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<document id="4D790059486B94BFFDF2E1602DD19A18" ID-CLB-Dataset="29652" ID-DOI="http://dx.doi.org/10.3897/zookeys.777.23212" ID-GBIF-Dataset="516d998c-117e-4cbb-9e21-e38843778702" ID-PMC="PMC6079114" ID-Pensoft-Pub="1313-2970-777-57" ID-PubMed="30100790" ID-ZooBank="D9F353643DCD4BA6B70D62FB275DEB1B" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-777-57" ModsDocOrigin="ZooKeys 777" ModsDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" checkinTime="1533035384019" checkinUser="pensoft" docAuthor="Schilthuizen, Menno, Perreau, Michel &amp; Njunjic, Iva" docDate="2018" docId="950D738AC4C264EB628B67886CC25358" docLanguage="en" docName="ZooKeys 777: 57-108" docOrigin="ZooKeys 777" docSource="http://dx.doi.org/10.3897/zookeys.777.23212" docTitle="Ptomaphaginus caroli Schilthuizen &amp; Perreau 2008" docType="treatment" docVersion="6" lastPageNumber="68" masterDocId="F179FFE1BF6AFFEEEF68FFF0FFB5FFBD" masterDocTitle="A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)" masterLastPageNumber="108" masterPageNumber="57" pageNumber="68" updateTime="1701383097238" updateUser="ExternalLinkService">
<mods:mods id="2879EE506E632DDA7320F64BB52F4019" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E484B701EF139FC88AE9CC683743266D">A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae)</mods:title>
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<mods:namePart id="E2FCBFF52392B6F739E1BB3AB24E7A07">Schilthuizen, Menno</mods:namePart>
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<mods:namePart id="DC57F0E16F96F7374A773921CBAE2DF9">Perreau, Michel</mods:namePart>
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<mods:namePart id="9F9098CAEB73C8F75781C4A67D903A16">Njunjic, Iva</mods:namePart>
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<mods:date id="15DDB3367FE2ACFDD542CEEA1F4A553B">2018</mods:date>
<mods:detail id="7084778C8247029ECC52D0992B66D8F0" type="volume">
<mods:number id="1B7C0A7A6CE50BE98C75D9F5BB125122">777</mods:number>
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<mods:url id="AA67D3067B2C8AD533325B005D728B05">http://dx.doi.org/10.3897/zookeys.777.23212</mods:url>
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<mods:identifier id="3657ADA783EAE749BEFC3E2C6C3481CB" type="DOI">http://dx.doi.org/10.3897/zookeys.777.23212</mods:identifier>
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<treatment id="950D738AC4C264EB628B67886CC25358" ID-GBIF-Taxon="146746174" LSID="urn:lsid:plazi:treatment:950D738AC4C264EB628B67886CC25358" httpUri="http://treatment.plazi.org/id/950D738AC4C264EB628B67886CC25358" lastPageNumber="68" pageId="11" pageNumber="68">
<subSubSection id="71AFA720AE5ED5275D3B6D287DC8B798" pageId="11" pageNumber="68" type="nomenclature">
<paragraph id="C3E3A4BC3EF4C7D34DF523D4674A4846" pageId="11" pageNumber="68">
<taxonomicName id="0BE39D1B39AC7E4F8D5FCF5356666FF0" ID-CoL="4Q98B" authority="Schilthuizen &amp; Perreau, 2008" authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaginus caroli" order="Coleoptera" pageId="11" pageNumber="68" phylum="Arthropoda" rank="species" species="caroli">Ptomaphaginus caroli Schilthuizen &amp; Perreau, 2008</taxonomicName>
Figures 4c, 8a, b
</paragraph>
</subSubSection>
<subSubSection id="5A3DC5800DFD034394194BFC05D04C71" pageId="11" pageNumber="68" type="reference_group">
<paragraph id="EF89DDC7FC6BEF6F9FD6861AE8635257" pageId="11" pageNumber="68">
<taxonomicName id="9978A9D86DE22D1114F5B8094F11CD3D" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaginus caroli" order="Coleoptera" pageId="11" pageNumber="68" phylum="Arthropoda" rank="species" species="caroli">Ptomaphaginus caroli</taxonomicName>
Schilthuizen &amp; Perreau, 2008: 193, figs 14-15, 26; type from Gunung Mas, Sabah, Borneo (in RMNH).
</paragraph>
</subSubSection>
<subSubSection id="3B3508EF1CE60302DD9CFF2036CE0C2C" pageId="11" pageNumber="68" type="description">
<paragraph id="D6C35088BEE4C0EF29549FE07E7247D2" pageId="11" pageNumber="68">Description.</paragraph>
<paragraph id="70C347A42C6291320C8BD5FC085D618F" pageId="11" pageNumber="68">
(Adapted from
<bibRefCitation id="EE0B0532233B784670EDED8FFADC4AAC" author="Schilthuizen, M" journalOrPublisher="Zoologische Mededelingen Leiden" pageId="37" pageNumber="94" pagination="189 - 210" title="New species and new records of Ptomaphaginus Portevin from the Sunda region (Coleoptera: Leiodidae: Cholevinae)." volume="82" year="2008">Schilthuizen and Perreau (2008)</bibRefCitation>
). Length 2.7 mm. Habitus relatively slender and narrow, flat. Pronotum 1.62 times as long as wide, slightly narrower than the elytra. Elytra 1.41 times as long as their combined width (length measured from the caudal tip of the scutellum to the apex of the elytra). Winged. Long setae on the ventral side of the male profemur and protibia absent. Aedeagus apically with two short
<normalizedToken id="F5D2C1A62FBDC6C7708BF063EDA7F055" originalValue="‘wings’">'wings'</normalizedToken>
and a very small, indistinct terminal processus. Spiculum gastrale long-triangular, the apex nearly truncate, with a small central projection; similar in shape to
<taxonomicName id="E46B35A3F1D717BF31F4D658159EACDC" lsidName="P. latimanus" pageId="11" pageNumber="68" rank="species" species="latimanus">P. latimanus</taxonomicName>
and
<taxonomicName id="B3E52F8BE45E1C66C6CA81541B7BAECB" lsidName="P. similipes" pageId="11" pageNumber="68" rank="species" species="similipes">P. similipes</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="101F294113078EBDCEF4DA91471C69B8" pageId="11" pageNumber="68" type="differential diagnosis">
<paragraph id="96E730DD4250324A244A4467F2AD4CF1" pageId="11" pageNumber="68">Differential diagnosis.</paragraph>
<paragraph id="A1BD7FFD368A89E5D589E0EB3BF7A2BB" pageId="11" pageNumber="68">
<taxonomicName id="E536335F139D6AE5E45A1D21BA378FFC" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphaginus caroli" order="Coleoptera" pageId="11" pageNumber="68" phylum="Arthropoda" rank="species" species="caroli">Ptomaphaginus caroli</taxonomicName>
has a similar aedeagus as
<taxonomicName id="A694DD3E6A7835C38748BFEF454A886F" lsidName="P. bryanti" pageId="11" pageNumber="68" rank="species" species="bryanti">P. bryanti</taxonomicName>
,
<taxonomicName id="F9219BB660010D63928D4E6EF078690F" lsidName="P. similipes" pageId="11" pageNumber="68" rank="species" species="similipes">P. similipes</taxonomicName>
, and
<taxonomicName id="7C691089618A80392B1D1261B4313109" lsidName="P. bryantioides" pageId="11" pageNumber="68" rank="species" species="bryantioides">P. bryantioides</taxonomicName>
. However, it differs in having a distinctly elongated habitus (elytral index of 1.41) and very short apical
<normalizedToken id="1DE00287A1BBA5D248BBE26F14BD5F3A" originalValue="‘wings’">'wings'</normalizedToken>
on the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="CDD14571B85C6E4AA63901D68C9DE528" pageId="11" pageNumber="68" type="habitat">
<paragraph id="8F431664C8768718FE57C05E5716AEDE" pageId="11" pageNumber="68">Habitat and distribution.</paragraph>
<paragraph id="756FC5F70FB75EED04F5531146949299" pageId="11" pageNumber="68">
So far, only known from the type specimen, collected in lower montane forest at 1350 m in the Crocker Range of Sabah. The aedeagal shape shows that it belongs within the &quot;
<taxonomicName id="20CB2DCCA2157736211DA43652C971A5" class="Magnoliopsida" family="Malvaceae" genus="Staphylinoidea" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Staphylinoidea bryanti" order="Malvales" pageId="11" pageNumber="68" phylum="Tracheophyta" rank="species" species="bryanti">bryanti</taxonomicName>
-group&quot;.
</paragraph>
</subSubSection>
<subSubSection id="AB4933E07CE80C9251DB39117C17F983" pageId="11" pageNumber="68" type="remarks">
<paragraph id="071201F7153EB829C529E1820698745F" pageId="11" pageNumber="68">Remarks.</paragraph>
<paragraph id="D3C025F7953F956256BBED52488A948D" pageId="11" pageNumber="68">
The aedeagus of the holotype has been lost shortly after it was first collected and studied (in 2000). Before the loss, sketches were made of the dorsal and lateral view of the aedeagus, which form the basis for the line drawing in Figure 8 and in
<bibRefCitation id="A6BC21F4F1359FC9C2231B63F96E2E8D" author="Schilthuizen, M" journalOrPublisher="Zoologische Mededelingen Leiden" pageId="37" pageNumber="94" pagination="189 - 210" title="New species and new records of Ptomaphaginus Portevin from the Sunda region (Coleoptera: Leiodidae: Cholevinae)." volume="82" year="2008">Schilthuizen and Perreau (2008)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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