217 lines
27 KiB
XML
217 lines
27 KiB
XML
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<mods:title id="AB062270C8F58A05A3193863F3EC4036">Euconnus Thomson of Japan: redescriptions of species established by Reitter, Sharp and Franz, new synonyms, and summary of current state of knowledge (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<paragraph id="8BD68A7DFF83FFEEFF7B2C6CFD21F8C6" blockId="24.[151,711,1851,1878]" box="[151,711,1851,1878]" pageId="24" pageNumber="25">
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<taxonomicName id="4C69F1FEFF83FFEEFF7B2C6CFD21F8C6" ID-CoL="9G8V6" authority="Hoshina" authorityName="Hoshina" authorityYear="2013" box="[151,711,1851,1878]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kumejimensis" subGenus="Euconnus">
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<emphasis id="B91D566FFF83FFEEFF7B2C6CFD21F8C6" bold="true" box="[151,711,1851,1878]" pageId="24" pageNumber="25">
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<emphasis id="B91D566FFF83FFEEFF7B2C6CFEF5F8C6" bold="true" box="[151,275,1851,1877]" italics="true" pageId="24" pageNumber="25">Euconnus</emphasis>
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(
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<emphasis id="B91D566FFF83FFEEFEC82C6CFE46F8C6" bold="true" box="[292,416,1851,1877]" italics="true" pageId="24" pageNumber="25">Euconnus</emphasis>
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)
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<emphasis id="B91D566FFF83FFEEFE5D2C6CFDB1F8C5" bold="true" box="[433,599,1851,1878]" italics="true" pageId="24" pageNumber="25">kumejimensis</emphasis>
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Hoshina
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C373D9F6FF83FFEEFF2B2CD7FC02F86A" pageId="24" pageNumber="25" type="reference_group">
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<paragraph id="8BD68A7DFF83FFEEFF2B2CD7FE2EF847" blockId="24.[199,1437,1919,2041]" pageId="24" pageNumber="25">
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<treatmentCitationGroup id="AB79AD53FF83FFEEFF2B2CD7FE2EF847" pageId="24" pageNumber="25">
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<treatmentCitation id="0AC8AC6CFF83FFEEFF2B2CD7FCCDF804" author="Hoshina, H." box="[199,811,1919,1944]" page="285" pageId="24" pageNumber="25" year="2013">
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<taxonomicName id="4C69F1FEFF83FFEEFF2B2CD7FCCDF804" ID-CoL="9G8V6" authority="Hoshina, 2013: 285" authorityName="Hoshina" authorityPageNumber="285" authorityYear="2013" box="[199,811,1919,1944]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kumejimensis" subGenus="Euconophron">
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<emphasis id="B91D566FFF83FFEEFF2B2CD7FECDF804" box="[199,299,1920,1943]" italics="true" pageId="24" pageNumber="25">Euconnus</emphasis>
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(
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||
<emphasis id="B91D566FFF83FFEEFED62CD7FE25F80B" box="[314,451,1920,1944]" italics="true" pageId="24" pageNumber="25">Euconophron</emphasis>
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)
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||
<emphasis id="B91D566FFF83FFEEFE3E2CD7FDBCF80B" box="[466,602,1920,1944]" italics="true" pageId="24" pageNumber="25">kumejimensis</emphasis>
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<bibRefCitation id="EFF8F78CFF83FFEEFD8C2CD7FCCDF804" author="Hoshina, H." box="[608,811,1919,1944]" pageId="24" pageNumber="25" pagination="285 - 288" refId="ref24786" refString="Hoshina, H. (2013) New record of the subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Kumejima Island, the Ryukyus, Japan, with description of a new species. Japanese Journal of Systematic Entomology, 19 (2), 285 - 288." type="journal article" year="2013">Hoshina, 2013: 285</bibRefCitation>
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</taxonomicName>
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. Aedeagus also illustrated in
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<bibRefCitation id="EFF8F78CFF83FFEEFBBE2CD7FB12F80B" author="Hoshina, H." box="[1106,1268,1920,1944]" pageId="24" pageNumber="25" pagination="49 - 65" refId="ref24877" refString="Hoshina, H. (2019 a) Seven new species of the subfamily Scydmaeninae (Coleoptera: Staphylinidae) from Japan. The Memoirs of the Research and Education Center for Regional Environment, Fukui University, 25, 49 - 65. [2018]" type="journal article" year="2019">Hoshina, 2019a</bibRefCitation>
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: figs 10‒12.
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<taxonomicName id="4C69F1FEFF83FFEEFF2B2CC8FD5FF825" ID-CoL="9G8V6" authority="Hoshina" authorityName="Hoshina" authorityYear="2013" box="[199,697,1950,1974]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kumejimensis" subGenus="Eupentarius">
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<emphasis id="B91D566FFF83FFEEFF2B2CC8FECDF825" box="[199,299,1951,1974]" italics="true" pageId="24" pageNumber="25">Euconnus</emphasis>
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||
(
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||
<emphasis id="B91D566FFF83FFEEFED22CC8FE5CF825" box="[318,442,1951,1974]" italics="true" pageId="24" pageNumber="25">Eupentarius</emphasis>
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||
)
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<emphasis id="B91D566FFF83FFEEFE212CC9FDB3F825" box="[461,597,1950,1974]" italics="true" pageId="24" pageNumber="25">kumejimensis</emphasis>
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Hoshina
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</taxonomicName>
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;
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<treatmentCitation id="0AC8AC6CFF83FFEEFD2B2CC9FC23F825" author="Jaloszynski, P." box="[711,965,1950,1974]" page="156" pageId="24" pageNumber="25" year="2021">
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<bibRefCitation id="EFF8F78CFF83FFEEFD2B2CC9FC23F825" author="Jaloszynski, P." box="[711,965,1950,1974]" pageId="24" pageNumber="25" pagination="145 - 160" refId="ref25844" refString="Jaloszynski, P. (2021 a) Taxonomy of " Euconnus complex ". Part XXII. Status of subgenera Allonapochus Franz, Dimorphoconnus Franz, Eupentarius Reitter, and Myrmecodytes Croissandeau (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4966 (2), 145 - 160. https: // doi. org / 10.11646 / zootaxa. 4966.2.3" type="journal article" year="2021">Jałoszyński, 2021a: 156</bibRefCitation>
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</treatmentCitation>
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||
, implied, by placing
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<taxonomicName id="4C69F1FEFF83FFEEFB402CC9FAD3F825" ID-CoL="9G88P" box="[1196,1333,1950,1974]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Euconophron">
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<emphasis id="B91D566FFF83FFEEFB402CC9FAD3F825" box="[1196,1333,1950,1974]" italics="true" pageId="24" pageNumber="25">Euconophron</emphasis>
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</taxonomicName>
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||
as junior synonym of
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||
<emphasis id="B91D566FFF83FFEEFEAA2CE9FE24F846" box="[326,450,1982,2005]" italics="true" pageId="24" pageNumber="25">Eupentarius</emphasis>
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||
.
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BD68A7DFF83FFEEFF2B2CB5FC02F86A" blockId="24.[199,1437,1919,2041]" box="[199,996,2017,2041]" pageId="24" pageNumber="25">
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||
<taxonomicName id="4C69F1FEFF83FFEEFF2B2CB5FD75F86A" ID-CoL="9G8V6" authority="Hoshina" authorityName="Hoshina" authorityYear="2013" box="[199,659,2017,2041]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kumejimensis" subGenus="Euconnus">
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||
<emphasis id="B91D566FFF83FFEEFF2B2CB5FECDF86A" box="[199,299,2018,2041]" italics="true" pageId="24" pageNumber="25">Euconnus</emphasis>
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||
(
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||
<emphasis id="B91D566FFF83FFEEFED62CB5FE78F86A" box="[314,414,2018,2041]" italics="true" pageId="24" pageNumber="25">Euconnus</emphasis>
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||
)
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<emphasis id="B91D566FFF83FFEEFE412CB6FDD3F86A" box="[429,565,2017,2041]" italics="true" pageId="24" pageNumber="25">kumejimensis</emphasis>
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Hoshina
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||
</taxonomicName>
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||
;
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<emphasis id="B91D566FFF83FFEEFD712CB6FC02F86A" bold="true" box="[669,996,2017,2041]" pageId="24" pageNumber="25">
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||
here placed in
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<taxonomicName id="4C69F1FEFF83FFEEFCD72CB6FC40F86B" ID-CoL="8VRNT" box="[827,934,2017,2040]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus" sensu="stricto">
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||
<emphasis id="B91D566FFF83FFEEFCD72CB6FC40F86B" bold="true" box="[827,934,2017,2040]" italics="true" pageId="24" pageNumber="25">Euconnus</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A22EEB14FF83FFEEFC402CB6FC02F86A" box="[940,996,2017,2041]" pageId="24" pageNumber="25" sensu="stricto">s. str.</taxonomicNameLabel>
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||
</emphasis>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C373D9F6FF82FFEFFF7B2BC0FC0CFAEA" pageId="25" pageNumber="26" type="discussion">
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<paragraph id="8BD68A7DFF82FFEFFF7B2BC0FA94FB82" blockId="25.[151,1437,151,1401]" pageId="25" pageNumber="26">
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<emphasis id="B91D566FFF82FFEFFF7B2BC0FEEDFF22" bold="true" box="[151,267,151,177]" pageId="25" pageNumber="26">Remarks.</emphasis>
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This species is known to occur on Kumejima, a small island near Okinawa-jima, the main island of the Ryukyu Archipelago. It belongs to the
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<taxonomicName id="4C69F1FEFF82FFEFFDAA2BECFD16FF46" authorityName="Vit" authorityYear="2011" box="[582,752,187,213]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="alesi">
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<emphasis id="B91D566FFF82FFEFFDAA2BECFD16FF46" box="[582,752,187,213]" italics="true" pageId="25" pageNumber="26">Euconnus alesi</emphasis>
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</taxonomicName>
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species group, which is broadly distributed throughout entire Ryukyuan Arch, from Koshikishima rettô near the western cost of Kyushu to Yonaguni-jima in the south, and
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<taxonomicName id="4C69F1FEFF82FFEFFAA62B88FEE6FE8E" authority="Vit, 2011" authorityName="Vit" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="alesi">
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<emphasis id="B91D566FFF82FFEFFAA62B88FA7AFF6A" box="[1354,1436,223,249]" italics="true" pageId="25" pageNumber="26">E. alesi</emphasis>
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||
<bibRefCitation id="EFF8F78CFF82FFEFFF7B2A54FEE6FE8E" author="Vit, S." box="[151,256,259,285]" pageId="25" pageNumber="26" pagination="443 - 448" refId="ref26567" refString="Vit, S. (2011) A description of Euconnus (Euconophron) alesi sp. nov., first Palaearctic Euconnus with cranial modifications (Coleoptera, Staphylinidae, Scydmaeninae). Studies and Reports, Taxonomical Series, 7 (1 - 2), 443 - 448." type="journal article" year="2011">Vít, 2011</bibRefCitation>
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||
</taxonomicName>
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is known to occur in the Chinese province of Fujian. This strongly suggests that similar species should be also expected to occur in
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<collectingCountry id="F37ECAEDFF82FFEFFE582A70FDECFED2" box="[436,522,295,321]" name="Taiwan" pageId="25" pageNumber="26">Taiwan</collectingCountry>
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. All these beetles share similar body form, conspicuous modifications of the head in males (a transverse glandular groove or impression that dorsally demarcates a slightly bulging median frontal region), distinctly tetramerous antennal clubs, and aedeagal structures. There are differences between samples from different islands of the Ryukyus that I have examined, including the shape of the male cephalic glandular structure and the aedeagus. However, there also seems to be some variability within populations inhabiting large islands. The taxonomic situation is complicated by several factors: (1) the aedeagus in this group, similarly as that in the
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<taxonomicName id="4C69F1FEFF82FFEFFF7B2AA8FEE9FD8A" baseAuthorityName="Sharp" baseAuthorityYear="1874" box="[151,271,511,537]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="fustiger">
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||
<emphasis id="B91D566FFF82FFEFFF7B2AA8FEE9FD8A" box="[151,271,511,537]" italics="true" pageId="25" pageNumber="26">
|
||
E.
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<emphasis id="B91D566FFF82FFEFFF552AA8FEE9FD8A" box="[185,271,511,537]" pageId="25" pageNumber="26">fustiger</emphasis>
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</emphasis>
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||
</taxonomicName>
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group, is thin-walled and highly prone to distortions during preparation; (2) the aedeagus of
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<taxonomicName id="4C69F1FEFF82FFEFFAAA2AA8FA7AFD8A" authorityName="Vit" authorityYear="2011" box="[1350,1436,511,537]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="alesi">
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||
<emphasis id="B91D566FFF82FFEFFAAA2AA8FA7AFD8A" box="[1350,1436,511,537]" italics="true" pageId="25" pageNumber="26">E. alesi</emphasis>
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||
</taxonomicName>
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was illustrated based on a slightly damaged specimen, with the median lobe cracked and lacking parameres; (3) the aedeagus of the unique male included in the
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<typeStatus id="54D234DFFF82FFEFFD522910FD08FDF2" box="[702,750,583,609]" pageId="25" pageNumber="26">type</typeStatus>
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||
series of
|
||
<taxonomicName id="4C69F1FEFF82FFEFFCB22910FBF0FDF2" authorityName="Hoshina" authorityYear="2013" box="[862,1046,583,609]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kumejimensis">
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<emphasis id="B91D566FFF82FFEFFCB22910FBF0FDF2" box="[862,1046,583,609]" italics="true" pageId="25" pageNumber="26">E. kumejimensis</emphasis>
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</taxonomicName>
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is fully erected, which means that the entire endophallus is everted outside the median lobe, with its sclerotized structures displaced, and the ventral apical plate is flipped ventrad to open the ostium; and (4) another member of this group,
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<taxonomicName id="4C69F1FEFF82FFEFFB6329D8FB08FD3A" box="[1167,1262,655,681]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="touko">
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||
<emphasis id="B91D566FFF82FFEFFB6329D8FB08FD3A" box="[1167,1262,655,681]" italics="true" pageId="25" pageNumber="26">E. touko</emphasis>
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</taxonomicName>
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, was described from Shimokoshiki Island and although its illustrated aedeagus is intact, it is impossible to compare it with the one illustrated for
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<taxonomicName id="4C69F1FEFF82FFEFFED42980FE09FD62" authorityName="Hoshina" authorityYear="2013" box="[312,495,727,753]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kumejimensis">
|
||
<emphasis id="B91D566FFF82FFEFFED42980FE09FD62" box="[312,495,727,753]" italics="true" pageId="25" pageNumber="26">E. kumejimensis</emphasis>
|
||
</taxonomicName>
|
||
. Moreover,
|
||
<bibRefCitation id="EFF8F78CFF82FFEFFD972980FCCDFD62" author="Hoshina, H." box="[635,811,727,753]" pageId="25" pageNumber="26" pagination="285 - 288" refId="ref24786" refString="Hoshina, H. (2013) New record of the subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Kumejima Island, the Ryukyus, Japan, with description of a new species. Japanese Journal of Systematic Entomology, 19 (2), 285 - 288." type="journal article" year="2013">Hoshina (2013)</bibRefCitation>
|
||
stated that “both parameres of aedeagus bent at a right {angle} in lateral view”, but in fact in the aedeagus illustrated in the lateral view in his fig. 5, the structure that appears as a paramere is a hybrid of a paramere (proximal half) with the dorsal apical plate, which gives an impression of a slender ‘paramere’ bent at a right angle. As illustrated in figures
|
||
<quantity id="4C912798FF82FFEFFC602814FC33FCCE" box="[908,981,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="8.89" metricValueMax="10.16" metricValueMin="7.62" pageId="25" pageNumber="26" unit="in" value="3.5" valueMax="4.0" valueMin="3.0">3‒4 in</quantity>
|
||
the same study, the parameres are short and do not reach the apex of median lobe.
|
||
<bibRefCitation id="EFF8F78CFF82FFEFFD952830FCD3FC12" author="Hoshina, H." box="[633,821,871,897]" pageId="25" pageNumber="26" pagination="49 - 65" refId="ref24877" refString="Hoshina, H. (2019 a) Seven new species of the subfamily Scydmaeninae (Coleoptera: Staphylinidae) from Japan. The Memoirs of the Research and Education Center for Regional Environment, Fukui University, 25, 49 - 65. [2018]" type="journal article" year="2019">Hoshina (2019a)</bibRefCitation>
|
||
illustrated again an aedeagus supposedly belonging to
|
||
<taxonomicName id="4C69F1FEFF82FFEFFF7B28DCFEABFC36" authorityName="Hoshina" authorityYear="2013" box="[151,333,907,933]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kumejimensis">
|
||
<emphasis id="B91D566FFF82FFEFFF7B28DCFEABFC36" box="[151,333,907,933]" italics="true" pageId="25" pageNumber="26">E. kumejimensis</emphasis>
|
||
</taxonomicName>
|
||
, but he did not explain where this male specimen came from. The aedeagus shown in that paper in ventral and dorsal views seems almost intact (but almost certainly highly simplified), whereas that in the lateral view is fully erected. These illustrations can hardly be compared with those in the original description, and may or may not show the same species. This only adds uncertainty regarding the taxonomy of the
|
||
<taxonomicName id="4C69F1FEFF82FFEFFBBB28A0FB4DFB82" authorityName="Vit" authorityYear="2011" box="[1111,1195,1015,1041]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="alesi">
|
||
<emphasis id="B91D566FFF82FFEFFBBB28A0FB4DFB82" box="[1111,1195,1015,1041]" italics="true" pageId="25" pageNumber="26">E. alesi</emphasis>
|
||
</taxonomicName>
|
||
species complex.
|
||
</paragraph>
|
||
<paragraph id="8BD68A7DFF82FFEFFF2B2F4CFBC3FB79" blockId="25.[151,1437,151,1401]" pageId="25" pageNumber="26">
|
||
Fortunately, Kumejima, where the male
|
||
<typeStatus id="54D234DFFF82FFEFFD7A2F4CFD1EFBA6" box="[662,760,1051,1077]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C69F1FEFF82FFEFFCCE2F4CFC3CFBA6" authorityName="Hoshina" authorityYear="2013" box="[802,986,1051,1077]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kumejimensis">
|
||
<emphasis id="B91D566FFF82FFEFFCCE2F4CFC3CFBA6" box="[802,986,1051,1077]" italics="true" pageId="25" pageNumber="26">E. kumejimensis</emphasis>
|
||
</taxonomicName>
|
||
comes from, is a small island with the longest diameter ~
|
||
<quantity id="4C912798FF82FFEFFE862F68FE3BFBCA" box="[362,477,1087,1113]" metricMagnitude="4" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="25" pageNumber="26" unit="km" value="12.5" valueMax="13.0" valueMin="12.0">12‒13 km</quantity>
|
||
and a rather limited forested area suitable for leaf-litter
|
||
<taxonomicName id="4C69F1FEFF82FFEFFB8E2F68FAEFFBCA" authorityName="Leach" authorityYear="1815" box="[1122,1289,1087,1113]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="36" phylum="Arthropoda" rank="subFamily" subFamily="Scydmaeninae">Scydmaeninae</taxonomicName>
|
||
. It should be possible to collect new material and properly redescribe this interesting species, which is a prerequisite to analyze populations from other islands of the Ryukyus, possibly
|
||
<collectingCountry id="F37ECAEDFF82FFEFFCF02FD0FC97FB32" box="[796,881,1159,1185]" name="Taiwan" pageId="25" pageNumber="26">Taiwan</collectingCountry>
|
||
, and the Chinese coast in Fujian. At the moment taxonomic treatment of this group is not possible, despite an interesting sample of specimens from various Ryukyuan islands that I have examined (and have not been able to identify to species level).
|
||
</paragraph>
|
||
<paragraph id="8BD68A7DFF82FFEFFF2B2FA4FC0CFAEA" blockId="25.[151,1437,151,1401]" pageId="25" pageNumber="26">
|
||
<bibRefCitation id="EFF8F78CFF82FFEFFF2B2FA4FEDFFA9E" author="Vit, S." box="[199,313,1267,1293]" pageId="25" pageNumber="26" pagination="443 - 448" refId="ref26567" refString="Vit, S. (2011) A description of Euconnus (Euconophron) alesi sp. nov., first Palaearctic Euconnus with cranial modifications (Coleoptera, Staphylinidae, Scydmaeninae). Studies and Reports, Taxonomical Series, 7 (1 - 2), 443 - 448." type="journal article" year="2011">Vít (2011)</bibRefCitation>
|
||
placed
|
||
<taxonomicName id="4C69F1FEFF82FFEFFE662FA4FE3DFA9E" authorityName="Vit" authorityYear="2011" box="[394,475,1267,1293]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="alesi">
|
||
<emphasis id="B91D566FFF82FFEFFE662FA4FE3DFA9E" box="[394,475,1267,1293]" italics="true" pageId="25" pageNumber="26">E.alesi</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="4C69F1FEFF82FFEFFE162FA4FD77FA9E" box="[506,657,1267,1293]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Euconophron">
|
||
<emphasis id="B91D566FFF82FFEFFE162FA4FD77FA9E" box="[506,657,1267,1293]" italics="true" pageId="25" pageNumber="26">Euconophron</emphasis>
|
||
</taxonomicName>
|
||
, and also
|
||
<bibRefCitation id="EFF8F78CFF82FFEFFD162FA4FC40FA9E" author="Hoshina, H." box="[762,934,1267,1293]" pageId="25" pageNumber="26" pagination="285 - 288" refId="ref24786" refString="Hoshina, H. (2013) New record of the subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Kumejima Island, the Ryukyus, Japan, with description of a new species. Japanese Journal of Systematic Entomology, 19 (2), 285 - 288." type="journal article" year="2013">Hoshina (2013)</bibRefCitation>
|
||
placed
|
||
<taxonomicName id="4C69F1FEFF82FFEFFC142FA4FB4DFA9E" authorityName="Hoshina" authorityYear="2013" box="[1016,1195,1267,1293]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="kumejimensis">
|
||
<emphasis id="B91D566FFF82FFEFFC142FA4FB4DFA9E" box="[1016,1195,1267,1293]" italics="true" pageId="25" pageNumber="26">E. kumejimensis</emphasis>
|
||
</taxonomicName>
|
||
in the same subgenus. However, as redefined by
|
||
<bibRefCitation id="EFF8F78CFF82FFEFFE282E40FD46FAA2" author="Jaloszynski, P." box="[452,672,1303,1329]" pageId="25" pageNumber="26" pagination="450 - 460" refId="ref25610" refString="Jaloszynski, P. (2017 b) Taxonomy of " Euconnus complex ". Part XI. Status of Euconophron and Scopophus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4236 (3), 450 - 460. https: // doi. org / 10.11646 / zootaxa. 4236.3.2" type="journal article" year="2017">Jałoszyński (2017b</bibRefCitation>
|
||
, d; 2021a),
|
||
<emphasis id="B91D566FFF82FFEFFCC62E40FC55FAA3" box="[810,947,1303,1328]" italics="true" pageId="25" pageNumber="26">Eupentarius</emphasis>
|
||
(a senior synonym of
|
||
<taxonomicName id="4C69F1FEFF82FFEFFB542E40FAA9FAA2" box="[1208,1359,1303,1329]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Euconophron">
|
||
<emphasis id="B91D566FFF82FFEFFB542E40FAA9FAA2" box="[1208,1359,1303,1329]" italics="true" pageId="25" pageNumber="26">Euconophron</emphasis>
|
||
</taxonomicName>
|
||
) has a median antebasal pronotal pit and a clearly bifurcate or deeply emarginate aedeagal apex, features that these two Far Eastern species lack. Their placement is here corrected to
|
||
<taxonomicName id="4C69F1FEFF82FFEFFCCD2E08FC69FAEB" box="[801,911,1375,1400]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B91D566FFF82FFEFFCCD2E08FC69FAEB" box="[801,911,1375,1400]" italics="true" pageId="25" pageNumber="26">Euconnus</emphasis>
|
||
</taxonomicName>
|
||
(s. str.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |