treatments-xml/data/CD/90/6B/CD906BED76E95CD593172534AA323D41.xml
2024-06-21 12:52:06 +02:00

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1033.60686" ID-GBIF-Dataset="920fdd6f-b0e8-4059-8ff8-dd401a03625f" ID-PMC="PMC8084852" ID-Pensoft-Pub="1313-2970-1033-81" ID-Pensoft-UUID="2E6467D3D0025F778A77798A4072BCB5" ID-PubMed="33958921" ID-ZooBank="BEFB015F4A354987AD10DFEB7D4F91DD" ModsDocID="1313-2970-1033-81" checkinTime="1619185013811" checkinUser="pensoft" docAuthor="Pinho, Luiz Carlos &amp; Andersen, Trond" docDate="2021" docId="CD906BED76E95CD593172534AA323D41" docLanguage="en" docName="ZooKeys 1033: 81-125" docOrigin="ZooKeys 1033" docPubDate="2021-04-22" docSource="http://dx.doi.org/10.3897/zookeys.1033.60686" docTitle="Nilothauma maya Pinho &amp; Andersen 2021, sp. nov." docType="treatment" docUuid="0165F284-D9E4-4FF1-B207-C4EE505A0E00" docUuidSource="ZooBank" docVersion="4" id="2E6467D3D0025F778A77798A4072BCB5" lastPageNumber="81" masterDocId="2E6467D3D0025F778A77798A4072BCB5" masterDocTitle="Neotropical Nilothauma Kieffer, 1921 (Diptera, Chironomidae): Key, eleven new species, re-descriptions, new combination and new records" masterLastPageNumber="125" masterPageNumber="81" pageNumber="81" updateTime="1668152459540" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Neotropical Nilothauma Kieffer, 1921 (Diptera, Chironomidae): Key, eleven new species, re-descriptions, new combination and new records</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pinho, Luiz Carlos</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9153-9997</mods:nameIdentifier>
<mods:affiliation>Laboratory of Systematics of Diptera, Department of Ecology and Zoology, Federal University of Santa Catarina, Campus Trindade, CEP 88040 - 900, Florianopolis, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">luizcarlospinho@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Andersen, Trond</mods:namePart>
<mods:affiliation>Department of Natural History, University Museum of Bergen, University of Bergen, P. O. Box 7800, NO- 5020, Bergen, Norway</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1033</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>81</mods:start>
<mods:end>125</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1033.60686</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1033.60686</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1033-81</mods:identifier>
<mods:identifier type="ZooBank">BEFB015F4A354987AD10DFEB7D4F91DD</mods:identifier>
<mods:identifier type="Pensoft-UUID">2E6467D3D0025F778A77798A4072BCB5</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="181534229" LSID="urn:lsid:zoobank.org:act:0165F284-D9E4-4FF1-B207-C4EE505A0E00" httpUri="http://treatment.plazi.org/id/CD906BED76E95CD593172534AA323D41" lastPageNumber="81" pageId="0" pageNumber="81">
<subSubSection pageId="0" pageNumber="81" type="nomenclature">
<paragraph pageId="0" pageNumber="81">
<taxonomicName LSID="http://zoobank.org/0165F284-D9E4-4FF1-B207-C4EE505A0E00" authority="Pinho &amp; Andersen, 2021" authorityName="Pinho &amp; Andersen" authorityYear="2021" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nilothauma maya" order="Diptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="maya" status="sp. nov.">Nilothauma maya</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="81">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Nilothauma maya sp. nov. adult male A hypopygium, dorsal view B hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right." figureDoi="10.3897/zookeys.1033.60686.figure10" httpUri="https://binary.pensoft.net/fig/534703" pageId="0" pageNumber="81">Figures 10A, B</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution maps of Neotropical Nilothauma species A N. hamadae sp. nov., N. jupau sp. nov., N. terena sp. nov., N. yekwana sp. nov., N. complicatum Mendes &amp; Andersen, 2009 B N. leccii sp. nov., N. marianoi sp. nov., N. mateusi sp. nov., N. txukuyana sp. nov., N. maya sp. nov. C N. karitiana sp. nov., N. duena Roback, 1960, N. calori Mendes &amp; Andersen, 2009 D N. werekena sp. nov., N. zitoi Mendes &amp; Andersen, 2009, N. strebulosum (Adam &amp; Saether, 2000)." figureDoi="10.3897/zookeys.1033.60686.figure16" httpUri="https://binary.pensoft.net/fig/534709" pageId="0" pageNumber="81">, 16B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="type material">
<paragraph pageId="0" pageNumber="81">Type material.</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Holotype</emphasis>
male, slide-mounted: Mexico, Campeche, Calacmul, Ejido Nuevo Becan, El Chorro,
<geoCoordinate degrees="18" direction="north" minutes="35" orientation="latitude" precision="15" seconds="26" value="18.590557">18°35'26&quot;N</geoCoordinate>
,
<geoCoordinate degrees="89" direction="west" minutes="15" orientation="longitude" precision="15" seconds="29" value="-89.25806">89°15'29&quot;W</geoCoordinate>
, 130 m a.s.l., 30.iv.1997, light trap, A. Contreras-Ramos et al. leg. (ZMBN).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="etymology">
<paragraph pageId="0" pageNumber="81">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="81">Named after the Maya people, who used to live in the area. The name is to be regarded as a noun in apposition.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="description">
<paragraph pageId="0" pageNumber="81">Diagnostic characters.</paragraph>
<paragraph pageId="0" pageNumber="81">The male can be distinguished from its congeners by the combination of: pale brown species; wing without markings; tergite IX without setose dorsal lobe(s) or spine; anal point spatulate; superior volsella slender, curved, tapering; gonostylus curved, with strong setae on protruberance on inner margin in basal one-third.</paragraph>
<caption doi="10.3897/zookeys.1033.60686.figure10" httpUri="https://binary.pensoft.net/fig/534703" pageId="0" pageNumber="81" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figure 10.</emphasis>
<taxonomicName authorityName="Pinho &amp; Andersen" authorityYear="2021" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nilothauma maya" order="Diptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="maya">
<emphasis italics="true" pageId="0" pageNumber="81">Nilothauma maya</emphasis>
</taxonomicName>
sp. nov. adult male
<emphasis bold="true" pageId="0" pageNumber="81">A</emphasis>
hypopygium, dorsal view
<emphasis bold="true" pageId="0" pageNumber="81">B</emphasis>
hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="description">
<paragraph pageId="0" pageNumber="81">Description.</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Male imago (n = 1).</emphasis>
Total length 2.43 mm. Wing length 1.12 mm. Total length/wing length 2.17. Wing length/length of profemur 2.15.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Colouration</emphasis>
.
</emphasis>
Pale brown. Wing membrane without dark markings.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Antenna</emphasis>
.
</emphasis>
AR = 0.18. Thirteenth flagellomere 112
<normalizedToken originalValue="µm">µm</normalizedToken>
long.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Head</emphasis>
.
</emphasis>
Temporal setae 4 in single row. Clypeus with 13 setae. Tentorium and stipes not measurable. Palp segment lengths (in
<normalizedToken originalValue="µm">µm</normalizedToken>
): 21, 28, 65, 117, 144. Third palpomere with 2 sensilla clavata subapically, longest 19
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Fifth palpomere/third palpomere 2.22.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Thorax</emphasis>
.
</emphasis>
Dorsocentrals 8 in single row, acrostichals 8, prealars 2. Scutellum with 2 setae.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Wing</emphasis>
.
</emphasis>
VR = 1.44. Brachiolum with 1 seta, R with 10 setae, R1 with 12 setae, R4+5 with 15 setae, remaining veins bare.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Legs</emphasis>
.
</emphasis>
Spur of fore tibia 48
<normalizedToken originalValue="µm">µm</normalizedToken>
long including 21
<normalizedToken originalValue="µm">µm</normalizedToken>
long scale. Mid-tibia with 1 spur, 26
<normalizedToken originalValue="µm">µm</normalizedToken>
long; hind tibia with 2 spurs, 25 and 33
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Combs of mid-tibia 12
<normalizedToken originalValue="µm">µm</normalizedToken>
long, of hind tibia 17
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Width at apex of fore tibia 36
<normalizedToken originalValue="µm">µm</normalizedToken>
, of mid-tibia 40
<normalizedToken originalValue="µm">µm</normalizedToken>
, of hind tibia 44
<normalizedToken originalValue="µm">µm</normalizedToken>
. Lengths and proportions of legs as in Table
<tableCitation captionStart="Table 9" captionStartId="T9" captionText="Table 9. Lengths (in μm) and proportions of leg segments in Nilothauma maya sp. nov., adult male (n = 1)." httpUri="http://table.plazi.org/id/F4C40C3B97AF330C2A17334A8FC43E24" pageId="0" pageNumber="81" tableUuid="F4C40C3B97AF330C2A17334A8FC43E24">9</tableCitation>
.
</paragraph>
<caption ID-Table-UUID="F4C40C3B97AF330C2A17334A8FC43E24" httpUri="http://table.plazi.org/id/F4C40C3B97AF330C2A17334A8FC43E24" pageId="0" pageNumber="81" start="Table 9" startId="T9">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Table 9.</emphasis>
Lengths (in
<normalizedToken originalValue="μm">μm</normalizedToken>
) and proportions of leg segments in
<taxonomicName authorityName="Pinho &amp; Andersen" authorityYear="2021" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nilothauma maya" order="Diptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="maya">
<emphasis italics="true" pageId="0" pageNumber="81">Nilothauma maya</emphasis>
</taxonomicName>
sp. nov., adult male (n = 1).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="81">
<table pageId="0" pageNumber="81">
<tr pageId="0" pageNumber="81">
<th colspan="1" isEmpty="true" pageId="0" pageNumber="81" rowspan="1">-</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">Fe</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">Ti</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">ta1</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">ta2</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">ta3</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">ta4</th>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">p1</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">520</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">372</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">p2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">484</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">308</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">208</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">76</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">56</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">32</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">p3</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">504</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">484</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">232</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">132</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">140</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">88</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" isEmpty="true" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">ta5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">LR</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">BV</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">SV</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">BR</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">p1</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">p2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">28</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">0.68</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">5.21</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3.81</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3.4</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">p3</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">48</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">0.48</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">2.99</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">4.26</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">4.7</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Hypopygium</emphasis>
(Fig.
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Nilothauma maya sp. nov. adult male A hypopygium, dorsal view B hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right." figureDoi="10.3897/zookeys.1033.60686.figure10" httpUri="https://binary.pensoft.net/fig/534703" pageId="0" pageNumber="81">10A, B</figureCitation>
). Tergite IX without lobes, with 8 setae above anal point and 5 somewhat weaker setae to each side of anal point. Anal point spatulate, 36
<normalizedToken originalValue="µm">µm</normalizedToken>
long, maximum width 11
<normalizedToken originalValue="µm">µm</normalizedToken>
. Tergite bands continuous. Laterosternite IX with 2 setae. Phallapodeme 48
<normalizedToken originalValue="µm">µm</normalizedToken>
long; transverse sternapodeme 27
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Gonocoxite 104
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Inferior volsella curved, 55
<normalizedToken originalValue="µm">µm</normalizedToken>
long, 14
<normalizedToken originalValue="µm">µm</normalizedToken>
wide subapically, with microtrichia and 4 strong apical setae. Superior volsella slender, curved, tapering, 54
<normalizedToken originalValue="µm">µm</normalizedToken>
long, 11
<normalizedToken originalValue="µm">µm</normalizedToken>
wide at base, 3
<normalizedToken originalValue="µm">µm</normalizedToken>
wide subapically, covered with microtrichia and with 2 weak apical setae. Median volsella consisting of two small tubercles, about 4
<normalizedToken originalValue="µm">µm</normalizedToken>
long, each with single setae at apex, longest 15
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Gonostylus 131
<normalizedToken originalValue="µm">µm</normalizedToken>
long, curved, with single, strong setae on protuberance on inner margin at 27
<normalizedToken originalValue="µm">µm</normalizedToken>
from base, setae 21 mm long. HR = 0.79. HV = 1.85.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="female adult and immatures">
<paragraph pageId="0" pageNumber="81">Female adult and immatures.</paragraph>
<paragraph pageId="0" pageNumber="81">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="remarks">
<paragraph pageId="0" pageNumber="81">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="81">The slide is distorted and the drawings are composites of left and right side.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="distribution">
<paragraph pageId="0" pageNumber="81">Distribution</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
(Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution maps of Neotropical Nilothauma species A N. hamadae sp. nov., N. jupau sp. nov., N. terena sp. nov., N. yekwana sp. nov., N. complicatum Mendes &amp; Andersen, 2009 B N. leccii sp. nov., N. marianoi sp. nov., N. mateusi sp. nov., N. txukuyana sp. nov., N. maya sp. nov. C N. karitiana sp. nov., N. duena Roback, 1960, N. calori Mendes &amp; Andersen, 2009 D N. werekena sp. nov., N. zitoi Mendes &amp; Andersen, 2009, N. strebulosum (Adam &amp; Saether, 2000)." figureDoi="10.3897/zookeys.1033.60686.figure16" httpUri="https://binary.pensoft.net/fig/534709" pageId="0" pageNumber="81">16B</figureCitation>
).
</emphasis>
Only known from Campeche State in Mexico.
</paragraph>
</subSubSection>
</treatment>
</document>