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<document id="52607EB4E02E9E942C5196F63D8DA4DF" ID-DOI="10.5281/zenodo.3699463" ID-GBIF-Dataset="8a1b577e-5fce-489a-9777-32c9564a26b2" ID-ISSN="1942-1354" ID-Zenodo-Dep="3699463" ID-ZooBank="938220AB-7950-4850-88DC-3C3D8AB72C89" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583500003806" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2018" docId="DB4387E1713FFFADFF410C920DB6FB6C" docLanguage="en" docName="InsectaMundi.0624.1-13.pdf.imf" docOrigin="Insecta Mundi 2018 (624)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Nigidius dawnae Gravely 1915" docType="treatment" docVersion="7" lastPageNumber="9" masterDocId="277AFF997137FFA7FF810A620E3FFFA9" masterDocTitle="Revision of the obesus species group of the stag beetle genus Nigidius MacLeay (Coleoptera: Lucanidae: Lucaninae: Figulini)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="7" updateTime="1698725587556" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="80D1D6756002AC3E57C82E1E9B48740C">Revision of the obesus species group of the stag beetle genus Nigidius MacLeay (Coleoptera: Lucanidae: Lucaninae: Figulini)</mods:title>
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<mods:namePart id="FFCFF6136F5E4FE02FC9A1816C405B50">Paulsen, M. J.</mods:namePart>
<mods:affiliation id="84570368610895AD5F610111A54E2FBE">Systematic Research Collections University of Nebraska State Museum W 436 Nebraska Hall Lincoln, NE 68588 - 0546</mods:affiliation>
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<mods:title id="478BAF63C48D47B476280972B5D14DAB">Insecta Mundi</mods:title>
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<treatment id="DB4387E1713FFFADFF410C920DB6FB6C" ID-DOI="http://doi.org/10.5281/zenodo.3704540" ID-GBIF-Taxon="162463115" ID-Zenodo-Dep="3704540" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFADFF410C920DB6FB6C" httpUri="http://treatment.plazi.org/id/DB4387E1713FFFADFF410C920DB6FB6C" lastPageId="10" lastPageNumber="9" pageId="8" pageNumber="7">
<subSubSection id="1BF0657C713FFFAFFF410C920CA0F8A0" box="[192,671,1776,1801]" pageId="8" pageNumber="7" type="nomenclature">
<paragraph id="535536F7713FFFAFFF410C920CA0F8A0" blockId="8.[192,671,1776,1801]" box="[192,671,1776,1801]" pageId="8" pageNumber="7">
<heading id="081D819B713FFFAFFF410C920CA0F8A0" bold="true" box="[192,671,1776,1801]" fontSize="10" level="2" pageId="8" pageNumber="7" reason="3">
<treatmentCitationGroup id="73FA11D9713FFFAFFF410C920CA0F8A0" box="[192,671,1776,1801]" pageId="8" pageNumber="7">
<treatmentCitation id="D24B10E6713FFFAFFF410C920CA0F8A0" author="Gravely" box="[192,671,1776,1801]" page="427" pageId="8" pageNumber="7" year="1915">
<taxonomicName id="94EA4D74713FFFAFFF410C920CA0F8A0" ID-CoL="73Y7V" authority="Gravely 1915: 427" authorityName="Gravely" authorityPageNumber="427" authorityYear="1915" box="[192,671,1776,1801]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="619EEAE5713FFFAFFF410C920CA0F8A0" bold="true" box="[192,671,1776,1801]" pageId="8" pageNumber="7">
<emphasis id="619EEAE5713FFFAFFF410C920F96F8A1" bold="true" box="[192,425,1776,1800]" italics="true" pageId="8" pageNumber="7">Nigidius dawnae</emphasis>
<bibRefCitation id="377B4B06713FFFAFFE300C920CA0F8A0" author="Gravely" box="[433,671,1776,1801]" firstAuthor="Gravely" pageId="8" pageNumber="7" pagination="407 - 431" refId="ref6786" refString="Gravely, F. H. 1915. A catalogue of the Lucanidae in the collection of the Indian Museum. Records of the Indian Museum 9 (6): 407 - 431." type="journal article" year="1915">Gravely 1915: 427</bibRefCitation>
</emphasis>
</taxonomicName>
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="1BF0657C713FFFAFFF410D420D24F89C" box="[192,795,1824,1846]" pageId="8" pageNumber="7" type="reference_group">
<paragraph id="535536F7713FFFAFFF410D420D24F89C" blockId="8.[192,795,1824,1846]" box="[192,795,1824,1846]" pageId="8" pageNumber="7">
<treatmentCitationGroup id="73FA11D9713FFFAFFF410D420D24F89C" box="[192,795,1824,1846]" pageId="8" pageNumber="7">
<treatmentCitation id="D24B10E6713FFFAFFF410D420C6EF89C" author="Lisle, M. O." box="[192,593,1824,1846]" page="" pageId="8" pageNumber="7" year="1974">
<taxonomicName id="94EA4D74713FFFAFFF410D420C6EF89C" ID-CoL="6S9VR" authority="de Lisle 1973: 797" authorityName="de Lisle" authorityPageNumber="797" authorityYear="1974" box="[192,593,1824,1846]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="larssoni">
<emphasis id="619EEAE5713FFFAFFF410D420F40F89C" box="[192,383,1824,1845]" italics="true" pageId="8" pageNumber="7">Nigidius larssoni</emphasis>
<bibRefCitation id="377B4B06713FFFAFFE070D420C6EF89C" author="Lisle, M. O." box="[390,593,1824,1846]" pageId="8" pageNumber="7" pagination="785 - 804" refId="ref6821" refString="Lisle, M. O. de 1974. Troisieme note sur quelques Coleoptera Lucanidae nouveaux ou peu connus. Revue Suisse de Zoologie 80 (4): 785 - 804." type="journal article" year="1974">de Lisle 1973: 797</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis id="619EEAE5713FFFAFFDDD0D420D24F89C" bold="true" box="[604,795,1824,1845]" pageId="8" pageNumber="7">
<taxonomicNameLabel id="7AAD579E713FFFAFFDDD0D420D29F89C" box="[604,790,1824,1845]" pageId="8" pageNumber="7">new synonymy</taxonomicNameLabel>
.
</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="1BF0657C713FFFADFF410D2D0F88FE4C" lastPageId="10" lastPageNumber="9" pageId="8" pageNumber="7" type="materials_examined">
<paragraph id="535536F7713FFFAEFF410D2D0CE8F821" blockId="8.[192,1440,1871,1929]" lastBlockId="9.[192,1440,1673,1928]" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">
<emphasis id="619EEAE5713FFFAFFF410D2D0FB4F8CE" bold="true" box="[192,395,1871,1895]" pageId="8" pageNumber="7">Type material.</emphasis>
<materialsCitation id="E3823CAA713FFFAEFE150D2D0C4FF96B" ID-GBIF-Occurrence="2574936469" collectingDate="2011-11-22" collectingDateMax="2011-11-30" collectingDateMin="2011-11-22" collectionCode="NHM, BMNH" collectorName="F. H. Gravely" country="Myanmar" elevation="671" lastPageId="9" lastPageNumber="8" location="Misty Hollow" municipality="Dawna Hills" pageId="8" pageNumber="7" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
Male*
<typeStatus id="8C518855713FFFAFFE690D2D0C69F8CE" box="[488,598,1871,1895]" pageId="8" pageNumber="7" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="94EA4D74713FFFAFFDFF0D2D0D6FF8CE" box="[638,848,1871,1895]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="619EEAE5713FFFAFFDFF0D2D0D6FF8CE" box="[638,848,1871,1895]" italics="true" pageId="8" pageNumber="7">Nigidius dawnae</emphasis>
</taxonomicName>
(
<collectionCode id="35FBAE32713FFFAFFCE30D320D95F8C1" box="[866,938,1872,1896]" pageId="8" pageNumber="7">NHM</collectionCode>
;
<figureCitation id="CBD12A72713FFFAFFC390D2D0A3FF8C1" box="[952,1024,1871,1896]" captionStart="Figures 34" captionStartId="5.[192,287,1506,1528]" captionTargetBox="[215,1371,238,1443]" captionTargetId="figure@5.[203,1383,233,1453]" captionTargetPageId="5" captionText="Figures 34. Nigidius dawnae Gravely. 3) Major male, dorsal habitus. 4) Lectotype male with labels." figureDoi="http://doi.org/10.5281/zenodo.3699469" httpUri="https://zenodo.org/record/3699469/files/figure.png" pageId="8" pageNumber="7">Fig. 4</figureCitation>
) labeled: a) red-bordered circular “Type”; b) “
<location id="5635602C713FFFAFFEC60D120FD3F821" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFADFF410C920DB6FB6C:5635602C713FFFAFFEC60D120FD3F821" box="[327,492,1904,1928]" municipality="Dawna Hills" name="Misty Hollow" pageId="8" pageNumber="7">Misty Hollow</location>
, / W. side of /
<collectingMunicipality id="B331AC8D713FFFAFFD0F0D130D19F821" box="[654,806,1904,1928]" pageId="8" pageNumber="7">Dawna Hills</collectingMunicipality>
/ c.
<quantity id="94129B12713FFFAFFCD50D130D8FF821" box="[852,944,1904,1929]" metricMagnitude="2" metricUnit="m" metricValue="6.7056" pageId="8" pageNumber="7" unit="ft" value="2200.0">
<elevation id="D8C7D1C4713FFFAFFCD50D130D8FF821" box="[852,944,1904,1929]" metricMagnitude="2" metricUnit="m" metricValue="6.7056" pageId="8" pageNumber="7" unit="ft" value="2200.0">2200 ft.</elevation>
</quantity>
/ 22-30.XI.11. / (along right margin)
<collectorName id="FE1F5321713FFFAEFAE80D130F1BF908" lastPageId="9" lastPageNumber="8" pageId="8" pageNumber="7">F.H. Gravely</collectorName>
”; c) “
<location id="5635602C713EFFAEFEE30CEB0FE3F908" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFADFF410C920DB6FB6C:5635602C713EFFAEFEE30CEB0FE3F908" box="[354,476,1673,1697]" municipality="Dawna Hills" name="Andrewes" pageId="9" pageNumber="8">Andrewes</location>
/
<collectingMunicipality id="B331AC8D713EFFAEFE730CE80C62F908" box="[498,605,1674,1697]" pageId="9" pageNumber="8">Bequest.</collectingMunicipality>
/ B.M. 1922-221”; d) handwritten “
<taxonomicName id="94EA4D74713EFFAEFB8A0CEB0A4AF908" box="[1035,1141,1673,1697]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="619EEAE5713EFFAEFB8A0CEB0A4AF908" box="[1035,1141,1673,1697]" italics="true" pageId="9" pageNumber="8">Nigidius</emphasis>
</taxonomicName>
(male symbol) /
<taxonomicName id="94EA4D74713EFFAEFAC00CEB0F1BF96B" authority="Gravely" authorityName="Gravely" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="619EEAE5713EFFAEFAC00CEB0B9FF908" box="[1345,1440,1673,1697]" italics="true" pageId="9" pageNumber="8">dawnae</emphasis>
Gravely
</taxonomicName>
”; e) “
<collectionCode id="35FBAE32713EFFAEFEE20CC90F81F96A" box="[355,446,1707,1731]" pageId="9" pageNumber="8">BMNH</collectionCode>
(E) / #607167”.
</materialsCitation>
<materialsCitation id="E3823CAA713EFFAEFD010CC80A78F8AD" ID-GBIF-Occurrence="2574936460" collectingDate="2011-11-22" collectingDateMax="2011-11-29" collectingDateMin="2011-11-22" collectionCode="NHM, BMNH" collectorName="F. H. Gravely" elevation="701" location="Misty Hollow" municipality="Sukli" pageId="9" pageNumber="8" specimenCount="4" typeStatus="paralectotype">
Female
<typeStatus id="8C518855713EFFAEFD620CC80DB6F96B" box="[739,905,1706,1730]" pageId="9" pageNumber="8" type="paralectotype">paralectotype</typeStatus>
(
<collectionCode id="35FBAE32713EFFAEFC180CC90DDFF96A" box="[921,992,1707,1731]" pageId="9" pageNumber="8">NHM</collectionCode>
) labeled: a) yellow-bordered circular “Co- / type”; b) “
<location id="5635602C713EFFAEFEFB0CA90C24F94A" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFADFF410C920DB6FB6C:5635602C713EFFAEFEFB0CA90C24F94A" box="[378,539,1739,1763]" municipality="Sukli" name="Misty Hollow" pageId="9" pageNumber="8">Misty Hollow</location>
/ to
<collectingMunicipality id="B331AC8D713EFFAEFDCA0CA90CB3F94A" box="[587,652,1739,1763]" pageId="9" pageNumber="8">Sukli</collectingMunicipality>
/
<quantity id="94129B12713EFFAEFD1F0CA90D02F94A" box="[670,829,1739,1763]" metricMagnitude="2" metricUnit="m" metricValue="7.0104" metricValueMax="7.62" metricValueMin="6.4008" pageId="9" pageNumber="8" unit="ft" value="2300.0" valueMax="2500.0" valueMin="2100.0">
<elevation id="D8C7D1C4713EFFAEFD1F0CA90D02F94A" box="[670,829,1739,1763]" metricMagnitude="2" metricUnit="m" metricValue="7.0104" metricValueMax="7.62" metricValueMin="6.4008" pageId="9" pageNumber="8" unit="ft" value="2300.0" valueMax="2500.0" valueMin="2100.0">2100-2500 ft.</elevation>
</quantity>
/ 22-29.XI.11. / (along right margin)
<collectorName id="FE1F5321713EFFAEFB700CAE0BAEF94A" box="[1265,1425,1739,1763]" pageId="9" pageNumber="8">F.H. Gravely</collectorName>
”; c) as type; d) as type, except (female symbol); e) “BMNH(E) / #607168”.
</materialsCitation>
<materialsCitation id="E3823CAA713EFFAEFBD00C8E0CAAF8EF" ID-GBIF-Occurrence="3878839305" collectionCode="NHM, BMNH" country="Myanmar" location="Burma" pageId="9" pageNumber="8" specimenCount="7" typeStatus="paralectotype">
Male
<typeStatus id="8C518855713EFFAEFB180C8E0B00F8AD" box="[1177,1343,1772,1796]" pageId="9" pageNumber="8" type="paralectotype">paralectotype</typeStatus>
(
<collectionCode id="35FBAE32713EFFAEFAD00C8F0BA7F8AC" box="[1361,1432,1773,1797]" pageId="9" pageNumber="8">NHM</collectionCode>
) labeled: a) green-bordered circular “Co- / type”; b) as type; c) as type; d) as type above, except (female symbol) [sic]; e) “BMNH(E) / #607169”
</materialsCitation>
. Type locality: “Lower
<collectingCountry id="2BFD7667713EFFAEFC290D4D0DC1F8EF" box="[936,1022,1839,1862]" name="Myanmar" pageId="9" pageNumber="8">Burma</collectingCountry>
… - near
<collectingMunicipality id="B331AC8D713EFFAEFBEF0D4C0B2EF8EF" box="[1134,1297,1838,1862]" pageId="9" pageNumber="8">Misty Hollow</collectingMunicipality>
and
<collectingMunicipality id="B331AC8D713EFFAEFACC0D4C0BAEF8EF" box="[1357,1425,1838,1862]" pageId="9" pageNumber="8">Sukli</collectingMunicipality>
”. [*The
<quantity id="94129B12713EFFAEFE8D0D2D0F60F8CE" box="[268,351,1871,1895]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="8" unit="mm" value="15.0">15 mm</quantity>
lectotype was not dissected to determine sex, but the shape of the basal mandibular tooth is consistent with a moderately sized
<specimenCount id="45ECFD7E713EFFAEFD0A0D120CF3F821" box="[651,716,1904,1928]" pageId="9" pageNumber="8" type="male">male</specimenCount>
.]
</paragraph>
<caption id="0795667F713EFFAEFF410C480A5FF9F4" ID-DOI="http://doi.org/10.5281/zenodo.3699477" ID-Zenodo-Dep="3699477" httpUri="https://zenodo.org/record/3699477/files/figure.png" pageId="9" pageNumber="8" startId="9.[192,287,1578,1600]" targetBox="[221,1399,218,1550]" targetPageId="9">
<paragraph id="535536F7713EFFAEFF410C480A5FF9F4" blockId="9.[192,1440,1578,1629]" pageId="9" pageNumber="8">
<emphasis id="619EEAE5713EFFAEFF410C480F51F996" bold="true" box="[192,366,1578,1600]" pageId="9" pageNumber="8">Figures 1013.</emphasis>
Male genitalia of
<taxonomicName id="94EA4D74713EFFAEFDB30C480C43F9E9" box="[562,636,1578,1600]" pageId="9" pageNumber="8" rank="species" species="obesus">obesus</taxonomicName>
group species.
<emphasis id="619EEAE5713EFFAEFCA00C480D79F9E9" bold="true" box="[801,838,1578,1600]" pageId="9" pageNumber="8">10)</emphasis>
<emphasis id="619EEAE5713EFFAEFCCD0C490D85F996" box="[844,954,1578,1600]" italics="true" pageId="9" pageNumber="8">
<taxonomicName id="94EA4D74713EFFAEFCCD0C490D88F996" authorityName="Parry" authorityYear="1864" box="[844,951,1578,1600]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="obesus">N. obesus</taxonomicName>
.
</emphasis>
<emphasis id="619EEAE5713EFFAEFC410C480DDBF9E9" bold="true" box="[960,996,1578,1600]" pageId="9" pageNumber="8">11)</emphasis>
<taxonomicName id="94EA4D74713EFFAEFC6B0C490A5FF996" box="[1002,1120,1578,1600]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="619EEAE5713EFFAEFC6B0C490A5FF996" box="[1002,1120,1578,1600]" italics="true" pageId="9" pageNumber="8">N. dawnae</emphasis>
</taxonomicName>
, arrow indicates small apical lobe.
<emphasis id="619EEAE5713EFFAEFF7B0C250F20F9F4" bold="true" box="[250,287,1607,1629]" pageId="9" pageNumber="8">12)</emphasis>
<taxonomicName id="94EA4D74713EFFAEFEA40C2A0FC0F9F4" authority="Paulsen" authorityName="Paulsen" authorityYear="2018" box="[293,511,1607,1629]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="gravelyi" status="sp. nov.">
<emphasis id="619EEAE5713EFFAEFEA40C2A0FA1F9F5" box="[293,414,1607,1629]" italics="true" pageId="9" pageNumber="8">N. gravelyi</emphasis>
Paulsen
</taxonomicName>
,
<taxonomicNameLabel id="7AAD579E713EFFAEFD8B0C2A0C79F9F4" box="[522,582,1608,1629]" pageId="9" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="619EEAE5713EFFAEFDCD0C250C4EF9F4" bold="true" box="[588,625,1607,1629]" pageId="9" pageNumber="8">13)</emphasis>
<taxonomicName id="94EA4D74713EFFAEFDF90C2A0CE1F9F5" box="[632,734,1607,1629]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="helleri">
<emphasis id="619EEAE5713EFFAEFDF90C2A0CE1F9F5" box="[632,734,1607,1629]" italics="true" pageId="9" pageNumber="8">N. helleri</emphasis>
</taxonomicName>
, arrow indicates large lateral lobe.
</paragraph>
</caption>
<paragraph id="535536F7713DFFADFF690ABF0B55FEFC" blockId="10.[192,1440,221,342]" pageId="10" pageNumber="9">
<materialsCitation id="E3823CAA713DFFADFF690ABF0B55FEFC" ID-GBIF-Occurrence="2574936437" collectingDate="1958-06-17" collectionCode="ZMUC" collectorName="B. Degenbol" country="Thailand" determinerName="M. O. de Lisle &amp; M. J. Paulsen" elevation="1100" location="Zoologisk Museum" pageId="10" pageNumber="9" specimenCount="2" typeStatus="holotype">
<typeStatus id="8C518855713DFFADFF690ABF0F69FF5C" box="[232,342,221,245]" pageId="10" pageNumber="9" type="holotype">Holotype</typeStatus>
female of
<taxonomicName id="94EA4D74713DFFADFE510ABF0C9DFF5C" authorityName="de Lisle" authorityYear="1974" box="[464,674,221,245]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="larssoni">
<emphasis id="619EEAE5713DFFADFE510ABF0C9DFF5C" box="[464,674,221,245]" italics="true" pageId="10" pageNumber="9">Nigidius larssoni</emphasis>
</taxonomicName>
(
<collectionCode id="35FBAE32713DFFADFD2E0ABF0D3BFF5C" box="[687,772,221,245]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="10" pageNumber="9">ZMUC</collectionCode>
) labeled: a) “
<location id="5635602C713DFFADFC1F0ABF0ABAFF5C" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFADFF410C920DB6FB6C:5635602C713DFFADFC1F0ABF0ABAFF5C" box="[926,1157,221,245]" country="Thailand" name="Zoologisk Museum" pageId="10" pageNumber="9">Zoologisk Museum</location>
, København. / Loc. [Doi Sutep, N.
<collectingCountry id="2BFD7667713DFFADFEBA0A9F0F95FEBC" box="[315,426,253,277]" name="Thailand" pageId="10" pageNumber="9">Thailand</collectingCountry>
/
<quantity id="94129B12713DFFADFE400A9F0C22FEBC" box="[449,541,253,277]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="9" unit="m" value="1100.0">
<elevation id="D8C7D1C4713DFFADFE400A9F0C22FEBC" box="[449,541,253,277]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="9" unit="m" value="1100.0">1100 m</elevation>
</quantity>
] / Dat.: [
<date id="27541037713DFFADFD070A9F0D3EFEBC" box="[646,769,253,277]" pageId="10" pageNumber="9" value="1958-06-17">
<collectingDate id="3710E9DF713DFFADFD070A9F0D3EFEBC" box="[646,769,253,277]" pageId="10" pageNumber="9" value="1958-06-17">17-6-1958</collectingDate>
</date>
] / Leg.: [
<collectorName id="FE1F5321713DFFADFCEA0A9C0A3EFEBC" box="[875,1025,253,277]" pageId="10" pageNumber="9">B. Degenbøl</collectorName>
] / Det. [Pr. 492] Journ. [4259]”; b) “[
<typeStatus id="8C518855713DFFADFF500B7F0F6FFE9C" box="[209,336,285,309]" pageId="10" pageNumber="9" type="holotype">Holotypus</typeStatus>
/
<taxonomicName id="94EA4D74713DFFADFEE20B7F0FF2FE9C" box="[355,461,285,309]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="619EEAE5713DFFADFEE20B7F0FF2FE9C" box="[355,461,285,309]" italics="true" pageId="10" pageNumber="9">Nigidius</emphasis>
</taxonomicName>
/
<taxonomicName id="94EA4D74713DFFADFE5E0B7F0C7EFE9C" authorityName="de Lisle" authorityYear="1974" box="[479,577,285,309]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="larssoni">
<emphasis id="619EEAE5713DFFADFE5E0B7F0C7EFE9C" box="[479,577,285,309]" italics="true" pageId="10" pageNumber="9">larssoni</emphasis>
</taxonomicName>
] /
<determinerName id="A689DA33713DFFADFDDD0B7F0CC3FE9C" box="[604,764,285,309]" pageId="10" pageNumber="9">M.O. de Lisle</determinerName>
det. 197[2]”; c) red paper, “TYPE”; d) “zmuc / 00020974”; e) “
<taxonomicName id="94EA4D74713DFFADFF690B5F0F6DFEFC" box="[232,338,317,341]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="619EEAE5713DFFADFF690B5F0F6DFEFC" box="[232,338,317,341]" italics="true" pageId="10" pageNumber="9">Nigidius</emphasis>
</taxonomicName>
/
<taxonomicName id="94EA4D74713DFFADFEE90B5F0FF8FEFC" box="[360,455,317,341]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="619EEAE5713DFFADFEE90B5F0FF8FEFC" box="[360,455,317,341]" italics="true" pageId="10" pageNumber="9">dawnae</emphasis>
</taxonomicName>
/ Gravely / det.
<determinerName id="A689DA33713DFFADFD0B0B5C0D14FEFC" box="[650,811,317,341]" pageId="10" pageNumber="9">M.J. Paulsen</determinerName>
2015”. Type locality: “Doi Sutep, N.
<collectingCountry id="2BFD7667713DFFADFB690B5F0B65FEFC" box="[1256,1370,317,341]" name="Thailand" pageId="10" pageNumber="9">Thailand</collectingCountry>
”.
</materialsCitation>
</paragraph>
<paragraph id="535536F7713DFFADFF410B0F0F88FE4C" blockId="10.[192,1441,365,485]" pageId="10" pageNumber="9">
<emphasis id="619EEAE5713DFFADFF410B0F0CE3FE2C" bold="true" box="[192,732,365,389]" pageId="10" pageNumber="9">
Additional material examined (
<figureCitation id="CBD12A72713DFFADFDEF0B0F0CF2FE2C" box="[622,717,365,389]" captionStart="Figure 14" captionStartId="11.[192,275,1877,1899]" captionTargetBox="[236,1395,288,1824]" captionTargetId="figure@11.[236,1395,288,1824]" captionTargetPageId="11" captionText="Figure 14. Distribution of obesus group species: N. obesus (triangles), N. dawnae (squares), N. gravelyi Paulsen, n. sp. (diamonds), N. helleri (circles)." figureDoi="http://doi.org/10.5281/zenodo.3699479" httpUri="https://zenodo.org/record/3699479/files/figure.png" pageId="10" pageNumber="9">Fig. 14</figureCitation>
).
</emphasis>
<materialsCitation id="E3823CAA713DFFADFD650B0F0ACDFE2C" ID-GBIF-Occurrence="2574936453" box="[740,1266,365,390]" collectionCode="CASC" country="Laos" location="Ban Tong" pageId="10" pageNumber="9" specimenCount="1" stateProvince="OUDOMXAY">
<emphasis id="619EEAE5713DFFADFD650B0F0D0CFE2C" bold="true" box="[740,819,365,389]" pageId="10" pageNumber="9">
<collectingCountry id="2BFD7667713DFFADFD650B0F0D0CFE2C" box="[740,819,365,389]" name="Laos" pageId="10" pageNumber="9">LAOS</collectingCountry>
</emphasis>
:
<collectingRegion id="912EF815713DFFADFCC30B0F0DD4FE2C" box="[834,1003,365,389]" country="Laos" name="Oudomxai" pageId="10" pageNumber="9">OUDOMXAY</collectingRegion>
:
<location id="5635602C713DFFADFC770B0C0A52FE2F" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFADFF410C920DB6FB6C:5635602C713DFFADFC770B0C0A52FE2F" box="[1014,1133,366,390]" country="Laos" name="Ban Tong" pageId="10" pageNumber="9" stateProvince="OUDOMXAY">Ban Tong</location>
(7,
<collectionCode id="35FBAE32713DFFADFB1A0B0F0AD5FE2C" box="[1179,1258,365,389]" collectionName="USA, California, San Francisco, California Academy of Sciences" pageId="10" pageNumber="9">CASC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="E3823CAA713DFFADFB7C0B0F0C79FE0C" ID-GBIF-Occurrence="2574936432" collectionCode="MJPC" country="Myanmar" location="Dawna Hills" pageId="10" pageNumber="9" specimenCount="1" stateProvince="KAYIN">
<emphasis id="619EEAE5713DFFADFB7C0B0F0BA7FE2C" bold="true" box="[1277,1432,365,389]" pageId="10" pageNumber="9">
<collectingCountry id="2BFD7667713DFFADFB7C0B0F0BA7FE2C" box="[1277,1432,365,389]" name="Myanmar" pageId="10" pageNumber="9">MYANMAR</collectingCountry>
</emphasis>
:
<collectingRegion id="912EF815713DFFADFF410BEF0F20FE0C" box="[192,287,397,421]" country="Myanmar" name="Kayin" pageId="10" pageNumber="9">KAYIN</collectingRegion>
:
<location id="5635602C713DFFADFEAA0BEC0FFAFE0C" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFADFF410C920DB6FB6C:5635602C713DFFADFEAA0BEC0FFAFE0C" box="[299,453,397,421]" country="Myanmar" name="Dawna Hills" pageId="10" pageNumber="9" stateProvince="KAYIN">Dawna Hills</location>
(2,
<collectionCode id="35FBAE32713DFFADFE740BEF0C79FE0C" box="[501,582,397,421]" collectionName="MJPC" pageId="10" pageNumber="9">MJPC</collectionCode>
</materialsCitation>
).
<materialsCitation id="E3823CAA713DFFADFDDB0BEF0AD9FE0C" ID-GBIF-Occurrence="2574936434" box="[602,1254,397,421]" collectionCode="MJPC" country="Thailand" location="Chiang Dao" pageId="10" pageNumber="9" specimenCount="1" stateProvince="CHIANG MAI">
<emphasis id="619EEAE5713DFFADFDDB0BEF0CC7FE0C" bold="true" box="[602,760,397,421]" pageId="10" pageNumber="9">
<collectingCountry id="2BFD7667713DFFADFDDB0BEF0CC7FE0C" box="[602,760,397,421]" name="Thailand" pageId="10" pageNumber="9">THAILAND</collectingCountry>
</emphasis>
:
<collectingRegion id="912EF815713DFFADFC860BEF0D83FE0C" box="[775,956,397,421]" country="Thailand" name="Chiang Mai" pageId="10" pageNumber="9">CHIANG MAI</collectingRegion>
:
<location id="5635602C713DFFADFC460BEF0A65FE0C" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFADFF410C920DB6FB6C:5635602C713DFFADFC460BEF0A65FE0C" box="[967,1114,397,421]" country="Thailand" name="Chiang Dao" pageId="10" pageNumber="9" stateProvince="CHIANG MAI">Chiang Dao</location>
(1,
<collectionCode id="35FBAE32713DFFADFB080BEF0AE5FE0C" box="[1161,1242,397,421]" collectionName="MJPC" pageId="10" pageNumber="9">MJPC</collectionCode>
).
</materialsCitation>
<materialsCitation id="E3823CAA713DFFADFB6F0BEF0FC6FE6C" ID-GBIF-Occurrence="2574936455" collectionCode="MJPC" location="Wiang Pa Pao" pageId="10" pageNumber="9" specimenCount="1" stateProvince="CHIANG RAI">
<collectingRegion id="912EF815713DFFADFB6F0BEF0BA3FE0C" box="[1262,1436,397,421]" country="Thailand" name="Chiang Rai" pageId="10" pageNumber="9">CHIANG RAI</collectingRegion>
:
<location id="5635602C713DFFADFF410BCF0F51FE6C" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFADFF410C920DB6FB6C:5635602C713DFFADFF410BCF0F51FE6C" box="[192,366,429,453]" name="Wiang Pa Pao" pageId="10" pageNumber="9" stateProvince="CHIANG RAI">Wiang Pa Pao</location>
(2,
<collectionCode id="35FBAE32713DFFADFE1D0BCF0FD2FE6C" box="[412,493,429,453]" collectionName="MJPC" pageId="10" pageNumber="9">MJPC</collectionCode>
).
</materialsCitation>
<materialsCitation id="E3823CAA713DFFADFD810BCF0AE9FE6C" ID-GBIF-Occurrence="2574936454" box="[512,1238,429,453]" collectionCode="MJPC" location="Thong Pha Phoom / Phu Yae" pageId="10" pageNumber="9" specimenCount="1" stateProvince="KANCHANABURI">
<collectingRegion id="912EF815713DFFADFD810BCF0CD1FE6C" box="[512,750,429,453]" country="Thailand" name="Kanchanaburi" pageId="10" pageNumber="9">KANCHANABURI</collectingRegion>
:
<location id="5635602C713DFFADFD780BCF0A74FE6C" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFADFF410C920DB6FB6C:5635602C713DFFADFD780BCF0A74FE6C" box="[761,1099,429,453]" name="Thong Pha Phoom / Phu Yae" pageId="10" pageNumber="9" stateProvince="KANCHANABURI">Thong Pha Phoom/Phu Yae</location>
(1,
<collectionCode id="35FBAE32713DFFADFBF80BCF0AF5FE6C" box="[1145,1226,429,453]" collectionName="MJPC" pageId="10" pageNumber="9">MJPC</collectionCode>
).
</materialsCitation>
<materialsCitation id="E3823CAA713DFFADFB5C0BCF0F88FE4C" ID-GBIF-Occurrence="2574936430" collectionCode="NHM" location="Umphang Village" pageId="10" pageNumber="9" specimenCount="450" stateProvince="TAK">
<collectingRegion id="912EF815713DFFADFB5C0BCF0B25FE6C" box="[1245,1306,429,453]" country="Thailand" name="Tak" pageId="10" pageNumber="9">TAK</collectingRegion>
:
<location id="5635602C713DFFADFAA70BCF0F29FE4C" LSID="urn:lsid:plazi:treatment:DB4387E1713FFFADFF410C920DB6FB6C:5635602C713DFFADFAA70BCF0F29FE4C" name="Umphang Village" pageId="10" pageNumber="9" stateProvince="TAK">Umphang Village</location>
(
<specimenCount id="45ECFD7E713DFFADFEA70BAF0F69FE4C" box="[294,342,461,485]" pageId="10" pageNumber="9" type="generic">450</specimenCount>
,
<collectionCode id="35FBAE32713DFFADFEE00BAF0F95FE4C" box="[353,426,461,485]" pageId="10" pageNumber="9">NHM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="1BF0657C713DFFADFF410B9F0D23FD3C" pageId="10" pageNumber="9" type="diagnosis">
<paragraph id="535536F7713DFFADFF410B9F0D23FD3C" blockId="10.[192,1441,509,661]" pageId="10" pageNumber="9">
<emphasis id="619EEAE5713DFFADFF410B9F0F70FDBC" bold="true" box="[192,335,509,533]" pageId="10" pageNumber="9">Diagnosis.</emphasis>
<taxonomicName id="94EA4D74713DFFADFED40B9F0C19FDBC" box="[341,550,509,533]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="619EEAE5713DFFADFED40B9F0C19FDBC" box="[341,550,509,533]" italics="true" pageId="10" pageNumber="9">Nigidius dawnae</emphasis>
</taxonomicName>
has male genitalia that are most similar to
<emphasis id="619EEAE5713DFFADFBBC0B9F0A87FDBC" box="[1085,1208,509,533]" italics="true" pageId="10" pageNumber="9">
<taxonomicName id="94EA4D74713DFFADFBBC0B9F0A8CFDBC" authorityName="Parry" authorityYear="1864" box="[1085,1203,509,533]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="obesus">N. obesus</taxonomicName>
,
</emphasis>
with a sinuous flagellum bearing a small lobe near the apex (
<figureCitation id="CBD12A72713DFFADFD46087F0D23FD9C" box="[711,796,541,565]" captionStart="Figures 1013" captionStartId="9.[192,287,1578,1600]" captionTargetBox="[221,1399,218,1550]" captionTargetPageId="9" captionText="Figures 1013. Male genitalia of obesus group species. 10) N. obesus. 11) N. dawnae, arrow indicates small apical lobe. 12) N. gravelyi Paulsen, n. sp. 13) N. helleri, arrow indicates large lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3699477" httpUri="https://zenodo.org/record/3699477/files/figure.png" pageId="10" pageNumber="9">Fig. 11</figureCitation>
). The flagellum is unique in having a greatly reduced lateral area that is enlarged in the remaining species (forming a large, lateral lobe in the Indonesian species). The elytral intervals are more irregularly punctate than in
<taxonomicName id="94EA4D74713DFFADFC6C083F0A61FDDC" authorityName="Parry" authorityYear="1864" box="[1005,1118,605,629]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="619EEAE5713DFFADFC6C083F0A61FDDC" box="[1005,1118,605,629]" italics="true" pageId="10" pageNumber="9">N. obesus</emphasis>
</taxonomicName>
, and the basal mandibular tooth of males is much broader than in
<taxonomicName id="94EA4D74713DFFADFD20081F0D2AFD3C" authorityName="Parry" authorityYear="1864" box="[673,789,637,661]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="619EEAE5713DFFADFD20081F0D2AFD3C" box="[673,789,637,661]" italics="true" pageId="10" pageNumber="9">N. obesus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="1BF0657C713DFFADFF4108CF0DB6FB6C" pageId="10" pageNumber="9" type="discussion">
<paragraph id="535536F7713DFFADFF4108CF0DB6FB6C" blockId="10.[192,1441,685,1221]" pageId="10" pageNumber="9">
<emphasis id="619EEAE5713DFFADFF4108CF0F7CFD6C" bold="true" box="[192,323,685,709]" pageId="10" pageNumber="9">Remarks.</emphasis>
The distribution of
<taxonomicName id="94EA4D74713DFFADFDB208CF0C8AFD6C" box="[563,693,685,709]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="619EEAE5713DFFADFDB208CF0C8AFD6C" box="[563,693,685,709]" italics="true" pageId="10" pageNumber="9">N. dawnae</emphasis>
</taxonomicName>
specimens examined is shown in
<figureCitation id="CBD12A72713DFFADFBCC08CF0A9BFD6C" box="[1101,1188,685,709]" captionStart="Figure 14" captionStartId="11.[192,275,1877,1899]" captionTargetBox="[236,1395,288,1824]" captionTargetId="figure@11.[236,1395,288,1824]" captionTargetPageId="11" captionText="Figure 14. Distribution of obesus group species: N. obesus (triangles), N. dawnae (squares), N. gravelyi Paulsen, n. sp. (diamonds), N. helleri (circles)." figureDoi="http://doi.org/10.5281/zenodo.3699479" httpUri="https://zenodo.org/record/3699479/files/figure.png" pageId="10" pageNumber="9">Fig. 14</figureCitation>
. Gravelys concept of Tenasserim included the Dawna Hills, so it cannot be assumed that the specimen labeled simply “Tenasserim” is from Tanintharyi state farther south, which is the current, more restricted interpretation of Tenasserim. A small series of
<taxonomicName id="94EA4D74713DFFADFDA2096F0C9BFC8C" box="[547,676,781,805]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="619EEAE5713DFFADFDA2096F0C9BFC8C" box="[547,676,781,805]" italics="true" pageId="10" pageNumber="9">N. dawnae</emphasis>
</taxonomicName>
from CASC are labeled as being from Ban Tong,
<collectingCountry id="2BFD7667713DFFADFB67096C0B1CFC8C" box="[1254,1315,782,805]" name="Laos" pageId="10" pageNumber="9">Laos</collectingCountry>
. A cursory general search online for the locality resulted in eleven such place-names.
<bibRefCitation id="377B4B06713DFFADFBD1094C0AC4FCEC" author="Burton" box="[1104,1275,813,837]" firstAuthor="Burton" pageId="10" pageNumber="9" refId="ref6674" refString="Burton, J. S. B. 1978. Tabanini of Thailand above the Isthmus of Kra (Diptera: Tabanidae). Entomological Reprint Specialists; Los Angeles, CA. 165 p." type="book" year="1978">Burton (1978)</bibRefCitation>
attempted to reconcile the labels of the collector in question, Vitalis de Salvaza, which he noted were overly specific to localized place-names and for the most part untraceable. He indicated that de Salvaza collected in Panghai,
<collectingCountry id="2BFD7667713DFFADFEB409EC0F50FC0C" box="[309,367,910,933]" name="Laos" pageId="10" pageNumber="9">Laos</collectingCountry>
on
<date id="27541037713DFFADFE1A09EF0C3AFC0C" box="[411,517,909,933]" pageId="10" pageNumber="9" value="1920-05-08">8.v.1920</date>
, which is about one month after the Ban Tong specimens date of
<date id="27541037713DFFADFAA809EF0BA3FC0C" box="[1321,1436,909,933]" pageId="10" pageNumber="9" value="1920-04-09">9.iv.1920</date>
. Panghai is in
<collectingRegion id="912EF815713DFFADFEF109CF0C59FC6C" box="[368,614,941,965]" country="Laos" name="Xaignabouli" pageId="10" pageNumber="9">Sainyabuli province</collectingRegion>
less than
<quantity id="94129B12713DFFADFD6A09CF0D06FC6C" box="[747,825,941,965]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="10" pageNumber="9" unit="km" value="14.0">14 km</quantity>
from
<collectingRegion id="912EF815713DFFADFC0709CF0ABBFC6C" box="[902,1156,941,965]" country="Thailand" name="Chiang Rai" pageId="10" pageNumber="9">Chiang Rai province</collectingRegion>
of
<collectingCountry id="2BFD7667713DFFADFB2F09CF0B1FFC6C" box="[1198,1312,941,965]" name="Thailand" pageId="10" pageNumber="9">Thailand</collectingCountry>
, where
<taxonomicName id="94EA4D74713DFFADFA0209CF0F20FC4C" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="dawnae">
<emphasis id="619EEAE5713DFFADFA0209CF0F20FC4C" italics="true" pageId="10" pageNumber="9">N. dawnae</emphasis>
</taxonomicName>
is known to occur. A further complication is that
<bibRefCitation id="377B4B06713DFFADFC0109AF0A2AFC4C" author="Beuk" box="[896,1045,973,997]" firstAuthor="Beuk" pageId="10" pageNumber="9" pagination="1 - 84" refId="ref6505" refString="Beuk, P. L. T. 1999. Revision of the cicadas of the Platylomia spinosa group (Homoptera: Cicadidae). Oriental Insects 33: 1 - 84." type="journal article" year="1999">Beuk (1999)</bibRefCitation>
lists a BMNH specimen label of “
<collectingCountry id="2BFD7667713DFFADFF4A098F0F03FBAC" box="[203,316,1005,1029]" name="Thailand" pageId="10" pageNumber="9">Thailand</collectingCountry>
, Ban Tong,
<date id="27541037713DFFADFE50098F0C7DFBAC" box="[465,578,1005,1029]" pageId="10" pageNumber="9" value="1920-04-09">9.iv.1920</date>
, R. V. de Salvaza”, giving the exact date but with a different country, perhaps due to the differently defined boundaries of the time. I have found no other instance of a Vitalis de Salvaza label attributed to
<collectingCountry id="2BFD7667713DFFADFDAE0E4F0C9EFBEC" box="[559,673,1069,1093]" name="Thailand" pageId="10" pageNumber="9">Thailand</collectingCountry>
, and he seems to have collected most frequently in present-day
<collectingCountry id="2BFD7667713DFFADFF410E2C0EC1FBCC" box="[192,254,1102,1125]" name="Laos" pageId="10" pageNumber="9">Laos</collectingCountry>
. The most likely Ban Tong locality is at
<geoCoordinate id="36DE5030713DFFADFD710E2F0D44FBCC" box="[752,891,1100,1125]" degrees="19" direction="north" minutes="57" orientation="latitude" pageId="10" pageNumber="9" precision="15" seconds="53" value="19.964724">19°5753″N</geoCoordinate>
,
<geoCoordinate id="36DE5030713DFFADFC090E2F0A20FBCC" box="[904,1055,1100,1125]" degrees="100" direction="east" minutes="57" orientation="longitude" pageId="10" pageNumber="9" precision="15" seconds="18" value="100.954994">100°5718″E</geoCoordinate>
, in
<collectingRegion id="912EF815713DFFADFBCC0E2F0B7AFBCC" box="[1101,1349,1101,1125]" country="Laos" name="Oudomxai" pageId="10" pageNumber="9">Oudomxay Province</collectingRegion>
not far removed (~
<quantity id="94129B12713DFFADFECB0E0F0FA6FB2C" box="[330,409,1133,1157]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="9" unit="km" value="45.0">45 km</quantity>
) from
<collectingRegion id="912EF815713DFFADFE6C0E0F0C43FB2C" box="[493,636,1133,1157]" country="Thailand" name="Chiang Rai" pageId="10" pageNumber="9">Chiang Rai</collectingRegion>
,
<collectingCountry id="2BFD7667713DFFADFD0B0E0F0CC6FB2C" box="[650,761,1133,1157]" name="Thailand" pageId="10" pageNumber="9">Thailand</collectingCountry>
or Panghai,
<collectingCountry id="2BFD7667713DFFADFC1A0E0C0DE6FB2C" box="[923,985,1134,1157]" name="Laos" pageId="10" pageNumber="9">Laos</collectingCountry>
. This locality has been indicated on the map with a question mark. Regardless, the specimens from
<collectingCountry id="2BFD7667713DFFADFC550EEC0A31FB0C" box="[980,1038,1166,1189]" name="Laos" pageId="10" pageNumber="9">Laos</collectingCountry>
constitute a new country record. Specimens examined are from April July and December.
</paragraph>
</subSubSection>
</treatment>
</document>