272 lines
28 KiB
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272 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.162.55550" ID-GBIF-Dataset="b9049e0e-c87a-45c3-a571-3008b0108fce" ID-PMC="PMC7557330" ID-Pensoft-Pub="1314-2003-162-45" ID-Pensoft-UUID="F6F902318CE951ED8048579E8D3BABF2" ID-PubMed="33110384" ModsDocID="1314-2003-162-45" checkinTime="1602130058990" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Soreng, Robert J., Sylvester, Mitsy D. P. V. & Clark, Vincent Ralph" docDate="2020" docId="DD93F5FD94F758D4A874531194B095E3" docLanguage="en" docName="PhytoKeys 162: 45-69" docOrigin="PhytoKeys 162" docSource="http://dx.doi.org/10.3897/phytokeys.162.55550" docTitle="Festuca caprina subsp. var. var. caprina var. caprina Nees, Fl. Afr. Austral. Ill. 443. 1841. Festuca nubigena subsp. caprina (Nees) St. - Yves, Rev. Bretonne Bot. Pure Appl. 2: 79. 1927" docType="treatment" docVersion="4" id="F6F902318CE951ED8048579E8D3BABF2" lastPageNumber="45" masterDocId="F6F902318CE951ED8048579E8D3BABF2" masterDocTitle="Festuca drakensbergensis (Poaceae): A common new species in the F. caprina complex from the Drakensberg Mountain Centre of Floristic Endemism, southern Africa, with key and notes on taxa in the complex including the overlooked F. exaristata" masterLastPageNumber="69" masterPageNumber="45" pageNumber="45" updateTime="1668140076684" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Festuca drakensbergensis (Poaceae): A common new species in the F. caprina complex from the Drakensberg Mountain Centre of Floristic Endemism, southern Africa, with key and notes on taxa in the complex including the overlooked F. exaristata</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sylvester, Steven P.</mods:namePart>
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<mods:affiliation>College of Biology and the Environment, Nanjing Forestry University, Long Pan Road No. 159, Nanjing, 210037, China</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5577-8782</mods:nameIdentifier>
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<mods:nameIdentifier type="email">steven_sylvester@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Soreng, Robert J.</mods:namePart>
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<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20013 - 7012, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-8358-4915</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sylvester, Mitsy D. P. V.</mods:namePart>
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<mods:affiliation>College of Biology and the Environment, Nanjing Forestry University, Long Pan Road No. 159, Nanjing, 210037, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Clark, Vincent Ralph</mods:namePart>
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<mods:affiliation>Afromontane Research Unit and Department of Geography, University of the Free State, Qwaqwa Campus, Phuthaditjhaba, 9866, South Africa</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5058-0742</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>162</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>45</mods:start>
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<mods:end>69</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.162.55550</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.162.55550</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-162-45</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F6F902318CE951ED8048579E8D3BABF2</mods:identifier>
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<treatment ID-GBIF-Taxon="168243620" LSID="urn:lsid:plazi:treatment:DD93F5FD94F758D4A874531194B095E3" httpUri="http://treatment.plazi.org/id/DD93F5FD94F758D4A874531194B095E3" lastPageNumber="45" pageId="0" pageNumber="45">
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<subSubSection pageId="0" pageNumber="45" type="nomenclature">
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<paragraph pageId="0" pageNumber="45">
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<taxonomicName LSID="DD93F5FD-94F7-58D4-A874-531194B095E3" authority="var. caprina Nees, Fl. Afr. Austral. Ill. 443. 1841. Festuca nubigena subsp. caprina (Nees) St. - Yves, Rev. Bretonne Bot. Pure Appl. 2: 79. 1927." class="Liliopsida" family="Poaceae" genus="Festuca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Festuca caprina subsp. var. var. caprina" order="Poales" pageId="0" pageNumber="45" phylum="Tracheophyta" rank="variety" species="caprina" subSpecies="var." variety="caprina">Festuca caprina var. caprina Nees, Fl. Afr. Austral. Ill. 443. 1841. Festuca nubigena subsp. caprina (Nees) St.-Yves, Rev. Bretonne Bot. Pure Appl. 2: 79. 1927.</taxonomicName>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Festuca caprina var. caprina. A Whole plant B abaxial leaf blade surface of tiller C, D spikelet, lateral view. A, B, D of S. P. Sylvester et al. 3492 (US) C of S. P. Sylvester et al. 3492 (PRE)." figureDoi="10.3897/phytokeys.162.55550.figure3" httpUri="https://binary.pensoft.net/fig/460435" pageId="0" pageNumber="45">Fig. 3</figureCitation>
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,
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<figureCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Differences in key morphological characters between the species of the Festuca caprina complex in southern Africa." pageId="0" pageNumber="45">Table 1</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="45" type="materials_examined">
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<paragraph pageId="0" pageNumber="45">Type.</paragraph>
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<paragraph pageId="0" pageNumber="45">
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South Africa. [Eastern Cape:] Table mountain, Queenstown Dev., Los-Tafelberg, 5000-6000 ft [1524-1829 m alt.], [1840], [flowering in December], D.F.
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<normalizedToken originalValue="Drége">Drege</normalizedToken>
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s.n. (lectotype, designated by
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<bibRefCitation author="Alexeev, EB" journalOrPublisher="Botaniĉeskij Zhurnal (Moscow & Leningrad)" pageId="0" pageNumber="45" pagination="1109 - 1117" refId="B1" refString="Alexeev, EB, 1986. New narrow-leaved Festuca (Poaceae) members from tropical and south Africa. Botaniĉeskij Zhurnal (Moscow & Leningrad) 71 (8): 1109 - 1117" title="New narrow-leaved Festuca (Poaceae) members from tropical and south Africa." volume="71" year="1986">Alexeev 1986</bibRefCitation>
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: 1115: K (K000345257 [image!]; isolectotypes: K (K000345258 [image!]), LE (LE00009757 [image!]); syntypes: South Africa. Plantes du Cap, Los Tafelberg, 5000-6000 ft [1524-1829 m alt.], 7 Dec 1832, D.F.
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<normalizedToken originalValue="Drége">Drege</normalizedToken>
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8.e.3920 (P (P00434763 [image!])); [South Africa] Afr. Austr. D.F.
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<normalizedToken originalValue="Drége">Drege</normalizedToken>
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s.n. (L (L1262355 [image!])); South Africa. Los Tafelberg, Dec 1826-1834, D.F.
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<normalizedToken originalValue="Drége">Drege</normalizedToken>
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s.n. (HAL (HAL0106999 [image!))).
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</paragraph>
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<paragraph pageId="0" pageNumber="45">
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=
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Festuca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Festuca caprina subsp. var. var. curvula" order="Poales" pageId="0" pageNumber="45" phylum="Tracheophyta" rank="variety" species="caprina" subSpecies="var." variety="curvula">Festuca caprina var. curvula</taxonomicName>
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Nees, Fl. Afr. Austral. Ill. 1: 443. 1841. Type: South Africa. [Eastern Cape:] [Monte] Los Tafelberg, an steinigen Oetern, 5000-6000 ft [1524-1829 m alt.], [flowering in December], D.F.
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<normalizedToken originalValue="Drége">Drege</normalizedToken>
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s.n. (lectotype,
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<emphasis bold="true" pageId="0" pageNumber="45">designated here</emphasis>
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: S (S-G-6704 right-hand plant annotated with
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<normalizedToken originalValue="‘b’">'b'</normalizedToken>
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[image!]); syntype: South Africa. Plantes du Cap, D.F.
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<normalizedToken originalValue="Drége">Drege</normalizedToken>
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8.e.3920? (P (P00434764 [image!])).
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</paragraph>
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<paragraph pageId="0" pageNumber="45">
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=
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<taxonomicName authorityName="Nees" authorityYear="1841" class="Liliopsida" family="Poaceae" genus="Festuca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Festuca costata subsp. var. var. longiseta" order="Poales" pageId="0" pageNumber="45" phylum="Tracheophyta" rank="variety" species="costata" subSpecies="var." variety="longiseta">Festuca costata var. longiseta</taxonomicName>
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Nees, Fl. Afr. Austral. Ill. 1: 447. 1841. Type: South Africa. [Eastern Cape:] Stockenstrom Division, Katberg, [4000-5000 ft; 1219-1524 m alt.], 1840, D.F.
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<normalizedToken originalValue="Drége">Drege</normalizedToken>
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s.n. (lectotype, designated by
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<bibRefCitation author="Alexeev, EB" journalOrPublisher="Botaniĉeskij Zhurnal (Moscow & Leningrad)" pageId="0" pageNumber="45" pagination="1109 - 1117" refId="B1" refString="Alexeev, EB, 1986. New narrow-leaved Festuca (Poaceae) members from tropical and south Africa. Botaniĉeskij Zhurnal (Moscow & Leningrad) 71 (8): 1109 - 1117" title="New narrow-leaved Festuca (Poaceae) members from tropical and south Africa." volume="71" year="1986">Alexeev 1986</bibRefCitation>
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: 1115: K (K000345256 [image!]); isolectotype: K (K000345255 [image!])).
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</paragraph>
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<caption doi="10.3897/phytokeys.162.55550.figure3" httpUri="https://binary.pensoft.net/fig/460435" pageId="0" pageNumber="45" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="45">
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<emphasis bold="true" pageId="0" pageNumber="45">Figure 3.</emphasis>
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<taxonomicName authority="var. caprina" class="Liliopsida" family="Poaceae" genus="Festuca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Festuca caprina subsp. var. var. caprina" order="Poales" pageId="0" pageNumber="45" phylum="Tracheophyta" rank="variety" species="caprina" subSpecies="var." variety="caprina">Festuca caprina var. caprina</taxonomicName>
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.
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<emphasis bold="true" pageId="0" pageNumber="45">A</emphasis>
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Whole plant
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<emphasis bold="true" pageId="0" pageNumber="45">B</emphasis>
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abaxial leaf blade surface of tiller
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<emphasis bold="true" pageId="0" pageNumber="45">C, D</emphasis>
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spikelet, lateral view.
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<emphasis bold="true" pageId="0" pageNumber="45">A, B, D</emphasis>
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of S.P. Sylvester et al. 3492 (US)
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<emphasis bold="true" pageId="0" pageNumber="45">C</emphasis>
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of S.P. Sylvester et al. 3492 (PRE).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="45" type="notes">
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<paragraph pageId="0" pageNumber="45">Notes.</paragraph>
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<paragraph pageId="0" pageNumber="45">
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<bibRefCitation author="Alexeev, EB" journalOrPublisher="Botaniĉeskij Zhurnal (Moscow & Leningrad)" pageId="0" pageNumber="45" pagination="1109 - 1117" refId="B1" refString="Alexeev, EB, 1986. New narrow-leaved Festuca (Poaceae) members from tropical and south Africa. Botaniĉeskij Zhurnal (Moscow & Leningrad) 71 (8): 1109 - 1117" title="New narrow-leaved Festuca (Poaceae) members from tropical and south Africa." volume="71" year="1986">Alexeev (1986)</bibRefCitation>
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separated
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<taxonomicName lsidName="F. caprina" pageId="0" pageNumber="45" rank="species" species="caprina">
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<emphasis italics="true" pageId="0" pageNumber="45">F. caprina</emphasis>
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</taxonomicName>
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from
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<taxonomicName lsidName="F. macra" pageId="0" pageNumber="45" rank="species" species="macra">
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<emphasis italics="true" pageId="0" pageNumber="45">F. macra</emphasis>
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</taxonomicName>
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(=
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<taxonomicName lsidName="F. caprina subsp. var. var. macra" pageId="0" pageNumber="45" rank="variety" species="caprina" subSpecies="var." variety="macra">F. caprina var. macra</taxonomicName>
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) based, in part, on the basal sheaths being fibrous. However, all type or original material of
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<taxonomicName lsidName="F. caprina" pageId="0" pageNumber="45" rank="species" species="caprina">
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<emphasis italics="true" pageId="0" pageNumber="45">F. caprina</emphasis>
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</taxonomicName>
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, including the lectotype of
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<taxonomicName authority="var. caprina" lsidName="F. caprina subsp. var. var. caprina" pageId="0" pageNumber="45" rank="variety" species="caprina" subSpecies="var." variety="caprina">F. caprina var. caprina</taxonomicName>
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designated by
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<bibRefCitation author="Alexeev, EB" journalOrPublisher="Botaniĉeskij Zhurnal (Moscow & Leningrad)" pageId="0" pageNumber="45" pagination="1109 - 1117" refId="B1" refString="Alexeev, EB, 1986. New narrow-leaved Festuca (Poaceae) members from tropical and south Africa. Botaniĉeskij Zhurnal (Moscow & Leningrad) 71 (8): 1109 - 1117" title="New narrow-leaved Festuca (Poaceae) members from tropical and south Africa." volume="71" year="1986">Alexeev (1986)</bibRefCitation>
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, had entire, often lustrous, basal sheaths apart from
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<taxonomicName lsidName="subsp. var. var. irrasa" pageId="0" pageNumber="45" rank="variety" species="caprina" subSpecies="var." variety="irrasa">var. irrasa</taxonomicName>
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, which were obviously fibrous. The protologue mentions basal sheaths to be fibrous and, as such,
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<bibRefCitation author="Alexeev, EB" journalOrPublisher="Botaniĉeskij Zhurnal (Moscow & Leningrad)" pageId="0" pageNumber="45" pagination="1109 - 1117" refId="B1" refString="Alexeev, EB, 1986. New narrow-leaved Festuca (Poaceae) members from tropical and south Africa. Botaniĉeskij Zhurnal (Moscow & Leningrad) 71 (8): 1109 - 1117" title="New narrow-leaved Festuca (Poaceae) members from tropical and south Africa." volume="71" year="1986">Alexeev (1986)</bibRefCitation>
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may have made an error in his choice of lectotype. Nevertheless, as only the type material of var.
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<taxonomicName lsidName="var. irrasa" pageId="0" pageNumber="45" rank="variety" species="caprina" variety="irrasa">irrasa</taxonomicName>
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, which was designated by
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<bibRefCitation author="Stapf, O" editor="Thiselton-Dyer, WT" journalOrPublisher="Lovell Reeve & Co. Ltd., London" pageId="0" pageNumber="45" pagination="310 - 765" refId="B33" refString="Stapf, O, 1900. Gramineae. In: Thiselton-Dyer, WT, Ed., Flora Capensis, Volume VII, Pontederiaceae to Gramineae. Lovell Reeve & Co. Ltd., London: 310 - 765" title="Gramineae." volumeTitle="Flora Capensis, Volume VII, Pontederiaceae to Gramineae." year="1900">Stapf (1900)</bibRefCitation>
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, has fibrous basal sheaths, this also raises questions over the accuracy of the description in the protologue for var.
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<taxonomicName lsidName="var. caprina" pageId="0" pageNumber="45" rank="variety" species="caprina" variety="caprina">caprina</taxonomicName>
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. If we treat
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<taxonomicName authority="var. caprina" lsidName="F. caprina subsp. var. var. caprina" pageId="0" pageNumber="45" rank="variety" species="caprina" subSpecies="var." variety="caprina">F. caprina var. caprina</taxonomicName>
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based on the K lectotype and isolectotype designated by
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<bibRefCitation author="Alexeev, EB" journalOrPublisher="Botaniĉeskij Zhurnal (Moscow & Leningrad)" pageId="0" pageNumber="45" pagination="1109 - 1117" refId="B1" refString="Alexeev, EB, 1986. New narrow-leaved Festuca (Poaceae) members from tropical and south Africa. Botaniĉeskij Zhurnal (Moscow & Leningrad) 71 (8): 1109 - 1117" title="New narrow-leaved Festuca (Poaceae) members from tropical and south Africa." volume="71" year="1986">Alexeev (1986)</bibRefCitation>
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then var.
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<taxonomicName lsidName="var. caprina" pageId="0" pageNumber="45" rank="variety" species="caprina" variety="caprina">caprina</taxonomicName>
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should be considered as having entire basal sheaths that do not split into fibres. Oddly, the inflorescences of all var.
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<taxonomicName lsidName="var. caprina" pageId="0" pageNumber="45" rank="variety" species="caprina" variety="caprina">caprina</taxonomicName>
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specimens studied had a distinct butter-like smell upon the opening of specimen press papers, which then quickly dissipated. This odour was barely to sometimes slightly susceptible in specimens of
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<taxonomicName lsidName="F. caprina subsp. var. var. macra" pageId="0" pageNumber="45" rank="variety" species="caprina" subSpecies="var." variety="macra">F. caprina var. macra</taxonomicName>
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or var.
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<taxonomicName lsidName="var. irrasa" pageId="0" pageNumber="45" rank="variety" species="caprina" variety="irrasa">irrasa</taxonomicName>
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or
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<taxonomicName lsidName="F. drakensbergensis" pageId="0" pageNumber="45" rank="species" species="drakensbergensis">
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<emphasis italics="true" pageId="0" pageNumber="45">F. drakensbergensis</emphasis>
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</taxonomicName>
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. It remains to be seen whether this character is diagnostic and what phytochemical compounds are involved.
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</paragraph>
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<paragraph pageId="0" pageNumber="45">
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<taxonomicName authority="var. caprina" class="Liliopsida" family="Poaceae" genus="Festuca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Festuca caprina subsp. var. var. caprina" order="Poales" pageId="0" pageNumber="45" phylum="Tracheophyta" rank="variety" species="caprina" subSpecies="var." variety="caprina">Festuca caprina var. caprina</taxonomicName>
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is more common at lower elevations in the Drakensberg Mountain Centre (
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<bibRefCitation DOI="https://doi.org/10.1016/j.sajb.2019.05.032" author="Carbutt, C" journalOrPublisher="South African Journal of Botany" pageId="0" pageNumber="45" pagination="508 - 529" refId="B4" refString="Carbutt, C, 2019. The Drakensberg Mountain Centre: A necessary revision of southern Africa's high-elevation centre of plant endemism. South African Journal of Botany 124: 508 - 529, DOI: https://doi.org/10.1016/j.sajb.2019.05.032" title="The Drakensberg Mountain Centre: A necessary revision of southern Africa's high-elevation centre of plant endemism." url="https://doi.org/10.1016/j.sajb.2019.05.032" volume="124" year="2019">Carbutt 2019</bibRefCitation>
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) and surrounding mountainous habitats of southern Africa and extends from southern Africa to Tanzania. The species appears to prefer more mesic Afro-alpine and Afro-montane grasslands and is outcompeted by
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<taxonomicName lsidName="F. caprina subsp. var. var. macra" pageId="0" pageNumber="45" rank="variety" species="caprina" subSpecies="var." variety="macra">F. caprina var. macra</taxonomicName>
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in the drier summit area of the high escarpment in the DMC. Of the 222 2 m
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<normalizedToken originalValue="×">x</normalizedToken>
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2 m plots studied for all vascular plants across the Afro-alpine DMC (Sylvester et al. unpubl. data),
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<emphasis italics="true" pageId="0" pageNumber="45">
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<taxonomicName lsidName="F. caprina" pageId="0" pageNumber="45" rank="species" species="caprina">F. caprina</taxonomicName>
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var.
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<taxonomicName lsidName="var. caprina" pageId="0" pageNumber="45" rank="variety" species="caprina" variety="caprina">caprina</taxonomicName>
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</emphasis>
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was rarely encountered, being found in only 13 plots from the Eastern Cape and Free State. The species was usually encountered in lower elevation Afro-montane transitioning to Afro-alpine grasslands at ca. 2500-2700 m alt. or exceptionally at higher elevations to 2981 m alt. in damper shaded sites, highlighting its very low frequency and commonality in the high-elevation xeric Afro-alpine zone of the DMC.
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</paragraph>
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<paragraph pageId="0" pageNumber="45">
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Festuca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Festuca caprina subsp. var. var. curvula" order="Poales" pageId="0" pageNumber="45" phylum="Tracheophyta" rank="variety" species="caprina" subSpecies="var." variety="curvula">Festuca caprina var. curvula</taxonomicName>
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is also herein lectotypified. In the protologue,
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.7585" author="Nees von Esenbeck, CGD" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="45" refId="B22" refString="Nees von Esenbeck, CGD, 1841. Florae Africae Australioris Illustrationes Monographicae: Gramineae. Palala Press. https://doi.org/10.5962/bhl.title.7585" title="Florae Africae Australioris Illustrationes Monographicae: Gramineae. Palala Press." url="https://doi.org/10.5962/bhl.title.7585" year="1841">Nees von Esenbeck (1841</bibRefCitation>
|
||
: 443) only cited a single
|
||
<normalizedToken originalValue="Drége">Drege</normalizedToken>
|
||
s.n. collection from monte Los-Tafelberg, 6000 ft (1829 m alt.), which is assumed to be the same type locality as var.
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Festuca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="var. caprina" order="Poales" pageId="0" pageNumber="45" phylum="Tracheophyta" rank="variety" species="caprina" variety="caprina">caprina</taxonomicName>
|
||
that was found in Los-Tafelberg of the Eastern Cape Province, near Queenstown.
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.7585" author="Nees von Esenbeck, CGD" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="45" refId="B22" refString="Nees von Esenbeck, CGD, 1841. Florae Africae Australioris Illustrationes Monographicae: Gramineae. Palala Press. https://doi.org/10.5962/bhl.title.7585" title="Florae Africae Australioris Illustrationes Monographicae: Gramineae. Palala Press." url="https://doi.org/10.5962/bhl.title.7585" year="1841">Nees von Esenbeck (1841</bibRefCitation>
|
||
: 443) labelled var.
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Festuca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="var. caprina" order="Poales" pageId="0" pageNumber="45" phylum="Tracheophyta" rank="variety" species="caprina" variety="caprina">caprina</taxonomicName>
|
||
and var.
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Festuca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="var. curvula" order="Poales" pageId="0" pageNumber="45" phylum="Tracheophyta" rank="variety" species="caprina" variety="curvula">curvula</taxonomicName>
|
||
<normalizedToken originalValue="‘a’">'a'</normalizedToken>
|
||
and
|
||
<normalizedToken originalValue="‘b’">'b'</normalizedToken>
|
||
, respectively, with the S-G-6704 right hand plant chosen as lectotype based on this matching the protologue information and being the only specimen sheet amongst the original material to be annotated with an
|
||
<normalizedToken originalValue="‘a’">'a'</normalizedToken>
|
||
and
|
||
<normalizedToken originalValue="‘b’">'b'</normalizedToken>
|
||
in Nees von
|
||
<normalizedToken originalValue="Esenbeck’s">Esenbeck's</normalizedToken>
|
||
cursive handwriting. The right-hand plant annotated with
|
||
<normalizedToken originalValue="‘b’">'b'</normalizedToken>
|
||
fitted the protologue description of var.
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Festuca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="var. curvula" order="Poales" pageId="0" pageNumber="45" phylum="Tracheophyta" rank="variety" species="caprina" variety="curvula">curvula</taxonomicName>
|
||
, with Nees differentiating the variety based on its shorter height, curved blades and subsecund panicle branches with few purplish spikelets. One specimen amongst the original material, D.F.
|
||
<normalizedToken originalValue="Drége">Drege</normalizedToken>
|
||
8.e.3920? (P00434764), also had
|
||
<normalizedToken originalValue="‘Curvula’">'Curvula'</normalizedToken>
|
||
written on the label but limited locality information aside from 'Plantes du
|
||
<normalizedToken originalValue="Cap’">Cap'</normalizedToken>
|
||
and is here considered a syntype of var.
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Festuca" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="var. curvula" order="Poales" pageId="0" pageNumber="45" phylum="Tracheophyta" rank="variety" species="caprina" variety="curvula">curvula</taxonomicName>
|
||
as it also fits the description given in the protologue. While the differentiating characters of
|
||
<taxonomicName lsidName="F. caprina subsp. var. var. curvula" pageId="0" pageNumber="45" rank="variety" species="caprina" subSpecies="var." variety="curvula">F. caprina var. curvula</taxonomicName>
|
||
are also found in
|
||
<taxonomicName lsidName="F. drakensbergensis" pageId="0" pageNumber="45" rank="species" species="drakensbergensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="45">F. drakensbergensis</emphasis>
|
||
</taxonomicName>
|
||
, we deduce that var.
|
||
<taxonomicName lsidName="var. curvula" pageId="0" pageNumber="45" rank="variety" species="drakensbergensis" variety="curvula">curvula</taxonomicName>
|
||
is a slight variation from the norm in
|
||
<taxonomicName lsidName="F. caprina" pageId="0" pageNumber="45" rank="species" species="caprina">
|
||
<emphasis italics="true" pageId="0" pageNumber="45">F. caprina</emphasis>
|
||
</taxonomicName>
|
||
as neither the type specimens designated herein, nor any of the other original material from the type locality, can be attributed to
|
||
<taxonomicName lsidName="F. drakensbergensis" pageId="0" pageNumber="45" rank="species" species="drakensbergensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="45">F. drakensbergensis</emphasis>
|
||
</taxonomicName>
|
||
based on their lacking extravaginal branching and cataphyllous shoots as well as having entire lustrous basal sheaths.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="45" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="45">Selected specimens examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="45">
|
||
<emphasis bold="true" pageId="0" pageNumber="45">South Africa.</emphasis>
|
||
Eastern Cape: Naudes Nek pass, near Rhodes,
|
||
<geoCoordinate degrees="30.764792" direction="south" orientation="latitude" precision="1" value="-30.764791">30.764792S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="28.105164" direction="east" orientation="longitude" precision="1" value="28.105164">28.105164E</geoCoordinate>
|
||
, 2588 m alt., Afro-alpine tussock grassland, low rock outcrop, 13 Feb 2020, S.P. Sylvester et al. 3492 (NU, PRE, US); Free State: Witsieshoek, at beginning of Sentinel trail by parking lot, path-side,
|
||
<geoCoordinate degrees="28.733181" direction="south" orientation="latitude" precision="1" value="-28.733181">28.733181S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="28.893296" direction="east" orientation="longitude" precision="1" value="28.893295">28.893296E</geoCoordinate>
|
||
, 2607 m alt., 5 Feb 2020, S.P. Sylvester et al. 3418 (US); Free State: Witsieshoek, Sentinel trail, along beginning of trail that leads to the chain ladders that take you up to Amphitheatre, path-side,
|
||
<geoCoordinate degrees="28.736207" direction="south" orientation="latitude" precision="1" value="-28.736206">28.736207S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="28.894084" direction="east" orientation="longitude" precision="1" value="28.894085">28.894084E</geoCoordinate>
|
||
, 2693 m alt., 5 Feb 2020, S.P. Sylvester et al. 3416 (US); Free State: Golden Gate National Park, summit of Wodehouse Peak, 14 Jan 1975, R.P. Ellis 2383 (PRE0464133); KwaZulu-Natal: Amphitheatre, slopes near the Tugela waterfall, Afro-alpine grassland,
|
||
<geoCoordinate degrees="28.750810" direction="south" orientation="latitude" precision="1" value="-28.75081">28.750810S</geoCoordinate>
|
||
, 28.888942 E, 2981 m alt., 5 Feb 2020, S.P. Sylvester et al. 3409a (US); KwaZulu-Natal: Sentinel Trail, off the main trail at the top of an east facing gully ca. 1 km from the chain ladders,
|
||
<geoCoordinate degrees="28.743162" direction="south" orientation="latitude" precision="1" value="-28.743162">28.743162S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="28.888205" direction="east" orientation="longitude" precision="1" value="28.888205">28.888205E</geoCoordinate>
|
||
, 2953 m alt., shaded Afro-montane grassland grading into Afro-alpine grassland, damp soil, rarely grazed, 6 Mar 2020, S.P. Sylvester et al. 3713 (NU, PRE, US).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |