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<document id="0874899C4DC7AAD16A107FADE3180E02" ID-DOI="10.11646/zootaxa.4033.3.1" ID-GBIF-Dataset="e9ff6a09-2779-4af0-8858-992f4b9a50e3" ID-ISSN="1175-5326" ID-Zenodo-Dep="253944" ID-ZooBank="2576327B-E225-467E-BBF4-B5E5A6A64649" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461297215894" checkinUser="plazi" docAuthor="Dow, Rory A., Hämäläinen, Matti &amp; Stokvis, Frank R." docDate="2015" docId="CE0B87FE5514FFAFFF7590C0CEECF97F" docLanguage="en" docName="zt04033p349.pdf" docOrigin="Zootaxa 4033 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Devadatta aran Dow, Hämäläinen &amp; Stokvis, 2015, spec. nov." docType="treatment" docVersion="9" lastPageNumber="324" masterDocId="3232FF86551AFFB8FFE29456CF0EFFFA" masterDocTitle="Revision of the genus Devadatta Kirby, 1890 in Borneo based on molecular and morphological methods, with descriptions of four new species (Odonata: Zygoptera: Devadattidae)" masterLastPageNumber="349" masterPageNumber="301" pageNumber="315" updateTime="1698616785481" updateUser="plazi">
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<mods:title id="116234C1A9C793454DAF4A8A6ECDFC76">Revision of the genus Devadatta Kirby, 1890 in Borneo based on molecular and morphological methods, with descriptions of four new species (Odonata: Zygoptera: Devadattidae)</mods:title>
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(
<figureCitation id="DE992A6D5514FFB6FF7D90EFCFE2FB28" box="[159,236,1209,1234]" captionStart="FIGURE 1" captionStartId="7.[151,250,1912,1934]" captionTargetBox="[160,1436,198,1884]" captionTargetId="figure@7.[151,1436,193,1890]" captionTargetPageId="7" captionText="FIGURE 1. COI gene tree for 73 specimens of Devadatta and one outgroup taxon, from Bayesian Inference analysis. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253945/files/figure.png" pageId="14" pageNumber="315">Figs. 1</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FF1890ECCE04FB28" box="[250,266,1210,1234]" captionStart="FIGURE 2" captionStartId="8.[151,250,1813,1835]" captionTargetBox="[151,1436,198,1786]" captionTargetId="figure@8.[151,1436,193,1792]" captionTargetPageId="8" captionText="FIGURE 2. COI gene tree for the 33 specimens of Devadatta also used in the analysis with other markers and one outgroup taxon. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253946/files/figure.png" pageId="14" pageNumber="315">2</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FEF490ECCE2BFB28" box="[278,293,1210,1234]" captionStart="FIGURE 3. 16 S" captionStartId="9.[151,250,1748,1770]" captionTargetBox="[159,1436,201,1719]" captionTargetId="figure@9.[151,1436,193,1727]" captionTargetPageId="9" captionText="FIGURE 3. 16 S gene tree for 33 specimens of Devadatta and one outgroup taxon, from Bayesian Inference analysis. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253947/files/figure.png" pageId="14" pageNumber="315">3</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FED090ECCE4CFB28" box="[306,322,1210,1234]" captionStart="FIGURE 4" captionStartId="10.[151,250,1769,1791]" captionTargetBox="[151,1436,198,1741]" captionTargetId="figure@10.[151,1436,193,1748]" captionTargetPageId="10" captionText="FIGURE 4. ITS gene tree for 33 specimens of Devadatta and one outgroup taxon, from Bayesian Inference analysis. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253949/files/figure.png" pageId="14" pageNumber="315">4</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FEAC90EFCE53FB28" box="[334,349,1209,1234]" captionStart="FIGURE 5. 28 S" captionStartId="11.[151,250,1830,1852]" captionTargetBox="[151,1436,200,1807]" captionTargetId="figure@11.[151,1436,193,1808]" captionTargetPageId="11" captionText="FIGURE 5. 28 S gene tree for 33 specimens of Devadatta and one outgroup taxon, from Bayesian Inference analysis. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253950/files/figure.png" pageId="14" pageNumber="315">5</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FE8890EFCE74FB28" box="[362,378,1209,1234]" captionStart="FIGURE 6" captionStartId="12.[151,250,1843,1865]" captionTargetBox="[157,1436,202,1816]" captionTargetId="figure@12.[151,1436,193,1821]" captionTargetPageId="12" captionText="FIGURE 6. Phylogenetic reconstruction for 33 specimens of Devadatta and one outgroup taxon from the combined COI + 16 S + ITS + 28 S data, using Bayesian Inference analysis. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253951/files/figure.png" pageId="14" pageNumber="315">6</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FE6490ECCE98FB2B" box="[390,406,1210,1233]" captionStart="FIGURE 7" captionStartId="13.[151,250,1743,1765]" captionTargetBox="[153,1436,202,1715]" captionTargetId="figure@13.[151,1436,193,1721]" captionTargetPageId="13" captionText="FIGURE 7. Maximum Likelihood best tree for 33 specimens of Devadatta and one outgroup taxon from the combined COI + 16 S + ITS + 28 S data set. Bootstrap support values below 100 are superimposed on the tree. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253952/files/figure.png" pageId="14" pageNumber="315">7</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FE4090ECCEBCFB28" box="[418,434,1210,1234]" captionStart="FIGURES 8 – 13" captionStartId="15.[151,264,1607,1629]" captionTargetBox="[151,1436,193,1585]" captionTargetId="figure@15.[151,1436,193,1586]" captionTargetPageId="15" captionText="FIGURES 8 – 13. Habitus of males of Devadatta species: (8) D. aran holotype; (9) D. basilanensis Davao City, Mindanao; (10) D. clavicauda holotype; (11) D. podolestoides male described in this paper; (12) D. somoh holotype; (13) D. tanduk holotype." httpUri="https://zenodo.org/record/253953/files/figure.png" pageId="14" pageNumber="315">8</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FE5C90ECCED2FB28" box="[446,476,1210,1234]" captionStart="FIGURES 14 – 19" captionStartId="16.[151,264,1803,1825]" captionTargetBox="[151,1436,733,1781]" captionTargetId="figure@16.[151,1436,733,1782]" captionTargetPageId="16" captionText="FIGURES 14 – 19. Heads of male Devadatta species: (14) D. aran holotype; (15) D. basilanensis Davao Oriental, Mindanao; (16) D. clavicauda holotype; (17) D. podolestoides male described in this paper; (18) D. somoh holotype; (19) D. tanduk holotype." httpUri="https://zenodo.org/record/253954/files/figure.png" pageId="14" pageNumber="315">14</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FE0A90ECCD08FB28" box="[488,518,1210,1234]" captionStart="FIGURES 20 – 25" captionStartId="17.[151,264,1919,1941]" captionTargetBox="[194,1386,193,1882]" captionTargetId="figure@17.[194,1387,193,1884]" captionTargetPageId="17" captionText="FIGURES 20 – 25. Synthorax of male Devadatta species, lateral view: (20) D. aran holotype; (21) D. basilanensis Davao Oriental, Mindanao; (22) D. clavicauda holotype; (23) D. podolestoides male described in this paper; (24) D. somoh holotype; (25) D. tanduk holotype." httpUri="https://zenodo.org/record/253955/files/figure.png" pageId="14" pageNumber="315">20</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FDF090EFCD3FFB28" box="[530,561,1209,1234]" captionStart="FIGURES 26 – 29" captionStartId="18.[151,264,1357,1379]" captionTargetBox="[151,1436,193,1335]" captionTargetId="figure@18.[151,1436,193,1335]" captionTargetPageId="18" captionText="FIGURES 26 – 29. Venter of synthorax of male Devadatta species: (26) D. aran holotype; (27) D. somoh holotype; (28) D. tanduk holotype; (29) D. tanduk paratype (SAB 12 _ AMP 29)." httpUri="https://zenodo.org/record/253956/files/figure.png" pageId="14" pageNumber="315">26</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FDDE90ECCD54FB28" box="[572,602,1210,1234]" captionStart="FIGURES 38 – 39" captionStartId="20.[151,264,1911,1933]" captionTargetBox="[151,1436,1261,1889]" captionTargetId="figure@20.[151,1436,1261,1889]" captionTargetPageId="20" captionText="FIGURES 38 – 39. Genital ligula of two Devadatta species, scale bars = 0.5 mm: (38) D. aran holotype; (39) D. podolestoides male described in this paper." httpUri="https://zenodo.org/record/253959/files/figure.png" pageId="14" pageNumber="315">38</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FD8590ECCD8BFB28" box="[615,645,1210,1234]" captionStart="FIGURES 40 – 45" captionStartId="21.[151,264,1699,1721]" captionTargetBox="[162,1425,203,1663]" captionTargetId="figure@21.[151,1436,193,1677]" captionTargetPageId="21" captionText="FIGURES 40 – 45. Male anal appendages, dorsal view: (40) D. aran holotype; (41) D. basilanensis Davao Oriental, Mindanao; (42) D. clavicauda holotype; (43) D. podolestoides Kubah NP, Sarawak; (44) D. somoh holotype; (45) D. tanduk holotype." httpUri="https://zenodo.org/record/253960/files/figure.png" pageId="14" pageNumber="315">40</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FD7290EFCDA0FB28" box="[656,686,1209,1234]" captionStart="FIGURES 46 – 51" captionStartId="22.[151,264,1951,1973]" captionTargetBox="[162,1418,393,1916]" captionTargetId="figure@22.[151,1436,373,1929]" captionTargetPageId="22" captionText="FIGURES 46 – 51. Male anal appendages, lateral view (scale bars 1 mm): (46) D. aran holotype; (47) D. basilanensis Davao Oriental, Mindanao; (48) D. clavicauda holotype; (49) D. podolestoides Kubah NP, Sarawak; (50) D. somoh holotype; (51) D. tanduk holotype." httpUri="https://zenodo.org/record/253961/files/figure.png" pageId="14" pageNumber="315">46</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FD5890EFCDD6FB28" box="[698,728,1209,1234]" captionStart="FIGURES 52 – 57" captionStartId="23.[151,264,1409,1431]" captionTargetBox="[151,1436,176,1394]" captionTargetId="figure@23.[151,1436,175,1395]" captionTargetPageId="23" captionText="FIGURES 52 – 57. Heads of Devadatta female specimens described in this paper: (52) D. aran; (53) D. basilanensis; (54) D. clavicauda; (55) D. podolestoides; (56) D. somoh; (57) D. tanduk." httpUri="https://zenodo.org/record/253962/files/figure.png" pageId="14" pageNumber="315">52</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FD0690EFCC0DFB28" box="[740,771,1209,1234]" captionStart="FIGURES 58 – 63" captionStartId="25.[151,264,1932,1954]" captionTargetBox="[151,1436,411,1910]" captionTargetId="figure@25.[151,1436,411,1910]" captionTargetPageId="25" captionText="FIGURES 58 – 63. Prothorax of Devadatta female specimens described in this paper, dorsal view: (58) D. aran; (59) D. basilanensis; (60) D. clavicauda; (61) D. podolestoides; (62) D. somoh; (63) D. tanduk." httpUri="https://zenodo.org/record/253963/files/figure.png" pageId="14" pageNumber="315">58</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FCED90EFCC58FB28" box="[783,854,1209,1234]" captionStart="FIGURES 64 – 67" captionStartId="26.[151,264,1535,1557]" captionTargetBox="[151,1436,193,1512]" captionTargetId="figure@26.[151,1436,193,1513]" captionTargetPageId="26" captionText="FIGURES 64 – 67. Prothorax of female Devadatta species with horns, lateral view: (64) D. aran female described in this paper; (65) D. aran Crocker Range, Sabah; (66) D. somoh female described in this paper; (67) D. tanduk female described in this paper." httpUri="https://zenodo.org/record/253964/files/figure.png" pageId="14" pageNumber="315">64, 65</figureCitation>
, 
<figureCitation id="DE992A6D5514FFB6FC8090EFCC8EFB28" box="[866,896,1209,1234]" captionStart="FIGURES 68 – 73" captionStartId="28.[151,264,1928,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@28.[151,1436,193,1907]" captionTargetPageId="28" captionText="FIGURES 68 – 73. Synthorax of Devadatta female specimens described in this paper, lateral view: (68) D. aran; (69) D. basilanensis; (70) D. clavicauda; (71) D. podolestoides; (72) D. somoh; (73) D. tanduk." httpUri="https://zenodo.org/record/253965/files/figure.png" pageId="14" pageNumber="315">68</figureCitation>
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<figureCitation id="DE992A6D5514FFB6FC6E90ECCCA7FB28" box="[908,937,1210,1234]" captionStart="FIGURE 74" captionStartId="29.[151,250,1608,1630]" captionTargetBox="[151,1436,193,1586]" captionTargetId="figure@29.[151,1436,193,1586]" captionTargetPageId="29" captionText="FIGURE 74. Distribution of D. aran (squares) and D. podolestoides (triangles)." httpUri="https://zenodo.org/record/253966/files/figure.png" pageId="14" pageNumber="315">74</figureCitation>
)
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<emphasis id="74D6EAFA5514FFB6FF7590ACCE7CFAEA" box="[151,370,1274,1296]" italics="true" pageId="14" pageNumber="315">Devadatta argyroides</emphasis>
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(sic) [nec (Selys, 1859)];— 
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<bibRefCitation id="22334B195514FFB6FD6B90ACCC13FAEA" author="Laidlaw" box="[649,797,1274,1296]" pageId="14" pageNumber="315" refString="Laidlaw, F. F. (1915) Contributions to a study of the dragonfly fauna of Borneo. Part III. A collection made on Mount Kina Balu by Mr J. C. Moulton in September and October 1913. Proceedings of the Zoological Society of London, 1915, 25 - 39." type="proceedings paper" year="1915">Laidlaw 1915</bibRefCitation>
: 25
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, 33 (Mount Kinabalu);— 
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<emphasis id="74D6EAFA5514FFB6FF75914FCE0FFAD5" box="[151,257,1305,1327]" italics="true" pageId="14" pageNumber="315">Devadatta</emphasis>
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species B;— 
<treatmentCitation id="C70310F95514FFB6FE67914CCDB5FAD5" author="Dow" box="[389,699,1305,1327]" page="9" pageId="14" pageNumber="315" year="2013">
<bibRefCitation id="22334B195514FFB6FE67914CCDAAFAD5" author="Dow" box="[389,676,1305,1327]" pageId="14" pageNumber="315" refString="Dow, R. A., Reels, G. T. &amp; Butler, S. G. (2013 a) Odonata of the Dulit Range in Sarawak, Malaysian Borneo. Notulae odonatologicae, 8 (1), 1 - 16." type="journal article" year="2013" yearSuffix="a">Dow, Reels &amp; Butler 2013a</bibRefCitation>
: 9
</treatmentCitation>
, 12 (Mount Dulit).
</treatmentCitationGroup>
</paragraph>
<paragraph id="461D36E85514FFB6FF75916ECEB3FA97" blockId="14.[151,1436,1274,1603]" pageId="14" pageNumber="315">
<treatmentCitationGroup id="66B211C65514FFB6FF75916ECEB3FA97" pageId="14" pageNumber="315">
<taxonomicName id="81A24D6B5514FFB6FF75916ECE0FFAB4" ID-CoL="43VM" box="[151,257,1336,1358]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA5514FFB6FF75916ECE0FFAB4" box="[151,257,1336,1358]" italics="true" pageId="14" pageNumber="315">Devadatta</emphasis>
</taxonomicName>
species C;— 
<treatmentCitation id="C70310F95514FFB6FE6B916ECD73FAB4" author="Dow" box="[393,637,1336,1358]" page="9" pageId="14" pageNumber="315" year="2014">
<bibRefCitation id="22334B195514FFB6FE6B916ECD6AFAB4" author="Dow" box="[393,612,1336,1358]" pageId="14" pageNumber="315" refString="Dow, R. A. &amp; Ngiam, R. W. J. (2014) Odonata from logged and unlogged forest in the Ulu Balui and Ulu Baleh, Kapit Division, Sarawak, in June and September 2013. International Dragonfly Fund Report, 73, 1 - 48." type="journal article" year="2014">Dow &amp; Ngiam 2014</bibRefCitation>
: 9
</treatmentCitation>
, 42, 44 (Ulu Balui, Ulu Baleh);— 
<treatmentCitation id="C70310F95514FFB6FC3D916ECBB2FAB4" author="Dow" box="[991,1212,1336,1358]" page="22" pageId="14" pageNumber="315" year="2015">
<bibRefCitation id="22334B195514FFB6FC3D916ECB97FAB4" author="Dow" box="[991,1177,1336,1358]" pageId="14" pageNumber="315" refString="Dow, R. A., Reels, G. T. &amp; Ngiam, R. W. J. (2015 a) Previously unpublished Odonata records from Sarawak, Borneo, Part III. Sri Aman, Sibu and Kapit Divisions. Faunistic Studies in South-East Asian and Pacific Island Odonata, 9, 1 - 34." type="journal article" year="2015" yearSuffix="a">
Dow 
<emphasis id="74D6EAFA5514FFB6FBF8916FCB46FAB4" box="[1050,1096,1336,1358]" italics="true" pageId="14" pageNumber="315">et al</emphasis>
. 2015a
</bibRefCitation>
: 22
</treatmentCitation>
(Kapit);— 
<treatmentCitation id="C70310F95514FFB6FACF916ECE22FA96" author="Dow" page="11" pageId="14" pageNumber="315" year="2015">
<bibRefCitation id="22334B195514FFB6FACF916ECE04FA96" author="Dow" pageId="14" pageNumber="315" refString="Dow, R. A., Reels, G. T. &amp; Ngiam, R. W. J. (2015 b) Odonata collected at Usun Apau National Park, Miri Division, Sarawak, Malaysia in April and May 2012. International Dragonfly Fund Report, 79, 1 - 17." type="journal article" year="2015" yearSuffix="b">
Dow 
<emphasis id="74D6EAFA5514FFB6FA85916FCA98FAB4" box="[1383,1430,1336,1358]" italics="true" pageId="14" pageNumber="315">et al</emphasis>
. 2015b
</bibRefCitation>
: 11
</treatmentCitation>
(Usun Apau).
</treatmentCitationGroup>
</paragraph>
<paragraph id="461D36E85514FFB6FF759123CA9EFA71" blockId="14.[151,1436,1274,1603]" box="[151,1424,1397,1419]" pageId="14" pageNumber="315">
<treatmentCitationGroup id="66B211C65514FFB6FF759123CA9EFA71" box="[151,1424,1397,1419]" pageId="14" pageNumber="315">
<taxonomicName id="81A24D6B5514FFB6FF759123CE0FFA71" ID-CoL="43VM" box="[151,257,1397,1419]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA5514FFB6FF759123CE0FFA71" box="[151,257,1397,1419]" italics="true" pageId="14" pageNumber="315">Devadatta</emphasis>
</taxonomicName>
sp.;— 
<treatmentCitation id="C70310F95514FFB6FEDD9123CD1EFA71" author="Laidlaw" box="[319,528,1397,1419]" page="550" pageId="14" pageNumber="315" year="1934">
<bibRefCitation id="22334B195514FFB6FEDD9123CEEEFA71" author="Laidlaw" box="[319,480,1397,1419]" pageId="14" pageNumber="315" refString="Laidlaw, F. F. (1934 b) A note on the dragonfly fauna (Odonata) of Mount Kinabalu and of some other mountain areas of Malaysia: with a description of some new or little known species. Journal of the Federated Malay States Museums, 17 (3), 549 - 561." type="journal article" year="1934" yearSuffix="b">Laidlaw 1934b</bibRefCitation>
: 550
</treatmentCitation>
(Part: Mount Kinabalu);— 
<bibRefCitation id="22334B195514FFB6FCC29123CCE2FA71" author="Dijkstra" box="[800,1004,1397,1419]" pageId="14" pageNumber="315" refString="Dijkstra, K. - D. B., Kalkman, V. J., Dow, R. A., Stokvis, F. R. &amp; van Tol, J. (2014) Redefining the damselfly families: the first comprehensive molecular phylogeny of Zygoptera (Odonata). Systematic Entomology, 39 (1), 68 - 96. http: // dx. doi. org / 10.1111 / syen. 12035" type="journal article" year="2014">
Dijkstra 
<emphasis id="74D6EAFA5514FFB6FC9E9120CCA1FA71" box="[892,943,1397,1419]" italics="true" pageId="14" pageNumber="315">et al.</emphasis>
2014
</bibRefCitation>
: Table 5 (
<collectingCountry id="3EB576785514FFB6FBAF9123CBA4FA71" box="[1101,1194,1397,1419]" name="Malaysia" pageId="14" pageNumber="315">Sarawak</collectingCountry>
: RMNH.INS.503484).
</treatmentCitationGroup>
</paragraph>
<paragraph id="461D36E85514FFB6FF7591C2CBBBFA32" blockId="14.[151,1436,1274,1603]" pageId="14" pageNumber="315">
<treatmentCitationGroup id="66B211C65514FFB6FF7591C2CBBBFA32" pageId="14" pageNumber="315">
<taxonomicName id="81A24D6B5514FFB6FF7591C2CE9FFA50" ID-CoL="6CPLH" box="[151,401,1428,1450]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA5514FFB6FF7591C2CE9FFA50" box="[151,401,1428,1450]" italics="true" pageId="14" pageNumber="315">Devadatta podolestoides</emphasis>
</taxonomicName>
[nec Laidlaw, 1934];— 
<treatmentCitation id="C70310F95514FFB6FD9E91C2CC33FA50" author="Kimmins" box="[636,829,1428,1450]" page="79" pageId="14" pageNumber="315" year="1936">
<bibRefCitation id="22334B195514FFB6FD9E91C2CC14FA50" author="Kimmins" box="[636,794,1428,1450]" pageId="14" pageNumber="315" refString="Kimmins, D. E. (1936) The Odonata of the Oxford University Sarawak expedition. Journal of the Federated Malay States Museum, 18, 65 - 108." type="journal article" year="1936">Kimmins 1936</bibRefCitation>
: 79
</treatmentCitation>
(Mount Dulit: part);— 
<treatmentCitation id="C70310F95514FFB6FBC691C3CBBBFA50" author="Dow" box="[1060,1205,1428,1450]" page="10" pageId="14" pageNumber="315" year="2006">
<bibRefCitation id="22334B195514FFB6FBC691C3CB9CFA50" author="Dow" box="[1060,1170,1428,1450]" pageId="14" pageNumber="315" refString="Dow, R. A. (2006) Fishing the road - Looking for Odonata in Sarawak's Kelabit Highlands. Agrion, 10 (1), 10." type="journal article" year="2006">Dow 2006</bibRefCitation>
: 10
</treatmentCitation>
(Tama Abu Range);— van 
<treatmentCitation id="C70310F95514FFB6FF1091E4CE7CFA32" author="Tol" box="[242,370,1458,1480]" page="13" pageId="14" pageNumber="315" year="2006">
<bibRefCitation id="22334B195514FFB6FF1091E4CE5EFA32" author="Tol" box="[242,336,1458,1480]" pageId="14" pageNumber="315" refString="van Tol, J. (2006) Odonata. In: van Tol, J. (Ed.), Gunung Lumut Biodiversity Assessment. National Museum of Natural History Naturalis, Leiden, pp. 12 - 15. Available from: https: // science. naturalis. nl / media / medialibrary / 2013 / 08 / 2005 _ gunung _ lumu t _ report _ pdf. pdf (accessed 15 September 2015)" type="journal article" year="2006">Tol 2006</bibRefCitation>
: 13
</treatmentCitation>
(Gunung Lumut, East Kalimantan);— 
<treatmentCitation id="C70310F95514FFB6FD1891E4CCEDFA32" author="Dow" box="[762,995,1458,1480]" page="13" pageId="14" pageNumber="315" year="2009">
<bibRefCitation id="22334B195514FFB6FD1891E4CCCFFA32" author="Dow" box="[762,961,1458,1480]" pageId="14" pageNumber="315" refString="Dow, R. A. &amp; Reels, G. T. (2009) Expedition to Mount Dulit, Sarawak, August - September 2008 - Odonata. International Dragonfly Fund Report, 19, 1 - 16." type="journal article" year="2009">Dow &amp; Reels 2009</bibRefCitation>
: 13
</treatmentCitation>
(Mount Dulit: part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="461D36E85514FFB6FF759187CE28F9FC" blockId="14.[151,1436,1274,1603]" pageId="14" pageNumber="315">
<treatmentCitationGroup id="66B211C65514FFB6FF759187CE28F9FC" pageId="14" pageNumber="315">
<taxonomicName id="81A24D6B5514FFB6FF759187CD2BFA1D" ID-CoL="6CPLH" box="[151,549,1489,1511]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="subSpecies" species="podolestoides" subSpecies="podolestoides">
<emphasis id="74D6EAFA5514FFB6FF759187CD2BFA1D" box="[151,549,1489,1511]" italics="true" pageId="14" pageNumber="315">Devadatta podolestoides podolestoides</emphasis>
</taxonomicName>
[nec Laidlaw, 1934];— 
<treatmentCitation id="C70310F95514FFB6FCF69187CCC5FA1D" author="Lieftinck" box="[788,971,1489,1511]" page="2" pageId="14" pageNumber="315" year="1954">
<bibRefCitation id="22334B195514FFB6FCF69187CCBDFA1D" author="Lieftinck" box="[788,947,1489,1511]" pageId="14" pageNumber="315" refString="Lieftinck, M. A. (1954) Handlist of Malaysian Odonata. A catalogue of the dragonflies of the Malay Peninsula, Sumatra, Java and Borneo, including the adjacent small islands. Treubia (Suppl.), 22, i - xiii + 1 - 202, 1 folded map excl." type="journal article" year="1954">Lieftinck 1954</bibRefCitation>
: 2
</treatmentCitation>
(part: Mount Kinabalu);— 
<treatmentCitation id="C70310F95514FFB6FB3C9187CFEEF9FC" author="Hamalainen" page="79" pageId="14" pageNumber="315" year="1994">
<bibRefCitation id="22334B195514FFB6FB3C9187CA96FA1D" author="Hamalainen" box="[1246,1432,1489,1511]" pageId="14" pageNumber="315" refString="Hamalainen, M. (1994) Dragonflies of Mount Kinabalu (the highest mountain in Borneo). Malangpo, 11, 77 - 81." type="journal article" year="1994">Hämäläinen 1994</bibRefCitation>
: 79
</treatmentCitation>
(part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="461D36E85514FFB6FF759258CCF7F9DE" blockId="14.[151,1436,1274,1603]" box="[151,1017,1550,1573]" pageId="14" pageNumber="315">
<taxonomicName id="81A24D6B5514FFB6FF759258CE0FF9DE" ID-CoL="43VM" box="[151,257,1550,1572]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA5514FFB6FF759258CE0FF9DE" box="[151,257,1550,1572]" italics="true" pageId="14" pageNumber="315">Devadatta</emphasis>
</taxonomicName>
cf 
<taxonomicName id="81A24D6B5514FFB6FEC39258CEA2F9DE" ID-CoL="6CPLH" box="[289,428,1550,1572]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA5514FFB6FEC39258CEA2F9DE" box="[289,428,1550,1572]" italics="true" pageId="14" pageNumber="315">podolestoides</emphasis>
</taxonomicName>
;— 
<bibRefCitation id="22334B195514FFB6FE2B9258CD9DF9DE" author="Dijkstra" box="[457,659,1550,1573]" pageId="14" pageNumber="315" refString="Dijkstra, K. - D. B., Kalkman, V. J., Dow, R. A., Stokvis, F. R. &amp; van Tol, J. (2014) Redefining the damselfly families: the first comprehensive molecular phylogeny of Zygoptera (Odonata). Systematic Entomology, 39 (1), 68 - 96. http: // dx. doi. org / 10.1111 / syen. 12035" type="journal article" year="2014">
Dijkstra 
<emphasis id="74D6EAFA5514FFB6FDC69246CD59F9DE" box="[548,599,1550,1573]" italics="true" pageId="14" pageNumber="315">et al.</emphasis>
2014
</bibRefCitation>
: 
<figureCitation id="DE992A6D5514FFB6FD7E9258CDECF9DE" box="[668,738,1550,1572]" captionStart="FIGURE 2" captionStartId="8.[151,250,1813,1835]" captionTargetBox="[151,1436,198,1786]" captionTargetId="figure@8.[151,1436,193,1792]" captionTargetPageId="8" captionText="FIGURE 2. COI gene tree for the 33 specimens of Devadatta also used in the analysis with other markers and one outgroup taxon. Posterior probability values are shown (as percentages) if less than 100 %. RMNH collection codes are shown for each specimen, with the RMNH. INS. prefix omitted for clarity." httpUri="https://zenodo.org/record/253946/files/figure.png" pageId="14" pageNumber="315">Figs. 2</figureCitation>
b, 5b (RMNH.INS.503484)
</paragraph>
<paragraph id="461D36E85514FFB6FF75927BCCF8F9B9" blockId="14.[151,1436,1274,1603]" box="[151,1014,1581,1603]" pageId="14" pageNumber="315">
<treatmentCitationGroup id="66B211C65514FFB6FF75927BCCF8F9B9" box="[151,1014,1581,1603]" pageId="14" pageNumber="315">
<taxonomicName id="81A24D6B5514FFB6FF75927BCE0FF9B9" ID-CoL="43VM" box="[151,257,1581,1603]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA5514FFB6FF75927BCE0FF9B9" box="[151,257,1581,1603]" italics="true" pageId="14" pageNumber="315">Devadatta</emphasis>
</taxonomicName>
spp. cf 
<taxonomicName id="81A24D6B5514FFB6FEB2927BCED5F9B9" ID-CoL="6CPLH" box="[336,475,1581,1603]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA5514FFB6FEB2927BCED5F9B9" box="[336,475,1581,1603]" italics="true" pageId="14" pageNumber="315">podolestoides</emphasis>
</taxonomicName>
;— 
<treatmentCitation id="C70310F95514FFB6FE189278CDE9F9B9" author="Dow" box="[506,743,1581,1603]" page="2" pageId="14" pageNumber="315" year="2012">
<bibRefCitation id="22334B195514FFB6FE189278CDDEF9B9" author="Dow" box="[506,720,1581,1603]" pageId="14" pageNumber="315" refString="Dow, R. A. &amp; Ngiam, R. W. J. (2012) Odonata collected in the Hose Mountains, Kapit Division, Sarawak, Malaysia in April 2011. International Dragonfly Fund Report, 44, 1 - 18." type="journal article" year="2012">Dow &amp; Ngiam 2012</bibRefCitation>
: 2
</treatmentCitation>
, 7 (Hose mountains: part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0EB865635514FFB6FF759239CB0CF90F" pageId="14" pageNumber="315" type="materials_examined">
<paragraph id="461D36E85514FFB6FF759239CA5AF956" blockId="14.[151,1436,1647,2033]" pageId="14" pageNumber="315">
<emphasis id="74D6EAFA5514FFB6FF759239CEC9F972" bold="true" box="[151,455,1647,1672]" pageId="14" pageNumber="315">
<typeStatus id="9919884A5514FFB6FF759239CFDCF972" box="[151,210,1647,1672]" pageId="14" pageNumber="315">Type</typeStatus>
material. 
<typeStatus id="9919884A5514FFB6FEB39239CECCF972" box="[337,450,1647,1672]" pageId="14" pageNumber="315" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCount id="50A4FD615514FFB6FE329226CEF3F97D" box="[464,509,1648,1672]" pageId="14" pageNumber="315" type="generic">1 ♂</specimenCount>
(SAR11_12_ 
<collectionCode id="20B3AE2D5514FFB6FD759226CDD6F972" box="[663,728,1648,1672]" httpUri="http://grbio.org/cool/90v0-7gw3" name="Australian Mycological Panel" pageId="14" pageNumber="315">AMP</collectionCode>
74, 
<collectionCode id="20B3AE2D5514FFB6FCE09239CC52F972" LSID="urn:lsid:biocol.org:col:34992" box="[770,860,1647,1672]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="315">RMNH</collectionCode>
.INS.506316), Pa’Gelawat and tributaries at 
<quantity id="815A9B0D5514FFB6FAB49226CFE9F957" metricMagnitude="3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="14" pageNumber="315" unit="m" value="1450.0" valueMax="1500.0" valueMin="1400.0">1400– 1500m</quantity>
, Pulong Tau National Park, Miri division, 
<collectingCountry id="3EB576785514FFB6FD2192C2CC28F957" box="[707,806,1684,1709]" name="Malaysia" pageId="14" pageNumber="315">Sarawak</collectingCountry>
, 
<collectingCountry id="3EB576785514FFB6FCD092C2CC92F957" box="[818,924,1684,1709]" name="Malaysia" pageId="14" pageNumber="315">Malaysia</collectingCountry>
, 
<date id="321C10285514FFB6FC4492C2CB2FF956" box="[934,1057,1684,1709]" pageId="14" pageNumber="315" value="2012-04-16">16 iv 2012</date>
, leg. R.A. Dow, in 
<collectionCode id="20B3AE2D5514FFB6FB1A92C3CA5EF956" LSID="urn:lsid:biocol.org:col:34992" box="[1272,1360,1685,1708]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="315">RMNH</collectionCode>
.
</paragraph>
<paragraph id="461D36E85514FFB6FF2592E1CB0CF90F" blockId="14.[151,1436,1647,2033]" pageId="14" pageNumber="315">
<typeStatus id="9919884A5514FFB6FF2592E1CE31F92A" box="[199,319,1719,1744]" pageId="14" pageNumber="315" type="paratype">
<emphasis id="74D6EAFA5514FFB6FF2592E1CE31F92A" bold="true" box="[199,319,1719,1744]" pageId="14" pageNumber="315">Paratypes</emphasis>
</typeStatus>
(
<specimenCount id="50A4FD615514FFB6FEB292EECE97F935" box="[336,409,1720,1744]" pageId="14" pageNumber="315" type="generic">139 ♂</specimenCount>
30 ♀). Female used in description: 1 ♀ (SAR06_ 
<collectionCode id="20B3AE2D5514FFB6FC3B92E1CB17F92A" box="[985,1049,1719,1744]" httpUri="http://grbio.org/cool/90v0-7gw3" name="Australian Mycological Panel" pageId="14" pageNumber="315">AMP</collectionCode>
21), trail to waterfall, Bario, Tama Abu Range, Miri division, 
<date id="321C10285514FFB6FE27928ACD4FF90F" box="[453,577,1756,1781]" pageId="14" pageNumber="315" value="2006-03-14">14 iii 2006</date>
, leg. RAD. See appendix for remainder.
</paragraph>
</subSubSection>
<subSubSection id="0EB865635514FFB6FF2592A9CEF9F89A" pageId="14" pageNumber="315" type="etymology">
<paragraph id="461D36E85514FFB6FF2592A9CEF9F89A" blockId="14.[151,1436,1647,2033]" pageId="14" pageNumber="315">
<emphasis id="74D6EAFA5514FFB6FF2592A9CE40F8E2" bold="true" box="[199,334,1791,1816]" pageId="14" pageNumber="315">Etymology.</emphasis>
<taxonomicName id="81A24D6B5514FFB6FEB59357CE82F8E2" box="[343,396,1793,1816]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="74D6EAFA5514FFB6FEB59357CE82F8E2" box="[343,396,1793,1816]" italics="true" pageId="14" pageNumber="315">aran</emphasis>
</taxonomicName>
, a noun in apposition; named in memory of the late Reddish (pronounced Raddish) 
<taxonomicName id="81A24D6B5514FFB6FA869356CA92F8E2" box="[1380,1436,1792,1816]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="344" phylum="Arthropoda" rank="species" species="aran">Aran</taxonomicName>
(born 
<date id="321C10285514FFB6FF389372CE51F8C7" box="[218,351,1828,1853]" pageId="14" pageNumber="315" value="1963-07-06">6 July 1963</date>
, died 
<date id="321C10285514FFB6FE439372CD75F8C6" box="[417,635,1828,1853]" pageId="14" pageNumber="315" value="2012-11-27">27 November 2012</date>
) of Bario in the Tama Abu Range, who accompanied the first author on his first trip to the 
<typeStatus id="9919884A5514FFB6FE84931ECE98F89A" box="[358,406,1864,1888]" pageId="14" pageNumber="315">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="0EB865635514FFB6FF25933ACCC9F80A" pageId="14" pageNumber="315" type="diagnosis">
<paragraph id="461D36E85514FFB6FF25933ACCC9F80A" blockId="14.[151,1436,1647,2033]" pageId="14" pageNumber="315">
<emphasis id="74D6EAFA5514FFB6FF25933ACE4FF87F" bold="true" box="[199,321,1900,1925]" pageId="14" pageNumber="315">Diagnosis.</emphasis>
A dark coloured 
<taxonomicName id="81A24D6B5514FFB6FDF6933ACD87F87E" box="[532,649,1900,1924]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA5514FFB6FDF6933ACD87F87E" box="[532,649,1900,1924]" italics="true" pageId="14" pageNumber="315">Devadatta</emphasis>
</taxonomicName>
species, in most known populations it can be separated from other species from the 
<taxonomicName id="81A24D6B5514FFB6FEBA93C6CEFCF852" box="[344,498,1936,1960]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA5514FFB6FEBA93C6CEFCF852" box="[344,498,1936,1960]" italics="true" pageId="14" pageNumber="315">podolestoides</emphasis>
</taxonomicName>
-group by having the venter of the synthorax mostly pale in both sexes and at least vestigial, but not long, horns on the middle pronotal lobe in females. Additionally, easily separated from 
<taxonomicName id="81A24D6B5514FFB6FA6393E3CE1CF80A" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="74D6EAFA5514FFB6FA6393E3CE1CF80A" italics="true" pageId="14" pageNumber="315">D. clavicauda</emphasis>
</taxonomicName>
by the less expanded ends of the cerci in the male of 
<taxonomicName id="81A24D6B5514FFB6FC8B938FCCCEF80A" box="[873,960,2009,2032]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="74D6EAFA5514FFB6FC8B938FCCCEF80A" box="[873,960,2009,2032]" italics="true" pageId="14" pageNumber="315">D. aran</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="12DD66605515FFB7FF759211CA9DF986" httpUri="https://zenodo.org/record/253953/files/figure.png" pageId="15" pageNumber="316" targetBox="[151,1436,193,1585]" targetPageId="15">
<paragraph id="461D36E85515FFB7FF759211CA9DF986" blockId="15.[151,1436,1607,1660]" pageId="15" pageNumber="316">
<emphasis id="74D6EAFA5515FFB7FF759211CE49F9A7" bold="true" box="[151,327,1607,1629]" pageId="15" pageNumber="316">FIGURES 8–13.</emphasis>
Habitus of males of 
<taxonomicName id="81A24D6B5515FFB7FDFA9211CD8FF9A7" box="[536,641,1607,1629]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA5515FFB7FDFA9211CD8FF9A7" box="[536,641,1607,1629]" italics="true" pageId="15" pageNumber="316">Devadatta</emphasis>
</taxonomicName>
species: (8) 
<taxonomicName id="81A24D6B5515FFB7FCE3921ECC41F9A7" box="[769,847,1608,1629]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="74D6EAFA5515FFB7FCE3921ECC41F9A7" box="[769,847,1608,1629]" italics="true" pageId="15" pageNumber="316">D. aran</emphasis>
</taxonomicName>
holotype; (9) 
<taxonomicName id="81A24D6B5515FFB7FC3C921ECB74F9A7" box="[990,1146,1607,1629]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="74D6EAFA5515FFB7FC3C921ECB74F9A7" box="[990,1146,1607,1629]" italics="true" pageId="15" pageNumber="316">D. basilanensis</emphasis>
</taxonomicName>
Davao City, Mindanao; (10) 
<taxonomicName id="81A24D6B5515FFB7FF759231CE2BF986" box="[151,293,1638,1660]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="74D6EAFA5515FFB7FF759231CE2BF986" box="[151,293,1638,1660]" italics="true" pageId="15" pageNumber="316">D. clavicauda</emphasis>
</taxonomicName>
holotype; (11) 
<taxonomicName id="81A24D6B5515FFB7FE239231CD65F986" box="[449,619,1638,1660]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA5515FFB7FE239231CD65F986" box="[449,619,1638,1660]" italics="true" pageId="15" pageNumber="316">D. podolestoides</emphasis>
</taxonomicName>
male described in this paper; (12) 
<taxonomicName id="81A24D6B5515FFB7FC299231CB22F986" box="[971,1068,1638,1660]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="74D6EAFA5515FFB7FC299231CB22F986" box="[971,1068,1638,1660]" italics="true" pageId="15" pageNumber="316">D. somoh</emphasis>
</taxonomicName>
holotype; (13) 
<taxonomicName id="81A24D6B5515FFB7FB2B9231CA23F986" box="[1225,1325,1638,1660]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="74D6EAFA5515FFB7FB2B9231CA23F986" box="[1225,1325,1638,1660]" italics="true" pageId="15" pageNumber="316">D. tanduk</emphasis>
</taxonomicName>
holotype.
</paragraph>
</caption>
<subSubSection id="0EB865635515FFAEFF2592F5CCBCFEBA" lastPageId="22" lastPageNumber="323" pageId="15" pageNumber="316" type="description">
<paragraph id="461D36E85515FFB7FF2592F5CCEAF8D2" blockId="15.[151,1437,1699,2012]" pageId="15" pageNumber="316">
<emphasis id="74D6EAFA5515FFB7FF2592F5CD20F946" bold="true" box="[199,558,1699,1724]" pageId="15" pageNumber="316">
Description of 
<typeStatus id="9919884A5515FFB7FE9E92F5CEEDF946" box="[380,483,1699,1724]" pageId="15" pageNumber="316" type="holotype">holotype</typeStatus>
male.
</emphasis>
Head: Labium mostly pale. Mandible bases and genae shining dark brown. Labrum, clypeus, frons and vertex to level of lateral ocelli black with metallic green reflection, remainder of dorsum of head matte black with indistinct pale area on protruding postocular lobes; no pale band joining these areas (
<figureCitation id="DE992A6D5515FFB7FF029359CE3DF8DD" box="[224,307,1807,1832]" captionStart="FIGURES 14 – 19" captionStartId="16.[151,264,1803,1825]" captionTargetBox="[151,1436,733,1781]" captionTargetId="figure@16.[151,1436,733,1782]" captionTargetPageId="16" captionText="FIGURES 14 – 19. Heads of male Devadatta species: (14) D. aran holotype; (15) D. basilanensis Davao Oriental, Mindanao; (16) D. clavicauda holotype; (17) D. podolestoides male described in this paper; (18) D. somoh holotype; (19) D. tanduk holotype." httpUri="https://zenodo.org/record/253954/files/figure.png" pageId="15" pageNumber="316">Fig. 14</figureCitation>
); underside extensively pale. Antennae almost entirely black.
</paragraph>
<paragraph id="461D36E85515FFA8FF259365CE62FE9F" blockId="15.[151,1437,1699,2012]" lastBlockId="16.[151,1437,151,680]" lastPageId="16" lastPageNumber="317" pageId="15" pageNumber="316">
Thorax: Pronotum and propleuron largely black, with pale streak below notopleural suture, pale lateral marks on anterior pronotal lobe, rear of middle pronotal lobe, laterally on posterior pronotal lobe. Dorsum of synthorax matte black with small pale mark between arms of anterior bifurcation of middorsal carina. Synthorax laterally with bright pale marks as follows (
<figureCitation id="DE992A6D5515FFB7FDC193C9CD77F84D" box="[547,633,1951,1976]" captionStart="FIGURES 20 – 25" captionStartId="17.[151,264,1919,1941]" captionTargetBox="[194,1386,193,1882]" captionTargetId="figure@17.[194,1387,193,1884]" captionTargetPageId="17" captionText="FIGURES 20 – 25. Synthorax of male Devadatta species, lateral view: (20) D. aran holotype; (21) D. basilanensis Davao Oriental, Mindanao; (22) D. clavicauda holotype; (23) D. podolestoides male described in this paper; (24) D. somoh holotype; (25) D. tanduk holotype." httpUri="https://zenodo.org/record/253955/files/figure.png" pageId="15" pageNumber="316">Fig. 20</figureCitation>
): along lower side of mesopleural suture a long streak and short streak near antealar carina, longer streak running whole length of interpleural suture, larger subtriangular area and smaller mark under metapleural suture, joined by fainter pale marking. Area between metapleural carina and venter pale. Venter of synthorax entirely pale with areas of greyish shadowing (
<figureCitation id="DE992A6D550AFFA8FC6794EACCD7FF2F" box="[901,985,188,213]" captionStart="FIGURES 26 – 29" captionStartId="18.[151,264,1357,1379]" captionTargetBox="[151,1436,193,1335]" captionTargetId="figure@18.[151,1436,193,1335]" captionTargetPageId="18" captionText="FIGURES 26 – 29. Venter of synthorax of male Devadatta species: (26) D. aran holotype; (27) D. somoh holotype; (28) D. tanduk holotype; (29) D. tanduk paratype (SAB 12 _ AMP 29)." httpUri="https://zenodo.org/record/253956/files/figure.png" pageId="16" pageNumber="317">Fig. 26</figureCitation>
). Legs (posterior pair removed for DNA extraction): Coxae and trochanters pale with brown areas. Femora obscurely brown and grey. Tibiae and tarsi dark brown. Wings with arculus ca level of Ax 4 (
<quantity id="815A9B0D550AFFA8FD7B9552CDC8FEE7" box="[665,710,260,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="16" pageNumber="317" unit="in" value="5.0">5 in</quantity>
left Hw), quadrilateral with 1 crossvein in Fw, 
<quantity id="815A9B0D550AFFA8FB019552CA01FEE7" box="[1251,1295,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="16" pageNumber="317" unit="in" value="2.0">2 in</quantity>
Hw, cubitoanal space free in all wings, pt brown. Wing membrane with brownish tint, extreme tips of both wings with slightly heavier brown tint.
</paragraph>
<paragraph id="461D36E8550AFFA8FF259539CE16FD7F" blockId="16.[151,1437,151,680]" pageId="16" pageNumber="317">
Abdomen: Black above and very dark brown laterally, becoming almost entirely black on terminal 3 segments, S3–7 with broad pale basal ring, brown dorsally, small remnant of the basal ring present laterally on S8. Genital ligula (
<figureCitation id="DE992A6D550AFFA8FF0F95E1CE48FE2A" box="[237,326,439,464]" captionStart="FIGURES 38 – 39" captionStartId="20.[151,264,1911,1933]" captionTargetBox="[151,1436,1261,1889]" captionTargetId="figure@20.[151,1436,1261,1889]" captionTargetPageId="20" captionText="FIGURES 38 – 39. Genital ligula of two Devadatta species, scale bars = 0.5 mm: (38) D. aran holotype; (39) D. podolestoides male described in this paper." httpUri="https://zenodo.org/record/253959/files/figure.png" pageId="16" pageNumber="317">Fig. 38</figureCitation>
) typical for 
<taxonomicName id="81A24D6B550AFFA8FE0495EECD8EFE2A" box="[486,640,440,464]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA550AFFA8FE0495EECD8EFE2A" box="[486,640,440,464]" italics="true" pageId="16" pageNumber="317">podolestoides</emphasis>
</taxonomicName>
-group, seminal vesicle brown. Anal appendages black. Paraprocts extremely short, rounded, dark basally, paler apically. Cerci longer than S10, in dorsal view (
<figureCitation id="DE992A6D550AFFA8FB35958ACA23FE0E" box="[1239,1325,476,501]" captionStart="FIGURES 40 – 45" captionStartId="21.[151,264,1699,1721]" captionTargetBox="[162,1425,203,1663]" captionTargetId="figure@21.[151,1436,193,1677]" captionTargetPageId="21" captionText="FIGURES 40 – 45. Male anal appendages, dorsal view: (40) D. aran holotype; (41) D. basilanensis Davao Oriental, Mindanao; (42) D. clavicauda holotype; (43) D. podolestoides Kubah NP, Sarawak; (44) D. somoh holotype; (45) D. tanduk holotype." httpUri="https://zenodo.org/record/253960/files/figure.png" pageId="16" pageNumber="317">Fig. 40</figureCitation>
) abruptly curved inward at ca mid length, apical half at just more than a right angle to proximal half, left cercus lying over right. In both dorsal and lateral (
<figureCitation id="DE992A6D550AFFA8FDF09672CD69FDC7" box="[530,615,548,573]" captionStart="FIGURES 46 – 51" captionStartId="22.[151,264,1951,1973]" captionTargetBox="[162,1418,393,1916]" captionTargetId="figure@22.[151,1436,373,1929]" captionTargetPageId="22" captionText="FIGURES 46 – 51. Male anal appendages, lateral view (scale bars 1 mm): (46) D. aran holotype; (47) D. basilanensis Davao Oriental, Mindanao; (48) D. clavicauda holotype; (49) D. podolestoides Kubah NP, Sarawak; (50) D. somoh holotype; (51) D. tanduk holotype." httpUri="https://zenodo.org/record/253961/files/figure.png" pageId="16" pageNumber="317">Fig. 46</figureCitation>
) views widest at base, narrowing towards inward turn then moderately expanded in apical half. Interior surface with deep slot-like excavation around middle part, brownish inside excavation.
</paragraph>
<paragraph id="461D36E8550AFFA8FF2596D9CB68FD52" blockId="16.[151,1437,151,680]" box="[199,1126,655,680]" pageId="16" pageNumber="317">
<emphasis id="74D6EAFA550AFFA8FF2596D9CE78FD52" bold="true" box="[199,374,655,680]" pageId="16" pageNumber="317">Measurements</emphasis>
[mm]: Abdomen without anal appendages 39.5, cerci ca 1, Hw 34.
</paragraph>
<caption id="12DD6660550AFFA8FF75935DCFF9F8A5" httpUri="https://zenodo.org/record/253954/files/figure.png" pageId="16" pageNumber="317" targetBox="[151,1436,733,1781]" targetPageId="16">
<paragraph id="461D36E8550AFFA8FF75935DCFF9F8A5" blockId="16.[151,1436,1803,1887]" pageId="16" pageNumber="317">
<emphasis id="74D6EAFA550AFFA8FF75935DCE5AF8DB" bold="true" box="[151,340,1803,1825]" pageId="16" pageNumber="317">FIGURES 14–19.</emphasis>
Heads of male 
<taxonomicName id="81A24D6B550AFFA8FE15935DCD6EF8DB" box="[503,608,1803,1825]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="317" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA550AFFA8FE15935DCD6EF8DB" box="[503,608,1803,1825]" italics="true" pageId="16" pageNumber="317">Devadatta</emphasis>
</taxonomicName>
species: (14) 
<taxonomicName id="81A24D6B550AFFA8FD13935ACC4EF8DB" box="[753,832,1804,1825]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="74D6EAFA550AFFA8FD13935ACC4EF8DB" box="[753,832,1804,1825]" italics="true" pageId="16" pageNumber="317">D. aran</emphasis>
</taxonomicName>
holotype; (15) 
<taxonomicName id="81A24D6B550AFFA8FC02935ACB73F8DB" box="[992,1149,1803,1825]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="74D6EAFA550AFFA8FC02935ACB73F8DB" box="[992,1149,1803,1825]" italics="true" pageId="16" pageNumber="317">D. basilanensis</emphasis>
</taxonomicName>
Davao Oriental, Mindanao; (16) 
<taxonomicName id="81A24D6B550AFFA8FF29937DCE52F8BA" box="[203,348,1834,1856]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="74D6EAFA550AFFA8FF29937DCE52F8BA" box="[203,348,1834,1856]" italics="true" pageId="16" pageNumber="317">D. clavicauda</emphasis>
</taxonomicName>
holotype; (17) 
<taxonomicName id="81A24D6B550AFFA8FDE1937DCDBFF8BA" box="[515,689,1834,1856]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA550AFFA8FDE1937DCDBFF8BA" box="[515,689,1834,1856]" italics="true" pageId="16" pageNumber="317">D. podolestoides</emphasis>
</taxonomicName>
male described in this paper; (18) 
<taxonomicName id="81A24D6B550AFFA8FBCB937DCB80F8BA" box="[1065,1166,1834,1856]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="74D6EAFA550AFFA8FBCB937DCB80F8BA" box="[1065,1166,1834,1856]" italics="true" pageId="16" pageNumber="317">D. somoh</emphasis>
</taxonomicName>
holotype; (19) 
<taxonomicName id="81A24D6B550AFFA8FAD6937DCA92F8BA" box="[1332,1436,1834,1856]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="74D6EAFA550AFFA8FAD6937DCA92F8BA" box="[1332,1436,1834,1856]" italics="true" pageId="16" pageNumber="317">D. tanduk</emphasis>
</taxonomicName>
holotype.
</paragraph>
</caption>
<caption id="12DD6660550BFFA9FF759329CE9FF828" httpUri="https://zenodo.org/record/253955/files/figure.png" pageId="17" pageNumber="318" targetBox="[194,1386,193,1882]" targetPageId="17">
<paragraph id="461D36E8550BFFA9FF759329CE9FF828" blockId="17.[151,1436,1919,2002]" pageId="17" pageNumber="318">
<emphasis id="74D6EAFA550BFFA9FF759329CE59F86F" bold="true" box="[151,343,1919,1941]" pageId="17" pageNumber="318">FIGURES 20–25.</emphasis>
Synthorax of male 
<taxonomicName id="81A24D6B550BFFA9FDCE9329CD9BF86F" box="[556,661,1919,1941]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA550BFFA9FDCE9329CD9BF86F" box="[556,661,1919,1941]" italics="true" pageId="17" pageNumber="318">Devadatta</emphasis>
</taxonomicName>
species, lateral view: (20) 
<taxonomicName id="81A24D6B550BFFA9FC5593D6CB04F86F" box="[951,1034,1920,1941]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="74D6EAFA550BFFA9FC5593D6CB04F86F" box="[951,1034,1920,1941]" italics="true" pageId="17" pageNumber="318">D. aran</emphasis>
</taxonomicName>
holotype; (21) 
<taxonomicName id="81A24D6B550BFFA9FB5393D6CA5EF86F" box="[1201,1360,1919,1941]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="74D6EAFA550BFFA9FB5393D6CA5EF86F" box="[1201,1360,1919,1941]" italics="true" pageId="17" pageNumber="318">D. basilanensis</emphasis>
</taxonomicName>
Davao Oriental, Mindanao; (22) 
<taxonomicName id="81A24D6B550BFFA9FE7E93C9CD24F849" box="[412,554,1949,1972]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="74D6EAFA550BFFA9FE7E93C9CD24F849" box="[412,554,1949,1972]" italics="true" pageId="17" pageNumber="318">D. clavicauda</emphasis>
</taxonomicName>
holotype; (23) 
<taxonomicName id="81A24D6B550BFFA9FD2A93C9CC7CF849" box="[712,882,1949,1972]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA550BFFA9FD2A93C9CC7CF849" box="[712,882,1949,1972]" italics="true" pageId="17" pageNumber="318">D. podolestoides</emphasis>
</taxonomicName>
male described in this paper; (24) 
<taxonomicName id="81A24D6B550BFFA9FB3693C9CA3BF849" box="[1236,1333,1949,1972]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="74D6EAFA550BFFA9FB3693C9CA3BF849" box="[1236,1333,1949,1972]" italics="true" pageId="17" pageNumber="318">D. somoh</emphasis>
</taxonomicName>
holotype; (25) 
<taxonomicName id="81A24D6B550BFFA9FF2593EBCE25F828" box="[199,299,1980,2002]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="74D6EAFA550BFFA9FF2593EBCE25F828" box="[199,299,1980,2002]" italics="true" pageId="17" pageNumber="318">D. tanduk</emphasis>
</taxonomicName>
holotype.
</paragraph>
</caption>
<caption id="12DD66605508FFAAFF75911BCC0EFA78" httpUri="https://zenodo.org/record/253956/files/figure.png" pageId="18" pageNumber="319" targetBox="[151,1436,193,1335]" targetPageId="18">
<paragraph id="461D36E85508FFAAFF75911BCC0EFA78" blockId="18.[151,1437,1357,1410]" pageId="18" pageNumber="319">
<emphasis id="74D6EAFA5508FFAAFF75911BCE58FA99" bold="true" box="[151,342,1357,1379]" pageId="18" pageNumber="319">FIGURES 26–29.</emphasis>
Venter of synthorax of male 
<taxonomicName id="81A24D6B5508FFAAFD68911BCDFAFA99" box="[650,756,1357,1379]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA5508FFAAFD68911BCDFAFA99" box="[650,756,1357,1379]" italics="true" pageId="18" pageNumber="319">Devadatta</emphasis>
</taxonomicName>
species: (26) 
<taxonomicName id="81A24D6B5508FFAAFC6A9118CCD7FA99" box="[904,985,1358,1379]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="74D6EAFA5508FFAAFC6A9118CCD7FA99" box="[904,985,1358,1379]" italics="true" pageId="18" pageNumber="319">D. aran</emphasis>
</taxonomicName>
holotype; (27) 
<taxonomicName id="81A24D6B5508FFAAFB9E9118CBEEFA99" box="[1148,1248,1357,1379]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="74D6EAFA5508FFAAFB9E9118CBEEFA99" box="[1148,1248,1357,1379]" italics="true" pageId="18" pageNumber="319">D. somoh</emphasis>
</taxonomicName>
holotype; (28) 
<taxonomicName id="81A24D6B5508FFAAFA619118CFD2FA78" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="74D6EAFA5508FFAAFA619118CFD2FA78" italics="true" pageId="18" pageNumber="319">D. tanduk</emphasis>
</taxonomicName>
holotype; (29) 
<taxonomicName id="81A24D6B5508FFAAFE9A913BCED2FA78" box="[376,476,1388,1410]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="319" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="74D6EAFA5508FFAAFE9A913BCED2FA78" box="[376,476,1388,1410]" italics="true" pageId="18" pageNumber="319">D. tanduk</emphasis>
</taxonomicName>
paratype (SAB12_AMP29).
</paragraph>
</caption>
<paragraph id="461D36E85508FFAAFF2591F8CFD3F9C8" blockId="18.[151,1437,1454,2019]" pageId="18" pageNumber="319">
<emphasis id="74D6EAFA5508FFAAFF2591F8CDACFA3D" bold="true" box="[199,674,1454,1479]" pageId="18" pageNumber="319">Description of female (SAR06_AMP21).</emphasis>
As male except as noted. Head. Basal ca 2/3 labrum brown. Most of mandible bases, genae and postclypeus brown, remainder of dorsum of head matte black with postocular lobes brown, joined by very distinct pale band along occipital ridge (
<figureCitation id="DE992A6D5508FFAAFCB691A0CCA6F9F5" box="[852,936,1526,1551]" captionStart="FIGURES 52 – 57" captionStartId="23.[151,264,1409,1431]" captionTargetBox="[151,1436,176,1394]" captionTargetId="figure@23.[151,1436,175,1395]" captionTargetPageId="23" captionText="FIGURES 52 – 57. Heads of Devadatta female specimens described in this paper: (52) D. aran; (53) D. basilanensis; (54) D. clavicauda; (55) D. podolestoides; (56) D. somoh; (57) D. tanduk." httpUri="https://zenodo.org/record/253962/files/figure.png" pageId="18" pageNumber="319">Fig. 52</figureCitation>
). Antennae with anterior yellowish streak on scape.
</paragraph>
<paragraph id="461D36E85508FFAAFF259268CDB1F8F0" blockId="18.[151,1437,1454,2019]" pageId="18" pageNumber="319">
Thorax: Pronotum (
<figureCitation id="DE992A6D5508FFAAFE4E9268CD0FF9AD" box="[428,513,1598,1623]" captionStart="FIGURES 58 – 63" captionStartId="25.[151,264,1932,1954]" captionTargetBox="[151,1436,411,1910]" captionTargetId="figure@25.[151,1436,411,1910]" captionTargetPageId="25" captionText="FIGURES 58 – 63. Prothorax of Devadatta female specimens described in this paper, dorsal view: (58) D. aran; (59) D. basilanensis; (60) D. clavicauda; (61) D. podolestoides; (62) D. somoh; (63) D. tanduk." httpUri="https://zenodo.org/record/253963/files/figure.png" pageId="18" pageNumber="319">Fig. 58</figureCitation>
) mottled brown, larger pale marks on anterior lobe, middle lobe bearing small projections, stump-like in lateral view (
<figureCitation id="DE992A6D5508FFAAFDBA9234CDA0F981" box="[600,686,1634,1659]" captionStart="FIGURES 64 – 67" captionStartId="26.[151,264,1535,1557]" captionTargetBox="[151,1436,193,1512]" captionTargetId="figure@26.[151,1436,193,1513]" captionTargetPageId="26" captionText="FIGURES 64 – 67. Prothorax of female Devadatta species with horns, lateral view: (64) D. aran female described in this paper; (65) D. aran Crocker Range, Sabah; (66) D. somoh female described in this paper; (67) D. tanduk female described in this paper." httpUri="https://zenodo.org/record/253964/files/figure.png" pageId="18" pageNumber="319">Fig. 64</figureCitation>
). Posterior lobe with yellow lateral patches. Dorsum of synthorax brown with middorsal carina black, small bluish pale mark between arms of anterior bifurcation of middorsal carina. Synthorax laterally brown with pale marks as in male (
<figureCitation id="DE992A6D5508FFAAFC8392FCCCB9F939" box="[865,951,1706,1731]" captionStart="FIGURES 68 – 73" captionStartId="28.[151,264,1928,1950]" captionTargetBox="[151,1436,193,1906]" captionTargetId="figure@28.[151,1436,193,1907]" captionTargetPageId="28" captionText="FIGURES 68 – 73. Synthorax of Devadatta female specimens described in this paper, lateral view: (68) D. aran; (69) D. basilanensis; (70) D. clavicauda; (71) D. podolestoides; (72) D. somoh; (73) D. tanduk." httpUri="https://zenodo.org/record/253965/files/figure.png" pageId="18" pageNumber="319">Fig. 68</figureCitation>
). Wings with arculus at Ax 
<quantity id="815A9B0D5508FFAAFB1A92FDCA2BF939" box="[1272,1317,1706,1731]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="18" pageNumber="319" unit="in" value="4.0">4 in</quantity>
all wings, quadrilateral with two crossvein in all wings, pt black proximally apical ca 2/3 mostly pale. Extreme tips of both wings slightly tinted brown, a little darker in Hw.
</paragraph>
<paragraph id="461D36E85508FFAAFF259340CC23F8A9" blockId="18.[151,1437,1454,2019]" pageId="18" pageNumber="319">Abdomen: Small pale lateral basal spot on S2. Cerci black, subtriangular, ca same length as S10. Ovipositor short, only reaching beyond S10 to ca level of tips of cerci.</paragraph>
<paragraph id="461D36E85508FFAAFF259308CB4DF88D" blockId="18.[151,1437,1454,2019]" box="[199,1091,1886,1911]" pageId="18" pageNumber="319">
<emphasis id="74D6EAFA5508FFAAFF259308CE78F88D" bold="true" box="[199,374,1886,1911]" pageId="18" pageNumber="319">Measurements</emphasis>
[mm]: abdomen without ovipositor and appendages 35, Hw 33.
</paragraph>
<paragraph id="461D36E85508FFABFF2593D4CD36FDE2" blockId="18.[151,1437,1454,2019]" lastBlockId="19.[151,1436,151,536]" lastPageId="19" lastPageNumber="320" pageId="18" pageNumber="319">
<emphasis id="74D6EAFA5508FFAAFF2593D4CED4F861" bold="true" box="[199,474,1922,1947]" pageId="18" pageNumber="319">
Variation in 
<typeStatus id="9919884A5508FFAAFEBF93D4CEDBF861" box="[349,469,1922,1947]" pageId="18" pageNumber="319" type="paratype">paratypes</typeStatus>
.
</emphasis>
Males: Apart from small differences in the brightness and extent of pale markings on the head, thorax and abdomen, there is little variation in most 
<typeStatus id="9919884A5508FFAAFCBD93F1CCC5F845" box="[863,971,1959,1983]" pageId="18" pageNumber="319" type="paratype">paratypes</typeStatus>
from 
<collectingCountry id="3EB576785508FFAAFBF093F0CB7CF845" box="[1042,1138,1958,1983]" name="Malaysia" pageId="18" pageNumber="319">Sarawak</collectingCountry>
and Sabah except in size. Teneral males frequently have a pale band joining the pale areas on the postocular lobes. Pale marks on the posterior pronotal lobe are absent in some individuals and more extensive in others. The arculus is at Ax 3–5, but in almost all individuals it is at Ax 4 or 
<quantity id="815A9B0D5509FFABFDDD94EACD65FF2F" box="[575,619,188,213]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="19" pageNumber="320" unit="in" value="5.0">5 in</quantity>
at least 2 wings; in only one specimen from the Hose Mountains it is at Ax 
<quantity id="815A9B0D5509FFABFF2294B6CFE5FF02" box="[192,235,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="19" pageNumber="320" unit="in" value="3.0">3 in</quantity>
all wings. Up to three crossveins in the quadrilateral. Rarely there is a crossvein in the cubito-anal space of one wing; in one individual from the Tama Abu range there are crossveins in two wings, and in one other there are single crossveins in two wings and two crossveins in one wing. The colour of pt varies, sometimes darker than in the 
<typeStatus id="9919884A5509FFABFF20951ACE29FE9F" box="[194,295,332,357]" pageId="19" pageNumber="320" type="holotype">holotype</typeStatus>
, sometimes with a pale central area; this variation appears to be associated with maturity. The wing tips are sometimes entirely hyaline. Rotation of one of the cerci, in one case through ca 90 degrees, is not uncommon and can make them appear different on a superficial view. The two male specimens from southeast Kalimantan have a slightly darker appearance than the others, with some shadowing on the venter of the synthorax near the abdomen. The cerci of all males (including the 
<typeStatus id="9919884A5509FFABFCFE958ACC8EFE0F" box="[796,896,476,501]" pageId="19" pageNumber="320" type="holotype">holotype</typeStatus>
) bear small spines; these are highly variable in size and position and non-diagnostic.
</paragraph>
<caption id="12DD66605509FFABFF7592ADCE31F8B4" httpUri="https://zenodo.org/record/253957/files/figure.png" pageId="19" pageNumber="320" targetBox="[151,1436,611,1753]" targetPageId="19">
<paragraph id="461D36E85509FFABFF7592ADCE31F8B4" blockId="19.[151,1437,1787,1870]" pageId="19" pageNumber="320">
<emphasis id="74D6EAFA5509FFABFF7592ADCE58F8EB" bold="true" box="[151,342,1787,1809]" pageId="19" pageNumber="320">FIGURES 30–33.</emphasis>
Venter of synthorax of male 
<taxonomicName id="81A24D6B5509FFABFD6892ADCDFAF8EB" box="[650,756,1787,1809]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA5509FFABFD6892ADCDFAF8EB" box="[650,756,1787,1809]" italics="true" pageId="19" pageNumber="320">Devadatta</emphasis>
</taxonomicName>
species: (30) 
<taxonomicName id="81A24D6B5509FFABFC6592AACB2BF8EB" box="[903,1061,1787,1809]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="74D6EAFA5509FFABFC6592AACB2BF8EB" box="[903,1061,1787,1809]" italics="true" pageId="19" pageNumber="320">D. basilanensis</emphasis>
</taxonomicName>
Davao Oriental, Mindanao; (31) 
<taxonomicName id="81A24D6B5509FFABFA6192AACE2CF8D5" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA5509FFABFA6192AACE2CF8D5" italics="true" pageId="19" pageNumber="320">D. podolestoides</emphasis>
</taxonomicName>
male described in this paper; (32) 
<taxonomicName id="81A24D6B5509FFABFD71934CCC2DF8D5" box="[659,803,1817,1839]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="74D6EAFA5509FFABFD71934CCC2DF8D5" box="[659,803,1817,1839]" italics="true" pageId="19" pageNumber="320">D. clavicauda</emphasis>
</taxonomicName>
holotype; (33) 
<taxonomicName id="81A24D6B5509FFABFC25934CCB59F8D5" box="[967,1111,1817,1839]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="320" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="74D6EAFA5509FFABFC25934CCB59F8D5" box="[967,1111,1817,1839]" italics="true" pageId="19" pageNumber="320">D. clavicauda</emphasis>
</taxonomicName>
paratype (SAR09_10_AMP40, Gunung Serapi).
</paragraph>
</caption>
<caption id="12DD6660550EFFACFF75903ECEF2FB41" httpUri="https://zenodo.org/record/253958/files/figure.png" pageId="20" pageNumber="321" targetBox="[151,1436,193,1106]" targetPageId="20">
<paragraph id="461D36E8550EFFACFF75903ECEF2FB41" blockId="20.[151,1436,1128,1211]" pageId="20" pageNumber="321">
<emphasis id="74D6EAFA550EFFACFF75903ECE5BFB84" bold="true" box="[151,341,1128,1150]" pageId="20" pageNumber="321">FIGURES 34–37.</emphasis>
Wings of some 
<taxonomicName id="81A24D6B550EFFACFE1F903ECD69FB84" box="[509,615,1128,1150]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA550EFFACFE1F903ECD69FB84" box="[509,615,1128,1150]" italics="true" pageId="20" pageNumber="321">Devadatta</emphasis>
</taxonomicName>
species: (34) 
<taxonomicName id="81A24D6B550EFFACFD14903CCC9DFB84" box="[758,915,1128,1151]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="74D6EAFA550EFFACFD14903CCC9DFB84" box="[758,915,1128,1151]" italics="true" pageId="20" pageNumber="321">D. basilanensis</emphasis>
</taxonomicName>
male Zamboanga, Mindanao; (35) 
<taxonomicName id="81A24D6B550EFFACFB1D903CCA92FB84" box="[1279,1436,1128,1151]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="74D6EAFA550EFFACFB1D903CCA92FB84" box="[1279,1436,1128,1151]" italics="true" pageId="20" pageNumber="321">D. basilanensis</emphasis>
</taxonomicName>
female Cotabato, Mindanao; (36) 
<taxonomicName id="81A24D6B550EFFACFDE190DECD98FB67" box="[515,662,1159,1181]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="74D6EAFA550EFFACFDE190DECD98FB67" box="[515,662,1159,1181]" italics="true" pageId="20" pageNumber="321">D. clavicauda</emphasis>
</taxonomicName>
paratype, Gunung Santubong, Sarawak; (37) 
<taxonomicName id="81A24D6B550EFFACFB6690DECA3AFB67" box="[1156,1332,1159,1181]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA550EFFACFB6690DECA3AFB67" box="[1156,1332,1159,1181]" italics="true" pageId="20" pageNumber="321">D. podolestoides</emphasis>
</taxonomicName>
paratype, Gunung Poteng, Kalimantan Barat.
</paragraph>
</caption>
<caption id="12DD6660550EFFACFF759321CEB7F851" httpUri="https://zenodo.org/record/253959/files/figure.png" pageId="20" pageNumber="321" targetBox="[151,1436,1261,1889]" targetPageId="20">
<paragraph id="461D36E8550EFFACFF759321CEB7F851" blockId="20.[151,1436,1911,1963]" pageId="20" pageNumber="321">
<emphasis id="74D6EAFA550EFFACFF759321CE5AF877" bold="true" box="[151,340,1911,1933]" pageId="20" pageNumber="321">FIGURES 38–39.</emphasis>
Genital ligula of two 
<taxonomicName id="81A24D6B550EFFACFDD49321CD91F877" box="[566,671,1911,1933]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="321" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA550EFFACFDD49321CD91F877" box="[566,671,1911,1933]" italics="true" pageId="20" pageNumber="321">Devadatta</emphasis>
</taxonomicName>
species, scale bars = 0.5mm: (38) 
<taxonomicName id="81A24D6B550EFFACFBE1932ECB5DF877" box="[1027,1107,1912,1933]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="74D6EAFA550EFFACFBE1932ECB5DF877" box="[1027,1107,1912,1933]" italics="true" pageId="20" pageNumber="321">D. aran</emphasis>
</taxonomicName>
holotype; (39) 
<taxonomicName id="81A24D6B550EFFACFB13932ECA92F877" box="[1265,1436,1911,1933]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA550EFFACFB13932ECA92F877" box="[1265,1436,1911,1933]" italics="true" pageId="20" pageNumber="321">D. podolestoides</emphasis>
</taxonomicName>
male described in this paper.
</paragraph>
</caption>
<caption id="12DD6660550FFFADFF7592F5CA7AF92D" httpUri="https://zenodo.org/record/253960/files/figure.png" pageId="21" pageNumber="322" targetBox="[162,1425,203,1663]" targetPageId="21">
<paragraph id="461D36E8550FFFADFF7592F5CA7AF92D" blockId="21.[151,1436,1699,1751]" pageId="21" pageNumber="322">
<emphasis id="74D6EAFA550FFFADFF7592F5CE5AF943" bold="true" box="[151,340,1699,1721]" pageId="21" pageNumber="322">FIGURES 40–45.</emphasis>
Male anal appendages, dorsal view: (40) 
<taxonomicName id="81A24D6B550FFFADFD1B92F2CC46F943" box="[761,840,1700,1721]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="74D6EAFA550FFFADFD1B92F2CC46F943" box="[761,840,1700,1721]" italics="true" pageId="21" pageNumber="322">D. aran</emphasis>
</taxonomicName>
holotype; (41) 
<taxonomicName id="81A24D6B550FFFADFC0692F2CB8EF943" box="[996,1152,1699,1721]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="74D6EAFA550FFFADFC0692F2CB8EF943" box="[996,1152,1699,1721]" italics="true" pageId="21" pageNumber="322">D. basilanensis</emphasis>
</taxonomicName>
Davao Oriental, Mindanao; (42) 
<taxonomicName id="81A24D6B550FFFADFF259294CE5BF92D" box="[199,341,1729,1751]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="74D6EAFA550FFFADFF259294CE5BF92D" box="[199,341,1729,1751]" italics="true" pageId="21" pageNumber="322">D. clavicauda</emphasis>
</taxonomicName>
holotype; (43) 
<taxonomicName id="81A24D6B550FFFADFE109294CDE9F92D" authority="Kubah" authorityName="Kubah" box="[498,743,1729,1751]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA550FFFADFE109294CD92F92D" box="[498,668,1729,1751]" italics="true" pageId="21" pageNumber="322">D. podolestoides</emphasis>
Kubah
</taxonomicName>
NP, Sarawak; (44) 
<taxonomicName id="81A24D6B550FFFADFC4E9294CB03F92D" box="[940,1037,1729,1751]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="74D6EAFA550FFFADFC4E9294CB03F92D" box="[940,1037,1729,1751]" italics="true" pageId="21" pageNumber="322">D. somoh</emphasis>
</taxonomicName>
holotype; (45) 
<taxonomicName id="81A24D6B550FFFADFB4B9294CA03F92D" box="[1193,1293,1729,1751]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="74D6EAFA550FFFADFB4B9294CA03F92D" box="[1193,1293,1729,1751]" italics="true" pageId="21" pageNumber="322">D. tanduk</emphasis>
</taxonomicName>
holotype.
</paragraph>
</caption>
<paragraph id="461D36E8550FFFAEFF2592A8CE75FF02" blockId="21.[151,1436,1790,2031]" lastBlockId="22.[151,1437,151,320]" lastPageId="22" lastPageNumber="323" pageId="21" pageNumber="322">
Females: The other female 
<typeStatus id="9919884A550FFFADFDE79356CD7FF8E2" box="[517,625,1792,1816]" pageId="21" pageNumber="322" type="paratype">paratypes</typeStatus>
show a range of variation in markings similar to that of males. In all specimens examined the arculus is at Ax 4 or 5, except one from the Hose Mountains where it is at Ax 
<quantity id="815A9B0D550FFFADFAA89375CA79F8C1" box="[1354,1399,1826,1851]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="21" pageNumber="322" unit="in" value="3.0">3 in</quantity>
all wings. Rarely with a crossvein in the cubito-anal space of one wing. In some specimens there is no or virtually no darkening of the wings tips, in a few it is more extensive and darker than in the female described. In some individuals the wing is entirely un-tinted; all of these appear teneral or just post-teneral. The most significant variation is in the horns on the middle pronotal lobe, which vary from the condition seen in the 
<typeStatus id="9919884A550FFFADFB1093E5CA2CF831" box="[1266,1314,1971,1995]" pageId="21" pageNumber="322">type</typeStatus>
to upright nipple-like horns (e.g. 
<figureCitation id="DE992A6D550FFFADFE7E9380CEFFF815" box="[412,497,2006,2031]" captionStart="FIGURES 64 – 67" captionStartId="26.[151,264,1535,1557]" captionTargetBox="[151,1436,193,1512]" captionTargetId="figure@26.[151,1436,193,1513]" captionTargetPageId="26" captionText="FIGURES 64 – 67. Prothorax of female Devadatta species with horns, lateral view: (64) D. aran female described in this paper; (65) D. aran Crocker Range, Sabah; (66) D. somoh female described in this paper; (67) D. tanduk female described in this paper." httpUri="https://zenodo.org/record/253964/files/figure.png" pageId="21" pageNumber="322">Fig. 65</figureCitation>
), which slope back slightly in a few individuals. Most individuals from the Tama Abu range have projections like that in the female described, but one is intermediate between this and the one shown in 
<figureCitation id="DE992A6D550CFFAEFEE594EACE54FF2F" box="[263,346,188,213]" captionStart="FIGURES 64 – 67" captionStartId="26.[151,264,1535,1557]" captionTargetBox="[151,1436,193,1512]" captionTargetId="figure@26.[151,1436,193,1513]" captionTargetPageId="26" captionText="FIGURES 64 – 67. Prothorax of female Devadatta species with horns, lateral view: (64) D. aran female described in this paper; (65) D. aran Crocker Range, Sabah; (66) D. somoh female described in this paper; (67) D. tanduk female described in this paper." httpUri="https://zenodo.org/record/253964/files/figure.png" pageId="22" pageNumber="323">Fig. 65</figureCitation>
, while both extremes are present in other areas; a rearward slant is more common in specimens from Kapit division.
</paragraph>
<paragraph id="461D36E8550CFFAEFF259552CCBCFEBA" blockId="22.[151,1437,151,320]" pageId="22" pageNumber="323">
<emphasis id="74D6EAFA550CFFAEFF259552CE78FEE7" bold="true" box="[199,374,260,285]" pageId="22" pageNumber="323">Measurements</emphasis>
[mm]: Males: abdomen without anal appendages 33.5–41.5, Hw 28.5–36.5. Females: abdomen without anal appendages or ovipositor 29–37, Hw 28.5–36.5.
</paragraph>
</subSubSection>
<caption id="12DD6660550CFFAEFF7593C9CE4CF808" httpUri="https://zenodo.org/record/253961/files/figure.png" pageId="22" pageNumber="323" targetBox="[162,1418,393,1916]" targetPageId="22">
<paragraph id="461D36E8550CFFAEFF7593C9CE4CF808" blockId="22.[151,1437,1951,2034]" pageId="22" pageNumber="323">
<emphasis id="74D6EAFA550CFFAEFF7593C9CE5BF84F" bold="true" box="[151,341,1951,1973]" pageId="22" pageNumber="323">FIGURES 46–51.</emphasis>
Male anal appendages, lateral view (scale bars 1 mm): (46) 
<taxonomicName id="81A24D6B550CFFAEFC2793F6CB1BF84F" box="[965,1045,1952,1973]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="74D6EAFA550CFFAEFC2793F6CB1BF84F" box="[965,1045,1952,1973]" italics="true" pageId="22" pageNumber="323">D. aran</emphasis>
</taxonomicName>
holotype; (47) 
<taxonomicName id="81A24D6B550CFFAEFB5793F6CA5CF84F" box="[1205,1362,1951,1973]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="74D6EAFA550CFFAEFB5793F6CA5CF84F" box="[1205,1362,1951,1973]" italics="true" pageId="22" pageNumber="323">D. basilanensis</emphasis>
</taxonomicName>
Davao Oriental, Mindanao; (48) 
<taxonomicName id="81A24D6B550CFFAEFE7E93E9CD24F829" box="[412,554,1981,2004]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="74D6EAFA550CFFAEFE7E93E9CD24F829" box="[412,554,1981,2004]" italics="true" pageId="22" pageNumber="323">D. clavicauda</emphasis>
</taxonomicName>
holotype; (49) 
<taxonomicName id="81A24D6B550CFFAEFD2A93E9CCB3F829" authority="Kubah" authorityName="Kubah" box="[712,957,1981,2004]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA550CFFAEFD2A93E9CC7DF829" box="[712,883,1981,2004]" italics="true" pageId="22" pageNumber="323">D. podolestoides</emphasis>
Kubah
</taxonomicName>
NP, Sarawak; (50) 
<taxonomicName id="81A24D6B550CFFAEFB6693E9CBEBF829" box="[1156,1253,1981,2004]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="74D6EAFA550CFFAEFB6693E9CBEBF829" box="[1156,1253,1981,2004]" italics="true" pageId="22" pageNumber="323">D. somoh</emphasis>
</taxonomicName>
holotype; (51) 
<taxonomicName id="81A24D6B550CFFAEFA6193E9CFD2F808" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="74D6EAFA550CFFAEFA6193E9CFD2F808" italics="true" pageId="22" pageNumber="323">D. tanduk</emphasis>
</taxonomicName>
holotype.
</paragraph>
</caption>
<caption id="12DD6660550DFFAFFF7591D7CC3AFA4F" httpUri="https://zenodo.org/record/253962/files/figure.png" pageId="23" pageNumber="324" targetBox="[151,1436,176,1394]" targetPageId="23">
<paragraph id="461D36E8550DFFAFFF7591D7CC3AFA4F" blockId="23.[151,1437,1409,1462]" pageId="23" pageNumber="324">
<emphasis id="74D6EAFA550DFFAFFF7591D7CE5BFA6D" bold="true" box="[151,341,1409,1431]" pageId="23" pageNumber="324">FIGURES 52–57.</emphasis>
Heads of 
<taxonomicName id="81A24D6B550DFFAFFE5D91D7CD26FA6D" box="[447,552,1409,1431]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA550DFFAFFE5D91D7CD26FA6D" box="[447,552,1409,1431]" italics="true" pageId="23" pageNumber="324">Devadatta</emphasis>
</taxonomicName>
female specimens described in this paper: (52) 
<taxonomicName id="81A24D6B550DFFAFFBF491D4CB68FA6D" box="[1046,1126,1410,1431]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="74D6EAFA550DFFAFFBF491D4CB68FA6D" box="[1046,1126,1410,1431]" italics="true" pageId="23" pageNumber="324">D. aran</emphasis>
</taxonomicName>
; (53) 
<taxonomicName id="81A24D6B550DFFAFFB4491D4CA4DFA6D" box="[1190,1347,1409,1431]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="basilanensis">
<emphasis id="74D6EAFA550DFFAFFB4491D4CA4DFA6D" box="[1190,1347,1409,1431]" italics="true" pageId="23" pageNumber="324">D. basilanensis</emphasis>
</taxonomicName>
; (54) 
<taxonomicName id="81A24D6B550DFFAFFA6191D4CE08FA4F" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="74D6EAFA550DFFAFFA6191D4CE08FA4F" italics="true" pageId="23" pageNumber="324">D. clavicauda</emphasis>
</taxonomicName>
; (55) 
<taxonomicName id="81A24D6B550DFFAFFEA691F7CEE3FA4F" box="[324,493,1439,1462]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="podolestoides">
<emphasis id="74D6EAFA550DFFAFFEA691F7CEE3FA4F" box="[324,493,1439,1462]" italics="true" pageId="23" pageNumber="324">D. podolestoides</emphasis>
</taxonomicName>
; (56) 
<taxonomicName id="81A24D6B550DFFAFFDCE91F7CD82FA4F" box="[556,652,1439,1462]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="74D6EAFA550DFFAFFDCE91F7CD82FA4F" box="[556,652,1439,1462]" italics="true" pageId="23" pageNumber="324">D. somoh</emphasis>
</taxonomicName>
; (57) 
<taxonomicName id="81A24D6B550DFFAFFD2991F7CC21FA4F" box="[715,815,1439,1462]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="74D6EAFA550DFFAFFD2991F7CC21FA4F" box="[715,815,1439,1462]" italics="true" pageId="23" pageNumber="324">D. tanduk</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="0EB86563550DFFAFFF25918BCEECF97F" pageId="23" pageNumber="324" type="discussion">
<paragraph id="461D36E8550DFFAFFF25918BCEECF97F" blockId="23.[151,1437,1501,1670]" pageId="23" pageNumber="324">
<emphasis id="74D6EAFA550DFFAFFF25918BCE35FA0C" bold="true" box="[199,315,1501,1526]" pageId="23" pageNumber="324">Remarks.</emphasis>
Across most of its known range 
<taxonomicName id="81A24D6B550DFFAFFD5E9188CC1BFA0F" box="[700,789,1502,1525]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="74D6EAFA550DFFAFFD5E9188CC1BFA0F" box="[700,789,1502,1525]" italics="true" pageId="23" pageNumber="324">D. aran</emphasis>
</taxonomicName>
is scarce at lower altitudes and more common above ca 
<specimenCount id="50A4FD61550DFFAFFF759257CFD0F9E3" box="[151,222,1537,1561]" pageId="23" pageNumber="324" type="generic">
<quantity id="815A9B0D550DFFAFFF759257CFD4F9E3" box="[151,218,1537,1561]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="23" pageNumber="324" unit="m" value="900.0">900m</quantity>
.
</specimenCount>
Although its distribution (
<figureCitation id="DE992A6D550DFFAFFDF09257CD68F9E3" box="[530,614,1537,1562]" captionStart="FIGURE 74" captionStartId="29.[151,250,1608,1630]" captionTargetBox="[151,1436,193,1586]" captionTargetId="figure@29.[151,1436,193,1586]" captionTargetPageId="29" captionText="FIGURE 74. Distribution of D. aran (squares) and D. podolestoides (triangles)." httpUri="https://zenodo.org/record/253966/files/figure.png" pageId="23" pageNumber="324">Fig. 74</figureCitation>
) broadly or entirely overlaps those of 
<taxonomicName id="81A24D6B550DFFAFFBC49254CBC8F9E3" box="[1062,1222,1537,1561]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="clavicauda">
<emphasis id="74D6EAFA550DFFAFFBC49254CBC8F9E3" box="[1062,1222,1537,1561]" italics="true" pageId="23" pageNumber="324">D. clavicauda</emphasis>
</taxonomicName>
, 
<taxonomicName id="81A24D6B550DFFAFFB359254CA4BF9E3" box="[1239,1349,1537,1561]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="somoh">
<emphasis id="74D6EAFA550DFFAFFB359254CA4BF9E3" box="[1239,1349,1537,1561]" italics="true" pageId="23" pageNumber="324">D. somoh</emphasis>
</taxonomicName>
and 
<taxonomicName id="81A24D6B550DFFAFFA639254CFEDF9C7" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="tanduk">
<emphasis id="74D6EAFA550DFFAFFA639254CFEDF9C7" italics="true" pageId="23" pageNumber="324">D. tanduk</emphasis>
</taxonomicName>
, above ca 
<quantity id="815A9B0D550DFFAFFEB99270CEA7F9C4" box="[347,425,1574,1598]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="324" unit="m" value="1000.0">1000m</quantity>
in the Tama Abu Range in 
<collectingCountry id="3EB57678550DFFAFFD3F9273CC33F9C4" box="[733,829,1573,1598]" name="Malaysia" pageId="23" pageNumber="324">Sarawak</collectingCountry>
and the adjacent Long Pasia highland area in Sabah it appears to be the only 
<taxonomicName id="81A24D6B550DFFAFFE41921FCD16F99B" box="[419,536,1609,1633]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="genus">
<emphasis id="74D6EAFA550DFFAFFE41921FCD16F99B" box="[419,536,1609,1633]" italics="true" pageId="23" pageNumber="324">Devadatta</emphasis>
</taxonomicName>
species present. 
<taxonomicName id="81A24D6B550DFFAFFD3D921FCC9DF99B" box="[735,915,1609,1633]" class="Insecta" family="Amphipterygidae" genus="Devadatta" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="aran">
<emphasis id="74D6EAFA550DFFAFFD3D921FCC9DF99B" box="[735,915,1609,1633]" italics="true" pageId="23" pageNumber="324">Devadatta aran</emphasis>
</taxonomicName>
occurs up to at least 
<quantity id="815A9B0D550DFFAFFB71921FCBEFF998" box="[1171,1249,1609,1634]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="324" unit="m" value="1500.0">1500m</quantity>
and is typically found at small forest streams.
</paragraph>
</subSubSection>
</treatment>
</document>