treatments-xml/data/D1/53/87/D15387F9FF90FF849CBB37ACFBDD094E.xml
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<document id="6612D39C1A975BC31BABBC649EAAF659" ID-CLB-Dataset="29265" ID-DOI="10.11646/zootaxa.4496.1.19" ID-GBIF-Dataset="1cf4a9f3-3009-4abe-a993-5bacf7508661" ID-ISSN="1175-5326" ID-Zenodo-Dep="1446783" ID-ZooBank="2BA480FD-2657-4740-9C71-4469FB27E3C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538747212016" checkinUser="plazi" docAuthor="Hong, Yong, Inkhavilay, Khamla &amp; James, Samuel W." docDate="2018" docId="D15387F9FF90FF849CBB37ACFBDD094E" docLanguage="en" docName="zootaxa.4496.1.19.pdf" docOrigin="Zootaxa 4496 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amynthas antethecus Hong &amp; Inkhavilay &amp; James 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="249" masterDocId="2D6AFF81FF99FF8F9C2C314DFF8E0C43" masterDocTitle="Nine new species of the &quot; aeruginosus &quot; - group in Amynthas (Clitellata: Megascolecidae) from Nam Et-Phouley National protected Area, Laos" masterLastPageNumber="250" masterPageNumber="238" pageNumber="247" updateTime="1698565735469" updateUser="ExternalLinkService">
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<mods:title id="10B2419BF33B7C6723ECF5DB0037C537">Nine new species of the &quot; aeruginosus &quot; - group in Amynthas (Clitellata: Megascolecidae) from Nam Et-Phouley National protected Area, Laos</mods:title>
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<mods:name id="297BB5B047E97F0B82178D59CCC807A0" type="personal">
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<mods:namePart id="0A3DB54012D2FEF00882859BBD5FA03F">Hong, Yong</mods:namePart>
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<mods:namePart id="9BA7D4C0EA123AC62C51A7631E68D3FF">Inkhavilay, Khamla</mods:namePart>
</mods:name>
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<mods:role id="7C535C8FBAAE147EDA02299292685C7D">
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<mods:namePart id="7543EC8E179479D3C8B56ABF4257BB66">James, Samuel W.</mods:namePart>
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<mods:title id="720DDB18DA9625CDD61C64A9CFAD4B2E">Zootaxa</mods:title>
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<mods:part id="293FCB78E874B3F5CD0035F7D7D1BE53">
<mods:date id="B76DDF3C101D1F2EFAA024666D98FFFE">2018</mods:date>
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<mods:number id="D297626960F297A9CB8E394DE8022673">2018-10-05</mods:number>
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<mods:number id="7E497B4DB9D646D23C3D5BF968631F38">4496</mods:number>
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<mods:identifier id="B5323605C5546C7BDFAE23737C0D9ABD" type="DOI">10.11646/zootaxa.4496.1.19</mods:identifier>
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<treatment id="D15387F9FF90FF849CBB37ACFBDD094E" ID-DOI="http://doi.org/10.5281/zenodo.5989261" ID-GBIF-Taxon="148639834" ID-Zenodo-Dep="5989261" LSID="urn:lsid:plazi:treatment:D15387F9FF90FF849CBB37ACFBDD094E" httpUri="http://treatment.plazi.org/id/D15387F9FF90FF849CBB37ACFBDD094E" lastPageId="11" lastPageNumber="249" pageId="9" pageNumber="247">
<subSubSection id="11E06564FF90FF869CBB37ACFEB80B5F" pageId="9" pageNumber="247" type="nomenclature">
<paragraph id="594536EFFF90FF869CBB37ACFE710AB8" blockId="9.[151,511,1761,1821]" box="[151,511,1761,1787]" pageId="9" pageNumber="247">
<heading id="020D8183FF90FF869CBB37ACFE710AB8" bold="true" box="[151,511,1761,1787]" fontSize="11" level="1" pageId="9" pageNumber="247" reason="1">
<emphasis id="6B8EEAFDFF90FF869CBB37ACFE710AB8" bold="true" box="[151,511,1761,1787]" pageId="9" pageNumber="247">
<taxonomicName id="9EFA4D6CFF90FF869CBB37ACFE140AB8" authority="Hong &amp; Inkhavilay &amp; James, 2018" authorityName="Hong &amp; Inkhavilay &amp; James" authorityYear="2018" box="[151,410,1761,1787]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="247" phylum="Annelida" rank="species" species="antethecus" status="sp. nov.">
<emphasis id="6B8EEAFDFF90FF869CBB37ACFE140AB8" bold="true" box="[151,410,1761,1787]" italics="true" pageId="9" pageNumber="247">Amynthas antethecus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70BD5786FF90FF869D8E37ACFE710AB8" box="[418,511,1761,1787]" pageId="9" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="594536EFFF90FF869CBB3649FEB80B5F" blockId="9.[151,511,1761,1821]" box="[151,310,1796,1821]" pageId="9" pageNumber="247">
(
<figureCitation id="C1C12A6AFF90FF869CB23649FEA20B5F" box="[158,300,1796,1821]" captionStart="FIGURE 3" captionStartId="10.[151,250,1969,1991]" captionTargetBox="[195,1404,209,1924]" captionTargetId="figure@10.[167,1419,205,1936]" captionTargetPageId="10" captionText="FIGURE 3. Amynthas fleischmani sp. nov. (A,B), Amynthas antethecus sp. nov. (C,D), and Amynthas elenabondae sp. nov. (E,F). A,C,E: ventral view; B,D,F: spermatheca. Scale bars 2 mm." httpUri="https://zenodo.org/record/1446789/files/figure.png" pageId="9" pageNumber="247">Figure 3C,D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="11E06564FF90FF869CBB360BFDB80BB1" pageId="9" pageNumber="247" type="materials_examined">
<paragraph id="594536EFFF90FF869CBB360BFDB80BB1" blockId="9.[151,1437,1862,2034]" pageId="9" pageNumber="247">
<materialsCitation id="E9923CB2FF90FF869CBB360BFF410B8D" ID-GBIF-Occurrence="1914265327" collectingDate="2005-07-21" collectionCode="NPA" collectorName="Phimphet &amp; Xay &amp; Ammaline &amp; Phoang &amp; K. Inkhavilay" country="Laos" county="Thad Ang" elevation="792" location="Hoaphane Province" municipality="Viengthong district" pageId="9" pageNumber="247" specimenCode="BDNUL0050" specimenCount="1" typeStatus="holotype">
<emphasis id="6B8EEAFDFF90FF869CBB360BFE880B1C" bold="true" box="[151,262,1862,1887]" pageId="9" pageNumber="247">Material.</emphasis>
<typeStatus id="8641884DFF90FF869D3F360BFE0E0B1C" box="[275,384,1862,1887]" pageId="9" pageNumber="247" type="holotype">Holotype</typeStatus>
: clitellate (
<specimenCode id="095C9E94FF90FF869E2A360BFD290B1C" box="[518,679,1862,1887]" collectionCode="BDNUL" pageId="9" pageNumber="247">BDNUL0050</specimenCode>
):
<collectingCountry id="21ED767FFF90FF869E93360AFD760B1D" box="[703,760,1863,1886]" name="Laos" pageId="9" pageNumber="247">Laos</collectingCountry>
,
<location id="5C256034FF90FF869F26360BFC600B1C" LSID="urn:lsid:plazi:treatment:D15387F9FF90FF849CBB37ACFBDD094E:5C256034FF90FF869F26360BFC600B1C" box="[778,1006,1862,1887]" country="Laos" county="Thad Ang" municipality="Viengthong district" name="Hoaphane Province" pageId="9" pageNumber="247">Hoaphane Province</location>
,
<location id="5C256034FF90FF86982C360AFB4C0B1C" LSID="urn:lsid:plazi:treatment:D15387F9FF90FF849CBB37ACFBDD094E:5C256034FF90FF86982C360AFB4C0B1C" box="[1024,1218,1862,1887]" country="Laos" county="Thad Ang" municipality="Viengthong district" name="Nam Et-Phouley" pageId="9" pageNumber="247">Nam Et-Phouley</location>
<collectionCode id="3FEBAE2AFF90FF8698FC360AFA860B1C" box="[1232,1288,1863,1887]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13690" name="Nanjing Institute of Geology and Paleontology, Academia Sinica" pageId="9" pageNumber="247">NPA</collectionCode>
,
<collectingMunicipality id="B921AC95FF90FF869935360BFF660BC5" pageId="9" pageNumber="247">Viengthong district</collectingMunicipality>
,
<collectingCounty id="B0244E63FF90FF869CD83620FEE80BC5" box="[244,358,1901,1926]" pageId="9" pageNumber="247">Thad Ang</collectingCounty>
waterfall area,
<elevation id="D2D7D1DCFF90FF869E363623FDEC0BC6" box="[538,610,1902,1926]" metricMagnitude="2" metricUnit="m" metricValue="7.92" pageId="9" pageNumber="247" unit="m" value="792.0">
<quantity id="9E029B0AFF90FF869E363623FDEC0BC6" box="[538,610,1902,1926]" metricMagnitude="2" metricUnit="m" metricValue="7.92" pageId="9" pageNumber="247" unit="m" value="792.0">792 m</quantity>
</elevation>
(20o 15. 446' N, 103o 08 921' E), good condition mossy forest with big trees, dark soil and thick litter layer,
<collectingDate id="3D00E9C7FF90FF869E6836DFFD520BE9" box="[580,732,1937,1962]" pageId="9" pageNumber="247" value="2005-07-21">
<date id="2D44102FFF90FF869E6836DFFD520BE9" box="[580,732,1937,1962]" pageId="9" pageNumber="247" value="2005-07-21">21 July 2005</date>
</collectingDate>
,
<collectorName id="F40F5339FF90FF869EC736DCFCD70BE9" box="[747,857,1937,1962]" pageId="9" pageNumber="247">Phimphet</collectorName>
,
<collectorName id="F40F5339FF90FF869F4B36DFFC190BE9" box="[871,919,1938,1962]" pageId="9" pageNumber="247">Xay</collectorName>
,
<collectorName id="F40F5339FF90FF869F8936DCFB900BE9" box="[933,1054,1937,1962]" pageId="9" pageNumber="247">Ammaline</collectorName>
,
<collectorName id="F40F5339FF90FF86980736DCFBF10BE9" box="[1067,1151,1937,1962]" pageId="9" pageNumber="247">Phoang</collectorName>
sone, and
<collectorName id="F40F5339FF90FF86992C36DFFA130BE9" box="[1280,1437,1937,1962]" pageId="9" pageNumber="247">K. Inkhavilay</collectorName>
colls
</materialsCitation>
.
<materialsCitation id="E9923CB2FF90FF869CCE36FBFB230B8D" ID-GBIF-Occurrence="1914265331" box="[226,1197,1973,1998]" collectingDate="2005-07-21" collectionCode="BDNUL" collectorName="Phimphet &amp; Xay &amp; Ammaline &amp; Phoang &amp; K. Inkhavilay" country="Laos" county="Thad Ang" elevation="792" location="Hoaphane Province" municipality="Viengthong district" pageId="9" pageNumber="247" specimenCode="BDNUL0051, BDNUL0052" specimenCount="2" typeStatus="paratype">
<typeStatus id="8641884DFF90FF869CCE36FBFEDD0B8D" box="[226,339,1974,1998]" pageId="9" pageNumber="247" type="paratype">Paratypes</typeStatus>
: 2 clitellates (
<specimenCode id="095C9E94FF90FF869E2836F8FD2F0B8D" box="[516,673,1973,1998]" collectionCode="BDNUL" pageId="9" pageNumber="247">BDNUL0051</specimenCode>
,
<specimenCode id="095C9E94FF90FF869E9836F8FCDF0B8D" box="[692,849,1973,1998]" collectionCode="BDNUL" pageId="9" pageNumber="247">BDNUL0052</specimenCode>
), same data as for
<typeStatus id="8641884DFF90FF86986B36F8FB230B8D" box="[1095,1197,1973,1998]" pageId="9" pageNumber="247" type="holotype">holotype</typeStatus>
</materialsCitation>
.
<materialsCitation id="E9923CB2FF90FF86989336F8FDBC0BB1" ID-GBIF-Occurrence="1914265337" collectingDate="2005-07-21" collectionCode="BDNUL" collectorName="Phimphet &amp; Xay &amp; Ammaline &amp; Phoang &amp; K. Inkhavilay" country="Laos" county="Thad Ang" elevation="792" location="Hoaphane Province" municipality="Viengthong district" pageId="9" pageNumber="247" specimenCount="1" typeStatus="holotype">
Other material: 25 clitellates, same data as for
<typeStatus id="8641884DFF90FF869DE03694FDBC0BB1" box="[460,562,2009,2034]" pageId="9" pageNumber="247" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="0D856667FF93FF859CBB36FCFCB60BA5" httpUri="https://zenodo.org/record/1446789/files/figure.png" pageId="10" pageNumber="248" startId="10.[151,250,1969,1991]" targetBox="[195,1404,209,1924]" targetPageId="10">
<paragraph id="594536EFFF93FF859CBB36FCFCB60BA5" blockId="10.[151,1435,1969,2022]" pageId="10" pageNumber="248">
<emphasis id="6B8EEAFDFF93FF859CBB36FCFE9A0B85" bold="true" box="[151,276,1969,1991]" pageId="10" pageNumber="248">FIGURE 3.</emphasis>
<taxonomicName id="9EFA4D6CFF93FF859D3036FCFE720B84" authority="Hong &amp; Inkhavilay &amp; James, 2018" authorityName="Hong &amp; Inkhavilay &amp; James" authorityYear="2018" box="[284,508,1969,1991]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="248" phylum="Annelida" rank="species" species="fleischmani" status="sp. nov.">
<emphasis id="6B8EEAFDFF93FF859D3036FCFE720B84" box="[284,508,1969,1991]" italics="true" pageId="10" pageNumber="248">Amynthas fleischmani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70BD5786FF93FF859E2836FCFDDB0B85" box="[516,597,1969,1990]" pageId="10" pageNumber="248" rank="species">
<emphasis id="6B8EEAFDFF93FF859E2836FCFDDB0B85" bold="true" box="[516,597,1969,1990]" pageId="10" pageNumber="248">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<emphasis id="6B8EEAFDFF93FF859E4A36FCFDF60B85" bold="true" box="[614,632,1969,1990]" pageId="10" pageNumber="248">A</emphasis>
,
<emphasis id="6B8EEAFDFF93FF859E5236FCFD010B85" bold="true" box="[638,655,1969,1990]" pageId="10" pageNumber="248">B</emphasis>
),
<taxonomicName id="9EFA4D6CFF93FF859E8A36FCFCF30B84" authority="Hong &amp; Inkhavilay &amp; James, 2018" authorityName="Hong &amp; Inkhavilay &amp; James" authorityYear="2018" box="[678,893,1969,1991]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="248" phylum="Annelida" rank="species" species="antethecus" status="sp. nov.">
<emphasis id="6B8EEAFDFF93FF859E8A36FCFCF30B84" box="[678,893,1969,1991]" italics="true" pageId="10" pageNumber="248">Amynthas antethecus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70BD5786FF93FF859FA936FCFC590B85" box="[901,983,1969,1990]" pageId="10" pageNumber="248" rank="species">
<emphasis id="6B8EEAFDFF93FF859FA936FCFC590B85" bold="true" box="[901,983,1969,1990]" pageId="10" pageNumber="248">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<emphasis id="6B8EEAFDFF93FF859FCB36FCFC770B84" bold="true" box="[999,1017,1969,1991]" pageId="10" pageNumber="248">C</emphasis>
,
<emphasis id="6B8EEAFDFF93FF85982C36FCFB9C0B85" bold="true" box="[1024,1042,1969,1990]" pageId="10" pageNumber="248">D</emphasis>
), and
<taxonomicName id="9EFA4D6CFF93FF85987936FCFACC0B84" authority="Hong &amp; Inkhavilay &amp; James, 2018" authorityName="Hong &amp; Inkhavilay &amp; James" authorityYear="2018" box="[1109,1346,1969,1991]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="10" pageNumber="248" phylum="Annelida" rank="species" species="elenabondae" status="sp. nov.">
<emphasis id="6B8EEAFDFF93FF85987936FCFACC0B84" box="[1109,1346,1969,1991]" italics="true" pageId="10" pageNumber="248">Amynthas elenabondae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70BD5786FF93FF85996636FCFA150B85" box="[1354,1435,1969,1990]" pageId="10" pageNumber="248" rank="species">
<emphasis id="6B8EEAFDFF93FF85996636FCFA150B85" bold="true" box="[1354,1435,1969,1990]" pageId="10" pageNumber="248">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<emphasis id="6B8EEAFDFF93FF859CB3369DFF3E0BA6" bold="true" box="[159,176,2000,2021]" pageId="10" pageNumber="248">E</emphasis>
,
<emphasis id="6B8EEAFDFF93FF859C9A369DFF4B0BA6" bold="true" box="[182,197,2000,2021]" pageId="10" pageNumber="248">F</emphasis>
). A,C,E: ventral view; B,D,F: spermatheca. Scale bars 2 mm.
</paragraph>
</caption>
<subSubSection id="11E06564FF92FF849CEB31D5FBCA0C97" pageId="11" pageNumber="249" type="etymology">
<paragraph id="594536EFFF92FF849CEB31D5FBCA0C97" blockId="11.[151,1437,151,1293]" pageId="11" pageNumber="249">
<emphasis id="6B8EEAFDFF92FF849CEB31D5FEC00CF3" bold="true" box="[199,334,152,176]" pageId="11" pageNumber="249">Etymology.</emphasis>
The species is named for the anteriorly shifted spermathecae, being in segments VII and VIII rather than VIII and IX as is usual for earthworms with spermathecal pores in 7/8/9.
</paragraph>
</subSubSection>
<subSubSection id="11E06564FF92FF849CEB3192FAF80DCB" pageId="11" pageNumber="249" type="diagnosis">
<paragraph id="594536EFFF92FF849CEB3192FAF80DCB" blockId="11.[151,1437,151,1293]" pageId="11" pageNumber="249">
<emphasis id="6B8EEAFDFF92FF849CEB3192FECF0CBB" bold="true" box="[199,321,223,248]" pageId="11" pageNumber="249">Diagnosis.</emphasis>
Secondary male pores near lateral margins of ventrum in XVIII at 10th setal lines, 0.31 circumference apart ventrally, sphincter-like; primary male pore on small penis within shallow parietal invagination; paired circular papillae within invaginations on 17/18, 18/19, medial to level of secondary male pores; spermathecal pores paired in 7/8/9 at 9th setal lines, pores large, 0.21 circumference apart ventrally; first dorsal pore in 5/6; very large fusiform muscular prostatic ducts in single loop bound together with tough fibers.
</paragraph>
</subSubSection>
<subSubSection id="11E06564FF92FF849CEB30DEFDBF081B" pageId="11" pageNumber="249" type="description">
<paragraph id="594536EFFF92FF849CEB30DEFEB10EB3" blockId="11.[151,1437,151,1293]" pageId="11" pageNumber="249">
<emphasis id="6B8EEAFDFF92FF849CEB30DEFED70DEF" bold="true" box="[199,345,403,428]" pageId="11" pageNumber="249">Description.</emphasis>
Dimensions
<quantity id="9E029B0AFF92FF849DDD30D9FDFA0DEF" box="[497,628,404,428]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.22" metricValueMin="0.88" pageId="11" pageNumber="249" unit="mm" value="105.0" valueMax="122.0" valueMin="88.0">88122mm</quantity>
, by 4.0
<quantity id="9E029B0AFF92FF849EF530D9FCBE0DEF" box="[729,816,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="11" pageNumber="249" unit="mm" value="4.1">4.1 mm</quantity>
at segment X,
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at segment XXX, 3.0
<quantity id="9E029B0AFF92FF84995630D9FF4D0D8C" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="11" pageNumber="249" unit="mm" value="3.8">3.8 mm</quantity>
at clitellum; body cylindrical, segments 82109. Setae slightly more crowded ventrally, numbering 5963 at VII, 5661 at XX, 1214 between male pores, setal formula AA:AB:ZZ:YZ= 1:1:3:2.5 at XIII. Female pore single in XIV, on a circular porophore
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in diameter. Prostomium epilobic with tongue open, clitellum light brown, anterior dorsal brown pigment, formalin preservation. First dorsal pore at 5/6. Clitellum annular XIVXVI, setae invisible externally. Secondary male pores near lateral margins of ventrum in XVIII at 10th setal lines, 0.31 circumference apart ventrally, opening surrounded by tubercular wrinkled epidermal ring; primary male pores on small cylindrical penis within shallow parietal invaginations. Paired circular papillae on 17/18, 18/19, medial to level of secondary male pores. Spermathecal pores paired in 7/8/9 at 9th setal lines, pores large, 0.21 circumference apart ventrally.
</paragraph>
<paragraph id="594536EFFF92FF849CEB33B1FDBF081B" blockId="11.[151,1437,151,1293]" pageId="11" pageNumber="249">
Septa 5/6/7/8 thinly muscular,
<date id="2D44102FFF92FF849E0333B1FDF90F57" box="[559,631,764,788]" pageId="11" pageNumber="249" value="2010-09-08">8/9/10</date>
absent, 10/11 thin,
<date id="2D44102FFF92FF849F4833B1FC4B0F57" box="[868,965,764,788]" pageId="11" pageNumber="249" value="2013-12-11">11/12/13</date>
/14 thinly muscular. Gizzard globular in VIIIX. Intestine enlarged from XV, lymph glands not found. Typhlosole vestigial from XXVII. Intestinal caeca originating in XXVII extending anteriorly about to XXIII, simple elongate triangle. Hearts esophageal XXIII, X smaller. Male sexual system holandric, testes and funnels in ventrally joined sacs in X, XI. Seminal vesicles paired, well-developed in XI, XII; dorsal lobes lacking. Prostates in XVIII, extending from XVIIXX; stout fusiform muscular ducts bent in one loop tightly bound together by tough fibers; glandular portions consisted of 1014 small digitate lobes. Dense sessile genital papilla glands corresponding to papillae of 17/18, 18/19. Ovaries in XIII. Spermathecae paired in VII, VIII; ampulla irregular, bent down and attached to side of duct, muscular ducts shorter than ampulla, iridescent seminal chamber ovate, with slender stalk shorter than ampulla; stalk joined duct near ampulla base, no nephridia on ducts.
</paragraph>
</subSubSection>
<subSubSection id="11E06564FF92FF849CEB3529FBDD094E" pageId="11" pageNumber="249" type="discussion">
<paragraph id="594536EFFF92FF849CEB3529FBDD094E" blockId="11.[151,1437,151,1293]" pageId="11" pageNumber="249">
<emphasis id="6B8EEAFDFF92FF849CEB3529FEB5083F" bold="true" box="[199,315,1124,1148]" pageId="11" pageNumber="249">Remarks.</emphasis>
This species is in the
<taxonomicName id="9EFA4D6CFF92FF849E143528FD4F083F" authorityName="Kinberg" authorityYear="1867" box="[568,705,1125,1148]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="11" pageNumber="249" phylum="Annelida" rank="species" species="aeruginosus">
<emphasis id="6B8EEAFDFF92FF849E143528FD4F083F" box="[568,705,1125,1148]" italics="true" pageId="11" pageNumber="249">aeruginosus</emphasis>
</taxonomicName>
-group Laotian section, generally characterized by long prostatic ducts often in mirror-image coils, first dorsal pores in 5/6 or nearby, and lack of pre-clitellar genital markings.
<taxonomicName id="9EFA4D6CFF92FF849CBB35E1FE060887" authority="Hong &amp; Inkhavilay &amp; James, 2018" authorityName="Hong &amp; Inkhavilay &amp; James" authorityYear="2018" box="[151,392,1196,1220]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="11" pageNumber="249" phylum="Annelida" rank="species" species="antethecus" status="sp. nov.">
<emphasis id="6B8EEAFDFF92FF849CBB35E1FE060887" box="[151,392,1196,1220]" italics="true" pageId="11" pageNumber="249">Amynthas antethecus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70BD5786FF92FF849DBE35E1FE600887" box="[402,494,1196,1220]" pageId="11" pageNumber="249" rank="species">
<emphasis id="6B8EEAFDFF92FF849DBE35E1FE600887" bold="true" box="[402,494,1196,1220]" pageId="11" pageNumber="249">sp. nov.</emphasis>
</taxonomicNameLabel>
has been compared to
<taxonomicName id="9EFA4D6CFF92FF849F2C35E1FC740887" authority="Hong &amp; Inkhavilay &amp; James, 2018" authorityName="Hong &amp; Inkhavilay &amp; James" authorityYear="2018" box="[768,1018,1196,1220]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="11" pageNumber="249" phylum="Annelida" rank="species" species="fleischmani" status="sp. nov.">
<emphasis id="6B8EEAFDFF92FF849F2C35E1FC740887" box="[768,1018,1196,1220]" italics="true" pageId="11" pageNumber="249">Amynthas fleischmani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70BD5786FF92FF84982835E1FBEC0887" box="[1028,1122,1196,1220]" pageId="11" pageNumber="249" rank="species">
<emphasis id="6B8EEAFDFF92FF84982835E1FBEC0887" bold="true" box="[1028,1122,1196,1220]" pageId="11" pageNumber="249">sp. nov.</emphasis>
</taxonomicNameLabel>
under the remarks for that species. The two are closely sympatric, found on the same site, with no intermediate forms. Our placement of it in
<taxonomicName id="9EFA4D6CFF92FF849CBB35B9FE88094F" authorityName="Hong &amp; Inkhavilay &amp; James" authorityYear="2018" box="[151,262,1268,1292]" class="Clitellata" family="Megascolecidae" genus="Amynthas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="11" pageNumber="249" phylum="Annelida" rank="genus">
<emphasis id="6B8EEAFDFF92FF849CBB35B9FE88094F" box="[151,262,1268,1292]" italics="true" pageId="11" pageNumber="249">Amynthas</emphasis>
</taxonomicName>
rather than in
<taxonomicName id="9EFA4D6CFF92FF849D8435B9FD92094F" authorityName="Sims &amp; Easton" authorityYear="1972" box="[424,540,1268,1292]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="11" pageNumber="249" phylum="Annelida" rank="genus">
<emphasis id="6B8EEAFDFF92FF849D8435B9FD92094F" box="[424,540,1268,1292]" italics="true" pageId="11" pageNumber="249">Metaphire</emphasis>
</taxonomicName>
is justified in the remarks on the previous species.
</paragraph>
</subSubSection>
</treatment>
</document>