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<mods:title id="57C4E8AFADF3639FCF8B0D9934C5656E">Revision of the genus Devadatta Kirby, 1890 in Borneo based on molecular and morphological methods, with descriptions of four new species (Odonata: Zygoptera: Devadattidae)</mods:title>
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<taxonomicName id="81A24D6B553AFF98FF7594C1CD61FF48" ID-CoL="6CPLH" authority="Laidlaw, 1934" authorityName="Laidlaw" authorityYear="1934" box="[151,623,151,178]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="podolestoides">
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<emphasis id="74D6EAFA553AFF98FF7594C1CEB7FF4B" bold="true" box="[151,441,151,177]" italics="true" pageId="32" pageNumber="333">Devadatta podolestoides</emphasis>
Laidlaw, 1934
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(
<figureCitation id="DE992A6D553AFF98FF7D94ECCFE2FF29" box="[159,236,186,211]" captionStart="FIGURE 1" captionStartId="7.[151,250,1912,1934]" captionTargetBox="[160,1436,198,1884]" captionTargetId="figure@7.[151,1436,193,1890]" captionTargetPageId="7" captionText="FIGURE 1. COI gene tree for 73 specimens of Devadatta and one outgroup taxon, from Bayesian Inference analysis. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253945/files/figure.png" pageId="32" pageNumber="333">Figs. 1</figureCitation>
,
<figureCitation id="DE992A6D553AFF98FF1894EDCE04FF29" box="[250,266,187,211]" captionStart="FIGURE 2" captionStartId="8.[151,250,1813,1835]" captionTargetBox="[151,1436,198,1786]" captionTargetId="figure@8.[151,1436,193,1792]" captionTargetPageId="8" captionText="FIGURE 2. COI gene tree for the 33 specimens of Devadatta also used in the analysis with other markers and one outgroup taxon. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253946/files/figure.png" pageId="32" pageNumber="333">2</figureCitation>
,
<figureCitation id="DE992A6D553AFF98FEF494EDCE2BFF29" box="[278,293,187,211]" captionStart="FIGURE 3. 16 S" captionStartId="9.[151,250,1748,1770]" captionTargetBox="[159,1436,201,1719]" captionTargetId="figure@9.[151,1436,193,1727]" captionTargetPageId="9" captionText="FIGURE 3. 16 S gene tree for 33 specimens of Devadatta and one outgroup taxon, from Bayesian Inference analysis. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253947/files/figure.png" pageId="32" pageNumber="333">3</figureCitation>
,
<figureCitation id="DE992A6D553AFF98FED094EDCE4CFF29" box="[306,322,187,211]" captionStart="FIGURE 4" captionStartId="10.[151,250,1769,1791]" captionTargetBox="[151,1436,198,1741]" captionTargetId="figure@10.[151,1436,193,1748]" captionTargetPageId="10" captionText="FIGURE 4. ITS gene tree for 33 specimens of Devadatta and one outgroup taxon, from Bayesian Inference analysis. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253949/files/figure.png" pageId="32" pageNumber="333">4</figureCitation>
,
<figureCitation id="DE992A6D553AFF98FEAC94ECCE53FF29" box="[334,349,186,211]" captionStart="FIGURE 5. 28 S" captionStartId="11.[151,250,1830,1852]" captionTargetBox="[151,1436,200,1807]" captionTargetId="figure@11.[151,1436,193,1808]" captionTargetPageId="11" captionText="FIGURE 5. 28 S gene tree for 33 specimens of Devadatta and one outgroup taxon, from Bayesian Inference analysis. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253950/files/figure.png" pageId="32" pageNumber="333">5</figureCitation>
,
<figureCitation id="DE992A6D553AFF98FE8894ECCE74FF29" box="[362,378,186,211]" captionStart="FIGURE 6" captionStartId="12.[151,250,1843,1865]" captionTargetBox="[157,1436,202,1816]" captionTargetId="figure@12.[151,1436,193,1821]" captionTargetPageId="12" captionText="FIGURE 6. Phylogenetic reconstruction for 33 specimens of Devadatta and one outgroup taxon from the combined COI + 16 S + ITS + 28 S data, using Bayesian Inference analysis. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253951/files/figure.png" pageId="32" pageNumber="333">6</figureCitation>
,
<figureCitation id="DE992A6D553AFF98FE6494EDCE98FF28" box="[390,406,187,210]" captionStart="FIGURE 7" captionStartId="13.[151,250,1743,1765]" captionTargetBox="[153,1436,202,1715]" captionTargetId="figure@13.[151,1436,193,1721]" captionTargetPageId="13" captionText="FIGURE 7. Maximum Likelihood best tree for 33 specimens of Devadatta and one outgroup taxon from the combined COI + 16 S + ITS + 28 S data set. Bootstrap support values below 100 are superimposed on the tree. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253952/files/figure.png" pageId="32" pageNumber="333">7</figureCitation>
,
<figureCitation id="DE992A6D553AFF98FE4094EDCEB1FF29" box="[418,447,187,211]" captionStart="FIGURES 8 13" captionStartId="15.[151,264,1607,1629]" captionTargetBox="[151,1436,193,1585]" captionTargetId="figure@15.[151,1436,193,1586]" captionTargetPageId="15" captionText="FIGURES 8 13. Habitus of males of Devadatta species: (8) D. aran holotype; (9) D. basilanensis Davao City, Mindanao; (10) D. clavicauda holotype; (11) D. podolestoides male described in this paper; (12) D. somoh holotype; (13) D. tanduk holotype." httpUri="https://zenodo.org/record/253953/files/figure.png" pageId="32" pageNumber="333">11</figureCitation>
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<figureCitation id="DE992A6D553AFF98FE2994EDCEE7FF29" box="[459,489,187,211]" captionStart="FIGURES 14 19" captionStartId="16.[151,264,1803,1825]" captionTargetBox="[151,1436,733,1781]" captionTargetId="figure@16.[151,1436,733,1782]" captionTargetPageId="16" captionText="FIGURES 14 19. Heads of male Devadatta species: (14) D. aran holotype; (15) D. basilanensis Davao Oriental, Mindanao; (16) D. clavicauda holotype; (17) D. podolestoides male described in this paper; (18) D. somoh holotype; (19) D. tanduk holotype." httpUri="https://zenodo.org/record/253954/files/figure.png" pageId="32" pageNumber="333">17</figureCitation>
,
<figureCitation id="DE992A6D553AFF98FE1794EDCD1DFF29" box="[501,531,187,211]" captionStart="FIGURES 20 25" captionStartId="17.[151,264,1919,1941]" captionTargetBox="[194,1386,193,1882]" captionTargetId="figure@17.[194,1387,193,1884]" captionTargetPageId="17" captionText="FIGURES 20 25. Synthorax of male Devadatta species, lateral view: (20) D. aran holotype; (21) D. basilanensis Davao Oriental, Mindanao; (22) D. clavicauda holotype; (23) D. podolestoides male described in this paper; (24) D. somoh holotype; (25) D. tanduk holotype." httpUri="https://zenodo.org/record/253955/files/figure.png" pageId="32" pageNumber="333">23</figureCitation>
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<figureCitation id="DE992A6D553AFF98FDFD94EDCD33FF29" box="[543,573,187,211]" captionStart="FIGURES 30 33" captionStartId="19.[151,264,1787,1809]" captionTargetBox="[151,1436,611,1753]" captionTargetId="figure@19.[151,1436,611,1754]" captionTargetPageId="19" captionText="FIGURES 30 33. Venter of synthorax of male Devadatta species: (30) D. basilanensis Davao Oriental, Mindanao; (31) D. podolestoides male described in this paper; (32) D. clavicauda holotype; (33) D. clavicauda paratype (SAR 09 _ 10 _ AMP 40, Gunung Serapi)." httpUri="https://zenodo.org/record/253957/files/figure.png" pageId="32" pageNumber="333">31</figureCitation>
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<figureCitation id="DE992A6D553AFF98FDAB94EDCD69FF29" box="[585,615,187,211]" captionStart="FIGURES 34 37" captionStartId="20.[151,264,1128,1150]" captionTargetBox="[151,1436,193,1106]" captionTargetId="figure@20.[151,1436,193,1107]" captionTargetPageId="20" captionText="FIGURES 34 37. Wings of some Devadatta species: (34) D. basilanensis male Zamboanga, Mindanao; (35) D. basilanensis female Cotabato, Mindanao; (36) D. clavicauda paratype, Gunung Santubong, Sarawak; (37) D. podolestoides paratype, Gunung Poteng, Kalimantan Barat." httpUri="https://zenodo.org/record/253958/files/figure.png" pageId="32" pageNumber="333">37</figureCitation>
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<figureCitation id="DE992A6D553AFF98FD9194EDCD9CFF29" box="[627,658,187,211]" captionStart="FIGURES 38 39" captionStartId="20.[151,264,1911,1933]" captionTargetBox="[151,1436,1261,1889]" captionTargetId="figure@20.[151,1436,1261,1889]" captionTargetPageId="20" captionText="FIGURES 38 39. Genital ligula of two Devadatta species, scale bars = 0.5 mm: (38) D. aran holotype; (39) D. podolestoides male described in this paper." httpUri="https://zenodo.org/record/253959/files/figure.png" pageId="32" pageNumber="333">39</figureCitation>
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<figureCitation id="DE992A6D553AFF98FD7F94EDCDB2FF29" box="[669,700,187,211]" captionStart="FIGURES 40 45" captionStartId="21.[151,264,1699,1721]" captionTargetBox="[162,1425,203,1663]" captionTargetId="figure@21.[151,1436,193,1677]" captionTargetPageId="21" captionText="FIGURES 40 45. Male anal appendages, dorsal view: (40) D. aran holotype; (41) D. basilanensis Davao Oriental, Mindanao; (42) D. clavicauda holotype; (43) D. podolestoides Kubah NP, Sarawak; (44) D. somoh holotype; (45) D. tanduk holotype." httpUri="https://zenodo.org/record/253960/files/figure.png" pageId="32" pageNumber="333">43</figureCitation>
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<figureCitation id="DE992A6D553AFF98FD2594EDCDEBFF29" box="[711,741,187,211]" captionStart="FIGURES 46 51" captionStartId="22.[151,264,1951,1973]" captionTargetBox="[162,1418,393,1916]" captionTargetId="figure@22.[151,1436,373,1929]" captionTargetPageId="22" captionText="FIGURES 46 51. Male anal appendages, lateral view (scale bars 1 mm): (46) D. aran holotype; (47) D. basilanensis Davao Oriental, Mindanao; (48) D. clavicauda holotype; (49) D. podolestoides Kubah NP, Sarawak; (50) D. somoh holotype; (51) D. tanduk holotype." httpUri="https://zenodo.org/record/253961/files/figure.png" pageId="32" pageNumber="333">49</figureCitation>
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<figureCitation id="DE992A6D553AFF98FD1394ECCC01FF29" box="[753,783,186,211]" captionStart="FIGURES 52 57" captionStartId="23.[151,264,1409,1431]" captionTargetBox="[151,1436,176,1394]" captionTargetId="figure@23.[151,1436,175,1395]" captionTargetPageId="23" captionText="FIGURES 52 57. Heads of Devadatta female specimens described in this paper: (52) D. aran; (53) D. basilanensis; (54) D. clavicauda; (55) D. podolestoides; (56) D. somoh; (57) D. tanduk." httpUri="https://zenodo.org/record/253962/files/figure.png" pageId="32" pageNumber="333">55</figureCitation>
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<figureCitation id="DE992A6D553AFF98FCF994ECCC37FF29" box="[795,825,186,211]" captionStart="FIGURES 58 63" captionStartId="25.[151,264,1932,1954]" captionTargetBox="[151,1436,411,1910]" captionTargetId="figure@25.[151,1436,411,1910]" captionTargetPageId="25" captionText="FIGURES 58 63. Prothorax of Devadatta female specimens described in this paper, dorsal view: (58) D. aran; (59) D. basilanensis; (60) D. clavicauda; (61) D. podolestoides; (62) D. somoh; (63) D. tanduk." httpUri="https://zenodo.org/record/253963/files/figure.png" pageId="32" pageNumber="333">61</figureCitation>
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<figureCitation id="DE992A6D553AFF98FCA794EDCC6DFF29" box="[837,867,187,211]" captionStart="FIGURES 68 73" captionStartId="28.[151,264,1928,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@28.[151,1436,193,1907]" captionTargetPageId="28" captionText="FIGURES 68 73. Synthorax of Devadatta female specimens described in this paper, lateral view: (68) D. aran; (69) D. basilanensis; (70) D. clavicauda; (71) D. podolestoides; (72) D. somoh; (73) D. tanduk." httpUri="https://zenodo.org/record/253965/files/figure.png" pageId="32" pageNumber="333">71</figureCitation>
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<emphasis id="74D6EAFA553AFF98FF759554CD26FEE2" box="[151,552,258,280]" italics="true" pageId="32" pageNumber="333">Devadatta podolestoides podolestoides</emphasis>
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(original description, part: specimens from Gunung Poteng);—
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</treatmentCitation>
(note on
<typeStatus id="9919884A553AFF98FC899577CCCAFECD" box="[875,964,289,311]" pageId="32" pageNumber="333" type="holotype">holotype</typeStatus>
and first described female); van
<treatmentCitation id="C70310F9553AFF98FAED9577CA92FECD" author="Tol" box="[1295,1436,289,311]" page="182" pageId="32" pageNumber="333" year="1992">
<bibRefCitation id="22334B19553AFF98FAED9577CA62FECD" author="Tol" box="[1295,1388,289,311]" pageId="32" pageNumber="333" refString="van Tol, J. (1992) An annotated index to names of Odonata used in publications by M. A. Lieftinck. Zoologische Verhandelingen, 279, 1 - 263." type="journal article" year="1992">Tol 1992</bibRefCitation>
: 182
</treatmentCitation>
(summary occurrences of name in publications by Lieftinck).
</treatmentCitationGroup>
</paragraph>
<paragraph id="461D36E8553AFF98FF759508CB46FE69" blockId="32.[151,1437,258,403]" pageId="32" pageNumber="333">
<treatmentCitationGroup id="66B211C6553AFF98FF759508CB46FE69" pageId="32" pageNumber="333">
<taxonomicName id="81A24D6B553AFF98FF759508CE9CFE8E" ID-CoL="6CPLH" box="[151,402,350,372]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="32" pageNumber="333" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA553AFF98FF759508CE9CFE8E" box="[151,402,350,372]" italics="true" pageId="32" pageNumber="333">Devadatta podolestoides</emphasis>
</taxonomicName>
;—
<treatmentCitation id="C70310F9553AFF98FE539509CD90FE8E" author="Dow" box="[433,670,350,372]" page="15" pageId="32" pageNumber="333" year="2010">
<bibRefCitation id="22334B19553AFF98FE539509CD75FE8E" author="Dow" box="[433,635,350,372]" pageId="32" pageNumber="333" refString="Dow, R. A. &amp; Reels, G. T. (2010) The Odonata of three national parks in Sarawak. Agrion, 14 (1), 14 - 19." type="journal article" year="2010">Dow &amp; Reels 2010</bibRefCitation>
: 15
</treatmentCitation>
(Kubah: part);—
<treatmentCitation id="C70310F9553AFF98FCAC9509CCD0FE8E" author="Dow" box="[846,990,350,372]" page="4" pageId="32" pageNumber="333" year="2012">
<bibRefCitation id="22334B19553AFF98FCAC9509CCC6FE8E" author="Dow" box="[846,968,350,372]" pageId="32" pageNumber="333" refString="Dow, R. A. (2012 a) Odonata collected around the Borneo Highlands Resort on Gunung Penrissen, Kuching Division, Sarawak, Malaysia in July 2012. International Dragonfly Fund Report, 50, 1 - 12." type="journal article" year="2012" yearSuffix="a">Dow 2012a</bibRefCitation>
: 4
</treatmentCitation>
(Gunung Penrissen);—Dow &amp; Reels 2013: 12 (records western
<collectingCountry id="3EB57678553AFF98FE77952BCEE1FE69" box="[405,495,381,403]" name="Malaysia" pageId="32" pageNumber="333">Sarawak</collectingCountry>
);—
<treatmentCitation id="C70310F9553AFF98FDF6952BCC5AFE69" author="Dow" box="[532,852,381,403]" page="11" pageId="32" pageNumber="333" year="2013">
<bibRefCitation id="22334B19553AFF98FDF6952BCC3CFE69" author="Dow" box="[532,818,381,403]" pageId="32" pageNumber="333" refString="Dow, R. A., Reels, G. T. &amp; Butler, S. G. (2013 b) Previously unpublished Odonata records from Sarawak, Borneo. Part II. Kubah National Park. Faunistic Studies in South-East Asian and Pacific Island Odonata, 6, 1 - 21." type="journal article" year="2013" yearSuffix="b">Dow, Reels &amp; Butler 2013b</bibRefCitation>
: 11
</treatmentCitation>
(Kubah National Park).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0EB86563553AFF98FF7595ECCB31FA1C" pageId="32" pageNumber="333" type="materials_examined">
<paragraph id="461D36E8553AFF98FF7595ECCE69FD09" blockId="32.[151,1437,442,2015]" pageId="32" pageNumber="333">
<emphasis id="74D6EAFA553AFF98FF7595ECCEB5FE29" bold="true" box="[151,443,442,467]" pageId="32" pageNumber="333">
Material.
<typeStatus id="9919884A553AFF98FEF395ECCE42FE29" box="[273,332,442,467]" pageId="32" pageNumber="333">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="9919884A553AFF98FE2F95ECCD34FE29" box="[461,570,442,467]" pageId="32" pageNumber="333" type="holotype">
<emphasis id="74D6EAFA553AFF98FE2F95ECCD34FE29" bold="true" box="[461,570,442,467]" pageId="32" pageNumber="333">Holotype</emphasis>
</typeStatus>
<specimenCount id="50A4FD61553AFF98FDA695EDCD7CFE28" box="[580,626,443,467]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
(JvT number 2084), Gunung Poteng,
<quantity id="815A9B0D553AFF98FBD095ECCB7BFE29" box="[1074,1141,442,467]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="32" pageNumber="333" unit="m" value="550.0">550m</quantity>
, Singkawang, W. Borneo [Kalimantan Barat, Kalimantan,
<collectingCountry id="3EB57678553AFF98FDFD9588CD9CFE0D" box="[543,658,478,503]" name="Indonesia" pageId="32" pageNumber="333">Indonesia</collectingCountry>
],
<date id="321C1028553AFF98FD4F9589CC47FE0D" box="[685,841,478,503]" pageId="32" pageNumber="333" value="1932-08-21">21 viii 1932</date>
, leg. L.C. de Ruiter, in
<collectionCode id="20B3AE2D553AFF98FB619589CBD5FE0C" LSID="urn:lsid:biocol.org:col:34992" box="[1155,1243,479,502]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="32" pageNumber="333">RMNH</collectionCode>
.
<emphasis id="74D6EAFA553AFF98FB0F9588CD0BFDE1" bold="true" pageId="32" pageNumber="333">
<typeStatus id="9919884A553AFF98FB0F9588CA6BFE0D" box="[1261,1381,478,503]" pageId="32" pageNumber="333" type="paratype">Paratypes</typeStatus>
(all Kalimantan Barat,
<collectingCountry id="3EB57678553AFF98FE9C9654CEF7FDE1" box="[382,505,514,539]" name="Indonesia" pageId="32" pageNumber="333">Indonesia</collectingCountry>
):
</emphasis>
<specimenCount id="50A4FD61553AFF98FDEE9655CD39FDE0" box="[524,567,515,539]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
, same location and collector as
<typeStatus id="9919884A553AFF98FC7A9654CCF3FDE1" box="[920,1021,514,539]" pageId="32" pageNumber="333" type="holotype">holotype</typeStatus>
,
<quantity id="815A9B0D553AFF98FBE59654CB44FDE1" box="[1031,1098,514,539]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="32" pageNumber="333" unit="m" value="550.0">550m</quantity>
,
<date id="321C1028553AFF98FBB79654CBCEFDE1" box="[1109,1216,514,539]" pageId="32" pageNumber="333" value="1931-11-05">5 xi 1931</date>
;
<specimenCount id="50A4FD61553AFF98FB299655CBF9FDE0" box="[1227,1271,515,539]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
, same location and collector,
<date id="321C1028553AFF98FED79671CEAEFDC5" box="[309,416,550,575]" pageId="32" pageNumber="333" value="1931-11-07">7 xi 1931</date>
, in
<collectionCode id="20B3AE2D553AFF98FE2A9671CD2EFDC4" box="[456,544,551,574]" pageId="32" pageNumber="333">BMNH</collectionCode>
;
<specimenCount id="50A4FD61553AFF98FDC99671CD59FDC4" box="[555,599,551,575]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
, all data as
<typeStatus id="9919884A553AFF98FD3B9670CC31FDC5" box="[729,831,550,575]" pageId="32" pageNumber="333" type="holotype">holotype</typeStatus>
;
<specimenCount id="50A4FD61553AFF98FCA89671CC78FDC4" box="[842,886,551,575]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
, same location and collector,
<quantity id="815A9B0D553AFF98FB239671CA0AFDC5" box="[1217,1284,551,575]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="32" pageNumber="333" unit="m" value="400.0">400m</quantity>
,
<date id="321C1028553AFF98FAED9671CA8DFDC5" box="[1295,1411,550,575]" pageId="32" pageNumber="333" value="1932-02-28">28 ii 1932</date>
;
<specimenCount id="50A4FD61553AFF98FA6C9671CFA0FD98" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
, same location and collector,
<date id="321C1028553AFF98FDE0961DCD75FD99" box="[514,635,586,611]" pageId="32" pageNumber="333" value="1932-02-28">28 ii 1932</date>
, in
<collectionCode id="20B3AE2D553AFF98FD44961DCDF0FD98" box="[678,766,587,610]" pageId="32" pageNumber="333">BMNH</collectionCode>
;
<specimenCount id="50A4FD61553AFF98FCE8961DCC36FD98" box="[778,824,587,611]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
, same location and collector,
<quantity id="815A9B0D553AFF98FB6F961CCBDEFD99" box="[1165,1232,586,611]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="32" pageNumber="333" unit="m" value="550.0">550m</quantity>
,
<date id="321C1028553AFF98FB3E961DCA52FD99" box="[1244,1372,586,611]" pageId="32" pageNumber="333" value="1932-03-13">13 iii 1932</date>
;
<specimenCount id="50A4FD61553AFF98FA88961DCA99FD98" box="[1386,1431,587,611]" pageId="32" pageNumber="333" type="generic">3 ♂</specimenCount>
, same location and collector,
<date id="321C1028553AFF98FE059639CD65FD7D" box="[487,619,622,647]" pageId="32" pageNumber="333" value="1932-03-13">13 iii 1932</date>
, in
<collectionCode id="20B3AE2D553AFF98FD799639CDE1FD7C" box="[667,751,623,646]" pageId="32" pageNumber="333">BMNH</collectionCode>
(two ex. Cowley collection);
<specimenCount id="50A4FD61553AFF98FBB39639CB71FD7C" box="[1105,1151,623,647]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
1 ♀, same location and collector,
<date id="321C1028553AFF98FEEA96C5CE81FD51" box="[264,399,658,683]" pageId="32" pageNumber="333" value="1932-07-17">17 vii 1932</date>
, in
<collectionCode id="20B3AE2D553AFF98FE5996C5CD1CFD50" box="[443,530,659,682]" pageId="32" pageNumber="333">BMNH</collectionCode>
;
<specimenCount id="50A4FD61553AFF98FDC296C5CD40FD50" box="[544,590,659,683]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
, same location and collector,
<quantity id="815A9B0D553AFF98FC4796C5CCE9FD51" box="[933,999,659,683]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="32" pageNumber="333" unit="m" value="300.0">300m</quantity>
,
<date id="321C1028553AFF98FC1796C5CB8AFD51" box="[1013,1156,658,683]" pageId="32" pageNumber="333" value="1932-08-21">21 viii 1932</date>
;
<specimenCount id="50A4FD61553AFF98FB7396C5CBCEFD50" box="[1169,1216,659,683]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
, same location and collector,
<date id="321C1028553AFF98FEE496E1CE8FFD35" box="[262,385,694,719]" pageId="32" pageNumber="333" value="1932-09-21">21 ix 1932</date>
;
<specimenCount id="50A4FD61553AFF98FE6C96E1CEB6FD34" box="[398,440,695,719]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
, same location and collector,
<date id="321C1028553AFF98FCE596E0CC75FD35" box="[775,891,694,719]" pageId="32" pageNumber="333" value="1932-10-16">16 x 1932</date>
, in
<collectionCode id="20B3AE2D553AFF98FC4196E1CCF4FD34" box="[931,1018,695,718]" pageId="32" pageNumber="333">BMNH</collectionCode>
; 1 ♀, same location and collector,
<date id="321C1028553AFF98FA6296E1CFE9FD09" pageId="32" pageNumber="333" value="1932-10-18">18 x 1932</date>
, in
<collectionCode id="20B3AE2D553AFF98FEEC968DCE6DFD09" box="[270,355,731,755]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="32" pageNumber="333">CUMZ</collectionCode>
.
</paragraph>
<paragraph id="461D36E8553AFF98FF2596A8CC1FFCA5" blockId="32.[151,1437,442,2015]" pageId="32" pageNumber="333">
<emphasis id="74D6EAFA553AFF98FF2596A8CE75FCED" bold="true" box="[199,379,766,791]" pageId="32" pageNumber="333">Other material</emphasis>
.
<emphasis id="74D6EAFA553AFF98FE6996A8CDE4FCED" bold="true" box="[395,746,766,791]" pageId="32" pageNumber="333">
Kalimantan Barat,
<collectingCountry id="3EB57678553AFF98FD9796A8CDE4FCED" box="[629,746,766,791]" name="Indonesia" pageId="32" pageNumber="333">Indonesia</collectingCountry>
</emphasis>
(all leg. L.C. de Ruiter, all in
<collectionCode id="20B3AE2D553AFF98FBA696A9CB90FCED" LSID="urn:lsid:biocol.org:col:34992" box="[1092,1182,767,791]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="32" pageNumber="333">RMNH</collectionCode>
): 1 ♀, Bagak, Gunung Raja Complex, Singkawang,
<date id="321C1028553AFF98FE059774CD6FFCC1" box="[487,609,802,827]" pageId="32" pageNumber="333" value="1932-02-26">26 ii 1932</date>
;
<specimenCount id="50A4FD61553AFF98FD8D9775CD92FCC0" box="[623,668,803,827]" pageId="32" pageNumber="333" type="generic">2 ♂</specimenCount>
, location as
<typeStatus id="9919884A553AFF98FCD09774CC99FCC1" box="[818,919,802,827]" pageId="32" pageNumber="333" type="holotype">holotype</typeStatus>
,
<date id="321C1028553AFF98FC469775CB19FCC1" box="[932,1047,802,827]" pageId="32" pageNumber="333" value="1934-04-01">1 iv 1934</date>
; 1 ♀ (prothorax destroyed), same location,
<quantity id="815A9B0D553AFF98FF1D9710CE4CFCA5" box="[255,322,838,863]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="32" pageNumber="333" unit="m" value="550.0">550m</quantity>
,
<date id="321C1028553AFF98FEAF9711CEB4FCA5" box="[333,442,838,863]" pageId="32" pageNumber="333" value="1934-04-01">1 iv 1934</date>
; ♂, same location,
<date id="321C1028553AFF98FD739711CC02FCA5" box="[657,780,838,863]" pageId="32" pageNumber="333" value="1939-06-30">30 vi 1939</date>
.
</paragraph>
<paragraph id="461D36E8553AFF98FF25973CCB31FA1C" blockId="32.[151,1437,442,2015]" pageId="32" pageNumber="333">
<emphasis id="74D6EAFA553AFF98FF25973CCEB7FC79" bold="true" box="[199,441,874,899]" pageId="32" pageNumber="333">
<collectingCountry id="3EB57678553AFF98FF25973CCE3DFC79" box="[199,307,874,899]" name="Malaysia" pageId="32" pageNumber="333">Sarawak</collectingCountry>
,
<collectingCountry id="3EB57678553AFF98FEA1973CCEBAFC79" box="[323,436,874,899]" name="Malaysia" pageId="32" pageNumber="333">Malaysia</collectingCountry>
:
</emphasis>
<specimenCount id="50A4FD61553AFF98FE26973DCEFCFC78" box="[452,498,875,899]" pageId="32" pageNumber="333" type="generic">2 ♂</specimenCount>
1 ♀, tributary to Sungai Annah Rais, Annah Rais, Gunung Penrissen, Kuching division,
<date id="321C1028553AFF98FF1D97D9CE7EFC5D" box="[255,368,910,935]" pageId="32" pageNumber="333" value="2005-05-19">19 v 2005</date>
, leg. R.A. Dow, in coll. Hämäläinen;
<specimenCount id="50A4FD61553AFF98FCF097D8CC33FC5C" box="[786,829,910,935]" pageId="32" pageNumber="333" type="generic">
<emphasis id="74D6EAFA553AFF98FCF097D8CC2EFC5D" bold="true" box="[786,800,910,935]" pageId="32" pageNumber="333">1</emphasis>
</specimenCount>
, tributary to Sungai Annah Rais, Annah Rais, Gunung Penrissen, Kuching division,
<date id="321C1028553AFF98FE0297E4CD43FC31" box="[480,589,946,971]" pageId="32" pageNumber="333" value="2006-01-26">26 i 2006</date>
, leg. G.T. Reels;
<specimenCount id="50A4FD61553AFF98FCE897E5CC3BFC30" box="[778,821,947,971]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
(
<collectionCode id="20B3AE2D553AFF98FCA197E4CC93FC31" LSID="urn:lsid:biocol.org:col:34992" box="[835,925,946,971]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="32" pageNumber="333">RMNH</collectionCode>
.INS.506808), forest stream running onto golf course, Borneo Highlands Resort, same mountain,
<date id="321C1028553AFF98FD309781CC5BFC15" box="[722,853,982,1007]" pageId="32" pageNumber="333" value="2012-07-21">21 vii 2012</date>
, leg. R.A. Dow;
<specimenCount id="50A4FD61553AFF98FBF09781CB30FC14" box="[1042,1086,983,1007]" pageId="32" pageNumber="333" type="generic">2 ♂</specimenCount>
, same data;
<specimenCount id="50A4FD61553AFF98FB2B9781CBFBFC14" box="[1225,1269,983,1007]" pageId="32" pageNumber="333" type="generic">2 ♂</specimenCount>
, same location and collector,
<date id="321C1028553AFF98FEDB97ADCEB6FBE9" box="[313,440,1018,1043]" pageId="32" pageNumber="333" value="2014-06-28">28 vi 2014</date>
;
<specimenCount id="50A4FD61553AFF98FE2797ADCEFEFBE8" box="[453,496,1019,1043]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
(
<collectionCode id="20B3AE2D553AFF98FDE397ACCD55FBE9" LSID="urn:lsid:biocol.org:col:34992" box="[513,603,1018,1043]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="32" pageNumber="333">RMNH</collectionCode>
.INS.506809), stream near forest trail, same resort,
<date id="321C1028553AFF98FB4097ADCA26FBE9" box="[1186,1320,1018,1043]" pageId="32" pageNumber="333" value="2012-07-22">22 vii 2012</date>
, leg. R.A. Dow;
<specimenCount id="50A4FD61553AFF98FF3E9048CE08FBCC" box="[220,262,1054,1079]" pageId="32" pageNumber="333" type="generic">5 ♂</specimenCount>
1 ♀, same data;
<specimenCount id="50A4FD61553AFF98FE219049CEE1FBCC" box="[451,495,1055,1079]" pageId="32" pageNumber="333" type="generic">3 ♂</specimenCount>
, forest stream, same resort,
<date id="321C1028553AFF98FCCB9049CCAAFBCD" box="[809,932,1054,1079]" pageId="32" pageNumber="333" value="2012-06-23">23 vi 2012</date>
, leg. R.A. Dow;
<specimenCount id="50A4FD61553AFF98FB829049CB82FBCC" box="[1120,1164,1055,1079]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
, forest stream near trail,
<date id="321C1028553AFF98FF759014CE16FBA1" box="[151,280,1090,1115]" pageId="32" pageNumber="333" value="2012-07-25">25 vii 2012</date>
, leg. R.A. Dow;
<specimenCount id="50A4FD61553AFF98FE319015CEF2FBA0" box="[467,508,1091,1115]" pageId="32" pageNumber="333" type="generic">3 ♂</specimenCount>
1 ♀, forest streams below Penrissen peak, same resort,
<date id="321C1028553AFF98FB8B9014CBE5FBA1" box="[1129,1259,1090,1115]" pageId="32" pageNumber="333" value="2012-07-26">26 vii 2012</date>
, leg. R.A. Dow, in coll. Hämäläinen;
<specimenCount id="50A4FD61553AFF98FE649031CEBAFB84" box="[390,436,1127,1151]" pageId="32" pageNumber="333" type="generic">2 ♂</specimenCount>
, same location and collector,
<date id="321C1028553AFF98FCEA9031CC71FB85" box="[776,895,1126,1151]" pageId="32" pageNumber="333" value="2014-07-01">1 vii 2014</date>
;
<specimenCount id="50A4FD61553AFF98FC6E9031CCB6FB84" box="[908,952,1127,1151]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
(SAR05_
<collectionCode id="20B3AE2D553AFF98FBCE9030CB63FB85" box="[1068,1133,1126,1151]" httpUri="http://grbio.org/cool/90v0-7gw3" name="Australian Mycological Panel" pageId="32" pageNumber="333">AMP</collectionCode>
8), 1 ♀ (SAR05_
<collectionCode id="20B3AE2D553AFF98FACE9030CA62FB85" box="[1324,1388,1126,1151]" httpUri="http://grbio.org/cool/90v0-7gw3" name="Australian Mycological Panel" pageId="32" pageNumber="333">AMP</collectionCode>
9; in tandem with the male), small stream, Gunung Serapi, Kubah National Park, Matang Range, Kuching division,
<date id="321C1028553AFF98FA6290DDCFFEFB3D" pageId="32" pageNumber="333" value="2005-04-13">13 iv 2005</date>
, leg. R.A. Dow;
<specimenCount id="50A4FD61553AFF98FE5590F9CEEDFB3C" box="[439,483,1199,1223]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
(
<collectionCode id="20B3AE2D553AFF98FE1690F8CD40FB3D" LSID="urn:lsid:biocol.org:col:34992" box="[500,590,1198,1223]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="32" pageNumber="333">RMNH</collectionCode>
.INS.506258), drain by road on Gunung Serapi, same national park,
<date id="321C1028553AFF98FABD90F9CFDCFB11" pageId="32" pageNumber="333" value="2012-03-30">30 iii 2012</date>
, leg. R.A. Dow; 1 ♀, same data;
<specimenCount id="50A4FD61553AFF98FDBE9085CD87FB10" box="[604,649,1235,1259]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
(
<collectionCode id="20B3AE2D553AFF98FD799084CDFBFB11" LSID="urn:lsid:biocol.org:col:34992" box="[667,757,1234,1259]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="32" pageNumber="333">RMNH</collectionCode>
.INS.506910), ♀ (
<collectionCode id="20B3AE2D553AFF98FC239084CB15FB11" LSID="urn:lsid:biocol.org:col:34992" box="[961,1051,1234,1259]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="32" pageNumber="333">RMNH</collectionCode>
.INS.506909), stream at ca
<quantity id="815A9B0D553AFF98FAB79085CA96FB11" box="[1365,1432,1235,1259]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="32" pageNumber="333" unit="m" value="400.0">400m</quantity>
, Gunung Serapi, same national park,
<date id="321C1028553AFF98FDD190A1CDBEFAF5" box="[563,688,1270,1295]" pageId="32" pageNumber="333" value="2012-09-30">30 ix 2012</date>
, leg. R.A. Dow;
<specimenCount id="50A4FD61553AFF98FC9290A1CC92FAF4" box="[880,924,1271,1295]" pageId="32" pageNumber="333" type="generic">2 ♂</specimenCount>
, same data; ♂, stream at
<quantity id="815A9B0D553AFF98FB5E90A0CA38FAF5" box="[1212,1334,1270,1295]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="32" pageNumber="333" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">600700m</quantity>
, Gunung Serapi, same area,
<date id="321C1028553AFF98FE88914DCED1FAC9" box="[362,479,1306,1331]" pageId="32" pageNumber="333" value="2013-07-01">1 vii 2013</date>
, leg. R.A. Dow;
<specimenCount id="50A4FD61553AFF98FD7E914DCDC6FAC8" box="[668,712,1307,1331]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
, waterfall trail, same national park,
<date id="321C1028553AFF98FBB9914DCBC1FAC9" box="[1115,1231,1306,1331]" pageId="32" pageNumber="333" value="2008-02-13">13 ii 2008</date>
, leg. G.T. Reels;
<specimenCount id="50A4FD61553AFF98FA6C914DCFA2FAAC" pageId="32" pageNumber="333" type="generic">4 ♂</specimenCount>
(PomS 7, 12, 13, 19a-2014), waterfall trail and waterfall at end of trail, same national park,
<date id="321C1028553AFF98FB069169CA6BFAAD" box="[1252,1381,1342,1367]" pageId="32" pageNumber="333" value="2014-03-13">13 iii 2014</date>
, leg. P.O.M. Steinhoff, in coll. Steinhoff;
<specimenCount id="50A4FD61553AFF98FDD59135CD6BFA80" box="[567,613,1379,1403]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
, stream above waterfall at end waterfall trail, same national park,
<date id="321C1028553AFF98FA839134CFDCFA65" pageId="32" pageNumber="333" value="2009-09-16">16 ix 2009</date>
, leg. R.A. Dow;
<specimenCount id="50A4FD61553AFF98FE7691D1CECEFA64" box="[404,448,1415,1439]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
1 ♀, stream above church, Gunung Singgai, same range,
<date id="321C1028553AFF98FBB491D1CBB3FA65" box="[1110,1213,1415,1439]" pageId="32" pageNumber="333" value="2008-10-02">2 x 2008</date>
, leg. R.A. Dow; ♂, stream by path to site of colonial era bungalow, Mount Matang, same range,
<date id="321C1028553AFF98FBFE91FDCB91FA39" box="[1052,1183,1450,1475]" pageId="32" pageNumber="333" value="2014-03-31">31 iii 2014</date>
, leg. R.A. Dow;
<specimenCount id="50A4FD61553AFF98FA8A91FDCA99FA38" box="[1384,1431,1451,1475]" pageId="32" pageNumber="333" type="generic">1 ♂</specimenCount>
, Ranchan Recreational Park, Serian, Samarahan division,
<date id="321C1028553AFF98FCF59199CC85FA1D" box="[791,907,1486,1511]" pageId="32" pageNumber="333" value="2008-02-14">14 ii 2008</date>
, leg. R.A. Dow.
</paragraph>
</subSubSection>
<subSubSection id="0EB86563553AFF99FF2591A4CDCDFA66" lastPageId="33" lastPageNumber="334" pageId="32" pageNumber="333" type="description">
<paragraph id="461D36E8553AFF98FF2591A4CE87F98D" blockId="32.[151,1437,442,2015]" pageId="32" pageNumber="333">
<emphasis id="74D6EAFA553AFF98FF2591A4CDFDF9F1" bold="true" box="[199,755,1522,1547]" pageId="32" pageNumber="333">Description of male (based on SAR05_AMP8).</emphasis>
Head (
<figureCitation id="DE992A6D553AFF98FCA791A4CC96F9F1" box="[837,920,1522,1547]" captionStart="FIGURES 14 19" captionStartId="16.[151,264,1803,1825]" captionTargetBox="[151,1436,733,1781]" captionTargetId="figure@16.[151,1436,733,1782]" captionTargetPageId="16" captionText="FIGURES 14 19. Heads of male Devadatta species: (14) D. aran holotype; (15) D. basilanensis Davao Oriental, Mindanao; (16) D. clavicauda holotype; (17) D. podolestoides male described in this paper; (18) D. somoh holotype; (19) D. tanduk holotype." httpUri="https://zenodo.org/record/253954/files/figure.png" pageId="32" pageNumber="333">Fig. 17</figureCitation>
). Labium mostly pale. Mandible bases, genae shining dark brown. Labrum, clypeus, frons and vertex to level of lateral ocelli black with metallic green reflection, remainder of dorsum of head matte black with indistinct brown area to rear; underside extensively pale. Antennae almost entirely black.
</paragraph>
<paragraph id="461D36E8553AFF98FF2592D4CDDBF825" blockId="32.[151,1437,442,2015]" pageId="32" pageNumber="333">
Thorax: Most of propleuron and pronotum black, with pale lateral marks on anterior pronotal lobe and rear of middle pronotal lobe, bronzy brown area centrally on posterior pronotal lobe. Dorsum of synthorax matte black with pair of small pale marks in antealar triangle and small pale mark between arms of anterior bifurcation of middorsal carina. Synthorax laterally with pale marks as follows (
<figureCitation id="DE992A6D553AFF98FC7A92B8CCFEF8FD" box="[920,1008,1774,1799]" captionStart="FIGURES 20 25" captionStartId="17.[151,264,1919,1941]" captionTargetBox="[194,1386,193,1882]" captionTargetId="figure@17.[194,1387,193,1884]" captionTargetPageId="17" captionText="FIGURES 20 25. Synthorax of male Devadatta species, lateral view: (20) D. aran holotype; (21) D. basilanensis Davao Oriental, Mindanao; (22) D. clavicauda holotype; (23) D. podolestoides male described in this paper; (24) D. somoh holotype; (25) D. tanduk holotype." httpUri="https://zenodo.org/record/253955/files/figure.png" pageId="32" pageNumber="333">Fig. 23</figureCitation>
): 2 short streaks along lower side of mesopleural suture, longer streak under interpleural suture for almost its entire length, a larger subtriangular area and smaller mark under metapleural suture. Area between metapleural carina and venter brown. Venter of synthorax largely pale (
<figureCitation id="DE992A6D553AFF98FE4E930CCD0AF889" box="[428,516,1882,1907]" captionStart="FIGURES 30 33" captionStartId="19.[151,264,1787,1809]" captionTargetBox="[151,1436,611,1753]" captionTargetId="figure@19.[151,1436,611,1754]" captionTargetPageId="19" captionText="FIGURES 30 33. Venter of synthorax of male Devadatta species: (30) D. basilanensis Davao Oriental, Mindanao; (31) D. podolestoides male described in this paper; (32) D. clavicauda holotype; (33) D. clavicauda paratype (SAR 09 _ 10 _ AMP 40, Gunung Serapi)." httpUri="https://zenodo.org/record/253957/files/figure.png" pageId="32" pageNumber="333">Fig. 31</figureCitation>
). Legs brownish grey, with indistinct darker and paler areas. White pruinosity sparsely present on coxae and adjacent parts of pro- and synthorax and venter of synthorax. Wings with arculus ca level of Ax 5, quadrilateral with two crossveins, cubito-anal space free in all wings, pt dark brown. Membrane with brownish tint, darker at extreme tips of both wings.
</paragraph>
<paragraph id="461D36E8553BFF99FF2594C1CC45FE9F" blockId="33.[151,1437,151,1977]" pageId="33" pageNumber="334">
Abdomen: Black above and very dark brown laterally, becoming almost entirely black on terminal 3 segments, S38 with a broad pale basal ring, widely interrupted dorsally. Genital ligula (
<figureCitation id="DE992A6D553BFF99FBDA94EACB80FF2E" box="[1080,1166,188,213]" captionStart="FIGURES 38 39" captionStartId="20.[151,264,1911,1933]" captionTargetBox="[151,1436,1261,1889]" captionTargetId="figure@20.[151,1436,1261,1889]" captionTargetPageId="20" captionText="FIGURES 38 39. Genital ligula of two Devadatta species, scale bars = 0.5 mm: (38) D. aran holotype; (39) D. podolestoides male described in this paper." httpUri="https://zenodo.org/record/253959/files/figure.png" pageId="33" pageNumber="334">Fig. 39</figureCitation>
) typical for the group, seminal vesicle pale. Anal appendages dark brown. Paraprocts extremely short. Cerci longer than S10, in dorsal view (
<figureCitation id="DE992A6D553BFF99FF3F9552CE20FEE6" box="[221,302,260,285]" captionStart="FIGURES 40 45" captionStartId="21.[151,264,1699,1721]" captionTargetBox="[162,1425,203,1663]" captionTargetId="figure@21.[151,1436,193,1677]" captionTargetPageId="21" captionText="FIGURES 40 45. Male anal appendages, dorsal view: (40) D. aran holotype; (41) D. basilanensis Davao Oriental, Mindanao; (42) D. clavicauda holotype; (43) D. podolestoides Kubah NP, Sarawak; (44) D. somoh holotype; (45) D. tanduk holotype." httpUri="https://zenodo.org/record/253960/files/figure.png" pageId="33" pageNumber="334">Fig. 43</figureCitation>
) broad at base, narrowing rather abruptly at ca one-quarter of their length from base, in-turned at less than a right angle at half length, apical half moderately expanded (
<figureCitation id="DE992A6D553BFF99FC609571CCD9FEBA" box="[898,983,295,320]" captionStart="FIGURES 46 51" captionStartId="22.[151,264,1951,1973]" captionTargetBox="[162,1418,393,1916]" captionTargetId="figure@22.[151,1436,373,1929]" captionTargetPageId="22" captionText="FIGURES 46 51. Male anal appendages, lateral view (scale bars 1 mm): (46) D. aran holotype; (47) D. basilanensis Davao Oriental, Mindanao; (48) D. clavicauda holotype; (49) D. podolestoides Kubah NP, Sarawak; (50) D. somoh holotype; (51) D. tanduk holotype." httpUri="https://zenodo.org/record/253961/files/figure.png" pageId="33" pageNumber="334">Fig. 49</figureCitation>
). Interiorly deeply excavated from point where narrows in dorsal view to tip, brown inside excavation.
</paragraph>
<paragraph id="461D36E8553BFF99FF259539CB49FE72" blockId="33.[151,1437,151,1977]" box="[199,1095,367,392]" pageId="33" pageNumber="334">
<emphasis id="74D6EAFA553BFF99FF259539CE78FE72" bold="true" box="[199,374,367,392]" pageId="33" pageNumber="334">Measurements</emphasis>
[mm]: Abdomen without anal appendages 32, cerci 1, Hw 28.5.
</paragraph>
<paragraph id="461D36E8553BFF99FF2595C2CC54FE0F" blockId="33.[151,1437,151,1977]" pageId="33" pageNumber="334">
<emphasis id="74D6EAFA553BFF99FF2595C2CC1CFE57" bold="true" box="[199,786,404,429]" pageId="33" pageNumber="334">Description of female (based on SAR05_AMP9).</emphasis>
As male except as noted. Head: Labrum mostly creamy brown. Most of frons dark brown, remainder of dorsum of head matte black with postocular lobes and a band joining them pale (
<figureCitation id="DE992A6D553BFF99FE88958ACEB0FE0F" box="[362,446,476,501]" captionStart="FIGURES 52 57" captionStartId="23.[151,264,1409,1431]" captionTargetBox="[151,1436,176,1394]" captionTargetId="figure@23.[151,1436,175,1395]" captionTargetPageId="23" captionText="FIGURES 52 57. Heads of Devadatta female specimens described in this paper: (52) D. aran; (53) D. basilanensis; (54) D. clavicauda; (55) D. podolestoides; (56) D. somoh; (57) D. tanduk." httpUri="https://zenodo.org/record/253962/files/figure.png" pageId="33" pageNumber="334">Fig. 55</figureCitation>
). Antennae with pedicel pale brown.
</paragraph>
<paragraph id="461D36E8553BFF99FF2595A9CFECFCEE" blockId="33.[151,1437,151,1977]" pageId="33" pageNumber="334">
Thorax: Pronotum (
<figureCitation id="DE992A6D553BFF99FE4E95A9CD0FFDE2" box="[428,513,511,536]" captionStart="FIGURES 58 63" captionStartId="25.[151,264,1932,1954]" captionTargetBox="[151,1436,411,1910]" captionTargetId="figure@25.[151,1436,411,1910]" captionTargetPageId="25" captionText="FIGURES 58 63. Prothorax of Devadatta female specimens described in this paper, dorsal view: (58) D. aran; (59) D. basilanensis; (60) D. clavicauda; (61) D. podolestoides; (62) D. somoh; (63) D. tanduk." httpUri="https://zenodo.org/record/253963/files/figure.png" pageId="33" pageNumber="334">Fig. 61</figureCitation>
) mottled brown, with raised part of anterior lobe extensively pale yellow, pale yellow band along free margin of posterior lobe, widest laterally, narrowly divided centrally. No horns on middle pronotal lobe. Dorsum of synthorax brown with middorsal carina black, small pale blue mark between arms of anterior bifurcation of middorsal carina. Synthorax laterally brown with pale marks as follows (
<figureCitation id="DE992A6D553BFF99FB02963ACA39FD7E" box="[1248,1335,620,645]" captionStart="FIGURES 68 73" captionStartId="28.[151,264,1928,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@28.[151,1436,193,1907]" captionTargetPageId="28" captionText="FIGURES 68 73. Synthorax of Devadatta female specimens described in this paper, lateral view: (68) D. aran; (69) D. basilanensis; (70) D. clavicauda; (71) D. podolestoides; (72) D. somoh; (73) D. tanduk." httpUri="https://zenodo.org/record/253965/files/figure.png" pageId="33" pageNumber="334">Fig. 71</figureCitation>
): 3 short streaks along lower side of mesopleural suture, one of these alongside mesokatepisternum, streak under interpleural suture broken and irregular. Wings with arculus ca level of Ax 4 (
<quantity id="815A9B0D553BFF99FC1F96E2CB29FD37" box="[1021,1063,692,717]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="33" pageNumber="334" unit="in" value="3.0">3 in</quantity>
right Fw), quadrilateral with two crossveins (one in left Fw), pt dark brown with pale area apically. Extreme tips of both wings tinted brown, darker in Hw.
</paragraph>
<paragraph id="461D36E8553BFF99FF259749CE4FFC7A" blockId="33.[151,1437,151,1977]" pageId="33" pageNumber="334">Abdomen: Brown, becoming darker on successive segments; basal ring, widely interrupted dorsally, also present on S2. Cerci black, subtriangular, shorter than S10. Ovipositor short, only reaching beyond S10 to ca level of tips of cerci.</paragraph>
<paragraph id="461D36E8553BFF99FF2597DACB19FC5F" blockId="33.[151,1437,151,1977]" box="[199,1047,908,933]" pageId="33" pageNumber="334">
<emphasis id="74D6EAFA553BFF99FF2597DACE78FC5F" bold="true" box="[199,374,908,933]" pageId="33" pageNumber="334">Measurements</emphasis>
[mm]: Abdomen without ovipositor or cerci 29.5, Hw 28.5.
</paragraph>
<paragraph id="461D36E8553BFF99FF2597F9CDF1FB87" blockId="33.[151,1437,151,1977]" pageId="33" pageNumber="334">
<emphasis id="74D6EAFA553BFF99FF2597F9CE3BFC32" bold="true" box="[199,309,943,968]" pageId="33" pageNumber="334">Variation</emphasis>
: Apart from minor differences in the extent of pale markings, there is little significant variation in both sexes apart from size and the following characters. In males occasionally there is no darkened area at all at tip in one or both pairs of wings; the degree of the general tinting of the wing membrane is variable. The arculus typically arises at Ax 4, occasionally at Ax 3 or Ax
<quantity id="815A9B0D553BFF99FD34904ACC0EFBCF" box="[726,768,1052,1077]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="33" pageNumber="334" unit="in" value="5.0">5 in</quantity>
one or more wings (
<figureCitation id="DE992A6D553BFF99FC07904ACB3BFBCE" box="[997,1077,1052,1077]" captionStart="FIGURES 34 37" captionStartId="20.[151,264,1128,1150]" captionTargetBox="[151,1436,193,1106]" captionTargetId="figure@20.[151,1436,193,1107]" captionTargetPageId="20" captionText="FIGURES 34 37. Wings of some Devadatta species: (34) D. basilanensis male Zamboanga, Mindanao; (35) D. basilanensis female Cotabato, Mindanao; (36) D. clavicauda paratype, Gunung Santubong, Sarawak; (37) D. podolestoides paratype, Gunung Poteng, Kalimantan Barat." httpUri="https://zenodo.org/record/253958/files/figure.png" pageId="33" pageNumber="334">Fig. 37</figureCitation>
shows the wings of a
<typeStatus id="9919884A553BFF99FAC9904ACA9FFBCE" box="[1323,1425,1052,1076]" pageId="33" pageNumber="334" type="paratype">paratype</typeStatus>
). Very rarely there is a single crossvein in the cubito-anal space of one wing. In females the pterostigma is sometimes entirely dark; this variation is associated with maturity.
</paragraph>
<paragraph id="461D36E8553BFF99FF2590D1CBBEFAAF" blockId="33.[151,1437,151,1977]" pageId="33" pageNumber="334">
The male described differs from the
<typeStatus id="9919884A553BFF99FD9590D1CDD7FB5A" box="[631,729,1159,1184]" pageId="33" pageNumber="334" type="holotype">holotype</typeStatus>
by being generally browner, in particular there are irregular brown marks between antennae and lateral ocelli; the brown patches at the rear of the head are joined by a brown band along the occipital ridge; the abdomen is more brown than in the male described, but the pale marks are faded and not very distinct. Additionally the
<typeStatus id="9919884A553BFF99FDA590A2CDA7FAF7" box="[583,681,1268,1293]" pageId="33" pageNumber="334" type="holotype">holotype</typeStatus>
lacks pale marks in the antealar triangle and the pale mark between the arms of the anterior bifurcation of the middorsal carina are very faint. The cerci of all males (including the
<typeStatus id="9919884A553BFF99FF75916ACFF5FAAF" box="[151,251,1340,1365]" pageId="33" pageNumber="334" type="holotype">holotype</typeStatus>
) bear small spines; these are highly variable in size and position and non-diagnostic.
</paragraph>
<paragraph id="461D36E8553BFF99FF259109CDCDFA66" blockId="33.[151,1437,151,1977]" pageId="33" pageNumber="334">
<emphasis id="74D6EAFA553BFF99FF259109CE78FA82" bold="true" box="[199,374,1375,1400]" pageId="33" pageNumber="334">Measurements</emphasis>
[mm]: Males: Abdomen without anal appendages 3240, Hw 28.532.5. Females: Abdomen without ovipositor or cerci 29.531.5, Hw 2829.
</paragraph>
</subSubSection>
<subSubSection id="0EB86563553BFF99FF2591F1CE71F843" pageId="33" pageNumber="334" type="discussion">
<paragraph id="461D36E8553BFF99FF2591F1CA2AF8FF" blockId="33.[151,1437,151,1977]" pageId="33" pageNumber="334">
<emphasis id="74D6EAFA553BFF99FF2591F1CE35FA3A" bold="true" box="[199,315,1447,1472]" pageId="33" pageNumber="334">Remarks.</emphasis>
The
<typeStatus id="9919884A553BFF99FE9491FFCEA8FA3B" box="[374,422,1449,1473]" pageId="33" pageNumber="334">type</typeStatus>
series of
<taxonomicName id="81A24D6B553BFF99FDF491FFCDDAFA3A" box="[534,724,1448,1472]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA553BFF99FDF491FFCDDAFA3A" box="[534,724,1448,1472]" italics="true" pageId="33" pageNumber="334">D. podolestoides</emphasis>
</taxonomicName>
is stated by
<bibRefCitation id="22334B19553BFF99FC8791F1CB11FA3A" author="Laidlaw" box="[869,1055,1447,1472]" pageId="33" pageNumber="334" refString="Laidlaw, F. F. (1934 a) Notes on some Oriental Dragonflies (Odonata), with description of a new species. Stylops, 3 (5), 99 - 103." type="journal article" year="1934" yearSuffix="a">Laidlaw (1934a)</bibRefCitation>
to consist of
<specimenCount id="50A4FD61553BFF99FB5991FECA2EFA3A" box="[1211,1312,1447,1472]" pageId="33" pageNumber="334" type="male">19 males</specimenCount>
(including the
<typeStatus id="9919884A553BFF99FF23919ACE2BFA1F" box="[193,293,1484,1509]" pageId="33" pageNumber="334" type="holotype">holotype</typeStatus>
) and four females collected at “Cjn” (presumably a misreading of Gn [Gunung] by Laidlaw) Poteng near Singkawang in present day Kalimantan Barat, “between
<date id="321C1028553BFF99FCB191A7CCC7F9F2" box="[851,969,1520,1544]" pageId="33" pageNumber="334" value="1931-11">Nov. 1931</date>
and
<date id="321C1028553BFF99FBE091A6CB78F9F2" box="[1026,1142,1520,1544]" pageId="33" pageNumber="334" value="1932-10">Oct. 1932</date>
”, five males and a female from “Kinakok river, [Mount] Kinabalu”, and two males collected at “Mampawa by Andjoengan” (also in the Singkawang area) by Coomans de Ruiter in
<date id="321C1028553BFF99FD799261CC27F9AA" box="[667,809,1591,1616]" pageId="33" pageNumber="334" value="1932-03">March 1932</date>
. A total of
<specimenCount id="50A4FD61553BFF99FC509261CB17F9AA" box="[946,1049,1591,1616]" pageId="33" pageNumber="334" type="male">15 males</specimenCount>
(including the
<typeStatus id="9919884A553BFF99FB2E9261CA3EF9AA" box="[1228,1328,1591,1616]" pageId="33" pageNumber="334" type="holotype">holotype</typeStatus>
) and one female from Gunung Poteng collected within the date range stated for the
<typeStatus id="9919884A553BFF99FBE8920BCB34F98F" box="[1034,1082,1629,1653]" pageId="33" pageNumber="334">type</typeStatus>
series were found in RMNH, BMNH and CUMZ, but not all were marked as
<typeStatus id="9919884A553BFF99FD2292D7CDF1F963" box="[704,767,1665,1689]" pageId="33" pageNumber="334">types</typeStatus>
; all have been listed as
<typeStatus id="9919884A553BFF99FBF192D7CB40F963" box="[1043,1102,1665,1689]" pageId="33" pageNumber="334">types</typeStatus>
here, leaving four males and three females unaccounted for. None of the specimens from Mount Kinabalu are
<taxonomicName id="81A24D6B553BFF99FBC892F3CBE9F946" box="[1066,1255,1700,1724]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA553BFF99FBC892F3CBE9F946" box="[1066,1255,1700,1724]" italics="true" pageId="33" pageNumber="334">D. podolestoides</emphasis>
</taxonomicName>
. Only one male
<taxonomicName id="81A24D6B553BFF99FF75929ECE02F91A" box="[151,268,1736,1760]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA553BFF99FF75929ECE02F91A" box="[151,268,1736,1760]" italics="true" pageId="33" pageNumber="334">Devadatta</emphasis>
</taxonomicName>
from Mampawa was located, in RMNH, collected on
<date id="321C1028553BFF99FC6A929ECB32F91A" box="[904,1084,1735,1760]" pageId="33" pageNumber="334" value="1932-03-19">19 March 1932</date>
; this is presumably one of the
<typeStatus id="9919884A553BFF99FF7592BBCE0BF8FF" box="[151,261,1773,1797]" pageId="33" pageNumber="334" type="paratype">paratypes</typeStatus>
, however it actually belongs to
<taxonomicName id="81A24D6B553BFF99FD8A92BBCC08F8FE" box="[616,774,1772,1796]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="74D6EAFA553BFF99FD8A92BBCC08F8FE" box="[616,774,1772,1796]" italics="true" pageId="33" pageNumber="334">D. clavicauda</emphasis>
</taxonomicName>
and is listed as a
<typeStatus id="9919884A553BFF99FC2992BBCB25F8FF" box="[971,1067,1773,1797]" pageId="33" pageNumber="334" type="paratype">paratype</typeStatus>
of that species above.
</paragraph>
<paragraph id="461D36E8553BFF99FF259346CE71F843" blockId="33.[151,1437,151,1977]" pageId="33" pageNumber="334">
<taxonomicName id="81A24D6B553BFF99FF259346CED3F8D2" box="[199,477,1808,1832]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA553BFF99FF259346CED3F8D2" box="[199,477,1808,1832]" italics="true" pageId="33" pageNumber="334">Devadatta podolestoides</emphasis>
</taxonomicName>
is locally common at some locations in southwestern
<collectingCountry id="3EB57678553BFF99FBA29359CBADF8D2" box="[1088,1187,1807,1832]" name="Malaysia" pageId="33" pageNumber="334">Sarawak</collectingCountry>
, and judging from the
<typeStatus id="9919884A553BFF99FF759363CFC9F8B7" box="[151,199,1845,1869]" pageId="33" pageNumber="334">type</typeStatus>
series, on Gunung Poteng near Singkawang in western Kalimantan. Above
<quantity id="815A9B0D553BFF99FBDF9362CB72F8B6" box="[1085,1148,1844,1868]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="33" pageNumber="334" unit="m" value="800.0">800m</quantity>
on Gunung Penrissen in
<collectingCountry id="3EB57678553BFF99FF759301CFF4F88A" box="[151,250,1879,1904]" name="Malaysia" pageId="33" pageNumber="334">Sarawak</collectingCountry>
, at the Indonesian border, it is the only
<taxonomicName id="81A24D6B553BFF99FD26930ECC37F88A" box="[708,825,1880,1904]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="334" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA553BFF99FD26930ECC37F88A" box="[708,825,1880,1904]" italics="true" pageId="33" pageNumber="334">Devadatta</emphasis>
</taxonomicName>
species that has been found, but at lower altitudes in
<collectingCountry id="3EB57678553BFF99FF75932ACFF9F86F" box="[151,247,1916,1941]" name="Malaysia" pageId="33" pageNumber="334">Sarawak</collectingCountry>
it overlaps with
<taxonomicName id="81A24D6B553BFF99FE22932BCD6FF86E" box="[448,609,1916,1940]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="33" pageNumber="334" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="74D6EAFA553BFF99FE22932BCD6FF86E" box="[448,609,1916,1940]" italics="true" pageId="33" pageNumber="334">D. clavicauda</emphasis>
</taxonomicName>
. The habitat of this species is typically small forest streams in hilly or mountainous terrain.
</paragraph>
</subSubSection>
</treatment>
</document>