassicaceae), a new species from southern Peru</mods:title>
</mods:titleInfo>
<mods:name id="B6EA7324442F489E0CD2BBDD831DAA27" type="personal">
<mods:role id="12816E3F816F31B9D67E122E83452E8B">
<mods:roleTerm id="47BF8AE24542E980316BD8926A877BDE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0717E405791DAAA173D4DF45F8E199A8">Al-Shehbaz, Ihsan A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="91EBC6D6507EE1CEA7D93F6AF6168DD6">text</mods:typeOfResource>
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<mods:title id="C8E2BAECEFD0DD4EB45B8FA263D11DE8">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="6A6B4BEF40FEA2CF4A1F6BB8162CA90D">
<mods:date id="A030102E905BF547F473210D4A2FD4D1">2023</mods:date>
<mods:detail id="B649A64C15F8E912F7685260E4DCC992" type="pubDate">
<mods:number id="200F7785F6EE99B6300180C6C1909891">2023-11-28</mods:number>
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<mods:number id="5E5B2E5EC1E0DF6D76AB65913DA0492D">626</mods:number>
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<mods:url id="747EAB8EA4DD6947712015F18E1E7CAC">http://dx.doi.org/10.11646/phytotaxa.626.4.7</mods:url>
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<mods:classification id="74BF92C687903957036295CE0548CF62">journal article</mods:classification>
<mods:identifier id="1A1516DE4033F8F3F284F919D04A4841" type="DOI">10.11646/phytotaxa.626.4.7</mods:identifier>
<mods:identifier id="FEDFAC96BEE2506D24A61AEFE1A70BB2" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="0168BA0A5C79F510BEA5258F6A80B9E3" type="Zenodo-Dep">13879337</mods:identifier>
</mods:mods>
<treatment id="BC0FC82EB35E277FFF73FB301291F9A8" LSID="urn:lsid:plazi:treatment:BC0FC82EB35E277FFF73FB301291F9A8" httpUri="http://treatment.plazi.org/id/BC0FC82EB35E277FFF73FB301291F9A8" lastPageId="2" lastPageNumber="299" pageId="0" pageNumber="297">
<subSubSection id="7CBC2AB3B35E277DFF73FB30123AFB67" box="[136,580,1209,1234]" pageId="0" pageNumber="297" type="nomenclature">
<paragraph id="34197938B35E277DFF73FB30123AFB67" blockId="0.[136,1452,1209,1378]" box="[136,580,1209,1234]" pageId="0" pageNumber="297">
<taxonomicName id="F3A602BBB35E277DFF73FB301197FB67" ID-CoL="C7K54" authority="Al-Shehbaz" authorityName="Al-Shehbaz" authorityYear="2023" box="[136,489,1209,1234]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="0" pageNumber="297" phylum="Tracheophyta" rank="species" species="weigendii" status="sp. nov.">
<emphasis id="06D2A52AB35E277DFF73FB30111FFB64" bold="true" box="[136,353,1209,1233]" italics="true" pageId="0" pageNumber="297">Lepidium weigendii</emphasis>
Al-Shehbaz
</taxonomicName>
,
<emphasis id="06D2A52AB35E277DFE08FB32123AFB67" box="[499,580,1211,1234]" italics="true" pageId="0" pageNumber="297">
<taxonomicNameLabel id="1DE11851B35E277DFE08FB32123AFB67" box="[499,580,1211,1234]" pageId="0" pageNumber="297" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="7CBC2AB3B35E277DFF73FB6915F6FAAF" pageId="0" pageNumber="297" type="diagnosis">
<paragraph id="34197938B35E277DFF73FB6915F6FAAF" blockId="0.[136,1452,1209,1378]" pageId="0" pageNumber="297">
Diagnosis:—
<taxonomicName id="F3A602BBB35E277DFEFCFB6911BBFB40" authorityName="Al-Shehbaz" authorityYear="2023" box="[263,453,1248,1269]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="0" pageNumber="297" phylum="Tracheophyta" rank="species" species="weigendii">
<emphasis id="06D2A52AB35E277DFEFCFB6911BBFB40" box="[263,453,1248,1269]" italics="true" pageId="0" pageNumber="297">Lepidium weigendii</emphasis>
</taxonomicName>
is easily distinguished from the related
<taxonomicName id="F3A602BBB35E277DFCBCFB6913CEFB40" box="[839,944,1248,1269]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="0" pageNumber="297" phylum="Tracheophyta" rank="species" species="crassius">
<emphasis id="06D2A52AB35E277DFCBCFB6913CEFB40" box="[839,944,1248,1269]" italics="true" pageId="0" pageNumber="297">L. crassius</emphasis>
</taxonomicName>
by having densely hirsute and notched (vs. glabrous and notchless) fruits, persistent (vs. caducous) nectar glands, and petals differentiated (vs. undifferentiated) into blade and claw.
</paragraph>
</subSubSection>
<subSubSection id="7CBC2AB3B35E277DFF73FAA114FFFAD4" pageId="0" pageNumber="297" type="materials_examined">
<paragraph id="34197938B35E277DFF73FAA114FFFAD4" blockId="0.[136,1452,1209,1378]" pageId="0" pageNumber="297">
<emphasis id="06D2A52AB35E277DFF73FAA110C4FA88" bold="true" box="[136,186,1320,1341]" pageId="0" pageNumber="297">
<typeStatus id="EB1DC79AB35E277DFF73FAA110C4FA88" box="[136,186,1320,1341]" pageId="0" pageNumber="297">Type</typeStatus>
</emphasis>
:—
<materialsCitation id="84CE7365B35E277DFF2CFAA11347FAD4" collectingDate="1997-10-04" collectionCode="MSB" collectorName="M. Weigend &amp; H. Forther" country="Peru" elevation="655" location="Lomas" municipality="Caravei" pageId="0" pageNumber="297" specimenCount="1" stateProvince="Arequipa" typeStatus="holotype">
<collectingCountry id="4CB139A8B35E277DFF2CFAA11167FA88" box="[215,281,1320,1341]" name="Peru" pageId="0" pageNumber="297">PERU</collectingCountry>
. Dept.
<collectingRegion id="F662B7DAB35E277DFEA0FAA111C6FA8B" box="[347,440,1320,1342]" country="Peru" name="Arequipa" pageId="0" pageNumber="297">Arequipa</collectingRegion>
: Prov.
<collectingMunicipality id="D47DE342B35E277DFE02FAA11238FA8B" box="[505,582,1320,1342]" pageId="0" pageNumber="297">Caravei</collectingMunicipality>
,
<location id="31792FE3B35E277DFDB4FAA112EEFA88" LSID="urn:lsid:plazi:treatment:BC0FC82EB35E277FFF73FB301291F9A8:31792FE3B35E277DFDB4FAA112EEFA88" box="[591,656,1320,1341]" country="Peru" municipality="Caravei" name="Lomas" pageId="0" pageNumber="297" stateProvince="Arequipa">Lomas</location>
ca.
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<quantity id="F35ED4DDB35E277DFD4CFAA11296FA8B" box="[695,744,1320,1342]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="297" unit="km" value="3.0">3 km</quantity>
E of km 655
</locationDeviation>
of
<location id="31792FE3B35E277DFC7FFAA1144DFA88" LSID="urn:lsid:plazi:treatment:BC0FC82EB35E277FFF73FB301291F9A8:31792FE3B35E277DFC7FFAA1144DFA88" box="[900,1075,1320,1342]" country="Peru" municipality="Caravei" name="Panamericana Sur" pageId="0" pageNumber="297" stateProvince="Arequipa">Panamericana Sur</location>
,
<quantity id="F35ED4DDB35E277DFBC6FAA114D5FA88" box="[1085,1195,1320,1342]" metricMagnitude="2" metricUnit="m" metricValue="6.55" metricValueMax="9.6" metricValueMin="3.5" pageId="0" pageNumber="297" unit="m" value="655.0" valueMax="960.0" valueMin="350.0">
<elevation id="BF8B9E0BB35E277DFBC6FAA114D5FA88" box="[1085,1195,1320,1342]" metricMagnitude="2" metricUnit="m" metricValue="6.55" metricValueMax="9.6" metricValueMin="3.5" pageId="0" pageNumber="297" unit="m" value="655.0" valueMax="960.0" valueMin="350.0">350–960 m</elevation>
</quantity>
, sandy and rocky places,
<date id="40185FF8B35E277DFA64FAA1113DFAD4" pageId="0" pageNumber="297" value="1997-10-04">
<collectingDate id="505CA610B35E277DFA64FAA1113DFAD4" pageId="0" pageNumber="297" value="1997-10-04">4 October 1997</collectingDate>
</date>
,
<emphasis id="06D2A52AB35E277DFEB6FAC512EBFAD4" box="[333,661,1356,1377]" italics="true" pageId="0" pageNumber="297">
<collectorName id="99531CEEB35E277DFEB6FAC511C1FAD4" box="[333,447,1356,1377]" pageId="0" pageNumber="297">M. Weigend</collectorName>
&amp;
<collectorName id="99531CEEB35E277DFE26FAC5123BFAD4" box="[477,581,1356,1377]" pageId="0" pageNumber="297">H. Förther</collectorName>
97/738.
</emphasis>
<typeStatus id="EB1DC79AB35E277DFD61FAC51288FAD7" box="[666,758,1356,1378]" pageId="0" pageNumber="297" type="holotype">Holotype</typeStatus>
,
<collectionCode id="52B7E1FDB35E277DFD04FAC5134BFAD4" box="[767,821,1356,1377]" collectionName="MSB" pageId="0" pageNumber="297">MSB</collectionCode>
;
</materialsCitation>
<materialsCitation id="84CE7365B35E277DFCC5FAC51381FAD4" box="[830,1023,1356,1378]" collectingDate="1997-10-04" collectionCode="USM" collectorName="M. Weigend &amp; H. Forther" country="Peru" elevation="655" location="Lomas" municipality="Caravei" pageId="0" pageNumber="297" specimenCount="1" stateProvince="Arequipa" typeStatus="isotype">
<typeStatus id="EB1DC79AB35E277DFCC5FAC513F8FAD7" box="[830,902,1356,1378]" pageId="0" pageNumber="297" type="isotype">isotype</typeStatus>
,
<collectionCode id="52B7E1FDB35E277DFC6BFAC513BAFAD4" box="[912,964,1356,1377]" collectionName="USM" pageId="0" pageNumber="297">USM</collectionCode>
(n.v.)
</materialsCitation>
.
<figureCitation id="AC9D65BDB35E277DFBF3FAC5141CFAD4" box="[1032,1122,1356,1378]" captionStart="FIGURE 1" captionStartId="1.[136,229,2047,2069]" captionTargetBox="[195,1393,242,2020]" captionTargetId="figure-53@1.[195,1393,242,2020]" captionTargetPageId="1" captionText="FIGURE 1. Holotype of Lepidium weigendii." figureDoi="http://doi.org/10.5281/zenodo.13879339" httpUri="https://zenodo.org/record/13879339/files/figure.png" pageId="0" pageNumber="297">Figures 1</figureCitation>
,
<figureCitation id="AC9D65BDB35E277DFB95FAC51402FAD4" box="[1134,1148,1356,1377]" captionStart="FIGURE 2" captionStartId="2.[136,229,895,917]" captionTargetId="figure-24@2.[336,1252,190,871]" captionTargetPageId="2" captionText="FIGURE 2. Lepidium weigendii.A. Flower. B. Fruit showing persistent nectaries." figureDoi="http://doi.org/10.5281/zenodo.13992433" httpUri="https://zenodo.org/record/13992433/files/figure.png" pageId="0" pageNumber="297">2</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7CBC2AB3B35E277DFF73FA1B1520F883" pageId="0" pageNumber="297" type="description">
<paragraph id="34197938B35E277DFF73FA1B1520F883" blockId="0.[136,1452,1426,2062]" pageId="0" pageNumber="297">
<emphasis id="06D2A52AB35E277DFF73FA1B1151FA1F" bold="true" box="[136,303,1426,1450]" pageId="0" pageNumber="297">Description:—</emphasis>
Herbs, annual, densely hirsute. Trichomes spreading, straight,
<quantity id="F35ED4DDB35E277DFC33FA1B1436FA1C" box="[968,1096,1426,1450]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" pageId="0" pageNumber="297" unit="mm" value="0.35" valueMax="0.6" valueMin="0.1">0.1–0.6 mm</quantity>
long. Stems ca.
<quantity id="F35ED4DDB35E277DFB02FA1B1545FA1C" box="[1273,1339,1426,1450]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="297" unit="cm" value="30.0">30 cm</quantity>
tall, erect, branched below middle and above, densely hirsute. Basal leaves fall off early; middle cauline leaves oblong to oblongovate or elliptic, 1.5–4 ×
<quantity id="F35ED4DDB35E277DFE6DFA531185FA44" box="[406,507,1498,1522]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="2.0" metricValueMin="0.5" pageId="0" pageNumber="297" unit="cm" value="1.25" valueMax="2.0" valueMin="0.5">0.5–2 cm</quantity>
, sessile, hirsute, base auriculate, margin serrate, with 3–6 teeth on each side, apex acute. Racemes lax, elongated considerably in fruit; rachis straight, densely hirsute with straight trichomes to
<quantity id="F35ED4DDB35E277DFB33FA771566F9A0" box="[1224,1304,1534,1558]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="0" pageNumber="297" unit="mm" value="0.6">0.6 mm</quantity>
long; fruiting pedicels slender, straight, divaricate-ascending,
<quantity id="F35ED4DDB35E277DFD71F9AB128EF98C" box="[650,752,1570,1594]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="0" pageNumber="297" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
long, densely hirsute all around, terete, not winged or margined,
<quantity id="F35ED4DDB35E277DFF73F9CF115DF9E8" box="[136,291,1606,1630]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" metricValueMax="3.3" metricValueMin="2.5" pageId="0" pageNumber="297" unit="mm" value="0.29" valueMax="0.33" valueMin="0.25">0.25–0.33 mm</quantity>
wide. Sepals oblong,
<quantity id="F35ED4DDB35E277DFDE8F9CF12EDF9E8" box="[531,659,1606,1630]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="0" pageNumber="297" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
long, caducous, margin and apex white, puberulent outside; petals white, obovate, 2–2.5 ×
<quantity id="F35ED4DDB35E277DFEBBF9E311D3F934" box="[320,429,1642,1666]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="0" pageNumber="297" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
, base attenuate, apex rounded; claw
<quantity id="F35ED4DDB35E277DFCD5F9E313E7F934" box="[814,921,1642,1666]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="0" pageNumber="297" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
long; stamens 4, 2 median and 2 lateral; filaments
<quantity id="F35ED4DDB35E277DFF73F907108CF910" box="[136,242,1678,1702]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="2.0" metricValueMin="1.2" pageId="0" pageNumber="297" unit="mm" value="1.6" valueMax="2.0" valueMin="1.2">1.2–2 mm</quantity>
long; anthers oblong,
<quantity id="F35ED4DDB35E277DFE26F9071225F910" box="[477,603,1678,1702]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="0" pageNumber="297" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
long; nectar glands 4, conical,
<quantity id="F35ED4DDB35E277DFC5AF9071461F910" box="[929,1055,1678,1702]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="0" pageNumber="297" unit="mm" value="0.55" valueMax="0.7" valueMin="0.4">0.4–0.7 mm</quantity>
long, alternating with filament bases, becoming spreading and persistent well after fruit maturity; ovules 2 per ovary. Fruits dehiscent, orbicular to suborbicular, 4.5–6 ×
<quantity id="F35ED4DDB35E277DFF19F95F112FF958" box="[226,337,1750,1774]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="0" pageNumber="297" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 mm</quantity>
, strongly angustiseptate, not inflated; valves papery, veinless, densely hirsute with spreading trichomes, smooth, strongly keeled, wingless; apical notch
<quantity id="F35ED4DDB35E277DFD74F9731371F8A4" box="[655,783,1786,1810]" metricMagnitude="-4" metricUnit="m" metricValue="3" metricValueMax="4.0" metricValueMin="2.0" pageId="0" pageNumber="297" unit="mm" value="0.3" valueMax="0.4" valueMin="0.2">0.2–0.4 mm</quantity>
deep; septum complete; style
<quantity id="F35ED4DDB35E277DFBA2F97314BBF8A4" box="[1113,1221,1786,1810]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="0" pageNumber="297" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
long, exserted above fruit notch; stigma entire. Seeds reddish brown, ovate, wingless, 2.5–3 ×
<quantity id="F35ED4DDB35E277DFC68F8971469F880" box="[915,1047,1822,1846]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="0" pageNumber="297" unit="mm" value="1.35" valueMax="1.5" valueMin="1.2">1.2–1.5 mm</quantity>
; cotyledons incumbent, entire.
</paragraph>
</subSubSection>
<subSubSection id="7CBC2AB3B35E277DFF46F8CB135CF8CB" pageId="0" pageNumber="297" type="etymology">
<paragraph id="34197938B35E277DFF46F8CB135CF8CB" blockId="0.[136,1452,1426,2062]" pageId="0" pageNumber="297">
<emphasis id="06D2A52AB35E277DFF46F8CB114AF8EF" bold="true" box="[189,308,1858,1882]" pageId="0" pageNumber="297">Eponymy:</emphasis>
—The species is named after one of its collectors, Maximilian Weigend (
<date id="40185FF8B35E277DFBBEF8CB1566F8EF" box="[1093,1304,1858,1882]" pageId="0" pageNumber="297" value="1968-09-25">25 September 1968</date>
–), director of the Bonn Botanic Garden, Systematic Botany, and Herbarium.
</paragraph>
</subSubSection>
<subSubSection id="7CBC2AB3B35E277DFF46F80312C9F814" box="[189,695,1930,1954]" pageId="0" pageNumber="297" type="biology_ecology">
<paragraph id="34197938B35E277DFF46F80312C9F814" blockId="0.[136,1452,1426,2062]" box="[189,695,1930,1954]" pageId="0" pageNumber="297">
<emphasis id="06D2A52AB35E277DFF46F8031160F817" bold="true" box="[189,286,1930,1954]" pageId="0" pageNumber="297">Habitat:</emphasis>
—Sandy and rocky areas;
<quantity id="F35ED4DDB35E277DFDCCF80312CDF814" box="[567,691,1930,1954]" metricMagnitude="2" metricUnit="m" metricValue="6.55" metricValueMax="9.6" metricValueMin="3.5" pageId="0" pageNumber="297" unit="m" value="655.0" valueMax="960.0" valueMin="350.0">350–960 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="7CBC2AB3B35E277DFF46F82712F6F870" box="[189,648,1966,1990]" pageId="0" pageNumber="297" type="distribution">
<paragraph id="34197938B35E277DFF46F82712F6F870" blockId="0.[136,1452,1426,2062]" box="[189,648,1966,1990]" pageId="0" pageNumber="297">
<emphasis id="06D2A52AB35E277DFF46F827112CF873" bold="true" box="[189,338,1966,1990]" pageId="0" pageNumber="297">Distribution:</emphasis>
—Endemic to southern
<collectingCountry id="4CB139A8B35E277DFDABF82712FAF870" box="[592,644,1966,1989]" name="Peru" pageId="0" pageNumber="297">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="7CBC2AB3B35E277FFF46F85B1291F9A8" lastPageId="2" lastPageNumber="299" pageId="0" pageNumber="297" type="discussion">
<paragraph id="34197938B35E277CFF46F85B1528FF6E" blockId="0.[136,1452,1426,2062]" lastBlockId="1.[136,1451,159,219]" lastPageId="1" lastPageNumber="298" pageId="0" pageNumber="297">
<emphasis id="06D2A52AB35E277DFF46F85B11F8F85F" bold="true" box="[189,390,2002,2026]" pageId="0" pageNumber="297">Taxonomic notes:</emphasis>
—
<taxonomicName id="F3A602BBB35E277DFE5AF85B120BF85F" authorityName="Al-Shehbaz" authorityYear="2023" box="[417,629,2002,2026]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="0" pageNumber="297" phylum="Tracheophyta" rank="species" species="weigendii">
<emphasis id="06D2A52AB35E277DFE5AF85B120BF85F" box="[417,629,2002,2026]" italics="true" pageId="0" pageNumber="297">Lepidium weigendii</emphasis>
</taxonomicName>
is known thus far only from the
<typeStatus id="EB1DC79AB35E277DFC22F85B1479F85F" box="[985,1031,2002,2026]" pageId="0" pageNumber="297">type</typeStatus>
locality. It is most closely related to
<emphasis id="06D2A52AB35E277DFA6DF85A1098F7BB" italics="true" pageId="0" pageNumber="297">
<taxonomicName id="F3A602BBB35E277DFA6DF85A109CF7BB" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="0" pageNumber="297" phylum="Tracheophyta" rank="species" species="crassius">L. crassius</taxonomicName>
,
</emphasis>
another South Peruvian endemic to Dept.
<collectingRegion id="F662B7DAB35E277DFD54F87F1368F7BB" box="[687,790,2038,2062]" country="Peru" name="Arequipa" pageId="0" pageNumber="297">Arequipa</collectingRegion>
. From the latter, the novelty is easily distinguished by having densely hirsute (vs. glabrous) fruits 4.5–6 ×
<quantity id="F35ED4DDB35F277CFD9DFF1612ADFF02" box="[614,723,159,183]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="1" pageNumber="298" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 mm</quantity>
(vs. 3–4.5 ×
<quantity id="F35ED4DDB35F277CFC9FFF1613ABFF02" box="[868,981,159,183]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="1" pageNumber="298" unit="mm" value="3.75" valueMax="4.5" valueMin="3.0">3–4.5 mm</quantity>
), well-developed (vs. lacking) apical notch, clawed (vs. clawless) petals, and conical nectar glands that persist (vs. fall off) well after (vs. before) fruit maturity.
</paragraph>
<caption id="60D929B0B35F277CFF73F876123FF7A1" ID-DOI="http://doi.org/10.5281/zenodo.13879339" ID-Zenodo-Dep="13879339" box="[136,577,2047,2069]" httpUri="https://zenodo.org/record/13879339/files/figure.png" pageId="1" pageNumber="298" startId="1.[136,229,2047,2069]" targetBox="[195,1393,242,2020]" targetPageId="1" targetType="figure">
<paragraph id="34197938B35F277CFF73F876123FF7A1" blockId="1.[136,577,2047,2069]" box="[136,577,2047,2069]" pageId="1" pageNumber="298">
<emphasis id="06D2A52AB35F277CFF73F8761080F7A1" bold="true" box="[136,254,2047,2069]" pageId="1" pageNumber="298">FIGURE 1.</emphasis>
Holotype of
<taxonomicName id="F3A602BBB35F277CFE86F8761244F7A1" authorityName="Al-Shehbaz" authorityYear="2023" box="[381,570,2047,2068]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="1" pageNumber="298" phylum="Tracheophyta" rank="species" species="weigendii">
<emphasis id="06D2A52AB35F277CFE86F8761244F7A1" box="[381,570,2047,2068]" italics="true" pageId="1" pageNumber="298">Lepidium weigendii</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="60D929B0B35C277FFF73FCF613E1FC20" ID-DOI="http://doi.org/10.5281/zenodo.13992433" ID-Zenodo-Dep="13992433" box="[136,927,895,917]" httpUri="https://zenodo.org/record/13992433/files/figure.png" pageId="2" pageNumber="299" startId="2.[136,229,895,917]" targetBox="[349,1216,224,866]" targetPageId="2" targetType="figure">
<paragraph id="34197938B35C277FFF73FCF613E1FC20" blockId="2.[136,927,895,917]" box="[136,927,895,917]" pageId="2" pageNumber="299">
<emphasis id="06D2A52AB35C277FFF73FCF61080FC21" bold="true" box="[136,254,895,917]" pageId="2" pageNumber="299">FIGURE 2.</emphasis>
<emphasis id="06D2A52AB35C277FFEF8FCF611B7FC21" box="[259,457,895,916]" italics="true" pageId="2" pageNumber="299">
<taxonomicName id="F3A602BBB35C277FFEF8FCF611BAFC21" authorityName="Al-Shehbaz" authorityYear="2023" box="[259,452,895,916]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="weigendii">Lepidium weigendii</taxonomicName>
.
</emphasis>
A. Flower. B. Fruit showing persistent nectaries.
</paragraph>
</caption>
<paragraph id="34197938B35C277FFF46FC4C1245FB6C" blockId="2.[136,1452,965,1566]" pageId="2" pageNumber="299">
<taxonomicName id="F3A602BBB35C277FFF46FC4C11EFFC68" authorityName="Al-Shehbaz" authorityYear="2023" box="[189,401,965,989]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="weigendii">
<emphasis id="06D2A52AB35C277FFF46FC4C11EFFC68" box="[189,401,965,989]" italics="true" pageId="2" pageNumber="299">Lepidium weigendii</emphasis>
</taxonomicName>
is unique among all of the 55 native South American congeners by the presence of densely hirsute (vs. glabrous or at most puberulent) fruits. Of the nine
<taxonomicName id="F3A602BBB35C277FFD1FFC6013E1FBB4" box="[740,927,1001,1025]" pageId="2" pageNumber="299">
<emphasis id="06D2A52AB35C277FFD1FFC601337FBB4" box="[740,841,1001,1025]" italics="true" pageId="2" pageNumber="299">Lepidium</emphasis>
species
</taxonomicName>
on that continent that have puberulent fruits,
<taxonomicName id="F3A602BBB35C277FFA6DFC63118EFB93" authority="Al-Shehbaz (2017: 403)" authorityName="Al-Shehbaz" authorityPageNumber="403" authorityYear="2017" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="arequipa">
<emphasis id="06D2A52AB35C277FFA6DFC631099FB90" italics="true" pageId="2" pageNumber="299">L. arequipa</emphasis>
<bibRefCitation id="503704C9B35C277FFF16FB84118EFB93" author="Al-Shehbaz, I. A." box="[237,496,1037,1062]" pageId="2" pageNumber="299" pagination="403 - 413" refId="ref1540" refString="Al-Shehbaz, I. A. (2017) Five new species of Lepidium (Brassicaceae): L. pabotii (Iran), L. arequipa (Peru), and L. lapazianum, L. linearilobum, and L. stephan-beckii (Bolivia). Novon 25: 403 - 413. https: // doi. org / 10.3417 / D- 17 - 00006" type="journal article" year="2017">Al-Shehbaz (2017: 403)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="F3A602BBB35C277FFE01FB87133DFB90" authority="Al-Shehbaz (1999: 5)" authorityName="Al-Shehbaz" authorityPageNumber="5" authorityYear="1999" box="[506,835,1037,1062]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="beckii">
<emphasis id="06D2A52AB35C277FFE01FB871228FB90" box="[506,598,1037,1061]" italics="true" pageId="2" pageNumber="299">L. beckii</emphasis>
<bibRefCitation id="503704C9B35C277FFDA7FB84133DFB90" author="Al-Shehbaz, I. A." box="[604,835,1037,1062]" pageId="2" pageNumber="299" pagination="5 - 7" refId="ref1475" refString="Al-Shehbaz, I. A. (1999) Lepidium beckii (Brassicaceae), a new species from Bolivia. Novon 9: 5 - 7. https: // doi. org / 10.2307 / 3392106" type="journal article" year="1999">Al-Shehbaz (1999: 5)</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="F3A602BBB35C277FFC81FB871572FB93" authority="Thellung (1906: 222)" authorityName="Thellung" authorityPageNumber="222" authorityYear="1906" box="[890,1292,1037,1062]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="ecuadoriense">
<emphasis id="06D2A52AB35C277FFC81FB87145AFB90" box="[890,1060,1037,1061]" italics="true" pageId="2" pageNumber="299">L. ecuadoriense</emphasis>
<bibRefCitation id="503704C9B35C277FFBD1FB841572FB93" author="Thellung, A." box="[1066,1292,1037,1062]" pageId="2" pageNumber="299" pagination="1 - 340" refId="ref1855" refString="Thellung, A. (1906) Die Gattung Lepidium (L.) R. Br. Eine monographische Studie. Neue Denkschriften der Schweizerischen Naturforschenden Gesellschaft 41 (1): 1 - 340. https: // doi. org / 10.5962 / bhl. title. 15476" type="journal article" year="1906">Thellung (1906: 222)</bibRefCitation>
</taxonomicName>
are puberulent on the entire valves, though the last may be so only when the fruits are young. In
<taxonomicName id="F3A602BBB35C277FFBEDFBBB15F6FBFF" authority="Thellung (1928: 9)" authorityName="Thellung" authorityPageNumber="9" authorityYear="1928" box="[1046,1416,1073,1098]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="argentinum">
<emphasis id="06D2A52AB35C277FFBEDFBBB14D1FBFC" box="[1046,1199,1074,1097]" italics="true" pageId="2" pageNumber="299">L. argentinum</emphasis>
<bibRefCitation id="503704C9B35C277FFB43FBB815F6FBFF" author="Thellung, A." box="[1208,1416,1073,1098]" pageId="2" pageNumber="299" pagination="8 - 11" refId="ref1906" refString="Thellung, A. (1928) Amarantus hybride et Lepidium nouveau de l'Argentine. Physis (Buenos Aires) 9: 8 - 11. https: // doi. org / 10.3406 / jatba. 1928.4660" type="journal article" year="1928">Thellung (1928: 9)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="F3A602BBB35C277FFA6DFBBB1181FBDB" authority="Al-Shehbaz (1989: 1189)" authorityName="Al-Shehbaz" authorityPageNumber="1189" authorityYear="1989" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="boelckei">
<emphasis id="06D2A52AB35C277FFA6DFBBB109FFBD8" italics="true" pageId="2" pageNumber="299">L. boelckei</emphasis>
<bibRefCitation id="503704C9B35C277FFF11FBDC1181FBDB" author="Al-Shehbaz, I. A." box="[234,511,1109,1134]" pageId="2" pageNumber="299" pagination="1189 - 1192" refId="ref1427" refString="Al-Shehbaz, I. A. (1989) Lepidium boelckei and L. jujuyanum (Brassicaceae), new species from Jujuy, Argentina. Annals of the Missouri Botanical Garden 76: 1189 - 1192. https: // doi. org / 10.2307 / 2399714" type="journal article" year="1989">Al-Shehbaz (1989: 1189)</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="F3A602BBB35C277FFDC0FBDF1430FBD8" authority="(Watson 1876: 46) Rattan (1888: 25)" authorityName="Rattan" authorityPageNumber="25" authorityYear="1888" baseAuthorityName="Watson" baseAuthorityPageNumber="46" baseAuthorityYear="1876" box="[571,1102,1109,1134]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="strictum">
<emphasis id="06D2A52AB35C277FFDC0FBDF12CFFBD8" box="[571,689,1110,1133]" italics="true" pageId="2" pageNumber="299">L. strictum</emphasis>
(
<bibRefCitation id="503704C9B35C277FFD3AFBDF1303FBD8" author="Watson, S." box="[705,893,1109,1134]" pageId="2" pageNumber="299" refId="ref1944" refString="Watson, S. (1876) Botany [of California], vol. 1. Welch, Bigelow, &amp; Co., University Press, Cambridge, Mass [achusetts]., 628 pp." type="book" year="1876">Watson 1876: 46</bibRefCitation>
)
<bibRefCitation id="503704C9B35C277FFC76FBDF1430FBD8" author="Rattan, V." box="[909,1102,1109,1134]" pageId="2" pageNumber="299" refId="ref1797" refString="Rattan, V. (1888) Analytical key to West Coast botany, containing descriptions of sixteen hundred species of flowering plants growing west of the Sierra Nevada and Cascade Crests, from San Diego to Puget Sound, ed. 2. A. L. Bancroft and Company, San Francisco, 128 pp." type="book" year="1888">Rattan (1888: 25)</bibRefCitation>
</taxonomicName>
the fruits are either glabrous or puberulent. Finally, only the valve keel is minutely puberulent in
<taxonomicName id="F3A602BBB35C277FFCA8FBF314B0FB27" authority="Desvaux (1815: 180)" authorityName="Desvaux" authorityPageNumber="180" authorityYear="1815" box="[851,1230,1145,1170]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="pubescens">
<emphasis id="06D2A52AB35C277FFCA8FBF313A1FB24" box="[851,991,1145,1169]" italics="true" pageId="2" pageNumber="299">L. pubescens</emphasis>
<bibRefCitation id="503704C9B35C277FFC1CFBF314B0FB27" author="Desvaux, N. A." box="[999,1230,1145,1170]" pageId="2" pageNumber="299" pagination="145 - 187" refId="ref1609" refString="Desvaux, N. A. (1815) Sur la famille des Plants Cruciferes. Journal de Botanique, Appliquee a l'Agriculture, a la Pharmacie, a la Medecine et aux Arts 3 (4): 145 - 187." type="journal article" year="1815">Desvaux (1815: 180)</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="F3A602BBB35C277FFB20FBF31095FB00" authority="Philippi (1891: 5)" authorityName="Philippi" authorityPageNumber="5" authorityYear="1891" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="rahmeri">
<emphasis id="06D2A52AB35C277FFB20FBF31531FB24" box="[1243,1359,1145,1169]" italics="true" pageId="2" pageNumber="299">L. rahmeri</emphasis>
<bibRefCitation id="503704C9B35C277FFAACFBF01095FB00" author="Philippi, R. A." pageId="2" pageNumber="299" refId="ref1721" refString="Philippi, R. A. (1891) Catalogus praevius plantarum in itinere ad Tarapaca a Friderico Philippi lectarum. Annales del Museo Nacional, Santiago de Chile 2: 1 - 6 [Cruciferae]. Also published as Verzeichniss der von Friedrich Philippi auf der Hohebene der Provinzen Antofagasta und Tarapaca gesammelten Pflanzenvaufgestellt von Dr. R. A. Philippi. F. A. Brockhaus, Leipzig, 96 pp." type="book" year="1891">Philippi (1891: 5)</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="F3A602BBB35C277FFED8FB17129AFB03" authority="Al-Shehbaz (2017: 412)" authorityName="Al-Shehbaz" authorityPageNumber="412" authorityYear="2017" box="[291,740,1181,1206]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="stephan-beckii">
<emphasis id="06D2A52AB35C277FFED8FB1711A4FB00" box="[291,474,1181,1205]" italics="true" pageId="2" pageNumber="299">L. stephan-beckii</emphasis>
<bibRefCitation id="503704C9B35C277FFE1BFB14129AFB03" author="Al-Shehbaz, I. A." box="[480,740,1181,1206]" pageId="2" pageNumber="299" pagination="403 - 413" refId="ref1540" refString="Al-Shehbaz, I. A. (2017) Five new species of Lepidium (Brassicaceae): L. pabotii (Iran), L. arequipa (Peru), and L. lapazianum, L. linearilobum, and L. stephan-beckii (Bolivia). Novon 25: 403 - 413. https: // doi. org / 10.3417 / D- 17 - 00006" type="journal article" year="2017">Al-Shehbaz (2017: 412)</bibRefCitation>
</taxonomicName>
. In addition,
<taxonomicName id="F3A602BBB35C277FFC96FB17138EFB00" authorityName="Al-Shehbaz" authorityYear="2023" box="[877,1008,1181,1205]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="weigendii">
<emphasis id="06D2A52AB35C277FFC96FB17138EFB00" box="[877,1008,1181,1205]" italics="true" pageId="2" pageNumber="299">L. weigendii</emphasis>
</taxonomicName>
differs from all these by its persistent (vs. caducous) nectar glands at the fruit base.
</paragraph>
<paragraph id="34197938B35C277FFF46FB6C1530FADC" blockId="2.[136,1452,965,1566]" pageId="2" pageNumber="299">
<taxonomicName id="F3A602BBB35C277FFF46FB6C11EDFB48" authorityName="Al-Shehbaz" authorityYear="2023" box="[189,403,1253,1277]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="weigendii">
<emphasis id="06D2A52AB35C277FFF46FB6C11EDFB48" box="[189,403,1253,1277]" italics="true" pageId="2" pageNumber="299">Lepidium weigendii</emphasis>
</taxonomicName>
may be confused with the Peruvian endemic
<taxonomicName id="F3A602BBB35C277FFC71FB6F144DFB48" box="[906,1075,1253,1277]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="cyclocarpum">
<emphasis id="06D2A52AB35C277FFC71FB6F144DFB48" box="[906,1075,1253,1277]" italics="true" pageId="2" pageNumber="299">L. cyclocarpum</emphasis>
</taxonomicName>
, which is also hirsute annual with orbicular, notched fruits, but it differs by the auriculate cauline leaves, four stamens, clawed petals, hirsute, non-reticulate fruits 4.5–6 ×
<quantity id="F35ED4DDB35C277FFEDAFAA411EFFAF0" box="[289,401,1325,1349]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="2" pageNumber="299" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 mm</quantity>
, and persistent nectar glands. By contrast,
<taxonomicName id="F3A602BBB35C277FFCA6FAA7147AFAF0" box="[861,1028,1325,1349]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="cyclocarpum">
<emphasis id="06D2A52AB35C277FFCA6FAA7147AFAF0" box="[861,1028,1325,1349]" italics="true" pageId="2" pageNumber="299">L. cyclocarpum</emphasis>
</taxonomicName>
has non-auriculate cauline leaves, only two median stamens, non-clawed petals, glabrous, reticulate fruits 3.5–4 ×
<quantity id="F35ED4DDB35C277FFC53FADB1469FADC" box="[936,1047,1362,1386]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.8" metricValueMin="3.0" pageId="2" pageNumber="299" unit="mm" value="3.4" valueMax="3.8" valueMin="3.0">3–3.8 mm</quantity>
, and caducous nectar glands.
</paragraph>
<paragraph id="34197938B35C277FFF46FAFC1291F9A8" blockId="2.[136,1452,965,1566]" pageId="2" pageNumber="299">
The Chilean-endemic
<taxonomicName id="F3A602BBB35C277FFE51FAFC1322FA3B" authority="Philippi (1860: 8)" authorityName="Philippi" authorityPageNumber="8" authorityYear="1860" box="[426,860,1397,1422]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="spathulatum">
<emphasis id="06D2A52AB35C277FFE51FAFC12E7FA38" box="[426,665,1397,1421]" italics="true" pageId="2" pageNumber="299">Lepidium spathulatum</emphasis>
<bibRefCitation id="503704C9B35C277FFD64FAFC1322FA3B" author="Philippi, R. A." box="[671,860,1397,1422]" pageId="2" pageNumber="299" refId="ref1674" refString="Philippi, R. A. (1860) Florula Atacamensis, seu enumeration plantarum, quas in itinere per desertum Atacamense. Sumptibus Eduardi Anton, Halis Saxonum, 62 pp. https: // doi. org / 10.5962 / bhl. title. 162865" type="book" year="1860">Philippi (1860: 8)</bibRefCitation>
</taxonomicName>
is a third species with orbicular fruits and hirsute stems that might also be confused with
<taxonomicName id="F3A602BBB35C277FFE1EFA131217FA04" authorityName="Al-Shehbaz" authorityYear="2023" box="[485,617,1433,1457]" class="Magnoliopsida" family="Brassicaceae" genus="Lepidium" kingdom="Plantae" order="Brassicales" pageId="2" pageNumber="299" phylum="Tracheophyta" rank="species" species="weigendii">
<emphasis id="06D2A52AB35C277FFE1EFA131217FA04" box="[485,617,1433,1457]" italics="true" pageId="2" pageNumber="299">L. weigendii</emphasis>
</taxonomicName>
. However, it differs from the present novelty by being a perennial (vs. annual), with exauriculate (vs. auriculate) cauline leaves, adaxially puberulent and abaxially glabrous fruiting pedicels (vs. densely hirsute all around), petals undifferentiated (vs. differentiated) into blade and claw, two median (vs. two lateral and two median) stamens, and glabrous (vs. densely hirsute) fruits
</paragraph>
</subSubSection>
</treatment>
</document>
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