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<document id="AE2B790BD1CDBF153496905BC8BABE21" ID-DOI="10.11646/zootaxa.3949.2.8" ID-GBIF-Dataset="bec6d0a9-b118-471e-aa1e-42c0e6628069" ID-ISSN="1175-5326" ID-Zenodo-Dep="235827" ID-ZooBank="A5F993A6-5E36-47DD-BED5-A407BE159A18" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461045776942" checkinUser="plazi" docAuthor="Sinitshenkova, Nina D., Aristov, Daniil S., Wegierek, Piotr &amp; Żyła, Dagmara" docDate="2015" docId="5C0987DBFFCAFFA3FF377CE0497CFBAD" docLanguage="en" docName="zt03949p288.pdf" docOrigin="Zootaxa 3949 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Triassolitophlebia Sinitshenkova &amp; Aristov, gen. nov." docType="treatment" docVersion="7" lastPageNumber="283" masterDocId="A030FFA3FFC8FFA1FFA07C3D481FFFE2" masterDocTitle="New mayfly genera from the Middle Triassic of Poland and their evolutionary and paleogeographic implications (Ephemerida: Litophlebiidae, Vogesonymphidae)" masterLastPageNumber="288" masterPageNumber="281" pageNumber="283" updateTime="1698576785429" updateUser="plazi">
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<mods:title id="3DEC7524C0F637B445C3A2DC09FF8FA9">New mayfly genera from the Middle Triassic of Poland and their evolutionary and paleogeographic implications (Ephemerida: Litophlebiidae, Vogesonymphidae)</mods:title>
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<mods:namePart id="423BE3971775266D215EE886F3446EEE">Sinitshenkova, Nina D.</mods:namePart>
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<mods:namePart id="98E8E837A7DCB0CD76D45696B7D50165">Aristov, Daniil S.</mods:namePart>
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<mods:namePart id="2B26BCD5A094A248113EFADD22FD3F51">Wegierek, Piotr</mods:namePart>
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<mods:namePart id="60CF24642FD7C0B550A5EC24EB8CE8B4">Żyła, Dagmara</mods:namePart>
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<treatment id="5C0987DBFFCAFFA3FF377CE0497CFBAD" ID-DOI="http://doi.org/10.5281/zenodo.6098316" ID-GBIF-Taxon="119610503" ID-Zenodo-Dep="6098316" LSID="urn:lsid:plazi:treatment:5C0987DBFFCAFFA3FF377CE0497CFBAD" httpUri="http://treatment.plazi.org/id/5C0987DBFFCAFFA3FF377CE0497CFBAD" lastPageNumber="283" pageId="2" pageNumber="283">
<subSubSection id="9CBA6546FFCAFFA3FF377CE04B62FF15" box="[151,893,221,247]" pageId="2" pageNumber="283" type="nomenclature">
<paragraph id="D41F36CDFFCAFFA3FF377CE04B62FF15" blockId="2.[151,893,221,247]" box="[151,893,221,247]" pageId="2" pageNumber="283">
<heading id="8F5781A1FFCAFFA3FF377CE04B62FF15" bold="true" box="[151,893,221,247]" fontSize="11" level="1" pageId="2" pageNumber="283" reason="1">
<emphasis id="E6D4EADFFFCAFFA3FF377CE04B62FF15" bold="true" box="[151,893,221,247]" pageId="2" pageNumber="283">
Genus
<taxonomicName id="13A04D4EFFCAFFA3FF4D7CE04B1DFF15" authority="Sinitshenkova &amp; Aristov" authorityName="Sinitshenkova &amp; Aristov" box="[237,770,221,247]" class="Insecta" family="Litophlebiidae" genus="Triassolitophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="E6D4EADFFFCAFFA3FF4D7CE049D9FF15" bold="true" box="[237,454,221,247]" italics="true" pageId="2" pageNumber="283">Triassolitophlebia</emphasis>
Sinitshenkova &amp; Aristov
</taxonomicName>
,
<taxonomicNameLabel id="FDE757A4FFCAFFA3FCAF7CE04B62FF15" box="[783,893,221,247]" pageId="2" pageNumber="283" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9CBA6546FFCAFFA3FF377D184C31FED8" box="[151,1070,292,314]" pageId="2" pageNumber="283" type="materials_examined">
<paragraph id="D41F36CDFFCAFFA3FF377D184C31FED8" blockId="2.[151,1070,292,314]" box="[151,1070,292,314]" pageId="2" pageNumber="283">
<typeStatus id="0B1B886FFFCAFFA3FF377D1848D6FED8" box="[151,201,293,314]" pageId="2" pageNumber="283">Type</typeStatus>
species.
<taxonomicName id="13A04D4EFFCAFFA3FE857D194B21FED8" authority="Sinitshenkova &amp; Aristov" authorityName="Sinitshenkova &amp; Aristov" box="[293,830,292,314]" class="Insecta" family="Litophlebiidae" genus="Triassolitophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="palegica" status="sp. nov.">
<emphasis id="E6D4EADFFFCAFFA3FE857D194A24FED8" box="[293,571,292,314]" italics="true" pageId="2" pageNumber="283">Triassolitophlebia palegica</emphasis>
Sinitshenkova &amp; Aristov
</taxonomicName>
,
<taxonomicNameLabel id="FDE757A4FFCAFFA3FCE87D184B86FED8" box="[840,921,293,314]" pageId="2" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="9CBA6546FFCAFFA3FF377D5B4B15FBE5" pageId="2" pageNumber="283" type="diagnosis">
<paragraph id="D41F36CDFFCAFFA3FF377D5B4D6DFE09" blockId="2.[151,1437,358,1103]" pageId="2" pageNumber="283">
<emphasis id="E6D4EADFFFCAFFA3FF377D5B490EFE9D" bold="true" box="[151,273,358,383]" pageId="2" pageNumber="283">Diagnosis.</emphasis>
The forewing is 3.2 times as long as wide, the apex widely rounded. SC does not reach the wing apex. In the RS system, there are not fewer than five longitudinal veins, and RS is not fused with MA stem at the wing base. MA branches in the basal half of the wing, and MP divides at the very wing base. CuA is long and reaches the wing margin distally of MA branching; CuP is long and reaches the wing margin in the basal third of the wing.
</paragraph>
<paragraph id="D41F36CDFFCAFFA3FF677DCB4B15FBE5" blockId="2.[151,1437,358,1103]" pageId="2" pageNumber="283">
<emphasis id="E6D4EADFFFCAFFA3FF677DCB497DFDED" bold="true" box="[199,354,502,527]" pageId="2" pageNumber="283">Comparison.</emphasis>
The new genus differs significantly from
<taxonomicName id="13A04D4EFFCAFFA3FC9D7DCB4C5EFDED" authority="Riek, 1976" authorityName="Riek" authorityYear="1976" box="[829,1089,502,527]" class="Insecta" family="Litophlebiidae" genus="Litophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="E6D4EADFFFCAFFA3FC9D7DCB4BA1FDED" box="[829,958,502,527]" italics="true" pageId="2" pageNumber="283">Litophlebia</emphasis>
<bibRefCitation id="B0314B3CFFCAFFA3FC657DCB4C5EFDED" author="Riek" box="[965,1089,502,527]" pageId="2" pageNumber="283" refString="Riek, E. F. (1976) An unusual mayfly (Insecta: Ephemeroptera) from the Triassic of South Africa. Palaeontologica Africana, 19, 149 - 151." type="journal article" year="1976">Riek, 1976</bibRefCitation>
</taxonomicName>
by the long CuA and CuP and by a shorter SC. In the original description of
<taxonomicName id="13A04D4EFFCAFFA3FD347E274B0AFDD1" box="[660,789,538,563]" class="Insecta" family="Litophlebiidae" genus="Litophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="E6D4EADFFFCAFFA3FD347E274B0AFDD1" box="[660,789,538,563]" italics="true" pageId="2" pageNumber="283">Litophlebia</emphasis>
</taxonomicName>
”RS appearing to arise as a forward branch of MA, in fact on the photo (
<bibRefCitation id="B0314B3CFFCAFFA3FE947E0349E3FDB5" author="Riek" box="[308,508,574,599]" pageId="2" pageNumber="283" refString="Riek, E. F. (1976) An unusual mayfly (Insecta: Ephemeroptera) from the Triassic of South Africa. Palaeontologica Africana, 19, 149 - 151." type="journal article" year="1976">Riek 1976: Fig. 3</bibRefCitation>
) it can be distinctly seen that the basal part of RS stem comes closely to the basal part of MA stem and runs near to it for some distance. Perhaps they are not fused, because at the very wing base the RS base is seen as a fine separate vein. For the
<taxonomicName id="13A04D4EFFCAFFA3FD1E7EBB4B76FD7D" box="[702,873,646,671]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="order">Ephemeroptera</taxonomicName>
the separate arising of RS from the wing base is characteristic. Some features of the new mayfly venation have a formal similarity to that of
<taxonomicName id="13A04D4EFFCAFFA3FAAE7E974BDEFD05" authority="Sinitshenkova &amp; Papier, 2005 (Sinitshenkova et al. 2005)" authorityName="Sinitshenkova &amp; Papier, 2005 (Sinitshenkova et al." authorityYear="2005" class="Insecta" family="Misthodotidae" genus="Triassodotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="vogesiacus">
<emphasis id="E6D4EADFFFCAFFA3FAAE7E97490DFD04" italics="true" pageId="2" pageNumber="283">Triassodotes vogesiacus</emphasis>
<bibRefCitation id="B0314B3CFFCAFFA3FEBD7EF34A66FD05" author="Sinitshenkova" box="[285,633,718,743]" pageId="2" pageNumber="283" refString="Sinitshenkova, N. D., Marchal-Papier, F., Grauvogel-Stamm, L. &amp; Gall, J. C. (2005) The Ephemeridea (Insecta) from the Gres Voltzia (early Middle Triassic) of the Vosges (NE France). Palaeontologische Zeitschrift, 79 (3), 377 - 397. http: // dx. doi. org / 10.1007 / BF 02991930" type="journal article" year="2005">Sinitshenkova &amp; Papier, 2005</bibRefCitation>
(
<bibRefCitation id="B0314B3CFFCAFFA3FD2B7EF34BA6FD05" author="Sinitshenkova" box="[651,953,718,743]" pageId="2" pageNumber="283" refString="Sinitshenkova, N. D., Marchal-Papier, F., Grauvogel-Stamm, L. &amp; Gall, J. C. (2005) The Ephemeridea (Insecta) from the Gres Voltzia (early Middle Triassic) of the Vosges (NE France). Palaeontologische Zeitschrift, 79 (3), 377 - 397. http: // dx. doi. org / 10.1007 / BF 02991930" type="journal article" year="2005">
Sinitshenkova
<emphasis id="E6D4EADFFFCAFFA3FC987EF24B6BFD05" box="[824,884,718,743]" italics="true" pageId="2" pageNumber="283">et al.</emphasis>
2005
</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="13A04D4EFFCAFFA3FBA07EF2494CFCE9" authority="Krzeminski &amp; Lombardo, 2001" authorityName="Krzeminski &amp; Lombardo" authorityYear="2001" class="Insecta" family="Tintorinidae" genus="Tintorina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="meridensis">
<emphasis id="E6D4EADFFFCAFFA3FBA07EF24CF2FD05" box="[1024,1261,718,743]" italics="true" pageId="2" pageNumber="283">Tintorina meridensis</emphasis>
<bibRefCitation id="B0314B3CFFCAFFA3FB587EF3494CFCE9" author="Krzeminski" pageId="2" pageNumber="283" refString="Krzeminski, W. &amp; Lombardo, C. (2001) New fossil Ephemeroptera and Coleoptera from the Ladinian (Middle Triassic) of Canton Ticino (Switzerland). Revista Italiana di Paleontologia e Stratigrafia, 107 (1), 69 - 78." type="journal article" year="2001">Krzeminski &amp; Lombardo, 2001</bibRefCitation>
</taxonomicName>
of the family
<taxonomicName id="13A04D4EFFCAFFA3FE567ECF4BE3FCE9" authority="Krzeminski &amp; Lombardo 2001" authorityName="Krzeminski &amp; Lombardo" authorityYear="2001" box="[502,1020,754,779]" class="Insecta" family="Tintorinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="family">
Tintorinidae (
<bibRefCitation id="B0314B3CFFCAFFA3FD307ECF4BECFCE9" author="Krzeminski" box="[656,1011,754,779]" pageId="2" pageNumber="283" refString="Krzeminski, W. &amp; Lombardo, C. (2001) New fossil Ephemeroptera and Coleoptera from the Ladinian (Middle Triassic) of Canton Ticino (Switzerland). Revista Italiana di Paleontologia e Stratigrafia, 107 (1), 69 - 78." type="journal article" year="2001">Krzeminski &amp; Lombardo 2001</bibRefCitation>
)
</taxonomicName>
. These features include the short SC and MP branching at the wing base. These are the proper characters that clearly distinguish the Triassic
<taxonomicName id="13A04D4EFFCAFFA3FF377F07493AFCB1" box="[151,293,826,851]" class="Insecta" family="Misthodotidae" genus="Triassodotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="E6D4EADFFFCAFFA3FF377F07493AFCB1" box="[151,293,826,851]" italics="true" pageId="2" pageNumber="283">Triassodotes</emphasis>
</taxonomicName>
from the Permian
<taxonomicName id="13A04D4EFFCAFFA3FE597F074B5BFCB1" authority="Tshernova, 1965" authorityName="Tshernova" authorityYear="1965" box="[505,836,826,851]" class="Insecta" family="Misthodotidae" genus="Misthodotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="E6D4EADFFFCAFFA3FE597F074A9DFCB1" box="[505,642,826,851]" italics="true" pageId="2" pageNumber="283">Misthodotes</emphasis>
<bibRefCitation id="B0314B3CFFCAFFA3FD297F074B5BFCB1" author="Tshernova" box="[649,836,826,851]" pageId="2" pageNumber="283" refString="Tshernova, O. A. (1965) Some fossil mayflies (Ephemeroptera, Misthodotidae) from Permian beds of the Ural. Entomologicheskoe Obozrenie, 44 (2), 353 - 361. [in Russian, English translation: Entomological Review, 44 (1), 202 - 207]" type="journal article" year="1965">Tshernova, 1965</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="B0314B3CFFCAFFA3FCF37F074C13FCB1" author="Tshernova" box="[851,1036,826,851]" pageId="2" pageNumber="283" refString="Tshernova, O. A. (1965) Some fossil mayflies (Ephemeroptera, Misthodotidae) from Permian beds of the Ural. Entomologicheskoe Obozrenie, 44 (2), 353 - 361. [in Russian, English translation: Entomological Review, 44 (1), 202 - 207]" type="journal article" year="1965">Tshernova 1965</bibRefCitation>
;
<bibRefCitation id="B0314B3CFFCAFFA3FBB87F074D2BFCB1" author="Kinzelbach" box="[1048,1332,826,851]" pageId="2" pageNumber="283" refString="Kinzelbach, R. &amp; Lutz, H. (1984) Eine neue Eintagsfliege Misthodotes stapfi n. sp. aus dem Rotliegenden des Nahe-Gebietes (Ephemeroptera: Permoplectoptera: Misthodotidae). Palaontologische Zeitschrift, 58 (3 / 4), 247 - 253. http: // dx. doi. org / 10.1007 / BF 02986063" type="journal article" year="1984">Kinzelbach &amp; Lutz 1984</bibRefCitation>
). In spite of this similarity, the new mayfly could not be referred to the
<taxonomicName id="13A04D4EFFCAFFA3FCE17F634BFCFC95" box="[833,995,862,887]" class="Insecta" family="Misthodotidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="family">Misthodotidae</taxonomicName>
because of the sharp corrugation that is not characteristic for this family but rather typical for the
<taxonomicName id="13A04D4EFFCAFFA3FCBF7FBF4BD9FC79" box="[799,966,898,923]" class="Insecta" family="Litophlebiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="family">Litophlebiidae</taxonomicName>
. The representatives of the
<taxonomicName id="13A04D4EFFCAFFA3FB5A7FBF4D83FC79" box="[1274,1436,898,923]" class="Insecta" family="Misthodotidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="family">Misthodotidae</taxonomicName>
have wider wings; they are flattened to some extent and never have a sharp corrugation. This feature is also the main difference between the
<taxonomicName id="13A04D4EFFCAFFA3FE4D7FF74A8EFC01" box="[493,657,970,995]" class="Insecta" family="Litophlebiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="family">Litophlebiidae</taxonomicName>
and the
<taxonomicName id="13A04D4EFFCAFFA3FD5D7FF74B95FC01" box="[765,906,970,995]" class="Insecta" family="Tintorinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="family">Tintorinidae</taxonomicName>
. Moreover, the wings of the
<taxonomicName id="13A04D4EFFCAFFA3FB457FF74D70FC01" box="[1253,1391,970,995]" class="Insecta" family="Tintorinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="family">Tintorinidae</taxonomicName>
are longer and narrower; they are 3.7 times as long as wide.
</paragraph>
</subSubSection>
<subSubSection id="9CBA6546FFCAFFA3FF67782F497CFBAD" pageId="2" pageNumber="283" type="etymology">
<paragraph id="D41F36CDFFCAFFA3FF67782F497CFBAD" blockId="2.[151,1437,358,1103]" pageId="2" pageNumber="283">
<emphasis id="E6D4EADFFFCAFFA3FF67782F4951FBC9" bold="true" box="[199,334,1042,1067]" pageId="2" pageNumber="283">Etymology.</emphasis>
The name is a combination of the period Triassic and
<taxonomicName id="13A04D4EFFCAFFA3FC62782F4C5CFBC9" box="[962,1091,1042,1067]" class="Insecta" family="Litophlebiidae" genus="Litophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="E6D4EADFFFCAFFA3FC62782F4C5CFBC9" box="[962,1091,1042,1067]" italics="true" pageId="2" pageNumber="283">Litophlebia</emphasis>
</taxonomicName>
the
<typeStatus id="0B1B886FFFCAFFA3FB27782E4CA8FBC9" box="[1159,1207,1043,1067]" pageId="2" pageNumber="283">type</typeStatus>
genus of the family. Gender: feminine.
</paragraph>
</subSubSection>
</treatment>
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