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<document id="59B89B6C5B9AE516C413E506FD1768CC" ID-CLB-Dataset="27502" ID-DOI="10.11646/zootaxa.4550.1.1" ID-GBIF-Dataset="291701d4-652e-440e-adab-f6b0201c0972" ID-ISSN="1175-5326" ID-Zenodo-Dep="2625030" ID-ZooBank="2F7D1EC4-D4ED-4FAE-B227-CF7B79EAE833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554274879827" checkinUser="plazi" docAuthor="Huber, Bernhard A." docDate="2019" docId="CA3B104CFF88FF85FF3DFD24FBBDE72C" docLanguage="en" docName="zootaxa.4550.1.1.pdf" docOrigin="Zootaxa 4550 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tissahamia kottawagamaensis" docType="treatment" docVersion="14" lastPageNumber="31" masterDocId="36026834FF95FF9BFFAAFFA1FFB6E16A" masterDocTitle="The pholcid spiders of Sri Lanka (Araneae: Pholcidae)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="30" updateTime="1698716853299" updateUser="ExternalLinkService">
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<mods:title id="B7182C604CA1AF185A6A416DE0D98C6B">The pholcid spiders of Sri Lanka (Araneae: Pholcidae)</mods:title>
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<mods:namePart id="F5495AD3942D57E591FD681DAFE7067F">Huber, Bernhard A.</mods:namePart>
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<mods:date id="3D1F2DAA1D73F98FEA40121B3CC773EE">2019</mods:date>
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<treatment id="CA3B104CFF88FF85FF3DFD24FBBDE72C" ID-DOI="http://doi.org/10.5281/zenodo.3510762" ID-GBIF-Taxon="154534138" ID-Zenodo-Dep="3510762" LSID="urn:lsid:plazi:treatment:CA3B104CFF88FF85FF3DFD24FBBDE72C" httpUri="http://treatment.plazi.org/id/CA3B104CFF88FF85FF3DFD24FBBDE72C" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<subSubSection id="0A88F2D1FF88FF86FF3DFD24FD6AE3F5" box="[151,732,645,671]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="422DA15AFF88FF86FF3DFD24FD6AE3F5" blockId="29.[151,732,645,704]" box="[151,732,645,671]" pageId="29" pageNumber="30">
<heading id="19651636FF88FF86FF3DFD24FD6AE3F5" bold="true" box="[151,732,645,671]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<taxonomicName id="8592DAD9FF88FF86FF3DFD24FD6AE3F5" ID-CoL="577S3" authority="(Yao &amp; Li, 2016)" baseAuthorityName="Yao &amp; Li" baseAuthorityYear="2016" box="[151,732,645,671]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kottawagamaensis">
<emphasis id="70E67D48FF88FF86FF3DFD24FD6AE3F5" bold="true" box="[151,732,645,671]" pageId="29" pageNumber="30">
<emphasis id="70E67D48FF88FF86FF3DFD24FDB0E3F5" bold="true" box="[151,518,645,671]" italics="true" pageId="29" pageNumber="30">Tissahamia kottawagamaensis</emphasis>
(
<bibRefCitation id="2603DCABFF88FF86FDBFFD24FD62E3F5" author="Yao, Z. &amp; Dong, T. &amp; Zheng, G. &amp; Fu, J. &amp; Li, S." box="[533,724,645,671]" pageId="29" pageNumber="30" pagination="35757" refId="ref29041" refString="Yao, Z., Dong, T., Zheng, G., Fu, J. &amp; Li, S. (2016) High endemism at cave entrances: a case study of spiders of the genus Uthina. Scientific Reports, 6, 35757. https: // doi. org / 10.1038 / srep 35757" type="journal article" year="2016">Yao &amp; Li, 2016</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FF88FF86FF3DFD06FDF3E3AA" box="[151,581,679,704]" pageId="29" pageNumber="30" type="description">
<paragraph id="422DA15AFF88FF86FF3DFD06FDF3E3AA" blockId="29.[151,732,645,704]" box="[151,581,679,704]" pageId="29" pageNumber="30">
<figureCitation id="DAA9BDDFFF88FF86FF3DFD06FE85E3AA" box="[151,307,679,704]" captionStart="FIGURES 93–102" captionStartId="25.[151,264,1918,1941]" captionTargetBox="[151,1436,418,1859]" captionTargetId="figure@25.[151,1436,418,1859]" captionTargetPageId="25" captionText="FIGURES 93–102. Tissahamia, live specimens. 93–96. T. ethagala (Huber, 2011), male from Ethagala, female with egg-sac from Kandalama, and females from Dematagala. 97–98. T. karuna sp. n., male and female with egg-sac from Labugama. 99– 100. T. maturata (Huber, 2011), male with mite and female with egg-sac from Hakgala. 101–102. T. kottawagamaensis (Yao &amp; Li, 2016), male from Kottawa, female from Viharekele." httpUri="https://zenodo.org/record/2625062/files/figure.png" pageId="29" pageNumber="30">Figs 101–102</figureCitation>
,
<figureCitation id="DAA9BDDFFF88FF86FE94FD06FE17E3AA" box="[318,417,679,704]" captionStart="FIGURES 110–115" captionStartId="28.[151,264,1879,1902]" captionTargetBox="[202,1418,342,1855]" captionTargetId="figure@28.[164,1427,332,1861]" captionTargetPageId="28" captionText="FIGURES 110–115. Left procursi in prolateral (upper row) and retrolateral views, all images at same scale. 110–111. Tissahamia karuna sp. n., from Labugama (ZFMK Ar 20066); asterisk marks area where males from Mitirigala have a small conical process. 112–113. T. maturata (Huber, 2011), from Hakgala (ZFMK Ar 20069); arrow points at distinctive prolateral process. 114–115. T. kottawagamaensis (Yao &amp; Li, 2016), from Kottawa (ZFMK Ar 20072). Abbreviations: bp, bifid process; hs, hinged sclerite; m, membrane; po, pocket; r, ridge; ta, tarsus. Scale line: 0.5 mm." httpUri="https://zenodo.org/record/2625066/files/figure.png" pageId="29" pageNumber="30">114–115</figureCitation>
,
<figureCitation id="DAA9BDDFFF88FF86FE06FD09FE6EE3AA" box="[428,472,680,704]" captionStart="FIGURES 116–119" captionStartId="29.[151,264,1920,1943]" captionTargetBox="[155,1435,1202,1900]" captionTargetId="figure@29.[151,1436,1193,1908]" captionTargetPageId="29" captionText="FIGURES 116–119. 116–117. Tissahamia karuna sp. n., from Labugama (ZFMK Ar 20066), male chelicerae, frontal view; and processes of left genital bulb, prolateral view. 118–119. T. maturata (Huber, 2011) from Hakgala (ZFMK Ar 20069) and T. kottawagamaensis (Yao &amp; Li, 2016) from Kottawa (ZFMK Ar 20072), processes of left genital bulbs in prolateral views (all genital bulbs at same scale). Abbreviations: a, ‘appendix’ (main bulbal process); e, embolus. Scale lines: 0.3 mm." httpUri="https://zenodo.org/record/2625070/files/figure.png" pageId="29" pageNumber="30">119</figureCitation>
,
<figureCitation id="DAA9BDDFFF88FF86FE49FD06FDF3E3AA" box="[483,581,679,704]" captionStart="FIGURES 120–125" captionStartId="30.[151,264,1133,1156]" captionTargetBox="[151,1436,193,1113]" captionTargetId="figure@30.[151,1436,193,1113]" captionTargetPageId="30" captionText="FIGURES 120–125. Epigyna in ventral views (upper row) and cleared female genitalia in dorsal views. 120–121. Tissahamia karuna sp. n., from Labugama (ZFMK Ar 20067); arrow points at internal arc (‘valve’). 122–123. T. maturata (Huber, 2011), from Hakgala (ZFMK Ar 20070). 124–125. T. kottawagamaensis (Yao &amp; Li, 2016), from Kottawa (ZFMK Ar 20073). Abbreviation: pp, pore plate. At various scales." httpUri="https://zenodo.org/record/2625072/files/figure.png" pageId="29" pageNumber="30">124–125</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FF88FF86FF3DFD4BFBECE275" pageId="29" pageNumber="30" type="reference_group">
<paragraph id="422DA15AFF88FF86FF3DFD4BFBE8E26A" blockId="29.[151,1118,746,799]" box="[151,1118,746,768]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="62828674FF88FF86FF3DFD4BFBE8E26A" box="[151,1118,746,768]" pageId="29" pageNumber="30">
<taxonomicName id="8592DAD9FF88FF86FF3DFD4BFCF8E26A" ID-CoL="94733" authority="Yao &amp; Li, 2016 in Dong et al. 2016: 208" box="[151,846,746,768]" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kottawagamaensis">
<emphasis id="70E67D48FF88FF86FF3DFD4BFE1CE26A" box="[151,426,746,768]" italics="true" pageId="29" pageNumber="30">Pholcus kottawagamaensis</emphasis>
<bibRefCitation id="2603DCABFF88FF86FE1AFD4AFDF8E26A" author="Yao, Z. &amp; Dong, T. &amp; Zheng, G. &amp; Fu, J. &amp; Li, S." box="[432,590,746,768]" pageId="29" pageNumber="30" pagination="35757" refId="ref29041" refString="Yao, Z., Dong, T., Zheng, G., Fu, J. &amp; Li, S. (2016) High endemism at cave entrances: a case study of spiders of the genus Uthina. Scientific Reports, 6, 35757. https: // doi. org / 10.1038 / srep 35757" type="journal article" year="2016">
Yao &amp; Li,
<quantity id="856A0CBFFF88FF86FDB1FD4BFDF8E26A" box="[539,590,746,768]" metricMagnitude="1" metricUnit="m" metricValue="5.12064" pageId="29" pageNumber="30" unit="in" value="2016.0">2016</quantity>
</bibRefCitation>
in
<treatmentCitation id="C333874BFF88FF86FDC4FD4AFCF8E26A" ID-DOI="http://doi.org/10.5281/zenodo.3508796" ID-Zenodo-Dep="3508796" author="Dong, T. &amp; Zheng, G. &amp; Yao, Z. &amp; Li, S." box="[622,846,746,768]" httpUri="http://treatment.plazi.org/id/110D0059FFDC2B14FF08FAB94F8844DD" page="208" pageId="29" pageNumber="30" year="2016">
<bibRefCitation id="2603DCABFF88FF86FDC4FD4AFCAAE26A" author="Dong, T. &amp; Zheng, G. &amp; Yao, Z. &amp; Li, S." box="[622,796,746,768]" pageId="29" pageNumber="30" pagination="201 - 246" refId="ref26568" refString="Dong, T., Zheng, G., Yao, Z. &amp; Li, S. (2016) Fifteen new species of the spider genus Pholcus (Araneae: Pholcidae) from Southeast Asia. Zootaxa, 4136 (2), 201 - 246. https: // doi. org / 10.11646 / zootaxa. 4136.2.1" type="journal article" year="2016">
Dong
<emphasis id="70E67D48FF88FF86FD06FD4AFD56E26A" box="[684,736,746,768]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2016
</bibRefCitation>
: 208
</treatmentCitation>
</taxonomicName>
, figs 5–6 (
<specimenCount id="54946AD3FF88FF86FC10FD4BFC67E26A" box="[954,977,746,768]" pageId="29" pageNumber="30" type="male">♂</specimenCount>
<specimenCount id="54946AD3FF88FF86FC7BFD4BFC57E26A" box="[977,993,746,768]" pageId="29" pageNumber="30" type="female">♀</specimenCount>
,
<collectingCountry id="3A85E1CAFF88FF86FC46FD4BFBE2E26A" box="[1004,1108,746,768]" name="Sri Lanka" pageId="29" pageNumber="30">Sri Lanka</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="422DA15AFF88FF86FF3DFCA8FBECE275" blockId="29.[151,1118,746,799]" box="[151,1114,777,799]" pageId="29" pageNumber="30">
<taxonomicName id="8592DAD9FF88FF86FF3DFCA8FE7DE275" ID-CoL="577S3" baseAuthorityName="Yao &amp; Li" baseAuthorityYear="2016" box="[151,459,777,799]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kottawagamaensis">
<emphasis id="70E67D48FF88FF86FF3DFCA8FE7DE275" box="[151,459,777,799]" italics="true" pageId="29" pageNumber="30">Tissahamia kottawagamaensis</emphasis>
</taxonomicName>
—
<bibRefCitation id="2603DCABFF88FF86FE4FFCA8FD2FE275" author="Eberle, J. &amp; Dimitrov, D. &amp; Valdez-Mondragon, A. &amp; Huber, B. A." box="[485,665,777,799]" pageId="29" pageNumber="30" pagination="141" refId="ref26630" refString="Eberle, J., Dimitrov, D., Valdez-Mondragon, A. &amp; Huber, B. A. (2018) Microhabitat change drives diversification in pholcid spiders. BMC Evolutionary Biology, 18, 141. https: // doi. org / 10.1186 / s 12862 - 018 - 1244 - 8" type="journal article" year="2018">
Eberle
<emphasis id="70E67D48FF88FF86FD87FCABFDD7E275" box="[557,609,777,799]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2018
</bibRefCitation>
(molecular data).
<bibRefCitation id="2603DCABFF88FF86FCF8FCA8FBB1E275" author="Huber, B. A. &amp; Eberle, J. &amp; Dimitrov, D." box="[850,1031,777,799]" pageId="29" pageNumber="30" pagination="51 - 101" refId="ref27545" refString="Huber, B. A., Eberle, J. &amp; Dimitrov, D. (2018) The phylogeny of pholcid spiders (Araneae, Pholcidae): a critical evaluation of relationships suggested by molecular data. ZooKeys, 789, 51 - 101. https: // doi. org / 10.3897 / zookeys. 789.22781" type="journal article" year="2018">
Huber
<emphasis id="70E67D48FF88FF86FC32FCABFC7DE275" box="[920,971,777,799]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2018
</bibRefCitation>
: fig. 10.
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FF88FF85FF3DFCE7FE85E4DC" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="422DA15AFF88FF86FF3DFCE7FD80E515" blockId="29.[151,1436,838,1151]" pageId="29" pageNumber="30">
<emphasis id="70E67D48FF88FF86FF3DFCE7FE79E235" bold="true" box="[151,463,838,863]" pageId="29" pageNumber="30">Diagnosis and Description</emphasis>
(amendments; see
<bibRefCitation id="2603DCABFF88FF86FD0CFCE6FCD0E235" author="Dong, T. &amp; Zheng, G. &amp; Yao, Z. &amp; Li, S." box="[678,870,838,863]" pageId="29" pageNumber="30" pagination="201 - 246" refId="ref26568" refString="Dong, T., Zheng, G., Yao, Z. &amp; Li, S. (2016) Fifteen new species of the spider genus Pholcus (Araneae: Pholcidae) from Southeast Asia. Zootaxa, 4136 (2), 201 - 246. https: // doi. org / 10.11646 / zootaxa. 4136.2.1" type="journal article" year="2016">
Dong
<emphasis id="70E67D48FF88FF86FD41FCE6FC95E235" box="[747,803,838,863]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2016
</bibRefCitation>
). Procursus elements in principle as in
<taxonomicName id="8592DAD9FF88FF86FAB0FCE6FA2CE234" authorityName="in Huber" authorityYear="2011" box="[1306,1434,839,862]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="maturata">
<emphasis id="70E67D48FF88FF86FAB0FCE6FA2CE234" box="[1306,1434,839,862]" italics="true" pageId="29" pageNumber="30">T. maturata</emphasis>
</taxonomicName>
and
<taxonomicName id="8592DAD9FF88FF86FF6CFCCDFE87E2E9" authorityName="Huber" authorityYear="2019" box="[198,305,874,899]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="karuna">
<emphasis id="70E67D48FF88FF86FF6CFCCDFE87E2E9" box="[198,305,874,899]" italics="true" pageId="29" pageNumber="30">T. karuna</emphasis>
</taxonomicName>
, with retrolateral membrane, bifid dorsal process, ventral ridge and pocket, and distal hinged sclerite (
<figureCitation id="DAA9BDDFFF88FF86FF35FC2FFE8AE2CD" box="[159,316,910,935]" captionStart="FIGURES 110–115" captionStartId="28.[151,264,1879,1902]" captionTargetBox="[202,1418,342,1855]" captionTargetId="figure@28.[164,1427,332,1861]" captionTargetPageId="28" captionText="FIGURES 110–115. Left procursi in prolateral (upper row) and retrolateral views, all images at same scale. 110–111. Tissahamia karuna sp. n., from Labugama (ZFMK Ar 20066); asterisk marks area where males from Mitirigala have a small conical process. 112–113. T. maturata (Huber, 2011), from Hakgala (ZFMK Ar 20069); arrow points at distinctive prolateral process. 114–115. T. kottawagamaensis (Yao &amp; Li, 2016), from Kottawa (ZFMK Ar 20072). Abbreviations: bp, bifid process; hs, hinged sclerite; m, membrane; po, pocket; r, ridge; ta, tarsus. Scale line: 0.5 mm." httpUri="https://zenodo.org/record/2625066/files/figure.png" pageId="29" pageNumber="30">Figs 114–115</figureCitation>
). Differs from both species by prolateral ridge narrow and ending in bifid process with one part very small; by very large retrolateral membrane and distal elements; by prolateral process between proximal and distal elements weakly sclerotized. Females differ from
<taxonomicName id="8592DAD9FF88FF86FD62FC76FCFDE284" authorityName="in Huber" authorityYear="2011" box="[712,843,983,1006]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="maturata">
<emphasis id="70E67D48FF88FF86FD62FC76FCFDE284" box="[712,843,983,1006]" italics="true" pageId="29" pageNumber="30">T. maturata</emphasis>
</taxonomicName>
and
<taxonomicName id="8592DAD9FF88FF86FC28FC76FC5AE285" authorityName="Huber" authorityYear="2019" box="[898,1004,982,1007]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="karuna">
<emphasis id="70E67D48FF88FF86FC28FC76FC5AE285" box="[898,1004,982,1007]" italics="true" pageId="29" pageNumber="30">T. karuna</emphasis>
</taxonomicName>
by large triangular (rather that oval or trapezoidal) epigynal plate and by large V-shaped (rather than evenly rounded) internal arc (‘valve’) visible through cuticle in front of epigynal plate (
<figureCitation id="DAA9BDDFFF88FF86FDC0FBBFFD7AE55D" box="[618,716,1054,1079]" captionStart="FIGURES 120–125" captionStartId="30.[151,264,1133,1156]" captionTargetBox="[151,1436,193,1113]" captionTargetId="figure@30.[151,1436,193,1113]" captionTargetPageId="30" captionText="FIGURES 120–125. Epigyna in ventral views (upper row) and cleared female genitalia in dorsal views. 120–121. Tissahamia karuna sp. n., from Labugama (ZFMK Ar 20067); arrow points at internal arc (‘valve’). 122–123. T. maturata (Huber, 2011), from Hakgala (ZFMK Ar 20070). 124–125. T. kottawagamaensis (Yao &amp; Li, 2016), from Kottawa (ZFMK Ar 20073). Abbreviation: pp, pore plate. At various scales." httpUri="https://zenodo.org/record/2625072/files/figure.png" pageId="29" pageNumber="30">Fig. 124</figureCitation>
). Tibia
<quantity id="856A0CBFFF88FF86FC8AFBBEFCFDE55D" box="[800,843,1054,1079]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="29" pageNumber="30" unit="in" value="1.0">1 in</quantity>
8 newly examined males: 8.1–11.0 (mean 9.6); in
<specimenCount id="54946AD3FF88FF86FA2AFBBEFF45E531" pageId="29" pageNumber="30" type="female">13 females</specimenCount>
: 6.7–8.4 (mean 7.8). Internal female genitalia long, with elongated pore plates surrounded by weakly sclerotized cuticular folds (
<figureCitation id="DAA9BDDFFF88FF86FE6DFBC7FD9CE515" box="[455,554,1126,1151]" captionStart="FIGURES 120–125" captionStartId="30.[151,264,1133,1156]" captionTargetBox="[151,1436,193,1113]" captionTargetId="figure@30.[151,1436,193,1113]" captionTargetPageId="30" captionText="FIGURES 120–125. Epigyna in ventral views (upper row) and cleared female genitalia in dorsal views. 120–121. Tissahamia karuna sp. n., from Labugama (ZFMK Ar 20067); arrow points at internal arc (‘valve’). 122–123. T. maturata (Huber, 2011), from Hakgala (ZFMK Ar 20070). 124–125. T. kottawagamaensis (Yao &amp; Li, 2016), from Kottawa (ZFMK Ar 20073). Abbreviation: pp, pore plate. At various scales." httpUri="https://zenodo.org/record/2625072/files/figure.png" pageId="29" pageNumber="30">Fig. 125</figureCitation>
).
</paragraph>
<caption id="16EDF1D2FF88FF86FF3DF821FAA2E698" httpUri="https://zenodo.org/record/2625070/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,264,1920,1943]" targetBox="[155,1435,1202,1900]" targetPageId="29">
<paragraph id="422DA15AFF88FF86FF3DF821FAA2E698" blockId="29.[151,1437,1920,2034]" pageId="29" pageNumber="30">
<emphasis id="70E67D48FF88FF86FF3DF821FED3E6FF" bold="true" box="[151,357,1920,1943]" pageId="29" pageNumber="30">FIGURES 116–119</emphasis>
. 116–117.
<taxonomicName id="8592DAD9FF88FF86FE73F821FD2AE6FC" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" box="[473,668,1920,1942]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="karuna" status="sp. nov.">
<emphasis id="70E67D48FF88FF86FE73F821FD2AE6FC" box="[473,668,1920,1942]" italics="true" pageId="29" pageNumber="30">Tissahamia karuna</emphasis>
</taxonomicName>
<emphasis id="70E67D48FF88FF86FD0EF820FD69E6FC" bold="true" box="[676,735,1921,1942]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="6BD5C033FF88FF86FD0EF820FD69E6FC" box="[676,735,1921,1942]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, from Labugama (ZFMK Ar 20066), male chelicerae, frontal view; and processes of left genital bulb, prolateral view. 118–119.
<taxonomicName id="8592DAD9FF88FF86FD52F801FBB4E6DF" authority="(Huber, 2011)" authorityName="Huber" authorityYear="2011" baseAuthorityName="Huber" baseAuthorityYear="2011" box="[760,1026,1951,1973]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="maturata">
<emphasis id="70E67D48FF88FF86FD52F801FCD8E6DF" box="[760,878,1952,1973]" italics="true" pageId="29" pageNumber="30">T. maturata</emphasis>
(Huber, 2011)
</taxonomicName>
from Hakgala (ZFMK Ar 20069) and
<taxonomicName id="8592DAD9FF88FF86FA23F801FDBAE6B9" authority="(Yao &amp; Li, 2016)" authorityName="Yao &amp; Li" authorityYear="2016" baseAuthorityName="Yao &amp; Li" baseAuthorityYear="2016" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kottawagamaensis">
<emphasis id="70E67D48FF88FF86FA23F801FEE4E6B9" italics="true" pageId="29" pageNumber="30">T. kottawagamaensis</emphasis>
(Yao &amp; Li, 2016)
</taxonomicName>
from Kottawa (ZFMK Ar 20072), processes of left genital bulbs in prolateral views (all genital bulbs at same scale). Abbreviations: a, ‘appendix’ (main bulbal process); e, embolus. Scale lines: 0.3 mm.
</paragraph>
</caption>
<caption id="16EDF1D2FF8BFF85FF3DFBCCFDC3E58A" httpUri="https://zenodo.org/record/2625072/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,264,1133,1156]" targetBox="[151,1436,193,1113]" targetPageId="30">
<paragraph id="422DA15AFF8BFF85FF3DFBCCFDC3E58A" blockId="30.[151,1437,1133,1248]" pageId="30" pageNumber="31">
<emphasis id="70E67D48FF8BFF85FF3DFBCCFED0E5E9" bold="true" box="[151,358,1133,1156]" pageId="30" pageNumber="31">FIGURES 120–125</emphasis>
. Epigyna in ventral views (upper row) and cleared female genitalia in dorsal views. 120–121.
<taxonomicName id="8592DAD9FF8BFF85FA83FBCFFF69E5C9" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="karuna" status="sp. nov.">
<emphasis id="70E67D48FF8BFF85FA83FBCFFF69E5C9" italics="true" pageId="30" pageNumber="31">Tissahamia karuna</emphasis>
</taxonomicName>
<emphasis id="70E67D48FF8BFF85FF4FFB2CFEA9E5C8" bold="true" box="[229,287,1165,1186]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="6BD5C033FF8BFF85FF4FFB2CFEA9E5C8" box="[229,287,1165,1186]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, from Labugama (ZFMK Ar 20067); arrow points at internal arc (‘valve’). 122–123.
<taxonomicName id="8592DAD9FF8BFF85FB20FB2FFA2EE5C9" authority="(Huber, 2011)" authorityName="Huber" authorityYear="2011" baseAuthorityName="Huber" baseAuthorityYear="2011" box="[1162,1432,1165,1187]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="maturata">
<emphasis id="70E67D48FF8BFF85FB20FB2FFAB6E5C9" box="[1162,1280,1166,1187]" italics="true" pageId="30" pageNumber="31">T. maturata</emphasis>
(Huber, 2011)
</taxonomicName>
, from Hakgala (ZFMK Ar 20070). 124–125.
<taxonomicName id="8592DAD9FF8BFF85FDD3FB0DFBA9E5AB" authority="(Yao &amp; Li, 2016)" authorityName="Yao &amp; Li" authorityYear="2016" baseAuthorityName="Yao &amp; Li" baseAuthorityYear="2016" box="[633,1055,1195,1217]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="kottawagamaensis">
<emphasis id="70E67D48FF8BFF85FDD3FB0DFCE4E5AB" box="[633,850,1195,1217]" italics="true" pageId="30" pageNumber="31">T. kottawagamaensis</emphasis>
(Yao &amp; Li, 2016)
</taxonomicName>
, from Kottawa (ZFMK Ar 20073). Abbreviation: pp, pore plate. At various scales.
</paragraph>
</caption>
<paragraph id="422DA15AFF8BFF85FF6DFAADFE85E4DC" blockId="30.[151,1437,1292,1606]" pageId="30" pageNumber="31">
<emphasis id="70E67D48FF8BFF85FF6DFAADFEE9E44F" bold="true" box="[199,351,1292,1317]" pageId="30" pageNumber="31">New records</emphasis>
.
<materialsCitation id="F2FAAB07FF8BFF85FEC5FAACFD71E44F" ID-GBIF-Occurrence="2234226363" box="[367,711,1293,1317]" collectionCode="NMSL" country="Sri Lanka" location="SRI LANKA" pageId="30" pageNumber="31" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingCountry id="3A85E1CAFF8BFF85FEC5FAACFDBCE44F" box="[367,522,1293,1317]" name="Sri Lanka" pageId="30" pageNumber="31">SRI LANKA</collectingCountry>
:
<specimenCount id="54946AD3FF8BFF85FDBCFAACFD8FE44F" box="[534,569,1293,1317]" pageId="30" pageNumber="31" type="male">1♂</specimenCount>
<specimenCount id="54946AD3FF8BFF85FDE8FAACFDD3E44F" box="[578,613,1293,1317]" pageId="30" pageNumber="31" type="female">1♀</specimenCount>
,
<collectionCode id="2483399FFF8BFF85FDDCFAACFD71E44F" box="[630,711,1293,1317]" country="Sri Lanka" httpUri="http://grbio.org/cool/9670-f832" name="National Museum of Sri Lanka" pageId="30" pageNumber="31">NMSL</collectionCode>
</materialsCitation>
,
<materialsCitation id="F2FAAB07FF8BFF85FD7EFAADFB85E44F" ID-GBIF-Occurrence="2234226369" box="[724,1075,1292,1317]" collectionCode="ZFMK" collectorName="Ar" country="Sri Lanka" pageId="30" pageNumber="31" specimenCount="10" specimenCount-female="5" specimenCount-male="5">
<specimenCount id="54946AD3FF8BFF85FD7EFAADFD40E44F" box="[724,758,1292,1317]" pageId="30" pageNumber="31" type="male">5♂</specimenCount>
<specimenCount id="54946AD3FF8BFF85FD55FAADFC92E44F" box="[767,804,1292,1317]" pageId="30" pageNumber="31" type="female">5♀</specimenCount>
,
<collectionCode id="2483399FFF8BFF85FC99FAAFFC37E44F" box="[819,897,1294,1317]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="30" pageNumber="31" type="Museum">ZFMK</collectionCode>
(
<collectorName id="EF67C48CFF8BFF85FC3BFAACFC06E44F" box="[913,944,1293,1317]" pageId="30" pageNumber="31">Ar</collectorName>
20072–73)
</materialsCitation>
and
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<specimenCount id="54946AD3FF8BFF85FBC4FAACFB27E44F" box="[1134,1169,1293,1317]" pageId="30" pageNumber="31" type="female">1♀</specimenCount>
2 juvs in pure ethanol,
<collectionCode id="2483399FFF8BFF85FF3DFA93FF53E423" box="[151,229,1330,1353]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="30" pageNumber="31" type="Museum">ZFMK</collectionCode>
(
<specimenCode id="12340921FF8BFF85FF59FA90FEF4E420" box="[243,322,1329,1354]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="30" pageNumber="31" type="Museum">SL158</specimenCode>
),
<collectingRegion id="80566FB8FF8BFF85FEFFFA90FD9EE420" box="[341,552,1329,1354]" country="Sri Lanka" name="Southern" pageId="30" pageNumber="31">Southern Province</collectingRegion>
,
<location id="474DF781FF8BFF85FD98FA93FD6BE423" LSID="urn:lsid:plazi:treatment:CA3B104CFF88FF85FF3DFD24FBBDE72C:474DF781FF8BFF85FD98FA93FD6BE423" box="[562,733,1330,1353]" country="Sri Lanka" latitude="6.097" longLatPrecision="78" longitude="80.308" name="Kottawa Forest" pageId="30" pageNumber="31" stateProvince="Southern">Kottawa Forest</location>
(
<geoCoordinate id="27A6C79DFF8BFF85FD41FA90FCF8E420" box="[747,846,1329,1354]" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="55" value="6.097">6.097°N</geoCoordinate>
,
<geoCoordinate id="27A6C79DFF8BFF85FCF2FA90FC70E423" box="[856,966,1329,1353]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="55" value="80.308">80.308°E</geoCoordinate>
),
<elevation id="C9BF4669FF8BFF85FC72FA90FBFBE420" box="[984,1101,1329,1354]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="30" pageNumber="31" unit="m" value="60.0">
<quantity id="856A0CBFFF8BFF85FC72FA90FBA7E423" box="[984,1041,1329,1354]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="30" pageNumber="31" unit="m" value="60.0">60 m</quantity>
a.s.l.
</elevation>
,
<date id="362C879AFF8BFF85FBF2FA90FB64E420" box="[1112,1234,1329,1354]" pageId="30" pageNumber="31" value="2017-03-17">
<collectingDate id="26687E72FF8BFF85FBF2FA90FB64E420" box="[1112,1234,1329,1354]" pageId="30" pageNumber="31" value="2017-03-17">17.iii.2017</collectingDate>
</date>
(
<collectorName id="EF67C48CFF8BFF85FB4BFA90FAD1E420" box="[1249,1383,1329,1354]" pageId="30" pageNumber="31">B.A. Huber</collectorName>
)
</materialsCitation>
.
<materialsCitation id="F2FAAB07FF8BFF85FAD3FA90FE97E407" ID-GBIF-Occurrence="2234226225" collectionCode="RMNH" country="Sri Lanka" pageId="30" pageNumber="31" specimenCount="5" specimenCount-female="2" specimenCount-male="3">
<specimenCount id="54946AD3FF8BFF85FAD3FA90FA2BE423" box="[1401,1437,1329,1353]" pageId="30" pageNumber="31" type="male">3♂</specimenCount>
<specimenCount id="54946AD3FF8BFF85FF3DFAF4FF0DE407" box="[151,187,1365,1389]" pageId="30" pageNumber="31" type="female">2♀</specimenCount>
,
<collectionCode id="2483399FFF8BFF85FF60FAF7FE97E407" box="[202,289,1366,1389]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="30" pageNumber="31" type="Museum">RMNH</collectionCode>
</materialsCitation>
,
<materialsCitation id="F2FAAB07FF8BFF85FE84FAF7FC23E404" ID-GBIF-Occurrence="2234226318" box="[302,917,1365,1390]" collectingDate="1982-10-15" collectionCode="RMNH" collectorName="F. Wanless" country="Sri Lanka" pageId="30" pageNumber="31" specimenCount="1">
same locality, “under leaves”,
<date id="362C879AFF8BFF85FD21FAF4FD4DE404" box="[651,763,1365,1390]" pageId="30" pageNumber="31" value="1982-10-15">
<collectingDate id="26687E72FF8BFF85FD21FAF4FD4DE404" box="[651,763,1365,1390]" pageId="30" pageNumber="31" value="1982-10-15">15.x.1982</collectingDate>
</date>
(
<collectorName id="EF67C48CFF8BFF85FCA7FAF4FC3BE404" box="[781,909,1365,1390]" pageId="30" pageNumber="31">F. Wanless</collectorName>
)
</materialsCitation>
.
<materialsCitation id="F2FAAB07FF8BFF85FC0BFAF4FB61E407" ID-GBIF-Occurrence="2234226119" box="[929,1239,1365,1389]" collectionCode="ZFMK" collectorName="Ar" country="Sri Lanka" pageId="30" pageNumber="31" specimenCount="11" specimenCount-female="8" specimenCount-male="3">
<specimenCount id="54946AD3FF8BFF85FC0BFAF4FC72E407" box="[929,964,1365,1389]" pageId="30" pageNumber="31" type="male">3♂</specimenCount>
<specimenCount id="54946AD3FF8BFF85FC67FAF4FC47E407" box="[973,1009,1365,1389]" pageId="30" pageNumber="31" type="female">8♀</specimenCount>
,
<collectionCode id="2483399FFF8BFF85FBAAFAF7FBF8E407" box="[1024,1102,1366,1389]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="30" pageNumber="31" type="Museum">ZFMK</collectionCode>
(
<collectorName id="EF67C48CFF8BFF85FBF5FAF4FBC8E407" box="[1119,1150,1365,1389]" pageId="30" pageNumber="31">Ar</collectorName>
20074)
</materialsCitation>
and
<materialsCitation id="F2FAAB07FF8BFF85FABBFAF4FE99E4DC" ID-GBIF-Occurrence="2234226135" collectingDate="2017-03-17" collectionCode="ZFMK" collectorName="B. A. Huber" country="Sri Lanka" elevation="120" latitude="6.099" location="Viharekele Forest" longLatPrecision="78" longitude="80.594" pageId="30" pageNumber="31" specimenCode="SL153" specimenCount="3" specimenCount-female="2" stateProvince="Southern">
<specimenCount id="54946AD3FF8BFF85FABBFAF4FA82E407" box="[1297,1332,1365,1389]" pageId="30" pageNumber="31" type="female">2♀</specimenCount>
<specimenCount id="54946AD3FF8BFF85FA97FAF4FAC8E404" box="[1341,1406,1365,1390]" pageId="30" pageNumber="31" type="generic">1 juv.</specimenCount>
in pure ethanol,
<collectionCode id="2483399FFF8BFF85FE9AFADBFEC8E4FB" box="[304,382,1402,1425]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="30" pageNumber="31" type="Museum">ZFMK</collectionCode>
(
<specimenCode id="12340921FF8BFF85FE24FAD8FE6AE4F8" box="[398,476,1401,1426]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="30" pageNumber="31" type="Museum">SL153</specimenCode>
),
<collectingRegion id="80566FB8FF8BFF85FE5BFAD8FD72E4F8" box="[497,708,1401,1426]" country="Sri Lanka" name="Southern" pageId="30" pageNumber="31">Southern Province</collectingRegion>
,
<location id="474DF781FF8BFF85FD65FAD8FC23E4FB" LSID="urn:lsid:plazi:treatment:CA3B104CFF88FF85FF3DFD24FBBDE72C:474DF781FF8BFF85FD65FAD8FC23E4FB" box="[719,917,1401,1426]" country="Sri Lanka" latitude="6.099" longLatPrecision="78" longitude="80.594" name="Viharekele Forest" pageId="30" pageNumber="31" stateProvince="Southern">Viharekele Forest</location>
(
<geoCoordinate id="27A6C79DFF8BFF85FC0EFAD8FBB1E4F8" box="[932,1031,1401,1426]" direction="north" orientation="latitude" pageId="30" pageNumber="31" precision="55" value="6.099">6.099°N</geoCoordinate>
,
<geoCoordinate id="27A6C79DFF8BFF85FBB8FAD8FBC9E4F8" box="[1042,1151,1401,1426]" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="55" value="80.594">80.594°E</geoCoordinate>
),
<elevation id="C9BF4669FF8BFF85FB38FAD8FAA0E4F8" box="[1170,1302,1401,1426]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="31" unit="m" value="120.0">
<quantity id="856A0CBFFF8BFF85FB38FAD8FB6CE4FB" box="[1170,1242,1401,1425]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="30" pageNumber="31" unit="m" value="120.0">120 m</quantity>
a.s.l.
</elevation>
,
<date id="362C879AFF8BFF85FA89FAD8FA2AE4F8" box="[1315,1436,1401,1426]" pageId="30" pageNumber="31" value="2017-03-17">
<collectingDate id="26687E72FF8BFF85FA89FAD8FA2AE4F8" box="[1315,1436,1401,1426]" pageId="30" pageNumber="31" value="2017-03-17">17.iii.2017</collectingDate>
</date>
(
<collectorName id="EF67C48CFF8BFF85FF0AFA3CFE91E4DC" box="[160,295,1437,1462]" pageId="30" pageNumber="31">B.A. Huber</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FF8BFF85FF6DFA60FCF5E748" pageId="30" pageNumber="31" type="description">
<paragraph id="422DA15AFF8BFF85FF6DFA60FCF5E748" blockId="30.[151,1437,1292,1606]" pageId="30" pageNumber="31">
<emphasis id="70E67D48FF8BFF85FF6DFA60FEC8E4B0" bold="true" box="[199,382,1473,1498]" pageId="30" pageNumber="31">Natural history</emphasis>
. The spiders built their domed webs with the apex connected to the undersides of live leaves. The webs were conspicuous (like in
<taxonomicName id="8592DAD9FF8BFF85FDECFA47FD71E494" baseAuthorityName="Huber" baseAuthorityYear="2011" box="[582,711,1509,1534]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="ethagala">
<emphasis id="70E67D48FF8BFF85FDECFA47FD71E494" box="[582,711,1509,1534]" italics="true" pageId="30" pageNumber="31">T. ethagala</emphasis>
</taxonomicName>
rather than in
<taxonomicName id="8592DAD9FF8BFF85FCD0FA47FC48E497" baseAuthorityName="Huber" baseAuthorityYear="2011" box="[890,1022,1510,1533]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="maturata">
<emphasis id="70E67D48FF8BFF85FCD0FA47FC48E497" box="[890,1022,1510,1533]" italics="true" pageId="30" pageNumber="31">T. maturata</emphasis>
</taxonomicName>
) and extended far beyond the leaf. Sometimes the webs were shared by many cecidomyiid flies.
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FF8BFF85FF6DF98CFBBDE72C" box="[199,1035,1581,1606]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="422DA15AFF8BFF85FF6DF98CFBBDE72C" blockId="30.[151,1437,1292,1606]" box="[199,1035,1581,1606]" pageId="30" pageNumber="31">
<emphasis id="70E67D48FF8BFF85FF6DF98CFEEFE72C" bold="true" box="[199,345,1581,1606]" pageId="30" pageNumber="31">Distribution</emphasis>
. Known from two localities in southern
<collectingCountry id="3A85E1CAFF8BFF85FCB5F98CFC3BE72C" box="[799,909,1581,1606]" name="Sri Lanka" pageId="30" pageNumber="31">Sri Lanka</collectingCountry>
(
<figureCitation id="DAA9BDDFFF8BFF85FC36F98CFC48E72C" box="[924,1022,1581,1606]" captionStart="FIGURES 224–225" captionStartId="52.[151,264,1873,1896]" captionTargetBox="[151,1436,1159,1861]" captionTargetId="figure@52.[151,1436,1159,1861]" captionTargetPageId="52" captionText="FIGURES 224–225. Known distribution of Tissahamia in Sri Lanka; “?”, female with uncertain identity from Eastern Sinharaja." httpUri="https://zenodo.org/record/2625102/files/figure.png" pageId="30" pageNumber="31">Fig. 225</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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