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<mods:title id="E3F4697BBC51915A2B2E663E2742F361">First Potwarmus from the Miocene of Saudi Arabia and the early phylogeny of murines (Rodentia: Muroidea)</mods:title>
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<paragraph id="B6103566324BBE20FCC3F891FAEEF8E7" blockId="5.[894,1333,1812,1833]" box="[894,1333,1812,1833]" pageId="5" pageNumber="669">
<heading id="ED58820A324BBE20FCC3F891FAEEF8E7" box="[894,1333,1812,1833]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="669" reason="2">
<taxonomicName id="71AF4EE5324BBE20FC3EF891FC26F8E6" authority="ASIATICUS LINDSAY, 1988" authorityName="JAEGER" authorityYear="1977" box="[899,1021,1812,1833]" class="Mammalia" family="Muridae" genus="Dakkamys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="5" pageNumber="669" phylum="Chordata" rank="genus">
<emphasis id="84DBE974324BBE20FC3EF891FC26F8E6" box="[899,1021,1812,1833]" italics="true" pageId="5" pageNumber="669">DAKKAMYS</emphasis>
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<bibRefCitation id="D23E4897324BBE20FB36F891FAEEF8E7" author="Lindsay EH" box="[1163,1333,1812,1833]" pageId="5" pageNumber="669" pagination="95 - 154" refId="ref11384" refString="Lindsay EH. 1988. Cricetid rodents from the Siwalik deposits near Chinji Village. Part I: Megacricetodontinae, Myocricetodontinae, and Dendromurinae. Palaeovertebrata 18: 95 - 154." type="journal article" year="1988">LINDSAY, 1988</bibRefCitation>
</heading>
</paragraph>
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This species was erected by
<bibRefCitation id="D23E4897324BBE20FB3EF8BFFAE8F89E" author="Lindsay EH" box="[1155,1331,1850,1872]" pageId="5" pageNumber="669" pagination="95 - 154" refId="ref11384" refString="Lindsay EH. 1988. Cricetid rodents from the Siwalik deposits near Chinji Village. Part I: Megacricetodontinae, Myocricetodontinae, and Dendromurinae. Palaeovertebrata 18: 95 - 154." type="journal article" year="1988">Lindsay (1988)</bibRefCitation>
on the basis of seven isolated cheek teeth from the Middle Miocene localities YGSP 76 (Chinji Formation), YGSP 259 (lower Nagri Formation), and
<collectingCountry id="CEB875F63248BE23FD85FF61FD45FF34" box="[568,670,228,250]" name="Pakistan" pageId="6" pageNumber="670">Pakistan</collectingCountry>
Museum of Natural History locality JAL-
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north-east of Jalalpur village, Potwar Plateau,
<collectingCountry id="CEB875F63248BE23FD23FEA7FCDCFEF6" box="[670,775,290,312]" name="Pakistan" pageId="6" pageNumber="670">Pakistan</collectingCountry>
. Recently, additional material has also been recorded in
<collectingCountry id="CEB875F63248BE23FF7EFEDAFEF2FEBB" box="[195,297,351,373]" name="Pakistan" pageId="6" pageNumber="670">Pakistan</collectingCountry>
from the lower Nagri Formation at locality Y797 (
<bibRefCitation id="D23E48973248BE23FE9AFEF8FD94FE5D" author="Flynn LJ &amp; Morgan ME" box="[295,591,381,403]" pageId="6" pageNumber="670" pagination="1 - 10" refId="ref10987" refString="Flynn LJ, Morgan ME. 2005. An unusual diatomyid rodent from an infrequently sampled late Miocene interval in the Siwaliks of Pakistan. Palaeontologia Electronica 8 17 A: 1 - 10." type="journal article" year="2005">Flynn &amp; Morgan, 2005</bibRefCitation>
). The
<typeStatus id="69148BC43248BE23FD17FEF8FCD0FE5D" box="[682,779,381,403]" pageId="6" pageNumber="670" type="holotype">holotype</typeStatus>
(YGSP 19541) is an isolated left M1 from locality YGSP 76 (
<bibRefCitation id="D23E48973248BE23FEA3FE3EFE18FE1E" author="Lindsay EH" box="[286,451,443,465]" pageId="6" pageNumber="670" pagination="95 - 154" refId="ref11384" refString="Lindsay EH. 1988. Cricetid rodents from the Siwalik deposits near Chinji Village. Part I: Megacricetodontinae, Myocricetodontinae, and Dendromurinae. Palaeovertebrata 18: 95 - 154." type="journal article" year="1988">Lindsay, 1988</bibRefCitation>
: pl. 9, fig. a).
</paragraph>
</subSubSection>
<subSubSection id="FEB566ED3248BE23FF01FE5CFE4AFBBF" pageId="6" pageNumber="670" type="description">
<paragraph id="B61035663248BE23FF01FE5CFE4AFBBF" blockId="6.[164,779,198,1138]" pageId="6" pageNumber="670">
Although the morphology of the M1 of
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<emphasis id="84DBE9743248BE23FD33FE5CFCDDFE20" box="[654,774,473,494]" italics="true" pageId="6" pageNumber="670">Dakkamys</emphasis>
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<emphasis id="84DBE9743248BE23FF19FE7DFED2FDC3" box="[164,265,504,525]" italics="true" pageId="6" pageNumber="670">asiaticus</emphasis>
is close to that of AJ7, there are some differences. For instance, in the M1 of
<emphasis id="84DBE9743248BE23FDCAFD92FD4AFDE2" box="[631,657,535,556]" italics="true" pageId="6" pageNumber="670">D.</emphasis>
<emphasis id="84DBE9743248BE23FD22FD92FCDFFDE2" box="[671,772,535,556]" italics="true" pageId="6" pageNumber="670">asiaticus</emphasis>
, the enterostyle is connected to the posterior side of the protocone and it is much larger than the isolated enterostyle of AJ7. In addition, the M1 of
<emphasis id="84DBE9743248BE23FD3CFDF6FD47FD46" box="[641,668,627,648]" italics="true" pageId="6" pageNumber="670">D.</emphasis>
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has a strong and long anterior arm of the protocone, and lacks an anterior cingulum, whereas the Arabian specimen has only a relict of the anterior arm of the protocone, and has an anterior cingulum. The M2s of
<emphasis id="84DBE9743248BE23FF14FC89FF18FCEF" box="[169,195,780,801]" italics="true" pageId="6" pageNumber="670">D.</emphasis>
<emphasis id="84DBE9743248BE23FF6BFC89FEE0FCEF" box="[214,315,780,801]" italics="true" pageId="6" pageNumber="670">asiaticus</emphasis>
are distinct from AJ11 and AJ
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having a very large enterostyle connected to the posterior side of the protocone, whereas it is much smaller and isolated in the Arabian specimens. Furthermore, the M2s of
<emphasis id="84DBE9743248BE23FE75FC03FE38FC55" box="[456,483,902,923]" italics="true" pageId="6" pageNumber="670">D.</emphasis>
<emphasis id="84DBE9743248BE23FE50FC03FD89FC55" box="[493,594,902,923]" italics="true" pageId="6" pageNumber="670">asiaticus</emphasis>
lack the lingual anterior cingulum, and have long and well-developed anterior arms of the lingual cusps. AJ11 and AJ12 have a well-developed anterior lingual cingulum and lack the anterior arms of the lingual cusps. The m1 of
<emphasis id="84DBE9743248BE23FF14FB9AFF18FBFA" box="[169,195,1055,1076]" italics="true" pageId="6" pageNumber="670">D.</emphasis>
<emphasis id="84DBE9743248BE23FF68FB9AFEE1FBFA" box="[213,314,1055,1076]" italics="true" pageId="6" pageNumber="670">asiaticus</emphasis>
has a single-cusped anteroconid, and high and long anterior arms of the labial cusps, which are lacking in AJ13.
</paragraph>
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