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<document id="7D08E22721051C3C08BB5410CC3F8C49" ID-DOI="10.3897/zookeys.1223.139346" ID-ZooBank="451749E9-009E-43E9-A6F1-892035BDF1B0" ID-publisher-id="139346" URI-arpha="6EA2BAFE-82AB-5570-95B8-EE1F5FA60D46" URI-zoobank="http://zoobank.org/451749E9-009E-43E9-A6F1-892035BDF1B0" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1736349736103" checkinUser="pensoft" docAuthor="Wesener, Thomas, Akkari, Nesrine &amp; Golovatch, Sergei I." docDate="2025" docId="B77BE40B1CC852998AFAF927C052F425" docLanguage="en" docName="ZooKeys 1223: 185-220" docOrigin="ZooKeys 1223" docSource="https://zookeys.pensoft.net/article/139346/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Dalodesmus kompantsevi Wesener, Akkari &amp; Golovatch, 2025, sp. nov." docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="220" masterDocId="6EA2BAFE82AB557095B8EE1F5FA60D46" masterDocTitle="Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida)" masterLastPageNumber="220" masterPageNumber="185" pageNumber="185" updateTime="1736355520980" updateUser="admin">
<mods:mods id="8D46145BD9BE5C9F77664D8ABFBB7B24" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E9FEB75EC31587C98D671323E885DFDB">
<mods:title id="759D1E495F2AA1E97CCA4A31B27DBE14">Revision of the millipede family Dalodesmidae in Madagascar, with descriptions of two new Malagasy species of Dalodesmus Cook, 1896 (Diplopoda, Polydesmida)</mods:title>
</mods:titleInfo>
<mods:name id="5C86DE946CDCEC510AFB1BCA220206B8" type="personal">
<mods:role id="3D545A39A88AD9A847699836CC9BE45F">
<mods:roleTerm id="FD31EE751350BB595DB4B81E77760FEE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0D3D0E35FC08AF6465716925C92ACC0F">Wesener, Thomas</mods:namePart>
<mods:nameIdentifier id="D22EA66F6B855EE181E45ECD06488388" type="ORCID">0000-0002-2028-3541</mods:nameIdentifier>
<mods:affiliation id="1398E9FF83DF0EDCDC2ED3D962B3CBAB">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, D- 53113 Bonn, Germany</mods:affiliation>
</mods:name>
<mods:name id="D4A009C8D8B95E5002D2038E0BD0B951" type="personal">
<mods:role id="BBFB6D25C4DCCD79019FE589B2000C27">
<mods:roleTerm id="3D70B5F34791D3B0154EA20401F9B136">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B543AC9028FDA97366047C530E182EF">Akkari, Nesrine</mods:namePart>
<mods:nameIdentifier id="F2A9A90F8773EE42D72088137857FA31" type="ORCID">0000-0001-5019-4833</mods:nameIdentifier>
<mods:affiliation id="28DD301E421053A218C7A06621C2B8FB">Third Zoological Department, Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
</mods:name>
<mods:name id="3E52ED9AF3087F666C2424AF8A8F74D0" type="personal">
<mods:role id="5D225298DC7C19070A4F10745D36CBFE">
<mods:roleTerm id="37D6E06178DBCED2F23B60181533FE4A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0FFEEAD0D200C0FA20457701C65656AD">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="88E0059C552A332784F8C28E09F8512E">Institute of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow 119071, Russia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8EA8611B4E9B050AD8B95CF2E7152D0B">text</mods:typeOfResource>
<mods:relatedItem id="76D0A23E0DC92BCFCA10EE798453A8A1" type="host">
<mods:titleInfo id="A2BAB0DA117907942CB2AEC6756819DF">
<mods:title id="599E8E808137F934BF9433067F7AF331">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="43468776A7F50E01D8A5A944C25B0A26">
<mods:date id="E952F69619D0F04DC4EB3C334EC9A23E">2025</mods:date>
<mods:detail id="E8E4641AAD3F8CD8F4349E2E540B53AC" type="pubDate">
<mods:number id="0AF9FEA47628067B178F2552398F62F5">2025-01-08</mods:number>
</mods:detail>
<mods:detail id="4DCE4D127FA8A2AE322F75D52B7B03B2" type="volume">
<mods:number id="061A7ED7178452BE3C8E7306DB354DE3">1223</mods:number>
</mods:detail>
<mods:extent id="A2F7EFF21C8E5EDA61D412AF775C62A2" unit="page">
<mods:start id="4BBB4CC6E77B813A0FD8B9A728D05A13">185</mods:start>
<mods:end id="8330416013EE710F0A5DFBDCB4A519E4">220</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="37DCC6F4A5BA76107240F5263D1D20CB">journal article</mods:classification>
<mods:identifier id="2D47F199C49B1EB739C9F39F7E24A8BB" type="DOI">10.3897/zookeys.1223.139346</mods:identifier>
<mods:identifier id="767E97918EEC9C91CF51D7604BD03ED5" type="ZooBank">451749E9-009E-43E9-A6F1-892035BDF1B0</mods:identifier>
</mods:mods>
<treatment id="B77BE40B1CC852998AFAF927C052F425" ID-DOI="http://doi.org/10.5281/zenodo.14616989" ID-Zenodo-Dep="14616989" ID-arpha="B77BE40B-1CC8-5299-8AFA-F927C052F425" ID-zoobank="https://zoobank.org/B8043509-6990-4E62-99B8-CDFE90E3B656" LSID="urn:lsid:plazi:treatment:B77BE40B1CC852998AFAF927C052F425" LSID-ZBK="urn:lsid:zoobank.org:act:B8043509-6990-4E62-99B8-CDFE90E3B656" httpUri="http://treatment.plazi.org/id/B77BE40B1CC852998AFAF927C052F425" scope_class="Diplopoda" scope_family="Dalodesmidae" scope_order="Polydesmida">
<subSubSection id="4AD31273B479293E38F2C6CA70C047B2" type="nomenclature">
<paragraph id="98E4CE12FAED82530E1DA7FB21BE480E">
<taxonomicName id="F345DB6BBA789B701FE6289C61DE25EC" authorityName="Wesener &amp; Akkari &amp; Golovatch" authorityYear="2025" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="kompantsevi" status="sp. nov.">
<emphasis id="C9675A98460F4F76147BECCD331A14A2" italics="true">Dalodesmus kompantsevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7044C4149BB455CAD331AC0C3432D416" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0BB59B052772AA1C9FED5ADE6B703AF0" type="description">
<paragraph id="3EB7BFEB314B708043212FBB0E3A74C3">
<figureCitation id="413ACFFB8811C2E805FA972D6123777E" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">Figs 13</figureCitation>
,
<figureCitation id="D69020DB3552499DD59609597ADDA2E0" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dalodesmus kompantsevi sp. nov., ♀ (MZUF Fi 30 A). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, paranotum with ozopore C detail of surface structures of prozonite and metazonite D ozopore E endotergum F midbody legs. Scale bars: 1000 µm (A); 100 µm (B, C); 10 µm (D, E); 200 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure14" httpUri="https://binary.pensoft.net/fig/1215838">14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EE6BG" type="type material">
<paragraph id="14806A4FFAC1C261B5175EECE91D7F26">
<heading id="EC13548C16DB2AD20C1F2FF9BB7CF7BB" reason="title">Type material.</heading>
</paragraph>
<paragraph id="EA236625B9F84537E37A9B5CF156FBE7">
<materialsCitation id="941A11E9DA26B05C86F760D22778BBA7" collectedFrom="tropical forest" collectingDate="2018-12-16" collectingDateMax="2018-12-18" collectingDateMin="2018-12-16" collectionCode="ZMUM" collectorName="A. Kompantsev" country="Madagascar" elevation="950" latitude="-12.51358" location="Parc National Montagne d'Ambre" longLatPrecision="1" longitude="49.183002" specimenCount="1" specimenCount-male="1" stateProvince="Antsiranana Prov." typeStatus="holotype">
<emphasis id="C096C28652CB4F1824B31862DA1906E1" bold="true" italics="true">
<typeStatus id="EBB4B63F40B20D0C5A21A0211364DF13" type="holotype">Holotype</typeStatus>
</emphasis>
•
<specimenCount id="1A3E3FC6225DC3A2468B3FDC6F00B734" count="1" type="male">♂</specimenCount>
(
<abbrev content-type="institution" id="ABBRID0EP6BG" xlink_title="Zoological Museum, State University of Moscow, Russia">
<collectionCode id="465FCCB503BECDFBDAB07F2C94D75313" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" type="">ZMUM</collectionCode>
</abbrev>
), Northern
<collectingCountry id="90BBB83442CA3B7E563C64420376A2C4" name="Madagascar">Madagascar</collectingCountry>
,
<collectingRegion id="81ECE0F09454EF7218DDDD9465731411" country="Madagascar" name="Antsiranana">Antsiranana Prov.</collectingRegion>
,
<locationDeviation id="87598D3C862DD9C0C919F9028A7FD0ED" location="Joffreville">
<quantity id="1C16C573AD750FBEF58E88311E9CFF16" metricMagnitude="3" metricUnit="m" metricValue="4.0" unit="km" value="4.0">4 km</quantity>
SW of Joffreville
</locationDeviation>
(= Ambohitra),
<location id="8CC3FAD4EE0729825E709F91A500E71C" LSID="urn:lsid:plazi:treatment:B77BE40B1CC852998AFAF927C052F425:8CC3FAD4EE0729825E709F91A500E71C" country="Madagascar" latitude="-12.51358" longLatPrecision="1" longitude="49.183002" municipality="Northern" name="Parc National Montagne d'Ambre" stateProvince="Antsiranana Prov.">Parc National Montagne d’Ambre</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EX6BG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[49.183001,-12.513580]}">
<geoCoordinate id="B365F25F55DF05E6A23A9B0FABFD0B54" degrees="12.51358" direction="south" orientation="latitude" precision="1" value="-12.51358">12.51358°S</geoCoordinate>
,
<geoCoordinate id="AB1A818C013EF413BD30BAAD42D6FD9E" degrees="49.183001" direction="east" orientation="longitude" precision="1" value="49.183002">49.183001°E</geoCoordinate>
</named-content>
,
<elevation id="7683E39BD1BAAEC89DA9874A8A3D7BC4" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">
<quantity id="FCE5F483B354EA33A8475BC7325BC698" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">900–1000 m</quantity>
a. s. l.
</elevation>
,
<collectedFrom id="18B9314AF64AD32B3C2D2FE98E7F2F0B">tropical forest</collectedFrom>
,
<date id="B0D828FAE4E9F139490492603A099525" value="2018-12-16" valueMax="2018-12-18" valueMin="2018-12-16">
<collectingDate id="EF3A5BDA19071D7FEEF980EBF961B5EF" value="2018-12-16" valueMax="2018-12-18" valueMin="2018-12-16">16–18. XII. 2018</collectingDate>
</date>
,
<collectorName id="2E9BCE6CD60622FB3AC5C1469D4BAA58">A. Kompantsev</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E36BG" type="material">
<paragraph id="827E69A574377B3636686A984E1FED2A">
<heading id="FF81EE4883B03B70D957B9EC998942A2" reason="title">Other material.</heading>
</paragraph>
<paragraph id="D578AE4098952C3D5B4A57FFB1CF81DF">
•
<materialsCitation id="8F24E93D5C51CD94B832521538951D43" collectedFrom="c/o grande cascade" collectingDate="1989-09-26" collectionCode="MZUF" collectorName="L. Bartolozzi &amp; S. Taiti" country="Madagascar" elevation="900" location="Montagne d'Ambre" specimenCode="MZUF Fi-30 A" specimenCount="1" specimenCount-female="1">
<specimenCount id="46F2D73B31876BA998B408A8068B4A6A" count="1" type="female">1 ♀</specimenCount>
, (
<specimenCode id="AA4E1AB8F01678BC6D6804B7F62F18BC">
<named-content id="F4F2E260EE14A88D15519A5E1C774E84" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-zoologico-la-specola-universita-di-firenze" xlink_title="Museum «La Specola», Florence, Italy">MZUF</named-content>
Fi-30 A
</specimenCode>
),
<collectingCountry id="B37F56D4E5FEDD6C272F7D1D0E0B0E7E" name="Madagascar">Madagascar</collectingCountry>
,
<location id="3279C7A2C3891B317048864C7A23FC10" LSID="urn:lsid:plazi:treatment:B77BE40B1CC852998AFAF927C052F425:3279C7A2C3891B317048864C7A23FC10" country="Madagascar" name="Montagne d'Ambre">Montagne d’Ambre</location>
,
<quantity id="E83C1CF0AD5A318A70AD7AB12A6DCB0B" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">
<elevation id="881E0E67237A56FEC68A1E4F8F7C7671" metricMagnitude="2" metricUnit="m" metricValue="9.0" unit="m" value="900.0">900 m</elevation>
</quantity>
,
<collectedFrom id="6FC8F915749A90835F5FBFF518BF4F38">c/o grande cascade</collectedFrom>
, leg.
<date id="BA51B2FF3921D84FCC658CCD313E762A" value="1989-09-26">
<collectingDate id="F20DD3386AC9FB1E91510D16445271C0" value="1989-09-26">26 Sept. 1989</collectingDate>
</date>
,
<collectorName id="73098625B26F26AB56F02E010C67D08A">L. Bartolozzi</collectorName>
&amp;
<collectorName id="9EAD6C95E4B4AA0618D21DB05C09396B">S. Taiti</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIAAI" type="diagnosis">
<paragraph id="487E37E7A2D4BB771EA54294BA125AA8">
<heading id="618F6EFE1B4F6CFFE223FB3174952446" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="6815182720318E09D1AE4F7D61566A53">
Tips of paraterga not projecting past posterior tergal margin (Fig.
<figureCitation id="4FE0636863CFD8F8A44D7915C3A18C28" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 A – C</figureCitation>
), like in
<taxonomicName id="A267814C98A059A7674E1EF46CC2176D" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" class="Diplopoda" family="Dalodesmidae" genus="Tubercularium" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="2414EDDAF636DDBE3894B986A7483928" italics="true">D. hova</emphasis>
</taxonomicName>
,
<taxonomicName id="64FC0D42E701EBCCD74002B8369A29F1" authorityName="Cook" authorityYear="1896" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="tectus">
<emphasis id="A96220BFF8B242F2D2DEB8DA0ED780EA" italics="true">D. tectus</emphasis>
</taxonomicName>
and
<taxonomicName id="10A8361203F2E3499EB0672C0FAD8643" authorityName="Wesener &amp; Akkari &amp; Golovatch" authorityYear="2025" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="speophilus" status="sp. nov.">
<emphasis id="902BE5FE33A9D92591D440794F1E6098" italics="true">D. speophilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9494D40C942CA5D9467C44E353C33CD3" rank="species">sp. nov.</taxonomicNameLabel>
, vs sharper and projecting beyond margin in
<taxonomicName id="26786EDE825C698BA931AA6303AE4DB5" baseAuthorityName="Brandt" baseAuthorityYear="1841" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="ACB76A5263B4C0E83320218B2F28A003" italics="true">D. hamatus</emphasis>
</taxonomicName>
,
<taxonomicName id="F6D676587F31BE65564E3A3E58C1338A" baseAuthorityName="Attems" baseAuthorityYear="1898" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="odontopezus">
<emphasis id="C436F41DD957D2491302DB93049F8890" italics="true">D. odontopezus</emphasis>
</taxonomicName>
and
<taxonomicName id="95CDE72F4337811C18B615B3C38E412F" authorityName="Hoffman" authorityYear="1974" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="orator">
<emphasis id="0902E2C5FCAB36CF9B85D3CD3957A5C4" italics="true">D. orator</emphasis>
</taxonomicName>
. Differs from all other species of the genus primarily by the light colouration, coupled with the unusually compact, short, and clearly trifid acropodite, this being divided into three subequally long and upright branches: a middle, subacuminate and non-sigmoid solenomere (sl) flanked by a securiform, axe-shaped mesal branch (mb) and a nearly finger-shaped lateral branch (lb) of the solenophore (Fig.
<figureCitation id="7E713CFEE1AC3CD28C197C72699FE6EC" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 E, F</figureCitation>
). See also Key below.
</paragraph>
<caption id="F784D1F7D241FBE179CBB6686BB40228" ID-DOI="10.3897/zookeys.1223.139346.figure13" ID-arpha="B81D76F5-097E-56EC-97B7-6059B4CB59D2" httpUri="https://binary.pensoft.net/fig/1215837" startId="F13">
<paragraph id="9EF0C8F10F6D8AF4A87BE8E8E58940A4">
<label id="E0F99BA2C6E1AD03819ADF132EBB44D7">Figure 13.</label>
</paragraph>
<paragraph id="1975A803FEA1AE75470FB4626C9AEFEB">
<taxonomicName id="8B9F3B8681F7909E7C05F9E88692CEB4" authorityName="Wesener &amp; Akkari &amp; Golovatch" authorityYear="2025" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="kompantsevi" status="sp. nov.">
<emphasis id="BB01AC64B829260485A4ABAD65D14740" italics="true">Dalodesmus kompantsevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9C3FFCE5AC457134F2D26727F89984E3" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ holotype:
<emphasis id="C5AD52FBF973401F4A74C4D1166DB47E" bold="true">A – C</emphasis>
habitus, dorsal, lateral, and ventral views, respectively
<emphasis id="2BE60066AFDFEE4E5DEDE8463BEE3536" bold="true">D</emphasis>
body ring 7 with gonopods in situ, ventral view
<emphasis id="1C27666BFFB84C2A03D3F63BC6158D4F" bold="true">E, F</emphasis>
gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EWDAI" type="etymology">
<paragraph id="74890CC50CFBE48FA705207CCE4C2AC5">
<heading id="5660E8DBC1B166F32EE75E2469E5C64A" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="50E248CCE5CF517F4C1A8C5C318EBD24">
To honour the late Aleksandr Kompantsev (
<collectingCountry id="DFFAD0F9653C20AAD9F06372880701A9" name="Russia">Russia</collectingCountry>
), the collector.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E2DAI" type="description">
<paragraph id="2D25E097FFFB89A76B272C896301AA61">
<heading id="6BB1B8DCB09839FF78862EFF0D52B9D7" reason="title">Description.</heading>
</paragraph>
<paragraph id="B64259F0E40683DE3B640F8C6C5D4712">
Length of
<typeStatus id="0237DC08C854C9D66D20F758AE94330E" type="holotype">holotype</typeStatus>
~
<quantity id="EF8D71D594224A2456F59BB0257A1697" metricMagnitude="-2" metricUnit="m" metricValue="2.3" unit="mm" value="23.0">23 mm</quantity>
, width of midbody pro- and metazona 1.5 and
<quantity id="7917C91483E354A5568000C8EBB18E3D" metricMagnitude="-3" metricUnit="m" metricValue="2.9" unit="mm" value="2.9">2.9 mm</quantity>
, respectively. Width of prozona
<quantity id="18A8FD7F5B1840906BE0E623081487F8" metricMagnitude="-3" metricUnit="m" metricValue="2.2" unit="mm" value="2.2">2.2 mm</quantity>
, of metazona
<quantity id="DBC95B8783D5ABB7061BFE4722512EE1" metricMagnitude="-3" metricUnit="m" metricValue="4.2" unit="mm" value="4.2">4.2 mm</quantity>
in
<specimenCount id="A707832501070B31EEDA333CA8E89573" count="1" type="female">♀</specimenCount>
non-type.
</paragraph>
<paragraph id="4C2B009EA7FCB708D1326411099A35B6">
Colouration in alcohol uniformly pale brown to beige, axial stripe on prozona thin, vague, and grey, antennae somewhat infuscate, increasingly brown distad (Fig.
<figureCitation id="3D220C37C1D9D491AC9FF8C95569A3AC" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 A – C</figureCitation>
).
</paragraph>
<paragraph id="71744E81709C2F7B9FD9C62BAD3C74C1">
Body with 20 rings. Tegument mainly dull to only slightly shining, microgranulate to microtuberculate throughout, even surfaces of prozona and of metazona below paraterga finely microgranulate, sterna granulate. Head also densely microtuberculate or granulate throughout, micropilose up to level of antennae; epicranial suture thin, but distinct; genae squarish, set off ventrally from gnathochilarial stipes by a small, but evident ridge (Fig.
<figureCitation id="7E791230CD152F81535D171C2A339E4D" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 A – C</figureCitation>
). Interantennal isthmus ~ 2 × diameter of antennal socket (Fig.
<figureCitation id="0FF04BD0373A32C01365D3CD6631D529" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 C</figureCitation>
). Antennae short and rather clavate, in situ reaching back past ring 3 when stretched dorsally, very densely setose and microgranulate. In length, antennomere 6&gt; 5&gt; 3&gt; 4&gt; 2&gt; 1 = 7; antennomere 6 the largest and the highest, antennomeres 5 and 6 each with a small, round, distodorsal knob. In width, collum &lt;head &lt;ring 3 &lt;2 = 4–16; thereafter body gradually tapering towards telson (Fig.
<figureCitation id="47DD8C5C411D9B0356E7BFB3D8624371" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 A</figureCitation>
). Collum transversely suboval, regularly and broadly rounded laterally, densely tuberculate, most tuberculations being slightly oblong-oval, evident, equipped with very short, mostly subclavate setae and arranged in 7–8 transverse, rather irregular, arcuate rows. Metaterga 2 and 3 each with four similar transverse arcuated rows of setiferous tubercles, following metaterga each largely with 5–6 such rows (Figs
<figureCitation id="06A6862100DF1B9765A6AFE0662C27CF" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 A</figureCitation>
,
<figureCitation id="1C02885F6E242A3081C500310FF133EB" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dalodesmus kompantsevi sp. nov., ♀ (MZUF Fi 30 A). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, paranotum with ozopore C detail of surface structures of prozonite and metazonite D ozopore E endotergum F midbody legs. Scale bars: 1000 µm (A); 100 µm (B, C); 10 µm (D, E); 200 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure14" httpUri="https://binary.pensoft.net/fig/1215838">14 A, B</figureCitation>
). Paraterga well-developed, set high (mostly at upper ¼ body), largely slightly upturned to subhorizontal, thus leaving the dorsum only faintly convex (Figs
<figureCitation id="0C5D9B4943503814F8790B187AA3CB5F" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 B</figureCitation>
,
<figureCitation id="42DC070BDA30477E67031DA1CC72FB40" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dalodesmus kompantsevi sp. nov., ♀ (MZUF Fi 30 A). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, paranotum with ozopore C detail of surface structures of prozonite and metazonite D ozopore E endotergum F midbody legs. Scale bars: 1000 µm (A); 100 µm (B, C); 10 µm (D, E); 200 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure14" httpUri="https://binary.pensoft.net/fig/1215838">14 B</figureCitation>
); anterior and posterior margins of paraterga 2 and 3 clearly drawn both forward and caudad, following paraterga drawn increasingly only caudad, but caudal corners produced past rear tergal margin only on rings 17–19; posterior margins clearly bisinuate, well concave behind paraterga (Fig.
<figureCitation id="FB2C7DD7BB47EE5C9D09FC3D25D7B1A1" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 A – C</figureCitation>
). Lateral and caudal margins of paraterga beset with numerous, similarly oblong, and usually subequal, setigerous tubercles / lobulations, caudolateral lobulation nearly sharp. Ozopores (Fig.
<figureCitation id="E099F05F06506780F978DBA55EAD02D4" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dalodesmus kompantsevi sp. nov., ♀ (MZUF Fi 30 A). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, paranotum with ozopore C detail of surface structures of prozonite and metazonite D ozopore E endotergum F midbody legs. Scale bars: 1000 µm (A); 100 µm (B, C); 10 µm (D, E); 200 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure14" httpUri="https://binary.pensoft.net/fig/1215838">14 B, D</figureCitation>
) inconspicuous, opening dorsally near penultimate lateral lobulation on pore-baring rings 5, 7, 9, 10, 12, 13, 15–19. Strictures between pro- and metazona narrow and rather deep, nearly smooth (Fig.
<figureCitation id="0C116C77E17981027B2E11AFF3D4A82C" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dalodesmus kompantsevi sp. nov., ♀ (MZUF Fi 30 A). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, paranotum with ozopore C detail of surface structures of prozonite and metazonite D ozopore E endotergum F midbody legs. Scale bars: 1000 µm (A); 100 µm (B, C); 10 µm (D, E); 200 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure14" httpUri="https://binary.pensoft.net/fig/1215838">14 C</figureCitation>
). Epiproct small, conical and subtruncate at tip. Hypoproct trapeziform, with 1 + 1 setae borne on distinct oblong knobs at caudal margin. Limbus very thin, small, and entire. Neither an axial line nor pleurosternal carinae (Fig.
<figureCitation id="D61751D39D2F60F92C7650C28DD3E9DA" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 A – C</figureCitation>
). Endotergum inconspicuous, posterior margin of metazona projecting into long, sharp, apically microdenticulate, triangular projections (Fig.
<figureCitation id="73CB154A8BAE7C5AFF4C17D7F1C34120" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dalodesmus kompantsevi sp. nov., ♀ (MZUF Fi 30 A). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, paranotum with ozopore C detail of surface structures of prozonite and metazonite D ozopore E endotergum F midbody legs. Scale bars: 1000 µm (A); 100 µm (B, C); 10 µm (D, E); 200 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure14" httpUri="https://binary.pensoft.net/fig/1215838">14 E</figureCitation>
).
</paragraph>
<caption id="DFEAF0C13D95CDD35A2A939920B08CAF" ID-DOI="10.3897/zookeys.1223.139346.figure14" ID-arpha="48491778-792C-5E67-B661-E613E9EB5A32" httpUri="https://binary.pensoft.net/fig/1215838" startId="F14">
<paragraph id="8A5818EA9D55BF37D120E6C11CB958EB">
<label id="6F13753781564525DC983EA850B7B5B8">Figure 14.</label>
</paragraph>
<paragraph id="9A1DAB9EEDCBAA8CC8158C723A07CF56">
<taxonomicName id="21344CF7121297EB4B5329017F2C6044" authorityName="Wesener &amp; Akkari &amp; Golovatch" authorityYear="2025" class="Diplopoda" family="Dalodesmidae" genus="Dalodesmus" kingdom="Animalia" order="Polydesmida" phylum="Arthropoda" rank="species" species="kompantsevi" status="sp. nov.">
<emphasis id="D68FC02046804FE3CBDEF92385C42B0B" italics="true">Dalodesmus kompantsevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9CB5980D6DC71D6AA093D23035473D19" rank="species">sp. nov.</taxonomicNameLabel>
, ♀ (
<named-content id="B61E0EA952A95C51F11C5D89C82C5C7B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/museo-zoologico-la-specola-universita-di-firenze" xlink_title="Museum «La Specola», Florence, Italy">MZUF</named-content>
Fi 30 A).
<abbrev id="ABBRID0EQGAI" xlink_title="scanning electron microscopy">SEM</abbrev>
micrographs
<emphasis id="720C72C754E004DEAA9E566FDD80FBCD" bold="true">A</emphasis>
midbody ring with pro- and metazonite, dorsal view
<emphasis id="6A5CD9A5F946B807D49B355E2ACB48A8" bold="true">B</emphasis>
midbody ring, paranotum with ozopore
<emphasis id="3C1491E975DE1632F43A4886A1DE33D8" bold="true">C</emphasis>
detail of surface structures of prozonite and metazonite
<emphasis id="48A5C9CC8880DA73C020CD0A4CD046B5" bold="true">D</emphasis>
ozopore
<emphasis id="B3632566846B10D14442CEA09270639D" bold="true">E</emphasis>
endotergum
<emphasis id="827367D9CF7D7588E6EBB6C3DA79F03E" bold="true">F</emphasis>
midbody legs. Scale bars: 1000 µm (
<emphasis id="14A371A2ACDED3A0CB8C6715B34B15CB" bold="true">A</emphasis>
); 100 µm (
<emphasis id="812EA993A42ACE00047629615E3AACEC" bold="true">B, C</emphasis>
); 10 µm (
<emphasis id="241B8710C0611C4E5EFED23BA3729C8A" bold="true">D, E</emphasis>
); 200 µm (
<emphasis id="F00260E963AF372ECDB20C9159F25CC8" bold="true">F</emphasis>
).
</paragraph>
</caption>
<paragraph id="E729B088C385DA57CBD67F56DCB86D24">
Sterna mostly densely setose, deep, and narrow between coxae 1–3, increasingly broad thereafter, clearly excavate between coxae 5 and 6 (
<specimenCount id="8D61A1F09D3366F9A78C5C8495950AA8" count="1" type="male">♂</specimenCount>
); postgonopodal sterna rather flat, devoid of modifications, cross-impressions faint (Fig.
<figureCitation id="A77275C196DE966B0023084B5B5F2F3A" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 C</figureCitation>
). Gonopodal aperture roundly pentagonal, relatively small, taking up ~ 1 / 3 width of metazonum 7, clearly open and drawn into metazonum 6 (Fig.
<figureCitation id="848A48A0A7442C515770D3A5E10A6983" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 D</figureCitation>
). Legs incrassate, rather long. 1.4–1.5 × as long as body height, with small, stout, abundant and usual curved setae (Fig.
<figureCitation id="85C75323B3ED7178281A9D1131702637" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Dalodesmus kompantsevi sp. nov., ♀ (MZUF Fi 30 A). SEM micrographs A midbody ring with pro- and metazonite, dorsal view B midbody ring, paranotum with ozopore C detail of surface structures of prozonite and metazonite D ozopore E endotergum F midbody legs. Scale bars: 1000 µm (A); 100 µm (B, C); 10 µm (D, E); 200 µm (F)." figureDoi="10.3897/zookeys.1223.139346.figure14" httpUri="https://binary.pensoft.net/fig/1215838">14 F</figureCitation>
) with admixture of sphaerotrichomes ventrally on all podomeres (
<specimenCount id="C74F0A01EBC9E41D5519BE5F92E42F9E" count="1" type="male">♂</specimenCount>
); gonopores on coxae 2 inconspicuous, each borne on a very small swelling (
<specimenCount id="31BAE75FA127FA9F327904BA60AC3521" count="1" type="male">♂</specimenCount>
); prefemora not bulged laterally; claws very small; in length, tarsus&gt; femur&gt; prefemur&gt; tibia&gt; postfemur&gt; coxa.
</paragraph>
<paragraph id="588FC51C5D7CF96167D3E3948F97C387">
Gonopods (Fig.
<figureCitation id="5AE0A760A01C4D0BFB2BDECAB2B7971C" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 D – F</figureCitation>
) very slender and long, tips in situ reaching anteriorly beyond coxae 6 (Fig.
<figureCitation id="D8419B1944BF4C4B23CA0441111EF391" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Dalodesmus kompantsevi sp. nov., ♂ holotype: A – C habitus, dorsal, lateral, and ventral views, respectively D body ring 7 with gonopods in situ, ventral view E, F gonopods, dorsal and ventral views, respectively. Abbreviations: fe femorite, lb lateral branch, mb mesal branch, sl solenomere. Photographs by K. Makarov, taken not to scale." figureDoi="10.3897/zookeys.1223.139346.figure13" httpUri="https://binary.pensoft.net/fig/1215837">13 C, D</figureCitation>
). Both coxites and clearly prefemoral (= densely setose) parts of telopodites equally very short and stout, fused medially, the former fully and the latter mostly hidden inside gonopodal aperture. Femorites (fe) contiguous medially in basal 1 / 3, basically bare and only one at base with a distinct lateral seta, both very faintly attenuating and diverging distad. Acropodites of both telopodites clearly diverging, but unusually compact, short, and clearly trifid, being divided into three subequally short and upright branches: a middle, subacuminate and non-sigmoid solenomere (sl) flanked by a solenophore represented by a securiform, axe-shaped mesal branch (mb) and a nearly finger-shaped lateral branch (lb) with a mesal knob parabasally.
</paragraph>
</subSubSection>
</treatment>
</document>
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