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<document id="211D44B93C3880AC50ABA8F201452B37" ID-CLB-Dataset="46681" ID-DOI="10.11646/zootaxa.3754.2.6" ID-GBIF-Dataset="2200bba5-8275-4fa3-ae01-d1c618202084" ID-ISSN="1175-5326" ID-Zenodo-Dep="227398" ID-ZooBank="995C32F3-15B0-4616-A69C-EA8FAA9BB6AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460368133557" checkinUser="plazi" docAuthor="Pilato, Giovanni, Sabella, Giorgio &amp; Lisi, Oscar" docDate="2014" docId="1F5E505CF21AA32894DDFCBC0AE24D9C" docLanguage="en" docName="zt03754p184.pdf" docOrigin="Zootaxa 3754 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrobiotus insuetus Pilato, Sabella &amp; Lisi, 2014, sp. nov." docType="treatment" docVersion="10" lastPageNumber="179" masterDocId="E3672824F21BA32E944AFD080B1F4E65" masterDocTitle="Two new tardigrade species from Sicily" masterLastPageNumber="184" masterPageNumber="173" pageNumber="174" updateTime="1698295434153" updateUser="ExternalLinkService">
<mods:mods id="99CB3176FA2F538F4302A0462F7B2A9E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B2C2E08C155911F9B21D9CA3A7B77D32">
<mods:title id="60FA4B2A227E2345417FCDC43F9C38E7">Two new tardigrade species from Sicily</mods:title>
</mods:titleInfo>
<mods:name id="4ECAEE736A5131EA96BA2798C45611CD" type="personal">
<mods:role id="C905B6AEB12F03D5F3CACF4A55D81097">
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<mods:namePart id="AB8E22C8148B32363834433EFEADB723">Pilato, Giovanni</mods:namePart>
</mods:name>
<mods:name id="F87777F825EBDFFC4234182A1C0BE901" type="personal">
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<mods:namePart id="78BF5C21556A7ABED6A16D3C0DBDEFFF">Sabella, Giorgio</mods:namePart>
</mods:name>
<mods:name id="DF3E1B50730A3BF0365FABB33FDE6837" type="personal">
<mods:role id="BCF2683B76AB0A2F0D5C42705D0DF720">
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<mods:namePart id="9416B93B2FB66BBD893BC7B25C908575">Lisi, Oscar</mods:namePart>
</mods:name>
<mods:typeOfResource id="B0025BF47AC0F0B8CB6805A0EDDB1768">text</mods:typeOfResource>
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<mods:title id="1D42E97F73A444B013F3BEC112F80A5A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4A8CA8DDB8E0D0912D1FACF388EAB0A2">
<mods:date id="4FF7447C61039EECDAB674446796B6F3">2014</mods:date>
<mods:detail id="086189506721CFC2B435B575D4F04ED7" type="volume">
<mods:number id="A2F3D48FFE6CF9C740AE94B0E4E7C6A5">3754</mods:number>
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<mods:detail id="9DA56F9A46A45B5D9EB37E79241A6A8B" type="issue">
<mods:number id="381F5AA45D696B6B9A16630764D93088">2</mods:number>
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<mods:classification id="5F2133F05F4053043D79A3CE70134F82">journal article</mods:classification>
<mods:identifier id="AF8C7D77F73F13B19953B19316D5FA85" type="CLB-Dataset">46681</mods:identifier>
<mods:identifier id="5DC48A9236073894BF6F1FA37AE2A767" type="DOI">10.11646/zootaxa.3754.2.6</mods:identifier>
<mods:identifier id="AF3CA30F53FA6AF90C48F456D693A800" type="GBIF-Dataset">2200bba5-8275-4fa3-ae01-d1c618202084</mods:identifier>
<mods:identifier id="2D771A72480BCCDD04D065C8E3767555" type="ISSN">1175-5326</mods:identifier>
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<treatment id="1F5E505CF21AA32894DDFCBC0AE24D9C" ID-DOI="http://doi.org/10.5281/zenodo.5623119" ID-GBIF-Taxon="119411190" ID-Zenodo-Dep="5623119" LSID="urn:lsid:plazi:treatment:1F5E505CF21AA32894DDFCBC0AE24D9C" httpUri="http://treatment.plazi.org/id/1F5E505CF21AA32894DDFCBC0AE24D9C" lastPageId="6" lastPageNumber="179" pageId="1" pageNumber="174">
<subSubSection id="DFEDB2C1F21AA32F94DDFCBC0BE04F95" pageId="1" pageNumber="174" type="nomenclature">
<paragraph id="9748E14AF21AA32F94DDFCBC091E4FAA" blockId="1.[151,513,436,496]" box="[151,513,436,463]" pageId="1" pageNumber="174">
<heading id="CC005626F21AA32F94DDFCBC091E4FAA" bold="true" box="[151,513,436,463]" fontSize="11" level="1" pageId="1" pageNumber="174" reason="1">
<emphasis id="A5833D58F21AA32F94DDFCBC091E4FAA" bold="true" box="[151,513,436,463]" pageId="1" pageNumber="174">
<taxonomicName id="50F79AC9F21AA32F94DDFCBC0A834FAB" box="[151,412,436,462]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="1" pageNumber="174" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F21AA32F94DDFCBC0A834FAB" bold="true" box="[151,412,436,462]" italics="true" pageId="1" pageNumber="174">Macrobiotus insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F21AA32F95EEFCBD091E4FAA" box="[420,513,437,463]" pageId="1" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9748E14AF21AA32F94DDFCDF0BE04F95" blockId="1.[151,513,436,496]" box="[151,255,471,496]" pageId="1" pageNumber="174">
<figureCitation id="0FCCFDCFF21AA32F94DDFCDF0BC34F95" box="[151,220,471,496]" captionStart="FIGURE 1" captionStartId="3.[151,255,1844,1867]" captionTargetBox="[180,1370,402,1819]" captionTargetId="figure@3.[174,1372,397,1822]" captionTargetPageId="3" captionText="FIGURE 1. A-D, Macrobiotus insuetus sp. nov. A, Holotype buccal-pharyngeal apparatus. B, Paratype buccal armature (slide No. 5225). C, Holotype claws of leg II. Arrow indicates the lunule at the base of the external claw. D, Paratype claws of leg IV (slide No. 1424). The long, common basal tract and short secondary branch of both posterior and anterior claws on leg IV are markedly different from the claws on legs I – III. Scale bar = 10 µm." httpUri="https://zenodo.org/record/227399/files/figure.png" pageId="1" pageNumber="174">Figs 1</figureCitation>
,
<figureCitation id="0FCCFDCFF21AA32F94A0FCD00BE44F95" box="[234,251,472,496]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1526,1549]" captionTargetBox="[156,1430,198,1500]" captionTargetId="figure@4.[151,1435,194,1504]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Macrobiotus insuetus sp. nov. A, B, Holotype claws of leg IV. In A the posterior claw is in focus (arrow), and B the anterior claw (arrow). The long, common basal tract for both posterior and anterior claws is very clear. C, D, Eggs (slide No. 5225). D shows the reticular ornamentation of the processes (arrows). Scale bar = 10 µm." httpUri="https://zenodo.org/record/227400/files/figure.png" pageId="1" pageNumber="174">2</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DFEDB2C1F21AA32F94DDFF1709174D82" pageId="1" pageNumber="174" type="materials_examined">
<paragraph id="9748E14AF21AA32F94DDFF17092B4CC0" blockId="1.[151,1436,543,2009]" pageId="1" pageNumber="174">
<emphasis id="A5833D58F21AA32F94DDFF170A614C5D" bold="true" box="[151,382,543,568]" pageId="1" pageNumber="174">Material examined</emphasis>
: Moss sample, Nelson Park, Maniace (Catania):
<typeStatus id="484C5FE8F21AA32F978DFF170F364C5D" box="[967,1065,543,568]" pageId="1" pageNumber="174" type="holotype">holotype</typeStatus>
(slide No. 5520), 10
<typeStatus id="484C5FE8F21AA32F917BFF280E834C5D" box="[1329,1436,544,568]" pageId="1" pageNumber="174" type="paratype">paratypes</typeStatus>
(slides Nos. 5521-5526) and
<specimenCount id="81F12AC3F21AA32F95A4FF4C09244C38" box="[494,571,580,605]" pageId="1" pageNumber="174" type="egg">3 eggs</specimenCount>
(slides 5521, 5224 and 5225) (collected in
<date id="E349C78AF21AA32F900CFF4C0E1F4C39" box="[1094,1280,580,605]" pageId="1" pageNumber="174" value="2001-11">November 2001</date>
by Giovanni Pilato). Moss sample, Contrada Bauly, Palazzolo Acreide (Siracusa): 1
<typeStatus id="484C5FE8F21AA32F979FFF600F2A4CE5" box="[981,1077,616,640]" pageId="1" pageNumber="174" type="paratype">paratype</typeStatus>
(slide No. 1424) (collected by Angelo Zappalà in
<date id="E349C78AF21AA32F9524FF8409384CC0" box="[366,551,652,677]" pageId="1" pageNumber="174" value="1985-11">November 1985</date>
).
</paragraph>
<paragraph id="9748E14AF21AA32F948DFFA70E034C88" blockId="1.[151,1436,543,2009]" pageId="1" pageNumber="174">
<emphasis id="A5833D58F21AA32F948DFFA70A974CAD" bold="true" box="[199,392,687,712]" pageId="1" pageNumber="174">
<typeStatus id="484C5FE8F21AA32F948DFFA70A1D4CAD" box="[199,258,687,712]" pageId="1" pageNumber="174">Type</typeStatus>
repository
</emphasis>
:
<typeStatus id="484C5FE8F21AA32F95D6FFA7091B4CAD" box="[412,516,687,712]" pageId="1" pageNumber="174" type="holotype">Holotype</typeStatus>
and
<typeStatus id="484C5FE8F21AA32F960FFFB809AF4CAD" box="[581,688,688,712]" pageId="1" pageNumber="174" type="paratype">paratypes</typeStatus>
are deposited in the Binda &amp; Pilato collection, Department of Biological, Geological and Environmental Sciences, Section of Animal Biology, University of Catania.
</paragraph>
<paragraph id="9748E14AF21AA32F948DFFFF09174D82" blockId="1.[151,1436,543,2009]" pageId="1" pageNumber="174">
<emphasis id="A5833D58F21AA32F948DFFFF0ABA4D75" bold="true" box="[199,421,759,784]" pageId="1" pageNumber="174">Specific diagnosis:</emphasis>
Colourless; cuticle smooth without pores; eye spots absent; buccal-pharyngeal apparatus of
<taxonomicName id="50F79AC9F21AA32F94FCFE140A5B4D51" box="[182,324,796,820]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="1" pageNumber="174" phylum="Tardigrada" rank="genus">
<emphasis id="A5833D58F21AA32F94FCFE140A5B4D51" box="[182,324,796,820]" italics="true" pageId="1" pageNumber="174">Macrobiotus</emphasis>
</taxonomicName>
<typeStatus id="484C5FE8F21AA32F9501FE150A604D50" box="[331,383,797,821]" pageId="1" pageNumber="174">type</typeStatus>
; pharyngeal bulb with three macroplacoids and microplacoid close to the third macroplacoid; claws of
<taxonomicName id="50F79AC9F21AA32F94B4FE480AE04D3D" box="[254,511,832,856]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="1" pageNumber="174" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="A5833D58F21AA32F94B4FE480AE04D3D" box="[254,511,832,856]" italics="true" pageId="1" pageNumber="174">Macrobiotus hufelandi</emphasis>
</taxonomicName>
<typeStatus id="484C5FE8F21AA32F964DFE4809284D3D" box="[519,567,832,856]" pageId="1" pageNumber="174">type</typeStatus>
but on the first three pairs of legs they have a short common tract, while on the hind legs primary and secondary branch are joined in an extended common tract and the secondary branch, markedly shorter than on legs I–III, forms a right angle with the main branch. Accessory points and lunules present. Eggs, freely laid, spherical with conical, reticulate processes; egg shell not areolated, with small dots around the base of the processes.
</paragraph>
</subSubSection>
<subSubSection id="DFEDB2C1F21AA328948DFEFC0AE24D9C" lastPageId="6" lastPageNumber="179" pageId="1" pageNumber="174" type="description">
<paragraph id="9748E14AF21AA32F948DFEFC08F34B49" blockId="1.[151,1436,543,2009]" pageId="1" pageNumber="174">
<emphasis id="A5833D58F21AA32F948DFEFC09164A68" bold="true" box="[199,521,1012,1037]" pageId="1" pageNumber="174">
Description of the
<typeStatus id="484C5FE8F21AA32F95E8FEFC09164A68" box="[418,521,1012,1037]" pageId="1" pageNumber="174" type="holotype">holotype</typeStatus>
</emphasis>
: Body length 334 µm; colourless; cuticle smooth without pores; eye spots absent; Buccal-pharyngeal apparatus of
<taxonomicName id="50F79AC9F21AA32F964EF910098D4A55" box="[516,658,1048,1072]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="1" pageNumber="174" phylum="Tardigrada" rank="genus">
<emphasis id="A5833D58F21AA32F964EF910098D4A55" box="[516,658,1048,1072]" italics="true" pageId="1" pageNumber="174">Macrobiotus</emphasis>
</taxonomicName>
<typeStatus id="484C5FE8F21AA32F96D3F91009D64A55" box="[665,713,1048,1072]" pageId="1" pageNumber="174">type</typeStatus>
(buccal tube rigid with ventral lamina); mouth terminal with 10 peribuccal lamellae (
<figureCitation id="0FCCFDCFF21AA32F95CCF9340AD74A31" box="[390,456,1084,1109]" captionStart="FIGURE 1" captionStartId="3.[151,255,1844,1867]" captionTargetBox="[180,1370,402,1819]" captionTargetId="figure@3.[174,1372,397,1822]" captionTargetPageId="3" captionText="FIGURE 1. A-D, Macrobiotus insuetus sp. nov. A, Holotype buccal-pharyngeal apparatus. B, Paratype buccal armature (slide No. 5225). C, Holotype claws of leg II. Arrow indicates the lunule at the base of the external claw. D, Paratype claws of leg IV (slide No. 1424). The long, common basal tract and short secondary branch of both posterior and anterior claws on leg IV are markedly different from the claws on legs I – III. Scale bar = 10 µm." httpUri="https://zenodo.org/record/227399/files/figure.png" pageId="1" pageNumber="174">Fig. 1</figureCitation>
A); buccal armature composed of an anterior band of small teeth, a posterior ring of triangular teeth and, more caudally, three dorsal and three ventral transverse ridges (
<figureCitation id="0FCCFDCFF21AA32F9007F9570F994A1D" box="[1101,1158,1119,1144]" captionStart="FIGURE 1" captionStartId="3.[151,255,1844,1867]" captionTargetBox="[180,1370,402,1819]" captionTargetId="figure@3.[174,1372,397,1822]" captionTargetPageId="3" captionText="FIGURE 1. A-D, Macrobiotus insuetus sp. nov. A, Holotype buccal-pharyngeal apparatus. B, Paratype buccal armature (slide No. 5225). C, Holotype claws of leg II. Arrow indicates the lunule at the base of the external claw. D, Paratype claws of leg IV (slide No. 1424). The long, common basal tract and short secondary branch of both posterior and anterior claws on leg IV are markedly different from the claws on legs I – III. Scale bar = 10 µm." httpUri="https://zenodo.org/record/227399/files/figure.png" pageId="1" pageNumber="174">Fig.1</figureCitation>
A, B). Rigid buccal tube 35.8 µm long and 6.5 µm wide externally (
<emphasis id="A5833D58F21AA32F96CEF98D09864AF9" box="[644,665,1157,1180]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 18.2); stylet supports inserted on the buccal tube at 79.2 % of its length (
<emphasis id="A5833D58F21AA32F94A7F9A10A1D4AA5" box="[237,258,1193,1216]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 79.2). Pharyngeal bulb with apophyses, three rod-shaped macroplacoids and microplacoid close to the third macroplacoid (
<figureCitation id="0FCCFDCFF21AA32F9536F9C40AA34A81" box="[380,444,1228,1253]" captionStart="FIGURE 1" captionStartId="3.[151,255,1844,1867]" captionTargetBox="[180,1370,402,1819]" captionTargetId="figure@3.[174,1372,397,1822]" captionTargetPageId="3" captionText="FIGURE 1. A-D, Macrobiotus insuetus sp. nov. A, Holotype buccal-pharyngeal apparatus. B, Paratype buccal armature (slide No. 5225). C, Holotype claws of leg II. Arrow indicates the lunule at the base of the external claw. D, Paratype claws of leg IV (slide No. 1424). The long, common basal tract and short secondary branch of both posterior and anterior claws on leg IV are markedly different from the claws on legs I – III. Scale bar = 10 µm." httpUri="https://zenodo.org/record/227399/files/figure.png" pageId="1" pageNumber="174">Fig. 1</figureCitation>
A); first macroplacoid 6.0 µm long (
<emphasis id="A5833D58F21AA32F9717F9C5086D4A81" box="[861,882,1229,1252]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 16.8); second 4.4 µm (
<emphasis id="A5833D58F21AA32F90D9F9C50FB64A81" box="[1171,1193,1229,1252]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 12.3); third, with a preterminal constriction, 5.4 µm (
<emphasis id="A5833D58F21AA32F966BF9F909284B6D" box="[545,567,1265,1288]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 15.1); microplacoid well developed 4.2 µm long (
<emphasis id="A5833D58F21AA32F90EDF9F90FA34B6D" box="[1191,1212,1265,1288]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 11.7); the entire placoid row 21.5 µm (
<emphasis id="A5833D58F21AA32F95DBF81D0AB84B49" box="[401,423,1301,1324]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 60.1); the macroplacoid row 17.5 µm (
<emphasis id="A5833D58F21AA32F973DF81D08924B49" box="[887,909,1301,1324]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 48.9).
</paragraph>
<paragraph id="9748E14AF21AA32F948DF83F0E3C48B9" blockId="1.[151,1436,543,2009]" pageId="1" pageNumber="174">
Claws well developed, of
<taxonomicName id="50F79AC9F21AA32F95A7F83009F14B35" box="[493,750,1336,1360]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="1" pageNumber="174" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="A5833D58F21AA32F95A7F83009F14B35" box="[493,750,1336,1360]" italics="true" pageId="1" pageNumber="174">Macrobiotus hufelandi</emphasis>
</taxonomicName>
<typeStatus id="484C5FE8F21AA32F96BFF831083A4B34" box="[757,805,1337,1361]" pageId="1" pageNumber="174">type</typeStatus>
but distinctive. Claws of the first three pairs of legs are typically formed with the primary and secondary branches joined along a short common basal tract and are clearly bifurcated, the external claws clearly longer than the internal (
<tableCitation id="DA75D4F1F21AA32F9730F87708CB4BFD" box="[890,980,1407,1432]" captionStart="TABLE 1" captionStartId="2.[151,239,293,316]" captionText="TABLE 1. Measurements (in µm) and pt values of some Macrobiotus insuetus sp. nov. structures." pageId="1" pageNumber="174">Table 1</tableCitation>
;
<figureCitation id="0FCCFDCFF21AA32F97ADF8770F344BFD" box="[999,1067,1407,1432]" captionStart="FIGURE 1" captionStartId="3.[151,255,1844,1867]" captionTargetBox="[180,1370,402,1819]" captionTargetId="figure@3.[174,1372,397,1822]" captionTargetPageId="3" captionText="FIGURE 1. A-D, Macrobiotus insuetus sp. nov. A, Holotype buccal-pharyngeal apparatus. B, Paratype buccal armature (slide No. 5225). C, Holotype claws of leg II. Arrow indicates the lunule at the base of the external claw. D, Paratype claws of leg IV (slide No. 1424). The long, common basal tract and short secondary branch of both posterior and anterior claws on leg IV are markedly different from the claws on legs I – III. Scale bar = 10 µm." httpUri="https://zenodo.org/record/227399/files/figure.png" pageId="1" pageNumber="174">Fig. 1</figureCitation>
C). The claws of hind legs are markedly different as the primary and secondary branch of both posterior and anterior claws are joined for a much longer basal tract with the secondary branch bifurcating late, making it appear shorter, and forming a right angle with the main branch (Fig, 1D;
<figureCitation id="0FCCFDCFF21AA32F95B0F8E409234861" box="[506,572,1516,1541]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1526,1549]" captionTargetBox="[156,1430,198,1500]" captionTargetId="figure@4.[151,1435,194,1504]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Macrobiotus insuetus sp. nov. A, B, Holotype claws of leg IV. In A the posterior claw is in focus (arrow), and B the anterior claw (arrow). The long, common basal tract for both posterior and anterior claws is very clear. C, D, Eggs (slide No. 5225). D shows the reticular ornamentation of the processes (arrows). Scale bar = 10 µm." httpUri="https://zenodo.org/record/227400/files/figure.png" pageId="1" pageNumber="174">Fig. 2</figureCitation>
A-B). The claws of the hind legs are slightly shorter than those of legs II and III (
<tableCitation id="DA75D4F1F21AA32F948BFB070A0B484D" box="[193,276,1551,1576]" captionStart="TABLE 1" captionStartId="2.[151,239,293,316]" captionText="TABLE 1. Measurements (in µm) and pt values of some Macrobiotus insuetus sp. nov. structures." pageId="1" pageNumber="174">Table 1</tableCitation>
), and the posterior claws slightly longer than the anterior, though the difference is less marked than that between external and internal claws of legs I–III. Basal portion of all claws with round lateral angles. Length of the external and internal claws of legs II = 12.9 µm (
<emphasis id="A5833D58F21AA32F96F6FB5109CE4815" box="[700,721,1625,1648]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 36.0) and 10.2 µm (
<emphasis id="A5833D58F21AA32F9785FB5108FA4815" box="[975,997,1625,1648]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 28.5) respectively; on legs III = 12.8 µm (
<emphasis id="A5833D58F21AA32F9487FB750BFC48F1" box="[205,227,1661,1684]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 35.8) and 10.2 µm (
<emphasis id="A5833D58F21AA32F95A9FB750AE648F1" box="[483,505,1661,1684]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 28.5) respectively; posterior and anterior claws of legs IV = 11.9 µm (
<emphasis id="A5833D58F21AA32F9167FB750E5C48F1" box="[1325,1347,1661,1684]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 32.2) and 10.9 µm (
<emphasis id="A5833D58F21AA32F9573FBA90A5048DD" box="[313,335,1697,1720]" italics="true" pageId="1" pageNumber="174">pt</emphasis>
= 30.4) respectively. All claw main branches with long, thin accessory points (
<figureCitation id="0FCCFDCFF21AA32F909CFB970E3048DD" box="[1238,1327,1695,1720]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1526,1549]" captionTargetBox="[156,1430,198,1500]" captionTargetId="figure@4.[151,1435,194,1504]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Macrobiotus insuetus sp. nov. A, B, Holotype claws of leg IV. In A the posterior claw is in focus (arrow), and B the anterior claw (arrow). The long, common basal tract for both posterior and anterior claws is very clear. C, D, Eggs (slide No. 5225). D shows the reticular ornamentation of the processes (arrows). Scale bar = 10 µm." httpUri="https://zenodo.org/record/227400/files/figure.png" pageId="1" pageNumber="174">Fig. 2B</figureCitation>
). Smooth lunules present on all legs (
<figureCitation id="0FCCFDCFF21AA32F9580FBCC091B48B8" box="[458,516,1732,1757]" captionStart="FIGURE 1" captionStartId="3.[151,255,1844,1867]" captionTargetBox="[180,1370,402,1819]" captionTargetId="figure@3.[174,1372,397,1822]" captionTargetPageId="3" captionText="FIGURE 1. A-D, Macrobiotus insuetus sp. nov. A, Holotype buccal-pharyngeal apparatus. B, Paratype buccal armature (slide No. 5225). C, Holotype claws of leg II. Arrow indicates the lunule at the base of the external claw. D, Paratype claws of leg IV (slide No. 1424). The long, common basal tract and short secondary branch of both posterior and anterior claws on leg IV are markedly different from the claws on legs I – III. Scale bar = 10 µm." httpUri="https://zenodo.org/record/227399/files/figure.png" pageId="1" pageNumber="174">Fig.1</figureCitation>
C), and a cuticular bar present under the lunules on legs I–III (
<figureCitation id="0FCCFDCFF21AA32F90F4FBCC0FE148B9" box="[1214,1278,1732,1757]" captionStart="FIGURE 1" captionStartId="3.[151,255,1844,1867]" captionTargetBox="[180,1370,402,1819]" captionTargetId="figure@3.[174,1372,397,1822]" captionTargetPageId="3" captionText="FIGURE 1. A-D, Macrobiotus insuetus sp. nov. A, Holotype buccal-pharyngeal apparatus. B, Paratype buccal armature (slide No. 5225). C, Holotype claws of leg II. Arrow indicates the lunule at the base of the external claw. D, Paratype claws of leg IV (slide No. 1424). The long, common basal tract and short secondary branch of both posterior and anterior claws on leg IV are markedly different from the claws on legs I – III. Scale bar = 10 µm." httpUri="https://zenodo.org/record/227399/files/figure.png" pageId="1" pageNumber="174">Fig. 1</figureCitation>
C).
</paragraph>
<paragraph id="9748E14AF21AA32C948DFBE10A774E9D" blockId="1.[151,1436,543,2009]" lastBlockId="2.[151,1436,151,248]" lastPageId="2" lastPageNumber="175" pageId="1" pageNumber="174">
In the sample studied we found only
<taxonomicName id="50F79AC9F21AA32F9629FBE0084B489A" box="[611,852,1768,1792]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="1" pageNumber="174" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F21AA32F9629FBE0084B489A" box="[611,852,1768,1792]" italics="true" pageId="1" pageNumber="174">Macrobiotus insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F21AA32F9710FBEF08AC4965" box="[858,947,1767,1792]" pageId="1" pageNumber="174" rank="species">
<emphasis id="A5833D58F21AA32F9710FBEF08AC4965" bold="true" box="[858,947,1767,1792]" pageId="1" pageNumber="174">sp. nov.</emphasis>
</taxonomicNameLabel>
and three eggs, which we believe belong to this species. The eggs, freely laid, are spherical with conical processes (
<figureCitation id="0FCCFDCFF21AA32F97F5FA040F244941" box="[959,1083,1804,1829]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1526,1549]" captionTargetBox="[156,1430,198,1500]" captionTargetId="figure@4.[151,1435,194,1504]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Macrobiotus insuetus sp. nov. A, B, Holotype claws of leg IV. In A the posterior claw is in focus (arrow), and B the anterior claw (arrow). The long, common basal tract for both posterior and anterior claws is very clear. C, D, Eggs (slide No. 5225). D shows the reticular ornamentation of the processes (arrows). Scale bar = 10 µm." httpUri="https://zenodo.org/record/227400/files/figure.png" pageId="1" pageNumber="174">Fig. 2C, D</figureCitation>
). We were able to measure two eggs (
<tableCitation id="DA75D4F1F21AA32F9496FA270A2A492D" box="[220,309,1839,1864]" captionStart="TABLE 2" captionStartId="2.[151,239,1110,1133]" captionTargetBox="[151,1437,1190,1762]" captionTargetPageId="2" captionText="TABLE 2. Comparable egg measurements (in µm) and number of egg processes for: Macrobiotus insuetus sp. nov., M. wuzhishanensis, M. diffusus and M. dimentmani." httpUri="http://table.plazi.org/id/C388B1C2F219A32C94DDF95E09834AE8" pageId="1" pageNumber="174" tableUuid="C388B1C2F219A32C94DDF95E09834AE8">Table 2</tableCitation>
) and, despite a marked difference in diameter (59 and 72.6 µm), the number of processes to the circumference (11 and 13), the number of processes per hemisphere (31 for both) and the process height and basal diameter were all similar (
<tableCitation id="DA75D4F1F21AA32F958FFA7F090049F5" box="[453,543,1911,1936]" captionStart="TABLE 2" captionStartId="2.[151,239,1110,1133]" captionTargetBox="[151,1437,1190,1762]" captionTargetPageId="2" captionText="TABLE 2. Comparable egg measurements (in µm) and number of egg processes for: Macrobiotus insuetus sp. nov., M. wuzhishanensis, M. diffusus and M. dimentmani." httpUri="http://table.plazi.org/id/C388B1C2F219A32C94DDF95E09834AE8" pageId="1" pageNumber="174" tableUuid="C388B1C2F219A32C94DDF95E09834AE8">Table 2</tableCitation>
). The processes have a short, flexible terminal portion, sometimes forked and often broken (
<figureCitation id="0FCCFDCFF21AA32F9570FA940A8949D1" box="[314,406,1948,1973]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1526,1549]" captionTargetBox="[156,1430,198,1500]" captionTargetId="figure@4.[151,1435,194,1504]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Macrobiotus insuetus sp. nov. A, B, Holotype claws of leg IV. In A the posterior claw is in focus (arrow), and B the anterior claw (arrow). The long, common basal tract for both posterior and anterior claws is very clear. C, D, Eggs (slide No. 5225). D shows the reticular ornamentation of the processes (arrows). Scale bar = 10 µm." httpUri="https://zenodo.org/record/227400/files/figure.png" pageId="1" pageNumber="174">Fig. 2C</figureCitation>
), while the surface has a reticular design with small, almost isodiametric mesh (
<figureCitation id="0FCCFDCFF21AA32F9176FA940E9049D0" box="[1340,1423,1948,1973]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1526,1549]" captionTargetBox="[156,1430,198,1500]" captionTargetId="figure@4.[151,1435,194,1504]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Macrobiotus insuetus sp. nov. A, B, Holotype claws of leg IV. In A the posterior claw is in focus (arrow), and B the anterior claw (arrow). The long, common basal tract for both posterior and anterior claws is very clear. C, D, Eggs (slide No. 5225). D shows the reticular ornamentation of the processes (arrows). Scale bar = 10 µm." httpUri="https://zenodo.org/record/227400/files/figure.png" pageId="1" pageNumber="174">Fig.2D</figureCitation>
). Unfortunately, observation of the egg surface was difficult, but it is clearly not areolated and though the egg shell seemed to be smooth, small dots and some thin radial striae around the base of the processes are present. The eggs are most similar to three other
<emphasis id="A5833D58F219A32C95B9FDB4099D4EB1" box="[499,642,188,212]" italics="true" pageId="2" pageNumber="175">harmsworthi</emphasis>
<typeStatus id="484C5FE8F219A32C96C0FDB509A54EB0" box="[650,698,189,213]" pageId="2" pageNumber="175">type</typeStatus>
species,
<emphasis id="A5833D58F219A32C9768FDB50F474EB1" box="[802,1112,188,212]" italics="true" pageId="2" pageNumber="175">
<taxonomicName id="50F79AC9F219A32C9768FDB508BD4EB1" box="[802,930,188,212]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="diffusus">M. diffusus</taxonomicName>
,
<taxonomicName id="50F79AC9F219A32C97E4FDB50F474EB1" box="[942,1112,188,212]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="dimentmani">M. dimentmani</taxonomicName>
</emphasis>
and
<taxonomicName id="50F79AC9F219A32C90DAFDB50A7B4E9D" authority="Yin, Wang" authorityName="Yin, Wang" authorityYear="2011" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="wuzhishanensis">
<emphasis id="A5833D58F219A32C90DAFDB50E7B4EB1" box="[1168,1380,188,212]" italics="true" pageId="2" pageNumber="175">M. wuzhishanensis</emphasis>
<bibRefCitation id="F3669CBBF219A32C9126FDB40A7B4E9D" author="Yin" pageId="2" pageNumber="175" refString="Yin, H., Wang, L. &amp; Li, X. (2011) Two new species of the genus Macrobiotus (Tardigrada: Macrobiotidae) from China. Proceedings of the Biological Society of Washington, 124, 3, 165 - 178." type="journal article" year="2011">Yin, Wang, &amp; Li, 2011</bibRefCitation>
</taxonomicName>
.
</paragraph>
<caption id="C388B1C2F219A32C94DDFC2D0FAE4F59" box="[151,1201,292,316]" pageId="2" pageNumber="175">
<paragraph id="9748E14AF219A32C94DDFC2D0FAE4F59" blockId="2.[151,1361,292,1042]" box="[151,1201,292,316]" pageId="2" pageNumber="175">
<emphasis id="A5833D58F219A32C94DDFC2D0A154F59" bold="true" box="[151,266,293,316]" pageId="2" pageNumber="175">TABLE 1.</emphasis>
Measurements (in µm) and
<emphasis id="A5833D58F219A32C9672FC2D09524F59" box="[568,589,293,316]" italics="true" pageId="2" pageNumber="175">pt</emphasis>
values of some
<taxonomicName id="50F79AC9F219A32C96B0FC2C08C14F59" box="[762,990,292,316]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F219A32C96B0FC2C08C14F59" box="[762,990,292,316]" italics="true" pageId="2" pageNumber="175">Macrobiotus insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F219A32C97AFFC2D0F244F59" box="[997,1083,293,316]" pageId="2" pageNumber="175" rank="species">
<emphasis id="A5833D58F219A32C97AFFC2D0F244F59" bold="true" box="[997,1083,293,316]" pageId="2" pageNumber="175">sp. nov.</emphasis>
</taxonomicNameLabel>
structures.
</paragraph>
</caption>
<paragraph id="9748E14AF219A32C9609FC5D086C4F0E" blockId="2.[151,1361,292,1042]" box="[579,883,341,363]" pageId="2" pageNumber="175">
<taxonomicName id="50F79AC9F219A32C9609FC5D08034F0E" box="[579,796,341,363]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F219A32C9609FC5D08034F0E" box="[579,796,341,363]" italics="true" pageId="2" pageNumber="175">Macrobiotus insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F219A32C9769FC5D086C4F0E" box="[803,883,341,363]" pageId="2" pageNumber="175" rank="species">
<emphasis id="A5833D58F219A32C9769FC5D086C4F0E" bold="true" box="[803,883,341,363]" pageId="2" pageNumber="175">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="9748E14AF219A32C94D6FC710FA84FEA" blockId="2.[151,1361,292,1042]" box="[156,1207,377,399]" pageId="2" pageNumber="175">Slide No. 5526 5520 5523</paragraph>
<paragraph id="9748E14AF219A32C9609FC910FC24FCB" blockId="2.[151,1361,292,1042]" box="[579,1245,408,430]" pageId="2" pageNumber="175">
<typeStatus id="484C5FE8F219A32C9609FC9109834FCB" box="[579,668,409,430]" pageId="2" pageNumber="175" type="paratype">Paratype</typeStatus>
<typeStatus id="484C5FE8F219A32C972EFC9008DD4FCB" box="[868,962,408,430]" pageId="2" pageNumber="175" type="holotype">Holotype</typeStatus>
<typeStatus id="484C5FE8F219A32C90CFFC910FC24FCB" box="[1157,1245,409,430]" pageId="2" pageNumber="175" type="paratype">Paratype</typeStatus>
</paragraph>
<paragraph id="9748E14AF219A32C9609FCB50E264FB7" blockId="2.[151,1361,292,1042]" box="[579,1337,444,466]" pageId="2" pageNumber="175">
µm (
<emphasis id="A5833D58F219A32C9691FCB509F04FB7" box="[731,751,445,466]" italics="true" pageId="2" pageNumber="175">pt</emphasis>
) µm (
<emphasis id="A5833D58F219A32C97B7FCB50F0E4FB7" box="[1021,1041,445,466]" italics="true" pageId="2" pageNumber="175">pt</emphasis>
) µm (
<emphasis id="A5833D58F219A32C9157FCB50E2E4FB7" box="[1309,1329,445,466]" italics="true" pageId="2" pageNumber="175">pt</emphasis>
)
</paragraph>
<paragraph id="9748E14AF219A32C94D6FCE80F8E4F93" blockId="2.[151,1361,292,1042]" box="[156,1169,480,502]" pageId="2" pageNumber="175">Body length 293 334?</paragraph>
<paragraph id="9748E14AF219A32C94D6FF0C0FAE4C7F" blockId="2.[151,1361,292,1042]" box="[156,1201,516,538]" pageId="2" pageNumber="175">Buccal tube length 33.1 35.8 41.1</paragraph>
<paragraph id="9748E14AF219A32C94D6FF20091349F6" blockId="2.[151,1361,292,1042]" lastBlockId="2.[151,1436,1807,2012]" pageId="2" pageNumber="175">
External buccal tube width 6.3 (
<emphasis id="A5833D58F219A32C9691FF2008184C5B" box="[731,775,552,574]" italics="true" pageId="2" pageNumber="175">19.0</emphasis>
) 6.5 (
<emphasis id="A5833D58F219A32C97B7FF200F364C5B" box="[1021,1065,552,574]" italics="true" pageId="2" pageNumber="175">18.2</emphasis>
) 8.1 (
<emphasis id="A5833D58F219A32C9157FF200E564C5B" box="[1309,1353,552,574]" italics="true" pageId="2" pageNumber="175">19.7</emphasis>
) Style supports insertion point (
<emphasis id="A5833D58F219A32C9691FF4408184C07" box="[731,775,588,610]" italics="true" pageId="2" pageNumber="175">79.0</emphasis>
) (
<emphasis id="A5833D58F219A32C97B7FF440F364C07" box="[1021,1065,588,610]" italics="true" pageId="2" pageNumber="175">79.2</emphasis>
) (
<emphasis id="A5833D58F219A32C9157FF440E564C07" box="[1309,1353,588,610]" italics="true" pageId="2" pageNumber="175">79.4</emphasis>
) First macroplacoid 5.1 (
<emphasis id="A5833D58F219A32C9691FF7808184CE3" box="[731,775,624,646]" italics="true" pageId="2" pageNumber="175">15.4</emphasis>
) 6.0 (
<emphasis id="A5833D58F219A32C97B7FF780F364CE3" box="[1021,1065,624,646]" italics="true" pageId="2" pageNumber="175">16.8</emphasis>
) 6.6 (
<emphasis id="A5833D58F219A32C9157FF780E564CE3" box="[1309,1353,624,646]" italics="true" pageId="2" pageNumber="175">16.1</emphasis>
) Second macroplacoid 4.2 (
<emphasis id="A5833D58F219A32C9691FF9C08184CCF" box="[731,775,660,682]" italics="true" pageId="2" pageNumber="175">12.7</emphasis>
) 4.4 (
<emphasis id="A5833D58F219A32C97B7FF9C0F364CCF" box="[1021,1065,660,682]" italics="true" pageId="2" pageNumber="175">12.3</emphasis>
) 5.6 (
<emphasis id="A5833D58F219A32C9157FF9C0E564CCF" box="[1309,1353,660,682]" italics="true" pageId="2" pageNumber="175">13.6</emphasis>
) Third macroplacoid 5.2 (
<emphasis id="A5833D58F219A32C9691FFB008184CAB" box="[731,775,696,718]" italics="true" pageId="2" pageNumber="175">15.7</emphasis>
) 5.4 (
<emphasis id="A5833D58F219A32C97B7FFB00F364CAB" box="[1021,1065,696,718]" italics="true" pageId="2" pageNumber="175">15.1</emphasis>
) 6.3 (
<emphasis id="A5833D58F219A32C9157FFB00E564CAB" box="[1309,1353,696,718]" italics="true" pageId="2" pageNumber="175">15.3</emphasis>
) Microplacoid 3.9 (
<emphasis id="A5833D58F219A32C9696FFD4081A4C97" box="[732,773,732,754]" italics="true" pageId="2" pageNumber="175">11. 8</emphasis>
) 4.2 (
<emphasis id="A5833D58F219A32C97B7FFD40F394C94" box="[1021,1062,732,754]" italics="true" pageId="2" pageNumber="175">11. 7</emphasis>
) 4.6 (
<emphasis id="A5833D58F219A32C9157FFD40E594C97" box="[1309,1350,732,754]" italics="true" pageId="2" pageNumber="175">11. 2</emphasis>
) Placoid row 19.6 (
<emphasis id="A5833D58F219A32C9696FE0808174D73" box="[732,776,768,790]" italics="true" pageId="2" pageNumber="175">59.2</emphasis>
) 21.5 (
<emphasis id="A5833D58F219A32C97B7FE080F364D73" box="[1021,1065,768,790]" italics="true" pageId="2" pageNumber="175">60.1</emphasis>
) 23.9 (
<emphasis id="A5833D58F219A32C9157FE080E564D73" box="[1309,1353,768,790]" italics="true" pageId="2" pageNumber="175">58.2</emphasis>
) Macroplacoid row 15.3 (
<emphasis id="A5833D58F219A32C9696FE2C08174D5F" box="[732,776,804,826]" italics="true" pageId="2" pageNumber="175">46.2</emphasis>
) 17.5 (
<emphasis id="A5833D58F219A32C97B7FE2C0F364D5F" box="[1021,1065,804,826]" italics="true" pageId="2" pageNumber="175">48.9</emphasis>
) 19.4 (
<emphasis id="A5833D58F219A32C9157FE2C0E564D5F" box="[1309,1353,804,826]" italics="true" pageId="2" pageNumber="175">47.2</emphasis>
) External claw II 11.0 (
<emphasis id="A5833D58F219A32C9696FE4008174D3B" box="[732,776,840,862]" italics="true" pageId="2" pageNumber="175">33.2</emphasis>
) 12.9 (
<emphasis id="A5833D58F219A32C97B7FE400F364D3B" box="[1021,1065,840,862]" italics="true" pageId="2" pageNumber="175">36.0</emphasis>
) 13.1 (
<emphasis id="A5833D58F219A32C9157FE400E564D3B" box="[1309,1353,840,862]" italics="true" pageId="2" pageNumber="175">31.9</emphasis>
) Internal claw II 9.2 (
<emphasis id="A5833D58F219A32C9696FE6408174DE7" box="[732,776,876,898]" italics="true" pageId="2" pageNumber="175">27.8</emphasis>
) 10.2 (
<emphasis id="A5833D58F219A32C97B7FE640F364DE7" box="[1021,1065,876,898]" italics="true" pageId="2" pageNumber="175">28.5</emphasis>
) 10.5 (
<emphasis id="A5833D58F219A32C9157FE640E564DE7" box="[1309,1353,876,898]" italics="true" pageId="2" pageNumber="175">25.5</emphasis>
) External claw III 11.5 (
<emphasis id="A5833D58F219A32C9696FE9808174DC3" box="[732,776,912,934]" italics="true" pageId="2" pageNumber="175">34.7</emphasis>
) 12.8 (
<emphasis id="A5833D58F219A32C97B7FE980F364DC3" box="[1021,1065,912,934]" italics="true" pageId="2" pageNumber="175">35.8</emphasis>
) 13.2 (
<emphasis id="A5833D58F219A32C9157FE980E564DC3" box="[1309,1353,912,934]" italics="true" pageId="2" pageNumber="175">32.1</emphasis>
) Internal claw III 9.5 (
<emphasis id="A5833D58F219A32C9696FEBC08174DAF" box="[732,776,948,970]" italics="true" pageId="2" pageNumber="175">28.7</emphasis>
) 10.2 (
<emphasis id="A5833D58F219A32C97B7FEBC0F364DAF" box="[1021,1065,948,970]" italics="true" pageId="2" pageNumber="175">28.5</emphasis>
) 11.0 (
<emphasis id="A5833D58F219A32C9157FEBC0E564DAF" box="[1309,1353,948,970]" italics="true" pageId="2" pageNumber="175">26.8</emphasis>
) Posterior claw IV?? 11.9 (
<emphasis id="A5833D58F219A32C97B7FED00F364D8B" box="[1021,1065,984,1006]" italics="true" pageId="2" pageNumber="175">33.2</emphasis>
) 12.6 (
<emphasis id="A5833D58F219A32C9157FED00E564D8B" box="[1309,1353,984,1006]" italics="true" pageId="2" pageNumber="175">30.7</emphasis>
) Anterior claw IV?? 10.9 (
<emphasis id="A5833D58F219A32C97B7FEF40F364A77" box="[1021,1065,1020,1042]" italics="true" pageId="2" pageNumber="175">30.4</emphasis>
) 11.3 (
<emphasis id="A5833D58F219A32C9157FEF40E564A77" box="[1309,1353,1020,1042]" italics="true" pageId="2" pageNumber="175">27.5</emphasis>
)
<emphasis id="A5833D58F219A32C948DFA070A2B494D" bold="true" box="[199,308,1807,1832]" pageId="2" pageNumber="175">Remarks</emphasis>
: The
<typeStatus id="484C5FE8F219A32C95C1FA180AE9494D" box="[395,502,1808,1832]" pageId="2" pageNumber="175" type="paratype">paratypes</typeStatus>
are similar to the
<typeStatus id="484C5FE8F219A32C96B3FA070844494D" box="[761,859,1807,1832]" pageId="2" pageNumber="175" type="holotype">holotype</typeStatus>
in both morphological and metric characters; morphometric charaters of the
<typeStatus id="484C5FE8F219A32C95BAFA3B094D4929" box="[496,594,1843,1868]" pageId="2" pageNumber="175" type="holotype">holotype</typeStatus>
and two
<typeStatus id="484C5FE8F219A32C96F3FA3C083A4929" box="[697,805,1844,1868]" pageId="2" pageNumber="175" type="paratype">paratypes</typeStatus>
with measurable characters are reported in
<tableCitation id="DA75D4F1F219A32C9144FA3B0E7E4929" box="[1294,1377,1843,1868]" captionStart="TABLE 1" captionStartId="2.[151,239,293,316]" captionText="TABLE 1. Measurements (in µm) and pt values of some Macrobiotus insuetus sp. nov. structures." pageId="2" pageNumber="175">Table 1</tableCitation>
.
<emphasis id="A5833D58F219A32C948DFA5F0A574915" bold="true" box="[199,328,1879,1904]" pageId="2" pageNumber="175">Etymology</emphasis>
: The name
<taxonomicName id="50F79AC9F219A32C959EFA50092F490A" box="[468,560,1880,1903]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="insuetus">
<emphasis id="A5833D58F219A32C959EFA50092F490A" box="[468,560,1880,1903]" italics="true" pageId="2" pageNumber="175">insuetus</emphasis>
</taxonomicName>
(Latin: unusual) refers to the uniqueness of the very different hind leg claw shape from those of the legs I–III
</paragraph>
<caption id="C388B1C2F219A32C94DDF95E09834AE8" ID-Table-UUID="C388B1C2F219A32C94DDF95E09834AE8" httpUri="http://table.plazi.org/id/C388B1C2F219A32C94DDF95E09834AE8" pageId="2" pageNumber="175" targetBox="[151,1437,1190,1762]" targetIsTable="true" targetPageId="2">
<paragraph id="9748E14AF219A32C94DDF95E09834AE8" blockId="2.[151,1437,1109,1165]" pageId="2" pageNumber="175">
<emphasis id="A5833D58F219A32C94DDF95E0A154A08" bold="true" box="[151,266,1110,1133]" pageId="2" pageNumber="175">TABLE 2.</emphasis>
Comparable egg measurements (in µm) and number of egg processes for:
<taxonomicName id="50F79AC9F219A32C9064F95D0E0B4A08" box="[1070,1300,1109,1133]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F219A32C9064F95D0E0B4A08" box="[1070,1300,1109,1133]" italics="true" pageId="2" pageNumber="175">Macrobiotus insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F219A32C9151F95E0E6E4A08" box="[1307,1393,1110,1133]" pageId="2" pageNumber="175" rank="species">
<emphasis id="A5833D58F219A32C9151F95E0E6E4A08" bold="true" box="[1307,1393,1110,1133]" pageId="2" pageNumber="175">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="50F79AC9F219A32C9135F95E0A224AE8" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="wuzhishanensis">
<emphasis id="A5833D58F219A32C9135F95E0A224AE8" italics="true" pageId="2" pageNumber="175">M. wuzhishanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="50F79AC9F219A32C9500F97E0ADF4AE8" box="[330,448,1141,1165]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="diffusus">
<emphasis id="A5833D58F219A32C9500F97E0ADF4AE8" box="[330,448,1141,1165]" italics="true" pageId="2" pageNumber="175">M. diffusus</emphasis>
</taxonomicName>
and
<taxonomicName id="50F79AC9F219A32C95BEF97E09894AE8" box="[500,662,1141,1165]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="dimentmani">
<emphasis id="A5833D58F219A32C95BEF97E09894AE8" box="[500,662,1141,1165]" italics="true" pageId="2" pageNumber="175">M. dimentmani</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="9748E14AF219A32C95F3F9AF0AE24887" pageId="2" pageNumber="175">
<table id="E5F713EAF2195CD194DDF9AE0E824887" box="[151,1437,1190,1762]" gridcols="4" gridrows="9" pageId="2" pageNumber="175">
<tr id="29C7E308F2195CD194DDF9AE0E824ABE" box="[151,1437,1190,1243]" gridrow="0" pageId="2" pageNumber="175">
<th id="6A168A74F2195CD194DDF9AE09314ABE" box="[151,558,1190,1243]" gridcol="0" gridrow="0" pageId="2" pageNumber="175">
<taxonomicName id="50F79AC9F219A32C95F3F9AF09314AD9" box="[441,558,1191,1212]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F219A32C95F3F9AF09314AD9" box="[441,558,1191,1212]" italics="true" pageId="2" pageNumber="175">M. insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F219A32C958AF9CD090F4ABE" box="[448,528,1221,1243]" pageId="2" pageNumber="175" rank="species">
<emphasis id="A5833D58F219A32C958AF9CD090F4ABE" bold="true" box="[448,528,1221,1243]" pageId="2" pageNumber="175">sp. nov.</emphasis>
</taxonomicNameLabel>
</th>
<th id="6A168A74F2195CD19603F9AE09A14ABE" box="[585,702,1190,1243]" gridcol="1" gridrow="0" pageId="2" pageNumber="175">
<taxonomicName id="50F79AC9F219A32C9600F9AF09A14AD9" box="[586,702,1191,1212]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F219A32C9600F9AF09A14AD9" box="[586,702,1191,1212]" italics="true" pageId="2" pageNumber="175">M. insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F219A32C9600F9CD09854ABE" box="[586,666,1221,1243]" pageId="2" pageNumber="175" rank="species">
<emphasis id="A5833D58F219A32C9600F9CD09854ABE" bold="true" box="[586,666,1221,1243]" pageId="2" pageNumber="175">sp. nov.</emphasis>
</taxonomicNameLabel>
</th>
<th id="6A168A74F2195CD1969DF9AE08BC4ABE" box="[727,931,1190,1243]" gridcol="2" gridrow="0" pageId="2" pageNumber="175">
<emphasis id="A5833D58F219A32C969DF9AF08BC4AD9" box="[727,931,1190,1212]" italics="true" pageId="2" pageNumber="175">
<taxonomicName id="50F79AC9F219A32C969DF9AF08864AD9" box="[727,921,1190,1212]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="wuzhishanensis">M. wuzhishanensis</taxonomicName>
*
</emphasis>
</th>
<th id="6A168A74F2195CD19788F9AE0E824ABE" box="[962,1437,1190,1243]" gridcol="3" gridrow="0" pageId="2" pageNumber="175">
<emphasis id="A5833D58F219A32C9788F9AF0E944AD9" box="[962,1419,1190,1212]" italics="true" pageId="2" pageNumber="175">
<taxonomicName id="50F79AC9F219A32C9788F9AF0F2E4AD9" box="[962,1073,1190,1212]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="diffusus">M. diffusus</taxonomicName>
<taxonomicName id="50F79AC9F219A32C90B8F9AF0E944AD9" box="[1266,1419,1190,1212]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="dimentmani">M. dimentmani</taxonomicName>
</emphasis>
</th>
</tr>
<tr id="29C7E308F2195CD194DDF9E10E824A9A" box="[151,1437,1257,1279]" gridrow="1" pageId="2" pageNumber="175" rowspan-2="1">
<th id="6A168A74F2195CD194DDF9E109314A9A" box="[151,558,1257,1279]" gridcol="0" gridrow="1" pageId="2" pageNumber="175">Slide No. 5524</th>
<td id="6A168A74F2195CD19603F9E109A14A9A" box="[585,702,1257,1279]" gridcol="1" gridrow="1" pageId="2" pageNumber="175">5525</td>
<td id="6A168A74F2195CD19788F9E10E824A9A" box="[962,1437,1257,1279]" gridcol="3" gridrow="1" pageId="2" pageNumber="175">3466 5359</td>
</tr>
<tr id="29C7E308F2195CD194DDF8050E824B27" box="[151,1437,1293,1346]" gridrow="2" pageId="2" pageNumber="175">
<th id="6A168A74F2195CD194DDF80509314B27" box="[151,558,1293,1346]" gridcol="0" gridrow="2" pageId="2" pageNumber="175">Diameter without 59 processes</th>
<td id="6A168A74F2195CD19603F80509A14B27" box="[585,702,1293,1346]" gridcol="1" gridrow="2" pageId="2" pageNumber="175">72.6</td>
<td id="6A168A74F2195CD1969DF80508BC4B27" box="[727,931,1293,1346]" gridcol="2" gridrow="2" pageId="2" pageNumber="175">85.8</td>
<td id="6A168A74F2195CD19788F8050E824B27" box="[962,1437,1293,1346]" gridcol="3" gridrow="2" pageId="2" pageNumber="175">58–81 79–90</td>
</tr>
<tr id="29C7E308F2195CD194DDF8470E824BC6" box="[151,1437,1359,1443]" gridrow="3" pageId="2" pageNumber="175">
<th id="6A168A74F2195CD194DDF84709314BC6" box="[151,558,1359,1443]" gridcol="0" gridrow="3" pageId="2" pageNumber="175">Diameter with 73.0 processes</th>
<td id="6A168A74F2195CD19603F84709A14BC6" box="[585,702,1359,1443]" gridcol="1" gridrow="3" pageId="2" pageNumber="175">86.2</td>
<td id="6A168A74F2195CD1969DF84708BC4BC6" box="[727,931,1359,1443]" gridcol="2" gridrow="3" pageId="2" pageNumber="175">114</td>
<td id="6A168A74F2195CD19788F8470E824BC6" box="[962,1437,1359,1443]" gridcol="3" gridrow="3" pageId="2" pageNumber="175">? 106–120 (the terminal filaments are often bent or broken)</td>
</tr>
<tr id="29C7E308F2195CD194DDF8B90E824B80" box="[151,1437,1457,1509]" gridrow="4" pageId="2" pageNumber="175">
<th id="6A168A74F2195CD194DDF8B909314B80" box="[151,558,1457,1509]" gridcol="0" gridrow="4" pageId="2" pageNumber="175">Number of processes in the 13 circumference</th>
<td id="6A168A74F2195CD19603F8B909A14B80" box="[585,702,1457,1509]" gridcol="1" gridrow="4" pageId="2" pageNumber="175">11</td>
<td id="6A168A74F2195CD1969DF8B908BC4B80" box="[727,931,1457,1509]" gridcol="2" gridrow="4" pageId="2" pageNumber="175">16</td>
<td id="6A168A74F2195CD19788F8B90E824B80" box="[962,1437,1457,1509]" gridcol="3" gridrow="4" pageId="2" pageNumber="175">15–23 15–17 (generally16–20)</td>
</tr>
<tr id="29C7E308F2195CD194DDF8FC0E82484D" box="[151,1437,1524,1576]" gridrow="5" pageId="2" pageNumber="175">
<th id="6A168A74F2195CD194DDF8FC0931484D" box="[151,558,1524,1576]" gridcol="0" gridrow="5" pageId="2" pageNumber="175">Number of processes in the 31 hemisphere</th>
<td id="6A168A74F2195CD19603F8FC09A1484D" box="[585,702,1524,1576]" gridcol="1" gridrow="5" pageId="2" pageNumber="175">31</td>
<td id="6A168A74F2195CD1969DF8FC08BC484D" box="[727,931,1524,1576]" gridcol="2" gridrow="5" pageId="2" pageNumber="175">?</td>
<td id="6A168A74F2195CD19788F8FC0E82484D" box="[962,1437,1524,1576]" gridcol="3" gridrow="5" pageId="2" pageNumber="175">41–63 56–71</td>
</tr>
<tr id="29C7E308F2195CD194DDFB3E0E8248EF" box="[151,1437,1590,1674]" gridrow="6" pageId="2" pageNumber="175">
<th id="6A168A74F2195CD194DDFB3E093148EF" box="[151,558,1590,1674]" gridcol="0" gridrow="6" pageId="2" pageNumber="175">Process height 8.6</th>
<td id="6A168A74F2195CD19603FB3E09A148EF" box="[585,702,1590,1674]" gridcol="1" gridrow="6" pageId="2" pageNumber="175">7.9</td>
<td id="6A168A74F2195CD1969DFB3E08BC48EF" box="[727,931,1590,1674]" gridcol="2" gridrow="6" pageId="2" pageNumber="175">18.5</td>
<td id="6A168A74F2195CD19788FB3E0E8248EF" box="[962,1437,1590,1674]" gridcol="3" gridrow="6" pageId="2" pageNumber="175">9–16 16–17 (the terminal filaments are often bent or broken)</td>
</tr>
<tr id="29C7E308F2195CD194DDFB9F0E8248C8" box="[151,1437,1687,1709]" gridrow="7" pageId="2" pageNumber="175">
<th id="6A168A74F2195CD194DDFB9F093148C8" box="[151,558,1687,1709]" gridcol="0" gridrow="7" pageId="2" pageNumber="175">Process basal diameter 11.1–12.8</th>
<td id="6A168A74F2195CD19603FB9F09A148C8" box="[585,702,1687,1709]" gridcol="1" gridrow="7" pageId="2" pageNumber="175">13.1–15.2</td>
<td id="6A168A74F2195CD1969DFB9F08BC48C8" box="[727,931,1687,1709]" gridcol="2" gridrow="7" pageId="2" pageNumber="175">10.6</td>
<td id="6A168A74F2195CD19788FB9F0E8248C8" box="[962,1437,1687,1709]" gridcol="3" gridrow="7" pageId="2" pageNumber="175">7.8–14.5 10–14</td>
</tr>
<tr id="29C7E308F2195CD194DDFBC20E824887" box="[151,1437,1738,1762]" gridrow="8" pageId="2" pageNumber="175" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="6A168A74F2195CD194DDFBC209314887" box="[151,558,1738,1762]" gridcol="0" gridrow="8" pageId="2" pageNumber="175">
*) According to Yin
<emphasis id="A5833D58F219A32C9539FBC30ABE4887" box="[371,417,1738,1762]" italics="true" pageId="2" pageNumber="175">et al</emphasis>
. (2011).
</th>
</tr>
</table>
</paragraph>
<paragraph id="9748E14AF219A32D948DFA970A264F01" blockId="2.[151,1436,1807,2012]" lastBlockId="3.[151,1436,151,357]" lastPageId="3" lastPageNumber="176" pageId="2" pageNumber="175">
<emphasis id="A5833D58F219A32C948DFA970AD949DD" bold="true" box="[199,454,1951,1976]" pageId="2" pageNumber="175">Differential diagnosis</emphasis>
: Based on the egg structure and the claw shapes of legs I-III,
<taxonomicName id="50F79AC9F219A32C90C8FAA80E6C49D2" box="[1154,1395,1952,1976]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F219A32C90C8FAA80E6C49D2" box="[1154,1395,1952,1976]" italics="true" pageId="2" pageNumber="175">Macrobiotus insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F219A32C9130FA970BD749B9" pageId="2" pageNumber="175" rank="species">
<emphasis id="A5833D58F219A32C9130FA970BD749B9" bold="true" pageId="2" pageNumber="175">sp. nov.</emphasis>
</taxonomicNameLabel>
is similar to three other known
<emphasis id="A5833D58F219A32C9665FACC09A149B9" box="[559,702,1988,2012]" italics="true" pageId="2" pageNumber="175">harmsworthi</emphasis>
<typeStatus id="484C5FE8F219A32C968CFACC09E949B9" box="[710,758,1988,2012]" pageId="2" pageNumber="175">type</typeStatus>
species:
<taxonomicName id="50F79AC9F219A32C9717FACC0FBE49B9" box="[861,1185,1988,2012]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="wuhzishanensis">
<emphasis id="A5833D58F219A32C9717FACC0FBE49B9" box="[861,1185,1988,2012]" italics="true" pageId="2" pageNumber="175">Macrobiotus wuhzishanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="50F79AC9F219A32C90E5FACC0E8349B9" box="[1199,1436,1988,2012]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="175" phylum="Tardigrada" rank="species" species="diffusus">
<emphasis id="A5833D58F219A32C90E5FACC0E8349B9" box="[1199,1436,1988,2012]" italics="true" pageId="2" pageNumber="175">Macrobiotus diffusus</emphasis>
</taxonomicName>
and
<taxonomicName id="50F79AC9F218A32D9480FD900AF74ED5" box="[202,488,152,176]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="176" phylum="Tardigrada" rank="species" species="dimentmani">
<emphasis id="A5833D58F218A32D9480FD900AF74ED5" box="[202,488,152,176]" italics="true" pageId="3" pageNumber="176">Macrobiotus dimentmani</emphasis>
</taxonomicName>
. It differs from
<taxonomicName id="50F79AC9F218A32D96E7FD9008EB4ED5" box="[685,1012,152,176]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="176" phylum="Tardigrada" rank="species" species="wuhzishanensis">
<emphasis id="A5833D58F218A32D96E7FD9008EB4ED5" box="[685,1012,152,176]" italics="true" pageId="3" pageNumber="176">Macrobiotus wuhzishanensis</emphasis>
</taxonomicName>
in: lacking eye spots, having wider buccal tube, higher
<emphasis id="A5833D58F218A32D953CFDB50A934EB1" box="[374,396,189,212]" italics="true" pageId="3" pageNumber="176">pt</emphasis>
index values relative to placoids and claws, differently shaped hind leg claws from those of legs I–III and the posterior hind leg claws shorter than the external claws of legs II–III (in
<taxonomicName id="50F79AC9F218A32D9144FDE80A5A4F79" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="176" phylum="Tardigrada" rank="species" species="wuhzishanensis">
<emphasis id="A5833D58F218A32D9144FDE80A5A4F79" italics="true" pageId="3" pageNumber="176">Macrobiotus wuhzishanensis</emphasis>
</taxonomicName>
they all the same length) (
<tableCitation id="DA75D4F1F218A32D963DFC0C09D04F79" box="[631,719,260,285]" captionStart="TABLE 3" captionStartId="6.[151,239,152,175]" captionText="TABLE 3. Comparable measurements (in µm) and pt values of the holotype Macrobiotus insuetus sp. nov. with similar sized (where possible) specimens of: M. wuzhishanensis, M. diffusus and M. dimentmani." pageId="3" pageNumber="176">Table 3</tableCitation>
). The eggs of the two species are very similar but those of the new species are: smaller, with shorter processes, fewer processes in the circumference and less evident ring of basal dots (
<tableCitation id="DA75D4F1F218A32D949EFC440A334F01" box="[212,300,332,357]" captionStart="TABLE 2" captionStartId="2.[151,239,1110,1133]" captionTargetBox="[151,1437,1190,1762]" captionTargetPageId="2" captionText="TABLE 2. Comparable egg measurements (in µm) and number of egg processes for: Macrobiotus insuetus sp. nov., M. wuzhishanensis, M. diffusus and M. dimentmani." httpUri="http://table.plazi.org/id/C388B1C2F219A32C94DDF95E09834AE8" pageId="3" pageNumber="176" tableUuid="C388B1C2F219A32C94DDF95E09834AE8">Table 2</tableCitation>
).
</paragraph>
<caption id="C388B1C2F218A32D94DDFA3C0F2449CE" httpUri="https://zenodo.org/record/227399/files/figure.png" pageId="3" pageNumber="176" targetBox="[180,1370,402,1819]" targetPageId="3">
<paragraph id="9748E14AF218A32D94DDFA3C0F2449CE" blockId="3.[151,1436,1843,1963]" pageId="3" pageNumber="176">
<emphasis id="A5833D58F218A32D94DDFA3C0A49492E" bold="true" box="[151,342,1844,1867]" pageId="3" pageNumber="176">FIGURE 1. A-D,</emphasis>
<taxonomicName id="50F79AC9F218A32D9516FA3B095D492E" box="[348,578,1843,1867]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="176" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F218A32D9516FA3B095D492E" box="[348,578,1843,1867]" italics="true" pageId="3" pageNumber="176">Macrobiotus insuetus</emphasis>
</taxonomicName>
<emphasis id="A5833D58F218A32D9602FA3C09A0492E" bold="true" box="[584,703,1844,1867]" pageId="3" pageNumber="176">
<taxonomicNameLabel id="BEB08023F218A32D9602FA3C0981492E" box="[584,670,1844,1867]" pageId="3" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
A,
</emphasis>
Holotype buccal-pharyngeal apparatus.
<emphasis id="A5833D58F218A32D9024FA3C0F98492E" bold="true" box="[1134,1159,1844,1867]" pageId="3" pageNumber="176">B,</emphasis>
Paratype buccal armature (slide No. 5225).
<emphasis id="A5833D58F218A32D951DFA5C0A6E490E" bold="true" box="[343,369,1876,1899]" pageId="3" pageNumber="176">C,</emphasis>
Holotype claws of leg II. Arrow indicates the lunule at the base of the external claw.
<emphasis id="A5833D58F218A32D9156FA5C0E28490E" bold="true" box="[1308,1335,1876,1899]" pageId="3" pageNumber="176">D,</emphasis>
Paratype claws of leg IV (slide No.1424). The long, common basal tract and short secondary branch of both posterior and anterior claws on leg IV are markedly different from the claws on legs I–III. Scale bar = 10 µm.
</paragraph>
</caption>
<caption id="C388B1C2F21FA32A94DDF8FE0E224828" httpUri="https://zenodo.org/record/227400/files/figure.png" pageId="4" pageNumber="177" targetBox="[156,1430,198,1500]" targetPageId="4">
<paragraph id="9748E14AF21FA32A94DDF8FE0E224828" blockId="4.[151,1437,1525,1613]" pageId="4" pageNumber="177">
<emphasis id="A5833D58F21FA32A94DDF8FE0A404868" bold="true" box="[151,351,1526,1549]" pageId="4" pageNumber="177">FIGURE 2. A–D,</emphasis>
<taxonomicName id="50F79AC9F21FA32A9522F8FD09504868" box="[360,591,1525,1549]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="177" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F21FA32A9522F8FD09504868" box="[360,591,1525,1549]" italics="true" pageId="4" pageNumber="177">Macrobiotus insuetus</emphasis>
</taxonomicName>
<emphasis id="A5833D58F21FA32A9613F8FE09EA4868" bold="true" box="[601,757,1526,1549]" pageId="4" pageNumber="177">
<taxonomicNameLabel id="BEB08023F21FA32A9613F8FE09AE4868" box="[601,689,1526,1549]" pageId="4" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
A, B,
</emphasis>
Holotype claws of leg IV. In
<emphasis id="A5833D58F21FA32A900AF8FE0F4C4868" bold="true" box="[1088,1107,1526,1549]" pageId="4" pageNumber="177">A</emphasis>
the posterior claw is in focus (arrow), and
<emphasis id="A5833D58F21FA32A956AFB1E0A2D4848" bold="true" box="[288,306,1558,1581]" pageId="4" pageNumber="177">B</emphasis>
the anterior claw (arrow). The long, common basal tract for both posterior and anterior claws is very clear.
<emphasis id="A5833D58F21FA32A94DDFB3E0BCD4828" bold="true" box="[151,210,1590,1613]" pageId="4" pageNumber="177">C, D,</emphasis>
Eggs (slide No. 5225).
<emphasis id="A5833D58F21FA32A9585FB3E0AFD4828" bold="true" box="[463,482,1590,1613]" pageId="4" pageNumber="177">D</emphasis>
shows the reticular ornamentation of the processes (arrows). Scale bar = 10 µm.
</paragraph>
</caption>
<paragraph id="9748E14AF21FA32A948DFB7E0F7C4903" blockId="4.[151,1436,1653,2003]" pageId="4" pageNumber="177">
<taxonomicName id="50F79AC9F21FA32A948DFB7E0AA548EB" box="[199,442,1654,1678]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="177" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F21FA32A948DFB7E0AA548EB" box="[199,442,1654,1678]" italics="true" pageId="4" pageNumber="177">Macrobiotus insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F21FA32A9589FB7D090148EB" box="[451,542,1653,1678]" pageId="4" pageNumber="177" rank="species">
<emphasis id="A5833D58F21FA32A9589FB7D090748EB" bold="true" box="[451,536,1653,1678]" pageId="4" pageNumber="177">sp. nov</emphasis>
.
</taxonomicNameLabel>
is also similar to
<taxonomicName id="50F79AC9F21FA32A96BAFB7E0F1448EB" box="[752,1035,1654,1678]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="177" phylum="Tardigrada" rank="species" species="dimentmani">
<emphasis id="A5833D58F21FA32A96BAFB7E0F1448EB" box="[752,1035,1654,1678]" italics="true" pageId="4" pageNumber="177">Macrobiotus dimentmani</emphasis>
</taxonomicName>
but differs in: lacking eye spots, a slightly narrower buccal tube (
<emphasis id="A5833D58F21FA32A95B3FB93091048D7" box="[505,527,1691,1714]" italics="true" pageId="4" pageNumber="177">pt</emphasis>
index values
<quantity id="500F4CAFF21FA32A96F8FB92085D48D6" box="[690,834,1690,1715]" metricMagnitude="-1" metricUnit="m" metricValue="4.8133" metricValueMax="5.0038" metricValueMin="4.6228" pageId="4" pageNumber="177" unit="in" value="18.95" valueMax="19.7" valueMin="18.2">18.2–19.7 in</quantity>
<taxonomicName id="50F79AC9F21FA32A9706FB9308CF48D7" box="[844,976,1691,1714]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="177" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F21FA32A9706FB93087C48D7" box="[844,867,1691,1714]" italics="true" pageId="4" pageNumber="177">M</emphasis>
.
<emphasis id="A5833D58F21FA32A973EFB9308CF48D7" box="[884,976,1691,1714]" italics="true" pageId="4" pageNumber="177">insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F21FA32A9793FB920F2A48D6" box="[985,1077,1690,1715]" pageId="4" pageNumber="177" rank="species">
<emphasis id="A5833D58F21FA32A9793FB920F2A48D6" bold="true" box="[985,1077,1690,1715]" pageId="4" pageNumber="177">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<quantity id="500F4CAFF21FA32A900CFB920FC948D6" box="[1094,1238,1690,1715]" metricMagnitude="-1" metricUnit="m" metricValue="5.3975" metricValueMax="5.892799999999999" metricValueMin="4.9022" pageId="4" pageNumber="177" unit="in" value="21.25" valueMax="23.2" valueMin="19.3">19.3–23.2 in</quantity>
<taxonomicName id="50F79AC9F21FA32A9095FB930E9448D7" box="[1247,1419,1690,1714]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="177" phylum="Tardigrada" rank="species" species="dimentmani">
<emphasis id="A5833D58F21FA32A9095FB930E9448D7" box="[1247,1419,1690,1714]" italics="true" pageId="4" pageNumber="177">M. dimentmani</emphasis>
</taxonomicName>
), stylet supports inserted on the buccal tube at a slightly more caudal position (
<emphasis id="A5833D58F21FA32A9056FBB70F2D48B3" box="[1052,1074,1727,1750]" italics="true" pageId="4" pageNumber="177">pt</emphasis>
index value = 79.0–
<quantity id="500F4CAFF21FA32A9169FBB60E6B48B3" box="[1315,1396,1725,1750]" metricMagnitude="0" metricUnit="m" metricValue="2.01676" pageId="4" pageNumber="177" unit="in" value="79.4">79.4 in</quantity>
<taxonomicName id="50F79AC9F21FA32A9137FBB70BEC489F" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="177" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F21FA32A9137FBB70BEC489F" italics="true" pageId="4" pageNumber="177">M. insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F21FA32A94B1FBEA0A4C489F" box="[251,339,1762,1787]" pageId="4" pageNumber="177" rank="species">
<emphasis id="A5833D58F21FA32A94B1FBEA0A51489E" bold="true" box="[251,334,1762,1787]" pageId="4" pageNumber="177">sp. nov</emphasis>
.
</taxonomicNameLabel>
,
<quantity id="500F4CAFF21FA32A952EFBEA0AEE489E" box="[356,497,1762,1787]" metricMagnitude="0" metricUnit="m" metricValue="1.94691" metricValueMax="1.9685" metricValueMin="1.92532" pageId="4" pageNumber="177" unit="in" value="76.65" valueMax="77.5" valueMin="75.8">75.8–77.5 in</quantity>
<taxonomicName id="50F79AC9F21FA32A95B3FBEB09BC489F" box="[505,675,1762,1786]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="177" phylum="Tardigrada" rank="species" species="dimentmani">
<emphasis id="A5833D58F21FA32A95B3FBEB09BC489F" box="[505,675,1762,1786]" italics="true" pageId="4" pageNumber="177">M. dimentmani</emphasis>
</taxonomicName>
); both anterior and posterior claws of legs IV with a long common basal tract and short secondary branches, while in
<taxonomicName id="50F79AC9F21FA32A969BFA0F0864497B" box="[721,891,1798,1822]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="177" phylum="Tardigrada" rank="species" species="dimentmani">
<emphasis id="A5833D58F21FA32A969BFA0F0864497B" box="[721,891,1798,1822]" italics="true" pageId="4" pageNumber="177">M. dimentmani</emphasis>
</taxonomicName>
only the posterior claws have this characteristic (
<figureCitation id="0FCCFDCFF21FA32A94D5FA220BF34927" box="[159,236,1834,1859]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1526,1549]" captionTargetBox="[156,1430,198,1500]" captionTargetId="figure@4.[151,1435,194,1504]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Macrobiotus insuetus sp. nov. A, B, Holotype claws of leg IV. In A the posterior claw is in focus (arrow), and B the anterior claw (arrow). The long, common basal tract for both posterior and anterior claws is very clear. C, D, Eggs (slide No. 5225). D shows the reticular ornamentation of the processes (arrows). Scale bar = 10 µm." httpUri="https://zenodo.org/record/227400/files/figure.png" pageId="4" pageNumber="177">Figs. 2</figureCitation>
A-B and 3A). Characters of the eggs differ in: smaller dimensions, fewer processes, shorter processes and more rarely forked (
<figureCitation id="0FCCFDCFF21FA32A953DFA450AEC4903" box="[375,499,1869,1894]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1526,1549]" captionTargetBox="[156,1430,198,1500]" captionTargetId="figure@4.[151,1435,194,1504]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Macrobiotus insuetus sp. nov. A, B, Holotype claws of leg IV. In A the posterior claw is in focus (arrow), and B the anterior claw (arrow). The long, common basal tract for both posterior and anterior claws is very clear. C, D, Eggs (slide No. 5225). D shows the reticular ornamentation of the processes (arrows). Scale bar = 10 µm." httpUri="https://zenodo.org/record/227400/files/figure.png" pageId="4" pageNumber="177">Figs 2C, D</figureCitation>
and
<figureCitation id="0FCCFDCFF21FA32A9660FA4609524903" box="[554,589,1870,1894]" captionStart="FIGURE 3. A, B" captionStartId="5.[151,255,1833,1856]" captionTargetBox="[155,1431,454,1809]" captionTargetId="figure@5.[150,1435,447,1809]" captionTargetPageId="5" captionText="FIGURE 3. A, B, Macrobiotus dimentmani. A, Claws of leg IV (slide No. 5359). It can been seen that the posterior and anterior claws are of different shapes, with only the posterior claws possessing the long, common basal tract (arrow). B, Detail of the egg (slide No. 5359). C, D, Macrobiotus diffusus. C, Claws of leg IV (slide No. 5527). As with M. dimentmani the claws are of different shapes, with only the posterior claws with a long, common basal tract (arrow). D, Detail of the egg (slide No. 3466). Arrows indicate some radial striae of the egg shell. Scale bar = 10 µm." httpUri="https://zenodo.org/record/227401/files/figure.png" pageId="4" pageNumber="177">3B</figureCitation>
;
<tableCitation id="DA75D4F1F21FA32A9613FA4509B14903" box="[601,686,1869,1894]" captionStart="TABLE 2" captionStartId="2.[151,239,1110,1133]" captionTargetBox="[151,1437,1190,1762]" captionTargetPageId="2" captionText="TABLE 2. Comparable egg measurements (in µm) and number of egg processes for: Macrobiotus insuetus sp. nov., M. wuzhishanensis, M. diffusus and M. dimentmani." httpUri="http://table.plazi.org/id/C388B1C2F219A32C94DDF95E09834AE8" pageId="4" pageNumber="177" tableUuid="C388B1C2F219A32C94DDF95E09834AE8">Table 2</tableCitation>
), and less visible dots on the egg shell.
</paragraph>
<paragraph id="9748E14AF21FA32B948DFA7A098D4FE2" blockId="4.[151,1436,1653,2003]" lastBlockId="5.[151,1436,151,392]" lastPageId="5" lastPageNumber="178" pageId="4" pageNumber="177">
<taxonomicName id="50F79AC9F21FA32A948DFA7A0AA549EF" box="[199,442,1906,1930]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="177" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F21FA32A948DFA7A0AA549EF" box="[199,442,1906,1930]" italics="true" pageId="4" pageNumber="177">Macrobiotus insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F21FA32A9589FA7A090049EE" box="[451,543,1906,1931]" pageId="4" pageNumber="177" rank="species">
<emphasis id="A5833D58F21FA32A9589FA7A090049EE" bold="true" box="[451,543,1906,1931]" pageId="4" pageNumber="177">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from the third similar species,
<taxonomicName id="50F79AC9F21FA32A9791FA7A0FDD49EF" box="[987,1218,1906,1930]" genus="Macrobotus" pageId="4" pageNumber="177" rank="species" species="diffusus">
<emphasis id="A5833D58F21FA32A9791FA7A0FDD49EF" box="[987,1218,1906,1930]" italics="true" pageId="4" pageNumber="177">Macrobotus diffusus</emphasis>
</taxonomicName>
, in some claw and egg characters. In the new species, both the anterior and posterior claws of leg IV have a long common basal tract (in
<taxonomicName id="50F79AC9F21FA32A9488FAB20AAC49B7" box="[194,435,1978,2002]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="4" pageNumber="177" phylum="Tardigrada" rank="species" species="diffusus">
<emphasis id="A5833D58F21FA32A9488FAB20AAC49B7" box="[194,435,1978,2002]" italics="true" pageId="4" pageNumber="177">Macrobiotus diffusus</emphasis>
</taxonomicName>
only the posterior claws have this characteristic) (
<figureCitation id="0FCCFDCFF21FA32A905AFAB20F6649B7" box="[1040,1145,1978,2003]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1526,1549]" captionTargetBox="[156,1430,198,1500]" captionTargetId="figure@4.[151,1435,194,1504]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Macrobiotus insuetus sp. nov. A, B, Holotype claws of leg IV. In A the posterior claw is in focus (arrow), and B the anterior claw (arrow). The long, common basal tract for both posterior and anterior claws is very clear. C, D, Eggs (slide No. 5225). D shows the reticular ornamentation of the processes (arrows). Scale bar = 10 µm." httpUri="https://zenodo.org/record/227400/files/figure.png" pageId="4" pageNumber="177">Figs. 2A</figureCitation>
, and
<figureCitation id="0FCCFDCFF21FA32A90F4FAB20FD349B7" box="[1214,1228,1978,2002]" captionStart="FIGURE 3. A, B" captionStartId="5.[151,255,1833,1856]" captionTargetBox="[155,1431,454,1809]" captionTargetId="figure@5.[150,1435,447,1809]" captionTargetPageId="5" captionText="FIGURE 3. A, B, Macrobiotus dimentmani. A, Claws of leg IV (slide No. 5359). It can been seen that the posterior and anterior claws are of different shapes, with only the posterior claws possessing the long, common basal tract (arrow). B, Detail of the egg (slide No. 5359). C, D, Macrobiotus diffusus. C, Claws of leg IV (slide No. 5527). As with M. dimentmani the claws are of different shapes, with only the posterior claws with a long, common basal tract (arrow). D, Detail of the egg (slide No. 3466). Arrows indicate some radial striae of the egg shell. Scale bar = 10 µm." httpUri="https://zenodo.org/record/227401/files/figure.png" pageId="4" pageNumber="177">3</figureCitation>
C); the secondary branches are clearly shorter (
<figureCitation id="0FCCFDCFF21EA32B9595FD9F09334ED5" box="[479,556,151,176]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1526,1549]" captionTargetBox="[156,1430,198,1500]" captionTargetId="figure@4.[151,1435,194,1504]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Macrobiotus insuetus sp. nov. A, B, Holotype claws of leg IV. In A the posterior claw is in focus (arrow), and B the anterior claw (arrow). The long, common basal tract for both posterior and anterior claws is very clear. C, D, Eggs (slide No. 5225). D shows the reticular ornamentation of the processes (arrows). Scale bar = 10 µm." httpUri="https://zenodo.org/record/227400/files/figure.png" pageId="5" pageNumber="178">Figs. 2</figureCitation>
A-B and 3C); the claws of legs I–III (particularly the external) have higher
<emphasis id="A5833D58F21EA32B91CCFD910E834ED5" box="[1414,1436,153,176]" italics="true" pageId="5" pageNumber="178">pt</emphasis>
index values (
<tableCitation id="DA75D4F1F21EA32B9578FDB40A974EB1" box="[306,392,188,213]" captionStart="TABLE 3" captionStartId="6.[151,239,152,175]" captionText="TABLE 3. Comparable measurements (in µm) and pt values of the holotype Macrobiotus insuetus sp. nov. with similar sized (where possible) specimens of: M. wuzhishanensis, M. diffusus and M. dimentmani." pageId="5" pageNumber="178">Table 3</tableCitation>
); leg IV posterior claws are slightly shorter than the external claws of legs II–III, while in
<taxonomicName id="50F79AC9F21EA32B9137FDB50BF14E9D" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="178" phylum="Tardigrada" rank="species" species="diffusus">
<emphasis id="A5833D58F21EA32B9137FDB50BF14E9D" italics="true" pageId="5" pageNumber="178">M. diffusus</emphasis>
</taxonomicName>
they are longer or of similar length (
<tableCitation id="DA75D4F1F21EA32B96DAFDD709F84E9D" box="[656,743,223,248]" captionStart="TABLE 3" captionStartId="6.[151,239,152,175]" captionText="TABLE 3. Comparable measurements (in µm) and pt values of the holotype Macrobiotus insuetus sp. nov. with similar sized (where possible) specimens of: M. wuzhishanensis, M. diffusus and M. dimentmani." pageId="5" pageNumber="178">Table 3</tableCitation>
). The eggs of
<taxonomicName id="50F79AC9F21EA32B97C1FDE90F124E92" box="[907,1037,224,248]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="178" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F21EA32B97C1FDE90F124E92" box="[907,1037,224,248]" italics="true" pageId="5" pageNumber="178">M. insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F21EA32B905EFDD70F714E92" box="[1044,1134,223,248]" pageId="5" pageNumber="178" rank="species">
<emphasis id="A5833D58F21EA32B905EFDD70F784E9D" bold="true" box="[1044,1127,223,248]" pageId="5" pageNumber="178">sp. nov</emphasis>
.
</taxonomicNameLabel>
differ from
<taxonomicName id="50F79AC9F21EA32B90B3FDE90E694E9D" box="[1273,1398,224,248]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="178" phylum="Tardigrada" rank="species" species="diffusus">
<emphasis id="A5833D58F21EA32B90B3FDE90E694E9D" box="[1273,1398,224,248]" italics="true" pageId="5" pageNumber="178">M. diffusus</emphasis>
</taxonomicName>
in: having fewer processes in both circumference and hemisphere (
<tableCitation id="DA75D4F1F21EA32B9720FC0C08DD4F79" box="[874,962,260,285]" captionStart="TABLE 2" captionStartId="2.[151,239,1110,1133]" captionTargetBox="[151,1437,1190,1762]" captionTargetPageId="2" captionText="TABLE 2. Comparable egg measurements (in µm) and number of egg processes for: Macrobiotus insuetus sp. nov., M. wuzhishanensis, M. diffusus and M. dimentmani." httpUri="http://table.plazi.org/id/C388B1C2F219A32C94DDF95E09834AE8" pageId="5" pageNumber="178" tableUuid="C388B1C2F219A32C94DDF95E09834AE8">Table 2</tableCitation>
), and processes terminal portions that are thin and flexible, not true filaments (
<figureCitation id="0FCCFDCFF21EA32B967AFC2F09AA4F5A" box="[560,693,295,320]" captionStart="FIGURE 2. A – D" captionStartId="4.[151,255,1526,1549]" captionTargetBox="[156,1430,198,1500]" captionTargetId="figure@4.[151,1435,194,1504]" captionTargetPageId="4" captionText="FIGURE 2. A – D, Macrobiotus insuetus sp. nov. A, B, Holotype claws of leg IV. In A the posterior claw is in focus (arrow), and B the anterior claw (arrow). The long, common basal tract for both posterior and anterior claws is very clear. C, D, Eggs (slide No. 5225). D shows the reticular ornamentation of the processes (arrows). Scale bar = 10 µm." httpUri="https://zenodo.org/record/227400/files/figure.png" pageId="5" pageNumber="178">Figs. 2C, D</figureCitation>
, and
<figureCitation id="0FCCFDCFF21EA32B96A5FC2009E24F25" box="[751,765,296,320]" captionStart="FIGURE 3. A, B" captionStartId="5.[151,255,1833,1856]" captionTargetBox="[155,1431,454,1809]" captionTargetId="figure@5.[150,1435,447,1809]" captionTargetPageId="5" captionText="FIGURE 3. A, B, Macrobiotus dimentmani. A, Claws of leg IV (slide No. 5359). It can been seen that the posterior and anterior claws are of different shapes, with only the posterior claws possessing the long, common basal tract (arrow). B, Detail of the egg (slide No. 5359). C, D, Macrobiotus diffusus. C, Claws of leg IV (slide No. 5527). As with M. dimentmani the claws are of different shapes, with only the posterior claws with a long, common basal tract (arrow). D, Detail of the egg (slide No. 3466). Arrows indicate some radial striae of the egg shell. Scale bar = 10 µm." httpUri="https://zenodo.org/record/227401/files/figure.png" pageId="5" pageNumber="178">3</figureCitation>
D). Although the egg shell characters of the new species are not clearly visible, it does appear similar to
<taxonomicName id="50F79AC9F21EA32B9637FC4509E64F01" box="[637,761,332,356]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="178" phylum="Tardigrada" rank="species" species="diffusus">
<emphasis id="A5833D58F21EA32B9637FC4509E64F01" box="[637,761,332,356]" italics="true" pageId="5" pageNumber="178">M. diffusus</emphasis>
</taxonomicName>
in lacking areolation and with few, small dots and some thin radial striae around the base of the processes.
</paragraph>
<caption id="C388B1C2F21EA32B94DDFA210FE349A5" httpUri="https://zenodo.org/record/227401/files/figure.png" pageId="5" pageNumber="178" targetBox="[155,1431,454,1809]" targetPageId="5">
<paragraph id="9748E14AF21EA32B94DDFA210FE349A5" blockId="5.[151,1437,1832,1984]" pageId="5" pageNumber="178">
<emphasis id="A5833D58F21EA32B94DDFA210A454925" bold="true" box="[151,346,1833,1856]" pageId="5" pageNumber="178">FIGURE 3. A, B,</emphasis>
<emphasis id="A5833D58F21EA32B952AFA20096C4925" box="[352,627,1832,1856]" italics="true" pageId="5" pageNumber="178">
<taxonomicName id="50F79AC9F21EA32B952AFA2009704925" box="[352,623,1832,1856]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="178" phylum="Tardigrada" rank="species" species="dimentmani">Macrobiotus dimentmani</taxonomicName>
.
</emphasis>
<emphasis id="A5833D58F21EA32B9630FA21098B4925" bold="true" box="[634,660,1833,1856]" pageId="5" pageNumber="178">A,</emphasis>
Claws of leg IV (slide No. 5359). It can been seen that the posterior and anterior claws are of different shapes, with only the posterior claws possessing the long, common basal tract (arrow).
<emphasis id="A5833D58F21EA32B91CEFA410E834905" bold="true" box="[1412,1436,1865,1888]" pageId="5" pageNumber="178">B,</emphasis>
Detail of the egg (slide No.5359).
<emphasis id="A5833D58F21EA32B9669FA61097D49E5" bold="true" box="[547,610,1897,1920]" pageId="5" pageNumber="178">C, D,</emphasis>
<emphasis id="A5833D58F21EA32B9624FA60084449E5" box="[622,859,1896,1920]" italics="true" pageId="5" pageNumber="178">
<taxonomicName id="50F79AC9F21EA32B9624FA60084749E5" box="[622,856,1896,1920]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="178" phylum="Tardigrada" rank="species" species="diffusus">Macrobiotus diffusus</taxonomicName>
.
</emphasis>
<emphasis id="A5833D58F21EA32B972CFA61089E49E5" bold="true" box="[870,897,1897,1920]" pageId="5" pageNumber="178">C,</emphasis>
Claws of leg IV (slide No. 5527). As with
<taxonomicName id="50F79AC9F21EA32B9135FA610A0A49C5" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="5" pageNumber="178" phylum="Tardigrada" rank="species" species="dimentmani">
<emphasis id="A5833D58F21EA32B9135FA610A0A49C5" italics="true" pageId="5" pageNumber="178">M. dimentmani</emphasis>
</taxonomicName>
the claws are of different shapes, with only the posterior claws with a long, common basal tract (arrow).
<emphasis id="A5833D58F21EA32B91C8FA810E8349C5" bold="true" box="[1410,1436,1929,1952]" pageId="5" pageNumber="178">D,</emphasis>
Detail of the egg (slide No. 3466). Arrows indicate some radial striae of the egg shell. Scale bar = 10 µm.
</paragraph>
</caption>
<caption id="C388B1C2F21DA32894DDFD900F4F4EB5" pageId="6" pageNumber="179">
<paragraph id="9748E14AF21DA32894DDFD900F4F4EB5" blockId="6.[151,1436,151,964]" pageId="6" pageNumber="179">
<emphasis id="A5833D58F21DA32894DDFD900A154ECA" bold="true" box="[151,266,152,175]" pageId="6" pageNumber="179">TABLE 3.</emphasis>
Comparable measurements (in µm) and
<emphasis id="A5833D58F21DA328968BFD9109C94ED5" box="[705,726,153,176]" italics="true" pageId="6" pageNumber="179">pt</emphasis>
values of the holotype
<taxonomicName id="50F79AC9F21DA3289798FD9F0FA84ECA" box="[978,1207,151,175]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="179" phylum="Tardigrada" rank="species" species="insuetus" status="sp. nov.">
<emphasis id="A5833D58F21DA3289798FD9F0FA84ECA" box="[978,1207,151,175]" italics="true" pageId="6" pageNumber="179">Macrobiotus insuetus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BEB08023F21DA32890F4FD900E0A4ECA" box="[1214,1301,152,175]" pageId="6" pageNumber="179" rank="species">
<emphasis id="A5833D58F21DA32890F4FD900E0A4ECA" bold="true" box="[1214,1301,152,175]" pageId="6" pageNumber="179">sp. nov.</emphasis>
</taxonomicNameLabel>
with similar sized (where possible) specimens of:
<emphasis id="A5833D58F21DA328966CFDB1086C4EB5" box="[550,883,184,208]" italics="true" pageId="6" pageNumber="179">
<taxonomicName id="50F79AC9F21DA328966CFDB109ED4EB5" box="[550,754,184,208]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="179" phylum="Tardigrada" rank="species" species="wuzhishanensis">M. wuzhishanensis</taxonomicName>
,
<taxonomicName id="50F79AC9F21DA32896B7FDB1086C4EB5" box="[765,883,184,208]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="179" phylum="Tardigrada" rank="species" species="diffusus">M. diffusus</taxonomicName>
</emphasis>
and
<taxonomicName id="50F79AC9F21DA32897EDFDB10F544EB5" box="[935,1099,184,208]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="179" phylum="Tardigrada" rank="species" species="dimentmani">M. dimentmani</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="9748E14AF21DA328959CFDE20FD44E9A" blockId="6.[151,1436,151,964]" box="[470,1227,233,255]" pageId="6" pageNumber="179">
<emphasis id="A5833D58F21DA328959CFDE20FD44E9A" box="[470,1227,233,255]" italics="true" pageId="6" pageNumber="179">
<taxonomicName id="50F79AC9F21DA328959CFDE209544E9A" box="[470,587,234,255]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="179" phylum="Tardigrada" rank="species" species="insuetus">M. insuetus</taxonomicName>
<taxonomicName id="50F79AC9F21DA32896C5FDE108544E9A" box="[655,843,233,255]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="179" phylum="Tardigrada" rank="species" species="wuzhishanensis">M.wuzhishanensis</taxonomicName>
*
<taxonomicName id="50F79AC9F21DA3289730FDE208F54E9A" box="[890,1002,233,255]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="179" phylum="Tardigrada" rank="species" species="diffusus">M. diffusus</taxonomicName>
<taxonomicName id="50F79AC9F21DA3289078FDE20FD44E9A" box="[1074,1227,233,255]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="6" pageNumber="179" phylum="Tardigrada" rank="species" species="dimentmani">M. dimentmani</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="9748E14AF21DA328959CFC0009384F7B" blockId="6.[151,1436,151,964]" box="[470,551,264,286]" pageId="6" pageNumber="179">
<emphasis id="A5833D58F21DA328959CFC0009384F7B" bold="true" box="[470,551,264,286]" pageId="6" pageNumber="179">sp. nov.</emphasis>
</paragraph>
<paragraph id="9748E14AF21DA32894D6FC240E014F27" blockId="6.[151,1436,151,964]" box="[156,1310,300,322]" pageId="6" pageNumber="179">Slide No. 5520 3471 5359 5359</paragraph>
<paragraph id="9748E14AF21DA328959CFC420E5C4F05" blockId="6.[151,1436,151,964]" box="[470,1347,330,352]" pageId="6" pageNumber="179">
<typeStatus id="484C5FE8F21DA328959CFC4208C74F05" box="[470,984,330,352]" pageId="6" pageNumber="179" type="holotype">Holotype Holotype Holotype</typeStatus>
<typeStatus id="484C5FE8F21DA3289078FC430E5C4F05" box="[1074,1347,331,352]" pageId="6" pageNumber="179" type="paratype">Paratype Paratype</typeStatus>
</paragraph>
<paragraph id="9748E14AF21DA328959CFC670E0E4FCD" blockId="6.[151,1436,151,964]" pageId="6" pageNumber="179">
µm (
<emphasis id="A5833D58F21DA3289671FC7809514FE0" box="[571,590,368,389]" italics="true" pageId="6" pageNumber="179">pt</emphasis>
) µm (
<emphasis id="A5833D58F21DA3289747FC78083E4FE0" box="[781,801,368,389]" italics="true" pageId="6" pageNumber="179">pt</emphasis>
) µm (
<emphasis id="A5833D58F21DA328979CFC7808F54FE0" box="[982,1002,368,389]" italics="true" pageId="6" pageNumber="179">pt</emphasis>
) µm (
<emphasis id="A5833D58F21DA32890C5FC780FBC4FE0" box="[1167,1187,368,389]" italics="true" pageId="6" pageNumber="179">pt</emphasis>
) µm (
<emphasis id="A5833D58F21DA328917DFC780E554FE0" box="[1335,1354,368,389]" italics="true" pageId="6" pageNumber="179">pt</emphasis>
) Body length 334 420 303 333 333
</paragraph>
<paragraph id="9748E14AF21DA32894D6FCBE0E084FA9" blockId="6.[151,1436,151,964]" box="[156,1303,438,460]" pageId="6" pageNumber="179">Buccal tube length 35.8 44.2 35.8 39.0 39.9</paragraph>
<paragraph id="9748E14AF21DA32894D6FCD20E634DA1" blockId="6.[151,1436,151,964]" pageId="6" pageNumber="179">
External buccal tube width 6.5 (
<emphasis id="A5833D58F21DA3289679FCD209784F95" box="[563,615,474,496]" italics="true" pageId="6" pageNumber="179">18.2)</emphasis>
5.3 (
<emphasis id="A5833D58F21DA3289747FCD208264F95" box="[781,825,474,496]" italics="true" pageId="6" pageNumber="179">12.0</emphasis>
) 7.6 (
<emphasis id="A5833D58F21DA328979CFCD20F1D4F95" box="[982,1026,474,496]" italics="true" pageId="6" pageNumber="179">21.2</emphasis>
) 8.0 (
<emphasis id="A5833D58F21DA3289034FCD20FB54F95" box="[1150,1194,474,496]" italics="true" pageId="6" pageNumber="179">20.5</emphasis>
) 8.1 (
<emphasis id="A5833D58F21DA328917DFCD20E7C4F95" box="[1335,1379,474,496]" italics="true" pageId="6" pageNumber="179">20.3</emphasis>
) Stylet supports insertion point (
<emphasis id="A5833D58F21DA3289679FCF609404C71" box="[563,607,510,532]" italics="true" pageId="6" pageNumber="179">79.2</emphasis>
) (
<emphasis id="A5833D58F21DA3289747FCF608264C71" box="[781,825,510,532]" italics="true" pageId="6" pageNumber="179">78.7</emphasis>
) (
<emphasis id="A5833D58F21DA328979CFCF60F1D4C71" box="[982,1026,510,532]" italics="true" pageId="6" pageNumber="179">78.3</emphasis>
) (
<emphasis id="A5833D58F21DA32890C5FCF70FA44C71" box="[1167,1211,511,532]" italics="true" pageId="6" pageNumber="179">77.5</emphasis>
)
<emphasis id="A5833D58F21DA3289175FCF60E6C4C71" box="[1343,1395,510,532]" italics="true" pageId="6" pageNumber="179">(77.0</emphasis>
) First macroplacoid 6.0 (
<emphasis id="A5833D58F21DA3289679FF2A09404C5D" box="[563,607,546,568]" italics="true" pageId="6" pageNumber="179">16.8</emphasis>
) 5.5 (
<emphasis id="A5833D58F21DA3289747FF2A08264C5D" box="[781,825,546,568]" italics="true" pageId="6" pageNumber="179">12.4</emphasis>
) 5.1 (
<emphasis id="A5833D58F21DA328979CFF2A0F1D4C5D" box="[982,1026,546,568]" italics="true" pageId="6" pageNumber="179">14.3</emphasis>
) 5.5 (
<emphasis id="A5833D58F21DA32890C5FF2A0FA44C5D" box="[1167,1211,546,568]" italics="true" pageId="6" pageNumber="179">14.1</emphasis>
) 5.6 (
<emphasis id="A5833D58F21DA328910DFF2A0E6C4C5D" box="[1351,1395,546,568]" italics="true" pageId="6" pageNumber="179">14.0</emphasis>
) Second macroplacoid 4.4 (
<emphasis id="A5833D58F21DA3289679FF4E09404C39" box="[563,607,582,604]" italics="true" pageId="6" pageNumber="179">12.3</emphasis>
) 4.4 (
<emphasis id="A5833D58F21DA3289747FF4E08264C39" box="[781,825,582,604]" italics="true" pageId="6" pageNumber="179">10.0</emphasis>
) 4.9 (
<emphasis id="A5833D58F21DA328979CFF4E0F1D4C39" box="[982,1026,582,604]" italics="true" pageId="6" pageNumber="179">13.7</emphasis>
) 5.0 (
<emphasis id="A5833D58F21DA32890C5FF4E0FA44C39" box="[1167,1211,582,604]" italics="true" pageId="6" pageNumber="179">12.8</emphasis>
) 5.3 (
<emphasis id="A5833D58F21DA328910DFF4E0E6C4C39" box="[1351,1395,582,604]" italics="true" pageId="6" pageNumber="179">13.3</emphasis>
) Third macroplacoid 5.4 (
<emphasis id="A5833D58F21DA3289679FF6209404CE5" box="[563,607,618,640]" italics="true" pageId="6" pageNumber="179">15.1</emphasis>
) 5.7 (
<emphasis id="A5833D58F21DA3289747FF6208264CE5" box="[781,825,618,640]" italics="true" pageId="6" pageNumber="179">12.9</emphasis>
) 5.5 (
<emphasis id="A5833D58F21DA328979CFF620F1D4CE5" box="[982,1026,618,640]" italics="true" pageId="6" pageNumber="179">15.4</emphasis>
) 5.1 (
<emphasis id="A5833D58F21DA32890C5FF620FA44CE5" box="[1167,1211,618,640]" italics="true" pageId="6" pageNumber="179">13.1</emphasis>
) 6.0 (
<emphasis id="A5833D58F21DA328910DFF620E6C4CE5" box="[1351,1395,618,640]" italics="true" pageId="6" pageNumber="179">15.0</emphasis>
) Microplacoid 4.2 (
<emphasis id="A5833D58F21DA3289679FF8609434CC1" box="[563,604,654,676]" italics="true" pageId="6" pageNumber="179">11.7</emphasis>
) 4.9 (
<emphasis id="A5833D58F21DA3289747FF8608294CC1" box="[781,822,654,677]" italics="true" pageId="6" pageNumber="179">11. 1</emphasis>
) 4.0 (
<emphasis id="A5833D58F21DA328979CFF8608E04CC1" box="[982,1023,654,676]" italics="true" pageId="6" pageNumber="179">11.2</emphasis>
) 4.5 (
<emphasis id="A5833D58F21DA32890C5FF860FA84CC1" box="[1167,1207,654,677]" italics="true" pageId="6" pageNumber="179">11.5</emphasis>
) 5.0 (
<emphasis id="A5833D58F21DA328910DFF860E6C4CC1" box="[1351,1395,654,676]" italics="true" pageId="6" pageNumber="179">12.5</emphasis>
) Placoid row 21.5 (
<emphasis id="A5833D58F21DA3289679FFBA09404CAD" box="[563,607,690,712]" italics="true" pageId="6" pageNumber="179">60.1</emphasis>
)?? 21.2 (
<emphasis id="A5833D58F21DA328979CFFBA0F1D4CAD" box="[982,1026,690,712]" italics="true" pageId="6" pageNumber="179">59.2</emphasis>
) 23.1 (
<emphasis id="A5833D58F21DA32890C5FFBA0FA44CAD" box="[1167,1211,690,712]" italics="true" pageId="6" pageNumber="179">59.2</emphasis>
) 22.9 (
<emphasis id="A5833D58F21DA328910DFFBA0E6C4CAD" box="[1351,1395,690,712]" italics="true" pageId="6" pageNumber="179">57.4</emphasis>
) Macroplacoid row 17.5 (
<emphasis id="A5833D58F21DA3289679FFDE09404C89" box="[563,607,726,748]" italics="true" pageId="6" pageNumber="179">48.9</emphasis>
) 18.2 (
<emphasis id="A5833D58F21DA3289747FFDE08264C89" box="[781,825,726,748]" italics="true" pageId="6" pageNumber="179">41.2</emphasis>
) 16.4 (
<emphasis id="A5833D58F21DA328979CFFDE0F1D4C89" box="[982,1026,726,748]" italics="true" pageId="6" pageNumber="179">45.8</emphasis>
) 18.2 (
<emphasis id="A5833D58F21DA32890C5FFDE0FA44C89" box="[1167,1211,726,748]" italics="true" pageId="6" pageNumber="179">46.7</emphasis>
) 18.9 (
<emphasis id="A5833D58F21DA328910DFFDE0E6C4C89" box="[1351,1395,726,748]" italics="true" pageId="6" pageNumber="179">47.4</emphasis>
) External claw II 12.9 (
<emphasis id="A5833D58F21DA3289679FFF209404D75" box="[563,607,762,784]" italics="true" pageId="6" pageNumber="179">36.0</emphasis>
) 13.0 (
<emphasis id="A5833D58F21DA3289747FFF208264D75" box="[781,825,762,784]" italics="true" pageId="6" pageNumber="179">29.4</emphasis>
) 9.9 (
<emphasis id="A5833D58F21DA328979CFFF20F1D4D75" box="[982,1026,762,784]" italics="true" pageId="6" pageNumber="179">27.7</emphasis>
) 14.5 (
<emphasis id="A5833D58F21DA32890C5FFF20FA44D75" box="[1167,1211,762,784]" italics="true" pageId="6" pageNumber="179">37.2</emphasis>
) 14.5 (
<emphasis id="A5833D58F21DA328910DFFF20E6C4D75" box="[1351,1395,762,784]" italics="true" pageId="6" pageNumber="179">36.3</emphasis>
) Internal claw II 10.2 (
<emphasis id="A5833D58F21DA3289679FE1609404D51" box="[563,607,798,820]" italics="true" pageId="6" pageNumber="179">28.5</emphasis>
) 9.1 (
<emphasis id="A5833D58F21DA3289747FE1608264D51" box="[781,825,798,820]" italics="true" pageId="6" pageNumber="179">20.6</emphasis>
) 9.1 (
<emphasis id="A5833D58F21DA328979CFE160F1D4D51" box="[982,1026,798,820]" italics="true" pageId="6" pageNumber="179">25.4</emphasis>
) 10.9 (
<emphasis id="A5833D58F21DA32890C5FE160FA44D51" box="[1167,1211,798,820]" italics="true" pageId="6" pageNumber="179">27.9</emphasis>
) 11.4 (
<emphasis id="A5833D58F21DA328910DFE160E6C4D51" box="[1351,1395,798,820]" italics="true" pageId="6" pageNumber="179">28.6</emphasis>
) External claw III 12.8 (
<emphasis id="A5833D58F21DA3289679FE4A09404D3D" box="[563,607,834,856]" italics="true" pageId="6" pageNumber="179">35.8</emphasis>
) 13.0 (
<emphasis id="A5833D58F21DA3289747FE4A08264D3D" box="[781,825,834,856]" italics="true" pageId="6" pageNumber="179">29.4</emphasis>
) 9.9 (
<emphasis id="A5833D58F21DA328979CFE4A0F1D4D3D" box="[982,1026,834,856]" italics="true" pageId="6" pageNumber="179">27.7</emphasis>
) 14.5 (
<emphasis id="A5833D58F21DA32890C5FE4A0FA44D3D" box="[1167,1211,834,856]" italics="true" pageId="6" pageNumber="179">37.2</emphasis>
) 14.5 (
<emphasis id="A5833D58F21DA328910DFE4A0E6C4D3D" box="[1351,1395,834,856]" italics="true" pageId="6" pageNumber="179">36.3</emphasis>
) Internal claw III 10.2 (
<emphasis id="A5833D58F21DA3289679FE6E09404D19" box="[563,607,870,892]" italics="true" pageId="6" pageNumber="179">28.5</emphasis>
) 9.1 (
<emphasis id="A5833D58F21DA3289747FE6E08264D19" box="[781,825,870,892]" italics="true" pageId="6" pageNumber="179">20.6</emphasis>
) 9.1 (
<emphasis id="A5833D58F21DA328979CFE6E0F1D4D19" box="[982,1026,870,892]" italics="true" pageId="6" pageNumber="179">25.4</emphasis>
) 10.9
<emphasis id="A5833D58F21DA32890CCFE6E0FA44D19" box="[1158,1211,870,892]" italics="true" pageId="6" pageNumber="179">(27.9</emphasis>
) 11.4 (28.6) Posterior claw IV 11.9 (
<emphasis id="A5833D58F21DA3289679FE8209404DC5" box="[563,607,906,928]" italics="true" pageId="6" pageNumber="179">33.2</emphasis>
) 13.0 (
<emphasis id="A5833D58F21DA3289747FE8208264DC5" box="[781,825,906,928]" italics="true" pageId="6" pageNumber="179">29.4</emphasis>
) 11.2 (
<emphasis id="A5833D58F21DA328979CFE820F1D4DC5" box="[982,1026,906,928]" italics="true" pageId="6" pageNumber="179">31.3</emphasis>
) 13.7 (
<emphasis id="A5833D58F21DA32890C5FE820FA44DC5" box="[1167,1211,906,928]" italics="true" pageId="6" pageNumber="179">35.1</emphasis>
) 14.0 (
<emphasis id="A5833D58F21DA328910DFE820E6C4DC5" box="[1351,1395,906,928]" italics="true" pageId="6" pageNumber="179">35.1</emphasis>
) Anterior claw IV 10.9 (
<emphasis id="A5833D58F21DA3289679FEA609404DA1" box="[563,607,942,964]" italics="true" pageId="6" pageNumber="179">30.4</emphasis>
) 10.4 (
<emphasis id="A5833D58F21DA3289747FEA608264DA1" box="[781,825,942,964]" italics="true" pageId="6" pageNumber="179">23.5</emphasis>
) 10.0 (
<emphasis id="A5833D58F21DA328979CFEA60F1D4DA1" box="[982,1026,942,964]" italics="true" pageId="6" pageNumber="179">27.9</emphasis>
) 13.2 (
<emphasis id="A5833D58F21DA32890C5FEA60FA44DA1" box="[1167,1211,942,964]" italics="true" pageId="6" pageNumber="179">33.8</emphasis>
) 13.5 (
<emphasis id="A5833D58F21DA328910DFEA60E6C4DA1" box="[1351,1395,942,964]" italics="true" pageId="6" pageNumber="179">33.8</emphasis>
)
</paragraph>
<paragraph id="9748E14AF21DA32894DDFEEA0AE24D9C" blockId="6.[151,509,993,1017]" box="[151,509,993,1017]" pageId="6" pageNumber="179">
*) According to
<bibRefCitation id="F3669CBBF21DA328950CFEEA0AE64D9C" box="[326,505,993,1017]" pageId="6" pageNumber="179" refString="Yin, H., Wang, L. &amp; Li, X. (2011) Two new species of the genus Macrobiotus (Tardigrada: Macrobiotidae) from China. Proceedings of the Biological Society of Washington, 124, 3, 165 - 178." type="journal article">
Yin
<emphasis id="A5833D58F21DA3289539FEEA0ABE4D9C" box="[371,417,993,1017]" italics="true" pageId="6" pageNumber="179">et al</emphasis>
. (2011)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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