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<mods:title id="3884CBD568E9F409883206D69B24A282">Redescription of the eel Ilyophis arx Robins, 1976 (Anguilliformes: Synaphobranchidae: Ilyophinae) with a description of a new species of Ilyophis, and a taxonomic synopsis of the Ilyophinae</mods:title>
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<mods:namePart id="2097E2B7B93B3151F2FB59E5B0EE9EF2">Tighe, Kenneth A.</mods:namePart>
<mods:affiliation id="CDDE011C1349341D484EC01A67EE234B">Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, D. C., U. S. A.</mods:affiliation>
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<mods:namePart id="2B5CBBB47AC29C4D7CE74ABFFD3B29F0">Smith, David G.</mods:namePart>
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<taxonomicName id="40DD4D0DFF99770CFF79F6DDFD3B02A1" authority="Tighe, Smith &amp; Merrett" authorityName="Tighe, Smith &amp; Merrett" authorityYear="2024" box="[151,683,151,178]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="9" pageNumber="44" phylum="Chordata" rank="species" species="maclainei" status="sp. nov.">Ilyophis maclainei Tighe, Smith &amp; Merrett</taxonomicName>
,
<taxonomicNameLabel id="AE9A57E7FF99770CFD59F6DDFC8502A1" box="[695,789,151,177]" pageId="9" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="1FE62A0BFF99770CFF79F695FE8F02E9" box="[151,287,223,249]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStart-4="FIGURE 8" captionStart-5="FIGURE 9" captionStartId-0="9.[151,250,851,875]" captionStartId-1="10.[151,250,1902,1926]" captionStartId-2="11.[151,250,1052,1076]" captionStartId-3="11.[151,250,1918,1942]" captionStartId-4="12.[151,250,1052,1076]" captionStartId-5="13.[151,250,1501,1525]" captionTargetBox-0="[151,1436,432,827]" captionTargetBox-1="[346,1240,1246,1872]" captionTargetBox-2="[349,1238,190,1019]" captionTargetBox-3="[347,1240,1214,1885]" captionTargetBox-4="[346,1242,185,1020]" captionTargetBox-5="[158,1429,385,1468]" captionTargetId-0="figure-91@9.[151,1436,432,827]" captionTargetId-1="figure-563@10.[340,1247,1239,1878]" captionTargetId-2="figure-19@11.[340,1247,181,1028]" captionTargetId-3="figure-51@11.[340,1247,1206,1893]" captionTargetId-4="figure-34@12.[340,1247,179,1026]" captionTargetId-5="figure-106@13.[151,1436,378,1475]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="11" captionTargetPageId-4="12" captionTargetPageId-5="13" captionText-0="FIGURE 4. Ilyophis maclainei sp. nov. Holotype, BMNH 1995.4.19.31 (male, 242 mm TL)" captionText-1="FIGURE 5. Ilyophis maclainei sp. nov. Referred Specimen, DISCOVERY Collection, Station 50517, 237 mm TL (A) Lateral view. (B) Head and cephalic lateralis system (from Merrett &amp; Saldanha 1985)" captionText-2="FIGURE 6. Ilyophis maclainei sp. nov. Holotype, BMNH 1995.4.19.31(DISCOVERY Collection, Station 50710), 240 mm TL (A) Snout ornamentation. (B) Lower jaw tip ornamentation (from Merrett &amp; Saldanha 1985)" captionText-3="FIGURE 7. Ilyophis maclainei sp. nov. Referred Specimen, DISCOVERY Collection, Station 50517, 237 mm TL (A) Dentition of upper jaw. (B) Dentition of lower jaw (from Merrett &amp; Saldanha 1985)" captionText-4="FIGURE 8. Ilyophis maclainei sp. nov. (A) Referred Specimen, DISCOVERY Collection, Station 50517. Distribution of type α and ss squamation on left and right side of specimen. (B) Same specimen. Detail of squamation from area of trunk region (scales of type α ventrally). (C) Same specimen. Detail of squamation from anal region (scales of type ss). (D) Holotype, BMNH 1995.4.19.31. Detail of squamation from region of end of caudal (scales of type ss) (modified from Merrett &amp; Saldanha 1985)" captionText-5="FIGURE 9. Distribution of Ilyophis maclainei in the eastern North Atlantic Ocean. Open star is the type locality. Closed stars are other study material." figureDoi-0="http://doi.org/10.5281/zenodo.13746109" figureDoi-1="http://doi.org/10.5281/zenodo.13746111" figureDoi-2="http://doi.org/10.5281/zenodo.13746113" figureDoi-3="http://doi.org/10.5281/zenodo.13746115" figureDoi-4="http://doi.org/10.5281/zenodo.13746119" figureDoi-5="http://doi.org/10.5281/zenodo.13746121" httpUri-0="https://zenodo.org/record/13746109/files/figure.png" httpUri-1="https://zenodo.org/record/13746111/files/figure.png" httpUri-2="https://zenodo.org/record/13746113/files/figure.png" httpUri-3="https://zenodo.org/record/13746115/files/figure.png" httpUri-4="https://zenodo.org/record/13746119/files/figure.png" httpUri-5="https://zenodo.org/record/13746121/files/figure.png" pageId="9" pageNumber="44">Figures 49</figureCitation>
,
<tableCitation id="CA5F0335FF99770CFEC4F695FE0802E9" box="[298,408,223,249]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="3.[152,245,151,175]" captionStartId-1="7.[152,245,468,494]" captionTargetPageId-0="3" captionTargetPageId-1="7" captionText-0="TABLE 1. Morphometric, meristic and morphological data of Ilyophis arx Robins, 1976 and Ilyophis maclainei sp. nov. For bilateral pore counts, the counts are recorded left/right." captionText-1="TABLE 2. Comparison of morphometric, meristic, and morphological data for all species of Ilyophis. For unique specimens (holotypes and paratypes), the bilateral pore counts are recorded left/right (if recorded in original description)." httpUri-0="http://table.plazi.org/id/D3A26606FF937706FF76F6DDFC9E02C3" httpUri-1="http://table.plazi.org/id/D3A26606FF977702FF76F79EFC960002" pageId="9" pageNumber="44" tableUuid-0="D3A26606FF937706FF76F6DDFC9E02C3" tableUuid-1="D3A26606FF977702FF76F79EFC960002">Table 12</tableCitation>
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<taxonomicName id="40DD4D0DFF99770CFF79F76EFE82032B" ID-CoL="6N6QN" authorityName="C. H. Robins" authorityYear="1976" box="[151,274,292,316]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="9" pageNumber="44" phylum="Chordata" rank="species" species="arx">Ilyophis arx</taxonomicName>
(not of
<bibRefCitation id="E34C4B7FFF99770CFE8AF76EFE60032B" author="Robins, C. H. &amp; Robins, C. R." box="[356,496,291,316]" pageId="9" pageNumber="44" pagination="293 - 335" refId="ref18601" refString="Robins, C. H. &amp; Robins, C. R. (1976) New genera and species of dysommine and synaphobranchine eels (Synaphobranchidae) with an analysis of the Dysomminae. Proceedings of the Academy of Natural Sciences of Philadelphia, 122, 293 - 335." type="journal article" year="1976">Robins, 1976</bibRefCitation>
).
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<bibRefCitation id="E34C4B7FFF99770CFDEFF76EFCA9032B" author="Saldanha, L. &amp; Merrett, N. R." box="[513,825,291,316]" pageId="9" pageNumber="44" pagination="623 - 636" refId="ref18728" refString="Saldanha, L. &amp; Merrett, N. R. (1982) A new species of the deep-sea eel genus llyophis Gilbert (Synaphobranchidae) from the eastern North Atlantic, with comments on its ecology and intrafamilial relationships. Journal of Fish Biology, 21, 623 - 636. https: // doi. org / 10.1111 / j. 1095 - 8649.1982. tb 02866. x" type="journal article" year="1982">Saldanha &amp; Merrett 1982: 633</bibRefCitation>
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(listing of
<specimenCount id="91DBFD07FF99770CFC44F76EFBA6032C" box="[938,1078,292,316]" count="1" pageId="9" pageNumber="44" type="generic">one specimen</specimenCount>
from Porcupine Sea-Bight, eastern North Atlantic).
<treatmentCitation id="067C109FFF99770CFE9CF709FD72034A" author="Merrett, N. R. &amp; Saldanha, L." box="[370,738,322,346]" page="730" pageId="9" pageNumber="44" year="1985">
<bibRefCitation id="E34C4B7FFF99770CFE9CF709FD72034A" author="Merrett, N. R. &amp; Saldanha, L." box="[370,738,322,346]" pageId="9" pageNumber="44" pagination="719 - 747" refId="ref18341" refString="Merrett, N. R. &amp; Saldanha, L. (1985) Aspects of the morphology and ecology of some unusual deep-sea eels (Synaphobranchidae, Derichthyidae and Nettastomatidae) from the eastern North Atlantic. Journal of Fish Biology, 27, 719 - 747. https: // doi. org / 10.1111 / j. 1095 - 8649.1985. tb 03216. x" type="journal article" year="1985">Merrett &amp; Saldanha 1985: 730735</bibRefCitation>
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(description of northeastern Atlantic specimens).
<treatmentCitation id="067C109FFF99770CFB06F708FEB70369" author="Robins, C. H. &amp; Robins, C. R." page="238" pageId="9" pageNumber="44" year="1989">
<bibRefCitation id="E34C4B7FFF99770CFB06F708FEB70369" author="Robins, C. H. &amp; Robins, C. R." pageId="9" pageNumber="44" pagination="207 - 253" refId="ref18652" refString="Robins, C. H. &amp; Robins, C. R. (1989) Family Synaphobranchidae. In: Bohlke, E. B. (Ed.), Fishes of the Western North Atlantic. Memoir of the Sears Foundation for Marine Research, 1 (Part 9), pp. 207 - 253. https: // doi. org / 10.2307 / j. ctvbcd 0 dm. 12" type="journal article" year="1989">Robins &amp; Robins 1989:238</bibRefCitation>
</treatmentCitation>
(suggestion that northeastern Atlantic population was not
<taxonomicName id="40DD4D0DFF99770CFC81F72BFC780369" ID-CoL="6N6QN" authorityName="C. H. Robins" authorityYear="1976" box="[879,1000,353,377]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="9" pageNumber="44" phylum="Chordata" rank="species" species="arx">Ilyophis arx</taxonomicName>
).
<treatmentCitation id="067C109FFF99770CFC12F72BFACB0369" author="Sulak, K. J. &amp; Shcherbachev, Y. N." box="[1020,1371,353,377]" page="1163" pageId="9" pageNumber="44" year="1997">
<bibRefCitation id="E34C4B7FFF99770CFC12F72BFACB0369" author="Sulak, K. J. &amp; Shcherbachev, Y. N." box="[1020,1371,353,377]" pageId="9" pageNumber="44" pagination="1158 - 1194" refId="ref19041" refString="Sulak, K. J. &amp; Shcherbachev, Y. N. (1997) Zoogeography and systematics of six deep-living genera of synaphobranchid eels, with a key to taxa and description of two new species of Ilyophis. Bulletin of Marine Science, 60 (3), 1158 - 1194." type="journal article" year="1997">Sulak &amp; Shcherbachev 1997: 1163</bibRefCitation>
</treatmentCitation>
, 1172, 1188 (key, identification as llyophis
<taxonomicName id="40DD4D0DFF99770CFDD9F7CAFDC90387" ID-CoL="6N6QN" authorityName="C. H. Robins" authorityYear="1976" box="[567,601,384,407]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="9" pageNumber="44" phylum="Chordata" rank="species" species="arx">arx</taxonomicName>
, distribution).
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FIGURE 4.
<taxonomicName id="40DD4D0DFF99770CFEF7F519FE45017B" authorityName="Tighe &amp; Smith &amp; Merrett &amp; Frable &amp; Zajonz" authorityYear="2024" box="[281,469,851,875]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="9" pageNumber="44" phylum="Chordata" rank="species" species="maclainei" status="sp. nov.">Ilyophis maclainei</taxonomicName>
<taxonomicNameLabel id="AE9A57E7FF99770CFE32F519FDBC017B" box="[476,556,851,875]" pageId="9" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
Holotype, BMNH 1995.4.19.31 (male, 242 mm TL)
</paragraph>
</caption>
<subSubSection id="CFC76505FF99770CFF79F5F7FD4706BF" pageId="9" pageNumber="44" type="materials_examined">
<paragraph id="8762368EFF99770CFF79F5F7FAA801EB" blockId="9.[151,1437,957,2028]" pageId="9" pageNumber="44">
<materialsCitation id="37B53CD3FF99770CFF79F5F7FAA801EB" collectingDate="1979-10-17" collectionCode="BMNH" collectorName="SW Ireland" elevation="1900" latitude="49.501667" location="Discovery Collection Station" longLatPrecision="18" longitude="13.331667" municipality="northeastern Atlantic Ocean" pageId="9" pageNumber="44" specimenCode="BMNH 1995.4" specimenCount="1" typeStatus="holotype">
<typeStatus id="5866882CFF99770CFF79F5F7FE9801C7" box="[151,264,957,983]" pageId="9" pageNumber="44" type="holotype">Holotype</typeStatus>
:
<specimenCode id="D77B9EF5FF99770CFEF9F5F4FE5601C8" box="[279,454,957,984]" collectionCode="BMNH" pageId="9" pageNumber="44">BMNH 1995.4</specimenCode>
.19.31 (male,
<quantity id="40259B6BFF99770CFD8CF5F7FD5101C8" box="[610,705,957,984]" metricMagnitude="-1" metricUnit="m" metricValue="2.42" pageId="9" pageNumber="44" unit="mm" value="242.0">242 mm</quantity>
TL),
<location id="82026055FF99770CFCE8F5F7FBC001C7" LSID="urn:lsid:plazi:treatment:0F748798FF997708FF79F6DDFD6B05B6:82026055FF99770CFCE8F5F7FBC001C7" box="[774,1104,957,983]" latitude="49.501667" longLatPrecision="18" longitude="13.331667" municipality="northeastern Atlantic Ocean" name="Discovery Collection Station" pageId="9" pageNumber="44">Discovery Collection Station</location>
50710, Goban Spur, off
<collectorName id="2A285358FF99770CFA9DF5F7FF7901EB" pageId="9" pageNumber="44">SW Ireland</collectorName>
,
<collectingMunicipality id="6706ACF4FF99770CFF1AF5ABFDA001EB" box="[244,560,993,1019]" pageId="9" pageNumber="44">northeastern Atlantic Ocean</collectingMunicipality>
,
<geoCoordinate id="E2E95049FF99770CFDD5F5ABFD4A01EC" box="[571,730,993,1020]" degrees="49" direction="north" minutes="30" orientation="latitude" pageId="9" pageNumber="44" precision="15" seconds="06" value="49.501667">49° 30 06” N</geoCoordinate>
,
<geoCoordinate id="E2E95049FF99770CFD0BF5ABFC1101EC" box="[741,897,993,1020]" degrees="13" direction="east" minutes="19" orientation="longitude" pageId="9" pageNumber="44" precision="15" seconds="54" value="13.331667">13° 19 54” E</geoCoordinate>
;
<quantity id="40259B6BFF99770CFC63F5ABFBBB01EC" box="[909,1067,993,1020]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="9" pageNumber="44" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">
<elevation id="0CF0D1BDFF99770CFC63F5ABFBBB01EC" box="[909,1067,993,1020]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="9" pageNumber="44" unit="m" value="1900.0" valueMax="2000.0" valueMin="1800.0">18002000 m</elevation>
</quantity>
; date:
<date id="F363104EFF99770CFB9AF5ABFAA301EB" box="[1140,1331,993,1019]" pageId="9" pageNumber="44" value="1979-10-17">
<collectingDate id="E327E9A6FF99770CFB9AF5ABFAA301EB" box="[1140,1331,993,1019]" pageId="9" pageNumber="44" value="1979-10-17">17 October 1979</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8762368EFF99770CFF29F24FFB7B0653" blockId="9.[151,1437,957,2028]" pageId="9" pageNumber="44">
<materialsCitation id="37B53CD3FF99770CFF29F24FFB770653" collectingDate="1979-10-18" collectionCode="BMNH" collectorName="SW Ireland" elevation="4788" latitude="49.883335" location="Discovery Collection Station" longLatPrecision="1100" longitude="15.6" municipality="northeastern Atlantic Ocean" pageId="9" pageNumber="44" specimenCode="BMNH 1995.3" specimenCount="1" typeStatus="paratype">
<typeStatus id="5866882CFF99770CFF29F24FFEA8060F" box="[199,312,1029,1055]" pageId="9" pageNumber="44" type="paratype">Paratype</typeStatus>
:
<specimenCode id="D77B9EF5FF99770CFEAAF24CFE7E0630" box="[324,494,1029,1056]" collectionCode="BMNH" pageId="9" pageNumber="44">BMNH 1995.3</specimenCode>
.29.1 (male,
<quantity id="40259B6BFF99770CFD98F24FFD430630" box="[630,723,1029,1056]" metricMagnitude="-1" metricUnit="m" metricValue="2.63" pageId="9" pageNumber="44" unit="mm" value="263.0">263 mm</quantity>
TL),
<location id="82026055FF99770CFCFFF24FFBC4060F" LSID="urn:lsid:plazi:treatment:0F748798FF997708FF79F6DDFD6B05B6:82026055FF99770CFCFFF24FFBC4060F" box="[785,1108,1029,1055]" latitude="49.883335" longLatPrecision="1100" longitude="15.6" municipality="northeastern Atlantic Ocean" name="Discovery Collection Station" pageId="9" pageNumber="44">Discovery Collection Station</location>
50711, Porcupine Sea-Bight, off
<collectorName id="2A285358FF99770CFF50F263FED10653" box="[190,321,1065,1091]" pageId="9" pageNumber="44">SW Ireland</collectorName>
,
<collectingMunicipality id="6706ACF4FF99770CFEA2F263FD180653" box="[332,648,1065,1091]" pageId="9" pageNumber="44">northeastern Atlantic Ocean</collectingMunicipality>
;
<geoCoordinate id="E2E95049FF99770CFD7DF263FC930654" box="[659,771,1065,1092]" degrees="49" direction="north" minutes="53" orientation="latitude" pageId="9" pageNumber="44" precision="925" value="49.883335">49° 53 N</geoCoordinate>
,
<geoCoordinate id="E2E95049FF99770CFCE0F263FCEB0654" box="[782,891,1065,1092]" degrees="15" direction="east" minutes="36" orientation="longitude" pageId="9" pageNumber="44" precision="925" value="15.6">15° 36 E</geoCoordinate>
;
<quantity id="40259B6BFF99770CFC68F263FC4E0654" box="[902,990,1065,1092]" metricMagnitude="3" metricUnit="m" metricValue="4.788" pageId="9" pageNumber="44" unit="m" value="4788.0">
<elevation id="0CF0D1BDFF99770CFC68F263FC4E0654" box="[902,990,1065,1092]" metricMagnitude="3" metricUnit="m" metricValue="4.788" pageId="9" pageNumber="44" unit="m" value="4788.0">4788 m</elevation>
</quantity>
; date:
<date id="F363104EFF99770CFBC9F263FB770653" box="[1063,1255,1065,1091]" pageId="9" pageNumber="44" value="1979-10-18">
<collectingDate id="E327E9A6FF99770CFBC9F263FB770653" box="[1063,1255,1065,1091]" pageId="9" pageNumber="44" value="1979-10-18">18 October 1979</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8762368EFF99770CFF29F207FD4706BF" blockId="9.[151,1437,957,2028]" pageId="9" pageNumber="44">
Referred Specimen:
<materialsCitation id="37B53CD3FF99770CFE52F207FD4206BF" collectingDate="1979-06-07" collectionCode="BMNH" collectorName="Goban Spur &amp; SW Ireland" elevation="1790" latitude="49.558334" location="Discovery Collection Station" longLatPrecision="18" longitude="13.466666" municipality="northeastern Atlantic Ocean" pageId="9" pageNumber="44" specimenCode="BMNH 1996.3" specimenCount="1">
<location id="82026055FF99770CFE52F207FC900677" LSID="urn:lsid:plazi:treatment:0F748798FF997708FF79F6DDFD6B05B6:82026055FF99770CFE52F207FC900677" box="[444,768,1101,1127]" latitude="49.558334" longLatPrecision="18" longitude="13.466666" municipality="northeastern Atlantic Ocean" name="Discovery Collection Station" pageId="9" pageNumber="44">Discovery Collection Station</location>
50517 (female,
<quantity id="40259B6BFF99770CFC56F207FB850678" box="[952,1045,1101,1128]" metricMagnitude="-1" metricUnit="m" metricValue="2.37" pageId="9" pageNumber="44" unit="mm" value="237.0">237 mm</quantity>
TL, portion of specimen cataloged as
<specimenCode id="D77B9EF5FF99770CFF58F238FEF3069B" box="[182,355,1137,1164]" collectionCode="BMNH" pageId="9" pageNumber="44">BMNH 1996.3</specimenCode>
.8.2, see remarks below),
<collectorName id="2A285358FF99770CFD6DF23BFC9D069B" box="[643,781,1137,1163]" pageId="9" pageNumber="44">Goban Spur</collectorName>
, off
<collectorName id="2A285358FF99770CFCAEF23BFC54069B" box="[832,964,1137,1163]" pageId="9" pageNumber="44">SW Ireland</collectorName>
,
<collectingMunicipality id="6706ACF4FF99770CFC3EF23BFA9D069B" box="[976,1293,1137,1163]" pageId="9" pageNumber="44">northeastern Atlantic Ocean</collectingMunicipality>
,
<geoCoordinate id="E2E95049FF99770CFAF7F23BFF3E06A0" degrees="49" direction="north" minutes="33" orientation="latitude" pageId="9" pageNumber="44" precision="15" seconds="30" value="49.558334">49° 33 30” N</geoCoordinate>
,
<geoCoordinate id="E2E95049FF99770CFF57F2DFFEC506A0" box="[185,341,1173,1200]" degrees="13" direction="east" minutes="28" orientation="longitude" pageId="9" pageNumber="44" precision="15" seconds="00" value="13.466666">13° 28 00” E</geoCoordinate>
;
<quantity id="40259B6BFF99770CFE8FF2DFFE6F06A0" box="[353,511,1173,1200]" metricMagnitude="3" metricUnit="m" metricValue="1.7894999999999999" metricValueMax="1.794" metricValueMin="1.785" pageId="9" pageNumber="44" unit="m" value="1789.5" valueMax="1794.0" valueMin="1785.0">
<elevation id="0CF0D1BDFF99770CFE8FF2DFFE6F06A0" box="[353,511,1173,1200]" metricMagnitude="3" metricUnit="m" metricValue="1.7894999999999999" metricValueMax="1.794" metricValueMin="1.785" pageId="9" pageNumber="44" unit="m" value="1789.5" valueMax="1794.0" valueMin="1785.0">17851794 m</elevation>
</quantity>
; date:
<date id="F363104EFF99770CFDA6F2DCFD4206BF" box="[584,722,1173,1200]" pageId="9" pageNumber="44" value="1979-06-07">
<collectingDate id="E327E9A6FF99770CFDA6F2DCFD4206BF" box="[584,722,1173,1200]" pageId="9" pageNumber="44" value="1979-06-07">7 June 1979</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CFC76505FF99770CFF29F2F3FC51042B" pageId="9" pageNumber="44" type="diagnosis">
<paragraph id="8762368EFF99770CFF29F2F3FC51042B" blockId="9.[151,1437,957,2028]" pageId="9" pageNumber="44">
Diagnosis. A species of
<taxonomicName id="40DD4D0DFF99770CFE02F2F3FDD606C3" authorityName="Gilbert" authorityYear="1891" box="[492,582,1209,1235]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="9" pageNumber="44" phylum="Chordata" rank="genus">Ilyophis</taxonomicName>
with the following characters. Body covered with scales in an irregular basket-weave pattern. Trunk moderate (about 20% of TL). Dorsal-fin origin above posterior half of pectoral fin. Gape moderate, extending at most to posterior half of eye. Teeth not compound. Intermaxillary teeth conical and acute with approximately 1618 teeth in the tooth patch. Vomerine teeth abutting the rear of intermaxillary patch with approximately 25 teeth irregularly biserial anteriorly becoming uniserial posteriorly. Both intermaxillary and vomerine teeth set in beds of papillose tissue. Maxillary teeth arranged in bands of one to four (medially) to one irregular row, with anterior teeth somewhat enlarged. Dentary teeth arranged in bands with roughly three to five series of teeth, with anterior ones (in two rows) the largest. Lateral line extends to approximately 7075% the length of body. Cephalic lateralis pores: supraorbital pores (SO) 3, infraorbital pores (IO) 6 (including adnasal pore just posterior to anterior nostril), preoperculomandibular pores (POM) 89, 78 pores in mandibular region and the eighth or ninth pore in a preopercular position. Total vertebrae 131134.
</paragraph>
</subSubSection>
<subSubSection id="CFC76505FF99770FFF29F00FFB6906B1" lastPageId="10" lastPageNumber="45" pageId="9" pageNumber="44" type="description">
<paragraph id="8762368EFF99770CFF29F00FFACB0504" blockId="9.[151,1437,957,2028]" pageId="9" pageNumber="44">
Description. Measurements of
<typeStatus id="5866882CFF99770CFDDEF00FFD06044F" box="[560,662,1605,1631]" pageId="9" pageNumber="44" type="holotype">holotype</typeStatus>
: total length
<quantity id="40259B6BFF99770CFCC0F00FFC1F0470" box="[814,911,1605,1632]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" pageId="9" pageNumber="44" unit="mm" value="240.0">240 mm</quantity>
, preanal length
<quantity id="40259B6BFF99770CFBA6F00FFB0C0470" box="[1096,1180,1605,1632]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="9" pageNumber="44" unit="mm" value="76.0">76 mm</quantity>
, predorsal length
<quantity id="40259B6BFF99770CFA85F00FFF550494" metricMagnitude="-2" metricUnit="m" metricValue="3.6399999999999997" pageId="9" pageNumber="44" unit="mm" value="36.4">36.4 mm</quantity>
, head length 30.0 mm, trunk length
<quantity id="40259B6BFF99770CFDB9F023FD370494" box="[599,679,1641,1668]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="9" pageNumber="44" unit="mm" value="46.0">46 mm</quantity>
, snout length
<quantity id="40259B6BFF99770CFCACF023FC360494" box="[834,934,1641,1668]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="9" pageNumber="44" unit="mm" value="11.8">11.8 mm</quantity>
, upper jaw length 14.0 mm, eye diameter
<quantity id="40259B6BFF99770CFA97F023FF5504B8" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="9" pageNumber="44" unit="mm" value="3.3">3.3 mm</quantity>
, interorbital width
<quantity id="40259B6BFF99770CFE75F0C7FE6304B8" box="[411,499,1677,1704]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="9" pageNumber="44" unit="mm" value="6.6">6.6 mm</quantity>
, gill opening
<quantity id="40259B6BFF99770CFD65F0C7FD7304B8" box="[651,739,1677,1704]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="9" pageNumber="44" unit="mm" value="2.8">2.8 mm</quantity>
, intrabronchial distance 5.8, pectoral fin length
<quantity id="40259B6BFF99770CFB15F0C7FAC204B8" box="[1275,1362,1677,1704]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="9" pageNumber="44" unit="mm" value="7.8">7.8 mm</quantity>
, depth at anus
<quantity id="40259B6BFF99770CFF01F0FBFED904DC" box="[239,329,1713,1740]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="9" pageNumber="44" unit="mm" value="9.9">9.9 mm</quantity>
. Meristics and proportional morphometrics of the
<typeStatus id="5866882CFF99770CFC6DF0FBFC7504DB" box="[899,997,1713,1739]" pageId="9" pageNumber="44" type="holotype">holotype</typeStatus>
and
<typeStatus id="5866882CFF99770CFBF0F0F8FBEE04DC" box="[1054,1150,1714,1740]" pageId="9" pageNumber="44" type="paratype">paratype</typeStatus>
are presented in
<tableCitation id="CA5F0335FF99770CFAAFF0FBFA0604DB" box="[1345,1430,1713,1739]" captionStart="TABLE 1" captionStartId="3.[152,245,151,175]" captionTargetPageId="3" captionText="TABLE 1. Morphometric, meristic and morphological data of Ilyophis arx Robins, 1976 and Ilyophis maclainei sp. nov. For bilateral pore counts, the counts are recorded left/right." httpUri="http://table.plazi.org/id/D3A26606FF937706FF76F6DDFC9E02C3" pageId="9" pageNumber="44" tableUuid="D3A26606FF937706FF76F6DDFC9E02C3">Table 1</tableCitation>
. Rest of description based on data presented by
<bibRefCitation id="E34C4B7FFF99770CFD4EF09CFC4204E0" author="Merrett, N. R. &amp; Saldanha, L." box="[672,978,1749,1776]" pageId="9" pageNumber="44" pagination="719 - 747" refId="ref18341" refString="Merrett, N. R. &amp; Saldanha, L. (1985) Aspects of the morphology and ecology of some unusual deep-sea eels (Synaphobranchidae, Derichthyidae and Nettastomatidae) from the eastern North Atlantic. Journal of Fish Biology, 27, 719 - 747. https: // doi. org / 10.1111 / j. 1095 - 8649.1985. tb 03216. x" type="journal article" year="1985">Merrett &amp; Saldanha (1985)</bibRefCitation>
. However, it should be noted that the
<specimenCount id="91DBFD07FF99770CFA9CF09CFE9C0503" count="2" pageId="9" pageNumber="44" type="generic">two specimens</specimenCount>
available for examination have shrunk since originally measured by
<bibRefCitation id="E34C4B7FFF99770CFBFFF0B0FAC70504" author="Merrett, N. R. &amp; Saldanha, L." box="[1041,1367,1785,1812]" pageId="9" pageNumber="44" pagination="719 - 747" refId="ref18341" refString="Merrett, N. R. &amp; Saldanha, L. (1985) Aspects of the morphology and ecology of some unusual deep-sea eels (Synaphobranchidae, Derichthyidae and Nettastomatidae) from the eastern North Atlantic. Journal of Fish Biology, 27, 719 - 747. https: // doi. org / 10.1111 / j. 1095 - 8649.1985. tb 03216. x" type="journal article" year="1985">Merrett and Saldanha (1985)</bibRefCitation>
.
</paragraph>
<paragraph id="8762368EFF99770FFF29F157FB6906B1" blockId="9.[151,1437,957,2028]" lastBlockId="10.[151,1437,150,1185]" lastPageId="10" lastPageNumber="45" pageId="9" pageNumber="44">
Body moderately elongate, laterally compressed posteriorly (
<figureCitation id="1FE62A0BFF99770CFC80F157FC580528" box="[878,968,1821,1848]" captionStart="FIGURE 5" captionStartId="10.[151,250,1902,1926]" captionTargetBox="[346,1240,1246,1872]" captionTargetId="figure-563@10.[340,1247,1239,1878]" captionTargetPageId="10" captionText="FIGURE 5. Ilyophis maclainei sp. nov. Referred Specimen, DISCOVERY Collection, Station 50517, 237 mm TL (A) Lateral view. (B) Head and cephalic lateralis system (from Merrett &amp; Saldanha 1985)" figureDoi="http://doi.org/10.5281/zenodo.13746111" httpUri="https://zenodo.org/record/13746111/files/figure.png" pageId="9" pageNumber="44">Fig. 5A</figureCitation>
). Body tapers gradually to caudal, deepest body depth anteriorly, near anus. Dorsal-fin origin near posterior half or just behind tip of pectoral fin, predorsal length ca. 1415 % TL. Trunk length moderate (ca. 1819 % TL), contained about 5 times in TL. Preanal length approximately 3 times in TL, preanal fin length about 31% of TL. Head length moderate (ca. 1012 % TL) (
<figureCitation id="1FE62A0BFF99770CFA81F1C3FF2B05D8" captionStart="FIGURE 5" captionStartId="10.[151,250,1902,1926]" captionTargetBox="[346,1240,1246,1872]" captionTargetId="figure-563@10.[340,1247,1239,1878]" captionTargetPageId="10" captionText="FIGURE 5. Ilyophis maclainei sp. nov. Referred Specimen, DISCOVERY Collection, Station 50517, 237 mm TL (A) Lateral view. (B) Head and cephalic lateralis system (from Merrett &amp; Saldanha 1985)" figureDoi="http://doi.org/10.5281/zenodo.13746111" httpUri="https://zenodo.org/record/13746111/files/figure.png" pageId="9" pageNumber="44">Fig. 5B</figureCitation>
). Gill slits crescentic and horizontal, located ventrally just anterior to and below pectoral fin base. Anterior nostril tubular, directed anterolaterally. Posterior nostrils round, just in front of eye. Eye relatively small (10.314.3 % HL). Pectoral fin moderate in size (3.23.7 % TL, 2536 % HL) with 15 rays. Gape extends to near posterior half of eye. Snout plicate (
<figureCitation id="1FE62A0BFF9A770FFE28F6F1FDB102C5" box="[454,545,187,213]" captionStart="FIGURE 6" captionStartId="11.[151,250,1052,1076]" captionTargetBox="[349,1238,190,1019]" captionTargetId="figure-19@11.[340,1247,181,1028]" captionTargetPageId="11" captionText="FIGURE 6. Ilyophis maclainei sp. nov. Holotype, BMNH 1995.4.19.31(DISCOVERY Collection, Station 50710), 240 mm TL (A) Snout ornamentation. (B) Lower jaw tip ornamentation (from Merrett &amp; Saldanha 1985)" figureDoi="http://doi.org/10.5281/zenodo.13746113" httpUri="https://zenodo.org/record/13746113/files/figure.png" pageId="10" pageNumber="45">Fig. 6A</figureCitation>
), three main plicae on each side of mid-line, outermost least developed. Outer margins of first and second plicae bear some four and eight papillae respectively, other minute papillae occur along all plical ridges. Tip of lower jaw also plicate (
<figureCitation id="1FE62A0BFF9A770FFD49F749FC90030D" box="[679,768,259,285]" captionStart="FIGURE 6" captionStartId="11.[151,250,1052,1076]" captionTargetBox="[349,1238,190,1019]" captionTargetId="figure-19@11.[340,1247,181,1028]" captionTargetPageId="11" captionText="FIGURE 6. Ilyophis maclainei sp. nov. Holotype, BMNH 1995.4.19.31(DISCOVERY Collection, Station 50710), 240 mm TL (A) Snout ornamentation. (B) Lower jaw tip ornamentation (from Merrett &amp; Saldanha 1985)" figureDoi="http://doi.org/10.5281/zenodo.13746113" httpUri="https://zenodo.org/record/13746113/files/figure.png" pageId="10" pageNumber="45">Fig. 6B</figureCitation>
), two main plicae on each side of mid-line, outermost least developed. Teeth conical, arranged in bands along maxillary, dentary and intermaxillary/vomer [
<figureCitation id="1FE62A0BFF9A770FFB38F76DFA8E0351" box="[1238,1310,295,321]" captionStart="FIGURE 7" captionStartId="11.[151,250,1918,1942]" captionTargetBox="[347,1240,1214,1885]" captionTargetId="figure-51@11.[340,1247,1206,1893]" captionTargetPageId="11" captionText="FIGURE 7. Ilyophis maclainei sp. nov. Referred Specimen, DISCOVERY Collection, Station 50517, 237 mm TL (A) Dentition of upper jaw. (B) Dentition of lower jaw (from Merrett &amp; Saldanha 1985)" figureDoi="http://doi.org/10.5281/zenodo.13746115" httpUri="https://zenodo.org/record/13746115/files/figure.png" pageId="10" pageNumber="45">Fig. 7</figureCitation>
]. Teeth not compound. Maxillary teeth in two to four rows (medially) reducing to one irregular row posteriorly, with anterior teeth somewhat enlarged. Dentary teeth in three to five irregular rows, with anterior ones (in two rows) larger. Intermaxillary teeth with 1618 strong teeth concentrically arranged around tooth patch with inner teeth slightly larger. Vomerine teeth abutting intermaxillary tooth patch with two irregular rows anteriorly (with 1011 teeth each) and converging to single row posteriorly (with three to four teeth), total number of vomerine teeth 2425. Both intermaxillary and vomerine teeth surrounded by papillae. Cephalic lateralis pores (
<figureCitation id="1FE62A0BFF9A770FFBB3F7B5FB280009" box="[1117,1208,510,537]" captionStart="FIGURE 5" captionStartId="10.[151,250,1902,1926]" captionTargetBox="[346,1240,1246,1872]" captionTargetId="figure-563@10.[340,1247,1239,1878]" captionTargetPageId="10" captionText="FIGURE 5. Ilyophis maclainei sp. nov. Referred Specimen, DISCOVERY Collection, Station 50517, 237 mm TL (A) Lateral view. (B) Head and cephalic lateralis system (from Merrett &amp; Saldanha 1985)" figureDoi="http://doi.org/10.5281/zenodo.13746111" httpUri="https://zenodo.org/record/13746111/files/figure.png" pageId="10" pageNumber="45">Fig. 5B</figureCitation>
): supraorbital pores three, first located between first and second plicae on snout. second on snout at level of the dorsal rim of anterior nostril, third on dorsal of snout above adnasal pore; infraorbital pores six, first (adnasal pore) just posterior to dorsal rim of anterior nostril, second on lip just posteroventral to base of anterior nostril, third on lip one third of the way between anterior and posterior nostrils, forth on lip anteroventral to posterior nostril, fifth on lip just ventral to the anterior margin of the eye, sixth on lip slightly posterior to posterior margin of eye; preoperculomandibular pores eight to nine, first very small and located within plicae at tip of lower jaw, remaining 67 mandibular pores spaced progressively far apart along jaw, last mandibular pore located approximately below rictus, 12 preopercular pores located posteriorly in the opercular region. Only the
<typeStatus id="5866882CFF9A770FFD3DF555FCA30129" box="[723,819,799,825]" pageId="10" pageNumber="45" type="paratype">paratype</typeStatus>
has 2 preopercular pores which are on the left side of the head and are set very close together in a pair. Lateral line incomplete, relatively long; pores extend to approximately 75% length of body. Three to six pores anterior to pectoral-fin base, 710 pores anterior to dorsal-fin origin, 2932 pores anterior to anus, 8894 total pores. Scales present all over body, except head, arranged broadly in classic basket-weave pattern (
<figureCitation id="1FE62A0BFF9A770FFE7CF5E5FD9F01D9" box="[402,527,943,969]" captionStart="FIGURE 8" captionStartId="12.[151,250,1052,1076]" captionTargetBox="[346,1242,185,1020]" captionTargetId="figure-34@12.[340,1247,179,1026]" captionTargetPageId="12" captionText="FIGURE 8. Ilyophis maclainei sp. nov. (A) Referred Specimen, DISCOVERY Collection, Station 50517. Distribution of type α and ss squamation on left and right side of specimen. (B) Same specimen. Detail of squamation from area of trunk region (scales of type α ventrally). (C) Same specimen. Detail of squamation from anal region (scales of type ss). (D) Holotype, BMNH 1995.4.19.31. Detail of squamation from region of end of caudal (scales of type ss) (modified from Merrett &amp; Saldanha 1985)" figureDoi="http://doi.org/10.5281/zenodo.13746119" httpUri="https://zenodo.org/record/13746119/files/figure.png" pageId="10" pageNumber="45">Fig. 8AD</figureCitation>
). Number, shape, and disposition variable. Scales generally elongate in shape, but length may vary between 23 scale width [type α scales (
<figureCitation id="1FE62A0BFF9A770FFCFFF599FCF701FD" box="[785,871,979,1005]" captionStart="FIGURE 8" captionStartId="12.[151,250,1052,1076]" captionTargetBox="[346,1242,185,1020]" captionTargetId="figure-34@12.[340,1247,179,1026]" captionTargetPageId="12" captionText="FIGURE 8. Ilyophis maclainei sp. nov. (A) Referred Specimen, DISCOVERY Collection, Station 50517. Distribution of type α and ss squamation on left and right side of specimen. (B) Same specimen. Detail of squamation from area of trunk region (scales of type α ventrally). (C) Same specimen. Detail of squamation from anal region (scales of type ss). (D) Holotype, BMNH 1995.4.19.31. Detail of squamation from region of end of caudal (scales of type ss) (modified from Merrett &amp; Saldanha 1985)" figureDoi="http://doi.org/10.5281/zenodo.13746119" httpUri="https://zenodo.org/record/13746119/files/figure.png" pageId="10" pageNumber="45">Fig. 8B</figureCitation>
) or 1015 or more times scale width [type β scales (
<figureCitation id="1FE62A0BFF9A770FFF71F5BDFE8B0601" box="[159,283,1015,1041]" captionStart="FIGURE 8" captionStartId="12.[151,250,1052,1076]" captionTargetBox="[346,1242,185,1020]" captionTargetId="figure-34@12.[340,1247,179,1026]" captionTargetPageId="12" captionText="FIGURE 8. Ilyophis maclainei sp. nov. (A) Referred Specimen, DISCOVERY Collection, Station 50517. Distribution of type α and ss squamation on left and right side of specimen. (B) Same specimen. Detail of squamation from area of trunk region (scales of type α ventrally). (C) Same specimen. Detail of squamation from anal region (scales of type ss). (D) Holotype, BMNH 1995.4.19.31. Detail of squamation from region of end of caudal (scales of type ss) (modified from Merrett &amp; Saldanha 1985)" figureDoi="http://doi.org/10.5281/zenodo.13746119" httpUri="https://zenodo.org/record/13746119/files/figure.png" pageId="10" pageNumber="45">Fig. 8CD</figureCitation>
). Disposition of scale types variable between specimens and not necessarily bilaterally symmetrical on an individual. Type α scales present in only
<specimenCount id="91DBFD07FF9A770FFD6CF251FC8C0625" box="[642,796,1051,1077]" count="1" pageId="10" pageNumber="45" type="generic">one specimen</specimenCount>
(
<figureCitation id="1FE62A0BFF9A770FFCC5F251FC140625" box="[811,900,1051,1077]" captionStart="FIGURE 8" captionStartId="12.[151,250,1052,1076]" captionTargetBox="[346,1242,185,1020]" captionTargetId="figure-34@12.[340,1247,179,1026]" captionTargetPageId="12" captionText="FIGURE 8. Ilyophis maclainei sp. nov. (A) Referred Specimen, DISCOVERY Collection, Station 50517. Distribution of type α and ss squamation on left and right side of specimen. (B) Same specimen. Detail of squamation from area of trunk region (scales of type α ventrally). (C) Same specimen. Detail of squamation from anal region (scales of type ss). (D) Holotype, BMNH 1995.4.19.31. Detail of squamation from region of end of caudal (scales of type ss) (modified from Merrett &amp; Saldanha 1985)" figureDoi="http://doi.org/10.5281/zenodo.13746119" httpUri="https://zenodo.org/record/13746119/files/figure.png" pageId="10" pageNumber="45">Fig. 8A</figureCitation>
), one patch on left side on belly posterior to left pectoral fin and two patches on right side, one between lateral line and dorsal profile, posterior to right pectoral fin, and second on beginning of caudal region between lateral line and ventral profile. Type β scales are most common and present on all
<specimenCount id="91DBFD07FF9A770FFE8AF2CDFD8B06B1" box="[356,539,1159,1185]" count="3" pageId="10" pageNumber="45" type="generic">three specimens</specimenCount>
. Vertebral numbers: predorsal 89, preanal 3132, total 131134.
</paragraph>
</subSubSection>
<caption id="D3A26606FF9A770FFF79F124FC3B05BA" ID-DOI="http://doi.org/10.5281/zenodo.13746111" ID-Zenodo-Dep="13746111" httpUri="https://zenodo.org/record/13746111/files/figure.png" pageId="10" pageNumber="45" startId="10.[151,250,1902,1926]" targetBox="[346,1240,1246,1872]" targetPageId="10" targetType="figure">
<paragraph id="8762368EFF9A770FFF79F124FC3B05BA" blockId="10.[151,1436,1902,1962]" pageId="10" pageNumber="45">
FIGURE 5.
<taxonomicName id="40DD4D0DFF9A770FFEF4F124FE460596" authorityName="Tighe &amp; Smith &amp; Merrett &amp; Frable &amp; Zajonz" authorityYear="2024" box="[282,470,1902,1926]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="10" pageNumber="45" phylum="Chordata" rank="species" species="maclainei" status="sp. nov.">Ilyophis maclainei</taxonomicName>
<taxonomicNameLabel id="AE9A57E7FF9A770FFE33F124FDBE0596" box="[477,558,1902,1926]" pageId="10" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
Referred Specimen, DISCOVERY Collection, Station 50517, 237 mm TL (A) Lateral view. (B) Head and cephalic lateralis system (from
<bibRefCitation id="E34C4B7FFF9A770FFD4EF1D8FC3405BA" author="Merrett, N. R. &amp; Saldanha, L." box="[672,932,1938,1962]" pageId="10" pageNumber="45" pagination="719 - 747" refId="ref18341" refString="Merrett, N. R. &amp; Saldanha, L. (1985) Aspects of the morphology and ecology of some unusual deep-sea eels (Synaphobranchidae, Derichthyidae and Nettastomatidae) from the eastern North Atlantic. Journal of Fish Biology, 27, 719 - 747. https: // doi. org / 10.1111 / j. 1095 - 8649.1985. tb 03216. x" type="journal article" year="1985">Merrett &amp; Saldanha 1985</bibRefCitation>
)
</paragraph>
</caption>
<caption id="D3A26606FF9B770EFF79F256FBD10648" ID-DOI="http://doi.org/10.5281/zenodo.13746113" ID-Zenodo-Dep="13746113" httpUri="https://zenodo.org/record/13746113/files/figure.png" pageId="11" pageNumber="46" startId="11.[151,250,1052,1076]" targetBox="[349,1238,190,1019]" targetPageId="11" targetType="figure">
<paragraph id="8762368EFF9B770EFF79F256FBD10648" blockId="11.[151,1436,1052,1112]" pageId="11" pageNumber="46">
FIGURE 6.
<taxonomicName id="40DD4D0DFF9B770EFEF6F256FE440624" authorityName="Tighe &amp; Smith &amp; Merrett &amp; Frable &amp; Zajonz" authorityYear="2024" box="[280,468,1052,1076]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="maclainei" status="sp. nov.">Ilyophis maclainei</taxonomicName>
<taxonomicNameLabel id="AE9A57E7FF9B770EFE34F256FDBA0624" box="[474,554,1052,1076]" pageId="11" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
Holotype, BMNH 1995.4.19.31(DISCOVERY Collection, Station 50710), 240 mm TL (A) Snout ornamentation. (B) Lower jaw tip ornamentation (from
<bibRefCitation id="E34C4B7FFF9B770EFCD8F20BFBAA0648" author="Merrett, N. R. &amp; Saldanha, L." box="[822,1082,1088,1112]" pageId="11" pageNumber="46" pagination="719 - 747" refId="ref18341" refString="Merrett, N. R. &amp; Saldanha, L. (1985) Aspects of the morphology and ecology of some unusual deep-sea eels (Synaphobranchidae, Derichthyidae and Nettastomatidae) from the eastern North Atlantic. Journal of Fish Biology, 27, 719 - 747. https: // doi. org / 10.1111 / j. 1095 - 8649.1985. tb 03216. x" type="journal article" year="1985">Merrett &amp; Saldanha 1985</bibRefCitation>
)
</paragraph>
</caption>
<caption id="D3A26606FF9B770EFF79F134FC1105AB" ID-DOI="http://doi.org/10.5281/zenodo.13746115" ID-Zenodo-Dep="13746115" httpUri="https://zenodo.org/record/13746115/files/figure.png" pageId="11" pageNumber="46" startId="11.[151,250,1918,1942]" targetBox="[347,1240,1214,1885]" targetPageId="11" targetType="figure">
<paragraph id="8762368EFF9B770EFF79F134FC1105AB" blockId="11.[151,1436,1918,1979]" pageId="11" pageNumber="46">
FIGURE 7.
<taxonomicName id="40DD4D0DFF9B770EFEF9F135FE420587" authorityName="Tighe &amp; Smith &amp; Merrett &amp; Frable &amp; Zajonz" authorityYear="2024" box="[279,466,1919,1943]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="11" pageNumber="46" phylum="Chordata" rank="species" species="maclainei" status="sp. nov.">Ilyophis maclainei</taxonomicName>
<taxonomicNameLabel id="AE9A57E7FF9B770EFE39F135FDB60587" box="[471,550,1919,1943]" pageId="11" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
Referred Specimen, DISCOVERY Collection, Station 50517, 237 mm TL (A) Dentition of upper jaw. (B) Dentition of lower jaw (from
<bibRefCitation id="E34C4B7FFF9B770EFD99F1E9FCEA05AB" author="Merrett, N. R. &amp; Saldanha, L." box="[631,890,1955,1979]" pageId="11" pageNumber="46" pagination="719 - 747" refId="ref18341" refString="Merrett, N. R. &amp; Saldanha, L. (1985) Aspects of the morphology and ecology of some unusual deep-sea eels (Synaphobranchidae, Derichthyidae and Nettastomatidae) from the eastern North Atlantic. Journal of Fish Biology, 27, 719 - 747. https: // doi. org / 10.1111 / j. 1095 - 8649.1985. tb 03216. x" type="journal article" year="1985">Merrett &amp; Saldanha 1985</bibRefCitation>
)
</paragraph>
</caption>
<caption id="D3A26606FF9C7709FF79F256FA1D06B1" ID-DOI="http://doi.org/10.5281/zenodo.13746119" ID-Zenodo-Dep="13746119" httpUri="https://zenodo.org/record/13746119/files/figure.png" pageId="12" pageNumber="47" startId="12.[151,250,1052,1076]" targetBox="[346,1242,185,1020]" targetPageId="12" targetType="figure">
<paragraph id="8762368EFF9C7709FF79F256FA1D06B1" blockId="12.[151,1437,1052,1185]" pageId="12" pageNumber="47">
FIGURE 8.
<taxonomicName id="40DD4D0DFF9C7709FEF4F257FE460625" authorityName="Tighe &amp; Smith &amp; Merrett &amp; Frable &amp; Zajonz" authorityYear="2024" box="[282,470,1053,1077]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="maclainei" status="sp. nov.">Ilyophis maclainei</taxonomicName>
<taxonomicNameLabel id="AE9A57E7FF9C7709FE32F257FDBD0625" box="[476,557,1053,1077]" pageId="12" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
(A) Referred Specimen, DISCOVERY Collection, Station 50517. Distribution of type α and ss squamation on left and right side of specimen. (B) Same specimen. Detail of squamation from area of trunk region (scales of type α ventrally). (C) Same specimen. Detail of squamation from anal region (scales of type ss). (D) Holotype, BMNH 1995.4.19.31. Detail of squamation from region of end of caudal (scales of type ss) (modified from
<bibRefCitation id="E34C4B7FFF9C7709FB6CF2C3FA1606B1" author="Merrett, N. R. &amp; Saldanha, L." box="[1154,1414,1161,1185]" pageId="12" pageNumber="47" pagination="719 - 747" refId="ref18341" refString="Merrett, N. R. &amp; Saldanha, L. (1985) Aspects of the morphology and ecology of some unusual deep-sea eels (Synaphobranchidae, Derichthyidae and Nettastomatidae) from the eastern North Atlantic. Journal of Fish Biology, 27, 719 - 747. https: // doi. org / 10.1111 / j. 1095 - 8649.1985. tb 03216. x" type="journal article" year="1985">Merrett &amp; Saldanha 1985</bibRefCitation>
)
</paragraph>
</caption>
<subSubSection id="CFC76505FF9C7709FF29F2B9FE630585" pageId="12" pageNumber="47" type="discussion">
<paragraph id="8762368EFF9C7709FF29F2B9FA0D07D1" blockId="12.[151,1437,1267,2013]" pageId="12" pageNumber="47">
Remarks. The Referred Specimen listed above (Discovery Collection Station 50517) was one of the original
<specimenCount id="91DBFD07FF9C7709FF79F35DFED80721" box="[151,328,1303,1329]" count="3" pageId="12" pageNumber="47" type="generic">three specimens</specimenCount>
described by
<bibRefCitation id="E34C4B7FFF9C7709FE35F35DFC960722" author="Merrett, N. R. &amp; Saldanha, L." box="[475,774,1303,1330]" pageId="12" pageNumber="47" pagination="719 - 747" refId="ref18341" refString="Merrett, N. R. &amp; Saldanha, L. (1985) Aspects of the morphology and ecology of some unusual deep-sea eels (Synaphobranchidae, Derichthyidae and Nettastomatidae) from the eastern North Atlantic. Journal of Fish Biology, 27, 719 - 747. https: // doi. org / 10.1111 / j. 1095 - 8649.1985. tb 03216. x" type="journal article" year="1985">Merrett &amp; Saldanha(1985)</bibRefCitation>
. This specimen was also examined by
<bibRefCitation id="E34C4B7FFF9C7709FB4CF35DFF750745" author="Sulak, K. J. &amp; Shcherbachev, Y. N." pageId="12" pageNumber="47" pagination="1158 - 1194" refId="ref19041" refString="Sulak, K. J. &amp; Shcherbachev, Y. N. (1997) Zoogeography and systematics of six deep-living genera of synaphobranchid eels, with a key to taxa and description of two new species of Ilyophis. Bulletin of Marine Science, 60 (3), 1158 - 1194." type="journal article" year="1997">Sulak&amp; Shcherbachev (1997)</bibRefCitation>
. Since then, most of the specimen has apparently been lost. Only the gonads of this female specimen are still in existence and are cataloged at the Natural History Museum as BMNH 1996.3.8.2. However, since much of the data on the northeastern Atlantic population provided by
<bibRefCitation id="E34C4B7FFF9C7709FD3BF3C9FB90078E" author="Merrett, N. R. &amp; Saldanha, L." box="[725,1024,1411,1438]" pageId="12" pageNumber="47" pagination="719 - 747" refId="ref18341" refString="Merrett, N. R. &amp; Saldanha, L. (1985) Aspects of the morphology and ecology of some unusual deep-sea eels (Synaphobranchidae, Derichthyidae and Nettastomatidae) from the eastern North Atlantic. Journal of Fish Biology, 27, 719 - 747. https: // doi. org / 10.1111 / j. 1095 - 8649.1985. tb 03216. x" type="journal article" year="1985">Merrett &amp; Saldanha (1985)</bibRefCitation>
was based on this specimen, we have included it in the description. If the body of this specimen is ever found, it should be reunited with BMNH 1996.3.8.2.
</paragraph>
<paragraph id="8762368EFF9C7709FF29F381FB190465" blockId="12.[151,1437,1267,2013]" pageId="12" pageNumber="47">
Comparative remarks.
<taxonomicName id="40DD4D0DFF9C7709FE38F381FD3107F5" authorityName="Tighe &amp; Smith &amp; Merrett &amp; Frable &amp; Zajonz" authorityYear="2024" box="[470,673,1483,1509]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="maclainei">Ilyophis maclainei</taxonomicName>
can be distinguished from all members of the genera
<taxonomicName id="40DD4D0DFF9C7709FB3DF381FA0507F5" authorityName="Robins &amp; Robins" authorityYear="1970" box="[1235,1429,1483,1509]" family="Synaphobranchidae" genus="Atractodenchelys" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="genus">Atractodenchelys</taxonomicName>
,
<taxonomicName id="40DD4D0DFF9C7709FF79F3BAFE970419" authorityName="Alcock" authorityYear="1889" box="[151,263,1520,1545]" family="Synaphobranchidae" genus="Dysomma" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="genus">Dysomma</taxonomicName>
<taxonomicName id="40DD4D0DFF9C7709FEE1F3BAFE080419" authorityName="Ginsburg" authorityYear="1951" box="[271,408,1520,1545]" family="Synaphobranchidae" genus="Dysommina" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="genus">Dysommina</taxonomicName>
, and
<taxonomicName id="40DD4D0DFF9C7709FE3AF3A5FDF60419" authorityName="Smith" authorityYear="1989" box="[468,614,1519,1545]" family="Synaphobranchidae" genus="Linkenchelys" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="genus">Linkenchelys</taxonomicName>
by the lack of compound teeth in the dentition (versus at least vomerine teeth compound).
<taxonomicName id="40DD4D0DFF9C7709FEB1F059FDBD043D" authorityName="Tighe &amp; Smith &amp; Merrett &amp; Frable &amp; Zajonz" authorityYear="2024" box="[351,557,1555,1581]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="maclainei">Ilyophis maclainei</taxonomicName>
is distinguished from
<taxonomicName id="40DD4D0DFF9C7709FCCDF059FC75043D" baseAuthorityName="Kamohara" baseAuthorityYear="1938" box="[803,997,1555,1581]" family="Synaphobranchidae" genus="Meadia" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="abyssalis">Meadia abyssalis</taxonomicName>
and
<taxonomicName id="40DD4D0DFF9C7709FBF6F05EFB11043D" authorityName="Mok, Lee &amp; Chan" authorityYear="1991" box="[1048,1153,1556,1581]" family="Synaphobranchidae" genus="Meadia" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="roseni">M. roseni</taxonomicName>
by its head length shorter than its trunk (versus head length longer than trunk). Finally, it can be distinguished from a third species in the genus
<taxonomicName id="40DD4D0DFF9C7709FF79F011FF7D0465" authorityName="Bohlke" authorityYear="1951" box="[151,237,1627,1653]" family="Synaphobranchidae" genus="Meadia" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="genus">Meadia</taxonomicName>
,
<taxonomicName id="40DD4D0DFF9C7709FF15F016FEF30465" authorityName="Vo &amp; Ho" authorityYear="2021" box="[251,355,1628,1653]" family="Synaphobranchidae" genus="Meadia" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="minor">M. minor</taxonomicName>
, by its higher vertebral number (131134 versus
<quantity id="40259B6BFF9C7709FC62F011FB9A0465" box="[908,1034,1627,1653]" metricMagnitude="0" metricUnit="m" metricValue="3.048" metricValueMax="3.0987999999999998" metricValueMin="2.9972" pageId="12" pageNumber="47" unit="in" value="120.0" valueMax="122.0" valueMin="118.0">118122 in</quantity>
<taxonomicName id="40DD4D0DFF9C7709FBFFF016FBE90465" authorityName="Vo &amp; Ho" authorityYear="2021" box="[1041,1145,1628,1653]" family="Synaphobranchidae" genus="Meadia" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="minor">M. minor</taxonomicName>
).
</paragraph>
<paragraph id="8762368EFF9C7709FF29F035FE630585" blockId="12.[151,1437,1267,2013]" pageId="12" pageNumber="47">
Morphometric, meristic, and morphological data for all described species of the genus
<taxonomicName id="40DD4D0DFF9C7709FB4AF035FB6E0489" authorityName="Gilbert" authorityYear="1891" box="[1188,1278,1663,1689]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="genus">Ilyophis</taxonomicName>
are presented in
<tableCitation id="CA5F0335FF9C7709FF5BF0E9FE9D04AD" box="[181,269,1699,1725]" captionStart="TABLE 2" captionStartId="7.[152,245,468,494]" captionTargetPageId="7" captionText="TABLE 2. Comparison of morphometric, meristic, and morphological data for all species of Ilyophis. For unique specimens (holotypes and paratypes), the bilateral pore counts are recorded left/right (if recorded in original description)." httpUri="http://table.plazi.org/id/D3A26606FF977702FF76F79EFC960002" pageId="12" pageNumber="47" tableUuid="D3A26606FF977702FF76F79EFC960002">Table 2</tableCitation>
. The presence of scales distinguishes
<taxonomicName id="40DD4D0DFF9C7709FD2EF0EEFCD704AD" authorityName="Tighe &amp; Smith &amp; Merrett &amp; Frable &amp; Zajonz" authorityYear="2024" box="[704,839,1699,1725]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="maclainei">I. maclainei</taxonomicName>
from other members of the genus
<taxonomicName id="40DD4D0DFF9C7709FB39F0E9FAA104AD" authorityName="Gilbert" authorityYear="1891" box="[1239,1329,1699,1725]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="genus">Ilyophis</taxonomicName>
except
<taxonomicName id="40DD4D0DFF9C7709FA62F0EEFE9204F1" authorityName="Gilbert" authorityYear="1891" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="brunneus">I. brunneus</taxonomicName>
, l.
<taxonomicName id="40DD4D0DFF9C7709FECCF08DFEE404F1" authorityName="Saldanha &amp; Merrett" authorityYear="1982" box="[290,372,1735,1761]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="blachei">blachei</taxonomicName>
and l.
<taxonomicName id="40DD4D0DFF9C7709FE51F08DFD9204F1" authorityName="Shcherbachev &amp; Sulak" authorityYear="1997" box="[447,514,1735,1761]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="nigeli">nigeli</taxonomicName>
.
<taxonomicName id="40DD4D0DFF9C7709FDE2F08DFD4A04F1" authorityName="Tighe &amp; Smith &amp; Merrett &amp; Frable &amp; Zajonz" authorityYear="2024" box="[524,730,1735,1761]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="maclainei">Ilyophis maclainei</taxonomicName>
is differentiated from those three species by its lower vertebrae number (131134 versus
<quantity id="40259B6BFF9C7709FE58F0A1FDA60515" box="[438,566,1771,1798]" metricMagnitude="0" metricUnit="m" metricValue="3.7592" metricValueMax="3.8354" metricValueMin="3.683" pageId="12" pageNumber="47" unit="in" value="148.0" valueMax="151.0" valueMin="145.0">145151 in</quantity>
<taxonomicName id="40DD4D0DFF9C7709FDD3F0A6FD500515" authorityName="Gilbert" authorityYear="1891" box="[573,704,1771,1797]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="brunneus">I. brunneus</taxonomicName>
,
<quantity id="40259B6BFF9C7709FD22F0A1FCDB0515" box="[716,843,1771,1797]" metricMagnitude="0" metricUnit="m" metricValue="4.6355" metricValueMax="4.7752" metricValueMin="4.4958" pageId="12" pageNumber="47" unit="in" value="182.5" valueMax="188.0" valueMin="177.0">177188 in</quantity>
l.
<taxonomicName id="40DD4D0DFF9C7709FC87F0A1FC2B0515" authorityName="Saldanha &amp; Merrett" authorityYear="1982" box="[873,955,1771,1797]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="blachei">blachei</taxonomicName>
and
<quantity id="40259B6BFF9C7709FC1DF0A1FBE20515" box="[1011,1138,1771,1798]" metricMagnitude="0" metricUnit="m" metricValue="3.7084" metricValueMax="3.8608" metricValueMin="3.556" pageId="12" pageNumber="47" unit="in" value="146.0" valueMax="152.0" valueMin="140.0">140152 in</quantity>
l.
<taxonomicName id="40DD4D0DFF9C7709FB7EF0A1FB400515" authorityName="Shcherbachev &amp; Sulak" authorityYear="1997" box="[1168,1232,1771,1797]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="nigeli">nigeli</taxonomicName>
).
<taxonomicName id="40DD4D0DFF9C7709FB06F0A6FAFD0515" authorityName="Tighe &amp; Smith &amp; Merrett &amp; Frable &amp; Zajonz" authorityYear="2024" box="[1256,1389,1771,1797]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="maclainei">I. maclainei</taxonomicName>
can be distinguished from
<taxonomicName id="40DD4D0DFF9C7709FE7DF15AFE400539" authorityName="C. H. Robins" authorityYear="1976" box="[403,464,1808,1833]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="arx">I. arx</taxonomicName>
with which it had previously been confused not only by the presence of scales (versus no scales in
<taxonomicName id="40DD4D0DFF9C7709FEF1F17EFECD055D" authorityName="C. H. Robins" authorityYear="1976" box="[287,349,1843,1869]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="arx">I. arx</taxonomicName>
), but also by its longer lateral line (ca. 7076% TL versus ca. 3546% TL in
<taxonomicName id="40DD4D0DFF9C7709FB5FF17EFB7D055D" authorityName="C. H. Robins" authorityYear="1976" box="[1201,1261,1844,1869]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="arx">I. arx</taxonomicName>
), the correlated higher number of lateral line pores (8894 versus
<quantity id="40259B6BFF9C7709FD37F11DFCAE0561" box="[729,830,1879,1905]" metricMagnitude="-1" metricUnit="m" metricValue="9.906" metricValueMax="11.937999999999999" metricValueMin="7.874" pageId="12" pageNumber="47" unit="in" value="39.0" valueMax="47.0" valueMin="31.0">3147 in</quantity>
<taxonomicName id="40DD4D0DFF9C7709FCA6F112FC180561" authorityName="C. H. Robins" authorityYear="1976" box="[840,904,1879,1905]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="arx">I. arx</taxonomicName>
), and a lower number of preanal fin vertebrae (3334 versus
<quantity id="40259B6BFF9C7709FED2F131FE0F0585" box="[316,415,1915,1941]" metricMagnitude="0" metricUnit="m" metricValue="1.0032999999999999" metricValueMax="1.0668" metricValueMin="0.9398" pageId="12" pageNumber="47" unit="in" value="39.5" valueMax="42.0" valueMin="37.0">3742 in</quantity>
<taxonomicName id="40DD4D0DFF9C7709FE48F136FE720585" authorityName="C. H. Robins" authorityYear="1976" box="[422,482,1916,1941]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="12" pageNumber="47" phylum="Chordata" rank="species" species="arx">I. arx</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="CFC76505FF9C7708FF29F1D5FC4A02C5" lastPageId="13" lastPageNumber="48" pageId="12" pageNumber="47" type="etymology">
<paragraph id="8762368EFF9C7708FF29F1D5FC4A02C5" blockId="12.[151,1437,1267,2013]" lastBlockId="13.[151,1437,151,357]" lastPageId="13" lastPageNumber="48" pageId="12" pageNumber="47">
Etymology. We take pleasure in naming this new species after our colleague James Maclaine, Senior Curator of Fishes at The Natural History Museum,
<collectingCountry id="FFCA761EFF9C7709FD68F189FCD905CD" box="[646,841,1987,2013]" name="United Kingdom" pageId="12" pageNumber="47">United Kingdom</collectingCountry>
, for his dedication to the collections under his care, and especially his hard work in incorporating the collections of the Institute of Oceanographic Sciences/National Oceanography Centre into the Natural History Museum,
<collectingCountry id="FFCA761EFF9D7708FCFBF6F1FC4502C5" box="[789,981,187,213]" name="United Kingdom" pageId="13" pageNumber="48">United Kingdom</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="CFC76505FF9D7708FF20F694FD8F0375" pageId="13" pageNumber="48" type="distribution">
<paragraph id="8762368EFF9D7708FF20F694FAFA030D" blockId="13.[151,1437,151,357]" pageId="13" pageNumber="48">
Distribution.
<taxonomicName id="40DD4D0DFF9D7708FE80F695FDAE02E9" authorityName="Tighe &amp; Smith &amp; Merrett &amp; Frable &amp; Zajonz" authorityYear="2024" box="[366,574,223,249]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="maclainei">Ilyophis maclainei</taxonomicName>
is known only from the
<specimenCount id="91DBFD07FF9D7708FCBFF695FB9502E9" box="[849,1029,223,249]" count="3" pageId="13" pageNumber="48" type="generic">three specimens</specimenCount>
which were collected in the vicinity of the Porcupine Sea-Bight and Goban Spur areas, off SW Ireland at a depth range of
<quantity id="40259B6BFF9D7708FBB6F748FA99030D" box="[1112,1289,258,285]" metricMagnitude="3" metricUnit="m" metricValue="3.2864999999999993" metricValueMax="4.788" metricValueMin="1.785" pageId="13" pageNumber="48" unit="m" value="3286.5" valueMax="4788.0" valueMin="1785.0">1785 to 4788 m</quantity>
(
<figureCitation id="1FE62A0BFF9D7708FAF6F749FACC030D" box="[1304,1372,259,285]" captionStart="FIGURE 9" captionStartId="13.[151,250,1501,1525]" captionTargetBox="[158,1429,385,1468]" captionTargetId="figure-106@13.[151,1436,378,1475]" captionTargetPageId="13" captionText="FIGURE 9. Distribution of Ilyophis maclainei in the eastern North Atlantic Ocean. Open star is the type locality. Closed stars are other study material." figureDoi="http://doi.org/10.5281/zenodo.13746121" httpUri="https://zenodo.org/record/13746121/files/figure.png" pageId="13" pageNumber="48">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8762368EFF9D7708FF29F76DFD8F0375" blockId="13.[151,1437,151,357]" pageId="13" pageNumber="48">Whether the species is endemic to this area or might be more widely distributed in the slope region of the eastern North Atlantic is unknown.</paragraph>
</subSubSection>
<caption id="D3A26606FF9D7708FF79F397FE1E040A" ID-DOI="http://doi.org/10.5281/zenodo.13746121" ID-Zenodo-Dep="13746121" httpUri="https://zenodo.org/record/13746121/files/figure.png" pageId="13" pageNumber="48" startId="13.[151,250,1501,1525]" targetBox="[158,1429,385,1468]" targetPageId="13" targetType="figure">
<paragraph id="8762368EFF9D7708FF79F397FE1E040A" blockId="13.[151,1436,1501,1562]" pageId="13" pageNumber="48">
FIGURE 9. Distribution of
<taxonomicName id="40DD4D0DFF9D7708FE59F394FDE407E6" authorityName="Tighe &amp; Smith &amp; Merrett &amp; Frable &amp; Zajonz" authorityYear="2024" box="[439,628,1502,1526]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="maclainei">Ilyophis maclainei</taxonomicName>
in the eastern North Atlantic Ocean. Open star is the type locality. Closed stars are other study material.
</paragraph>
</caption>
<subSubSection id="CFC76505FF9D7708FF29F026FD6B05B6" pageId="13" pageNumber="48" type="materials_examined">
<paragraph id="8762368EFF9D7708FF29F026FD6B05B6" blockId="13.[151,1437,1644,1958]" pageId="13" pageNumber="48">
Comparative Specimens Examined.
<taxonomicName id="40DD4D0DFF9D7708FD9CF026FCA90496" authorityName="Gilbert" authorityYear="1891" box="[626,825,1644,1670]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="brunneus">Ilyophis brunneus</taxonomicName>
:
<materialsCitation id="37B53CD3FF9D7708FCA8F026FD8D04DE" collectionCode="USNM" country="San Cristobal Island" county="Albatross station" elevation="1160" latitude="-0.60833335" location="Eastern Pacific Ocean" longLatPrecision="21" longitude="-89.316666" municipality="Near" pageId="13" pageNumber="48" specimenCode="USNM 44403" specimenCount="1" typeStatus="holotype">
<specimenCode id="D77B9EF5FF9D7708FCA8F026FC720496" box="[838,994,1644,1670]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="48" type="Museum">USNM 44403</specimenCode>
(
<typeStatus id="5866882CFF9D7708FC01F026FBC50496" box="[1007,1109,1644,1670]" pageId="13" pageNumber="48" type="holotype">holotype</typeStatus>
,
<quantity id="40259B6BFF9D7708FBB1F026FB2A0496" box="[1119,1210,1644,1670]" metricMagnitude="-1" metricUnit="m" metricValue="3.8" pageId="13" pageNumber="48" unit="mm" value="380.0">380 mm</quantity>
TL),
<location id="82026055FF9D7708FB1BF026FF7104BA" LSID="urn:lsid:plazi:treatment:0F748798FF997708FF79F6DDFD6B05B6:82026055FF9D7708FB1BF026FF7104BA" country="San Cristobal Island" county="Albatross station" latitude="-0.60833335" longLatPrecision="21" longitude="-89.316666" municipality="Near" name="Eastern Pacific Ocean" pageId="13" pageNumber="48">Eastern Pacific Ocean</location>
,
<location id="82026055FF9D7708FF05F0DAFE2904BA" LSID="urn:lsid:plazi:treatment:0F748798FF997708FF79F6DDFD6B05B6:82026055FF9D7708FF05F0DAFE2904BA" box="[235,441,1680,1706]" country="San Cristobal Island" county="Albatross station" latitude="-0.60833335" longLatPrecision="21" longitude="-89.316666" municipality="Near" name="Galapagos Islands" pageId="13" pageNumber="48">Galápagos Islands</location>
,
<collectingMunicipality id="6706ACF4FF9D7708FE2CF0DAFE6804BA" box="[450,504,1680,1706]" pageId="13" pageNumber="48">Near</collectingMunicipality>
<collectingCountry id="FFCA761EFF9D7708FE10F0DAFD4D04BA" box="[510,733,1680,1706]" name="San Cristobal Island" pageId="13" pageNumber="48">San Cristóbal Island</collectingCountry>
[Chatham Island],
<geoCoordinate id="E2E95049FF9D7708FC5FF0DAFBA404BA" box="[945,1076,1680,1706]" degrees="0" direction="south" minutes="36" orientation="latitude" pageId="13" pageNumber="48" precision="15" seconds="30" value="-0.60833335">0° 36' 30&quot; S</geoCoordinate>
,
<geoCoordinate id="E2E95049FF9D7708FBD1F0DAFB4904BA" box="[1087,1241,1680,1706]" degrees="89" direction="west" minutes="19" orientation="longitude" pageId="13" pageNumber="48" precision="15" seconds="00" value="-89.316666">89° 19' 00&quot; W</geoCoordinate>
,
<collectingCounty id="6E034E02FF9D7708FB0FF0DAFA0D04BA" box="[1249,1437,1680,1706]" pageId="13" pageNumber="48">Albatross station</collectingCounty>
2808, depth 634 fathoms (
<quantity id="40259B6BFF9D7708FE55F0FEFD8404DE" box="[443,532,1716,1742]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="13" pageNumber="48" unit="m" value="1160.0">
<elevation id="0CF0D1BDFF9D7708FE55F0FEFD8404DE" box="[443,532,1716,1742]" metricMagnitude="3" metricUnit="m" metricValue="1.16" pageId="13" pageNumber="48" unit="m" value="1160.0">1160 m</elevation>
</quantity>
)
</materialsCitation>
.
<materialsCitation id="37B53CD3FF9D7708FDC9F0FEFD5904E2" collectionCode="USNM" country="San Cristobal Island" county="Oregon station" elevation="1143" latitude="28.966667" location="Gulf of Mexico" longLatPrecision="1229" longitude="-88.183334" municipality="Mississippi Delta" pageId="13" pageNumber="48" specimenCode="USNM 185665" specimenCount="1" typeStatus="holotype">
<specimenCode id="D77B9EF5FF9D7708FDC9F0FEFD4204DF" box="[551,722,1716,1743]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="48" type="Museum">USNM 185665</specimenCode>
(
<quantity id="40259B6BFF9D7708FD0AF0FEFCCA04DE" box="[740,858,1716,1742]" metricMagnitude="0" metricUnit="m" metricValue="1.3119999999999998" pageId="13" pageNumber="48" unit="mm" value="1312.0">1, 312 mm</quantity>
TL),
<location id="82026055FF9D7708FC77F0FEFBD904DE" LSID="urn:lsid:plazi:treatment:0F748798FF997708FF79F6DDFD6B05B6:82026055FF9D7708FC77F0FEFBD904DE" box="[921,1097,1716,1742]" country="San Cristobal Island" county="Oregon station" latitude="28.966667" longLatPrecision="1229" longitude="-88.183334" municipality="Mississippi Delta" name="Gulf of Mexico" pageId="13" pageNumber="48">Gulf of Mexico</location>
, off
<collectingMunicipality id="6706ACF4FF9D7708FB95F0FEFAD004DE" box="[1147,1344,1716,1742]" pageId="13" pageNumber="48">Mississippi Delta</collectingMunicipality>
,
<geoCoordinate id="E2E95049FF9D7708FAA5F0FEFF3E04E2" degrees="28" direction="north" minutes="58" orientation="latitude" pageId="13" pageNumber="48" precision="925" value="28.966667">28° 58 N</geoCoordinate>
,
<geoCoordinate id="E2E95049FF9D7708FF54F092FEBD04E2" box="[186,301,1752,1778]" degrees="88" direction="west" minutes="11" orientation="longitude" pageId="13" pageNumber="48" precision="925" value="-88.183334">88° 11 W</geoCoordinate>
,
<collectingCounty id="6E034E02FF9D7708FED6F092FE4F04E2" box="[312,479,1752,1778]" pageId="13" pageNumber="48">Oregon station</collectingCounty>
2822, depth
<quantity id="40259B6BFF9D7708FD9CF092FD5904E2" box="[626,713,1752,1778]" metricMagnitude="3" metricUnit="m" metricValue="1.143" pageId="13" pageNumber="48" unit="m" value="1143.0">
<elevation id="0CF0D1BDFF9D7708FD9CF092FD5904E2" box="[626,713,1752,1778]" metricMagnitude="3" metricUnit="m" metricValue="1.143" pageId="13" pageNumber="48" unit="m" value="1143.0">1143 m</elevation>
</quantity>
</materialsCitation>
.
<materialsCitation id="37B53CD3FF9D7708FD3BF092FCB80506" collectionCode="USNM" country="San Cristobal Island" county="Oregon station" elevation="1646" latitude="28.916666" location="Gulf of Mexico" longLatPrecision="1230" longitude="-87.816666" municipality="Alabama" pageId="13" pageNumber="48" specimenCode="USNM 195901" specimenCount="1" typeStatus="holotype">
<specimenCode id="D77B9EF5FF9D7708FD3BF092FC1104E3" box="[725,897,1752,1779]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="48" type="Museum">USNM 195901</specimenCode>
(
<quantity id="40259B6BFF9D7708FC7AF092FB9C04E2" box="[916,1036,1752,1778]" metricMagnitude="0" metricUnit="m" metricValue="1.414" pageId="13" pageNumber="48" unit="mm" value="1414.0">1, 414 mm</quantity>
TL),
<location id="82026055FF9D7708FBA2F092FB6E04E2" LSID="urn:lsid:plazi:treatment:0F748798FF997708FF79F6DDFD6B05B6:82026055FF9D7708FBA2F092FB6E04E2" box="[1100,1278,1752,1778]" country="San Cristobal Island" county="Oregon station" latitude="28.916666" longLatPrecision="1230" longitude="-87.816666" municipality="Alabama" name="Gulf of Mexico" pageId="13" pageNumber="48">Gulf of Mexico</location>
, off
<collectingMunicipality id="6706ACF4FF9D7708FADEF092FA0804E2" box="[1328,1432,1752,1778]" pageId="13" pageNumber="48">Alabama</collectingMunicipality>
,
<geoCoordinate id="E2E95049FF9D7708FF79F0B6FE980506" box="[151,264,1788,1815]" degrees="28" direction="north" minutes="55" orientation="latitude" pageId="13" pageNumber="48" precision="925" value="28.916666">28° 55 N</geoCoordinate>
,
<geoCoordinate id="E2E95049FF9D7708FEFAF0B6FE180506" box="[276,392,1788,1814]" degrees="87" direction="west" minutes="49" orientation="longitude" pageId="13" pageNumber="48" precision="925" value="-87.816666">87° 49 W</geoCoordinate>
,
<collectingCounty id="6E034E02FF9D7708FE7AF0B6FDAB0506" box="[404,571,1788,1814]" pageId="13" pageNumber="48">Oregon station</collectingCounty>
2022, depth
<quantity id="40259B6BFF9D7708FD21F0B6FCB80506" box="[719,808,1788,1814]" metricMagnitude="3" metricUnit="m" metricValue="1.646" pageId="13" pageNumber="48" unit="m" value="1646.0">
<elevation id="0CF0D1BDFF9D7708FD21F0B6FCB80506" box="[719,808,1788,1814]" metricMagnitude="3" metricUnit="m" metricValue="1.646" pageId="13" pageNumber="48" unit="m" value="1646.0">1646 m</elevation>
</quantity>
</materialsCitation>
.
<materialsCitation id="37B53CD3FF9D7708FCDDF0B6FBF2052A" collectionCode="USNM" country="San Cristobal Island" latitude="24.4" location="North Atlantic Ocean" longLatPrecision="1251" longitude="-80.0" municipality="Cay Sal Bank." pageId="13" pageNumber="48" specimenCode="USNM 197115" specimenCount="1" typeStatus="holotype">
<specimenCode id="D77B9EF5FF9D7708FCDDF0B6FC4F0507" box="[819,991,1788,1815]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="48" type="Museum">USNM 197115</specimenCode>
(
<quantity id="40259B6BFF9D7708FC1CF0B6FBFA0506" box="[1010,1130,1788,1814]" metricMagnitude="0" metricUnit="m" metricValue="1.403" pageId="13" pageNumber="48" unit="mm" value="1403.0">1, 403 mm</quantity>
TL)
<location id="82026055FF9D7708FB4AF0B6FA080506" LSID="urn:lsid:plazi:treatment:0F748798FF997708FF79F6DDFD6B05B6:82026055FF9D7708FB4AF0B6FA080506" box="[1188,1432,1788,1814]" country="San Cristobal Island" latitude="24.4" longLatPrecision="1251" longitude="-80.0" municipality="Cay Sal Bank." name="North Atlantic Ocean" pageId="13" pageNumber="48">North Atlantic Ocean</location>
, Straits of Florida, north of
<collectingMunicipality id="6706ACF4FF9D7708FE51F16AFDCF052A" box="[447,607,1824,1850]" pageId="13" pageNumber="48">Cay Sal Bank.</collectingMunicipality>
<geoCoordinate id="E2E95049FF9D7708FD8AF16AFD42052A" box="[612,722,1824,1850]" degrees="24" direction="north" minutes="24" orientation="latitude" pageId="13" pageNumber="48" precision="925" value="24.4">24° 24 N</geoCoordinate>
,
<geoCoordinate id="E2E95049FF9D7708FD32F16AFCDD052A" box="[732,845,1824,1850]" degrees="80" direction="west" minutes="00" orientation="longitude" pageId="13" pageNumber="48" precision="925" value="-80.0">80° 00 W</geoCoordinate>
, depth 400400 fathoms
</materialsCitation>
.
<materialsCitation id="37B53CD3FF9D7708FB85F16AFBE5054E" collectionCode="USNM" country="Venezuela" elevation="1006" latitude="11.733334" location="Southern Caribbean Sea" longLatPrecision="1295" longitude="-68.71667" pageId="13" pageNumber="48" specimenCode="USNM 212185" specimenCount="1" typeStatus="holotype">
<specimenCode id="D77B9EF5FF9D7708FB85F16AFA86052B" box="[1131,1302,1824,1851]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="48" type="Museum">USNM 212185</specimenCode>
(
<quantity id="40259B6BFF9D7708FAC6F16AFA0D052A" box="[1320,1437,1824,1850]" metricMagnitude="0" metricUnit="m" metricValue="1.262" pageId="13" pageNumber="48" unit="mm" value="1262.0">1, 262 mm</quantity>
TL),
<location id="82026055FF9D7708FF21F10EFE4E054E" LSID="urn:lsid:plazi:treatment:0F748798FF997708FF79F6DDFD6B05B6:82026055FF9D7708FF21F10EFE4E054E" box="[207,478,1860,1886]" country="Venezuela" latitude="11.733334" longLatPrecision="1295" longitude="-68.71667" name="Southern Caribbean Sea" pageId="13" pageNumber="48">Southern Caribbean Sea</location>
, off western
<collectingCountry id="FFCA761EFF9D7708FD85F10EFD71054E" box="[619,737,1860,1886]" name="Venezuela" pageId="13" pageNumber="48">Venezuela</collectingCountry>
,
<geoCoordinate id="E2E95049FF9D7708FD04F10EFCC6054E" box="[746,854,1860,1886]" degrees="11" direction="north" minutes="44" orientation="latitude" pageId="13" pageNumber="48" precision="925" value="11.733334">11° 44 N</geoCoordinate>
,
<geoCoordinate id="E2E95049FF9D7708FC8EF10EFC41054E" box="[864,977,1860,1886]" degrees="68" direction="west" minutes="43" orientation="longitude" pageId="13" pageNumber="48" precision="925" value="-68.71667">68° 43 W</geoCoordinate>
, depth
<quantity id="40259B6BFF9D7708FBF0F10EFBE5054E" box="[1054,1141,1860,1886]" metricMagnitude="3" metricUnit="m" metricValue="1.0059999999999998" pageId="13" pageNumber="48" unit="m" value="1006.0">
<elevation id="0CF0D1BDFF9D7708FBF0F10EFBE5054E" box="[1054,1141,1860,1886]" metricMagnitude="3" metricUnit="m" metricValue="1.0059999999999998" pageId="13" pageNumber="48" unit="m" value="1006.0">1006 m</elevation>
</quantity>
</materialsCitation>
.
<taxonomicName id="40DD4D0DFF9D7708FB91F10EFADD054E" authorityName="Karmovskaya &amp; Parin" authorityYear="1999" box="[1151,1357,1860,1886]" family="Synaphobranchidae" genus="Ilyophis" kingdom="Animalia" order="Anguilliformes" pageId="13" pageNumber="48" phylum="Chordata" rank="species" species="saldanhai">Ilyophis saldanhai</taxonomicName>
:
<materialsCitation id="37B53CD3FF9D7708FAB5F10EFE5F0592" collectionCode="MMF" country="Venezuela" pageId="13" pageNumber="48" specimenCode="MMF 27080" specimenCount="1" typeStatus="holotype">
<specimenCode id="D77B9EF5FF9D7708FAB5F10EFF4D0592" collectionCode="MMF" country="Portugal" httpUri="http://grbio.org/cool/mme0-gx6r" name="Museu Municipal do Funchal" pageId="13" pageNumber="48">MMF 27080</specimenCode>
(2, 354
<quantity id="40259B6BFF9D7708FEAEF122FE0C0592" box="[320,412,1896,1923]" metricMagnitude="-1" metricUnit="m" metricValue="3.82" pageId="13" pageNumber="48" unit="mm" value="382.0">382 mm</quantity>
TL)
</materialsCitation>
,
<materialsCitation id="37B53CD3FF9D7708FE37F122FD6705B6" collectionCode="TL" country="Venezuela" county="Broken Spur Hydrothermal Vent Field." elevation="3020" latitude="29.133333" location="North Atlantic Ocean" municipality="Mid-Atlantic Ridge" pageId="13" pageNumber="48" specimenCode="MMF 27081" specimenCount="1" typeStatus="holotype">
<specimenCode id="D77B9EF5FF9D7708FE37F122FDF70592" box="[473,615,1896,1922]" collectionCode="MMF" country="Portugal" httpUri="http://grbio.org/cool/mme0-gx6r" name="Museu Municipal do Funchal" pageId="13" pageNumber="48">MMF 27081</specimenCode>
(
<quantity id="40259B6BFF9D7708FD96F122FD7D0592" box="[632,749,1896,1922]" metricMagnitude="0" metricUnit="m" metricValue="1.409" pageId="13" pageNumber="48" unit="mm" value="1409.0">1, 409 mm</quantity>
<collectionCode id="E1CCAE4BFF9D7708FD1DF122FC8A0592" box="[755,794,1896,1922]" pageId="13" pageNumber="48">TL</collectionCode>
);
<location id="82026055FF9D7708FCC2F122FB8C0592" LSID="urn:lsid:plazi:treatment:0F748798FF997708FF79F6DDFD6B05B6:82026055FF9D7708FCC2F122FB8C0592" box="[812,1052,1896,1922]" country="Venezuela" county="Broken Spur Hydrothermal Vent Field." latitude="29.133333" municipality="Mid-Atlantic Ridge" name="North Atlantic Ocean" pageId="13" pageNumber="48">North Atlantic Ocean</location>
, Mid-Atlantic Ridge, Broken Spur Hydrothermal Vent Field. 29 08N, 43 13” W.
<quantity id="40259B6BFF9D7708FD71F1C6FD6705B6" box="[671,759,1932,1958]" metricMagnitude="3" metricUnit="m" metricValue="3.02" pageId="13" pageNumber="48" unit="m" value="3020.0">
<elevation id="0CF0D1BDFF9D7708FD71F1C6FD6705B6" box="[671,759,1932,1958]" metricMagnitude="3" metricUnit="m" metricValue="3.02" pageId="13" pageNumber="48" unit="m" value="3020.0">3020 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>