treatments-xml/data/B7/28/35/B72835F5547853F79B2490B5F70739C2.xml

<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.156.49137" ID-GBIF-Dataset="c0b681b1-f311-4bed-a025-4c1793cc7645" ID-PMC="PMC7455574" ID-Pensoft-Pub="1314-2003-156-103" ID-Pensoft-UUID="AAD4849367965E749B4876EC50F920E0" ID-PubMed="32913411" ModsDocID="1314-2003-156-103" checkinTime="1598112938897" checkinUser="pensoft" docAuthor="Sun, Jiao, Zhang, Dai-Gui, Huang, Xian-Han, Tojibaev, Komiljon, Yang, Jing-Yuan, Wang, Heng-Chang &amp; Deng, Tao" docDate="2020" docId="B72835F5547853F79B2490B5F70739C2" docLanguage="en" docName="PhytoKeys 156: 103-112" docOrigin="PhytoKeys 156" docSource="http://dx.doi.org/10.3897/phytokeys.156.49137" docTitle="Primula sunhangii T. Deng, D. G. Zhang &amp; Jiao Sun 2020, sp. nov." docType="treatment" docVersion="4" id="AAD4849367965E749B4876EC50F920E0" lastPageNumber="103" masterDocId="AAD4849367965E749B4876EC50F920E0" masterDocTitle="Primula sunhangii (Primulaceae): a new species from Hubei, Central China" masterLastPageNumber="112" masterPageNumber="103" pageNumber="103" updateTime="1668139928596" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Primula sunhangii (Primulaceae): a new species from Hubei, Central China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sun, Jiao</mods:namePart>
<mods:affiliation>CAS Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, Hubei 430074, China &amp; CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China &amp; University of Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5028-8132</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Dai-Gui</mods:namePart>
<mods:affiliation>CAS Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, Hubei 430074, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Huang, Xian-Han</mods:namePart>
<mods:affiliation>CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tojibaev, Komiljon</mods:namePart>
<mods:affiliation>Central Herbarium of Uzbekistan, Institute of Botany, Academy Sciences of Uzbekistan, Tashkent 100025, Uzbekistan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Jing-Yuan</mods:namePart>
<mods:affiliation>Administration of Shennongjia National Park, Shennongjia, Hubei 44241, China</mods:affiliation>
<mods:nameIdentifier type="email">290753238@qq.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Heng-Chang</mods:namePart>
<mods:affiliation>CAS Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan, Hubei 430074, China</mods:affiliation>
<mods:nameIdentifier type="email">hcwang@wbgcas.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Deng, Tao</mods:namePart>
<mods:affiliation>CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
<mods:nameIdentifier type="email">dengtao@mail.kib.ac.cn</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>PhytoKeys</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>156</mods:number>
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<mods:extent unit="page">
<mods:start>103</mods:start>
<mods:end>112</mods:end>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.156.49137</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.156.49137</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-156-103</mods:identifier>
<mods:identifier type="Pensoft-UUID">AAD4849367965E749B4876EC50F920E0</mods:identifier>
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<treatment ID-GBIF-Taxon="167292933" LSID="urn:lsid:plazi:treatment:B72835F5547853F79B2490B5F70739C2" httpUri="http://treatment.plazi.org/id/B72835F5547853F79B2490B5F70739C2" lastPageNumber="103" pageId="0" pageNumber="103">
<subSubSection pageId="0" pageNumber="103" type="nomenclature">
<paragraph pageId="0" pageNumber="103">
<taxonomicName LSID="B72835F5-5478-53F7-9B24-90B5F70739C2" authority="T. Deng, D. G. Zhang &amp; Jiao Sun" authorityName="T. Deng, D. G. Zhang &amp; Jiao Sun" authorityYear="2020" class="Magnoliopsida" family="Primulaceae" genus="Primula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Primula sunhangii" order="Ericales" pageId="0" pageNumber="103" phylum="Tracheophyta" rank="species" species="sunhangii" status="sp. nov.">Primula sunhangii T. Deng, D. G. Zhang &amp; Jiao Sun</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="103">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="materials_examined">
<paragraph pageId="0" pageNumber="103">Type.</paragraph>
<paragraph pageId="0" pageNumber="103">
China, Hubei Province, Shennongjia Forestry District, Hongping, Dashuping. 
<geoCoordinate degrees="31" direction="north" minutes="26.67" orientation="latitude" precision="9" value="31.4445">31°26.67'N</geoCoordinate>
, 
<geoCoordinate degrees="110" direction="east" minutes="16.01" orientation="longitude" precision="9" value="110.26683">110°16.01'E</geoCoordinate>
Alt. 2877 m. 5 Jun 2011, D. G. Zhang et al. Zdg20110605023 (Holotype 
<emphasis bold="true" pageId="0" pageNumber="103">KUN</emphasis>
!) (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Holotype of Primula sunhangii T. Deng, D. G. Zhang &amp; Jiao Sun, sp. nov." figureDoi="10.3897/phytokeys.156.49137.figure1" httpUri="https://binary.pensoft.net/fig/444819" pageId="0" pageNumber="103">1</figureCitation>
). - 
<emphasis bold="true" italics="true" pageId="0" pageNumber="103">Isotype</emphasis>
: China, Hubei Province, Shennongjia Forestry District, Hongping, Dashuping. 
<geoCoordinate degrees="31" direction="north" minutes="26.67" orientation="latitude" precision="9" value="31.4445">31°26.67'N</geoCoordinate>
, 
<geoCoordinate degrees="110" direction="east" minutes="16.01" orientation="longitude" precision="9" value="110.26683">110°16.01'E</geoCoordinate>
Alt. 2877 m. 5 Jun 2011, D. G. Zhang et al. Zdg20110605023 (
<emphasis bold="true" pageId="0" pageNumber="103">JIU</emphasis>
!).
</paragraph>
<caption doi="10.3897/phytokeys.156.49137.figure1" httpUri="https://binary.pensoft.net/fig/444819" pageId="0" pageNumber="103" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">Figure 1.</emphasis>
Holotype of 
<taxonomicName authorityName="T. Deng, D. G. Zhang &amp; Jiao Sun" authorityYear="2020" class="Magnoliopsida" family="Primulaceae" genus="Primula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Primula sunhangii" order="Ericales" pageId="0" pageNumber="103" phylum="Tracheophyta" rank="species" species="sunhangii">
<emphasis italics="true" pageId="0" pageNumber="103">Primula sunhangii</emphasis>
</taxonomicName>
T. Deng, D. G. Zhang &amp; Jiao Sun, sp. nov.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="diagnosis">
<paragraph pageId="0" pageNumber="103">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="103">
<taxonomicName authorityName="T. Deng, D. G. Zhang &amp; Jiao Sun" authorityYear="2020" class="Magnoliopsida" family="Primulaceae" genus="Primula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Primula sunhangii" order="Ericales" pageId="0" pageNumber="103" phylum="Tracheophyta" rank="species" species="sunhangii">
<emphasis italics="true" pageId="0" pageNumber="103">Primula sunhangii</emphasis>
</taxonomicName>
is similar to 
<taxonomicName lsidName="P. involucrata" pageId="0" pageNumber="103" rank="species" species="involucrata">
<emphasis italics="true" pageId="0" pageNumber="103">P. involucrata</emphasis>
</taxonomicName>
in glabrous, efarinose, ovate leaves, and lanceolate bracts base prolonged below into auriculate appendage and its length. But it differs from 
<taxonomicName lsidName="P. involucrata" pageId="0" pageNumber="103" rank="species" species="involucrata">
<emphasis italics="true" pageId="0" pageNumber="103">P. involucrata</emphasis>
</taxonomicName>
in the following characters: glabrous sepal, cylindrical calyx and capsule, truncate or acute leaves base, and shorter scape (at most 3 times the leaves).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="description">
<paragraph pageId="0" pageNumber="103">Description.</paragraph>
<paragraph pageId="0" pageNumber="103">Herbs perennial, glabrous, efarinose. Leaves 5-10, all basal in a rosette; petiole green, basal white to pink, 1-5.6 cm long, 0.2-0.4 cm diam.; leaf blade green, ovate or oblong, 0.8-4.8 cm long, 0.5-2 cm wide, papery, base truncate or acute, margin entire or slightly denticulate, apex obtuse to rounded. Scapes 5-19 cm in length; umbels 2-4-flowered; bracts 5, lanceolate, 5-10 mm long, 1-3 mm wide, membranous, base prolonged below into 3-6 mm auriculate appendage. Pedicel 0.5-3.5 cm. Calyx 5-7 mm in length, 3 mm diam., cylindrical, parted to 1/4 to 1/3; lobes lanceolate, apex narrowly acute. Corolla funnel-form, pink or purple; tube 8-10 mm in length, 8-10 mm diam., limb 1-2 cm diam.; lobes obovate, deeply emarginated apex.</paragraph>
<paragraph pageId="0" pageNumber="103">Pin flowers: corolla tube ca. 8.5 mm; stamens ca. 3.5 mm above base of corolla tube; the style is not or slightly exerted.</paragraph>
<paragraph pageId="0" pageNumber="103">Thrum flowers: corolla tube ca. 1 cm; stamens toward apex of corolla tube; the style is slightly shorter than calyx.</paragraph>
<paragraph pageId="0" pageNumber="103">
Capsule cylindric, apex irregular dehisces. (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Images of live Primula sunhangii T. Deng, D. G. Zhang &amp; Jiao Sun, sp. nov. A habitat B inflorescence C leaves D flower, front and back side E toot. Scale bars: 1 cm in (C, D, E). Photographer: Qun Liu." figureDoi="10.3897/phytokeys.156.49137.figure2" httpUri="https://binary.pensoft.net/fig/444820" pageId="0" pageNumber="103">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="distribution">
<paragraph pageId="0" pageNumber="103">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="103">
Fissures of rocks on mountain slopes; ca. 2,800 m. Shennongding (Hongping, Shennongjia, Hubei), Laojunshan (Muyu, Shennongjia, Hubei). (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of Prunus sunhangii T. Deng, D. G. Zhang &amp; Jiao Sun." figureDoi="10.3897/phytokeys.156.49137.figure3" httpUri="https://binary.pensoft.net/fig/444821" pageId="0" pageNumber="103">3</figureCitation>
)
</paragraph>
<caption doi="10.3897/phytokeys.156.49137.figure2" httpUri="https://binary.pensoft.net/fig/444820" pageId="0" pageNumber="103" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">Figure 2.</emphasis>
Images of live 
<taxonomicName authorityName="T. Deng, D. G. Zhang &amp; Jiao Sun" authorityYear="2020" class="Magnoliopsida" family="Primulaceae" genus="Primula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Primula sunhangii" order="Ericales" pageId="0" pageNumber="103" phylum="Tracheophyta" rank="species" species="sunhangii">
<emphasis italics="true" pageId="0" pageNumber="103">Primula sunhangii</emphasis>
</taxonomicName>
T. Deng, D. G. Zhang &amp; Jiao Sun, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="103">A</emphasis>
habitat 
<emphasis bold="true" pageId="0" pageNumber="103">B</emphasis>
inflorescence 
<emphasis bold="true" pageId="0" pageNumber="103">C</emphasis>
leaves 
<emphasis bold="true" pageId="0" pageNumber="103">D</emphasis>
flower, front and back side 
<emphasis bold="true" pageId="0" pageNumber="103">E</emphasis>
toot. Scale bars: 1 cm in (
<emphasis bold="true" pageId="0" pageNumber="103">C, D, E</emphasis>
). Photographer: Qun Liu.
</paragraph>
</caption>
<caption doi="10.3897/phytokeys.156.49137.figure3" httpUri="https://binary.pensoft.net/fig/444821" pageId="0" pageNumber="103" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">Figure 3.</emphasis>
Distribution of 
<taxonomicName authorityName="D.G.Zhang &amp; T.Deng" authorityYear="2019" class="Magnoliopsida" family="Rosaceae" genus="Prunus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Prunus sunhangii" order="Rosales" pageId="0" pageNumber="103" phylum="Tracheophyta" rank="species" species="sunhangii">
<emphasis italics="true" pageId="0" pageNumber="103">Prunus sunhangii</emphasis>
</taxonomicName>
T. Deng, D. G. Zhang &amp; Jiao Sun.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="phenology">
<paragraph pageId="0" pageNumber="103">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="103">
<taxonomicName authorityName="T. Deng, D. G. Zhang &amp; Jiao Sun" authorityYear="2020" class="Magnoliopsida" family="Primulaceae" genus="Primula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Primula sunhangii" order="Ericales" pageId="0" pageNumber="103" phylum="Tracheophyta" rank="species" species="sunhangii">
<emphasis italics="true" pageId="0" pageNumber="103">Primula sunhangii</emphasis>
</taxonomicName>
flowers from June to July.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="etymology">
<paragraph pageId="0" pageNumber="103">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="103">The specific epithet refers to the Chinese botanist Hang Sun.</paragraph>
<paragraph pageId="0" pageNumber="103">
Vernacular name: Simplified Chinese: 
<normalizedToken originalValue="神农架报春">神农架报春</normalizedToken>
; Chinese Pinyin: 
<normalizedToken originalValue="Shénnóngjià">Shennongjia</normalizedToken>
<normalizedToken originalValue="Bàochūn">Baochūn</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="conservation status">
<paragraph pageId="0" pageNumber="103">Conservation status and IUCN preliminary assessment.</paragraph>
<paragraph pageId="0" pageNumber="103">The new species has only two known populations described thus far. The populations are both in subareas of Shennongjia, one in Shennongding and another in Laojunshan. Each population has ca. 40 individuals. The habitats are situated in a tourist attraction zone, with an average of 20,000 tourists visiting daily. According to IUCN red list categories and criteria, conservation status of this species should be Critically Endangered (CR) (B2abiii).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="103" type="relationship with related species">
<paragraph pageId="0" pageNumber="103">Relationship with related species.</paragraph>
<paragraph pageId="0" pageNumber="103">
Based on its glabrous, efarinose, ovate leaves, lanceolate bracts base prolonged below into auriculate appendage, and its length, 
<taxonomicName lsidName="P. sunhangii" pageId="0" pageNumber="103" rank="species" species="sunhangii">
<emphasis italics="true" pageId="0" pageNumber="103">P. sunhangii</emphasis>
</taxonomicName>
is most morphologically similar to 
<taxonomicName lsidName="P. involucrata" pageId="0" pageNumber="103" rank="species" species="involucrata">
<emphasis italics="true" pageId="0" pageNumber="103">P. involucrata</emphasis>
</taxonomicName>
(
<bibRefCitation author="Hu, CM" editor="Chen, FH" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="103" pagination="1 - 245" refId="B4" refString="Hu, CM, 1990. Primula. In: Chen, FH, Hu, CM, Eds., Flora Reipublicae Popularis Sinicae, Chapter 59. Science Press, Beijing: 1 - 245" title="Primula." volumeTitle="Flora Reipublicae Popularis Sinicae, Chapter 59." year="1990">Hu 1990</bibRefCitation>
; 
<bibRefCitation author="Hu, CM" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="103" refId="B5" refString="Hu, CM, Kelso, S, 1996. Primulaceae. In: Wu ZY, Raven PH (Eds) Flora of China. Science Press &amp; Missouri Botanical Garden Press, Beijing &amp; St. Louis." title="Primulaceae. In: Wu ZY, Raven PH (Eds) Flora of China. Science Press &amp; Missouri Botanical Garden Press, Beijing &amp; St. Louis." year="1996">Hu and Kelso 1996</bibRefCitation>
). But 
<taxonomicName lsidName="P. sunhangii" pageId="0" pageNumber="103" rank="species" species="sunhangii">
<emphasis italics="true" pageId="0" pageNumber="103">P. sunhangii</emphasis>
</taxonomicName>
differs from 
<taxonomicName lsidName="P. involucrata" pageId="0" pageNumber="103" rank="species" species="involucrata">
<emphasis italics="true" pageId="0" pageNumber="103">P. involucrata</emphasis>
</taxonomicName>
in glabrous sepal, cylindrical calyx and capsule, truncate or acute leaves base, and shorter scape (at most 3 times the leaves). (Table 
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Diagnostic morphological characters comparison between Primula sunhangii and P. involucrata." httpUri="http://table.plazi.org/id/C6B088D5FF10ECD9F468E9255E901FE5" pageId="0" pageNumber="103" tableUuid="C6B088D5FF10ECD9F468E9255E901FE5">1</tableCitation>
).
</paragraph>
<caption ID-Table-UUID="C6B088D5FF10ECD9F468E9255E901FE5" httpUri="http://table.plazi.org/id/C6B088D5FF10ECD9F468E9255E901FE5" pageId="0" pageNumber="103" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">Table 1.</emphasis>
Diagnostic morphological characters comparison between 
<taxonomicName authorityName="T. Deng, D. G. Zhang &amp; Jiao Sun" authorityYear="2020" class="Magnoliopsida" family="Primulaceae" genus="Primula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Primula sunhangii" order="Ericales" pageId="0" pageNumber="103" phylum="Tracheophyta" rank="species" species="sunhangii">
<emphasis italics="true" pageId="0" pageNumber="103">Primula sunhangii</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="P. involucrata" pageId="0" pageNumber="103" rank="species" species="involucrata">
<emphasis italics="true" pageId="0" pageNumber="103">P. involucrata</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="103">
<table pageId="0" pageNumber="103">
<tr pageId="0" pageNumber="103">
<th colspan="1" pageId="0" pageNumber="103" rowspan="1">Characteristics</th>
<th colspan="1" pageId="0" pageNumber="103" rowspan="1">
<taxonomicName lsidName="P. sunhangii" pageId="0" pageNumber="103" rank="species" species="sunhangii">
<emphasis italics="true" pageId="0" pageNumber="103">P. sunhangii</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="103" rowspan="1">
<taxonomicName lsidName="P. involucrata" pageId="0" pageNumber="103" rank="species" species="involucrata">
<emphasis italics="true" pageId="0" pageNumber="103">P. involucrata</emphasis>
</taxonomicName>
</th>
</tr>
<tr pageId="0" pageNumber="103">
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">Base of leaves</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">truncate or acute</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">cuneate, rounded to slightly cordate</td>
</tr>
<tr pageId="0" pageNumber="103">
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">Sepal</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">glabrous</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">glandular ciliolate</td>
</tr>
<tr pageId="0" pageNumber="103">
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">Calyx shape</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">cylindrical</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">campanulate</td>
</tr>
<tr pageId="0" pageNumber="103">
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">Capsule shape</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">cylindrical</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">oblong</td>
</tr>
<tr pageId="0" pageNumber="103">
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">Ratio of scape to leaves</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">less than 3 times</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">3-5 times</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="103">
We performed phylogenetic analyses using nuclear ITS as well as 
<emphasis italics="true" pageId="0" pageNumber="103">rbcL</emphasis>
and 
<emphasis italics="true" pageId="0" pageNumber="103">matK</emphasis>
of 
<taxonomicName lsidName="P. sunhangii" pageId="0" pageNumber="103" rank="species" species="sunhangii">
<emphasis italics="true" pageId="0" pageNumber="103">P. sunhangii</emphasis>
</taxonomicName>
and related species. Separate analyses for all three markers get six trees (ML and Bayesian) with similar structures. Combined molecular phylogenetic analyses shows that 
<taxonomicName lsidName="P. sunhangii" pageId="0" pageNumber="103" rank="species" species="sunhangii">
<emphasis italics="true" pageId="0" pageNumber="103">P. sunhangii</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="P. involucrata" pageId="0" pageNumber="103" rank="species" species="involucrata">
<emphasis italics="true" pageId="0" pageNumber="103">P. involucrata</emphasis>
</taxonomicName>
are sister taxa, with high support (Fig. 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Bayesian tree of selected species in Primula, focused on Sect. Aleuritia. Based on a combined sequences from nuclear ITS and rbcL, matK of cpDNA genes. Numbers above the branches represent the Bayesian posterior probabilities, and below showing the maximum likelihood values. (* multifurcation in Maximum-likelihood tree)." figureDoi="10.3897/phytokeys.156.49137.figure4" httpUri="https://binary.pensoft.net/fig/444822" pageId="0" pageNumber="103">4</figureCitation>
). They have 19 different sites in the 2020 bp nucleotide sequence.
</paragraph>
<paragraph pageId="0" pageNumber="103">
These genetic results and morphological data clearly support that 
<taxonomicName authorityName="T. Deng, D. G. Zhang &amp; Jiao Sun" authorityYear="2020" class="Magnoliopsida" family="Primulaceae" genus="Primula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Primula sunhangii" order="Ericales" pageId="0" pageNumber="103" phylum="Tracheophyta" rank="species" species="sunhangii">
<emphasis italics="true" pageId="0" pageNumber="103">Primula sunhangii</emphasis>
</taxonomicName>
be recognized as a distinct new species.
</paragraph>
<caption doi="10.3897/phytokeys.156.49137.figure4" httpUri="https://binary.pensoft.net/fig/444822" pageId="0" pageNumber="103" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">Figure 4.</emphasis>
Bayesian tree of selected species in 
<taxonomicName class="Magnoliopsida" family="Primulaceae" genus="Primula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Primula" order="Ericales" pageId="0" pageNumber="103" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="103">Primula</emphasis>
</taxonomicName>
, focused on Sect. 
<taxonomicName class="Magnoliopsida" family="Primulaceae" genus="Primula" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Ericales" pageId="0" pageNumber="103" phylum="Tracheophyta" rank="section" section="Aleuritia">Aleuritia</taxonomicName>
. Based on a combined sequences from nuclear ITS and 
<emphasis italics="true" pageId="0" pageNumber="103">rbcL</emphasis>
, 
<emphasis italics="true" pageId="0" pageNumber="103">matK</emphasis>
of cpDNA genes. Numbers above the branches represent the Bayesian posterior probabilities, and below showing the maximum likelihood values. (* multifurcation in Maximum-likelihood tree).
</paragraph>
</caption>
<caption ID-Table-UUID="EEB9DFB49B0CD25CEB54EC34BB41A4F7" httpUri="http://table.plazi.org/id/EEB9DFB49B0CD25CEB54EC34BB41A4F7" pageId="0" pageNumber="103" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="103">
<emphasis bold="true" pageId="0" pageNumber="103">Table 2.</emphasis>
Primers used in PCR amplification and sequencing.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="103">
<table pageId="0" pageNumber="103">
<tr pageId="0" pageNumber="103">
<th colspan="1" pageId="0" pageNumber="103" rowspan="1">DNA fragments</th>
<th colspan="1" pageId="0" pageNumber="103" rowspan="1">Primer name</th>
<th colspan="1" pageId="0" pageNumber="103" rowspan="1">5'-3' Sequences</th>
</tr>
<tr pageId="0" pageNumber="103">
<td colspan="1" pageId="0" pageNumber="103" rowspan="2">ITS</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">ITS1</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">GTCCACTGAACCTTATCATTTAG</td>
</tr>
<tr pageId="0" pageNumber="103">
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">ITS4</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">TCCTCCGCTTATTGATATGC</td>
</tr>
<tr pageId="0" pageNumber="103">
<td colspan="1" pageId="0" pageNumber="103" rowspan="2">rbcL</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">rbcLa_f</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">ATGTCACCACAAACAGAGACTAAAGC</td>
</tr>
<tr pageId="0" pageNumber="103">
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">724R</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">TCGCATGTACCTGCAGTAGC</td>
</tr>
<tr pageId="0" pageNumber="103">
<td colspan="1" pageId="0" pageNumber="103" rowspan="2">matK</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">3F-KIM</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">CGTACAGTACTTTTGTGTTTACGAG</td>
</tr>
<tr pageId="0" pageNumber="103">
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">XF</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">TAATTTACGATCAATTCATTC</td>
</tr>
<tr pageId="0" pageNumber="103">
<td colspan="1" pageId="0" pageNumber="103" rowspan="2">trnH-psbA</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">trnH-05</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">CGCGCATGGTGGATTCACAAATC</td>
</tr>
<tr pageId="0" pageNumber="103">
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">psbA3</td>
<td colspan="1" pageId="0" pageNumber="103" rowspan="1">GTTATGCATGAACGTAATGCTC</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>