treatments-xml/data/2B/3A/87/2B3A8784FFB7FFBEEBEAFBEC5442FE47.xml

<document id="3E9030C6AFF59AAE50320392AFEE71B7" ID-CLB-Dataset="119273" ID-DOI="10.5852/ejt.2022.834.1899" ID-GBIF-Dataset="8357241b-f19c-4873-9cb2-ae523b668b94" ID-ISSN="2118-9773" ID-Zenodo-Dep="6981051" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1660147112228" checkinUser="felipe" docAuthor="Trovó, Marcelo &amp; Echternacht, Lívia" docDate="2022" docId="2B3A8784FFB7FFBEEBEAFBEC5442FE47" docLanguage="en" docName="ejt-834_58-93.pdf" docOrigin="European Journal of Taxonomy 834" docSource="http://dx.doi.org/10.5852/ejt.2022.834.1899" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Paepalanthus irwinii Trovo &amp; Echtern. 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="68" masterDocId="D703FFFCFFB0FFB5EA15FF915621FFF4" masterDocTitle="The Paepalanthoideae (Eriocaulaceae) of the Chapada dos Veadeiros National Park, Brazil: taxonomic novelties, identification key, and illustrated list of species" masterLastPageNumber="93" masterPageNumber="58" pageNumber="65" updateTime="1699365320394" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="ADB392B627F692B9E7F80D3DC6047D35">The Paepalanthoideae (Eriocaulaceae) of the Chapada dos Veadeiros National Park, Brazil: taxonomic novelties, identification key, and illustrated list of species</mods:title>
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<mods:namePart id="B392E7F19C576FEE52E861B674D77E06">Trovó, Marcelo</mods:namePart>
<mods:affiliation id="A3B8352E6EB0D4538A6F34E92CBDB6C4">Departamento de Botânica, Instituto de Biologia, Universidade Federal do Rio de Janeiro, Avenida Carlos Chagas Filho 373, Rio de Janeiro, RJ 21941 - 590, Brazil.</mods:affiliation>
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<mods:namePart id="C24F2AB0C2889B81CDF9343A18D0031A">Echternacht, Lívia</mods:namePart>
<mods:affiliation id="3BF44752E606BBAD9E66CE862EC3050F">Departamento de Biodiversidade, Evolução e Meio Ambiente, Universidade Federal de Ouro Preto, Campus Morro do Cruzeiro, ICEB III, Ouro Preto, MG 35400 - 000, Brazil.</mods:affiliation>
<mods:nameIdentifier id="86543595710C82FE5F68C5F3D1F6080A" type="email">livia.echter@ufop.edu.br</mods:nameIdentifier>
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<mods:title id="3B67040B1186A2B1A321985EF85799D1">European Journal of Taxonomy</mods:title>
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<mods:date id="08B1FE0D01A9D61C4C344993DD8EBB84">2022</mods:date>
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<treatment id="2B3A8784FFB7FFBEEBEAFBEC5442FE47" ID-DOI="http://doi.org/10.5281/zenodo.6981262" ID-GBIF-Taxon="199013281" ID-Zenodo-Dep="6981262" LSID="urn:lsid:plazi:treatment:2B3A8784FFB7FFBEEBEAFBEC5442FE47" httpUri="http://treatment.plazi.org/id/2B3A8784FFB7FFBEEBEAFBEC5442FE47" lastPageId="11" lastPageNumber="68" pageId="7" pageNumber="65">
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<heading id="F86481FEFFB7FFB2EBEAFBEC5215FB6C" box="[511,1076,1149,1176]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="65" reason="2">
<taxonomicName id="64934D11FFB7FFB2EBEAFBEC55F2FB6C" authority="Trovo &amp; Echtern." authorityName="Trovo &amp; Echtern." authorityYear="2022" box="[511,979,1149,1176]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="7" pageNumber="65" phylum="Tracheophyta" rank="species" species="irwinii" status="sp. nov.">
<emphasis id="91E7EA80FFB7FFB2EBEAFBEC54DCFB6C" bold="true" box="[511,765,1149,1176]" italics="true" pageId="7" pageNumber="65">Paepalanthus irwinii</emphasis>
Trovó &amp; Echtern.
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<taxonomicNameLabel id="8AD457FBFFB7FFB2E9CFFBEF5215FB6C" box="[986,1076,1150,1176]" pageId="7" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="A32C3692FFB7FFB2E852FB3355CCFB48" blockId="7.[511,1076,1149,1250]" box="[583,1005,1186,1212]" pageId="7" pageNumber="65">urn:lsid:ipni.org:names:77302849-1</paragraph>
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<figureCitation id="3BA82A17FFB7FFB2E8DBFB565516FB16" box="[718,823,1223,1250]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[189,232,1892,1918]" captionStartId-1="9.[189,232,1963,1989]" captionTargetBox-0="[227,1361,312,1823]" captionTargetBox-1="[208,1380,265,1922]" captionTargetId-0="figure-85@8.[189,1399,288,1840]" captionTargetId-1="figure-49@9.[208,1380,265,1922]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 4. Paepalanthus irwinii Trovó &amp; Echtern. sp. nov. A. Habit. B. Inflorescence and reproductive apex detail. C. Capitulum detail. D. Involucral bract, abaxial surface. E. Floral bract, abaxial surface. F. Staminate flower. G. Staminate flower with sepals removed and corolla opened. H. Pistillate flower. I. Petal of the pistillate flower, abaxial surface. J. Gynoecium. (M.L.O. Trovó et al. 706 – RB)." captionText-1="Fig. 5. Paepalanthus irwinii Trovó &amp; Echtern. sp. nov. A. Habitat. B. Habitat detail. C. Habit. D. Reproductive axis detail. E. Sprouting rosettes. F. Rosette detail." figureDoi-0="http://doi.org/10.5281/zenodo.6981065" figureDoi-1="http://doi.org/10.5281/zenodo.6981069" httpUri-0="https://zenodo.org/record/6981065/files/figure.png" httpUri-1="https://zenodo.org/record/6981069/files/figure.png" pageId="7" pageNumber="65">Figs 4–5</figureCitation>
, 
<figureCitation id="3BA82A17FFB7FFB2E956FB565546FB15" box="[835,871,1223,1249]" captionStart="Fig" captionStartId="20.[189,232,1892,1918]" captionTargetBox="[228,1362,265,1869]" captionTargetId="figure-71@20.[228,1362,265,1869]" captionTargetPageId="20" captionText="Fig. 7. Species of Paepalanthoideae (Eriocaulaceae) from the Chapada dos Veadeiros National Park. A. Paepalanthus cassiae Trovó. B. P. chiquitensis Herzog. C. P. cordatus Ruhland. D. P. echinoides Trovó. E. P. flaccidus (Bong.) Kunth. F. P. irwinii Trovó &amp; Echtern. sp. nov. G. P. koernickei (Ruhland) Trovó. H. P. longibracteatus (Moldenke) Trovó." figureDoi="http://doi.org/10.5281/zenodo.6981079" httpUri="https://zenodo.org/record/6981079/files/figure.png" pageId="7" pageNumber="65">7A</figureCitation>
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<emphasis id="91E7EA80FFB7FFB2EAA8FA985717FAD7" bold="true" box="[189,310,1289,1315]" pageId="7" pageNumber="65">Diagnosis</emphasis>
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Differs from 
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<emphasis id="91E7EA80FFB7FFB2EB48FAA8549DFAA7" box="[349,700,1337,1363]" italics="true" pageId="7" pageNumber="65">Paepalanthus acanthophyllus</emphasis>
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by being a more gracile plant with shorter, linear leaves, shorter and narrower reproductive axis, shorter, linear to narrowly lanceolate axis bracts, fewer scapes per plant, capitula usually narrower, and petals of the pistillate flower with obtuse to mucronate apex.
</paragraph>
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<emphasis id="91E7EA80FFB7FFB2EAA8FA545765FA2B" bold="true" box="[189,324,1477,1503]" pageId="7" pageNumber="65">Etymology</emphasis>
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The epithet “ 
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<emphasis id="91E7EA80FFB7FFB2EB47FA675781F9FB" box="[338,416,1526,1551]" italics="true" pageId="7" pageNumber="65">irwinii</emphasis>
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” honors Howard Samuel Irwin, the coordinator of the Planalto Expeditions Program, which collected over 225 000 specimens and over 30 000 numbers in Central 
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by 1972 (
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). The 
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set of specimens from the “Planalto Expeditions” is precious, encompassing many 
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specimens.
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<emphasis id="91E7EA80FFB7FFB2EAA8F934578BF94B" bold="true" box="[189,426,1701,1727]" pageId="7" pageNumber="65">Material examined</emphasis>
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<typeStatus id="7C288830FFB7FFB2EAFFF9445706F91B" box="[234,295,1749,1775]" pageId="7" pageNumber="65">Type</typeStatus>
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• 
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, 
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, 
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, GO 239, 
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; 
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, 
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; 
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</date>
; 
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et al. 706
</emphasis>
; 
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: 
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[
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: 
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[
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], 
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.
</paragraph>
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<materialsCitation id="13FB3CCFFFB7FFB2EAFFF814529BF830" ID-GBIF-Occurrence="3886967314" collectingDate="1987-05-09" collectionCode="RB, SPF" collectorName="N. L. Menezes" country="Brazil" location="Alto Paraiso de Goias" pageId="7" pageNumber="65" specimenCount="1" stateProvince="Goias" typeStatus="paratype">
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</emphasis>
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• 
<collectingRegion id="6157F870FFB7FFB2EB51F83B57AAF830" box="[324,395,1962,1988]" country="Brazil" name="Goias" pageId="7" pageNumber="65">Goiás</collectingRegion>
, 
<location id="A64C6049FFB7FFB2EB8DF83B5485F830" LSID="urn:lsid:plazi:treatment:2B3A8784FFB7FFBEEBEAFBEC5442FE47:A64C6049FFB7FFB2EB8DF83B5485F830" box="[408,676,1962,1988]" country="Brazil" name="Alto Paraiso de Goias" pageId="7" pageNumber="65" stateProvince="Goias">Alto Paraíso de Goiás</location>
; 
<date id="D72D1052FFB7FFB2E8A6F83B5568F830" box="[691,841,1962,1988]" pageId="7" pageNumber="65" value="1987-05-09">
<collectingDate id="C769E9BAFFB7FFB2E8A6F83B5568F830" box="[691,841,1962,1988]" pageId="7" pageNumber="65" value="1987-05-09">9 May 1987</collectingDate>
</date>
; 
<emphasis id="91E7EA80FFB7FFB2E94DF83A521EF830" box="[856,1087,1962,1988]" italics="true" pageId="7" pageNumber="65">
<collectorName id="0E665344FFB7FFB2E94DF83A55DBF830" box="[856,1018,1963,1988]" pageId="7" pageNumber="65">N.L. Menezes</collectorName>
1217
</emphasis>
; 
<collectionCode id="C582AE57FFB7FFB2EE44F83B525AF830" box="[1105,1147,1962,1988]" country="Brazil" lsid="urn:lsid:biocol.org:col:15528" name="Jardim Botânico do Rio de Janeiro" pageId="7" pageNumber="65" type="Herbarium">RB</collectionCode>
, 
<collectionCode id="C582AE57FFB7FFB2EE9CF83B529BF830" box="[1161,1210,1962,1988]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15730" lsid="urn:lsid:biocol.org:col:15730" name="Universidade de São Paulo" pageId="7" pageNumber="65" type="Herbarium">SPF</collectionCode>
</materialsCitation>
• 
<materialsCitation id="13FB3CCFFFB7FFB2EECDF83B5247F81C" ID-GBIF-Occurrence="3886967312" collectingDate="1994-05-20" collectionCode="RB, SPF" collectorName="M. Aparecida-da-Silva" country="Brazil" location="Alto Paraiso" pageId="7" pageNumber="65" specimenCount="1" stateProvince="Goias" typeStatus="paratype">
same locality as for preceding; 
<date id="D72D1052FFB7FFB2EB81F85F541AF81C" box="[404,571,1998,2025]" pageId="7" pageNumber="65" value="1994-05-20">
<collectingDate id="C769E9BAFFB7FFB2EB81F85F541AF81C" box="[404,571,1998,2025]" pageId="7" pageNumber="65" value="1994-05-20">20 May 1994</collectingDate>
</date>
; 
<emphasis id="91E7EA80FFB7FFB2E85EF85E55C8F81C" box="[587,1001,1998,2024]" italics="true" pageId="7" pageNumber="65">
<collectorName id="0E665344FFB7FFB2E85EF85E557BF81C" box="[587,858,1998,2024]" pageId="7" pageNumber="65">M. Aparecida-da-Silva</collectorName>
et al. 1921
</emphasis>
; 
<collectionCode id="C582AE57FFB7FFB2E9E9F85E5207F81D" box="[1020,1062,1999,2025]" country="Brazil" lsid="urn:lsid:biocol.org:col:15528" name="Jardim Botânico do Rio de Janeiro" pageId="7" pageNumber="65" type="Herbarium">RB</collectionCode>
, 
<collectionCode id="C582AE57FFB7FFB2EE20F85F5247F81C" box="[1077,1126,1998,2024]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15730" lsid="urn:lsid:biocol.org:col:15730" name="Universidade de São Paulo" pageId="7" pageNumber="65" type="Herbarium">SPF</collectionCode>
</materialsCitation>
• 
<materialsCitation id="13FB3CCFFFB7FFBFEE92F85E55B5FED0" ID-GBIF-Occurrence="3886967303" collectingDate="2009-04-27" collectionCode="HUEFS, RB" collectorName="D. Cardoso" country="Brazil" lastPageId="10" lastPageNumber="68" location="Alto Paraiso" pageId="7" pageNumber="65" specimenCount="1" stateProvince="Goias" typeStatus="paratype">
same locality as for preceding; 
<date id="D72D1052FFBAFFBFEB55FE9857FAFED7" box="[320,475,265,291]" pageId="10" pageNumber="68" value="2009-04-27">
<collectingDate id="C769E9BAFFBAFFBFEB55FE9857FAFED7" box="[320,475,265,291]" pageId="10" pageNumber="68" value="2009-04-27">27 Apr. 2009</collectingDate>
</date>
; 
<emphasis id="91E7EA80FFBAFFBFEBF2FE9B54D2FED7" box="[487,755,265,291]" italics="true" pageId="10" pageNumber="68">
<collectorName id="0E665344FFBAFFBFEBF2FE9B544EFED7" box="[487,623,265,291]" pageId="10" pageNumber="68">D. Cardoso</collectorName>
et al. 2600
</emphasis>
; 
<collectionCode id="C582AE57FFBAFFBFE917FE985543FED7" box="[770,866,265,291]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15506" lsid="urn:lsid:biocol.org:col:15506" name="Universidade Estadual de Feira de Santana" pageId="10" pageNumber="68" type="Herbarium">HUEFS</collectionCode>
, 
<collectionCode id="C582AE57FFBAFFBFE978FE9B55B5FED0" box="[877,916,266,292]" country="Brazil" lsid="urn:lsid:biocol.org:col:15528" name="Jardim Botânico do Rio de Janeiro" pageId="10" pageNumber="68" type="Herbarium">RB</collectionCode>
</materialsCitation>
• 
<materialsCitation id="13FB3CCFFFBAFFBFE9B8FE9B54AEFEB3" ID-GBIF-Occurrence="3886967313" collectingDate="2012-03-22" collectionCode="RB, SPF" collectorName="M. Watanabe" country="Brazil" location="Alto Paraiso" pageId="10" pageNumber="68" specimenCount="1" stateProvince="Goias" typeStatus="paratype">
same locality as for preceding; 
<date id="D72D1052FFBAFFBFEF0EFE9856DAFEB3" pageId="10" pageNumber="68" value="2012-03-22">
<collectingDate id="C769E9BAFFBAFFBFEF0EFE9856DAFEB3" pageId="10" pageNumber="68" value="2012-03-22">22 Mar. 2012</collectingDate>
</date>
; 
<emphasis id="91E7EA80FFBAFFBFEB12FEBF5439FEB3" box="[263,536,301,327]" italics="true" pageId="10" pageNumber="68">
<collectorName id="0E665344FFBAFFBFEB12FEBF5783FEB3" box="[263,418,301,327]" pageId="10" pageNumber="68">M. Watanabe</collectorName>
et al. 227
</emphasis>
; 
<collectionCode id="C582AE57FFBAFFBFE832FEBC5470FEB3" box="[551,593,301,327]" country="Brazil" lsid="urn:lsid:biocol.org:col:15528" name="Jardim Botânico do Rio de Janeiro" pageId="10" pageNumber="68" type="Herbarium">RB</collectionCode>
, 
<collectionCode id="C582AE57FFBAFFBFE848FEBC54AEFEB3" box="[605,655,301,327]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15730" lsid="urn:lsid:biocol.org:col:15730" name="Universidade de São Paulo" pageId="10" pageNumber="68" type="Herbarium">SPF</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="F7EC661AFFB8FFBDEAA8F8F55331F81C" ID-DOI="http://doi.org/10.5281/zenodo.6981065" ID-Zenodo-Dep="6981065" httpUri="https://zenodo.org/record/6981065/files/figure.png" pageId="8" pageNumber="66" startId="8.[189,232,1892,1918]" targetBox="[227,1361,312,1823]" targetPageId="8">
<paragraph id="A32C3692FFB8FFBDEAA8F8F55331F81C" blockId="8.[189,1399,1892,2025]" pageId="8" pageNumber="66">
<emphasis id="91E7EA80FFB8FFBDEAA8F8F5572FF88B" bold="true" box="[189,270,1892,1919]" pageId="8" pageNumber="66">Fig. 4.</emphasis>
<taxonomicName id="64934D11FFB8FFBDEB0DF8F454D2F88B" authority="Trovo &amp; Echtern." authorityName="Trovo &amp; Echtern." authorityYear="2022" box="[280,755,1893,1919]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="8" pageNumber="66" phylum="Tracheophyta" rank="species" species="irwinii" status="sp. nov.">
<emphasis id="91E7EA80FFB8FFBDEB0DF8F45433F88B" box="[280,530,1893,1919]" italics="true" pageId="8" pageNumber="66">Paepalanthus irwinii</emphasis>
Trovó &amp; Echtern.
</taxonomicName>
<taxonomicNameLabel id="8AD457FBFFB8FFBDE8E8F8F4557AF88B" box="[765,859,1893,1919]" pageId="8" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="91E7EA80FFB8FFBDE973F8F5555AF88A" bold="true" box="[870,891,1892,1918]" pageId="8" pageNumber="66">A</emphasis>
. Habit. 
<emphasis id="91E7EA80FFB8FFBDE9F5F8F455D5F88B" bold="true" box="[992,1012,1893,1919]" pageId="8" pageNumber="66">B</emphasis>
. Inflorescence and reproductive apex detail. 
<emphasis id="91E7EA80FFB8FFBDEB45F8195745F856" bold="true" box="[336,356,1928,1954]" pageId="8" pageNumber="66">C</emphasis>
. Capitulum detail. 
<emphasis id="91E7EA80FFB8FFBDE859F8195441F856" bold="true" box="[588,608,1928,1954]" pageId="8" pageNumber="66">D</emphasis>
. Involucral bract, abaxial surface. 
<emphasis id="91E7EA80FFB8FFBDEE15F8195233F856" bold="true" box="[1024,1042,1928,1954]" pageId="8" pageNumber="66">E</emphasis>
. Floral bract, abaxial surface. 
<emphasis id="91E7EA80FFB8FFBDEAABF83A56EFF831" bold="true" box="[190,206,1963,1989]" pageId="8" pageNumber="66">F</emphasis>
. Staminate flower. 
<emphasis id="91E7EA80FFB8FFBDEBA6F83A57EBF831" bold="true" box="[435,458,1963,1989]" pageId="8" pageNumber="66">G</emphasis>
. Staminate flower with sepals removed and corolla opened. 
<emphasis id="91E7EA80FFB8FFBDEE84F83A5289F831" bold="true" box="[1169,1192,1963,1989]" pageId="8" pageNumber="66">H</emphasis>
. Pistillate flower. 
<emphasis id="91E7EA80FFB8FFBDEAA8F85F56E9F81C" bold="true" box="[189,200,1998,2024]" pageId="8" pageNumber="66">I</emphasis>
. Petal of the pistillate flower, abaxial surface. 
<emphasis id="91E7EA80FFB8FFBDE8FDF85F54D6F81C" bold="true" box="[744,759,1998,2024]" pageId="8" pageNumber="66">J</emphasis>
. Gynoecium. (
<emphasis id="91E7EA80FFB8FFBDE9B2F85E529AF81C" box="[935,1211,1998,2025]" italics="true" pageId="8" pageNumber="66">M.L.O. Trovó et al. 706</emphasis>
– RB).
</paragraph>
</caption>
<caption id="F7EC661AFFB9FFBCEAA8F83A55FAF81C" ID-DOI="http://doi.org/10.5281/zenodo.6981069" ID-Zenodo-Dep="6981069" httpUri="https://zenodo.org/record/6981069/files/figure.png" pageId="9" pageNumber="67" startId="9.[189,232,1963,1989]" targetBox="[208,1380,265,1922]" targetPageId="9">
<paragraph id="A32C3692FFB9FFBCEAA8F83A55FAF81C" blockId="9.[189,1399,1963,2025]" pageId="9" pageNumber="67">
<emphasis id="91E7EA80FFB9FFBCEAA8F83A5730F831" bold="true" box="[189,273,1963,1989]" pageId="9" pageNumber="67">Fig. 5.</emphasis>
<taxonomicName id="64934D11FFB9FFBCEB35F83A5528F831" authority="Trovo &amp; Echtern." authorityName="Trovo &amp; Echtern." authorityYear="2022" box="[288,777,1963,1989]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="9" pageNumber="67" phylum="Tracheophyta" rank="species" species="irwinii" status="sp. nov.">
<emphasis id="91E7EA80FFB9FFBCEB35F83A543FF831" box="[288,542,1963,1989]" italics="true" pageId="9" pageNumber="67">Paepalanthus irwinii</emphasis>
Trovó &amp; Echtern.
</taxonomicName>
<taxonomicNameLabel id="8AD457FBFFB9FFBCE902F83D5558F832" box="[791,889,1964,1990]" pageId="9" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="91E7EA80FFB9FFBCE992F83A55BDF831" bold="true" box="[903,924,1963,1989]" pageId="9" pageNumber="67">A</emphasis>
. Habitat. 
<emphasis id="91E7EA80FFB9FFBCEE0BF83A5213F831" bold="true" box="[1054,1074,1963,1989]" pageId="9" pageNumber="67">B</emphasis>
. Habitat detail. 
<emphasis id="91E7EA80FFB9FFBCEF16F83A5339F831" bold="true" box="[1283,1304,1963,1989]" pageId="9" pageNumber="67">C</emphasis>
. Habit. 
<emphasis id="91E7EA80FFB9FFBCEAA8F85F56F3F81C" bold="true" box="[189,210,1998,2024]" pageId="9" pageNumber="67">D</emphasis>
. Reproductive axis detail. 
<emphasis id="91E7EA80FFB9FFBCE818F85F5400F81C" bold="true" box="[525,545,1998,2024]" pageId="9" pageNumber="67">E</emphasis>
. Sprouting rosettes. 
<emphasis id="91E7EA80FFB9FFBCE907F85F5505F81C" bold="true" box="[786,804,1998,2024]" pageId="9" pageNumber="67">F</emphasis>
. Rosette detail.
</paragraph>
</caption>
<subSubSection id="EB896519FFBAFFBFEAA8FEFF54E4FB3E" pageId="10" pageNumber="68" type="description">
<paragraph id="A32C3692FFBAFFBFEAA8FEFF576FFE7C" blockId="10.[189,334,366,392]" box="[189,334,366,392]" pageId="10" pageNumber="68">
<heading id="F86481FEFFBAFFBFEAA8FEFF576FFE7C" bold="true" box="[189,334,366,392]" fontSize="11" level="3" pageId="10" pageNumber="68" reason="3">
<emphasis id="91E7EA80FFBAFFBFEAA8FEFF576FFE7C" bold="true" box="[189,334,366,392]" pageId="10" pageNumber="68">Description</emphasis>
</heading>
</paragraph>
<paragraph id="A32C3692FFBAFFBFEAA8FE0F54E4FB3E" blockId="10.[189,1400,414,1226]" pageId="10" pageNumber="68">
Herbs perennial, solitary or forming small clumps of ramets, 45.0−60.0 cm tall; rhizome short, erect, usually sprouting; rosette stem short, ca 
<quantity id="646B9B77FFBAFFBFE89AFE505532FE28" box="[655,787,449,476]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="10" pageNumber="68" unit="cm" value="1.0" valueMax="1.5" valueMin="0.5">0.5−1.5 cm</quantity>
long. Leaves usually deciduous in fertile specimens, membranaceous to chartaceous, linear, greenish, flat, patent to recurved, 1.0−3.0 × 
<quantity id="646B9B77FFBAFFBFEE71FE7452C8FDF4" box="[1124,1257,485,512]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="10" pageNumber="68" unit="cm" value="0.15" valueMax="0.2" valueMin="0.1">0.1−0.2 cm</quantity>
,glabrescent on both surfaces, margins with long, sparse trichomes, apex acute. Reproductive axis erect, 30.0−45.0 cm long, ca 
<quantity id="646B9B77FFBAFFBFEB36FDBC5755FDB3" box="[291,372,557,583]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="68" unit="cm" value="0.3">0.3 cm</quantity>
diameter, glabrous; all axis bracts with similar shape, amplexicaul, chartaceous, linear to narrowly lanceolate, patent to recurved, 0.5−1.5 × 
<quantity id="646B9B77FFBAFFBFE93AFDC05597FD9F" box="[815,950,593,619]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="10" pageNumber="68" unit="cm" value="0.2" valueMax="0.3" valueMin="0.1">0.1−0.3 cm</quantity>
, apex acute, usually entirely glabrous, the younger with long trichomes towards the margin. Spathes 
<quantity id="646B9B77FFBAFFBFE9B5FDE55211FD7B" box="[928,1072,628,655]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="10" pageNumber="68" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5−4.5 mm</quantity>
long, glabrous, apex acute. Scapes 5−25, arranged at the reproductive axis apex, 12.0−28.0 cm long, glabrous or rarely with sparse trichomes. Capitula 
<quantity id="646B9B77FFBAFFBFEBA5FD2D5462FD22" box="[432,579,700,727]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="10" pageNumber="68" unit="mm" value="0.7" valueMax="0.9" valueMin="0.5">0.5−0.9 mm</quantity>
diam., hemispherical to spherical, whitish; involucral bracts in 2−4 series, dark castaneous, deltate, ca 
<quantity id="646B9B77FFBAFFBFE873FD4E54E2FD0E" box="[614,707,735,762]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="68" unit="mm" value="2.5">2.5 mm</quantity>
long, glabrous or rarely with sparse trichomes abaxially, sparsely ciliate, apex obtuse. Flowers dimerous, ca 80 per capitulum, disposed in concentric rings; floral bracts linear to narrowly oblanceolate, castaneous with central whitish stripe, flat, 1.5−2.0 mm long, mostly glabrous or sparsely pilose distally on the abaxial surface, margin ciliate toward the acute apex, glabrescent. Staminate flowers ca 
<quantity id="646B9B77FFBAFFBFE84EFCFF5499FC7D" box="[603,696,878,905]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="68" unit="mm" value="3.5">3.5 mm</quantity>
long; pedicel with ca 
<quantity id="646B9B77FFBAFFBFE9D9FCFF5208FC7D" box="[972,1065,878,905]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="68" unit="mm" value="0.5">0.5 mm</quantity>
long; sepals free, narrowly obovate, castaneous with central whitish stripe, ca 
<quantity id="646B9B77FFBAFFBFE93CFC0355A6FC59" box="[809,903,914,941]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="68" unit="mm" value="2.5">2.5 mm</quantity>
long, mostly glabrous or sparsely pilose distally on the abaxial surface, densely ciliate toward the acute apex, glabrescent; anthophore fleshy, elongated, ca 
<quantity id="646B9B77FFBAFFBFEB4AFC4B5798FC00" box="[351,441,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="68" unit="mm" value="1.5">1.5 mm</quantity>
long; petals fused into a tube, whitish, membranaceous, ca 
<quantity id="646B9B77FFBAFFBFEE61FC4B52ECFC00" box="[1140,1229,986,1013]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="68" unit="mm" value="0.5">0.5 mm</quantity>
long; stamens ca 
<quantity id="646B9B77FFBAFFBFEAF5FC6F571DFBEC" box="[224,316,1022,1048]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="68" unit="mm" value="0.8">0.8 mm</quantity>
long, cream-colored; carpellodes 3, papillose. Pistillate flowers ca 
<quantity id="646B9B77FFBAFFBFEE4AFC6C529AFBEC" box="[1119,1211,1021,1048]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="68" unit="mm" value="2.5">2.5 mm</quantity>
long, sessile or subsessile; sepals free, dolabriform, whitish, ca 2.0 mm long, densely pilose abaxially, densely ciliate toward the obtuse apex, glabrescent; petals free, dolabriform, whitish, ca 
<quantity id="646B9B77FFBAFFBFEE36FBD4525FFB94" box="[1059,1150,1093,1120]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="68" unit="mm" value="1.5">1.5 mm</quantity>
long, sparsely pilose abaxially, ciliate toward the obtuse to mucronate apex, glabrescent; gynoecium 
<quantity id="646B9B77FFBAFFBFEE72FBF852E0FB77" box="[1127,1217,1129,1156]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="68" unit="mm" value="2.5">2.5 mm</quantity>
long, stigmatic branches bifid at the apex, twice as long as the nectariferous branches, staminodes not seen. Fruit a loculicidal capsule. Seeds ellipsoid, reddish.
</paragraph>
</subSubSection>
<subSubSection id="EB896519FFBAFFBFEAA8FB63523EF9C1" pageId="10" pageNumber="68" type="distribution">
<paragraph id="A32C3692FFBAFFBFEAA8FB635485FAF8" blockId="10.[189,676,1266,1292]" box="[189,676,1266,1292]" pageId="10" pageNumber="68">
<heading id="F86481FEFFBAFFBFEAA8FB635485FAF8" bold="true" box="[189,676,1266,1292]" fontSize="11" level="3" pageId="10" pageNumber="68" reason="3">
<emphasis id="91E7EA80FFBAFFBFEAA8FB635485FAF8" bold="true" box="[189,676,1266,1292]" pageId="10" pageNumber="68">Distribution, habitat, and conservation</emphasis>
</heading>
</paragraph>
<paragraph id="A32C3692FFBAFFBFEAA8FAB0523EF9C1" blockId="10.[187,1399,1313,1589]" pageId="10" pageNumber="68">
Populations of 
<taxonomicName id="64934D11FFBAFFBFEB79FAB05440FACF" authority="Trovó &amp; Echternacht, 2022" authorityName="Trovó &amp; Echternacht" authorityYear="2022" box="[364,609,1313,1339]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="10" pageNumber="68" phylum="Tracheophyta" rank="species" species="irwinii" status="sp. nov.">
<emphasis id="91E7EA80FFBAFFBFEB79FAB05440FACF" box="[364,609,1313,1339]" italics="true" pageId="10" pageNumber="68">Paepalanthus irwinii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8AD457FBFFBAFFBFE87DFAB354E0FAC8" box="[616,705,1314,1340]" pageId="10" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
with ca 10−75 flowering individuals were frequently found in the Chapada dos Veadeiros National Park. Different from most of the endemic species of 
<taxonomicName id="64934D11FFBAFFBFEEC7FAD45356FAAB" authorityName="Martinov" authorityYear="1820" box="[1234,1399,1349,1375]" class="Liliopsida" family="Eriocaulaceae" kingdom="Plantae" order="Poales" pageId="10" pageNumber="68" phylum="Tracheophyta" rank="family">Eriocaulaceae</taxonomicName>
to the area, which are locally restricted, the species is distributed from the Rio Preto Valley in the vicinity of São Jorge village to the Pouso Alto Region. Populations occur in different phytophysiognomies, from open fields to rupestrian savannahs, but the individuals grow only over shallow, usually wet, rocky soils. It is also worth mentioning that many individuals were seen in flower after a severe fire event in 2017. Based on the 
<bibRefCitation id="C7024B63FFBAFFBFEBBDFA695464F9E6" author="IUCN" box="[424,581,1528,1554]" pageId="10" pageNumber="68" refId="ref13396" refString="IUCN 2019. Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Available from: http: // www. iucn. org [accessed 8 Jun. 2021]." type="url" year="2019">IUCN (2019)</bibRefCitation>
distribution criteria, EN: B1ab (i,ii,iii,iv), and its occurrence inside a National Conservation Unit, the species should be considered Vulnerable.
</paragraph>
</subSubSection>
<subSubSection id="EB896519FFBAFFBEEAA8F9CF5442FE47" lastPageId="11" lastPageNumber="69" pageId="10" pageNumber="68" type="discussion">
<paragraph id="A32C3692FFBAFFBFEAA8F9CF5722F98C" blockId="10.[189,259,1630,1656]" box="[189,259,1630,1656]" pageId="10" pageNumber="68">
<heading id="F86481FEFFBAFFBFEAA8F9CF5722F98C" bold="true" box="[189,259,1630,1656]" fontSize="11" level="3" pageId="10" pageNumber="68" reason="3">
<emphasis id="91E7EA80FFBAFFBFEAA8F9CF5722F98C" bold="true" box="[189,259,1630,1656]" pageId="10" pageNumber="68">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="A32C3692FFBAFFBEEAA8F91C5442FE47" blockId="10.[188,1400,1677,2025]" lastBlockId="11.[188,1399,265,435]" lastPageId="11" lastPageNumber="69" pageId="10" pageNumber="68">
<taxonomicName id="64934D11FFBAFFBFEAA8F91C5799F952" authority="Trovó &amp; Echternacht, 2022" authorityName="Trovó &amp; Echternacht" authorityYear="2022" box="[189,440,1677,1703]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="10" pageNumber="68" phylum="Tracheophyta" rank="species" species="irwinii" status="sp. nov.">
<emphasis id="91E7EA80FFBAFFBFEAA8F91C5799F952" box="[189,440,1677,1703]" italics="true" pageId="10" pageNumber="68">Paepalanthus irwinii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8AD457FBFFBAFFBFEBD0F91C5405F953" box="[453,548,1677,1703]" pageId="10" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
belongs to 
<taxonomicName id="64934D11FFBAFFBFE8AFF91C528EF953" authority="Ruhland" authorityName="Ruhland" box="[698,1199,1677,1703]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="10" pageNumber="68" phylum="Tracheophyta" rank="section" section="Diphyomene">
<emphasis id="91E7EA80FFBAFFBFE8AFF91C557AF953" box="[698,859,1677,1703]" italics="true" pageId="10" pageNumber="68">Paepalanthus</emphasis>
sect. 
<emphasis id="91E7EA80FFBAFFBFE9BDF91C521DF953" box="[936,1084,1677,1703]" italics="true" pageId="10" pageNumber="68">Diphyomene</emphasis>
Ruhland
</taxonomicName>
, one of the few categories of 
<taxonomicName id="64934D11FFBAFFBFEB74F9215423F93E" box="[353,514,1712,1738]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="10" pageNumber="68" phylum="Tracheophyta" rank="genus">
<emphasis id="91E7EA80FFBAFFBFEB74F9215423F93E" box="[353,514,1712,1738]" italics="true" pageId="10" pageNumber="68">Paepalanthus</emphasis>
</taxonomicName>
mostly diversified in the Chapada dos Veadeiros Region (
<bibRefCitation id="C7024B63FFBAFFBFEEDCF92156D9F91A" author="Trovo M. &amp; Sano P. T." pageId="10" pageNumber="68" pagination="49 - 55" refId="ref14645" refString="Trovo M. &amp; Sano P. T. 2010. Taxonomic survey of Paepalanthus section Diphyomene (Eriocaulaceae). Phytotaxa 14: 49 - 55. https: // doi. org / 10.11646 / phytotaxa. 14.1.4" type="journal article" year="2010">Trovó &amp; Sano 2010</bibRefCitation>
; 
<bibRefCitation id="C7024B63FFBAFFBFEB1DF94557FFF91A" author="Trovo M. &amp; Echternacht L. A. &amp; Sano P. T." box="[264,478,1748,1774]" pageId="10" pageNumber="68" pagination="195 - 206" refId="ref14685" refString="Trovo M., Echternacht L. A. &amp; Sano P. T. 2013. Distribution and conservation of Paepalanthus Mart. sect. Diphyomene Ruhland (Eriocaulaceae) in Neotropical savannas. Adansonia 35: 195 - 206. https: // doi. org / 10.5252 / a 2013 n 2 a 2" type="journal article" year="2013">
Trovó 
<emphasis id="91E7EA80FFBAFFBFEB4FF94457B7F91A" box="[346,406,1748,1774]" italics="true" pageId="10" pageNumber="68">et al.</emphasis>
2013
</bibRefCitation>
). The species is morphologically most similar to 
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<emphasis id="91E7EA80FFBAFFBFEE3BF9445321F91A" box="[1070,1280,1748,1774]" italics="true" pageId="10" pageNumber="68">P. acanthophyllus</emphasis>
</taxonomicName>
due to its general habit, deciduous leaves, patent and spiny axis bracts, and the shape and color of the capitula and involucral bracts. 
<taxonomicName id="64934D11FFBAFFBFEB99F88A54A0F8C1" authorityName="Trovó &amp; Echternacht" authorityYear="2022" box="[396,641,1819,1845]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="10" pageNumber="68" phylum="Tracheophyta" rank="species" species="irwinii">
<emphasis id="91E7EA80FFBAFFBFEB99F88A54A0F8C1" box="[396,641,1819,1845]" italics="true" pageId="10" pageNumber="68">Paepalanthus irwinii</emphasis>
</taxonomicName>
is distinguished by being a more gracile plant up to 
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tall (vs up to 
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); with shorter and narrower, linear leaves, 1.0−3.0 × 
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(vs lanceolate, 3.5−8.0 × 
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); shorter and narrower reproductive axis, ca 45.0 cm tall and 
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diam. (vs longer and wider, ca 100.0 cm tall and 
<quantity id="646B9B77FFBAFFBFEBE2F8165467F855" box="[503,582,1927,1953]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="68" unit="cm" value="0.6">0.6 cm</quantity>
diam.); linear to narrowly lanceolate axis bracts, 0.5−1.5 × 
<quantity id="646B9B77FFBAFFBFEEE6F8165356F855" box="[1267,1399,1927,1953]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="10" pageNumber="68" unit="cm" value="0.2" valueMax="0.3" valueMin="0.1">0.1−0.3 cm</quantity>
(vs lanceolate, 1.0−5.5 × 
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); up to 25 scapes per plant (vs up to 80 scapes per plant); narrower capitula, ca 
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diam. (vs wider, ca 
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diam.); and the petals of the pistillate flower with obtuse to mucronate apex (vs petals of the pistillate flower with obtuse to rounded apex). Although sympatric, these species grow in different habitats and flower at different times. While 
<taxonomicName id="64934D11FFBBFFBEEE7BFEBC52FCFEB2" authorityName="Trovó &amp; Echternacht" authorityYear="2022" box="[1134,1245,301,326]" class="Liliopsida" family="Eriocaulaceae" genus="Paepalanthus" kingdom="Plantae" order="Poales" pageId="11" pageNumber="69" phylum="Tracheophyta" rank="species" species="irwinii">
<emphasis id="91E7EA80FFBBFFBEEE7BFEBC52FCFEB2" box="[1134,1245,301,326]" italics="true" pageId="11" pageNumber="69">P. irwinii</emphasis>
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occurs over shallow, rocky soils, 
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<emphasis id="91E7EA80FFBBFFBEEBA4FEC154A1FE9D" box="[433,640,335,361]" italics="true" pageId="11" pageNumber="69">P. acanthophyllus</emphasis>
</taxonomicName>
occurs over a more consolidated argillaceous soil. Individuals of the new species are usually seen in flower and fruit from November to May, while 
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<emphasis id="91E7EA80FFBBFFBEEE9EFEE2537AFE78" box="[1163,1371,370,396]" italics="true" pageId="11" pageNumber="69">P. acanthophyllus</emphasis>
</taxonomicName>
is usually fertile from April to August.
</paragraph>
</subSubSection>
</treatment>
</document>