164 lines
17 KiB
XML
164 lines
17 KiB
XML
<document id="8E082226C25C332E74A91E952ECE3B31" ID-CLB-Dataset="45627" ID-DOI="10.5281/zenodo.211674" ID-GBIF-Dataset="d13a52a2-c0aa-45e9-9c6a-58613b627afb" ID-ISSN="1175-5326" ID-Zenodo-Dep="211674" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460429898246" checkinUser="plazi" docAuthor="Tilbrook, Kevin J." docDate="2012" docId="5C370C4EFFBFFFA9FF646E751EB39DA3" docLanguage="en" docName="zt03165p063.pdf" docOrigin="Zootaxa 3165" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Bryopesanser ecphymatotes Tilbrook, 2012, n. sp." docType="treatment" docVersion="9" lastPageNumber="55" masterDocId="A00E7436FFB1FFB9FFF369001A099F6F" masterDocTitle="Review of the bryozoan genus Bryopesanser Tilbrook, 2006 (Escharinidae: Cheilostomata) with the description of 11 new species" masterLastPageNumber="63" masterPageNumber="39" pageNumber="53" updateTime="1698331236171" updateUser="ExternalLinkService">
|
||
<mods:mods id="2A65570FABBD0CC845B80FC48AB9F65B" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="215B148A3FE0C6C84516285722D5E7A3">
|
||
<mods:title id="09AE217E31A02AEA9A8B78F65F3FBE82">Review of the bryozoan genus Bryopesanser Tilbrook, 2006 (Escharinidae: Cheilostomata) with the description of 11 new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="AAB61D5C42E562320526B39FE9B69DE4" type="personal">
|
||
<mods:role id="B352BF6C3E34CF6FBDAC60FD05FCDFA1">
|
||
<mods:roleTerm id="91A5DCE22B1CCFF73CE43132A13B0B47">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DF2ECA362BB10666B91673DCFF4DC03F">Tilbrook, Kevin J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="EE22ECA54C7AAD929B26C7353BDA9187">text</mods:typeOfResource>
|
||
<mods:relatedItem id="061658E69375540E01035A88CCA69786" type="host">
|
||
<mods:titleInfo id="C75D2C5CC6314AE5A079B473D193CAC3">
|
||
<mods:title id="FDACE668B20A04620281C99346201070">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="E3E2679D3D119A1A1CA354AB17764020">
|
||
<mods:date id="7C0E64F8292BB385C31BCD58B9405016">2012</mods:date>
|
||
<mods:detail id="280E0D593E6910DF4BB30EDD12E74402" type="volume">
|
||
<mods:number id="205A57E484B71017FE9CBCAC85F17021">3165</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="BFDEF21050509C0F9FADB69E5D9220FD" unit="page">
|
||
<mods:start id="DCFB0C7D279E186C6FD97B162EB6BE68">39</mods:start>
|
||
<mods:end id="910655D5F46F06B883C029718F120E56">63</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="E0DC4B6185B853A4319FF387110652A9">journal article</mods:classification>
|
||
<mods:identifier id="B26EA70D2233758DA61D3121A14F9B34" type="CLB-Dataset">45627</mods:identifier>
|
||
<mods:identifier id="5C5B1AFCCF9FC7B164A7F4993AE7CB67" type="DOI">10.5281/zenodo.211674</mods:identifier>
|
||
<mods:identifier id="E5EEBAF8FD7BD7B09556F421648C8176" type="GBIF-Dataset">d13a52a2-c0aa-45e9-9c6a-58613b627afb</mods:identifier>
|
||
<mods:identifier id="BF206C50756CAD3487A4D74E12B942F2" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="517374FACEB76DEDBE567CF0582D1123" type="Zenodo-Dep">211674</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="5C370C4EFFBFFFA9FF646E751EB39DA3" ID-DOI="http://doi.org/10.5281/zenodo.6493219" ID-GBIF-Taxon="119429742" ID-Zenodo-Dep="6493219" LSID="urn:lsid:plazi:treatment:5C370C4EFFBFFFA9FF646E751EB39DA3" httpUri="http://treatment.plazi.org/id/5C370C4EFFBFFFA9FF646E751EB39DA3" lastPageId="16" lastPageNumber="55" pageId="14" pageNumber="53">
|
||
<subSubSection id="9C84EED3FFBFFFB7FF646E751BA298DF" pageId="14" pageNumber="53" type="nomenclature">
|
||
<paragraph id="D421BD58FFBFFFB7FF646E75182698E0" blockId="14.[151,559,1909,1968]" box="[151,559,1909,1935]" pageId="14" pageNumber="53">
|
||
<heading id="8F690A34FFBFFFB7FF646E75182698E0" bold="true" box="[151,559,1909,1935]" fontSize="11" level="1" pageId="14" pageNumber="53" reason="1">
|
||
<emphasis id="E6EA614AFFBFFFB7FF646E75182698E0" bold="true" box="[151,559,1909,1935]" pageId="14" pageNumber="53">
|
||
<taxonomicName id="139EC6DBFFBFFFB7FF646E751BEF98E0" box="[151,486,1909,1935]" class="Gymnolaemata" family="Escharinidae" genus="Bryopesanser" kingdom="Animalia" order="Cheilostomatida" pageId="14" pageNumber="53" phylum="Bryozoa" rank="species" species="ecphymatotes" status="sp. nov.">
|
||
<emphasis id="E6EA614AFFBFFFB7FF646E751BEF98E0" bold="true" box="[151,486,1909,1935]" italics="true" pageId="14" pageNumber="53">Bryopesanser ecphymatotes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDD9DC31FFBFFFB7FE1F6E75182698E0" box="[492,559,1909,1935]" pageId="14" pageNumber="53" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D421BD58FFBFFFB7FF646E981BA298DF" blockId="14.[151,559,1909,1968]" box="[151,427,1944,1968]" pageId="14" pageNumber="53">
|
||
(
|
||
<figureCitation id="4CA5A1DDFFBFFFB7FF6D6E981B4A98DF" box="[158,323,1944,1968]" captionStart="FIGURES 35 – 37" captionStartId="15.[151,263,1717,1740]" captionTargetBox="[171,1415,193,1694]" captionTargetId="figure@15.[171,1416,193,1696]" captionTargetPageId="15" captionText="FIGURES 35 – 37, Bryopesanser ecphymatotes n. sp. Holotype specimen, NHMUK 1889.8. 21.19, South China Sea. Figures 38 – 40, Bryopesanser hebelomaia n. sp. Holotype specimen, NHMUK ex 1936.12.30.60 B, Sri Lanka. Scale bars: 35, 500 µm; 36, 200 µm; 37, 100 µm; 38, 100 µm; 39, 200 µm; 40, 100 µm." httpUri="https://zenodo.org/record/211682/files/figure.png" pageId="14" pageNumber="53">Figures 35–37</figureCitation>
|
||
, Table 1)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="80E1EDD0FFBEFFB6FF646FB519139867" httpUri="https://zenodo.org/record/211682/files/figure.png" pageId="15" pageNumber="54" targetBox="[171,1415,193,1694]" targetPageId="15">
|
||
<paragraph id="D421BD58FFBEFFB6FF646FB519139867" blockId="15.[151,1436,1717,1800]" pageId="15" pageNumber="54">
|
||
<emphasis id="E6EA614AFFBEFFB6FF646FB518B699A4" bold="true" box="[151,703,1717,1740]" pageId="15" pageNumber="54">
|
||
FIGURES 35–37,
|
||
<taxonomicName id="139EC6DBFFBEFFB6FEA86FB6187499A4" box="[347,637,1717,1739]" class="Gymnolaemata" family="Escharinidae" genus="Bryopesanser" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="54" phylum="Bryozoa" rank="species" species="ecphymatotes" status="sp. nov.">
|
||
<emphasis id="E6EA614AFFBEFFB6FEA86FB6187499A4" bold="true" box="[347,637,1717,1739]" italics="true" pageId="15" pageNumber="54">Bryopesanser ecphymatotes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDD9DC31FFBEFFB6FD776FB618B699A4" box="[644,703,1718,1739]" pageId="15" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Holotype specimen, NHMUK 1889.8.21.19, South China Sea.
|
||
<emphasis id="E6EA614AFFBEFFB6FAB86FB518399986" bold="true" pageId="15" pageNumber="54">
|
||
Figures 38–40,
|
||
<taxonomicName id="139EC6DBFFBEFFB6FF106FD51BF99985" box="[227,496,1748,1770]" class="Gymnolaemata" family="Escharinidae" genus="Bryopesanser" kingdom="Animalia" order="Cheilostomatida" pageId="15" pageNumber="54" phylum="Bryozoa" rank="species" species="hebelomaia" status="sp. nov.">
|
||
<emphasis id="E6EA614AFFBEFFB6FF106FD51BF99985" bold="true" box="[227,496,1748,1770]" italics="true" pageId="15" pageNumber="54">Bryopesanser hebelomaia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDD9DC31FFBEFFB6FE046FD418399986" box="[503,560,1748,1769]" pageId="15" pageNumber="54" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Holotype specimen, NHMUK ex 1936.12.30.60B, Sri Lanka. Scale bars:
|
||
<emphasis id="E6EA614AFFBEFFB6FAD36FD41F309986" bold="true" box="[1312,1337,1748,1769]" pageId="15" pageNumber="54">35</emphasis>
|
||
, 500 µm;
|
||
<emphasis id="E6EA614AFFBEFFB6FF646FF31AB99867" bold="true" box="[151,176,1779,1800]" pageId="15" pageNumber="54">36</emphasis>
|
||
, 200 µm;
|
||
<emphasis id="E6EA614AFFBEFFB6FEEA6FF31B3B9867" bold="true" box="[281,306,1779,1800]" pageId="15" pageNumber="54">37</emphasis>
|
||
, 100 µm;
|
||
<emphasis id="E6EA614AFFBEFFB6FE686FF31BBD9867" bold="true" box="[411,436,1779,1800]" pageId="15" pageNumber="54">38</emphasis>
|
||
, 100 µm;
|
||
<emphasis id="E6EA614AFFBEFFB6FDEF6FF3183C9867" bold="true" box="[540,565,1779,1800]" pageId="15" pageNumber="54">39</emphasis>
|
||
, 200 µm;
|
||
<emphasis id="E6EA614AFFBEFFB6FD6D6FF318BE9867" bold="true" box="[670,695,1779,1800]" pageId="15" pageNumber="54">40</emphasis>
|
||
, 100 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="9C84EED3FFBEFFB6FF646E351E53981E" pageId="15" pageNumber="54" type="materials_examined">
|
||
<paragraph id="D421BD58FFBEFFB6FF646E351E53981E" blockId="15.[151,1436,1845,2016]" pageId="15" pageNumber="54">
|
||
<emphasis id="E6EA614AFFBEFFB6FF646E351B0F9821" bold="true" box="[151,262,1845,1870]" pageId="15" pageNumber="54">Material.</emphasis>
|
||
<typeStatus id="0B2503FAFFBEFFB6FEFD6E351B7C9822" box="[270,373,1845,1869]" pageId="15" pageNumber="54" type="holotype">
|
||
<emphasis id="E6EA614AFFBEFFB6FEFD6E351B7C9822" box="[270,373,1845,1869]" italics="true" pageId="15" pageNumber="54">Holotype</emphasis>
|
||
</typeStatus>
|
||
(here selected):
|
||
<collectionCode id="B28F259DFFBEFFB6FDC26E3618929822" LSID="urn:lsid:biocol.org:col:34665" box="[561,667,1846,1869]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="15" pageNumber="54">NHMUK</collectionCode>
|
||
1889.8.21.19, Tizard Reef, South
|
||
<collectingCountry id="AC89FDC8FFBEFFB6FBD16E351E6C9822" box="[1058,1125,1845,1869]" name="China" pageId="15" pageNumber="54">China</collectingCountry>
|
||
Sea. [Tizard Bank is a reef area in the northern part of the Spratly Group of islands,
|
||
<geoCoordinate id="B1AADB9FFFBEFFB6FCE06E5919AC981E" box="[787,933,1881,1905]" direction="north" orientation="latitude" pageId="15" pageNumber="54" precision="15" value="10.264167">10°15’51” N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="B1AADB9FFFBEFFB6FC426E591E46981E" box="[945,1103,1881,1905]" direction="east" orientation="longitude" pageId="15" pageNumber="54" precision="15" value="114.40305">114°24’11” E</geoCoordinate>
|
||
.]
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84EED3FFBEFFA9FF346E7F1BBB9E0B" lastPageId="16" lastPageNumber="55" pageId="15" pageNumber="54" type="description">
|
||
<paragraph id="D421BD58FFBEFFA9FF346E7F1BBB9E0B" blockId="15.[151,1436,1845,2016]" lastBlockId="16.[151,1437,151,716]" lastPageId="16" lastPageNumber="55" pageId="15" pageNumber="54">
|
||
<emphasis id="E6EA614AFFBEFFB6FF346E7F1B5098F7" bold="true" box="[199,345,1919,1944]" pageId="15" pageNumber="54">Description.</emphasis>
|
||
Colony large, ca
|
||
<quantity id="136610BDFFBEFFB6FDD56E80187F98F7" box="[550,630,1920,1944]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="54" unit="mm" value="15.0">15 mm</quantity>
|
||
<quantity id="136610BDFFBEFFB6FD856E7D189498F7" box="[630,669,1917,1944]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="15" pageNumber="54" unit="in" value="2.0">2 in</quantity>
|
||
area. Autozooids 0.60–0.80 x
|
||
<quantity id="136610BDFFBEFFB6FC0D6E801EAC98F7" box="[1022,1189,1920,1944]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="15" pageNumber="54" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.50–0.60 mm</quantity>
|
||
, hexagonal or irregularly polygonal, distinct, separated by grooves. Frontal shield covered by small ‘pimples’, slightly convex, evenly perforated by numerous pores, with small areolar septular pores at each lateral and proximal angle. Primary orifice longer than wide, ca 0.10 x
|
||
<quantity id="136610BDFFA1FFA9FE17699718459FC0" box="[484,588,151,175]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="16" pageNumber="55" unit="mm" value="0.09">0.09 mm</quantity>
|
||
excluding sinus, anter deeply arched, proximal border straight, condyles smooth, sinus drop-shaped, as wide as long. Oral spines 7, evenly spaced. Proximal peristome flared, not developed into a median mucro. Avicularia originating lateral to second pair of spines, rostra medium-sized, open-ended distally, crossbar complete, distomedially directed, mandibles fan-shaped. Ovicell hyperstomial, producing a thickened proximal edge, a raised frontal process lacking. Ovicellate zooids with 6 oral spines, the most distal pair incorporated into ovicell.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84EED3FFA1FFA9FF3468701F709EE8" box="[199,1401,367,392]" pageId="16" pageNumber="55" type="etymology">
|
||
<paragraph id="D421BD58FFA1FFA9FF3468701F709EE8" blockId="16.[151,1437,151,716]" box="[199,1401,367,392]" pageId="16" pageNumber="55">
|
||
<emphasis id="E6EA614AFFA1FFA9FF3468701B4F9EE7" bold="true" box="[199,326,368,392]" pageId="16" pageNumber="55">Etymology</emphasis>
|
||
. Greek
|
||
<emphasis id="E6EA614AFFA1FFA9FE6C6870182A9EE7" box="[415,547,368,392]" italics="true" pageId="16" pageNumber="55">ecphymatos</emphasis>
|
||
, eruption of pimples, alluding to the nature of the frontal shield sculpturing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84EED3FFA1FFA9FF3468941E1A9DC8" pageId="16" pageNumber="55" type="discussion">
|
||
<paragraph id="D421BD58FFA1FFA9FF3468941BF89D53" blockId="16.[151,1437,151,716]" pageId="16" pageNumber="55">
|
||
<emphasis id="E6EA614AFFA1FFA9FF3468941B329EC3" bold="true" box="[199,315,404,428]" pageId="16" pageNumber="55">Remarks.</emphasis>
|
||
<taxonomicName id="139EC6DBFFA1FFA9FEB66895188B9EC3" box="[325,642,404,428]" class="Gymnolaemata" family="Escharinidae" genus="Bryopesanser" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="55" phylum="Bryozoa" rank="species" species="ecphymatotes" status="sp. nov.">
|
||
<emphasis id="E6EA614AFFA1FFA9FEB66895188B9EC3" box="[325,642,404,428]" italics="true" pageId="16" pageNumber="55">Bryopesanser ecphymatotes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDD9DC31FFA1FFA9FD7E689418C69EC3" box="[653,719,404,428]" pageId="16" pageNumber="55" rank="species">
|
||
<emphasis id="E6EA614AFFA1FFA9FD7E689418C69EC3" bold="true" box="[653,719,404,428]" pageId="16" pageNumber="55">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is characterised by the details of the primary orifice with its straight proximal border, smooth condyles, sinus as wide as long, and flared peristome. The ‘pimpled’ surface of the frontal shield is also characteristic, although not unique among the
|
||
<taxonomicName id="139EC6DBFFA1FFA9FC4D68DD1E519E9B" box="[958,1112,477,500]" class="Gymnolaemata" family="Escharinidae" genus="Bryopesanser" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="55" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="E6EA614AFFA1FFA9FC4D68DD1E519E9B" box="[958,1112,477,500]" italics="true" pageId="16" pageNumber="55">Bryopesanser</emphasis>
|
||
</taxonomicName>
|
||
species described herein;
|
||
<taxonomicName id="139EC6DBFFA1FFA9FA7768DD1AFF9D77" class="Gymnolaemata" family="Escharinidae" genus="Bryopesanser" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="55" phylum="Bryozoa" rank="species" species="lobiones" status="sp. nov.">
|
||
<emphasis id="E6EA614AFFA1FFA9FA7768DD1AFF9D77" italics="true" pageId="16" pageNumber="55">B. lobiones</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDD9DC31FFA1FFA9FF0C6B001B489D77" box="[255,321,512,536]" pageId="16" pageNumber="55" rank="species">
|
||
<emphasis id="E6EA614AFFA1FFA9FF0C6B001B489D77" bold="true" box="[255,321,512,536]" pageId="16" pageNumber="55">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
also possesses a similar frontal-shield morphology but this species produces multiporous frontal pores and a peristomial mucro.
|
||
</paragraph>
|
||
<paragraph id="D421BD58FFA1FFA9FF346B491E1A9DC8" blockId="16.[151,1437,151,716]" pageId="16" pageNumber="55">
|
||
<taxonomicName id="139EC6DBFFA1FFA9FF346B4918099D0F" box="[199,512,584,608]" class="Gymnolaemata" family="Escharinidae" genus="Bryopesanser" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="55" phylum="Bryozoa" rank="species" species="ecphymatotes" status="sp. nov.">
|
||
<emphasis id="E6EA614AFFA1FFA9FF346B4918099D0F" box="[199,512,584,608]" italics="true" pageId="16" pageNumber="55">Bryopesanser ecphymatotes</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FDD9DC31FFA1FFA9FDF46B48184C9D0F" box="[519,581,584,608]" pageId="16" pageNumber="55" rank="species">
|
||
<emphasis id="E6EA614AFFA1FFA9FDF46B48184C9D0F" bold="true" box="[519,581,584,608]" pageId="16" pageNumber="55">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from the other species described herein in the shape of its oral proximal border, more specifically in its smooth oral condyles, and the ‘pimpled’ frontal shield; it also produces a flared peristome but lacks both a peristomial mucro and frontal process on the ovicell.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9C84EED3FFA1FFA9FF346BB31EB39DA3" box="[199,1210,691,716]" pageId="16" pageNumber="55" type="distribution">
|
||
<paragraph id="D421BD58FFA1FFA9FF346BB31EB39DA3" blockId="16.[151,1437,151,716]" box="[199,1210,691,716]" pageId="16" pageNumber="55">
|
||
<emphasis id="E6EA614AFFA1FFA9FF346BB31B699DA3" bold="true" box="[199,352,691,716]" pageId="16" pageNumber="55">Distribution.</emphasis>
|
||
The species is known from a single specimen found in the South
|
||
<collectingCountry id="AC89FDC8FFA1FFA9FBB36BB41E8A9DA3" box="[1088,1155,692,716]" name="China" pageId="16" pageNumber="55">China</collectingCountry>
|
||
Sea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |