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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.284.3983" ID-PMC="PMC3677377" ID-Pensoft-Pub="1313-2970-284-1" ID-Pensoft-UUID="192EFF970E2BFFDF3161FFE53C4BFFAA" ID-PubMed="23794843" ID-Zenodo-Dep="578323" ID-ZooBank="33B15A74746D4A82A865EA1E7E55A9BB" ModsDocID="1313-2970-284-1" checkinTime="1451247469560" checkinUser="pensoft" docAuthor="Shi, Hongliang, Zhou, Hongzhang &amp; Liang, Hongbin" docDate="2013" docId="AFFE33DF9B72A09D4DA2F30555B8D155" docLanguage="en" docName="ZooKeys 284: 1-129" docOrigin="ZooKeys 284" docPubDate="2013-04-04" docSource="http://dx.doi.org/10.3897/zookeys.284.3983" docTitle="Anchista brunnea" docType="treatment" docVersion="5" id="192EFF970E2BFFDF3161FFE53C4BFFAA" lastPageNumber="24" masterDocId="192EFF970E2BFFDF3161FFE53C4BFFAA" masterDocTitle="Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera" masterLastPageNumber="129" masterPageNumber="1" pageNumber="22" updateTime="1668155586682" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Taxonomic synopsis of the subtribe Physoderina (Coleoptera, Carabidae, Lebiini), with species revisions of eight genera</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shi, Hongliang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; Graduate School of Chinese Academy of Sciences, Beijing 100039, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhou, Hongzhang</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liang, Hongbin</mods:namePart>
<mods:affiliation>Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
<mods:nameIdentifier type="email">lianghb@ioz.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-04-04</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>284</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>129</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.284.3983</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.284.3983</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-284-1</mods:identifier>
<mods:identifier type="ZooBank">33B15A74746D4A82A865EA1E7E55A9BB</mods:identifier>
<mods:identifier type="Pensoft-UUID">192EFF970E2BFFDF3161FFE53C4BFFAA</mods:identifier>
<mods:identifier type="Zenodo-Dep">578323</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152042247" LSID="urn:lsid:plazi:treatment:AFFE33DF9B72A09D4DA2F30555B8D155" httpUri="http://treatment.plazi.org/id/AFFE33DF9B72A09D4DA2F30555B8D155" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
<subSubSection pageId="21" pageNumber="22" type="nomenclature">
<paragraph pageId="21" pageNumber="22">
<taxonomicName LSID="AFFE33DF-9B72-A09D-4DA2-F30555B8D155" authority="(Wiedemann, 1823)" baseAuthorityName="Wiedemann" baseAuthorityYear="1823" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Anchista brunnea" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="brunnea">Anchista brunnea (Wiedemann, 1823)</taxonomicName>
<figureCitation captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Habitus, scale bars = 2.0 mm: 1 Paraphaea binotata (Dejean), a specimen from Yunnan 2 Paraphaea formosana (Jedlicka), a specimen from Taiwan 3 Paraphaea minor sp. n., a paratype from Hainan 4 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir from India 5 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal 6 Anchista fenestrata fenestrata (Schmidt-Goebel), a specimen from Nepal." httpUri="https://binary.pensoft.net/fig/29670" pageId="21" pageNumber="22">Habitus: Figs 4</figureCitation>
<figureCitation captionStart="Figures 37–42" captionStartId="F7" captionText="Figures 37 - 42. Type materials, scale bars = 2.0 mm: 37 Lectotype of Lebia brunnea Wiedemann [= Anchista brunnea (Wiedemann)] 38 Lectotype of Anchista picea Chaudoir [= Anchista brunnea (Wiedemann)] 39 Holotype of Plochionus fenestratus Schmidt-Goebel [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 40 Holotype of Anchista glabra Chaudoir [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 41 Lectotype of Anchista subpubescens Chaudoir [= Anchista fenestrata subpubescens Chaudoir] 42 Lectotype of Anchista nubila Andrewes." httpUri="https://binary.pensoft.net/fig/29680" pageId="21" pageNumber="22">37, 38</figureCitation>
<figureCitation captionStart="Figure 70" captionStartId="F15" captionText="Figure 70. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir." httpUri="https://binary.pensoft.net/fig/29688" pageId="21" pageNumber="22">male genitalia: Figs 70</figureCitation>
<figureCitation captionStart="Figures 95–110" captionStartId="F40" captionText="Figures 95 - 110. Left and right parameres of male genitalia, scale bars = 0.2 mm: 95 Paraphaea binotata (Dejean), a specimen from Guangxi 96 Paraphaea minor sp. n., holotype 97 Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir 98 Anchista pilosa sp. n., holotype 99 Metallanchista laticollis sp. n., holotype 100 Physodera amplicollis van de Poll, a specimen from Taiwan 101 Diamella cupreomicans (Oberthuer), a specimen from Yunnan 102 Diamella arrowi (Jedlicka), holotype 103 Allocota aurata (Bates), a specimen from Tonkin 104 Allocota aurata (Bates), a specimen from Shaanxi 105 Allocota bicolor sp. n., holotype 106 Lachnoderma asperum Bates, a specimen from Taiwan 107 Dasiosoma ivorense Basilewsky, a paratype from Cote d'Ivoire 108 Dasiosoma quadraticolle sp. n., holotype 109 Orionella kathmanduensis (Kirschenhofer), a specimen from Nepal 110 Endynomena pradieri (Fairmaire), a specimen from Fujian." httpUri="https://binary.pensoft.net/fig/29716" pageId="21" pageNumber="22">97</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="21" pageNumber="22" type="reference_group">
<paragraph pageId="21" pageNumber="22">
<taxonomicName baseAuthorityName="Wiedemann" baseAuthorityYear="1823" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="brunnea">Anchista brunnea</taxonomicName>
<bibRefCitation author="Wiedemann, CRW" journalOrPublisher="Notes from the Leyden Museum" pageId="89" pageNumber="90" refId="B87" refString="Wiedemann, CRW, 1823. Zweihundert neue Kaefer von Java, Bengalen und dem Vorgebirge der guten Hoffnung. Zoologisches Magazin2: 1-135, 162-164." title="Zweihundert neue Kaefer von Java, Bengalen und dem Vorgebirge der guten Hoffnung. Zoologisches Magazin 2: 1 - 135, 162 - 164." year="1823">Wiedemann 1823</bibRefCitation>
: 59 (original:
<taxonomicName class="Hexapoda" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="21" pageNumber="22">Lebia</emphasis>
</taxonomicName>
; type locality: Bengalia; lectotype deposited in ZMUC);
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Transactions of the Entomological Society of London" pageId="84" pageNumber="85" pagination="145 - 195" refId="B2" refString="Andrewes, HE, 1921. Notes on synonymy and on some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195" title="Notes on synonymy and on some types of Oriental Carabidae in various foreign collections." volume="1921" year="1921">Andrewes 1921</bibRefCitation>
: 172 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="21" pageNumber="22">Anchista</emphasis>
</taxonomicName>
);
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Zoologische Mededeelingen, Leiden" pageId="85" pageNumber="86" refId="B14" refString="Andrewes, HE, 1930d. Catalogue of Indian Insects, Part 18-Carabidae. Calcutta. Government of India Central Publication Branch, 389 pp." title="Catalogue of Indian Insects, Part 18 - Carabidae. Calcutta. Government of India Central Publication Branch, 389 pp." year="1930 d">Andrewes 1930d</bibRefCitation>
: 22 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="21" pageNumber="22">Anchista</emphasis>
</taxonomicName>
, catalogue);
<bibRefCitation author="Csiki, E" editor="Junk, W" journalOrPublisher="Volumen III. Carabidae III. Berlin" pageId="86" pageNumber="87" pagination="1279 - 1598" refId="B37" refString="Csiki, E, 1932. Carabidae: Harpalinae VII (Pars 124). In: Junk, W, Schenkling, S, Eds., Coleopterorum catalogus. Volumen III. Carabidae III. Berlin: 1279 - 1598" title="Carabidae: Harpalinae VII (Pars 124)." volumeTitle="Coleopterorum catalogus." year="1932">Csiki 1932</bibRefCitation>
: 1456 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="21" pageNumber="22">Anchista</emphasis>
</taxonomicName>
, catalogue).
</paragraph>
<paragraph pageId="21" pageNumber="22">
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="modesta">Anchista modesta</taxonomicName>
<bibRefCitation author="Nietner, J" journalOrPublisher="Etudes Entomologiques" pageId="88" pageNumber="89" refId="B74" refString="Nietner, J, 1856. Entomological papers, being descriptions of new Ceylon Coleoptera with such observations on their habits as appear in any way interesting. Journal of the Asiatic Society of Bengal 25 [1856]: 381-394, 523-554." title="Entomological papers, being descriptions of new Ceylon Coleoptera with such observations on their habits as appear in any way interesting. Journal of the Asiatic Society of Bengal 25 [1856]: 381 - 394, 523 - 554." year="1856">Nietner 1856</bibRefCitation>
: 523 (type locality: Ceylon; syntype deposited in MZPW);
<bibRefCitation author="Nietner, J" journalOrPublisher="Etudes Entomologiques" pageId="88" pageNumber="89" publicationUrl="10.1080/00222935708697726" refId="B75" refString="Nietner, J, 1857. Descriptions of new Ceylon Coleoptera. The Annals and Magazine of Natural History (2) 19: 241-249, 374-388. ., 10.1080/00222935708697726" title="Descriptions of new Ceylon Coleoptera. The Annals and Magazine of Natural History (2) 19: 241 - 249, 374 - 388." url="10.1080/00222935708697726" year="1857">Nietner 1857</bibRefCitation>
: 375 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="21" pageNumber="22">Anchista</emphasis>
</taxonomicName>
, redundant publication);
<bibRefCitation author="Chaudoir, BM" journalOrPublisher="Bulletin de la Societe Imperiale des Naturalistes de Moscou" pageId="86" pageNumber="87" pagination="188 - 268" refId="B35" refString="Chaudoir, BM, 1877. Genres nouveaux et especes inedites de la famille des Carabiques. Bulletin de la Societe Imperiale des Naturalistes de Moscou 52 (1): 188 - 268" title="Genres nouveaux et especes inedites de la famille des Carabiques." volume="52" year="1877">Chaudoir 1877</bibRefCitation>
: 239 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="21" pageNumber="22">Anchista</emphasis>
</taxonomicName>
);
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Spolia Zeylanica Colombo" pageId="84" pageNumber="85" pagination="129 - 141" refId="B4" refString="Andrewes, HE, 1924a. Description of some new Carabidae from Ceylon. Spolia Zeylanica Colombo 13: 129 - 141" title="Description of some new Carabidae from Ceylon." volume="13" year="1924 a">Andrewes 1924a</bibRefCitation>
: 140 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="21" pageNumber="22">Anchista</emphasis>
</taxonomicName>
, synonymized with
<emphasis italics="true" pageId="21" pageNumber="22">brunnea</emphasis>
Wiedemann);
<bibRefCitation author="Andrewes, HE" journalOrPublisher="London, The Trustees of the British Museum" pageId="84" pageNumber="85" publicationUrl="10.1080/00222932708633575" refId="B8" refString="Andrewes, HE, 1927b. Papers on Oriental Carabidae. XIX. The Annals and Magazine of Natural History (9) 19: 97-111. ., 10.1080/00222932708633575" title="Papers on Oriental Carabidae. XIX. The Annals and Magazine of Natural History (9) 19: 97 - 111." url="10.1080/00222932708633575" year="1927 b">Andrewes 1927b</bibRefCitation>
: 103 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="21" pageNumber="22">Anchista</emphasis>
</taxonomicName>
). [Synonym]
</paragraph>
<paragraph pageId="21" pageNumber="22">
<taxonomicName authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="picea">Anchista picea</taxonomicName>
<bibRefCitation author="Chaudoir, BM" journalOrPublisher="Bulletin de la Societe Imperiale des Naturalistes de Moscou" pageId="86" pageNumber="87" pagination="188 - 268" refId="B35" refString="Chaudoir, BM, 1877. Genres nouveaux et especes inedites de la famille des Carabiques. Bulletin de la Societe Imperiale des Naturalistes de Moscou 52 (1): 188 - 268" title="Genres nouveaux et especes inedites de la famille des Carabiques." volume="52" year="1877">Chaudoir 1877</bibRefCitation>
: 238 (type locality: Deccan; lectotype deposited in MNHN);
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Transactions of the Entomological Society of London" pageId="84" pageNumber="85" pagination="145 - 195" refId="B2" refString="Andrewes, HE, 1921. Notes on synonymy and on some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195" title="Notes on synonymy and on some types of Oriental Carabidae in various foreign collections." volume="1921" year="1921">Andrewes 1921</bibRefCitation>
: 172 (
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="21" pageNumber="22">Anchista</emphasis>
</taxonomicName>
, synonymized with
<emphasis italics="true" pageId="21" pageNumber="22">brunnea</emphasis>
Wiedemann). [Synonym]
</paragraph>
</subSubSection>
<subSubSection lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="type examined">
<paragraph pageId="21" pageNumber="22">Type examined.</paragraph>
<paragraph lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis bold="true" pageId="21" pageNumber="22">Lectotype</emphasis>
of
<taxonomicName class="Hexapoda" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis italics="true" pageId="21" pageNumber="22">Lebia brunnea</emphasis>
</taxonomicName>
Wiedemann, designated herein (ZMUC): female, body length = 7.4 mm, pin mounted, &quot;Mus. / Westerm.&quot;;
<normalizedToken originalValue="“Type”">&quot;Type&quot;</normalizedToken>
[red label];
<pageBreakToken pageId="22" pageNumber="23" start="start">&quot;</pageBreakToken>
<emphasis italics="true" pageId="22" pageNumber="23">Bengal. / Mai 1809. / D: brunneus. /Lebia brun / nea Wied</emphasis>
. &quot;; &quot;LECTOTYPE ♀/ Lebia brunnea / Wiedemann, 1823 / Des. SHI H. L. 2011&quot; [red label][Fig. 37].
<emphasis bold="true" pageId="22" pageNumber="23">Lectotype</emphasis>
of
<taxonomicName authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="picea">
<emphasis italics="true" pageId="22" pageNumber="23">Anchista picea</emphasis>
</taxonomicName>
Chaudoir, designated herein (MNHN), male, body length = 7.2 mm, pin mounted, &quot;Ex Musaeo, Chaudoir&quot; [red letter];
<normalizedToken originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
[red label]; &quot;MUSEUM PARIS / 1952 / COLL. R.
<normalizedToken originalValue="OBERTHÜR”">OBERTHUER&quot;</normalizedToken>
; &quot;
<emphasis italics="true" pageId="22" pageNumber="23">picea Chaudoir</emphasis>
,
<emphasis italics="true" pageId="22" pageNumber="23">Deccan</emphasis>
,
<emphasis italics="true" pageId="22" pageNumber="23">Stevens</emphasis>
&quot; [large box label but pinned under specimen]; &quot;LECTOTYPE ♂/ Anchista picea / Chaudoir, 1877 / Des. SHI H. L. 2011&quot; [red label][Fig. 38].
<emphasis bold="true" pageId="22" pageNumber="23">Paralectotype</emphasis>
of
<taxonomicName authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="picea">
<emphasis italics="true" pageId="22" pageNumber="23">Anchista picea</emphasis>
</taxonomicName>
Chaudoir (MNHN): 4 males, &quot;Ex Musaeo, Chaudoir&quot; [red letter];
<normalizedToken originalValue="“PARATYPE”">&quot;PARATYPE&quot;</normalizedToken>
[red label]; &quot;MUSEUM PARIS / 1952 / COLL. R.
<normalizedToken originalValue="OBERTHÜR”">OBERTHUER&quot;</normalizedToken>
; &quot;PARALECTOTYPE ♂/ Anchista picea / Chaudoir, 1877 / Des. SHI H. L. 2011&quot; [red label] [Figs 4, 70, 97].
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="notes on types">
<paragraph pageId="22" pageNumber="23">Notes on types.</paragraph>
<paragraph pageId="22" pageNumber="23">
<emphasis bold="true" pageId="22" pageNumber="23">
<taxonomicName class="Hexapoda" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis italics="true" pageId="22" pageNumber="23">Lebia brunnea</emphasis>
</taxonomicName>
Wiedemann
</emphasis>
: In the original literature, Wiedemann
<normalizedToken originalValue="didn’t">didn't</normalizedToken>
mention how many specimens belonged to the type series. We borrowed one female (
<figureCitation captionStart="Figures 37–42" captionStartId="F7" captionText="Figures 37 - 42. Type materials, scale bars = 2.0 mm: 37 Lectotype of Lebia brunnea Wiedemann [= Anchista brunnea (Wiedemann)] 38 Lectotype of Anchista picea Chaudoir [= Anchista brunnea (Wiedemann)] 39 Holotype of Plochionus fenestratus Schmidt-Goebel [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 40 Holotype of Anchista glabra Chaudoir [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 41 Lectotype of Anchista subpubescens Chaudoir [= Anchista fenestrata subpubescens Chaudoir] 42 Lectotype of Anchista nubila Andrewes." httpUri="https://binary.pensoft.net/fig/29680" pageId="22" pageNumber="23">Fig. 37</figureCitation>
) from ZMUC with a red label
<normalizedToken originalValue="“Type”">&quot;Type&quot;</normalizedToken>
and
<normalizedToken originalValue="Wiedemann’s">Wiedemann's</normalizedToken>
hand-written label. We herein designate this specimen as lectotype for taxonomic purpose of fixing the name to unique name-bearing type.
</paragraph>
<paragraph pageId="22" pageNumber="23">
<emphasis bold="true" pageId="22" pageNumber="23">
<taxonomicName authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="picea">
<emphasis italics="true" pageId="22" pageNumber="23">Anchista picea</emphasis>
</taxonomicName>
Chaudoir
</emphasis>
: Five specimens were mentioned in the original description. In the collection of MNHN, these specimens used to be in
<normalizedToken originalValue="Chaudoir’s">Chaudoir's</normalizedToken>
box, but were moved to
<taxonomicName authorityName="Bonelli" authorityYear="1810" lsidName="" pageId="22" pageNumber="23" rank="tribe" tribe="Lebiini">Lebiini</taxonomicName>
sorted boxes by J. Mateu. Obviously, Mateu selected the first one, and put
<normalizedToken originalValue="Chaudoir’s">Chaudoir's</normalizedToken>
box label and a red
<normalizedToken originalValue="“type”">&quot;type&quot;</normalizedToken>
label under it. We herein designate this specimen (
<figureCitation captionStart="Figures 37–42" captionStartId="F7" captionText="Figures 37 - 42. Type materials, scale bars = 2.0 mm: 37 Lectotype of Lebia brunnea Wiedemann [= Anchista brunnea (Wiedemann)] 38 Lectotype of Anchista picea Chaudoir [= Anchista brunnea (Wiedemann)] 39 Holotype of Plochionus fenestratus Schmidt-Goebel [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 40 Holotype of Anchista glabra Chaudoir [= Anchista fenestrata fenestrata (Schmidt-Goebel)] 41 Lectotype of Anchista subpubescens Chaudoir [= Anchista fenestrata subpubescens Chaudoir] 42 Lectotype of Anchista nubila Andrewes." httpUri="https://binary.pensoft.net/fig/29680" pageId="22" pageNumber="23">Fig. 38</figureCitation>
) with such labels as lectotype for this species for taxonomic purpose of fixing the name to unique name-bearing type.
</paragraph>
<paragraph pageId="22" pageNumber="23">
<emphasis bold="true" pageId="22" pageNumber="23">Non-type material examined</emphasis>
(Total 7 specimens from India). 1 specimen (MNHN), &quot;Chota Nagpore, Nowatoli, R. P. Cardon, XI-XII 1896&quot;. 1 specimen (MNHN), &quot;Chota Nagpore, Nowatoli, R. P. Cardon, VIII-IX 1896&quot;. 2 specimens (NHML), &quot;India Nevinson Coll. 1918-14.&quot;. 1 specimen (NHML), &quot;Fyzabad, Unt. Prov., India. R. W. G. Hingston.&quot;; &quot;Anchista brunnea Wied (= picea Chaud.), H. E. Andrewes det.&quot;. 1 specimen (NHML), &quot;Nagpur., C. P. India, 1000ft.&quot;; &quot;Anchista brunnea Wied, = picea Chaud., Compared with 2 types H. E. A.&quot;. 1 female (NMPC), &quot;Dabhalwala, Dehra Dun, India&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="23" type="diagnosis">
<paragraph pageId="22" pageNumber="23">Diagnosis.</paragraph>
<paragraph pageId="22" pageNumber="23">Head and pronotum glabrous; pronotum widest at middle, lateral margins slightly angulate in middle; elytra uniform brownish; 3rd interval of elytra with two pores, 5th interval with one pore near base.</paragraph>
<paragraph pageId="22" pageNumber="23">
Based on the glabrous surface and number of setigerous pores on 3rd and 5th intervals, this species can be readily distinguished from all the other species of
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="22" pageNumber="23">Anchista</emphasis>
</taxonomicName>
. Other species with the same pore number all belong to different genera (
<taxonomicName authorityName="Shi &amp; Zhou &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Paraphaea" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="22" pageNumber="23">Paraphaea</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Shi &amp; Liang" authorityYear="2013" class="Insecta" family="Carabidae" genus="Metallanchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="22" pageNumber="23">Metallanchista</emphasis>
</taxonomicName>
gen. n.). They are different in elytral pattern and pronotum shape and have other generic differences.
</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph pageId="22" pageNumber="23">Description.</paragraph>
<paragraph lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
Body length 7.2-7.8 mm; head and pronotum reddish brown; antennae and mouthparts yellowish brown or reddish brown, 1st antennomere and apices of terminal maxillary and labial palpomeres slightly paler; pronotum with lateral explanate area yellowish brown; elytra dark brown, without distinct pattern, but usually with basal two-thirds of disc slightly paler; ventral side same color as dorsal side; femora yellow, tarsi and tibiae yellowish brown.
<emphasis bold="true" pageId="22" pageNumber="23">Head</emphasis>
glabrous, without punc
<pageBreakToken pageId="23" pageNumber="24" start="start">tures</pageBreakToken>
or microsculpture.
<emphasis bold="true" pageId="23" pageNumber="24">Pronotum</emphasis>
glabrous,cordiform, widest slightly before middle; ratio PW/PL 1.38 to 1.41; pronotal base briefly but distinctly lobed; disc moderately convex, microsculpture indistinct, without punctures; front angles widened and rounded, not projected, with a few short setae; lateral margins slightly angulate in middle, strongly sinuate before hind angles; lateral explanate areas wide and even, with a few coarse punctures; hind angles sharp, rectangular or slightly acute, with a few very short accessory setae; basal foveae moderately deep, with a few punctures; median line distinct, strongly depressed and punctate anteriorly and subbasally.
<emphasis bold="true" pageId="23" pageNumber="24">Elytra</emphasis>
with striae shallow, with fine punctures; intervals hardly convex, without accessory setae, sparsely and finely punctate; microsculpture distinct, isodiametric; 3rd interval with two setigerous pores, basal one placed at basal one-third approximately, adjacent to 3rd interval, apical one placed at apical one-eighth approximately, adjacent to 2nd stria; 5th interval with one setigerous pore near base, adjacent to 5th stria; basal margination nearly complete. Males with adhesive hairs well developed (two whole rows) on 1st to 3rd mesotarsomeres.
<emphasis bold="true" pageId="23" pageNumber="24">Male genitalia</emphasis>
with median lobe of aedeagus straight and flat, slightly expanded to apex; left-lateral margin curved in middle; apical orifice opened dorsally, basal margin of apical orifice setose; apical lamella placed on right side, flat, short and rounded in dorsal view, slightly curved upward dorsally in lateral view; internal sac with two elongate and sinuate weakly sclerotized sclerites near apical orifice, basal part of internal sac finely scaled (
<figureCitation captionStart="Figure 70" captionStartId="F15" captionText="Figure 70. Median lobe of male genitalia, right-lateral, ventral, left-lateral, dorsal views of Anchista brunnea (Wiedemann), a paralectotype of picea Chaudoir." httpUri="https://binary.pensoft.net/fig/29688" pageId="23" pageNumber="24">Fig. 70</figureCitation>
). Female genitalia not studied.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="24" type="distribution">
<paragraph pageId="23" pageNumber="24">Distribution</paragraph>
<paragraph pageId="23" pageNumber="24">
(
<figureCitation captionStart="Map 3" captionStartId="F47" captionText="Map 3. Known distribution of genus Anchista: ◆ Anchista brunnea (Wiedemann) ★ Anchista nubila Andrewes ● Anchista pilosa sp. n. △ Anchista fenestrata fenestrata (Schmidt-Goebel) ▼ Anchista fenestrata subpubescens Chaudoir." httpUri="https://binary.pensoft.net/fig/29725" pageId="23" pageNumber="24">Map 3</figureCitation>
). India, Bengal, Sri Lanka.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="24" type="notes on synonym">
<paragraph pageId="23" pageNumber="24">Notes on synonym.</paragraph>
<paragraph pageId="23" pageNumber="24">
<bibRefCitation author="Andrewes, HE" journalOrPublisher="Spolia Zeylanica Colombo" pageId="84" pageNumber="85" pagination="129 - 141" refId="B4" refString="Andrewes, HE, 1924a. Description of some new Carabidae from Ceylon. Spolia Zeylanica Colombo 13: 129 - 141" title="Description of some new Carabidae from Ceylon." volume="13" year="1924 a">Andrewes (1924a</bibRefCitation>
,
<bibRefCitation author="Andrewes, HE" journalOrPublisher="London, The Trustees of the British Museum" pageId="84" pageNumber="85" publicationUrl="10.1080/00222932708633575" refId="B8" refString="Andrewes, HE, 1927b. Papers on Oriental Carabidae. XIX. The Annals and Magazine of Natural History (9) 19: 97-111. ., 10.1080/00222932708633575" title="Papers on Oriental Carabidae. XIX. The Annals and Magazine of Natural History (9) 19: 97 - 111." url="10.1080/00222932708633575" year="1927 b">1927b</bibRefCitation>
) indicated that
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="modesta">
<emphasis italics="true" pageId="23" pageNumber="24">Anchista modesta</emphasis>
</taxonomicName>
Nietner is a synonym of
<taxonomicName class="Hexapoda" family="Carabidae" genus="Lebia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="brunnea">
<emphasis italics="true" pageId="23" pageNumber="24">Lebia brunnea</emphasis>
</taxonomicName>
Wiedemann. We
<normalizedToken originalValue="didn’t">didn't</normalizedToken>
examine the type of
<taxonomicName authorityName="Nietner" authorityYear="1856" class="Insecta" family="Carabidae" genus="Anchista" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="modesta">
<emphasis italics="true" pageId="23" pageNumber="24">Anchista modesta</emphasis>
</taxonomicName>
, which should be in the Warsaw Museum, nor any material from Sri Lanka. We follow
<normalizedToken originalValue="Andrewes’">Andrewes'</normalizedToken>
treatment in the present paper.
</paragraph>
</subSubSection>
</treatment>
</document>
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