treatments-xml/data/03/CD/BE/03CDBE749327B502CE38FA94FEE9F813.xml

<document id="4C48E58138A0F1C2D41264051997531A" ID-DOI="10.5252/zoosystema2023v45a9" ID-ISSN="1638-9387" ID-Zenodo-Dep="8071253" ID-ZooBank="urn:lsid:zoobank.org:pub:69C34731-8C25-4A1E-B336-B222CD3CBAC3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1686747199180" checkinUser="felipe" docAuthor="Guinot, Danièle" docDate="2023" docId="03CDBE749327B502CE38FA94FEE9F813" docLanguage="en" docName="Zoosystema.45.9.225-372.pdf" docOrigin="Zoosystema 45 (9)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.5:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="5" docTitle="Dorippe trilobata : Manning 1993" docType="treatment" docVersion="2" lastPageNumber="261" masterDocId="FFF4C60C9304B524CD15FFC0FF9AFFF8" masterDocTitle="A new subfamily classification of the highly diversified Dorippidae H. Milne Edwards, 1837 (Crustacea, Decapoda, Brachyura, Dorippoidea), using morphological, molecular and palaeotonlogical data, with special emphasis on its unique female reproductive system" masterLastPageNumber="372" masterPageNumber="225" pageNumber="258" updateTime="1687453794845" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="598F0FC4D9A1CF7ED43B06A4EEEC7CF9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="29A1142861D5ABBA0884F38945091710">
<mods:title id="6F6ADEE3B96A35E200169D485E64F29F">A new subfamily classification of the highly diversified Dorippidae H. Milne Edwards, 1837 (Crustacea, Decapoda, Brachyura, Dorippoidea), using morphological, molecular and palaeotonlogical data, with special emphasis on its unique female reproductive system</mods:title>
</mods:titleInfo>
<mods:name id="7D686BCA13B202A7208DCA56B9650E9B" type="personal">
<mods:role id="CDF497EA78486563D64228E1460DA424">
<mods:roleTerm id="9BE57A052825830DAEDE21FBEBDE5349">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3E89AC9904E5027BF078175E4A951565">Guinot, Danièle</mods:namePart>
<mods:affiliation id="8C5DDF62CAFFBAE893A69656AE6F2BB8">Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national d’Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) daniele. guinot @ mnhn. fr Dedicated to the memory of my colleague and dearest friend Ngan Kee NG (1966 - 2022)</mods:affiliation>
<mods:nameIdentifier id="E0081980A1B826F523391196C6F8BA12" type="email">guinot@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="7B7050A53E022FA3E88F630FCDAE1EB0">text</mods:typeOfResource>
<mods:relatedItem id="ECFA5CF997A51804F7B4ADFE6FFC6196" type="host">
<mods:titleInfo id="CB45DC9EFA51EF9322BF38CFFE3C4718">
<mods:title id="93F94D040E7FF67498237D9517DDAA50">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="BA707B665043CC17F3DAFF1E4ADAC5B6">
<mods:date id="FB5C712B4C6E15EDD7B067FEE30FF921">2023</mods:date>
<mods:detail id="DAA42C11E6A1893FB30817FD74266DCD" type="pubDate">
<mods:number id="C929C545DD3DE8453EE38BD77F2AB252">2023-06-05</mods:number>
</mods:detail>
<mods:detail id="D02325D8BA41900E5705FA34F2F60987" type="volume">
<mods:number id="C8480A3E6288A9EA4589183E2D00B0CA">45</mods:number>
</mods:detail>
<mods:detail id="12F6E1B47F04F04C4A2C37508E5BD333" type="issue">
<mods:number id="83FF09CA0DCE3C65232DD48AAECA3895">9</mods:number>
</mods:detail>
<mods:extent id="AAE7016A833FBEEFA18D425AB64A0060" unit="page">
<mods:start id="F2A8720F0566DBE4DF064B3A6B83E1C9">225</mods:start>
<mods:end id="1F41591476821E1680058C3B13E17BC3">372</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="88B92E4235619CE88878CD17586438B3">journal article</mods:classification>
<mods:identifier id="1350A012B4C77B9CAE56CADE00582AF0" type="DOI">10.5252/zoosystema2023v45a9</mods:identifier>
<mods:identifier id="4D0361384D41DEF8C10CC18A79861062" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="48A98E7AF700D335B14D7FFBD3945CF1" type="Zenodo-Dep">8071253</mods:identifier>
<mods:identifier id="C73E66EAEBA3217300F3F92A0AA2C480" type="ZooBank">urn:lsid:zoobank.org:pub:69C34731-8C25-4A1E-B336-B222CD3CBAC3</mods:identifier>
</mods:mods>
<treatment id="03CDBE749327B502CE38FA94FEE9F813" LSID="urn:lsid:plazi:treatment:03CDBE749327B502CE38FA94FEE9F813" httpUri="http://treatment.plazi.org/id/03CDBE749327B502CE38FA94FEE9F813" lastPageId="38" lastPageNumber="261" pageId="35" pageNumber="258">
<subSubSection id="C37E5CE99327B507CE38FA94FB50FA75" pageId="35" pageNumber="258" type="nomenclature">
<paragraph id="8BDB0F629327B507CE38FA94FB50FA75" blockId="35.[811,1456,1363,2028]" pageId="35" pageNumber="258">
<smallCapsWord id="8D3D99BE9327B507CE38FA94FC0BFA94" baselines="1383,1383" box="[813,913,1364,1390]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="35" pageNumber="258">REMARKS</smallCapsWord>
<smallCapsWord id="8D3D99BE9327B507CE8DFA99FC23FA94" baselines="1383" box="[920,953,1369,1388]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="on" pageId="35" pageNumber="258">ON</smallCapsWord>
<taxonomicName id="4C6474E19327B507CED5FA94FAD1FA95" ID-CoL="6DGLV" authority="MANNING, 1993" authorityName=": Manning" authorityYear="1993" box="[960,1355,1363,1390]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="258" phylum="Arthropoda" rank="species" species="trilobata">
<emphasis id="B910D3709327B507CED5FA94FB0AFA94" box="[960,1168,1364,1389]" italics="true" pageId="35" pageNumber="258">
<smallCapsWord id="8D3D99BE9327B507CED5FA94FB81FA94" baselines="1383,1383" box="[960,1051,1364,1389]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Dorippe" pageId="35" pageNumber="258">DORIPPE</smallCapsWord>
<smallCapsWord id="8D3D99BE9327B507C937FA99FB0AFA94" baselines="1383" box="[1058,1168,1369,1388]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="trilobata" pageId="35" pageNumber="258">TRILOBATA</smallCapsWord>
</emphasis>
<bibRefCitation id="EFF572939327B507C983FA94FAD1FA95" author="MANNING" box="[1174,1355,1363,1390]" firstAuthor="MANNING" pageId="35" pageNumber="258" pagination="1 - 4" refId="ref131161" refString="MANNING R. B. 1993. - Two new dorippid crabs from Australia (Crustacea: Decapoda: Dorippidae). Records from the Australian Museum 45: 1 - 4. https: // doi. org / 10.3853 / j. 0067 - 1975.45.1993.124" type="journal article" year="1993">
<smallCapsWord id="8D3D99BE9327B507C983FA94FA9FFA94" baselines="1383,1383" box="[1174,1285,1364,1390]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Manning" pageId="35" pageNumber="258">MANNING</smallCapsWord>
, 1993
</bibRefCitation>
</taxonomicName>
<smallCapsWord id="8D3D99BE9327B507CE38FAB9FCC1FA74" baselines="1415" box="[813,859,1401,1420]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="and" pageId="35" pageNumber="258">AND</smallCapsWord>
<taxonomicName id="4C6474E19327B507CE77FAB4FB50FA75" authority="MANNING, 1993" authorityName=": Manning" authorityYear="1993" box="[866,1226,1395,1422]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="258" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="B910D3709327B507CE77FAB4FB95FA74" box="[866,1039,1396,1421]" italics="true" pageId="35" pageNumber="258">
<smallCapsWord id="8D3D99BE9327B507CE77FAB4FC26FA74" baselines="1415,1415" box="[866,956,1396,1421]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Dorippe" pageId="35" pageNumber="258">DORIPPE</smallCapsWord>
<smallCapsWord id="8D3D99BE9327B507CED6FAB9FB95FA74" baselines="1415" box="[963,1039,1401,1420]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="glabra" pageId="35" pageNumber="258">GLABRA</smallCapsWord>
</emphasis>
<bibRefCitation id="EFF572939327B507C900FAB4FB50FA75" author="MANNING" box="[1045,1226,1395,1422]" firstAuthor="MANNING" pageId="35" pageNumber="258" pagination="1 - 4" refId="ref131161" refString="MANNING R. B. 1993. - Two new dorippid crabs from Australia (Crustacea: Decapoda: Dorippidae). Records from the Australian Museum 45: 1 - 4. https: // doi. org / 10.3853 / j. 0067 - 1975.45.1993.124" type="journal article" year="1993">
<smallCapsWord id="8D3D99BE9327B507C900FAB4FB1EFA74" baselines="1415,1415" box="[1045,1156,1396,1422]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Manning" pageId="35" pageNumber="258">MANNING</smallCapsWord>
, 1993
</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C37E5CE99327B502CE38FA54FEE9F813" lastPageId="38" lastPageNumber="261" pageId="35" pageNumber="258" type="discussion">
<paragraph id="8BDB0F629327B507CE38FA54FBE7F914" blockId="35.[811,1456,1363,2028]" pageId="35" pageNumber="258">
Two 
<emphasis id="B910D3709327B507CE71FA54FC34FA55" box="[868,942,1428,1453]" italics="true" pageId="35" pageNumber="258">Doripp</emphasis>
e from 
<collectingCountry id="F3734FF29327B507C914FA53FBFAFA55" box="[1025,1120,1427,1453]" name="Australia" pageId="35" pageNumber="258">Australia</collectingCountry>
described by 
<bibRefCitation id="EFF572939327B507C9EFFA53FA32FA55" author="Manning" box="[1274,1448,1427,1453]" firstAuthor="Manning" pageId="35" pageNumber="258" pagination="1 - 4" refId="ref131161" refString="MANNING R. B. 1993. - Two new dorippid crabs from Australia (Crustacea: Decapoda: Dorippidae). Records from the Australian Museum 45: 1 - 4. https: // doi. org / 10.3853 / j. 0067 - 1975.45.1993.124" type="journal article" year="1993">Manning (1993</bibRefCitation>
; see 
<bibRefCitation id="EFF572939327B507CE47FA73FB93FA36" author="DAVIE P. J. F." box="[850,1033,1459,1486]" pageId="35" pageNumber="258" refId="ref114947" refString="DAVIE P. J. F. 2002. - Crustacea: Malacostraca. ‰ ucarida (Part 2). Decapoda Anomura, Brachyura, in WELLS A. &amp; HOUSTON W. W. K. (eds), Zological Catalogue of Australia. Vol. 19.3 B. CSIR ̿ Publishing, Melbourne, xiv + 641 p." type="book" year="2002">Davie 2002: 156</bibRefCitation>
), each known only by its 
<typeStatus id="54DFB1C09327B507C803FA73FADEFA35" box="[1302,1348,1459,1485]" pageId="35" pageNumber="258">type</typeStatus>
specimen and thus by a single sex, and never reported since, are problematic. The characters used to distinguish them from other known species are tenuous and mainly concern the more or less marked ornamentation and especially the setation and size of the legs. Previously, only two dorippids were known from 
<collectingCountry id="F3734FF29327B507CE39F9B2FC15F974" box="[812,911,1650,1676]" name="Australia" pageId="35" pageNumber="258">Australia</collectingCountry>
: 
<taxonomicName id="4C6474E19327B507CE89F9B3FBF6F974" authorityName=": Holthuis &amp; Manning" authorityYear="1990" box="[924,1132,1650,1676]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="258" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B910D3709327B507CE89F9B3FBF6F974" box="[924,1132,1650,1676]" italics="true" pageId="35" pageNumber="258">Dorippe quadridens</emphasis>
</taxonomicName>
and 
<emphasis id="B910D3709327B507C9B3F9B2FA2AF974" box="[1190,1456,1650,1676]" italics="true" pageId="35" pageNumber="258">
<taxonomicName id="4C6474E19327B507C9B3F9B2FA36F974" baseAuthorityName="Miers" baseAuthorityYear="1884" box="[1190,1452,1650,1676]" class="Malacostraca" family="Dorippidae" genus="Paradorippe" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="258" phylum="Arthropoda" rank="species" species="australiensis">Paradorippe australiensis</taxonomicName>
.
</emphasis>
Thanks to photographs of the 
<typeStatus id="54DFB1C09327B507C962F953FB34F955" box="[1143,1198,1683,1709]" pageId="35" pageNumber="258">types</typeStatus>
of each of these species deposited in the 
<collectionCode id="ED7597A79327B507CEFDF972FB8CF934" box="[1000,1046,1714,1740]" pageId="35" pageNumber="258">AM</collectionCode>
, kindly sent to us by Shane Ahyong, some remarks can be provided.
</paragraph>
<paragraph id="8BDB0F629327B502CE56F933FE4CFB76" blockId="35.[811,1456,1363,2028]" lastBlockId="38.[130,777,215,2028]" lastPageId="38" lastPageNumber="261" pageId="35" pageNumber="258">
<taxonomicName id="4C6474E19327B507CE56F933FC6BF8F4" authorityName=": Manning" authorityYear="1993" box="[835,1009,1778,1804]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="258" phylum="Arthropoda" rank="species" species="trilobata">
<emphasis id="B910D3709327B507CE56F933FC6BF8F4" box="[835,1009,1778,1804]" italics="true" pageId="35" pageNumber="258">Dorippe trilobata</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFF572939327B507CEEAF932FB2DF8F4" author="Manning" box="[1023,1207,1778,1804]" firstAuthor="Manning" page="3" pageId="35" pageNumber="258" pagination="1 - 4" refId="ref131161" refString="MANNING R. B. 1993. - Two new dorippid crabs from Australia (Crustacea: Decapoda: Dorippidae). Records from the Australian Museum 45: 1 - 4. https: // doi. org / 10.3853 / j. 0067 - 1975.45.1993.124" type="journal article" year="1993">Manning 1993: 3</bibRefCitation>
, 4, fig. 4; cited by 
<bibRefCitation id="EFF572939327B507C864F932FC00F8D4" author="DAVIE P. J. F." pageId="35" pageNumber="258" refId="ref114947" refString="DAVIE P. J. F. 2002. - Crustacea: Malacostraca. ‰ ucarida (Part 2). Decapoda Anomura, Brachyura, in WELLS A. &amp; HOUSTON W. W. K. (eds), Zological Catalogue of Australia. Vol. 19.3 B. CSIR ̿ Publishing, Melbourne, xiv + 641 p." type="book" year="2002">Davie 2002: 156</bibRefCitation>
), described from Western Autralia (Admiralty 
<collectingRegion id="49A0C1809327B507C866F8D2FA3EF8D4" box="[1395,1444,1810,1836]" country="Papua New Guinea" name="Gulf" pageId="35" pageNumber="258">Gulf</collectingRegion>
) on the basis of the single 
<typeStatus id="54DFB1C09327B507C92EF8F1FB07F8B3" box="[1083,1181,1841,1867]" pageId="35" pageNumber="258">holotype</typeStatus>
, a male 20.5 × 
<quantity id="4C9CA2879327B507C857F8F2FA33F8B4" box="[1346,1449,1842,1868]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="35" pageNumber="258" unit="mm" value="21.1">21.1 mm</quantity>
, was considered to be close to 
<taxonomicName id="4C6474E19327B507C979F892FAF4F894" authority="Chen, 1980" authorityName=": Chen" authorityYear="1980" box="[1132,1390,1873,1900]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="258" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B910D3709327B507C979F892FB78F893" box="[1132,1250,1874,1899]" italics="true" pageId="35" pageNumber="258">D. tenuipes</emphasis>
<bibRefCitation id="EFF572939327B507C9FCF891FAF4F894" author="Chen" box="[1257,1390,1873,1900]" firstAuthor="Chen" pageId="35" pageNumber="258" pagination="154 - 160" refId="ref112715" refString="CHEN H. 1980. - Two new species of crabs from South China Sea. Oceanologia et Limnologia Sinica 11 (2): 154 - 160, pls 1 - 2 (In Chinese with ‰ nglish abstract)." type="journal article" year="1980">Chen, 1980</bibRefCitation>
</taxonomicName>
. Both species were regarded as “different from all other species of 
<taxonomicName id="4C6474E19327B507CE38F852FC1BF853" authorityName="Fabricius" authorityYear="1798" box="[813,897,1938,1963]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="258" phylum="Arthropoda" rank="genus">
<emphasis id="B910D3709327B507CE38F852FC1BF853" box="[813,897,1938,1963]" italics="true" pageId="35" pageNumber="258">Dorippe</emphasis>
</taxonomicName>
in having elongate P2 and P3, with the P3 merus six or more times longer than high”, although to a lesser degree in 
<taxonomicName id="4C6474E19327B507CE5EF812FC58F813" authorityName=": Chen" authorityYear="1980" box="[843,962,2002,2027]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="258" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B910D3709327B507CE5EF812FC58F813" box="[843,962,2002,2027]" italics="true" pageId="35" pageNumber="258">D. tenuipes</emphasis>
</taxonomicName>
. In 
<taxonomicName id="4C6474E19327B507CEE7F812FBF5F813" authorityName=": Manning" authorityYear="1993" box="[1010,1135,2001,2027]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="35" pageNumber="258" phylum="Arthropoda" rank="species" species="trilobata">
<emphasis id="B910D3709327B507CEE7F812FBF5F813" box="[1010,1135,2001,2027]" italics="true" pageId="35" pageNumber="258">D. trilobata</emphasis>
</taxonomicName>
the P2 and P3 (
<figureCitation id="135F13E79327B507C833F811FA3DF813" box="[1318,1447,2001,2028]" captionStart="FIG" captionStartId="37.[132,143,1963,1981]" captionTargetBox="[135,1453,216,1873]" captionTargetId="figure-102@37.[731,818,214,1264]" captionTargetPageId="37" captionText="FIG. 15. — Dorippinae H. Milne Edwards,1837 n. stat.: A-D, J, Dorippe trilobata Manning,1993,holotype, ♂ 20.5 × 21.1 mm, off Mitchell River,Admiralty Gulf,AM P27124: A, habitus; B, ventral surface; C, carapace; D, pleon; J, callosity. E-I: Dorippe glabra Manning, 1993, holotype, ovigerous ♀ 24.3 × 26.7 mm, Chambers Bay, Australia, AM P13363: E, habitus with detached right cheliped, right P2 and P3, left P3; F, carapace; G, ventral surface; H, pleon; I, callosity." figureDoi="http://doi.org/10.5281/zenodo.8071291" httpUri="https://zenodo.org/record/8071291/files/figure.png" pageId="35" pageNumber="258">Fig. 15A, B</figureCitation>
) are longer and thinner than those of most 
<emphasis id="B910D3709322B502CF52FF18FD0BFF09" box="[583,657,216,241]" italics="true" pageId="38" pageNumber="261">Doripp</emphasis>
e (
<figureCitation id="135F13E79322B502CFBFFF17FD6DFF09" box="[682,759,215,241]" captionStart="FIG" captionStartId="30.[132,143,1277,1295]" captionTargetBox="[133,1453,215,1227]" captionTargetId="figure-572@30.[132,1455,214,1227]" captionTargetPageId="30" captionText="FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed." figureDoi="http://doi.org/10.5281/zenodo.8071279" httpUri="https://zenodo.org/record/8071279/files/figure.png" pageId="38" pageNumber="261">Fig. 10</figureCitation>
), thus quite similar to those of 
<taxonomicName id="4C6474E19322B502CCACFF38FDB4FEE9" authorityName=": Chen" authorityYear="1980" box="[441,558,248,273]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B910D3709322B502CCACFF38FDB4FEE9" box="[441,558,248,273]" italics="true" pageId="38" pageNumber="261">D. tenuipes</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CF2EFF37FD15FEE9" box="[571,655,247,273]" captionStart="FIG" captionStartId="26.[132,143,1214,1232]" captionTargetBox="[140,1456,215,1172]" captionTargetId="figure-502@26.[140,800,691,1172]" captionTargetPageId="26" captionText="FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area." figureDoi="http://doi.org/10.5281/zenodo.8071277" httpUri="https://zenodo.org/record/8071277/files/figure.png" pageId="38" pageNumber="261">Fig. 9D</figureCitation>
) but not as thin and elongated as in the latter. The long meri of P2 and P3 of 
<taxonomicName id="4C6474E19322B502CDD0FEF8FEDDFEA9" authorityName=": Manning" authorityYear="1993" box="[197,327,311,337]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="trilobata">
<emphasis id="B910D3709322B502CDD0FEF8FEDDFEA9" box="[197,327,311,337]" italics="true" pageId="38" pageNumber="261">D. trilobata</emphasis>
</taxonomicName>
are naked and the dactyli of P2 and especially P3 are enlarged. A characteristic feature of 
<taxonomicName id="4C6474E19322B502CF90FE98FD65FE89" authorityName=": Manning" authorityYear="1993" box="[645,767,343,369]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="trilobata">
<emphasis id="B910D3709322B502CF90FE98FD65FE89" box="[645,767,343,369]" italics="true" pageId="38" pageNumber="261">D. trilobata</emphasis>
</taxonomicName>
, namely the pleonal somites 2 and 3 with three sharp teeth, is not found in 
<taxonomicName id="4C6474E19322B502CC0DFE57FE15FE48" authorityName=": Chen" authorityYear="1980" box="[280,399,407,432]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B910D3709322B502CC0DFE57FE15FE48" box="[280,399,407,432]" italics="true" pageId="38" pageNumber="261">D. tenuipes</emphasis>
</taxonomicName>
where there are only low rounded and granular elevations. In 
<taxonomicName id="4C6474E19322B502CCABFE77FDDAFE28" authorityName=": Manning" authorityYear="1993" box="[446,576,438,464]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="trilobata">
<emphasis id="B910D3709322B502CCABFE77FDDAFE28" box="[446,576,438,464]" italics="true" pageId="38" pageNumber="261">D. trilobata</emphasis>
</taxonomicName>
, the anterolateral margin is unarmed anterior to epibranchial region and then shows low tubercles, whereas in 
<taxonomicName id="4C6474E19322B502CCF7FE37FDC3FDE8" authorityName=": Chen" authorityYear="1980" box="[482,601,503,528]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B910D3709322B502CCF7FE37FDC3FDE8" box="[482,601,503,528]" italics="true" pageId="38" pageNumber="261">D. tenuipes</emphasis>
</taxonomicName>
it bears numerous denticles. Most significantly, the callosity of 
<taxonomicName id="4C6474E19322B502CF9EFDD7FC9CFDC8" authorityName=": Manning" authorityYear="1993" box="[651,774,534,560]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="trilobata">
<emphasis id="B910D3709322B502CF9EFDD7FC9CFDC8" box="[651,774,534,560]" italics="true" pageId="38" pageNumber="261">D. trilobata</emphasis>
</taxonomicName>
is simply hemicircular (
<figureCitation id="135F13E79322B502CC95FDF6FD83FDA8" box="[384,537,566,593]" captionStart="FIG" captionStartId="37.[132,143,1963,1981]" captionTargetBox="[135,1453,216,1873]" captionTargetId="figure-102@37.[731,818,214,1264]" captionTargetPageId="37" captionText="FIG. 15. — Dorippinae H. Milne Edwards,1837 n. stat.: A-D, J, Dorippe trilobata Manning,1993,holotype, ♂ 20.5 × 21.1 mm, off Mitchell River,Admiralty Gulf,AM P27124: A, habitus; B, ventral surface; C, carapace; D, pleon; J, callosity. E-I: Dorippe glabra Manning, 1993, holotype, ovigerous ♀ 24.3 × 26.7 mm, Chambers Bay, Australia, AM P13363: E, habitus with detached right cheliped, right P2 and P3, left P3; F, carapace; G, ventral surface; H, pleon; I, callosity." figureDoi="http://doi.org/10.5281/zenodo.8071291" httpUri="https://zenodo.org/record/8071291/files/figure.png" pageId="38" pageNumber="261">Fig. 15A, C, J</figureCitation>
), contrasting with the complex callosity in the form of a double calcified bulge with a whitish central area in 
<taxonomicName id="4C6474E19322B502CC98FDB7FD9EFD68" authorityName=": Chen" authorityYear="1980" box="[397,516,631,656]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B910D3709322B502CC98FDB7FD9EFD68" box="[397,516,631,656]" italics="true" pageId="38" pageNumber="261">D. tenuipes</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CF07FDB6FDF6FD68" box="[530,620,630,656]" captionStart="FIG" captionStartId="26.[132,143,1214,1232]" captionTargetBox="[140,1456,215,1172]" captionTargetId="figure-502@26.[140,800,691,1172]" captionTargetPageId="26" captionText="FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area." figureDoi="http://doi.org/10.5281/zenodo.8071277" httpUri="https://zenodo.org/record/8071277/files/figure.png" pageId="38" pageNumber="261">Figs 9D</figureCitation>
; 
<figureCitation id="135F13E79322B502CF6DFDB6FD54FD68" box="[632,718,630,656]" captionStart="FIG" captionStartId="87.[132,143,1829,1847]" captionTargetBox="[347,1239,216,1787]" captionTargetId="figure-107@87.[790,881,1000,1790]" captionTargetPageId="87" captionText="FIG. 33. — Callosities in Dorippoidinae n. subfam. (Dorippoides) and Dorippinae n. stat. (Dorippe): A, callosity reduced to a thin calcified band: Dorippoides facchino (Herbst, 1785), ♂ 25.3 × 32.0 mm, Malaysia, Johore, Pontian, ZRC 1991.6672. B, no apparent callosity in male Dorippoides nudipes Manning &amp; Holthuis, 1986, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227. C, callosity as a thick, elongated bulge: Dorippe quadridens (Fabricius, 1793), ♂ 39.0 × 41.4 mm, Thailand, Pattani Province, ZRC 2003.0126. D-F, callosity as a hemicircular structure: close-up views from different angles: D, E. Dorippe sinica Chen, 1980. D, ♀ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; E, ♂ 36.6 × 38.6 mm, Japan, Kochi, Shikoku, SMF 57855. F, D. frascone (Herbst, 1785), ♂ 29.7 × 31.0 mm, Philippines, Exp. Panglao 2004, ZRC 2008.0076. G, H, callosity as an arched double cup: D. tenuipes Chen, 1980, South China Sea, ZRC 1999.0009: G, ovigerous ♀ 17.3 × 19.6 mm; H, ♂ 13.2 × 14.2 mm. Abbreviations: b, branchiostegite; c, callosity; cx2-cx5, P2-P5 coxae; f, sulcated part of coxa; l, onepiece lateroventral part of coxa; m, membrane; p, pleon; pr, process of retention of female pleonal somite 2; pl6, exposed pleurite 6; P2-P5, pereiopods 2-5; r, rim of carapace posterior margin; s, strip along posterior rim; 1, 2, pleonal somites 1, 2; t, bottom of callosity with special texture; 7, 8, thoracic sternites 7, 8." figureDoi="http://doi.org/10.5281/zenodo.8071333" httpUri="https://zenodo.org/record/8071333/files/figure.png" pageId="38" pageNumber="261">33G, H</figureCitation>
).The anterolateral margin of the carapace without spines, tubercles or denticles in front of the epibranchial region is as in 
<taxonomicName id="4C6474E19322B502CD91FD16FF7BFD17" authorityName=": Chen" authorityYear="1980" box="[132,225,726,751]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="B910D3709322B502CD91FD16FF7BFD17" box="[132,225,726,751]" italics="true" pageId="38" pageNumber="261">D. sinica</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CDFBFD15FEE9FD08" box="[238,371,725,752]" captionStart="FIG" captionStartId="30.[132,143,1277,1295]" captionTargetBox="[133,1453,215,1227]" captionTargetId="figure-572@30.[132,1455,214,1227]" captionTargetPageId="30" captionText="FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed." figureDoi="http://doi.org/10.5281/zenodo.8071279" httpUri="https://zenodo.org/record/8071279/files/figure.png" pageId="38" pageNumber="261">Fig. 10C, D</figureCitation>
); but the three detached spines on the lower margin of the orbit contrast with the cluster in 
<taxonomicName id="4C6474E19322B502CFDEFD36FF2AFCD7" authorityName=": Chen" authorityYear="1980" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="B910D3709322B502CFDEFD36FF2AFCD7" italics="true" pageId="38" pageNumber="261">D. sinica</emphasis>
</taxonomicName>
and the higher number of teeth there in 
<taxonomicName id="4C6474E19322B502CF64FCD6FC9CFCD7" authorityName=": Holthuis &amp; Manning" authorityYear="1990" box="[625,774,789,815]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B910D3709322B502CF64FCD6FC9CFCD7" box="[625,774,789,815]" italics="true" pageId="38" pageNumber="261">D. quadridens</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CD99FCF5FF70FCB7" box="[140,234,821,847]" captionStart="FIG" captionStartId="32.[132,143,1707,1725]" captionTargetBox="[132,1455,215,1665]" captionTargetId="figure-169@32.[132,1455,215,1665]" captionTargetPageId="32" captionText="FIG. 12. — Dorippinae H. Milne Edwards,1837 n. stat.. Dorippe quadridens (Fabricius,1793):A-D, ovigerous ♀ 38.1 × 33.7 mm, South China Sea,Hee Huat,ZRC 1984.6307:A, habitus; B, anterior region, ventral view; C, thoracic sternum and pleon; D, vulvae. E, ovigerous ♀ 28.0 × 29.0 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279): thoracic sternum and vulvae." figureDoi="http://doi.org/10.5281/zenodo.8071285" httpUri="https://zenodo.org/record/8071285/files/figure.png" pageId="38" pageNumber="261">Fig. 12B</figureCitation>
) and may approximate the 3-5 teeth of 
<taxonomicName id="4C6474E19322B502CF81FCF6FC9DFCB7" authorityName=": Chen" authorityYear="1980" box="[660,775,821,847]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="frascone">
<emphasis id="B910D3709322B502CF81FCF6FC9DFCB7" box="[660,775,821,847]" italics="true" pageId="38" pageNumber="261">D. frascone</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CD99FC95FF46FC97" box="[140,220,853,879]" captionStart="FIG" captionStartId="36.[133,144,1962,1980]" captionTargetBox="[132,1455,216,1920]" captionTargetId="figure-16@36.[132,1455,216,1931]" captionTargetPageId="36" captionText="FIG. 14. — Dorippinae H. Milne Edwards,1837 n. stat.:A-G: Dorippe frascone (Herbst,1785),Philippines,Panglao Island,ZRC 2008.0076:A-D, ♂ 29.7 × 31.0 mm, A, habitus; B, ventral view; C, thoracic sternum with pleon; D, G1; E-G, prepubertal ♀ 23.3 × 24.4 mm: E, habitus;F, ventral view; G, vulvae; H, I, Dorippe irrorata Manning &amp; Holthuis, 1986, holotype, ♂ 21.5 × 22.0 mm, Andaman Sea, south of Mergui Archipelago, USNM 172495: H, habitus; I, ventral surface." figureDoi="http://doi.org/10.5281/zenodo.8071289" httpUri="https://zenodo.org/record/8071289/files/figure.png" pageId="38" pageNumber="261">Fig. 14</figureCitation>
A-D). 
<taxonomicName id="4C6474E19322B502CC32FC96FE7AFC97" authorityName=": Manning" authorityYear="1993" box="[295,480,853,879]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="trilobata">
<emphasis id="B910D3709322B502CC32FC96FE7AFC97" box="[295,480,853,879]" italics="true" pageId="38" pageNumber="261">Dorippe trilobata</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CCE4FC95FDDBFC88" box="[497,577,853,880]" captionStart="FIG" captionStartId="37.[132,143,1963,1981]" captionTargetBox="[135,1453,216,1873]" captionTargetId="figure-102@37.[731,818,214,1264]" captionTargetPageId="37" captionText="FIG. 15. — Dorippinae H. Milne Edwards,1837 n. stat.: A-D, J, Dorippe trilobata Manning,1993,holotype, ♂ 20.5 × 21.1 mm, off Mitchell River,Admiralty Gulf,AM P27124: A, habitus; B, ventral surface; C, carapace; D, pleon; J, callosity. E-I: Dorippe glabra Manning, 1993, holotype, ovigerous ♀ 24.3 × 26.7 mm, Chambers Bay, Australia, AM P13363: E, habitus with detached right cheliped, right P2 and P3, left P3; F, carapace; G, ventral surface; H, pleon; I, callosity." figureDoi="http://doi.org/10.5281/zenodo.8071291" httpUri="https://zenodo.org/record/8071291/files/figure.png" pageId="38" pageNumber="261">Fig. 15</figureCitation>
A-D) differs from all other species of 
<taxonomicName id="4C6474E19322B502CC45FCB6FE3EFC77" authorityName="Fabricius" authorityYear="1798" box="[336,420,886,911]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B910D3709322B502CC45FCB6FE3EFC77" box="[336,420,886,911]" italics="true" pageId="38" pageNumber="261">Dorippe</emphasis>
</taxonomicName>
in having a short, triangular and straight-edged pleon, instead of the longer and irregular-edged pleon of other 
<taxonomicName id="4C6474E19322B502CC3DFC75FEE6FC36" authorityName="Fabricius" authorityYear="1798" box="[296,380,949,974]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B910D3709322B502CC3DFC75FEE6FC36" box="[296,380,949,974]" italics="true" pageId="38" pageNumber="261">Dorippe</emphasis>
</taxonomicName>
; in addition, the three sharp tubercles on pleonal somites 2 and 3 are much more acute than in 
<taxonomicName id="4C6474E19322B502CDB4FC35FEAFFBF6" authorityName=": Holthuis &amp; Manning" authorityYear="1990" box="[161,309,1012,1038]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B910D3709322B502CDB4FC35FEAFFBF6" box="[161,309,1012,1038]" italics="true" pageId="38" pageNumber="261">D. quadridens</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CC56FC34FE3CFBF7" box="[323,422,1012,1039]" captionStart="FIG" captionStartId="30.[132,143,1277,1295]" captionTargetBox="[133,1453,215,1227]" captionTargetId="figure-572@30.[132,1455,214,1227]" captionTargetPageId="30" captionText="FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed." figureDoi="http://doi.org/10.5281/zenodo.8071279" httpUri="https://zenodo.org/record/8071279/files/figure.png" pageId="38" pageNumber="261">Figs 10A</figureCitation>
; 
<figureCitation id="135F13E79322B502CCA7FC35FE78FBF7" box="[434,482,1013,1039]" captionStart="FIG" captionStartId="31.[132,143,1307,1325]" captionTargetBox="[132,1455,215,1257]" captionTargetId="figure-351@31.[131,1455,207,1259]" captionTargetPageId="31" captionText="FIG. 11. — Dorippinae H. Milne Edwards, 1837 n. stat.. Dorippe quadridens (Fabricius, 1793): A-C, ♂ 38.1 × 36.3 mm, China Sea near Singapore, Hee Huat, ZRC 1984.6308. A, anterior region; B, thoracic sternum with pleon; C, thoracic sternum without pleon, G1. D, ♂ 16.6 × 17.0 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279): thoracic sternum without pleon, G1 and G2." figureDoi="http://doi.org/10.5281/zenodo.8071283" httpUri="https://zenodo.org/record/8071283/files/figure.png" pageId="38" pageNumber="261">11B</figureCitation>
), 
<taxonomicName id="4C6474E19322B502CCE0FC35FDC8FBF6" authorityName=": Chen" authorityYear="1980" box="[501,594,1013,1038]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="B910D3709322B502CCE0FC35FDC8FBF6" box="[501,594,1013,1038]" italics="true" pageId="38" pageNumber="261">D. sinica</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CF75FC34FD5FFBF7" box="[608,709,1012,1039]" captionStart="FIG" captionStartId="30.[132,143,1277,1295]" captionTargetBox="[133,1453,215,1227]" captionTargetId="figure-572@30.[132,1455,214,1227]" captionTargetPageId="30" captionText="FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed." figureDoi="http://doi.org/10.5281/zenodo.8071279" httpUri="https://zenodo.org/record/8071279/files/figure.png" pageId="38" pageNumber="261">Figs 10C</figureCitation>
; 
<figureCitation id="135F13E79322B502CFC5FC35FD65FBF7" box="[720,767,1013,1039]" captionStart="FIG" captionStartId="34.[133,144,1255,1273]" captionTargetBox="[132,1455,214,1213]" captionTargetId="figure-432@34.[132,1455,207,1220]" captionTargetPageId="34" captionText="FIG. 13. — Dorippinae H. Milne Edwards, 1837 n. stat.. Dorippe sinica Chen, 1980: A, B, ♂ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470: A, ventral view, thoracic sternum and pleon; B, thoracic sternum,G1 and G2; C, D, ♀ 34.8 × 38.9 mm, same data as A, B; C, thoracic sternum and pleon; D, thoracic sternum and vulvae." figureDoi="http://doi.org/10.5281/zenodo.8071287" httpUri="https://zenodo.org/record/8071287/files/figure.png" pageId="38" pageNumber="261">13A</figureCitation>
) and 
<taxonomicName id="4C6474E19322B502CDA0FBD5FEB2FBD6" authorityName=": Chen" authorityYear="1980" box="[181,296,1044,1070]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="frascone">
<emphasis id="B910D3709322B502CDA0FBD5FEB2FBD6" box="[181,296,1044,1070]" italics="true" pageId="38" pageNumber="261">D. frascone</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CC2DFBD4FE1CFBD6" box="[312,390,1044,1070]" captionStart="FIG" captionStartId="36.[133,144,1962,1980]" captionTargetBox="[132,1455,216,1920]" captionTargetId="figure-16@36.[132,1455,216,1931]" captionTargetPageId="36" captionText="FIG. 14. — Dorippinae H. Milne Edwards,1837 n. stat.:A-G: Dorippe frascone (Herbst,1785),Philippines,Panglao Island,ZRC 2008.0076:A-D, ♂ 29.7 × 31.0 mm, A, habitus; B, ventral view; C, thoracic sternum with pleon; D, G1; E-G, prepubertal ♀ 23.3 × 24.4 mm: E, habitus;F, ventral view; G, vulvae; H, I, Dorippe irrorata Manning &amp; Holthuis, 1986, holotype, ♂ 21.5 × 22.0 mm, Andaman Sea, south of Mergui Archipelago, USNM 172495: H, habitus; I, ventral surface." figureDoi="http://doi.org/10.5281/zenodo.8071289" httpUri="https://zenodo.org/record/8071289/files/figure.png" pageId="38" pageNumber="261">Fig. 14</figureCitation>
A-D). The relatively small size 20.5 × 
<quantity id="4C9CA2879322B502CD8EFBF4FE9BFBB7" box="[155,257,1076,1103]" metricMagnitude="-2" metricUnit="m" metricValue="2.11" pageId="38" pageNumber="261" unit="mm" value="21.1">21.1 mm</quantity>
of the 
<typeStatus id="54DFB1C09322B502CC46FBF4FE28FBB6" box="[339,434,1076,1102]" pageId="38" pageNumber="261">holotype</typeStatus>
of 
<taxonomicName id="4C6474E19322B502CCCCFBF5FDCCFBB6" authorityName=": Manning" authorityYear="1993" box="[473,598,1076,1102]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="trilobata">
<emphasis id="B910D3709322B502CCCCFBF5FDCCFBB6" box="[473,598,1076,1102]" italics="true" pageId="38" pageNumber="261">D. trilobata</emphasis>
</taxonomicName>
cannot account for these differences, so until more material is available this species can be considered valid.
</paragraph>
<caption id="DF1B5FEA9320B500CD90F86AFADCF812" ID-DOI="http://doi.org/10.5281/zenodo.8071289" ID-Zenodo-Dep="8071289" httpUri="https://zenodo.org/record/8071289/files/figure.png" pageId="36" pageNumber="259" startId="36.[133,144,1962,1980]" targetBox="[132,1455,216,1920]" targetPageId="36" targetType="figure">
<paragraph id="8BDB0F629320B500CD90F86AFADCF812" blockId="36.[133,1457,1962,2028]" pageId="36" pageNumber="259">
<smallCapsWord id="8D3D99BE9320B500CD90F86AFF07F843" baselines="1975,1976" box="[133,157,1962,1980]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="36" pageNumber="259">FIG</smallCapsWord>
. 14. — 
<taxonomicName id="4C6474E19320B500CDC9F86AFE67F844" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" box="[220,509,1962,1980]" class="Malacostraca" family="Dorippidae" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="259" phylum="Arthropoda" rank="subFamily" status="n. stat." subFamily="Dorippinae">Dorippinae H. Milne Edwards,1837</taxonomicName>
<taxonomicNameLabel id="A2236E0B9320B500CF17F86AFDA7F844" box="[514,573,1962,1980]" pageId="36" pageNumber="259">n. stat.</taxonomicNameLabel>
: 
<emphasis id="B910D3709320B500CF57F86AFDFEF844" bold="true" box="[578,612,1962,1980]" pageId="36" pageNumber="259">A -G</emphasis>
: 
<taxonomicName id="4C6474E19320B500CF7BF86AFCECF844" authority="(Herbst, 1785)" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[622,886,1962,1980]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="259" phylum="Arthropoda" rank="species" species="frascone">
<emphasis id="B910D3709320B500CF7BF86AFD61F844" box="[622,763,1962,1980]" italics="true" pageId="36" pageNumber="259">Dorippe frascone</emphasis>
(Herbst,1785)
</taxonomicName>
,Philippines,Panglao Island,ZRC 2008.0076: 
<emphasis id="B910D3709320B500C9F8F86AFA95F844" bold="true" box="[1261,1295,1962,1980]" pageId="36" pageNumber="259">A -D</emphasis>
, ♂ 29.7 × 31.0 mm, 
<emphasis id="B910D3709320B500CD90F801FF08F82B" bold="true" box="[133,146,1985,2003]" pageId="36" pageNumber="259">A</emphasis>
, habitus; 
<emphasis id="B910D3709320B500CDF7F801FF75F82B" bold="true" box="[226,239,1985,2003]" pageId="36" pageNumber="259">B</emphasis>
, ventral view; 
<emphasis id="B910D3709320B500CC70F801FEE9F82B" bold="true" box="[357,371,1985,2003]" pageId="36" pageNumber="259">C</emphasis>
, thoracic sternum with pleon; 
<emphasis id="B910D3709320B500CF78F801FDE1F82B" bold="true" box="[621,635,1985,2003]" pageId="36" pageNumber="259">D</emphasis>
, G1; 
<emphasis id="B910D3709320B500CFB3F801FD52F82B" bold="true" box="[678,712,1985,2003]" pageId="36" pageNumber="259">E -G</emphasis>
, prepubertal ♀ 23.3 × 24.4 mm: 
<emphasis id="B910D3709320B500CEC6F801FC45F82B" bold="true" box="[979,991,1985,2003]" pageId="36" pageNumber="259">E</emphasis>
, habitus; 
<emphasis id="B910D3709320B500C93AF801FBA0F82B" bold="true" box="[1071,1082,1985,2003]" pageId="36" pageNumber="259">F</emphasis>
, ventral view; 
<emphasis id="B910D3709320B500C9A5F801FB24F82B" bold="true" box="[1200,1214,1985,2003]" pageId="36" pageNumber="259">G</emphasis>
, vulvae; 
<emphasis id="B910D3709320B500C812F801FA8FF82B" bold="true" box="[1287,1301,1985,2003]" pageId="36" pageNumber="259">H</emphasis>
, 
<emphasis id="B910D3709320B500C80AF801FABFF82B" bold="true" box="[1311,1317,1985,2003]" pageId="36" pageNumber="259">I</emphasis>
, 
<taxonomicName id="4C6474E19320B500C83BF801FEC4F812" authority="Manning &amp; Holthuis, 1986" authorityName="Manning &amp; Holthuis" authorityYear="1986" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="36" pageNumber="259" phylum="Arthropoda" rank="species" species="irrorata">
<emphasis id="B910D3709320B500C83BF801FA35F82B" box="[1326,1455,1985,2003]" italics="true" pageId="36" pageNumber="259">Dorippe irrorata</emphasis>
<bibRefCitation id="EFF572939320B500CD90F818FEC4F812" author="Manning &amp; Holthuis" box="[133,350,2008,2026]" firstAuthor="Manning" pageId="36" pageNumber="259" pagination="363 - 365" refId="ref131244" refString="MANNING R. B. &amp; HOLTHUIS L. B. 1986. - Preliminary descriptions of four new species of dorippid crabs from the Indo-west Pacific region (Crustacea: Decapoda: Brachyura). Proceedings of the biological Society of Washington 99 (2): 363 - 365." type="journal article" year="1986">Manning &amp; Holthuis, 1986</bibRefCitation>
</taxonomicName>
, holotype, ♂ 21.5 × 22.0 mm, Andaman Sea, south of Mergui Archipelago, USNM 172495: 
<emphasis id="B910D3709320B500C941F818FBF8F812" bold="true" box="[1108,1122,2008,2026]" pageId="36" pageNumber="259">H</emphasis>
, habitus; 
<emphasis id="B910D3709320B500C9A1F818FB20F812" bold="true" box="[1204,1210,2008,2026]" pageId="36" pageNumber="259">I</emphasis>
, ventral surface.
</paragraph>
</caption>
<caption id="DF1B5FEA9321B501CD91F86BFAA6F813" ID-DOI="http://doi.org/10.5281/zenodo.8071291" ID-Zenodo-Dep="8071291" httpUri="https://zenodo.org/record/8071291/files/figure.png" pageId="37" pageNumber="260" startId="37.[132,143,1963,1981]" targetBox="[135,1453,216,1873]" targetPageId="37" targetType="figure">
<paragraph id="8BDB0F629321B501CD91F86BFAA6F813" blockId="37.[132,1454,1963,2027]" pageId="37" pageNumber="260">
<smallCapsWord id="8D3D99BE9321B501CD91F86BFF06F844" baselines="1976,1977" box="[132,156,1963,1981]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="37" pageNumber="260">FIG</smallCapsWord>
. 15. — 
<taxonomicName id="4C6474E19321B501CDC9F86BFE65F845" authority="H. Milne Edwards, 1837" authorityName="H. Milne Edwards" authorityYear="1837" box="[220,511,1963,1981]" class="Malacostraca" family="Dorippidae" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="260" phylum="Arthropoda" rank="subFamily" status="n. stat." subFamily="Dorippinae">Dorippinae H. Milne Edwards,1837</taxonomicName>
<taxonomicNameLabel id="A2236E0B9321B501CF16F86BFDA5F845" box="[515,575,1963,1981]" pageId="37" pageNumber="260">n. stat.</taxonomicNameLabel>
: 
<emphasis id="B910D3709321B501CF51F86BFDFCF845" bold="true" box="[580,614,1963,1981]" pageId="37" pageNumber="260">A -D</emphasis>
, 
<emphasis id="B910D3709321B501CF7AF86BFDE3F845" bold="true" box="[623,633,1963,1981]" pageId="37" pageNumber="260">J</emphasis>
, 
<taxonomicName id="4C6474E19321B501CF96F86BFC14F845" authority="Manning, 1993" authorityName=": Manning" authorityYear="1993" box="[643,910,1963,1981]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="260" phylum="Arthropoda" rank="species" species="trilobata">
<emphasis id="B910D3709321B501CF96F86BFC96F845" box="[643,780,1963,1981]" italics="true" pageId="37" pageNumber="260">Dorippe trilobata</emphasis>
<bibRefCitation id="EFF572939321B501CE04F86BFC14F845" author="Manning" box="[785,910,1963,1981]" firstAuthor="Manning" pageId="37" pageNumber="260" pagination="1 - 4" refId="ref131161" refString="MANNING R. B. 1993. - Two new dorippid crabs from Australia (Crustacea: Decapoda: Dorippidae). Records from the Australian Museum 45: 1 - 4. https: // doi. org / 10.3853 / j. 0067 - 1975.45.1993.124" type="journal article" year="1993">Manning,1993</bibRefCitation>
</taxonomicName>
,holotype, ♂ 20.5 × 21.1 mm, off Mitchell River,Admiralty Gulf,AM P27124: 
<emphasis id="B910D3709321B501CDDBF802FF41F82C" bold="true" box="[206,219,1986,2004]" pageId="37" pageNumber="260">A</emphasis>
, habitus; 
<emphasis id="B910D3709321B501CC39F802FEA3F82C" bold="true" box="[300,313,1986,2004]" pageId="37" pageNumber="260">B</emphasis>
, ventral surface; 
<emphasis id="B910D3709321B501CCDDF802FE4CF82C" bold="true" box="[456,470,1986,2004]" pageId="37" pageNumber="260">C</emphasis>
, carapace; 
<emphasis id="B910D3709321B501CF22F802FDDFF82C" bold="true" box="[567,581,1986,2004]" pageId="37" pageNumber="260">D</emphasis>
, pleon; 
<emphasis id="B910D3709321B501CF9DF802FD08F82C" bold="true" box="[648,658,1986,2004]" pageId="37" pageNumber="260">J</emphasis>
, callosity. 
<emphasis id="B910D3709321B501CFFCF802FC98F82C" bold="true" box="[745,770,1986,2004]" pageId="37" pageNumber="260">E -I</emphasis>
: 
<taxonomicName id="4C6474E19321B501CE1EF802FB90F82C" authority="Manning, 1993" authorityName=": Manning" authorityYear="1993" box="[779,1034,1986,2004]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="37" pageNumber="260" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="B910D3709321B501CE1EF802FC1DF82C" box="[779,903,1986,2004]" italics="true" pageId="37" pageNumber="260">Dorippe glabra</emphasis>
<bibRefCitation id="EFF572939321B501CE9EF802FB90F82C" author="Manning" box="[907,1034,1986,2004]" firstAuthor="Manning" pageId="37" pageNumber="260" pagination="1 - 4" refId="ref131161" refString="MANNING R. B. 1993. - Two new dorippid crabs from Australia (Crustacea: Decapoda: Dorippidae). Records from the Australian Museum 45: 1 - 4. https: // doi. org / 10.3853 / j. 0067 - 1975.45.1993.124" type="journal article" year="1993">Manning, 1993</bibRefCitation>
</taxonomicName>
, holotype, ovigerous ♀ 24.3 × 26.7 mm, Chambers Bay, Australia, AM P13363: 
<emphasis id="B910D3709321B501CC7EF819FEEDF813" bold="true" box="[363,375,2009,2027]" pageId="37" pageNumber="260">E</emphasis>
, habitus with detached right cheliped, right P2 and P3, left P3; 
<emphasis id="B910D3709321B501CE92F819FC08F813" bold="true" box="[903,914,2009,2027]" pageId="37" pageNumber="260">F</emphasis>
, carapace; 
<emphasis id="B910D3709321B501CEE1F818FB98F812" bold="true" box="[1012,1026,2008,2026]" pageId="37" pageNumber="260">G</emphasis>
, ventral surface; 
<emphasis id="B910D3709321B501C987F819FB3AF813" bold="true" box="[1170,1184,2009,2027]" pageId="37" pageNumber="260">H</emphasis>
, pleon; 
<emphasis id="B910D3709321B501C9F6F819FB73F813" bold="true" box="[1251,1257,2009,2027]" pageId="37" pageNumber="260">I</emphasis>
, callosity.
</paragraph>
</caption>
<paragraph id="8BDB0F629322B502CD8EFB55FEE9F813" blockId="38.[130,777,215,2028]" pageId="38" pageNumber="261">
<taxonomicName id="4C6474E19322B502CD8EFB55FEAEFB56" authorityName=": Manning" authorityYear="1993" box="[155,308,1172,1198]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="B910D3709322B502CD8EFB55FEAEFB56" box="[155,308,1172,1198]" italics="true" pageId="38" pageNumber="261">Dorippe glabra</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFF572939322B502CC73FB54FDBBFB56" author="Manning" box="[358,545,1172,1198]" firstAuthor="Manning" page="1" pageId="38" pageNumber="261" pagination="1 - 4" refId="ref131161" refString="MANNING R. B. 1993. - Two new dorippid crabs from Australia (Crustacea: Decapoda: Dorippidae). Records from the Australian Museum 45: 1 - 4. https: // doi. org / 10.3853 / j. 0067 - 1975.45.1993.124" type="journal article" year="1993">Manning 1993: 1</bibRefCitation>
, fig. 1; cited by 
<bibRefCitation id="EFF572939322B502CFDDFB54FF6FFB37" author="DAVIE P. J. F." pageId="38" pageNumber="261" refId="ref114947" refString="DAVIE P. J. F. 2002. - Crustacea: Malacostraca. ‰ ucarida (Part 2). Decapoda Anomura, Brachyura, in WELLS A. &amp; HOUSTON W. W. K. (eds), Zological Catalogue of Australia. Vol. 19.3 B. CSIR ̿ Publishing, Melbourne, xiv + 641 p." type="book" year="2002">Davie 2002: 156</bibRefCitation>
), from the north of the 
<collectingRegion id="49A0C1809322B502CCFFFB74FD29FB36" box="[490,691,1204,1230]" country="Australia" name="Northern Territory" pageId="38" pageNumber="261">Northern Territory</collectingRegion>
, known only by an ovigerous female measuring 24.3 × 
<quantity id="4C9CA2879322B502CF89FB14FC9FFB16" box="[668,773,1236,1262]" metricMagnitude="-2" metricUnit="m" metricValue="2.67" pageId="38" pageNumber="261" unit="mm" value="26.7">26.7 mm</quantity>
(deposited in the 
<collectionCode id="ED7597A79322B502CC44FB34FE1AFAF6" box="[337,384,1268,1294]" pageId="38" pageNumber="261">AM</collectionCode>
, see 
<bibRefCitation id="EFF572939322B502CCADFB33FD64FAF6" author="Springthorpe &amp; Lowry" box="[440,766,1267,1294]" firstAuthor="Springthorpe" pageId="38" pageNumber="261" pagination="1 - 134" refId="ref143235" refString="SPRINGTHORPE R. &amp; LOWRY J. 1994. - Catalogue of crustacean type specimens in the Australian Museum: Malacostraca. Technical Reports of the Australian Museum (11): 1 - 134." type="journal article" year="1994">Springthorpe &amp; Lowry 1994</bibRefCitation>
) and found mixed in a large collection identified as 
<taxonomicName id="4C6474E19322B502CFB2FAD4FF58FAB5" authorityName=": Holthuis &amp; Manning" authorityYear="1990" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B910D3709322B502CFB2FAD4FF58FAB5" italics="true" pageId="38" pageNumber="261">D. quadridens</emphasis>
</taxonomicName>
, “could be distinguished on sight from all of the other specimens in that lot by the naked meri of its P2 and P3”, the most significant feature according to 
<bibRefCitation id="EFF572939322B502CF2CFAB3FC98FA75" author="Manning" box="[569,770,1395,1422]" firstAuthor="Manning" page="2" pageId="38" pageNumber="261" pagination="1 - 4" refId="ref131161" refString="MANNING R. B. 1993. - Two new dorippid crabs from Australia (Crustacea: Decapoda: Dorippidae). Records from the Australian Museum 45: 1 - 4. https: // doi. org / 10.3853 / j. 0067 - 1975.45.1993.124" type="journal article" year="1993">Manning (1993: 2</bibRefCitation>
, 4). In photographs of the 
<typeStatus id="54DFB1C09322B502CCB4FA53FD99FA55" box="[417,515,1427,1453]" pageId="38" pageNumber="261">holotype</typeStatus>
, a male, the meri of the attached left P2, detached left P3 and detached right P3 are naked, but the merus of the detached right P2 (
<figureCitation id="135F13E79322B502CF9CFA13FD72FA15" box="[649,744,1491,1518]" captionStart="FIG" captionStartId="37.[132,143,1963,1981]" captionTargetBox="[135,1453,216,1873]" captionTargetId="figure-102@37.[731,818,214,1264]" captionTargetPageId="37" captionText="FIG. 15. — Dorippinae H. Milne Edwards,1837 n. stat.: A-D, J, Dorippe trilobata Manning,1993,holotype, ♂ 20.5 × 21.1 mm, off Mitchell River,Admiralty Gulf,AM P27124: A, habitus; B, ventral surface; C, carapace; D, pleon; J, callosity. E-I: Dorippe glabra Manning, 1993, holotype, ovigerous ♀ 24.3 × 26.7 mm, Chambers Bay, Australia, AM P13363: E, habitus with detached right cheliped, right P2 and P3, left P3; F, carapace; G, ventral surface; H, pleon; I, callosity." figureDoi="http://doi.org/10.5281/zenodo.8071291" httpUri="https://zenodo.org/record/8071291/files/figure.png" pageId="38" pageNumber="261">Fig. 15E</figureCitation>
) is covered with dense setae on the posterior margin. In 
<taxonomicName id="4C6474E19322B502CFDEFA33FF2AF9D4" authorityName=": Chen" authorityYear="1980" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="B910D3709322B502CFDEFA33FF2AF9D4" italics="true" pageId="38" pageNumber="261">D. sinica</emphasis>
</taxonomicName>
the meri of P2 and P3 are naked or barely pubescent in females (
<figureCitation id="135F13E79322B502CDF1F9F2FED2F9B5" box="[228,328,1586,1613]" captionStart="FIG" captionStartId="30.[132,143,1277,1295]" captionTargetBox="[133,1453,215,1227]" captionTargetId="figure-572@30.[132,1455,214,1227]" captionTargetPageId="30" captionText="FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed." figureDoi="http://doi.org/10.5281/zenodo.8071279" httpUri="https://zenodo.org/record/8071279/files/figure.png" pageId="38" pageNumber="261">Fig. 10D</figureCitation>
) and setose in males (
<figureCitation id="135F13E79322B502CF2CF9F2FD06F9B5" box="[569,668,1586,1613]" captionStart="FIG" captionStartId="30.[132,143,1277,1295]" captionTargetBox="[133,1453,215,1227]" captionTargetId="figure-572@30.[132,1455,214,1227]" captionTargetPageId="30" captionText="FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed." figureDoi="http://doi.org/10.5281/zenodo.8071279" httpUri="https://zenodo.org/record/8071279/files/figure.png" pageId="38" pageNumber="261">Fig. 10C</figureCitation>
), whereas they are covered with setae in both sexes of 
<taxonomicName id="4C6474E19322B502CF7AF993FC9DF994" authorityName=": Holthuis &amp; Manning" authorityYear="1990" box="[623,775,1618,1644]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="quadridens">
<emphasis id="B910D3709322B502CF7AF993FC9DF994" box="[623,775,1618,1644]" italics="true" pageId="38" pageNumber="261">D. quadridens</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CD9EF9B2FE8FF975" box="[139,277,1650,1677]" captionStart="FIG" captionStartId="30.[132,143,1277,1295]" captionTargetBox="[133,1453,215,1227]" captionTargetId="figure-572@30.[132,1455,214,1227]" captionTargetPageId="30" captionText="FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed." figureDoi="http://doi.org/10.5281/zenodo.8071279" httpUri="https://zenodo.org/record/8071279/files/figure.png" pageId="38" pageNumber="261">Figs 10A, B</figureCitation>
; 
<figureCitation id="135F13E79322B502CC36F9B2FECCF974" box="[291,342,1650,1676]" captionStart="FIG" captionStartId="32.[132,143,1707,1725]" captionTargetBox="[132,1455,215,1665]" captionTargetId="figure-169@32.[132,1455,215,1665]" captionTargetPageId="32" captionText="FIG. 12. — Dorippinae H. Milne Edwards,1837 n. stat.. Dorippe quadridens (Fabricius,1793):A-D, ovigerous ♀ 38.1 × 33.7 mm, South China Sea,Hee Huat,ZRC 1984.6307:A, habitus; B, anterior region, ventral view; C, thoracic sternum and pleon; D, vulvae. E, ovigerous ♀ 28.0 × 29.0 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279): thoracic sternum and vulvae." figureDoi="http://doi.org/10.5281/zenodo.8071285" httpUri="https://zenodo.org/record/8071285/files/figure.png" pageId="38" pageNumber="261">12A</figureCitation>
). The P2, P3 of 
<taxonomicName id="4C6474E19322B502CF06F9B3FDE0F974" authorityName=": Manning" authorityYear="1993" box="[531,634,1650,1676]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="B910D3709322B502CF06F9B3FDE0F974" box="[531,634,1650,1676]" italics="true" pageId="38" pageNumber="261">D. glabra</emphasis>
</taxonomicName>
(female) are sligthly longer and thinner than in most 
<taxonomicName id="4C6474E19322B502CF45F953FD3FF954" authorityName="Fabricius" authorityYear="1798" box="[592,677,1683,1708]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B910D3709322B502CF45F953FD3FF954" box="[592,677,1683,1708]" italics="true" pageId="38" pageNumber="261">Dorippe</emphasis>
</taxonomicName>
, but not so elongated and thin than those of 
<taxonomicName id="4C6474E19322B502CF07F973FD15F934" authorityName=": Manning" authorityYear="1993" box="[530,655,1714,1740]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="trilobata">
<emphasis id="B910D3709322B502CF07F973FD15F934" box="[530,655,1714,1740]" italics="true" pageId="38" pageNumber="261">D. trilobata</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CFB5F972FF02F914" captionStart="FIG" captionStartId="37.[132,143,1963,1981]" captionTargetBox="[135,1453,216,1873]" captionTargetId="figure-102@37.[731,818,214,1264]" captionTargetPageId="37" captionText="FIG. 15. — Dorippinae H. Milne Edwards,1837 n. stat.: A-D, J, Dorippe trilobata Manning,1993,holotype, ♂ 20.5 × 21.1 mm, off Mitchell River,Admiralty Gulf,AM P27124: A, habitus; B, ventral surface; C, carapace; D, pleon; J, callosity. E-I: Dorippe glabra Manning, 1993, holotype, ovigerous ♀ 24.3 × 26.7 mm, Chambers Bay, Australia, AM P13363: E, habitus with detached right cheliped, right P2 and P3, left P3; F, carapace; G, ventral surface; H, pleon; I, callosity." figureDoi="http://doi.org/10.5281/zenodo.8071291" httpUri="https://zenodo.org/record/8071291/files/figure.png" pageId="38" pageNumber="261">Fig. 15A, B</figureCitation>
), and those of 
<taxonomicName id="4C6474E19322B502CC2BF913FE2FF914" authorityName=": Chen" authorityYear="1980" box="[318,437,1747,1772]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B910D3709322B502CC2BF913FE2FF914" box="[318,437,1747,1772]" italics="true" pageId="38" pageNumber="261">D. tenuipes</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CCD0F912FD81F914" box="[453,539,1746,1772]" captionStart="FIG" captionStartId="26.[132,143,1214,1232]" captionTargetBox="[140,1456,215,1172]" captionTargetId="figure-502@26.[140,800,691,1172]" captionTargetPageId="26" captionText="FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area." figureDoi="http://doi.org/10.5281/zenodo.8071277" httpUri="https://zenodo.org/record/8071277/files/figure.png" pageId="38" pageNumber="261">Fig. 9D</figureCitation>
) that are granular. In 
<taxonomicName id="4C6474E19322B502CD91F933FF7DF8F4" authorityName=": Manning" authorityYear="1993" box="[132,231,1778,1804]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="B910D3709322B502CD91F933FF7DF8F4" box="[132,231,1778,1804]" italics="true" pageId="38" pageNumber="261">D. glabra</emphasis>
</taxonomicName>
the merus and carpus of the cheliped (
<figureCitation id="135F13E79322B502CF9FF932FD72F8F4" box="[650,744,1778,1805]" captionStart="FIG" captionStartId="37.[132,143,1963,1981]" captionTargetBox="[135,1453,216,1873]" captionTargetId="figure-102@37.[731,818,214,1264]" captionTargetPageId="37" captionText="FIG. 15. — Dorippinae H. Milne Edwards,1837 n. stat.: A-D, J, Dorippe trilobata Manning,1993,holotype, ♂ 20.5 × 21.1 mm, off Mitchell River,Admiralty Gulf,AM P27124: A, habitus; B, ventral surface; C, carapace; D, pleon; J, callosity. E-I: Dorippe glabra Manning, 1993, holotype, ovigerous ♀ 24.3 × 26.7 mm, Chambers Bay, Australia, AM P13363: E, habitus with detached right cheliped, right P2 and P3, left P3; F, carapace; G, ventral surface; H, pleon; I, callosity." figureDoi="http://doi.org/10.5281/zenodo.8071291" httpUri="https://zenodo.org/record/8071291/files/figure.png" pageId="38" pageNumber="261">Fig. 15E</figureCitation>
) is covered with dense granules, which eliminates 
<taxonomicName id="4C6474E19322B502CF71F8D2FD48F8D4" baseAuthorityName="Herbst" baseAuthorityYear="1785" box="[612,722,1810,1836]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="frascone">
<emphasis id="B910D3709322B502CF71F8D2FD48F8D4" box="[612,722,1810,1836]" italics="true" pageId="38" pageNumber="261">D. frascone</emphasis>
</taxonomicName>
with a smooth carpus. In 
<taxonomicName id="4C6474E19322B502CC4AF8F2FE58F8B3" authorityName=": Manning" authorityYear="1993" box="[351,450,1841,1867]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="B910D3709322B502CC4AF8F2FE58F8B3" box="[351,450,1841,1867]" italics="true" pageId="38" pageNumber="261">D. glabra</emphasis>
</taxonomicName>
the coxa of P3 is flanked by a high hemicircular calcified callosity (
<figureCitation id="135F13E79322B502CF1CF891FD0EF894" box="[521,660,1873,1900]" captionStart="FIG" captionStartId="37.[132,143,1963,1981]" captionTargetBox="[135,1453,216,1873]" captionTargetId="figure-102@37.[731,818,214,1264]" captionTargetPageId="37" captionText="FIG. 15. — Dorippinae H. Milne Edwards,1837 n. stat.: A-D, J, Dorippe trilobata Manning,1993,holotype, ♂ 20.5 × 21.1 mm, off Mitchell River,Admiralty Gulf,AM P27124: A, habitus; B, ventral surface; C, carapace; D, pleon; J, callosity. E-I: Dorippe glabra Manning, 1993, holotype, ovigerous ♀ 24.3 × 26.7 mm, Chambers Bay, Australia, AM P13363: E, habitus with detached right cheliped, right P2 and P3, left P3; F, carapace; G, ventral surface; H, pleon; I, callosity." figureDoi="http://doi.org/10.5281/zenodo.8071291" httpUri="https://zenodo.org/record/8071291/files/figure.png" pageId="38" pageNumber="261">Fig. 15E, F, I</figureCitation>
) similar to that of 
<taxonomicName id="4C6474E19322B502CDC0F8B2FEAEF873" authorityName=": Chen" authorityYear="1980" box="[213,308,1906,1931]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="sinica">
<emphasis id="B910D3709322B502CDC0F8B2FEAEF873" box="[213,308,1906,1931]" italics="true" pageId="38" pageNumber="261">D. sinica</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CC56F8B1FE06F873" box="[323,412,1905,1931]" captionStart="FIG" captionStartId="26.[132,143,1214,1232]" captionTargetBox="[140,1456,215,1172]" captionTargetId="figure-502@26.[140,800,691,1172]" captionTargetPageId="26" captionText="FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area." figureDoi="http://doi.org/10.5281/zenodo.8071277" httpUri="https://zenodo.org/record/8071277/files/figure.png" pageId="38" pageNumber="261">Figs 9C</figureCitation>
; 
<figureCitation id="135F13E79322B502CCBCF8B1FD9AF874" box="[425,512,1905,1932]" captionStart="FIG" captionStartId="30.[132,143,1277,1295]" captionTargetBox="[133,1453,215,1227]" captionTargetId="figure-572@30.[132,1455,214,1227]" captionTargetPageId="30" captionText="FIG. 10. — Dorippinae H. Milne Edwards, 1837 n. stat.: habitus: A, B, Dorippe quadridens (Fabricius,1793):A, ♂ 36.3 × 38.1 mm, China Sea near Singapore, Hee Huat, ZRC 1984. 6308; B, ♀ 29.5 × 30 mm, NW Madagascar, Ambaro Bay, MNHN-IU-2018-5193 (= MNHN-B18279). C, D, Dorippe sinica Chen, 1980, China, Guangdong, Nanao Island, ZRC 1999.0470: C, ♂ 36.2 × 39.5 mm; D, ♀ 34.8 × 38.9 mm, specimen brushed." figureDoi="http://doi.org/10.5281/zenodo.8071279" httpUri="https://zenodo.org/record/8071279/files/figure.png" pageId="38" pageNumber="261">10C, D</figureCitation>
; 
<figureCitation id="135F13E79322B502CF18F8B1FDFBF873" box="[525,609,1905,1932]" captionStart="FIG" captionStartId="87.[132,143,1829,1847]" captionTargetBox="[347,1239,216,1787]" captionTargetId="figure-107@87.[790,881,1000,1790]" captionTargetPageId="87" captionText="FIG. 33. — Callosities in Dorippoidinae n. subfam. (Dorippoides) and Dorippinae n. stat. (Dorippe): A, callosity reduced to a thin calcified band: Dorippoides facchino (Herbst, 1785), ♂ 25.3 × 32.0 mm, Malaysia, Johore, Pontian, ZRC 1991.6672. B, no apparent callosity in male Dorippoides nudipes Manning &amp; Holthuis, 1986, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227. C, callosity as a thick, elongated bulge: Dorippe quadridens (Fabricius, 1793), ♂ 39.0 × 41.4 mm, Thailand, Pattani Province, ZRC 2003.0126. D-F, callosity as a hemicircular structure: close-up views from different angles: D, E. Dorippe sinica Chen, 1980. D, ♀ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; E, ♂ 36.6 × 38.6 mm, Japan, Kochi, Shikoku, SMF 57855. F, D. frascone (Herbst, 1785), ♂ 29.7 × 31.0 mm, Philippines, Exp. Panglao 2004, ZRC 2008.0076. G, H, callosity as an arched double cup: D. tenuipes Chen, 1980, South China Sea, ZRC 1999.0009: G, ovigerous ♀ 17.3 × 19.6 mm; H, ♂ 13.2 × 14.2 mm. Abbreviations: b, branchiostegite; c, callosity; cx2-cx5, P2-P5 coxae; f, sulcated part of coxa; l, onepiece lateroventral part of coxa; m, membrane; p, pleon; pr, process of retention of female pleonal somite 2; pl6, exposed pleurite 6; P2-P5, pereiopods 2-5; r, rim of carapace posterior margin; s, strip along posterior rim; 1, 2, pleonal somites 1, 2; t, bottom of callosity with special texture; 7, 8, thoracic sternites 7, 8." figureDoi="http://doi.org/10.5281/zenodo.8071333" httpUri="https://zenodo.org/record/8071333/files/figure.png" pageId="38" pageNumber="261">33D, E</figureCitation>
), quite distinct from the double callosity of 
<taxonomicName id="4C6474E19322B502CCA2F852FDB7F853" authorityName=": Chen" authorityYear="1980" box="[439,557,1938,1963]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="tenuipes">
<emphasis id="B910D3709322B502CCA2F852FDB7F853" box="[439,557,1938,1963]" italics="true" pageId="38" pageNumber="261">D. tenuipes</emphasis>
</taxonomicName>
(
<figureCitation id="135F13E79322B502CF2EF851FD0FF853" box="[571,661,1937,1963]" captionStart="FIG" captionStartId="26.[132,143,1214,1232]" captionTargetBox="[140,1456,215,1172]" captionTargetId="figure-502@26.[140,800,691,1172]" captionTargetPageId="26" captionText="FIG. 9. — Particular structures of some dorippids: A, spur-like process on ischium of P3 and sometimes P2: Dorippoides facchino (Herbst, 1785), postpubertal ♀ 19.0 × 22.0 mm,Pondicherry Bay,Maindron coll.1882,MNHN-IU-2018-5199 (= MNHN-B19817).B, erect spine on sternite 8: Neodorippe callida (Fabricius,1795), ovigerous ♀ 13.5 × 14.3 mm, Amoy, MNHN-IU-2021-8738 (= MNHN-B11170). C, D, callosity at base of P3: C, Dorippe sinica Chen, 1980, ♀ 34.8 × 38.9 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; D, Dorippe tenuipes Chen, 1980, ♂ 13.2 × 14.2 mm, South China Sea, ZRC 1999.0009 (the granules on P2 and P3 are not visible in the photograph). Abbreviations: c, callosity; cx2-cx5, coxae of P2-P5; e, erect spine; m, articulating membrane; p, process of sternite 8; pl6, exposed pleurite 6; p2-p5, pereiopods 2-5; r, rim; s, strip; sp, spur-like process; t, special texture area." figureDoi="http://doi.org/10.5281/zenodo.8071277" httpUri="https://zenodo.org/record/8071277/files/figure.png" pageId="38" pageNumber="261">Figs 9D</figureCitation>
; 
<figureCitation id="135F13E79322B502CFB4F851FD63F853" box="[673,761,1937,1963]" captionStart="FIG" captionStartId="87.[132,143,1829,1847]" captionTargetBox="[347,1239,216,1787]" captionTargetId="figure-107@87.[790,881,1000,1790]" captionTargetPageId="87" captionText="FIG. 33. — Callosities in Dorippoidinae n. subfam. (Dorippoides) and Dorippinae n. stat. (Dorippe): A, callosity reduced to a thin calcified band: Dorippoides facchino (Herbst, 1785), ♂ 25.3 × 32.0 mm, Malaysia, Johore, Pontian, ZRC 1991.6672. B, no apparent callosity in male Dorippoides nudipes Manning &amp; Holthuis, 1986, ♂ 16.2 × 19.4 mm, Iran, ZRC 2017.1227. C, callosity as a thick, elongated bulge: Dorippe quadridens (Fabricius, 1793), ♂ 39.0 × 41.4 mm, Thailand, Pattani Province, ZRC 2003.0126. D-F, callosity as a hemicircular structure: close-up views from different angles: D, E. Dorippe sinica Chen, 1980. D, ♀ 36.2 × 39.5 mm, China, Guangdong, Nanao Island, ZRC 1999.0470; E, ♂ 36.6 × 38.6 mm, Japan, Kochi, Shikoku, SMF 57855. F, D. frascone (Herbst, 1785), ♂ 29.7 × 31.0 mm, Philippines, Exp. Panglao 2004, ZRC 2008.0076. G, H, callosity as an arched double cup: D. tenuipes Chen, 1980, South China Sea, ZRC 1999.0009: G, ovigerous ♀ 17.3 × 19.6 mm; H, ♂ 13.2 × 14.2 mm. Abbreviations: b, branchiostegite; c, callosity; cx2-cx5, P2-P5 coxae; f, sulcated part of coxa; l, onepiece lateroventral part of coxa; m, membrane; p, pleon; pr, process of retention of female pleonal somite 2; pl6, exposed pleurite 6; P2-P5, pereiopods 2-5; r, rim of carapace posterior margin; s, strip along posterior rim; 1, 2, pleonal somites 1, 2; t, bottom of callosity with special texture; 7, 8, thoracic sternites 7, 8." figureDoi="http://doi.org/10.5281/zenodo.8071333" httpUri="https://zenodo.org/record/8071333/files/figure.png" pageId="38" pageNumber="261">33G, H</figureCitation>
). A direct comparison between 
<taxonomicName id="4C6474E19322B502CCA9F872FD87F833" authorityName=": Manning" authorityYear="1993" box="[444,541,1969,1995]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="B910D3709322B502CCA9F872FD87F833" box="[444,541,1969,1995]" italics="true" pageId="38" pageNumber="261">D. glabra</emphasis>
</taxonomicName>
and the other 
<taxonomicName id="4C6474E19322B502CFA6F872FC9CF833" authorityName="Fabricius" authorityYear="1798" box="[691,774,1970,1995]" class="Malacostraca" family="Dorippidae" genus="Dorippe" kingdom="Animalia" order="Decapoda" pageId="38" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="B910D3709322B502CFA6F872FC9CF833" box="[691,774,1970,1995]" italics="true" pageId="38" pageNumber="261">Dorippe</emphasis>
</taxonomicName>
would be most useful.
</paragraph>
</subSubSection>
</treatment>
</document>