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<mods:title id="29FD6FACEFFB1C42B6EDC699CFEAEEE6">The subgenus Empidideicus (Anomaloptilus) (Diptera: Mythicomyiidae) in Madagascar</mods:title>
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<mods:namePart id="9CBDCCFD9F3E7BE668CC8B8D98D7186B">Evenhuis, Neal L.</mods:namePart>
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<mods:affiliation id="26E285FCB61D6592A7E1C4F30A8AC0A0">J. Linsley Gressitt Center for Research in Entomology, Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii 96817 - 2704, USA neale @ bishopmuseum. org; https: // orcid. org / 0000 - 0002 - 1314 - 755 X</mods:affiliation>
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<mods:affiliation id="039D32175D5BAD86D674884B7C23D760">Plant Protection Research Department, Qazvin Agricultural and Natural Resources Research and Education Center, AREEO, Qazvin, Iran. bgaharaei @ yahoo. com; https: // orcid. org / 0000 - 0003 - 2490 - 156 X</mods:affiliation>
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<taxonomicName id="EEFDD1922618FFA4FF60FBD51D55FBAC" authority="Evenhuis &amp; Gharali" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[151,881,1086,1113]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="hauseri" status="sp. nov." subGenus="Anomaloptilus">
<emphasis id="1B8976032618FFA4FF60FBD51C4FFBAD" bold="true" box="[151,619,1086,1113]" italics="true" pageId="7" pageNumber="408">Empidideicus (Anomaloptilus) hauseri</emphasis>
Evenhuis &amp; Gharali
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,
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(
<figureCitation id="B1C6B6942618FFA4FF68FB881EC3FB89" box="[159,231,1123,1149]" captionStart="FIGURES 510" captionStartId="4.[151,264,1768,1792]" captionTargetBox="[151,1435,181,1745]" captionTargetId="figure-17@4.[151,1435,181,1745]" captionTargetPageId="4" captionText="FIGURES 510. Malagasy Empidideicus (Anomaloptilus), habitus, lateral view. 5, E. (A.) andriambahomananii, n. sp., male; 6, E. (A.) andriambahomananii, n. sp., female; 7, E. (A.) contrasticus, n. sp., male; 8, E. (A.) hauseri, n. sp., male; 9, E. (A.) irwini, n. sp., male; 10, E. (A.) nanus, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471746" httpUri="https://zenodo.org/record/7471746/files/figure.png" pageId="7" pageNumber="408">Figs 8</figureCitation>
,
<figureCitation id="B1C6B6942618FFA4FF03FB881F18FB89" box="[244,316,1123,1149]" captionStart="FIGURES 2533" captionStartId="13.[151,264,1878,1902]" captionTargetBox="[151,1435,273,1855]" captionTargetId="figure-65@13.[151,1435,273,1855]" captionTargetPageId="13" captionText="FIGURES 2533. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 25, E. (A.) andriambahomananii, n. sp., male; 26, E. (A.) contrasticus, n. sp., male; 27, E. (A.) contrasticus, n. sp., female; 28, E. (A.) hauseri, n. sp., male; 29, E. (A.) hauseri, n. sp., female; 30, E. (A.) irwini, n. sp., male; 31, E. (A.) irwini, n. sp., female; 32, E. (A.) nanus, n. sp., male; 33, E. (A.) nanus, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471758" httpUri="https://zenodo.org/record/7471758/files/figure.png" pageId="7" pageNumber="408">28, 29</figureCitation>
,
<figureCitation id="B1C6B6942618FFA4FEBFFB881F43FB89" box="[328,359,1123,1149]" captionStart="FIGURES 4047" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 4047. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="7" pageNumber="408">42</figureCitation>
,
<figureCitation id="B1C6B6942618FFA4FE85FB891F9EFB89" box="[370,442,1122,1149]" captionStart="FIGURES 5058" captionStartId="18.[151,264,1889,1913]" captionTargetBox="[239,1348,181,1865]" captionTargetId="figure-17@18.[239,1348,181,1865]" captionTargetPageId="18" captionText="FIGURES 5058. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 50, E. (A.) andriambahomananii, n. sp., female; 51, E. (A.) contrasticus, n. sp., male; 52, E. (A.) contrasticus, n. sp., female; 53, E. (A.) hauseri, n. sp., male; 54, E. (A.) hauseri, n. sp., female; 55, E. (A.) irwini, n. sp., male; 56, E. (A.) irwini, n. sp., female; 57, E. (A.) nanus, n. sp., male; 58, E. (A.) regius, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471772" httpUri="https://zenodo.org/record/7471772/files/figure.png" pageId="7" pageNumber="408">53, 54</figureCitation>
,
<figureCitation id="B1C6B6942618FFA4FE31FB881C29FB89" box="[454,525,1123,1149]" captionStart="FIGURES 6768" captionStartId="20.[152,270,1775,1799]" captionTargetBox="[161,1430,1091,1747]" captionTargetId="figure-59@20.[151,1436,1091,1751]" captionTargetPageId="20" captionText="FIGURES 6768. Empidideicus (Anomaloptilus) hauseri, n. sp., genitalia. 67, male, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, epiphallic membrane, dorsal view, E, gonostylus, detail; 68, female. Abbreviations: cer = cercus; mem= membrane associated with epandrium." figureDoi="http://doi.org/10.5281/zenodo.7471784" httpUri="https://zenodo.org/record/7471784/files/figure.png" pageId="7" pageNumber="408">67, 68</figureCitation>
)
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.
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♂ from
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:
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<specimenCount id="3FFB61982618FFA4FE0BFAFC1C05FAC5" box="[508,545,1303,1329]" pageId="7" pageNumber="408" type="female">2♀</specimenCount>
, same data except
<date id="5D438CD12618FFA4FD09FAFC1D88FAC5" box="[766,940,1303,1329]" pageId="7" pageNumber="408" value="2002-02-08" valueMax="2002-02-14" valueMin="2002-02-08">
<collectingDate id="4D0775392618FFA4FD09FAFC1D88FAC5" box="[766,940,1303,1329]" pageId="7" pageNumber="408" value="2002-02-08" valueMax="2002-02-14" valueMin="2002-02-08">814 Feb 2002</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FC4AFAFC1AAAFAC5" box="[957,1166,1303,1329]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019837</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FB53FAFC1C16FAA1" ID-GBIF-Occurrence="4003029366" collectingDate="2002-04-12" collectingDateMax="2002-04-23" collectingDateMin="2002-04-12" collectionCode="CASLOT" country="Madagascar" elevation="840" location="Zombitse National Park" municipality="Office" pageId="7" pageNumber="408" specimenCode="CASLOT 019818" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FB53FAFC1AECFAC5" box="[1188,1224,1303,1329]" pageId="7" pageNumber="408" type="female">2♀</specimenCount>
, same data except
<date id="5D438CD12618FFA4FF60FAD01F69FAA1" box="[151,333,1339,1365]" pageId="7" pageNumber="408" value="2002-04-12" valueMax="2002-04-23" valueMin="2002-04-12">
<collectingDate id="4D0775392618FFA4FF60FAD01F69FAA1" box="[151,333,1339,1365]" pageId="7" pageNumber="408" value="2002-04-12" valueMax="2002-04-23" valueMin="2002-04-12">1223 Apr 2002</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FEACFAD01C0EFAA1" box="[347,554,1339,1365]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019818</specimenCode>
)
</materialsCitation>
; 1♂,
<materialsCitation id="9995A04C2618FFA4FD99FAD01B03FAA1" ID-GBIF-Occurrence="4003029350" box="[622,1319,1339,1365]" collectingDate="2002-02-28" collectingDateMax="2002-03-06" collectingDateMin="2002-02-28" collectionCode="CASLOT" country="Madagascar" elevation="840" location="Zombitse National Park" municipality="Office" pageId="7" pageNumber="408" specimenCode="CASLOT 019822" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FD99FAD01CB6FAA1" box="[622,658,1339,1365]" pageId="7" pageNumber="408" type="female">4♀</specimenCount>
, same data except
<date id="5D438CD12618FFA4FC92FAD01A66FAA1" box="[869,1090,1339,1365]" pageId="7" pageNumber="408" value="2002-02-28" valueMax="2002-03-06" valueMin="2002-02-28">
<collectingDate id="4D0775392618FFA4FC92FAD01A66FAA1" box="[869,1090,1339,1365]" pageId="7" pageNumber="408" value="2002-02-28" valueMax="2002-03-06" valueMin="2002-02-28">28 Feb6 Mar 2002</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FBA7FAD01B3BFAA1" box="[1104,1311,1339,1365]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019822</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FAC5FAD01C8EFA8D" ID-GBIF-Occurrence="4003029332" collectingDate="2001-11-19" collectingDateMax="2001-11-26" collectingDateMin="2001-11-19" collectionCode="CASLOT" country="Madagascar" elevation="840" location="Zombitse National Park" municipality="Office" pageId="7" pageNumber="408" specimenCode="CASLOT 01934" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FAC5FAD01B71FAA1" box="[1330,1365,1339,1365]" pageId="7" pageNumber="408" type="female">1♀</specimenCount>
, same data except
<date id="5D438CD12618FFA4FEECFAB41FF1FA8D" box="[283,469,1375,1401]" pageId="7" pageNumber="408" value="2001-11-19" valueMax="2001-11-26" valueMin="2001-11-19">
<collectingDate id="4D0775392618FFA4FEECFAB41FF1FA8D" box="[283,469,1375,1401]" pageId="7" pageNumber="408" value="2001-11-19" valueMax="2001-11-26" valueMin="2001-11-19">1926 Nov 2001</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FE15FAB41C86FA8D" box="[482,674,1375,1401]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 01934</specimenCode>
)
</materialsCitation>
;;
<materialsCitation id="9995A04C2618FFA4FD36FAB41B4CFA8D" ID-GBIF-Occurrence="4003029330" box="[705,1384,1374,1401]" collectingDate="2002-01-26" collectingDateMax="2002-02-05" collectingDateMin="2002-01-26" collectionCode="CASLOT" country="Madagascar" elevation="840" location="Zombitse National Park" municipality="Office" pageId="7" pageNumber="408" specimenCode="CASLOT 019821" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FD36FAB41CC0FA8D" box="[705,740,1375,1401]" pageId="7" pageNumber="408" type="female">1♀</specimenCount>
, same data except
<date id="5D438CD12618FFA4FC43FAB41AA2FA8D" box="[948,1158,1374,1401]" pageId="7" pageNumber="408" value="2002-01-26" valueMax="2002-02-05" valueMin="2002-01-26">
<collectingDate id="4D0775392618FFA4FC43FAB41AA2FA8D" box="[948,1158,1374,1401]" pageId="7" pageNumber="408" value="2002-01-26" valueMax="2002-02-05" valueMin="2002-01-26">26 Jan5 Feb 2002</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FB64FAB41B44FA8D" box="[1171,1376,1375,1401]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019821</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FA85FA8B1D3AFA69" ID-GBIF-Occurrence="4003029364" collectingDate="2002-08-25" collectingDateMax="2002-09-03" collectingDateMin="2002-08-25" collectionCode="CASLOT" country="Madagascar" elevation="840" location="Zombitse National Park" municipality="Office" pageId="7" pageNumber="408" specimenCode="CASLOT 019816" specimenCount="1" typeStatus="paratype">
2♂, same data except
<date id="5D438CD12618FFA4FEAAFA691C1EFA69" box="[349,570,1410,1437]" pageId="7" pageNumber="408" value="2002-08-25" valueMax="2002-09-03" valueMin="2002-08-25">
<collectingDate id="4D0775392618FFA4FEAAFA691C1EFA69" box="[349,570,1410,1437]" pageId="7" pageNumber="408" value="2002-08-25" valueMax="2002-09-03" valueMin="2002-08-25">25 Aug3 Sep 2002</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FDBFFA681D32FA69" box="[584,790,1411,1437]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019816</specimenCode>
)
</materialsCitation>
; 9♂
<materialsCitation id="9995A04C2618FFA4FCB8FA6F1EDCFA35" ID-GBIF-Occurrence="4003029306" collectingDate="2002-02-28" collectingDateMax="2002-03-10" collectingDateMin="2002-02-28" collectionCode="CASLOT" country="Madagascar" elevation="840" location="Zombitse National Park" municipality="Office" pageId="7" pageNumber="408" specimenCode="CASLOT 019844" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FCB8FA6F1D46FA6B" box="[847,866,1412,1439]" pageId="7" pageNumber="408" type="female"></specimenCount>
, same data except
<date id="5D438CD12618FFA4FBC2FA681B04FA69" box="[1077,1312,1411,1437]" pageId="7" pageNumber="408" value="2002-02-28" valueMax="2002-03-10" valueMin="2002-02-28">
<collectingDate id="4D0775392618FFA4FBC2FA681B04FA69" box="[1077,1312,1411,1437]" pageId="7" pageNumber="408" value="2002-02-28" valueMax="2002-03-10" valueMin="2002-02-28">28 Feb10 Mar 2002</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FAD9FA681ED4FA35" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019844</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FEF1FA4C1D8BFA35" ID-GBIF-Occurrence="4003029316" box="[262,943,1447,1473]" collectingDate="2001-12-22" collectingDateMax="2001-12-29" collectingDateMin="2001-12-22" collectionCode="CASLOT" country="Madagascar" elevation="840" location="Zombitse National Park" municipality="Office" pageId="7" pageNumber="408" specimenCode="CASLOT 019819" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FEF1FA4C1F0DFA35" box="[262,297,1447,1473]" pageId="7" pageNumber="408" type="female">1♀</specimenCount>
, same data except
<date id="5D438CD12618FFA4FDF0FA4C1CE1FA35" box="[519,709,1447,1473]" pageId="7" pageNumber="408" value="2001-12-22" valueMax="2001-12-29" valueMin="2001-12-22">
<collectingDate id="4D0775392618FFA4FDF0FA4C1CE1FA35" box="[519,709,1447,1473]" pageId="7" pageNumber="408" value="2001-12-22" valueMax="2001-12-29" valueMin="2001-12-22">2229 Dec 2001</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FD21FA4C1D83FA35" box="[726,935,1447,1473]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019819</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FC4BFA4C1F90FA11" ID-GBIF-Occurrence="4003029318" collectingDate="2001-12-29" collectingDateMax="2002-01-05" collectingDateMin="2001-12-29" collectionCode="CASLOT" country="Madagascar" elevation="840" location="Zombitse National Park" municipality="Office" pageId="7" pageNumber="408" specimenCode="CASLOT 019832" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FC4BFA4C1DC4FA35" box="[956,992,1447,1473]" pageId="7" pageNumber="408" type="female">3♀</specimenCount>
, same data except
<date id="5D438CD12618FFA4FB4AFA4C1B76FA35" box="[1213,1362,1446,1473]" pageId="7" pageNumber="408" value="2001-12-29">
<collectingDate id="4D0775392618FFA4FB4AFA4C1B76FA35" box="[1213,1362,1446,1473]" pageId="7" pageNumber="408" value="2001-12-29">29 Dec 2001</collectingDate>
</date>
<date id="5D438CD12618FFA4FA96FA4D1EEBFA11" pageId="7" pageNumber="408" value="2002-01-05">
<collectingDate id="4D0775392618FFA4FA96FA4D1EEBFA11" pageId="7" pageNumber="408" value="2002-01-05">5 Jan 2002</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FF29FA201F88FA11" box="[222,428,1483,1509]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019832</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FE37FA271C3CF9FD" ID-GBIF-Occurrence="4003029363" collectingDate="2001-12-07" collectingDateMax="2001-12-14" collectingDateMin="2001-12-07" collectionCode="R" collectorName="Harin'Hala" country="Madagascar" latitude="-22.8405" location="Zombitse National Park" longitude="44.681859" pageId="7" pageNumber="408" specimenCode="CASLOT 019824" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
4♂,
<specimenCount id="3FFB61982618FFA4FE1AFA271C29FA13" box="[493,525,1484,1511]" pageId="7" pageNumber="408" type="female">1♀</specimenCount>
,
<location id="2C22FCCA2618FFA4FDECFA201D0DFA11" LSID="urn:lsid:plazi:treatment:A1541B072618FFAAFF60FBD51B60FB31:2C22FCCA2618FFA4FDECFA201D0DFA11" box="[539,809,1483,1509]" country="Madagascar" latitude="-22.8405" name="Zombitse National Park" pageId="7" pageNumber="408">Zombitse National Park</location>
, national road, 22°50.43S, 44·43.87E,
<date id="5D438CD12618FFA4FB1DFA201BBCFA11" box="[1258,1432,1483,1509]" pageId="7" pageNumber="408" value="2001-12-07" valueMax="2001-12-14" valueMin="2001-12-07">
<collectingDate id="4D0775392618FFA4FB1DFA201BBCFA11" box="[1258,1432,1483,1509]" pageId="7" pageNumber="408" value="2001-12-07" valueMax="2001-12-14" valueMin="2001-12-07">714 Dec 2001</collectingDate>
</date>
,
<collectionCode id="4FEC32D42618FFA4FF60FA041E89F9FD" box="[151,173,1519,1545]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="408">R</collectionCode>
.
<collectorName id="8408CFC72618FFA4FF40FA041F11F9FD" box="[183,309,1519,1545]" pageId="7" pageNumber="408">HarinHala</collectorName>
(
<specimenCode id="795B026A2618FFA4FEB5FA041C34F9FD" box="[322,528,1519,1545]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019824</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FDD5FA041A81F9FD" ID-GBIF-Occurrence="4003029303" box="[546,1189,1519,1545]" collectingDate="2001-11-09" collectingDateMax="2001-11-19" collectingDateMin="2001-11-09" collectionCode="CASLOT" collectorName="Harin'Hala" country="Madagascar" latitude="-22.8405" location="Zombitse National Park" pageId="7" pageNumber="408" specimenCode="CASLOT 019829" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FDD5FA041C61F9FD" box="[546,581,1519,1545]" pageId="7" pageNumber="408" type="female">1♀</specimenCount>
, same data except
<date id="5D438CD12618FFA4FCE2FA041DE6F9FD" box="[789,962,1519,1545]" pageId="7" pageNumber="408" value="2001-11-09" valueMax="2001-11-19" valueMin="2001-11-09">
<collectingDate id="4D0775392618FFA4FCE2FA041DE6F9FD" box="[789,962,1519,1545]" pageId="7" pageNumber="408" value="2001-11-09" valueMax="2001-11-19" valueMin="2001-11-09">919 Nov 2001</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FC38FA041AB9F9FD" box="[975,1181,1519,1545]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019829</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FB58FA041C0AF9D9" ID-GBIF-Occurrence="4003029311" collectingDate="2002-07-13" collectingDateMax="2002-07-25" collectingDateMin="2002-07-13" collectionCode="CASLOT" collectorName="Harin'Hala" country="Madagascar" latitude="-22.8405" location="Zombitse National Park" pageId="7" pageNumber="408" specimenCode="CASLOT 019833" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FB58FA041AF6F9FD" box="[1199,1234,1519,1545]" pageId="7" pageNumber="408" type="female">1♀</specimenCount>
, same data except
<date id="5D438CD12618FFA4FF60F9F91F62F9D9" box="[151,326,1554,1581]" pageId="7" pageNumber="408" value="2002-07-13" valueMax="2002-07-25" valueMin="2002-07-13">
<collectingDate id="4D0775392618FFA4FF60F9F91F62F9D9" box="[151,326,1554,1581]" pageId="7" pageNumber="408" value="2002-07-13" valueMax="2002-07-25" valueMin="2002-07-13">1325 Jul 2002</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FEA1F9F81C02F9D9" box="[342,550,1555,1581]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019833</specimenCode>
)
</materialsCitation>
; 4♂
<materialsCitation id="9995A04C2618FFA4FD97F9FF1AFCF9D9" ID-GBIF-Occurrence="4003029305" box="[608,1240,1555,1583]" collectingDate="2002-05-01" collectingDateMax="2002-05-09" collectingDateMin="2002-05-01" collectionCode="CASLOT" collectorName="Harin'Hala" country="Madagascar" latitude="-22.8405" location="Zombitse National Park" pageId="7" pageNumber="408" specimenCode="CASLOT 019846" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FD97F9FF1C57F9DB" box="[608,627,1556,1583]" pageId="7" pageNumber="408" type="female"></specimenCount>
, same data except
<date id="5D438CD12618FFA4FCBCF9F81DD5F9D9" box="[843,1009,1555,1581]" pageId="7" pageNumber="408" value="2002-05-01" valueMax="2002-05-09" valueMin="2002-05-01">
<collectingDate id="4D0775392618FFA4FCBCF9F81DD5F9D9" box="[843,1009,1555,1581]" pageId="7" pageNumber="408" value="2002-05-01" valueMax="2002-05-09" valueMin="2002-05-01">19 May 2002</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FBF6F9F81AF4F9D9" box="[1025,1232,1555,1581]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019846</specimenCode>
)
</materialsCitation>
; 4♂
<materialsCitation id="9995A04C2618FFA4FAFCF9FF1C4DF9A5" ID-GBIF-Occurrence="4003029343" collectingDate="2002-01-05" collectingDateMax="2002-01-12" collectingDateMin="2002-01-05" collectionCode="CASLOT" collectorName="Harin'Hala" country="Madagascar" latitude="-22.8405" location="Zombitse National Park" pageId="7" pageNumber="408" specimenCode="CASLOT 019820" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FAFCF9FF1B3AF9DB" box="[1291,1310,1556,1583]" pageId="7" pageNumber="408" type="female"></specimenCount>
, same data except
<date id="5D438CD12618FFA4FF11F9DD1FA3F9A5" box="[230,391,1590,1617]" pageId="7" pageNumber="408" value="2002-01-05" valueMax="2002-01-12" valueMin="2002-01-05">
<collectingDate id="4D0775392618FFA4FF11F9DD1FA3F9A5" box="[230,391,1590,1617]" pageId="7" pageNumber="408" value="2002-01-05" valueMax="2002-01-12" valueMin="2002-01-05">512 Jan 2002</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FE63F9DC1C45F9A5" box="[404,609,1591,1617]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019820</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FD84F9DC1B0DF9A5" ID-GBIF-Occurrence="4003029368" box="[627,1321,1590,1617]" collectingDate="2002-08-25" collectingDateMax="2002-09-03" collectingDateMin="2002-08-25" collectionCode="CASLOT" collectorName="Harin'Hala" country="Madagascar" latitude="-22.8405" location="Zombitse National Park" pageId="7" pageNumber="408" specimenCode="CASLOT 019816" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FD84F9DC1CB3F9A5" box="[627,663,1591,1617]" pageId="7" pageNumber="408" type="female">3♀</specimenCount>
, same data except:
<date id="5D438CD12618FFA4FC9AF9DD1A63F9A5" box="[877,1095,1590,1617]" pageId="7" pageNumber="408" value="2002-08-25" valueMax="2002-09-03" valueMin="2002-08-25">
<collectingDate id="4D0775392618FFA4FC9AF9DD1A63F9A5" box="[877,1095,1590,1617]" pageId="7" pageNumber="408" value="2002-08-25" valueMax="2002-09-03" valueMin="2002-08-25">25 Aug3 Sep 2002</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FBA3F9DC1B05F9A5" box="[1108,1313,1591,1617]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019816</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FAC4F9D31CE6F981" ID-GBIF-Occurrence="4003029317" collectingDate="2001-12-22" collectingDateMax="2001-12-29" collectingDateMin="2001-12-22" collectionCode="CASLOT" collectorName="Harin'Hala" country="Madagascar" latitude="-22.8405" location="Zombitse National Park" pageId="7" pageNumber="408" specimenCode="CASLOT 019826" specimenCount="1" typeStatus="paratype">
4♂, same data except
<date id="5D438CD12618FFA4FEE8F9B01FFFF981" box="[287,475,1627,1653]" pageId="7" pageNumber="408" value="2001-12-22" valueMax="2001-12-29" valueMin="2001-12-22">
<collectingDate id="4D0775392618FFA4FEE8F9B01FFFF981" box="[287,475,1627,1653]" pageId="7" pageNumber="408" value="2001-12-22" valueMax="2001-12-29" valueMin="2001-12-22">2229 Dec 2001</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FE1CF9B01C9EF981" box="[491,698,1627,1653]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019826</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FD38F9B71BBCF981" ID-GBIF-Occurrence="4003029331" box="[719,1432,1627,1655]" collectingDate="2001-11-26" collectingDateMax="2001-12-02" collectingDateMin="2001-11-26" collectionCode="CASLOT" collectorName="Harin'Hala" country="Madagascar" latitude="-22.8405" location="Zombitse National Park" pageId="7" pageNumber="408" specimenCode="CASLOT 019838" specimenCount="1" typeStatus="paratype">
4♂, same data except
<date id="5D438CD12618FFA4FC3CF9B01A95F981" box="[971,1201,1627,1653]" pageId="7" pageNumber="408" value="2001-11-26" valueMax="2001-12-02" valueMin="2001-11-26">
<collectingDate id="4D0775392618FFA4FC3CF9B01A95F981" box="[971,1201,1627,1653]" pageId="7" pageNumber="408" value="2001-11-26" valueMax="2001-12-02" valueMin="2001-11-26">26 Nov2 Dec 2001</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FB36F9B01BB4F981" box="[1217,1424,1627,1653]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 019838</specimenCode>
)
</materialsCitation>
; 1♂,
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<specimenCount id="3FFB61982618FFA4FF3FF9941ECFF96D" box="[200,235,1663,1689]" pageId="7" pageNumber="408" type="female">1♀</specimenCount>
, same data except
<date id="5D438CD12618FFA4FE4AF9951CB4F96D" box="[445,656,1662,1689]" pageId="7" pageNumber="408" value="2002-07-25" valueMax="2002-08-01" valueMin="2002-07-25">
<collectingDate id="4D0775392618FFA4FE4AF9951CB4F96D" box="[445,656,1662,1689]" pageId="7" pageNumber="408" value="2002-07-25" valueMax="2002-08-01" valueMin="2002-07-25">25 Jul1 Aug 2002</collectingDate>
</date>
(
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)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FC88F96B1BBCF949" ID-GBIF-Occurrence="4003029321" collectingDate="2003-07-27" collectingDateMax="2003-08-03" collectingDateMin="2003-07-27" collectionCode="R" collectorName="Harin'Hala, M. E &amp; Irwin" country="Madagascar" latitude="-22.903667" location="Mikea Forest" longLatPrecision="12" longitude="48.4755" municipality="Malaise" pageId="7" pageNumber="408" specimenCode="CASLOT 014402" specimenCount="1" typeStatus="paratype">
1♂,
<locationDeviation id="38F0B3332618FFA4FC58F9941AB3F96D" box="[943,1175,1663,1689]" location="Manambo" pageId="7" pageNumber="408">
<quantity id="EE0507F42618FFA4FC58F9941DCCF96D" box="[943,1000,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="408" unit="km" value="4.0">4 km</quantity>
NW Manambo
</locationDeviation>
,
<location id="2C22FCCA2618FFA4FB55F9941B1DF96D" LSID="urn:lsid:plazi:treatment:A1541B072618FFAAFF60FBD51B60FB31:2C22FCCA2618FFA4FB55F9941B1DF96D" box="[1186,1337,1663,1689]" country="Madagascar" latitude="-22.903667" longLatPrecision="12" longitude="48.4755" municipality="Malaise" name="Mikea Forest" pageId="7" pageNumber="408">Mikea Forest</location>
,
<collectingMunicipality id="C926306B2618FFA4FAB3F9941BB9F96D" box="[1348,1437,1663,1689]" pageId="7" pageNumber="408">Malaise</collectingMunicipality>
in deciduous forest,
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,
<geoCoordinate id="4CC9CCD62618FFA4FDFCF94F1CAFF94B" box="[523,651,1700,1727]" degrees="48" direction="east" minutes="28.53" orientation="longitude" pageId="7" pageNumber="408" precision="9" value="48.4755">48°28.53E</geoCoordinate>
,
<date id="5D438CD12618FFA4FD6EF9481D51F949" box="[665,885,1699,1725]" pageId="7" pageNumber="408" value="2003-07-27" valueMax="2003-08-03" valueMin="2003-07-27">
<collectingDate id="4D0775392618FFA4FD6EF9481D51F949" box="[665,885,1699,1725]" pageId="7" pageNumber="408" value="2003-07-27" valueMax="2003-08-03" valueMin="2003-07-27">27 Jul3 Aug 2003</collectingDate>
</date>
,
<collectionCode id="4FEC32D42618FFA4FC75F9481DBCF949" box="[898,920,1699,1725]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="408">R</collectionCode>
.
<collectorName id="8408CFC72618FFA4FC52F9481A43F949" box="[933,1127,1699,1725]" pageId="7" pageNumber="408">HarinHala, M.E</collectorName>
.
<collectorName id="8408CFC72618FFA4FB84F9481A94F949" box="[1139,1200,1699,1725]" pageId="7" pageNumber="408">Irwin</collectorName>
(
<specimenCode id="795B026A2618FFA4FB37F9481BB4F949" box="[1216,1424,1699,1725]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 014402</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FF60F92C1D01F915" ID-GBIF-Occurrence="4003029346" box="[151,805,1735,1761]" collectingDate="2002-07-02" collectingDateMax="2002-07-13" collectingDateMin="2002-07-02" collectionCode="CASLOT" collectorName="Harin'Hala, M. E &amp; Irwin" country="Madagascar" latitude="-22.903667" location="Mikea Forest" longLatPrecision="12" longitude="48.4755" municipality="Malaise" pageId="7" pageNumber="408" specimenCode="CASLOT 024698" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FF60F92C1E9EF915" box="[151,186,1735,1761]" pageId="7" pageNumber="408" type="female">1♀</specimenCount>
, same data except
<date id="5D438CD12618FFA4FE6FF92C1C18F915" box="[408,572,1735,1761]" pageId="7" pageNumber="408" value="2002-07-02" valueMax="2002-07-13" valueMin="2002-07-02">
<collectingDate id="4D0775392618FFA4FE6FF92C1C18F915" box="[408,572,1735,1761]" pageId="7" pageNumber="408" value="2002-07-02" valueMax="2002-07-13" valueMin="2002-07-02">213 Jul 2002</collectingDate>
</date>
(
<specimenCode id="795B026A2618FFA4FDBBF92C1D39F915" box="[588,797,1735,1761]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 024698</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FCC4F92C1EDCF8F1" ID-GBIF-Occurrence="4003029319" collectingDate="2002-07-02" collectingDateMax="2002-07-13" collectingDateMin="2002-07-02" collectionCode="CASLOT" collectorName="Jul &amp; Aug" country="Madagascar" latitude="-22.903667" location="Mikea Forest" longLatPrecision="12" longitude="48.4755" municipality="Malaise" pageId="7" pageNumber="408" specimenCode="CASLOT 024700" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FCC4F92C1D73F915" box="[819,855,1735,1761]" pageId="7" pageNumber="408" type="female">2♀</specimenCount>
, same data except 23
<collectorName id="8408CFC72618FFA4FBAEF92C1A5EF915" box="[1113,1146,1735,1761]" pageId="7" pageNumber="408">Jul</collectorName>
6
<collectorName id="8408CFC72618FFA4FB6AF92C1AE9F915" box="[1181,1229,1735,1761]" pageId="7" pageNumber="408">Aug</collectorName>
21002 (
<specimenCode id="795B026A2618FFA4FAD9F92C1ED4F8F1" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 024700</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="9995A04C2618FFA4FEF4F9001D3AF8DD" ID-GBIF-Occurrence="4003029329" collectingDate="2002-12-09" collectingDateMax="2002-12-15" collectingDateMin="2002-12-09" collectionCode="R" collectorName="M. Irwin &amp; F. Parker &amp; Harin'Hala" country="Madagascar" elevation="37" latitude="-25.587667" location="Cape Ste. Marie Special Reserve" longLatPrecision="12" longitude="45.163" pageId="7" pageNumber="408" specimenCode="CASLOT 024734" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="3FFB61982618FFA4FEF4F9001F03F8F1" box="[259,295,1771,1797]" pageId="7" pageNumber="408" type="female">2♀</specimenCount>
,
<location id="2C22FCCA2618FFA4FEC4F9001C80F8F1" LSID="urn:lsid:plazi:treatment:A1541B072618FFAAFF60FBD51B60FB31:2C22FCCA2618FFA4FEC4F9001C80F8F1" box="[307,676,1771,1797]" country="Madagascar" latitude="-25.587667" longLatPrecision="12" longitude="45.163" name="Cape Ste. Marie Special Reserve" pageId="7" pageNumber="408">Cape Ste. Marie Special Reserve</location>
,
<geoCoordinate id="4CC9CCD62618FFA4FD58F9071D0AF8F3" box="[687,814,1772,1799]" degrees="25" direction="south" minutes="35.26" orientation="latitude" pageId="7" pageNumber="408" precision="9" value="-25.587667">25°35.26S</geoCoordinate>
,
<geoCoordinate id="4CC9CCD62618FFA4FCCEF9071D9DF8F3" box="[825,953,1772,1799]" degrees="45" direction="east" minutes="09.78" orientation="longitude" pageId="7" pageNumber="408" precision="9" value="45.163">45°09.78E</geoCoordinate>
,
<quantity id="EE0507F42618FFA4FC33F9001DDAF8F1" box="[964,1022,1771,1797]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="408" unit="m" value="37.0">
<elevation id="A2D04D222618FFA4FC33F9001DDAF8F1" box="[964,1022,1771,1797]" metricMagnitude="1" metricUnit="m" metricValue="3.7" pageId="7" pageNumber="408" unit="m" value="37.0">37 m</elevation>
</quantity>
,
<date id="5D438CD12618FFA4FBFDF9011A93F8F1" box="[1034,1207,1770,1797]" pageId="7" pageNumber="408" value="2002-12-09" valueMax="2002-12-15" valueMin="2002-12-09">
<collectingDate id="4D0775392618FFA4FBFDF9011A93F8F1" box="[1034,1207,1770,1797]" pageId="7" pageNumber="408" value="2002-12-09" valueMax="2002-12-15" valueMin="2002-12-09">915 Dec 2002</collectingDate>
</date>
,
<collectorName id="8408CFC72618FFA4FB35F9001B03F8F1" box="[1218,1319,1771,1797]" pageId="7" pageNumber="408">M. Irwin</collectorName>
,
<collectorName id="8408CFC72618FFA4FAC5F9001BBCF8F1" box="[1330,1432,1771,1797]" pageId="7" pageNumber="408">F. Parker</collectorName>
,
<collectionCode id="4FEC32D42618FFA4FF60F8E41E89F8DD" box="[151,173,1807,1833]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="408">R</collectionCode>
.
<collectorName id="8408CFC72618FFA4FF4FF8E41F1DF8DD" box="[184,313,1807,1833]" pageId="7" pageNumber="408">HarinHala</collectorName>
,
<collectorName id="8408CFC72618FFA4FEB3F8E41FB9F8DD" box="[324,413,1807,1833]" pageId="7" pageNumber="408">Malaise</collectorName>
in spiny bush (
<specimenCode id="795B026A2618FFA4FDB0F8E41D31F8DD" box="[583,789,1807,1833]" collectionCode="CASLOT" pageId="7" pageNumber="408">CASLOT 024734</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="9995A04C2618FFA4FCDFF8E41DCAF8DD" ID-GBIF-Occurrence="4003029325" box="[808,1006,1807,1833]" collectionCode="CAS" country="Madagascar" pageId="7" pageNumber="408" specimenCount="1" typeStatus="holotype">
<typeStatus id="F64614B32618FFA4FCDFF8E41DB4F8DD" box="[808,912,1807,1833]" pageId="7" pageNumber="408" type="holotype">Holotype</typeStatus>
in
<collectionCode id="4FEC32D42618FFA4FC43F8E41DCAF8DD" box="[948,1006,1807,1833]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="7" pageNumber="408" type="Museum">CAS</collectionCode>
</materialsCitation>
.
<materialsCitation id="9995A04C2618FFA4FC0FF8E41B63F8DD" ID-GBIF-Occurrence="4003029309" box="[1016,1351,1807,1833]" collectionCode="CAS, BPBM" country="Madagascar" pageId="7" pageNumber="408" specimenCount="1" typeStatus="paratype">
<typeStatus id="F64614B32618FFA4FC0FF8E41A41F8DD" box="[1016,1125,1807,1833]" pageId="7" pageNumber="408" type="paratype">Paratypes</typeStatus>
in
<collectionCode id="4FEC32D42618FFA4FB7EF8E41A9BF8DD" box="[1161,1215,1807,1833]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="7" pageNumber="408" type="Museum">CAS</collectionCode>
and
<collectionCode id="4FEC32D42618FFA4FB01F8E41B63F8DD" box="[1270,1351,1807,1833]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="7" pageNumber="408" type="Museum">BPBM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A2618FFA4FF30F8D91C36F829" pageId="7" pageNumber="408" type="diagnosis">
<paragraph id="2942AA112618FFA4FF30F8D91C36F829" blockId="7.[151,1437,1195,2013]" pageId="7" pageNumber="408">
<emphasis id="1B8976032618FFA4FF30F8D91F1EF8B8" bold="true" box="[199,314,1842,1868]" pageId="7" pageNumber="408">Diagnosis</emphasis>
. Similar to
<emphasis id="1B8976032618FFA4FE3FF8D81FFDF8B8" box="[456,473,1843,1868]" italics="true" pageId="7" pageNumber="408">E</emphasis>
.
<taxonomicName id="EEFDD1922618FFA4FE1DF8D81D36F8B9" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[490,786,1843,1869]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="andriambahomananii" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032618FFA4FE03F8D81C21F8B9" box="[500,517,1843,1869]" italics="true" pageId="7" pageNumber="408">A</emphasis>
.)
<emphasis id="1B8976032618FFA4FDE8F8D81D36F8B9" box="[543,786,1843,1869]" italics="true" pageId="7" pageNumber="408">andriambahomananii</emphasis>
</taxonomicName>
<emphasis id="1B8976032618FFA4FCEBF8D81D7AF8B9" bold="true" box="[796,862,1843,1869]" pageId="7" pageNumber="408">
<taxonomicNameLabel id="00BACB782618FFA4FCEBF8D81D7AF8B9" box="[796,862,1843,1869]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by both having the second antennal flagellomere more than 1/3 the length of the first flagellomere and a closed anal cell, but can be easily separated from it by the all yellow scutellum (scutellum with brown basomedially in
<emphasis id="1B8976032618FFA4FC8DF8901DAFF860" box="[890,907,1915,1940]" italics="true" pageId="7" pageNumber="408">E</emphasis>
.
<taxonomicName id="EEFDD1922618FFA4FC6CF8901AE6F861" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[923,1218,1915,1941]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="andriambahomananii" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032618FFA4FC53F8901D91F861" box="[932,949,1915,1941]" italics="true" pageId="7" pageNumber="408">A</emphasis>
.)
<emphasis id="1B8976032618FFA4FC38F8901AE6F861" box="[975,1218,1915,1941]" italics="true" pageId="7" pageNumber="408">andriambahomananii</emphasis>
</taxonomicName>
<emphasis id="1B8976032618FFA4FB3DF8901B2FF861" bold="true" box="[1226,1291,1915,1941]" pageId="7" pageNumber="408">
<taxonomicNameLabel id="00BACB782618FFA4FB3DF8901B2FF861" box="[1226,1291,1915,1941]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
), the yellow prescutellar area (black
<emphasis id="1B8976032618FFA4FE50F8741FFAF84C" box="[423,478,1951,1976]" italics="true" pageId="7" pageNumber="408">in E.</emphasis>
<taxonomicName id="EEFDD1922618FFA4FE11F8741D28F84D" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[486,780,1951,1977]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="andriambahomananii" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032618FFA4FE18F8741C24F84D" box="[495,512,1951,1977]" italics="true" pageId="7" pageNumber="408">A</emphasis>
.)
<emphasis id="1B8976032618FFA4FDEEF8741D28F84D" box="[537,780,1951,1977]" italics="true" pageId="7" pageNumber="408">andriambahomananii</emphasis>
</taxonomicName>
<emphasis id="1B8976032618FFA4FCE3F8741D70F84D" bold="true" box="[788,852,1951,1977]" pageId="7" pageNumber="408">
<taxonomicNameLabel id="00BACB782618FFA4FCE3F8741D70F84D" box="[788,852,1951,1977]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
), and by the brown sternites (sternites yellow in
<emphasis id="1B8976032618FFA4FA73F8741BB1F84C" box="[1412,1429,1951,1976]" italics="true" pageId="7" pageNumber="408">E</emphasis>
.
<taxonomicName id="EEFDD1922618FFA4FF60F8281F98F829" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[151,444,1987,2013]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="andriambahomananii" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032618FFA4FF56F8281E96F829" box="[161,178,1987,2013]" italics="true" pageId="7" pageNumber="408">A</emphasis>
.)
<emphasis id="1B8976032618FFA4FF3EF8281F98F829" box="[201,444,1987,2013]" italics="true" pageId="7" pageNumber="408">andriambahomananii</emphasis>
</taxonomicName>
<emphasis id="1B8976032618FFA4FE34F8281C25F829" bold="true" box="[451,513,1987,2013]" pageId="7" pageNumber="408">
<taxonomicNameLabel id="00BACB782618FFA4FE34F8281C25F829" box="[451,513,1987,2013]" pageId="7" pageNumber="408" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A2617FFAAFF30FF7D1D8DFB89" lastPageId="9" lastPageNumber="410" pageId="8" pageNumber="409" type="description">
<paragraph id="2942AA112617FFABFF30FF7D1F7EFF45" blockId="8.[151,1437,150,717]" box="[199,346,150,177]" pageId="8" pageNumber="409">
<emphasis id="1B8976032617FFABFF30FF7D1F75FF44" bold="true" box="[199,337,150,176]" pageId="8" pageNumber="409">Description</emphasis>
:
</paragraph>
<paragraph id="2942AA112617FFABFF30FF501D5EFE59" blockId="8.[151,1437,150,717]" pageId="8" pageNumber="409">
<emphasis id="1B8976032617FFABFF30FF501F37FF20" bold="true" box="[199,275,186,213]" pageId="8" pageNumber="409">Male (</emphasis>
<figureCitation id="B1C6B6942617FFABFEE3FF501F73FF21" box="[276,343,187,213]" captionStart="FIGURES 510" captionStartId="4.[151,264,1768,1792]" captionTargetBox="[151,1435,181,1745]" captionTargetId="figure-17@4.[151,1435,181,1745]" captionTargetPageId="4" captionText="FIGURES 510. Malagasy Empidideicus (Anomaloptilus), habitus, lateral view. 5, E. (A.) andriambahomananii, n. sp., male; 6, E. (A.) andriambahomananii, n. sp., female; 7, E. (A.) contrasticus, n. sp., male; 8, E. (A.) hauseri, n. sp., male; 9, E. (A.) irwini, n. sp., male; 10, E. (A.) nanus, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471746" httpUri="https://zenodo.org/record/7471746/files/figure.png" pageId="8" pageNumber="409">Fig. 8</figureCitation>
). Length:
<quantity id="EE0507F42617FFABFE3EFF511C4BFF21" box="[457,623,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.625" metricValueMax="1.8" metricValueMin="1.45" pageId="8" pageNumber="409" unit="mm" value="1.625" valueMax="1.8" valueMin="1.45">1.451.80 mm</quantity>
. Wing: 1.502.00 mm.
<emphasis id="1B8976032617FFABFC83FF501D95FF21" box="[884,945,187,213]" italics="true" pageId="8" pageNumber="409">Head</emphasis>
: longer than high; eyes dichoptic, separated at vertex by 1.5× distance between lateral ocelli; occiput and postgena black; frons with upper half black, lower half white; face white with sparse minute black hairs, tip of oral margin white; antennae (cf.
<figureCitation id="B1C6B6942617FFABFB46FEE81B26FEE9" box="[1201,1282,259,285]" captionStart="FIGURES 2324" captionStartId="10.[151,264,1309,1333]" captionTargetBox="[151,1435,181,1284]" captionTargetId="figure-17@10.[151,1435,181,1285]" captionTargetPageId="10" captionText="FIGURES 2324. Malagasy Empidideicus (Anomaloptilus), antennae. 23, E. (A.) andriambahomananii, n. sp.; 24, E. (A.) nanus, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471754" httpUri="https://zenodo.org/record/7471754/files/figure.png" pageId="8" pageNumber="409">Fig. 23</figureCitation>
) black; scape short, subtrapezoidal; pedicel subconical, wider than long; first flagellomere lanceolate, length 2.5× width; second flagellomere one-half length of first flagellomere, with minute transparent apical style; mentum black; clypeus brown, length one-half oral margin; proboscis brown, length three-fourths head length; labrum sclerotized, stiff, pointed apically, length subequal to proboscis; palpus not evident.
</paragraph>
<paragraph id="2942AA112617FFABFF30FE5C1C6AFD5D" blockId="8.[151,1437,150,717]" pageId="8" pageNumber="409">
<emphasis id="1B8976032617FFABFF30FE5C1F3CFE25" box="[199,280,439,465]" italics="true" pageId="8" pageNumber="409">Thorax</emphasis>
(
<figureCitation id="B1C6B6942617FFABFED2FE5C1F53FE25" box="[293,375,439,465]" captionStart="FIGURES 2533" captionStartId="13.[151,264,1878,1902]" captionTargetBox="[151,1435,273,1855]" captionTargetId="figure-65@13.[151,1435,273,1855]" captionTargetPageId="13" captionText="FIGURES 2533. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 25, E. (A.) andriambahomananii, n. sp., male; 26, E. (A.) contrasticus, n. sp., male; 27, E. (A.) contrasticus, n. sp., female; 28, E. (A.) hauseri, n. sp., male; 29, E. (A.) hauseri, n. sp., female; 30, E. (A.) irwini, n. sp., male; 31, E. (A.) irwini, n. sp., female; 32, E. (A.) nanus, n. sp., male; 33, E. (A.) nanus, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471758" httpUri="https://zenodo.org/record/7471758/files/figure.png" pageId="8" pageNumber="409">Fig. 28</figureCitation>
). Mesonotum matte black with yellow markings in following areas: subquadrate prescutellar area extending anteriorly for to mid scutum as admedian vittae; interhumeral marks extending posteriorly as admedian vittae (vittae interrupted in middle of scutum), dorsolaterally from humeral callus to postalar callus and coalescing with prescutellar mark; scutellum yellow; scutum and dorsum of scutellum with short fine white hairs; pleura bare; pleura dark brown with pale yellow to white marking on following: posterodorsal corner of anepisternum, upper margin of katepisternum, upper three-fourths of anepimeron, laterally on propleuron, upper three-fourths of meron; halter stem grayish brown, knob white.
</paragraph>
<paragraph id="2942AA112617FFABFF30FD581BA8FD39" blockId="8.[151,1437,150,717]" box="[199,1420,690,717]" pageId="8" pageNumber="409">
<emphasis id="1B8976032617FFABFF30FD581EDEFD38" box="[199,250,691,716]" italics="true" pageId="8" pageNumber="409">Legs</emphasis>
. Coxae brown; rest of legs yellow, except basal three-fourths of hind femur and tarsal segments 25 brown.
</paragraph>
<caption id="7D82FA992617FFABFF60FA851A45FA5E" ID-DOI="http://doi.org/10.5281/zenodo.7471750" ID-Zenodo-Dep="7471750" httpUri="https://zenodo.org/record/7471750/files/figure.png" pageId="8" pageNumber="409" startId="8.[151,264,1390,1414]" targetBox="[151,1435,741,1367]" targetPageId="8">
<paragraph id="2942AA112617FFABFF60FA851A45FA5E" blockId="8.[151,1436,1390,1451]" pageId="8" pageNumber="409">
<emphasis id="1B8976032617FFABFF60FA851F75FA72" bold="true" box="[151,337,1390,1414]" pageId="8" pageNumber="409">FIGURES 1719</emphasis>
. Malagasy
<taxonomicName id="EEFDD1922617FFABFE3AFA851D22FA73" box="[461,774,1390,1415]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="subGenus" subGenus="Anomaloptilus">
<emphasis id="1B8976032617FFABFE3AFA851C73FA72" box="[461,599,1390,1414]" italics="true" pageId="8" pageNumber="409">Empidideicus</emphasis>
(
<emphasis id="1B8976032617FFABFD9EFA851CD9FA72" box="[617,765,1390,1414]" italics="true" pageId="8" pageNumber="409">Anomaloptilus</emphasis>
)
</taxonomicName>
, heads.
<emphasis id="1B8976032617FFABFCA9FA851D53FA72" bold="true" box="[862,887,1390,1414]" pageId="8" pageNumber="409">17</emphasis>
,
<emphasis id="1B8976032617FFABFC7FFA841DB3FA72" box="[904,919,1391,1414]" italics="true" pageId="8" pageNumber="409">E</emphasis>
.
<taxonomicName id="EEFDD1922617FFABFC50FA851A97FA72" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[935,1203,1390,1415]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="andriambahomananii" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032617FFABFC47FA841D9BFA72" box="[944,959,1391,1414]" italics="true" pageId="8" pageNumber="409">A</emphasis>
.)
<emphasis id="1B8976032617FFABFC20FA851A97FA72" box="[983,1203,1390,1414]" italics="true" pageId="8" pageNumber="409">andriambahomananii</emphasis>
</taxonomicName>
,
<emphasis id="1B8976032617FFABFB34FA851ADBFA72" bold="true" box="[1219,1279,1390,1414]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="00BACB782617FFABFB34FA851ADBFA72" box="[1219,1279,1390,1414]" pageId="8" pageNumber="409" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male, oblique view;
<emphasis id="1B8976032617FFABFF21FA791ECBFA5E" bold="true" box="[214,239,1426,1450]" pageId="8" pageNumber="409">18</emphasis>
,
<emphasis id="1B8976032617FFABFF0BFA781F2FFA5E" box="[252,267,1427,1450]" italics="true" pageId="8" pageNumber="409">E</emphasis>
.
<taxonomicName id="EEFDD1922617FFABFEEFFA791FA6FA5E" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[280,386,1426,1451]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="irwini" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032617FFABFED7FA781F0BFA5E" box="[288,303,1427,1450]" italics="true" pageId="8" pageNumber="409">A</emphasis>
.)
<emphasis id="1B8976032617FFABFEB2FA781FA6FA5E" box="[325,386,1427,1450]" italics="true" pageId="8" pageNumber="409">irwini</emphasis>
</taxonomicName>
,
<emphasis id="1B8976032617FFABFE79FA791FE3FA5E" bold="true" box="[398,455,1426,1450]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="00BACB782617FFABFE79FA791FE3FA5E" box="[398,455,1426,1450]" pageId="8" pageNumber="409" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female, dorsal view;
<emphasis id="1B8976032617FFABFD5FFA791CE5FA5E" bold="true" box="[680,705,1426,1450]" pageId="8" pageNumber="409">19</emphasis>
,
<emphasis id="1B8976032617FFABFD39FA781CF9FA5E" box="[718,733,1427,1450]" italics="true" pageId="8" pageNumber="409">E</emphasis>
.
<taxonomicName id="EEFDD1922617FFABFD1DFA791D71FA5E" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[746,853,1426,1451]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="rinhai" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032617FFABFD04FA781D26FA5E" box="[755,770,1427,1450]" italics="true" pageId="8" pageNumber="409">A</emphasis>
.)
<emphasis id="1B8976032617FFABFCE0FA791D71FA5E" box="[791,853,1426,1450]" italics="true" pageId="8" pageNumber="409">rinhai</emphasis>
</taxonomicName>
,
<emphasis id="1B8976032617FFABFC95FA791DBFFA5E" bold="true" box="[866,923,1426,1450]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="00BACB782617FFABFC95FA791DBFFA5E" box="[866,923,1426,1450]" pageId="8" pageNumber="409" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male, dorsal view.
</paragraph>
</caption>
<caption id="7D82FA992617FFABFF60F8461A76F81E" ID-DOI="http://doi.org/10.5281/zenodo.7471752" ID-Zenodo-Dep="7471752" httpUri="https://zenodo.org/record/7471752/files/figure.png" pageId="8" pageNumber="409" startId="8.[151,264,1965,1989]" targetBox="[151,1363,1475,1942]" targetPageId="8">
<paragraph id="2942AA112617FFABFF60F8461A76F81E" blockId="8.[151,1436,1965,2026]" pageId="8" pageNumber="409">
<emphasis id="1B8976032617FFABFF60F8461F6AF831" bold="true" box="[151,334,1965,1989]" pageId="8" pageNumber="409">FIGURES 2022</emphasis>
. Malagasy
<taxonomicName id="EEFDD1922617FFABFE31F8451CDAF832" box="[454,766,1966,1990]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="subGenus" subGenus="Anomaloptilus">
<emphasis id="1B8976032617FFABFE31F8451C74F832" box="[454,592,1966,1990]" italics="true" pageId="8" pageNumber="409">Empidideicus</emphasis>
(
<emphasis id="1B8976032617FFABFD97F8451CD0F832" box="[608,756,1966,1990]" italics="true" pageId="8" pageNumber="409">Anomaloptilus</emphasis>
)
</taxonomicName>
, heads.
<emphasis id="1B8976032617FFABFCA5F8461D4FF831" bold="true" box="[850,875,1965,1989]" pageId="8" pageNumber="409">20</emphasis>
,
<emphasis id="1B8976032617FFABFC8EF8451DACF831" box="[889,904,1966,1989]" italics="true" pageId="8" pageNumber="409">E</emphasis>
.
<taxonomicName id="EEFDD1922617FFABFC61F8451A64F831" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[918,1088,1966,1990]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="contrasticus" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032617FFABFC69F8451D89F831" box="[926,941,1966,1989]" italics="true" pageId="8" pageNumber="409">A</emphasis>
.)
<emphasis id="1B8976032617FFABFC33F8451A64F831" box="[964,1088,1966,1989]" italics="true" pageId="8" pageNumber="409">contrasticus</emphasis>
</taxonomicName>
,
<emphasis id="1B8976032617FFABFBB9F8451AACF832" bold="true" box="[1102,1160,1966,1990]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="00BACB782617FFABFBB9F8451AACF832" box="[1102,1160,1966,1990]" pageId="8" pageNumber="409" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male, frontal view;
<emphasis id="1B8976032617FFABFAA8F8461B5CF831" bold="true" box="[1375,1400,1965,1989]" pageId="8" pageNumber="409">21</emphasis>
,
<emphasis id="1B8976032617FFABFA71F8451BB1F831" box="[1414,1429,1966,1989]" italics="true" pageId="8" pageNumber="409">E</emphasis>
.
<taxonomicName id="EEFDD1922617FFABFF60F8391F25F81D" authority="Evenhuis &amp; Gharali, 2022" authorityName="Evenhuis &amp; Gharali" authorityYear="2022" box="[151,257,2002,2026]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="409" phylum="Arthropoda" rank="species" species="irwini" status="sp. nov." subGenus="Anomaloptilus">
(
<emphasis id="1B8976032617FFABFF57F8391E8BF81D" box="[160,175,2002,2025]" italics="true" pageId="8" pageNumber="409">A</emphasis>
.)
<emphasis id="1B8976032617FFABFF33F8391F25F81D" box="[196,257,2002,2025]" italics="true" pageId="8" pageNumber="409">irwini</emphasis>
</taxonomicName>
,
<emphasis id="1B8976032617FFABFEFAF8391F63F81E" bold="true" box="[269,327,2002,2026]" pageId="8" pageNumber="409">
<taxonomicNameLabel id="00BACB782617FFABFEFAF8391F63F81E" box="[269,327,2002,2026]" pageId="8" pageNumber="409" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male, oblique view;
<emphasis id="1B8976032617FFABFDD5F83A1C1FF81D" bold="true" box="[546,571,2001,2025]" pageId="8" pageNumber="409">22</emphasis>
,
<taxonomicName id="EEFDD1922617FFABFDBFF8391DA8F81E" authority="Evenhuis" authorityName="Evenhuis" box="[584,908,2002,2026]" class="Insecta" family="Bombyliidae" genus="Empidideicus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="403" phylum="Arthropoda" rank="species" species="rhinoclypeatus" subGenus="Anomaloptilus">
<emphasis id="1B8976032617FFABFDBFF8391C73F81D" box="[584,599,2002,2025]" italics="true" pageId="8" pageNumber="409">E</emphasis>
. (
<emphasis id="1B8976032617FFABFD9AF8391C58F81D" box="[621,636,2002,2025]" italics="true" pageId="8" pageNumber="409">A</emphasis>
.)
<emphasis id="1B8976032617FFABFD66F8391D03F81E" box="[657,807,2002,2026]" italics="true" pageId="8" pageNumber="409">rhinoclypeatus</emphasis>
Evenhuis
</taxonomicName>
, male, dorsal view.
</paragraph>
</caption>
<paragraph id="2942AA112616FFAAFF30FF7C1BB2FEB5" blockId="9.[151,1437,151,1221]" pageId="9" pageNumber="410">
<emphasis id="1B8976032616FFAAFF30FF7C1F25FF44" box="[199,257,151,176]" italics="true" pageId="9" pageNumber="410">Wing</emphasis>
(
<figureCitation id="B1C6B6942616FFAAFEF8FF7C1F45FF45" box="[271,353,151,177]" captionStart="FIGURES 4047" captionStartId="15.[151,264,1607,1631]" captionTargetBox="[170,1416,309,1582]" captionTargetId="figure-81@15.[170,1416,309,1584]" captionTargetPageId="15" captionText="FIGURES 4047. Malagasy Empidideicus (Anomaloptilus), wings. 40, E. (A.) andriambahomananii, n. sp.; 41, E. (A.) contrasticus, n. sp.; 42, E. (A.) hauseri, n. sp.; 43, E. (A.) irwini, n. sp.; 44, E. (A.) nanus, n. sp.; 45, E. (A.) regius, n. sp.; 46, E. (A.) rhinoclypeatus Evenhuis; 47, E. (A.) rinhai, n. sp." figureDoi="http://doi.org/10.5281/zenodo.7471764" httpUri="https://zenodo.org/record/7471764/files/figure.png" pageId="9" pageNumber="410">Fig. 42</figureCitation>
). Elongate, length 3× width; subhyaline with pale brownish tinge; veins brown; costa ends beyond R
<subScript id="B579A8542616FFAAFF5DFF261EE0FF2F" attach="left" box="[170,196,205,219]" fontSize="6" pageId="9" pageNumber="410">4+5</subScript>
about three-fourths way to M
<subScript id="B579A8542616FFAAFDD2FF261C09FF2F" attach="left" box="[549,557,205,219]" fontSize="6" pageId="9" pageNumber="410">1</subScript>
; vein Sc incomplete; Rs fading at connection with R
<subScript id="B579A8542616FFAAFB57FF261A8CFF2F" attach="left" box="[1184,1192,205,219]" fontSize="6" pageId="9" pageNumber="410">1</subScript>
; R
<subScript id="B579A8542616FFAAFB3AFF261AC3FF2F" attach="left" box="[1229,1255,205,219]" fontSize="6" pageId="9" pageNumber="410">4+5</subScript>
slightly curved upward at wing margin; vein M
<subScript id="B579A8542616FFAAFE01FF1A1FDAFF0B" attach="left" box="[502,510,241,255]" fontSize="6" pageId="9" pageNumber="410">1</subScript>
and M
<subScript id="B579A8542616FFAAFDBBFF1A1C70FF0B" attach="left" box="[588,596,241,255]" fontSize="6" pageId="9" pageNumber="410">2</subScript>
curved to wing margin; cell dm closed by apical crossvein, crossvein width ca. three-fourths width of crossvein r-m; A
<subScript id="B579A8542616FFAAFD86FEFE1C5DFED7" attach="left" box="[625,633,277,291]" fontSize="6" pageId="9" pageNumber="410">1</subScript>
thin, relatively straight to wing margin; anal cell closed at wing margin; fringe of hair on posterior margin of wing well shortest at wing tip becoming longer and more sparse toward base.
</paragraph>
<paragraph id="2942AA112616FFAAFF30FEA01FD3FE59" blockId="9.[151,1437,151,1221]" pageId="9" pageNumber="410">
<emphasis id="1B8976032616FFAAFF30FEA01F15FE91" box="[199,305,331,357]" italics="true" pageId="9" pageNumber="410">Abdomen</emphasis>
(
<figureCitation id="B1C6B6942616FFAAFEB3FEA01FBFFE91" box="[324,411,330,357]" captionStart="FIGURES 5058" captionStartId="18.[151,264,1889,1913]" captionTargetBox="[239,1348,181,1865]" captionTargetId="figure-17@18.[239,1348,181,1865]" captionTargetPageId="18" captionText="FIGURES 5058. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 50, E. (A.) andriambahomananii, n. sp., female; 51, E. (A.) contrasticus, n. sp., male; 52, E. (A.) contrasticus, n. sp., female; 53, E. (A.) hauseri, n. sp., male; 54, E. (A.) hauseri, n. sp., female; 55, E. (A.) irwini, n. sp., male; 56, E. (A.) irwini, n. sp., female; 57, E. (A.) nanus, n. sp., male; 58, E. (A.) regius, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471772" httpUri="https://zenodo.org/record/7471772/files/figure.png" pageId="9" pageNumber="410">Fig. 53</figureCitation>
). Dorsum dull brown medially, white laterally, brown at extreme lateral tips of tergites; medial brown on tergites becoming successively less on succeeding segments; sternites dark brown; tergites with sparsely scattered minute hairs.
</paragraph>
<paragraph id="2942AA112616FFAAFF30FE5C1C44FCE1" blockId="9.[151,1437,151,1221]" pageId="9" pageNumber="410">
<emphasis id="1B8976032616FFAAFF30FE5C1F15FE25" box="[199,305,439,465]" italics="true" pageId="9" pageNumber="410">Genitalia</emphasis>
(
<figureCitation id="B1C6B6942616FFAAFEB6FE5C1FB2FE25" box="[321,406,439,465]" captionStart="FIGURES 6768" captionStartId="20.[152,270,1775,1799]" captionTargetBox="[161,1430,1091,1747]" captionTargetId="figure-59@20.[151,1436,1091,1751]" captionTargetPageId="20" captionText="FIGURES 6768. Empidideicus (Anomaloptilus) hauseri, n. sp., genitalia. 67, male, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, epiphallic membrane, dorsal view, E, gonostylus, detail; 68, female. Abbreviations: cer = cercus; mem= membrane associated with epandrium." figureDoi="http://doi.org/10.5281/zenodo.7471784" httpUri="https://zenodo.org/record/7471784/files/figure.png" pageId="9" pageNumber="410">Fig. 67</figureCitation>
). Hypopygium shiny brown. In lateral view with epandrium subquadrate with thin, pointed apicoventral process (
<figureCitation id="B1C6B6942616FFAAFE7DFE301FD6FE01" box="[394,498,475,501]" captionStart="FIGURES 6768" captionStartId="20.[152,270,1775,1799]" captionTargetBox="[161,1430,1091,1747]" captionTargetId="figure-59@20.[151,1436,1091,1751]" captionTargetPageId="20" captionText="FIGURES 6768. Empidideicus (Anomaloptilus) hauseri, n. sp., genitalia. 67, male, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, epiphallic membrane, dorsal view, E, gonostylus, detail; 68, female. Abbreviations: cer = cercus; mem= membrane associated with epandrium." figureDoi="http://doi.org/10.5281/zenodo.7471784" httpUri="https://zenodo.org/record/7471784/files/figure.png" pageId="9" pageNumber="410">Fig. 67A</figureCitation>
); cerci large, slightly sclerotized, with large membranous sheath ventrally associated with parameral sheath (
<figureCitation id="B1C6B6942616FFAAFE6CFE141C04FDED" box="[411,544,511,537]" captionStart="FIGURES 6768" captionStartId="20.[152,270,1775,1799]" captionTargetBox="[161,1430,1091,1747]" captionTargetId="figure-59@20.[151,1436,1091,1751]" captionTargetPageId="20" captionText="FIGURES 6768. Empidideicus (Anomaloptilus) hauseri, n. sp., genitalia. 67, male, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, epiphallic membrane, dorsal view, E, gonostylus, detail; 68, female. Abbreviations: cer = cercus; mem= membrane associated with epandrium." figureDoi="http://doi.org/10.5281/zenodo.7471784" httpUri="https://zenodo.org/record/7471784/files/figure.png" pageId="9" pageNumber="410">Fig. 67C, D</figureCitation>
) (membrane in ventral view with lightly sclerotized medial rib with pair of arms curving outward toward to connect to epandrium ventrally); in ventral view with gonocoxites rounded, tapering to apex, with thin, basal sclerotization, with medial cleft shallow, broad (
<figureCitation id="B1C6B6942616FFAAFC4AFDAC1A01FD95" box="[957,1061,583,609]" captionStart="FIGURES 6768" captionStartId="20.[152,270,1775,1799]" captionTargetBox="[161,1430,1091,1747]" captionTargetId="figure-59@20.[151,1436,1091,1751]" captionTargetPageId="20" captionText="FIGURES 6768. Empidideicus (Anomaloptilus) hauseri, n. sp., genitalia. 67, male, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, epiphallic membrane, dorsal view, E, gonostylus, detail; 68, female. Abbreviations: cer = cercus; mem= membrane associated with epandrium." figureDoi="http://doi.org/10.5281/zenodo.7471784" httpUri="https://zenodo.org/record/7471784/files/figure.png" pageId="9" pageNumber="410">Fig. 67B</figureCitation>
); gonostylus consisting of apical pointed portion and larger, semi-trapezoidal, basal portion; lateral arms of parameral sheath extending well below gonocoxa, rounded, only slightly flared apically; aedeagal bulb very large, rounded, four-fifths length of lateral parameral arms (
<figureCitation id="B1C6B6942616FFAAFEA7FD581F92FD39" box="[336,438,691,717]" captionStart="FIGURES 6768" captionStartId="20.[152,270,1775,1799]" captionTargetBox="[161,1430,1091,1747]" captionTargetId="figure-59@20.[151,1436,1091,1751]" captionTargetPageId="20" captionText="FIGURES 6768. Empidideicus (Anomaloptilus) hauseri, n. sp., genitalia. 67, male, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, epiphallic membrane, dorsal view, E, gonostylus, detail; 68, female. Abbreviations: cer = cercus; mem= membrane associated with epandrium." figureDoi="http://doi.org/10.5281/zenodo.7471784" httpUri="https://zenodo.org/record/7471784/files/figure.png" pageId="9" pageNumber="410">Fig. 67C</figureCitation>
); aedeagal apodeme subequal in length to aedeagal bulb with long, pointed ventral portion; lateral rami thin; parameral sheath cylindrical distally, angled ventrally, truncate apically; thin aedeagus extending slightly beyond end of parameral sheath.
</paragraph>
<paragraph id="2942AA112616FFAAFF30FCF41D8DFB89" blockId="9.[151,1437,151,1221]" pageId="9" pageNumber="410">
<emphasis id="1B8976032616FFAAFF30FCF41F3AFCCD" bold="true" box="[199,286,799,825]" pageId="9" pageNumber="410">Female</emphasis>
: As in male except dark coloration of mesonotum and abdomen paler brown. Proboscis yellow; mentum yellowish white.Thorax (
<figureCitation id="B1C6B6942616FFAAFE47FCA81FDBFCA9" box="[432,511,835,861]" captionStart="FIGURES 2533" captionStartId="13.[151,264,1878,1902]" captionTargetBox="[151,1435,273,1855]" captionTargetId="figure-65@13.[151,1435,273,1855]" captionTargetPageId="13" captionText="FIGURES 2533. Malagasy Empidideicus (Anomaloptilus), thoraces, dorsal view. 25, E. (A.) andriambahomananii, n. sp., male; 26, E. (A.) contrasticus, n. sp., male; 27, E. (A.) contrasticus, n. sp., female; 28, E. (A.) hauseri, n. sp., male; 29, E. (A.) hauseri, n. sp., female; 30, E. (A.) irwini, n. sp., male; 31, E. (A.) irwini, n. sp., female; 32, E. (A.) nanus, n. sp., male; 33, E. (A.) nanus, n. sp., female." figureDoi="http://doi.org/10.5281/zenodo.7471758" httpUri="https://zenodo.org/record/7471758/files/figure.png" pageId="9" pageNumber="410">Fig. 29</figureCitation>
) with mesonotum with admedian vittae connected medially. Pleura all yellow except lower half of katepisternum and meron darker yellow to brown. Fore coxae yellow, mid and hind coxae brown; rest of legs all yellow. Abdomen (
<figureCitation id="B1C6B6942616FFAAFE29FC601C0BFC51" box="[478,559,906,933]" captionStart="FIGURES 5058" captionStartId="18.[151,264,1889,1913]" captionTargetBox="[239,1348,181,1865]" captionTargetId="figure-17@18.[239,1348,181,1865]" captionTargetPageId="18" captionText="FIGURES 5058. Malagasy Empidideicus (Anomaloptilus), abdomens, dorsal view. 50, E. (A.) andriambahomananii, n. sp., female; 51, E. (A.) contrasticus, n. sp., male; 52, E. (A.) contrasticus, n. sp., female; 53, E. (A.) hauseri, n. sp., male; 54, E. (A.) hauseri, n. sp., female; 55, E. (A.) irwini, n. sp., male; 56, E. (A.) irwini, n. sp., female; 57, E. (A.) nanus, n. sp., male; 58, E. (A.) regius, n. sp., male." figureDoi="http://doi.org/10.5281/zenodo.7471772" httpUri="https://zenodo.org/record/7471772/files/figure.png" pageId="9" pageNumber="410">Fig. 54</figureCitation>
) with tergites with small, paired spots in white lateral area. Genitalia (
<figureCitation id="B1C6B6942616FFAAFACCFC601BABFC51" box="[1339,1423,907,933]" captionStart="FIGURES 6768" captionStartId="20.[152,270,1775,1799]" captionTargetBox="[161,1430,1091,1747]" captionTargetId="figure-59@20.[151,1436,1091,1751]" captionTargetPageId="20" captionText="FIGURES 6768. Empidideicus (Anomaloptilus) hauseri, n. sp., genitalia. 67, male, A, lateral view, B, gonocoxae, ventral view, C, phallic complex, dorsal view, D, epiphallic membrane, dorsal view, E, gonostylus, detail; 68, female. Abbreviations: cer = cercus; mem= membrane associated with epandrium." figureDoi="http://doi.org/10.5281/zenodo.7471784" httpUri="https://zenodo.org/record/7471784/files/figure.png" pageId="9" pageNumber="410">Fig. 68</figureCitation>
): Genital fork U-shaped, with medial portion of “U” thin, evanescent in some specimens; with large opaque, round, black structure each side of furca, with narrow opaque black basal arm meeting at vaginal opening; spermatheca spheroid, sclerotized brown, slightly wider than high, without striations on basal half, shallow invagination apically, with striations except at apical margin; apical spermathecal duct ca. 3/4 length of spermatheca, thin, transparent; sperm pump 2× length of apical duct, transparent, not sclerotized apically or basally, slightly flared and rounded apically; basal duct confluent with sperm pump; common duct absent.
</paragraph>
</subSubSection>
<subSubSection id="61E7F99A2616FFAAFF30FB6C1B60FB31" pageId="9" pageNumber="410" type="etymology">
<paragraph id="2942AA112616FFAAFF30FB6C1B60FB31" blockId="9.[151,1437,151,1221]" pageId="9" pageNumber="410">
<emphasis id="1B8976032616FFAAFF30FB6C1F6CFB55" bold="true" box="[199,328,1159,1185]" pageId="9" pageNumber="410">Etymology</emphasis>
. The specific epithet honors Martin Hauser, collector of Malagasy mythicomyiids and for his generosity in supplying material of interesting flies to the senior author for study over the past many years.
</paragraph>
</subSubSection>
</treatment>
</document>