treatments-xml/data/03/C0/3B/03C03B6BFF88FFE3FF7B2E9CFBE0FDF2.xml
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<document id="8DA42C8D794CD0474A972FB9F8497E74" ID-CLB-Dataset="2431" ID-DOI="10.11646/zootaxa.5093.1.1" ID-GBIF-Dataset="0faf68be-3777-4466-8cd5-9eb8ca72a239" ID-ISSN="1175-5326" ID-Zenodo-Dep="5901006" ID-ZooBank="C2F5E368-CB15-4207-9944-E52C190FBF20" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1643101690477" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2022" docId="03C03B6BFF88FFE3FF7B2E9CFBE0FDF2" docLanguage="en" docName="zootaxa.5093.1.1.pdf" docOrigin="Zootaxa 5093 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euconnus (Euconnus) raucus Sharp 1886" docType="treatment" docVersion="8" lastPageNumber="22" masterDocId="FFF94313FF9BFFF6FFEC2B57FFE6FF93" masterDocTitle="Euconnus Thomson of Japan: redescriptions of species established by Reitter, Sharp and Franz, new synonyms, and summary of current state of knowledge (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="20" updateTime="1699292035325" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="19398EF186D4F231E3A216DD6358393D">Euconnus Thomson of Japan: redescriptions of species established by Reitter, Sharp and Franz, new synonyms, and summary of current state of knowledge (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<treatment id="03C03B6BFF88FFE3FF7B2E9CFBE0FDF2" ID-DOI="http://doi.org/10.5281/zenodo.5901026" ID-GBIF-Taxon="192597248" ID-Zenodo-Dep="5901026" LSID="urn:lsid:plazi:treatment:03C03B6BFF88FFE3FF7B2E9CFBE0FDF2" httpUri="http://treatment.plazi.org/id/03C03B6BFF88FFE3FF7B2E9CFBE0FDF2" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20">
<subSubSection id="C373D9F6FF88FFE5FF7B2E9CFDBFFA77" box="[151,601,1482,1509]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8BD68A7DFF88FFE5FF7B2E9CFDBFFA77" blockId="19.[151,601,1482,1509]" box="[151,601,1482,1509]" pageId="19" pageNumber="20">
<heading id="D09E3D11FF88FFE5FF7B2E9CFDBFFA77" bold="true" box="[151,601,1482,1509]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<taxonomicName id="4C69F1FEFF88FFE5FF7B2E9CFDBFFA77" ID-CoL="9G95N" authority="Sharp" authorityName="Sharp" authorityYear="1886" box="[151,601,1482,1509]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="raucus" subGenus="Euconnus">
<emphasis id="B91D566FFF88FFE5FF7B2E9CFDBFFA77" bold="true" box="[151,601,1482,1509]" pageId="19" pageNumber="20">
<emphasis id="B91D566FFF88FFE5FF7B2E9CFEF5FA76" bold="true" box="[151,275,1483,1509]" italics="true" pageId="19" pageNumber="20">Euconnus</emphasis>
(
<emphasis id="B91D566FFF88FFE5FEC82E9CFE46FA76" bold="true" box="[292,416,1483,1509]" italics="true" pageId="19" pageNumber="20">Euconnus</emphasis>
)
<emphasis id="B91D566FFF88FFE5FE5D2E9CFDE2FA76" bold="true" box="[433,516,1483,1509]" italics="true" pageId="19" pageNumber="20">raucus</emphasis>
Sharp
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C373D9F6FF88FFE5FF2B2D47FEC1F9D8" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="8BD68A7DFF88FFE5FF2B2D47FC90F9B4" blockId="19.[151,886,1551,1611]" box="[199,886,1551,1575]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="AB79AD53FF88FFE5FF2B2D47FC90F9B4" box="[199,886,1551,1575]" pageId="19" pageNumber="20">
<taxonomicName id="4C69F1FEFF88FFE5FF2B2D47FE1CF9B4" ID-CoL="9G95N" authority="Sharp, 1886" authorityName="Sharp" authorityYear="1886" box="[199,506,1551,1575]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="raucus">
<emphasis id="B91D566FFF88FFE5FF2B2D47FE90F9B4" box="[199,374,1552,1575]" italics="true" pageId="19" pageNumber="20">Euconnus raucus</emphasis>
<bibRefCitation id="EFF8F78CFF88FFE5FE912D58FE1CF9B4" author="Sharp, D." box="[381,506,1551,1575]" pageId="19" pageNumber="20" pagination="46 - 51" refId="ref26383" refString="Sharp, D. (1886) The Scydmaenidae of Japan. Entomologist's Monthly Magazine, 23, 46 - 51." type="journal article" year="1886">Sharp, 1886</bibRefCitation>
</taxonomicName>
: Redescribed by
<treatmentCitation id="0AC8AC6CFF88FFE5FD442D58FC94F9B4" author="Hoshina, H." box="[680,882,1551,1575]" page="99" pageId="19" pageNumber="20" year="2019">
<bibRefCitation id="EFF8F78CFF88FFE5FD442D58FC94F9B4" author="Hoshina, H." box="[680,882,1551,1575]" pageId="19" pageNumber="20" pagination="97 - 100" refId="ref24924" refString="Hoshina, H. (2019 b) Taxonomic note on the genus Euconnus (Coleoptera: Staphylinidae: Scydmaeninae) from Japan (II). Japanese Journal of Systematic Entomology, 25 (1), 97 - 100." type="journal article" year="2019">Hoshina, 2019b: 99</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD68A7DFF88FFE5FF7B2D66FEC1F9D8" blockId="19.[151,886,1551,1611]" box="[151,295,1585,1611]" pageId="19" pageNumber="20">
(
<figureCitation id="135296F8FF88FFE5FF732D66FEF9F9D8" box="[159,287,1585,1611]" captionStart="FIGURES 7071" captionStartId="20.[151,264,1621,1645]" captionTargetBox="[171,1426,925,1591]" captionTargetId="figure-283@20.[151,1436,921,1598]" captionTargetPageId="20" captionText="FIGURES 7071. Euconnus raucus, female. Dorsal habitus (70) and annotated mounting card with historical labels (71) of holotype." figureDoi="http://doi.org/10.5281/zenodo.5901034" httpUri="https://zenodo.org/record/5901034/files/figure.png" pageId="19" pageNumber="20">Figs 7071</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C373D9F6FF88FFE5FF7B2D24FD0FF88E" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="8BD68A7DFF88FFE5FF7B2D24FB06F946" blockId="19.[151,1437,1650,2037]" pageId="19" pageNumber="20">
<emphasis id="B91D566FFF88FFE5FF7B2D24FE78F91F" bold="true" box="[151,414,1650,1677]" pageId="19" pageNumber="20">Type material studied</emphasis>
.
<materialsCitation id="3B018020FF88FFE5FE422D24FB06F946" ID-GBIF-Occurrence="3454958301" collectionCode="NHM" collectorName="G. Lewis." country="Japan" location="Nagasaki" pageId="19" pageNumber="20" specimenCount="1" specimenCount-female="1" stateProvince="Nagasaki" typeStatus="holotype">
<emphasis id="B91D566FFF88FFE5FE422D24FDFDF91E" bold="true" box="[430,539,1651,1677]" pageId="19" pageNumber="20">
<typeStatus id="54D234DFFF88FFE5FE422D24FDFDF91E" box="[430,539,1651,1677]" pageId="19" pageNumber="20" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="9D6F41F4FF88FFE5FDC62D24FDD9F91E" box="[554,575,1651,1677]" pageId="19" pageNumber="20" type="female"></specimenCount>
(
<figureCitation id="135296F8FF88FFE5FDBC2D24FD42F91E" box="[592,676,1651,1677]" captionStart="FIGURES 7071" captionStartId="20.[151,264,1621,1645]" captionTargetBox="[171,1426,925,1591]" captionTargetId="figure-283@20.[151,1436,921,1598]" captionTargetPageId="20" captionText="FIGURES 7071. Euconnus raucus, female. Dorsal habitus (70) and annotated mounting card with historical labels (71) of holotype." figureDoi="http://doi.org/10.5281/zenodo.5901034" httpUri="https://zenodo.org/record/5901034/files/figure.png" pageId="19" pageNumber="20">Fig. 70</figureCitation>
), mounted on thick card with annotation: “
<taxonomicName id="4C69F1FEFF88FFE5FB662D24FB1DF91E" box="[1162,1275,1651,1677]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Euconnus</taxonomicName>
/ raucus Type / D.S. /
<collectingRegion id="49AD449FFF88FFE5FF032DC0FEBDF922" box="[239,347,1687,1713]" country="Japan" name="Nagasaki" pageId="19" pageNumber="20">Nagasaki</collectingRegion>
. / 26.3.81. Lewis.”, with two labels (
<figureCitation id="135296F8FF88FFE5FD142DC0FCABF922" box="[760,845,1687,1713]" captionStart="FIGURES 7071" captionStartId="20.[151,264,1621,1645]" captionTargetBox="[171,1426,925,1591]" captionTargetId="figure-283@20.[151,1436,921,1598]" captionTargetPageId="20" captionText="FIGURES 7071. Euconnus raucus, female. Dorsal habitus (70) and annotated mounting card with historical labels (71) of holotype." figureDoi="http://doi.org/10.5281/zenodo.5901034" httpUri="https://zenodo.org/record/5901034/files/figure.png" pageId="19" pageNumber="20">Fig. 71</figureCitation>
): “
<collectingCountry id="F37ECAEDFF88FFE5FC872DC0FC48F922" box="[875,942,1687,1713]" name="Japan" pageId="19" pageNumber="20">Japan</collectingCountry>
/
<collectorName id="269CEFABFF88FFE5FC282DC0FBD5F922" box="[964,1075,1687,1713]" pageId="19" pageNumber="20">G. Lewis.</collectorName>
/ 1910-320” [white with orange horizontal line, printed], and circular label “Type / H.T.” [white with red margin, printed] (
<collectionCode id="ED7812B8FF88FFE5FB612DECFB33F946" box="[1165,1237,1723,1749]" pageId="19" pageNumber="20">NHM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BD68A7DFF88FFE5FF2B2D88FD0FF88E" blockId="19.[151,1437,1650,2037]" pageId="19" pageNumber="20">
<emphasis id="B91D566FFF88FFE5FF2B2D88FE50F96B" bold="true" box="[199,438,1758,1785]" pageId="19" pageNumber="20">Emended diagnosis.</emphasis>
The only known female of this species matches diagnosis of
<taxonomicName id="4C69F1FEFF88FFE5FB9D2D88FB01F96A" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[1137,1255,1759,1785]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF88FFE5FB9D2D88FB01F96A" box="[1137,1255,1759,1785]" italics="true" pageId="19" pageNumber="20">
E.
<emphasis id="B91D566FFF88FFE5FB7D2D88FB01F96A" box="[1169,1255,1759,1785]" pageId="19" pageNumber="20">fustiger</emphasis>
</emphasis>
</taxonomicName>
, with exception of distinctly larger body: BL
<quantity id="4C912798FF88FFE5FE312C55FDA2F88E" box="[477,580,1794,1821]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="19" pageNumber="20" unit="mm" value="1.75">1.75 mm</quantity>
. See Remarks.
</paragraph>
</subSubSection>
<subSubSection id="C373D9F6FF88FFE2FF2B2C71FDCBFCCD" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="description">
<paragraph id="8BD68A7DFF88FFE5FF2B2C71FCB9F8F6" blockId="19.[151,1437,1650,2037]" pageId="19" pageNumber="20">
<emphasis id="B91D566FFF88FFE5FF2B2C71FE93F8D3" bold="true" box="[199,373,1830,1856]" pageId="19" pageNumber="20">Redescription.</emphasis>
Body of female (
<figureCitation id="135296F8FF88FFE5FDAF2C70FD71F8D2" box="[579,663,1831,1857]" captionStart="FIGURES 7071" captionStartId="20.[151,264,1621,1645]" captionTargetBox="[171,1426,925,1591]" captionTargetId="figure-283@20.[151,1436,921,1598]" captionTargetPageId="20" captionText="FIGURES 7071. Euconnus raucus, female. Dorsal habitus (70) and annotated mounting card with historical labels (71) of holotype." figureDoi="http://doi.org/10.5281/zenodo.5901034" httpUri="https://zenodo.org/record/5901034/files/figure.png" pageId="19" pageNumber="20">Fig. 70</figureCitation>
) moderately slender, strongly convex, BL
<quantity id="4C912798FF88FFE5FB682C71FB0BF8D2" box="[1156,1261,1830,1857]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="19" pageNumber="20" unit="mm" value="1.75">1.75 mm</quantity>
; cuticle glossy, pigmentation dark brown, vestiture of setae lighter than cuticle.
</paragraph>
<paragraph id="8BD68A7DFF88FFE2FF2B2C38FD06FE1A" blockId="19.[151,1437,1650,2037]" lastBlockId="20.[151,1437,151,897]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
Head rhomboidal, distinctly elongate and broadest at eyes, HL
<quantity id="4C912798FF88FFE5FC4B2C38FBF7F81A" box="[935,1041,1903,1929]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="19" pageNumber="20" unit="mm" value="0.4">0.40 mm</quantity>
, HW
<quantity id="4C912798FF88FFE5FBB42C39FB25F81A" box="[1112,1219,1902,1929]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="19" pageNumber="20" unit="mm" value="0.35">0.35 mm</quantity>
; tempora in dorsal view about 2.5 times as long as eyes and strongly converging posterad; vertex and frons confluent, weakly and evenly convex, posterior margin of vertex rounded, slightly bulging posterodorsad; supraantennal tubercles barely marked; frons between antennae steeply declining; clypeus unmodified. Eyes moderately large, finely faceted, distinctly but not strongly projecting laterad from the head silhouette, in lateral view oval. Punctures on vertex and frons inconspicuous, fine and sparse; setae short and sparse, suberect, tempora densely covered with thick bristles directed laterocaudad. Antennae moderately slender, with compact proximal portion of funicle and moderately loosely assembled, strongly broadened and sharply delimited tetramerous club, AnL
<quantity id="4C912798FF8FFFE2FBA62A54FB57FE8E" box="[1098,1201,259,285]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="20" pageNumber="21" unit="mm" value="0.7">0.70 mm</quantity>
; antennomeres 1 and 2 each elongate, 36 each about as long as broad, 7 slightly narrower than 6, indistinctly elongate, 8 much broader than 7, slightly transverse, 9 and 10 each strongly transverse and indistinctly broader, but distinctly shorter than 8, 11 indistinctly narrower than 10, slightly transverse.
</paragraph>
<paragraph id="8BD68A7DFF8FFFE2FF2B2AC4FE0BFDF2" blockId="20.[151,1437,151,897]" pageId="20" pageNumber="21">
Pronotum subconical with weakly rounded sides, broadest at base, PL
<quantity id="4C912798FF8FFFE2FC372AC5FBA7FE3E" box="[987,1089,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="20" pageNumber="21" unit="mm" value="0.45">0.45 mm</quantity>
, PW
<quantity id="4C912798FF8FFFE2FB912AC4FB02FE3E" box="[1149,1252,403,429]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="20" pageNumber="21" unit="mm" value="0.48">0.48 mm</quantity>
; anterior margin nearly straight and much shorter than posterior margin, anterior corners weakly marked, sides of pronotum weakly, evenly rounded and strongly converging anterad; posterior corners distinctly obtuse-angled, well-marked; posterior margin nearly straight. Pronotal base with two lateral pairs of tiny and shallow pits, transverse groove and sublateral carinae lacking. Disc dorsally covered with fine, inconspicuous punctures and moderately dense thin suberect setae, sides with dense thick bristles.
</paragraph>
<paragraph id="8BD68A7DFF8FFFE2FF2B293CFBF9FD62" blockId="20.[151,1437,151,897]" pageId="20" pageNumber="21">
Elytra oval, broadest distinctly in front of middle, EL
<quantity id="4C912798FF8FFFE2FCAF293CFC4AFD16" box="[835,940,619,645]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="21" unit="mm" value="0.9">0.90 mm</quantity>
, EW
<quantity id="4C912798FF8FFFE2FC1D293CFBBDFD16" box="[1009,1115,619,645]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="20" pageNumber="21" unit="mm" value="0.73">0.73 mm</quantity>
, EI 1.24; basal impressions short and shallow, humeral calli moderately strongly elevated, elongate, each delimited from adscutellar region by shallow elongate impression running posterolaterad, elytral apices separately rounded. Punctures on elytral disc fine, inconspicuous; setae long, moderately dense, suberect. Hind wings lacking.
</paragraph>
<paragraph id="8BD68A7DFF8FFFE2FF2B29ACFD3EFC86" blockId="20.[151,1437,151,897]" box="[199,728,763,789]" pageId="20" pageNumber="21">Legs moderately long and slender, unmodified.</paragraph>
<paragraph id="8BD68A7DFF8FFFE2FF2B2848FE20FCAA" blockId="20.[151,1437,151,897]" box="[199,454,799,825]" pageId="20" pageNumber="21">Abdomen unmodified.</paragraph>
<paragraph id="8BD68A7DFF8FFFE2FF2B2814FDCBFCCD" blockId="20.[151,1437,151,897]" box="[199,557,835,862]" pageId="20" pageNumber="21">Male. Unknown (see Remarks).</paragraph>
</subSubSection>
<subSubSection id="C373D9F6FF8FFFE2FF2B2831FDF3FC12" box="[199,533,870,897]" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8BD68A7DFF8FFFE2FF2B2831FDF3FC12" blockId="20.[151,1437,151,897]" box="[199,533,870,897]" pageId="20" pageNumber="21">
<emphasis id="B91D566FFF8FFFE2FF2B2831FEBFFC13" bold="true" box="[199,345,870,896]" pageId="20" pageNumber="21">Distribution</emphasis>
.
<collectingCountry id="F37ECAEDFF8FFFE2FE8B2830FE4DFC12" box="[359,427,871,897]" name="Japan" pageId="20" pageNumber="21">Japan</collectingCountry>
: Kyushu.
</paragraph>
</subSubSection>
<caption id="DF16DAF5FF8FFFE2FF7B2D02FF11F902" ID-DOI="http://doi.org/10.5281/zenodo.5901034" ID-Zenodo-Dep="5901034" httpUri="https://zenodo.org/record/5901034/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,264,1621,1645]" targetBox="[171,1426,925,1591]" targetPageId="20">
<paragraph id="8BD68A7DFF8FFFE2FF7B2D02FF11F902" blockId="20.[151,1436,1621,1681]" pageId="20" pageNumber="21">
<emphasis id="B91D566FFF8FFFE2FF7B2D02FEB0F9FE" bold="true" box="[151,342,1621,1645]" pageId="20" pageNumber="21">FIGURES 7071.</emphasis>
<taxonomicName id="4C69F1FEFF8FFFE2FEB22D01FDE9F9FE" authorityName="Sharp" authorityYear="1886" box="[350,527,1622,1645]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="raucus">
<emphasis id="B91D566FFF8FFFE2FEB22D01FDE9F9FE" box="[350,527,1622,1645]" italics="true" pageId="20" pageNumber="21">Euconnus raucus</emphasis>
</taxonomicName>
, female. Dorsal habitus (70) and annotated mounting card with historical labels (71) of holotype.
</paragraph>
</caption>
<subSubSection id="C373D9F6FF8FFFE3FF2B2D97FBE0FDF2" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="discussion">
<paragraph id="8BD68A7DFF8FFFE2FF2B2D97FB76F869" blockId="20.[151,1437,1727,2042]" pageId="20" pageNumber="21">
<emphasis id="B91D566FFF8FFFE2FF2B2D97FEDDF949" bold="true" box="[199,315,1728,1754]" pageId="20" pageNumber="21">Remarks.</emphasis>
This species was redescribed by
<bibRefCitation id="EFF8F78CFF8FFFE2FD362DE8FC78F949" author="Hoshina, H." box="[730,926,1727,1754]" pageId="20" pageNumber="21" pagination="97 - 100" refId="ref24924" refString="Hoshina, H. (2019 b) Taxonomic note on the genus Euconnus (Coleoptera: Staphylinidae: Scydmaeninae) from Japan (II). Japanese Journal of Systematic Entomology, 25 (1), 97 - 100." type="journal article" year="2019">Hoshina (2019b)</bibRefCitation>
. However, the single known type specimen preserved at NHM is a rather unremarkable female and it remains unknown how a male has been unambiguously assigned to this species. I have seen many similar specimens from Kyushu and Honshu, mostly from mountainous areas, matching the body length and form of the
<typeStatus id="54D234DFFF8FFFE2FD0A2C7CFCAEF8D6" box="[742,840,1835,1861]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
female of
<taxonomicName id="4C69F1FEFF8FFFE2FC3C2C7BFBA6F8D6" authorityName="Sharp" authorityYear="1886" box="[976,1088,1836,1861]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="raucus">
<emphasis id="B91D566FFF8FFFE2FC3C2C7BFBA6F8D6" box="[976,1088,1836,1861]" italics="true" pageId="20" pageNumber="21">E. raucus</emphasis>
</taxonomicName>
, but they represented several species whose males differ in genital structures. However, it is possible that these differences will turn out to be a morphocline, as seems to be the case for the smaller-bodied and broadly distributed
<taxonomicName id="4C69F1FEFF8FFFE2FBBB2C23FB2BF81D" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[1111,1229,1908,1934]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF8FFFE2FBBB2C23FB2BF81D" box="[1111,1229,1908,1934]" italics="true" pageId="20" pageNumber="21">
E.
<emphasis id="B91D566FFF8FFFE2FB9B2C23FB2BF81D" box="[1143,1229,1908,1934]" pageId="20" pageNumber="21">fustiger</emphasis>
</emphasis>
</taxonomicName>
. The female of
<taxonomicName id="4C69F1FEFF8FFFE2FA682C23FF05F822" authorityName="Sharp" authorityYear="1886" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="raucus">
<emphasis id="B91D566FFF8FFFE2FA682C23FF05F822" italics="true" pageId="20" pageNumber="21">E. raucus</emphasis>
</taxonomicName>
seems to differ from females of
<taxonomicName id="4C69F1FEFF8FFFE2FDBC2CCFFD23F821" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[592,709,1944,1970]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF8FFFE2FDBC2CCFFD23F821" box="[592,709,1944,1970]" italics="true" pageId="20" pageNumber="21">
E.
<emphasis id="B91D566FFF8FFFE2FD832CCFFD23F821" box="[623,709,1944,1970]" pageId="20" pageNumber="21">fustiger</emphasis>
</emphasis>
</taxonomicName>
in a slightly stouter head and a shorter antennomere 11 (which is slightly transverse, whereas in most females of
<taxonomicName id="4C69F1FEFF8FFFE2FD482CEBFCFEF845" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[676,792,1980,2006]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF8FFFE2FD482CEBFCFEF845" box="[676,792,1980,2006]" italics="true" pageId="20" pageNumber="21">
E.
<emphasis id="B91D566FFF8FFFE2FD2E2CEBFCFEF845" box="[706,792,1980,2006]" pageId="20" pageNumber="21">fustiger</emphasis>
</emphasis>
</taxonomicName>
it is at least as long as broad). A larger sample of similarly large-bodied specimens from many localities is necessary to clarify the status of
<taxonomicName id="4C69F1FEFF8FFFE2FBF22CB7FB6FF86A" authorityName="Sharp" authorityYear="1886" box="[1054,1161,2016,2041]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="raucus">
<emphasis id="B91D566FFF8FFFE2FBF22CB7FB6FF86A" box="[1054,1161,2016,2041]" italics="true" pageId="20" pageNumber="21">E. raucus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD68A7DFF8EFFE3FF2B2BC0FB16FD8A" blockId="21.[151,1437,151,609]" pageId="21" pageNumber="22">
If the
<bibRefCitation id="EFF8F78CFF8EFFE3FEE52BC0FE3FFF21" author="Hoshina, H." box="[265,473,151,178]" pageId="21" pageNumber="22" pagination="97 - 100" refId="ref24924" refString="Hoshina, H. (2019 b) Taxonomic note on the genus Euconnus (Coleoptera: Staphylinidae: Scydmaeninae) from Japan (II). Japanese Journal of Systematic Entomology, 25 (1), 97 - 100." type="journal article" year="2019">Hoshinas (2019b)</bibRefCitation>
identification is correct, then the aedeagus of
<taxonomicName id="4C69F1FEFF8EFFE3FC372BC0FBA0FF23" authorityName="Sharp" authorityYear="1886" box="[987,1094,151,176]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="raucus">
<emphasis id="B91D566FFF8EFFE3FC372BC0FBA0FF23" box="[987,1094,151,176]" italics="true" pageId="21" pageNumber="22">E. raucus</emphasis>
</taxonomicName>
differs from that of
<taxonomicName id="4C69F1FEFF8EFFE3FAC42BC0FA7BFF22" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[1320,1437,151,177]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF8EFFE3FAC42BC0FA7BFF22" box="[1320,1437,151,177]" italics="true" pageId="21" pageNumber="22">
E.
<emphasis id="B91D566FFF8EFFE3FAAB2BC0FA7BFF22" box="[1351,1437,151,177]" pageId="21" pageNumber="22">fustiger</emphasis>
</emphasis>
</taxonomicName>
in a long median, asymmetrical pointed endophallic sclerite with a subapical projection. However, this illustration of Hoshina also shows a genital organ distorted by improper preparation, with the endophallus partly extruded, which may in future make identifications based on non-distorted preparations difficult, if not impossible. Hoshina later described several extremely similar, large-bodied
<taxonomicName id="4C69F1FEFF8EFFE3FCFE2A70FC3DFED2" box="[786,987,295,321]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B91D566FFF8EFFE3FCFE2A70FC66FED3" box="[786,896,295,320]" italics="true" pageId="21" pageNumber="22">Euconnus</emphasis>
species
</taxonomicName>
with similar (and often also distorted) aedeagi. I found it impossible to clarify whether these are distinct species or some are identical with
<taxonomicName id="4C69F1FEFF8EFFE3FAE62A1CFA90FEF7" authorityName="Sharp" authorityYear="1886" box="[1290,1398,331,356]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="raucus">
<emphasis id="B91D566FFF8EFFE3FAE62A1CFA90FEF7" box="[1290,1398,331,356]" italics="true" pageId="21" pageNumber="22">E. raucus</emphasis>
</taxonomicName>
, or what actually
<taxonomicName id="4C69F1FEFF8EFFE3FED42A38FE40FE1B" authorityName="Sharp" authorityYear="1886" box="[312,422,367,392]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="raucus">
<emphasis id="B91D566FFF8EFFE3FED42A38FE40FE1B" box="[312,422,367,392]" italics="true" pageId="21" pageNumber="22">E. raucus</emphasis>
</taxonomicName>
is (i.e., whether it is a distinct species or the
<typeStatus id="54D234DFFF8EFFE3FC582A38FBF0FE1A" box="[948,1046,367,393]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
female is only an unusually large specimen of the variable
<taxonomicName id="4C69F1FEFF8EFFE3FE5F2AC4FDCEFE3E" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[435,552,403,429]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF8EFFE3FE5F2AC4FDCEFE3E" box="[435,552,403,429]" italics="true" pageId="21" pageNumber="22">
E.
<emphasis id="B91D566FFF8EFFE3FE3E2AC4FDCEFE3E" box="[466,552,403,429]" pageId="21" pageNumber="22">fustiger</emphasis>
</emphasis>
</taxonomicName>
). None of the large-bodied,
<taxonomicName id="4C69F1FEFF8EFFE3FC8B2AC4FC3BFE3E" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[871,989,403,429]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fustiger">
<emphasis id="B91D566FFF8EFFE3FC8B2AC4FC3BFE3E" box="[871,989,403,429]" italics="true" pageId="21" pageNumber="22">
E.
<emphasis id="B91D566FFF8EFFE3FC6B2AC4FC3BFE3E" box="[903,989,403,429]" pageId="21" pageNumber="22">fustiger</emphasis>
</emphasis>
</taxonomicName>
-like species in my collection match the aedeagal structures illustrated by Hoshina for
<taxonomicName id="4C69F1FEFF8EFFE3FD492AE0FCF6FE43" authorityName="Sharp" authorityYear="1886" box="[677,784,439,464]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="raucus">
<emphasis id="B91D566FFF8EFFE3FD492AE0FCF6FE43" box="[677,784,439,464]" italics="true" pageId="21" pageNumber="22">E. raucus</emphasis>
</taxonomicName>
and his other similar species, suggesting that this species complex is large and taxonomically challenging. The current situation caused by carelessly describing new species based on distorted aedeagi may already pose problems impossible to solve by morphological study.
</paragraph>
<paragraph id="8BD68A7DFF8EFFE3FF2B2974FBE0FDF2" blockId="21.[151,1437,151,609]" pageId="21" pageNumber="22">
<taxonomicName id="4C69F1FEFF8EFFE3FF2B2974FE61FDAF" authorityName="Sharp" authorityYear="1886" box="[199,391,547,572]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="raucus">
<emphasis id="B91D566FFF8EFFE3FF2B2974FE61FDAF" box="[199,391,547,572]" italics="true" pageId="21" pageNumber="22">Euconnus raucus</emphasis>
</taxonomicName>
is here placed in
<taxonomicName id="4C69F1FEFF8EFFE3FDAA2974FD52FDAF" box="[582,692,547,572]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis id="B91D566FFF8EFFE3FDAA2974FD52FDAF" box="[582,692,547,572]" italics="true" pageId="21" pageNumber="22">Euconnus</emphasis>
</taxonomicName>
(s. str.), based on the tetramerous antennal club, the pronotum bellshaped, broadest behind middle, and lack of the median antebasal pronotal pit.
</paragraph>
</subSubSection>
</treatment>
</document>