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<document ID-DOI="10.5281/zenodo.174866" ID-GBIF-Dataset="15363d41-dba5-4484-addb-9734f619bcf1" ID-ISSN="1175-5326" ID-Zenodo-Dep="174866" checkinTime="1459790442371" checkinUser="plazi" docAuthor="Short, Andrew Edward Z. &amp; Torres, Phillip J." docDate="2006" docId="8D7A87A6FFE3FFA81F50BA55AA6381D4" docLanguage="en" docName="zt01369p017.pdf" docOrigin="Zootaxa 1369" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@632cf7d3" docStyleName="zootaxa.0000.journal_article" docTitle="Hemiosus maculatus Sharp 1882" docType="treatment" docVersion="5" lastPageId="5" lastPageNumber="6" masterDocId="7143FFDEFFE0FFAD1E58BC33AB088002" masterDocTitle="Review of the Hemiosus Sharp and Derallus Sharp of North America, Central America, and the Caribbean (Coleoptera: Hydrophilidae: Berosini)" masterLastPageNumber="17" masterPageNumber="1" pageId="3" pageNumber="4" updateTime="1569536254438" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the Hemiosus Sharp and Derallus Sharp of North America, Central America, and the Caribbean (Coleoptera: Hydrophilidae: Berosini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Short, Andrew Edward Z.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Torres, Phillip J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2006</mods:date>
<mods:detail type="volume">
<mods:number>1369</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>17</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.174866</mods:identifier>
<mods:identifier type="GBIF-Dataset">15363d41-dba5-4484-addb-9734f619bcf1</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">174866</mods:identifier>
<mods:identifier type="ZooBank">7675CC72-B6D3-4EF6-B2EB-FE37BD42CEC1</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:8D7A87A6FFE3FFA81F50BA55AA6381D4" httpUri="http://treatment.plazi.org/id/8D7A87A6FFE3FFA81F50BA55AA6381D4" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
<subSubSection pageId="3" pageNumber="4" type="nomenclature">
<paragraph blockId="3.[264,674,1638,1699]" box="[264,674,1638,1664]" pageId="3" pageNumber="4">
<heading bold="true" box="[264,674,1638,1664]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<taxonomicName authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[264,674,1638,1664]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis bold="true" box="[264,674,1638,1664]" pageId="3" pageNumber="4">
<emphasis bold="true" box="[264,517,1638,1664]" italics="true" pageId="3" pageNumber="4">Hemiosus maculatus</emphasis>
<bibRefCitation author="Sharp" box="[524,674,1638,1664]" pageId="3" pageNumber="17" refString="Sharp, D. (1882). Fam. Hydrophylidae. In: Biologia Centrali-Americana. Insecta, Coleoptera, Vol. 1, Part 2. Taylor &amp; Francis, London, pp. 53 - 116. [Available from URL: http: // www. sil. si. edu / digitalcollections / bca /]" type="book chapter" year="1882">
Sharp,
<number box="[616,674,1638,1664]" pageId="3" pageNumber="4" value="1882.0">1882</number>
</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="3.[264,674,1638,1699]" box="[264,392,1673,1699]" pageId="3" pageNumber="4">
(
<figureCitation box="[273,383,1673,1699]" captionStart="FIGURES 1 – 2" captionStartId="5.[264,382,1180,1204]" captionTargetBox="[431,1172,528,1139]" captionTargetId="figure@5.[396,1192,497,1154]" captionTargetPageId="5" captionText="FIGURES 1 – 2. Dorsal and lateral habitus of Hemiosus maculatus Sharp (specimen from Costa Rica). Scale bar 2.1 mm." httpUri="https://zenodo.org/record/174867/files/figure.png" pageId="3" pageNumber="4">
Figs.
<number box="[339,383,1673,1699]" pageId="3" pageNumber="4" value="1.5" valueMax="2.0" valueMin="1.0">1–2</number>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[264,675,1753,1777]" pageId="3" pageNumber="4" type="reference_group">
<paragraph blockId="3.[264,675,1753,1777]" box="[264,675,1753,1777]" pageId="3" pageNumber="4">
<treatmentCitationGroup box="[264,675,1753,1777]" pageId="3" pageNumber="4">
<taxonomicName authority="Sharp, 1882: 85" authorityName="Sharp" authorityPageNumber="85" authorityYear="1882" box="[264,672,1753,1777]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis box="[264,486,1754,1777]" italics="true" pageId="3" pageNumber="4">Hemiosus maculatus</emphasis>
<treatmentCitation author="Sharp" box="[492,672,1753,1776]" page="85" pageId="3" pageNumber="4" year="1882">
<bibRefCitation author="Sharp" box="[492,625,1753,1776]" pageId="3" pageNumber="17" refString="Sharp, D. (1882). Fam. Hydrophylidae. In: Biologia Centrali-Americana. Insecta, Coleoptera, Vol. 1, Part 2. Taylor &amp; Francis, London, pp. 53 - 116. [Available from URL: http: // www. sil. si. edu / digitalcollections / bca /]" type="book chapter" year="1882">
Sharp,
<number box="[568,625,1753,1776]" pageId="3" pageNumber="4" value="1882.0">1882</number>
</bibRefCitation>
:
<number box="[642,672,1753,1776]" pageId="3" pageNumber="4" value="85.0">85</number>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="materials_examined">
<paragraph blockId="3.[264,1322,1825,1891]" pageId="3" pageNumber="4">
<emphasis bold="true" box="[264,577,1825,1851]" pageId="3" pageNumber="4">
<typeStatus box="[264,325,1825,1851]" pageId="3" pageNumber="4">Type</typeStatus>
material examined.
</emphasis>
<typeStatus box="[595,702,1825,1851]" pageId="3" pageNumber="4">Syntypes</typeStatus>
(
<number box="[720,735,1825,1851]" pageId="3" pageNumber="4" value="8.0">8</number>
examined): “
<taxonomicName box="[904,1024,1825,1851]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">Hemiosus</taxonomicName>
macula-/ tus.
<typeStatus box="[1195,1253,1825,1851]" pageId="3" pageNumber="4">Type</typeStatus>
D.S./
<collectingCountry box="[264,391,1865,1891]" name="Guatemala" pageId="3" pageNumber="4">Guatemala</collectingCountry>
Champion”, “
<typeStatus box="[561,622,1865,1891]" pageId="3" pageNumber="4">Type</typeStatus>
[red disc]”, “
<collectingCountry box="[784,913,1865,1891]" name="Guatemala" pageId="3" pageNumber="4">Guatemala</collectingCountry>
/ Champion.”, “Sharp Coll./
<number box="[1256,1318,1865,1891]" pageId="3" pageNumber="4" value="1905.0">1905</number>
.
</paragraph>
<paragraph blockId="4.[264,1323,284,1906]" pageId="4" pageNumber="5">
–
<number box="[281,328,284,310]" pageId="4" pageNumber="5" value="313.0">313</number>
”, “B.C.A. Col.
<date box="[510,554,284,310]" pageId="4" pageNumber="5">
I.
<number box="[534,550,284,310]" pageId="4" pageNumber="5" value="2.0">2</number>
.
</date>
/
<taxonomicName box="[572,688,284,310]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Hemiosus</taxonomicName>
/
<taxonomicName authority="Sharp." authorityName="Sharp." box="[705,916,284,310]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">maculatus, Sharp.</taxonomicName>
” (
<collectionCode box="[942,1037,284,310]" pageId="4" pageNumber="5">BMNH</collectionCode>
); [
<specimenCount box="[1065,1211,284,310]" pageId="4" pageNumber="5" type="generic">
<number box="[1065,1082,284,310]" pageId="4" pageNumber="5" value="2.0">2</number>
specimens
</specimenCount>
on card], “Pantaleon,/
<quantity box="[421,513,324,350]" metricMagnitude="2" metricUnit="m" metricValue="5.1816" pageId="4" pageNumber="5" unit="ft" value="1700.0">1700 ft</quantity>
./ Champion.”, “B.C.A. Col.
<date box="[874,925,324,350]" pageId="4" pageNumber="5">
I.
<number box="[905,921,324,350]" pageId="4" pageNumber="5" value="2.0">2</number>
.
</date>
/
<taxonomicName box="[948,1064,324,350]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Hemiosus</taxonomicName>
/
<taxonomicName authority="Sharp." authorityName="Sharp." box="[1087,1307,324,350]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">maculatus, Sharp.</taxonomicName>
”, “
<taxonomicName box="[274,527,364,390]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">Hemiosus maculatus</taxonomicName>
”, “SYN-TYPE [light blue disc]” (
<collectionCode box="[932,1026,364,390]" pageId="4" pageNumber="5">BMNH</collectionCode>
); “
<taxonomicName box="[1058,1177,364,390]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Hemiosus</taxonomicName>
/
<taxonomicName box="[1195,1313,364,390]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">maculatus</taxonomicName>
/ D.S./ Pantaleon
<quantity box="[464,555,403,429]" metricMagnitude="2" metricUnit="m" metricValue="5.1816" pageId="4" pageNumber="5" unit="ft" value="1700.0">1700 ft</quantity>
./ Champion [same card as
<specimenCount box="[895,1051,403,429]" pageId="4" pageNumber="5" type="generic">
<number box="[895,909,403,429]" pageId="4" pageNumber="5" value="2.0">2</number>
specimens
</specimenCount>
]”, “Sharp Coll./
<number box="[1257,1319,403,429]" pageId="4" pageNumber="5" value="1905.0">1905</number>
. –
<number box="[281,328,443,469]" pageId="4" pageNumber="5" value="313.0">313</number>
”, “Pantaleon,/
<quantity box="[517,610,443,469]" metricMagnitude="2" metricUnit="m" metricValue="5.1816" pageId="4" pageNumber="5" unit="ft" value="1700.0">1700 ft</quantity>
./ Champion.”, “B.C.A. Col.
<date box="[979,1030,443,469]" pageId="4" pageNumber="5">
I.
<number box="[1011,1027,443,469]" pageId="4" pageNumber="5" value="2.0">2</number>
.
</date>
/
<taxonomicName box="[1056,1172,443,469]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Hemiosus</taxonomicName>
/
<taxonomicName authority="Sharp." authorityName="Sharp." class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">maculatus, Sharp.</taxonomicName>
”, “SYN-TYPE [light blue disc]” (
<collectionCode box="[745,840,483,509]" pageId="4" pageNumber="5">BMNH</collectionCode>
); “[male symbol]”, “Pantaleon,/
<quantity box="[1223,1310,483,509]" metricMagnitude="2" metricUnit="m" metricValue="5.1816" pageId="4" pageNumber="5" unit="ft" value="1700.0">1700 ft</quantity>
./ Champion.”, “B.C.A. Col.
<date box="[583,628,523,549]" pageId="4" pageNumber="5">
I.
<number box="[608,624,523,549]" pageId="4" pageNumber="5" value="2.0">2</number>
.
</date>
/
<taxonomicName box="[646,763,523,549]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Hemiosus</taxonomicName>
/
<taxonomicName authority="Sharp." authorityName="Sharp." box="[780,995,523,549]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">maculatus, Sharp.</taxonomicName>
”, “Para-type [orange card]” (
<collectionCode box="[272,365,563,589]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="4" pageNumber="5">IRSNB</collectionCode>
); “Pantaleon,/
<quantity box="[545,635,563,589]" metricMagnitude="2" metricUnit="m" metricValue="5.1816" pageId="4" pageNumber="5" unit="ft" value="1700.0">1700 ft</quantity>
./ Champion.”, “B.C.A. Col.
<date box="[989,1037,563,589]" pageId="4" pageNumber="5">
I.
<number box="[1018,1034,563,589]" pageId="4" pageNumber="5" value="2.0">2</number>
.
</date>
/
<taxonomicName box="[1059,1175,563,589]" class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Hemiosus</taxonomicName>
/
<taxonomicName authority="Sharp." authorityName="Sharp." class="Insecta" family="Hydrophilidae" genus="Hemiosus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">maculatus, Sharp.</taxonomicName>
”, “Para-type [orange card]” (
<collectionCode box="[685,778,603,629]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="4" pageNumber="5">IRSNB</collectionCode>
).
</paragraph>
<paragraph blockId="4.[264,1323,284,1906]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[312,901,643,669]" pageId="4" pageNumber="5">
Additional material examined (
<number box="[704,752,643,669]" pageId="4" pageNumber="5" value="262.0">262</number>
).
<collectingCountry box="[782,895,643,669]" name="Belize" pageId="4" pageNumber="5">BELIZE</collectingCountry>
:
</emphasis>
<collectingCountry box="[917,992,643,669]" name="Belize" pageId="4" pageNumber="5">Belize</collectingCountry>
District: Western Highway near Zoo, light trap in pine grassland,
<date box="[706,804,683,709]" pageId="4" pageNumber="5" value="2003-01-07">
<number box="[706,721,683,709]" pageId="4" pageNumber="5" value="7.0">7</number>
.i.
<number box="[737,804,683,709]" pageId="4" pageNumber="5" value="2003.0">2003</number>
</date>
, C.R. Bartlett leg. (
<number box="[1032,1049,683,709]" pageId="4" pageNumber="5" value="3.0">3</number>
:
<collectionCode box="[1060,1138,683,709]" pageId="4" pageNumber="5">AEZS</collectionCode>
); Cayo District, near Teakettle Bank, Pook’s Hill, light trap at river, (
<number box="[906,952,723,749]" pageId="4" pageNumber="5" value="5.5" valueMax="6.0" valueMin="5.0">5–6</number>
).
<date box="[965,1045,723,749]" pageId="4" pageNumber="5" value="2003-01">
i.
<number box="[978,1045,723,749]" pageId="4" pageNumber="5" value="2003.0">2003</number>
</date>
, C.R. Bartlett leg. (
<number box="[1285,1318,723,749]" pageId="4" pageNumber="5" value="12.0">12</number>
:
<collectionCode box="[264,342,763,789]" pageId="4" pageNumber="5">AEZS</collectionCode>
).
<emphasis bold="true" box="[367,705,762,788]" pageId="4" pageNumber="5">
<collectingCountry box="[367,558,762,788]" name="Costa Rica" pageId="4" pageNumber="5">COSTA RICA</collectingCountry>
: Alajuela:
</emphasis>
near San Ramon, roadside pools near Quebrada Negra,
<date box="[350,486,802,828]" pageId="4" pageNumber="5" value="1999-07-13">
<number box="[350,379,802,828]" pageId="4" pageNumber="5" value="13.0">13</number>
.vii.
<number box="[421,486,802,828]" pageId="4" pageNumber="5" value="1999.0">1999</number>
</date>
, R.E. Roughly leg. (
<number box="[728,742,802,828]" pageId="4" pageNumber="5" value="1.0">1</number>
: UMC);
<emphasis bold="true" box="[846,1002,802,828]" pageId="4" pageNumber="5">Guanacaste:</emphasis>
Rio Guacimal at Highway
<quantity box="[264,370,842,868]" metricMagnitude="2" metricUnit="m" metricValue="4.2672" pageId="4" pageNumber="5" unit="ft" value="1400.0">1, 400 ft</quantity>
.,
<date box="[387,515,842,868]" pageId="4" pageNumber="5" value="2003-06-17">
<number box="[387,416,842,868]" pageId="4" pageNumber="5" value="17.0">17</number>
.vi.
<number box="[449,515,842,868]" pageId="4" pageNumber="5" value="2003.0">2003</number>
</date>
, A.E.Z. Short leg. (
<number box="[748,782,842,868]" pageId="4" pageNumber="5" value="37.0">37</number>
:
<collectionCode box="[794,870,842,868]" pageId="4" pageNumber="5">AEZS</collectionCode>
,
<collectionCode box="[883,961,842,868]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="4" pageNumber="5">INBio</collectionCode>
); Rio Carmona, near Carmona at Finca Aqua Fria, HG-vapor light,
<quantity box="[712,779,882,908]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="5" unit="m" value="30.0">30 m</quantity>
,
<date box="[793,907,882,908]" pageId="4" pageNumber="5" value="2003-01-15">
<number box="[793,822,882,908]" pageId="4" pageNumber="5" value="15.0">15</number>
.i.
<number box="[839,907,882,908]" pageId="4" pageNumber="5" value="2003.0">2003</number>
</date>
, A.E.Z. Short, R. Roughley &amp; W. Porras leg. (
<number box="[406,439,922,948]" pageId="4" pageNumber="5" value="17.0">17</number>
:
<collectionCode box="[451,527,922,948]" pageId="4" pageNumber="5">AEZS</collectionCode>
,
<collectionCode box="[538,616,922,948]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="4" pageNumber="5">INBio</collectionCode>
); same locality but
<date box="[843,969,922,948]" pageId="4" pageNumber="5" value="2002-11-26">
<number box="[843,874,922,948]" pageId="4" pageNumber="5" value="26.0">26</number>
.xi.
<number box="[905,969,922,948]" pageId="4" pageNumber="5" value="2002.0">2002</number>
</date>
(
<number box="[986,1004,922,948]" pageId="4" pageNumber="5" value="2.0">2</number>
:
<collectionCode box="[1016,1094,922,948]" pageId="4" pageNumber="5">AEZS</collectionCode>
); Nandayure, Finca Aqua Fria,
<quantity box="[409,522,962,988]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="4" pageNumber="5" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100 m</quantity>
,
<date box="[540,654,962,988]" pageId="4" pageNumber="5" value="2002-01-19">
<number box="[540,570,962,988]" pageId="4" pageNumber="5" value="19.0">19</number>
.i.
<number box="[586,654,962,988]" pageId="4" pageNumber="5" value="2002.0">2002</number>
</date>
, W. Porras leg. (
<number box="[876,890,962,988]" pageId="4" pageNumber="5" value="1.0">1</number>
:
<collectionCode box="[909,987,962,988]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="4" pageNumber="5">INBio</collectionCode>
);
<emphasis bold="true" box="[1014,1167,962,988]" pageId="4" pageNumber="5">Puntarenas:</emphasis>
Corcovado National Park, Sirena Station,
<quantity box="[628,678,1002,1028]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="5" unit="m" value="5.0">5 m</quantity>
,
<date box="[692,805,1002,1028]" pageId="4" pageNumber="5" value="1992-06-01">
<number box="[692,702,1002,1028]" pageId="4" pageNumber="5" value="1.0">1</number>
.vi.
<number box="[737,805,1002,1028]" pageId="4" pageNumber="5" value="1992.0">1992</number>
</date>
, UV light, Z.H. Falin leg. (
<number box="[1139,1156,1002,1028]" pageId="4" pageNumber="5" value="3.0">3</number>
:
<collectionCode box="[1170,1245,1002,1028]" httpUri="http://grbio.org/cool/sxrc-6x8s" name="Cornell University Insect Collection" pageId="4" pageNumber="5">CUIC</collectionCode>
); N of Cirvelas, Rio Seco at Highway
<number box="[651,666,1042,1068]" pageId="4" pageNumber="5" value="1.0">1</number>
, pools along river,
<date box="[905,1041,1042,1068]" pageId="4" pageNumber="5" value="2003-07-17">
<number box="[905,934,1042,1068]" pageId="4" pageNumber="5" value="17.0">17</number>
.vii.
<number box="[976,1041,1042,1068]" pageId="4" pageNumber="5" value="2003.0">2003</number>
</date>
, A.E.Z. Short leg. (
<number box="[1284,1318,1042,1068]" pageId="4" pageNumber="5" value="70.0">70</number>
:
<collectionCode box="[264,340,1082,1108]" pageId="4" pageNumber="5">AEZS</collectionCode>
,
<collectionCode box="[354,432,1082,1108]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="4" pageNumber="5">INBio</collectionCode>
); Golfito, Sector El Tajo,
<quantity box="[744,882,1082,1108]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="5" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100–200 m</quantity>
,
<date box="[896,1022,1082,1108]" pageId="4" pageNumber="5" value="2004-04-28">
<number box="[896,926,1082,1108]" pageId="4" pageNumber="5" value="28.0">28</number>
.iv.
<number box="[958,1022,1082,1108]" pageId="4" pageNumber="5" value="2004.0">2004</number>
</date>
, light trap, W. Porras, B. Gamboa, D. Briceno &amp; M. Morgaga leg. (
<number box="[770,788,1122,1148]" pageId="4" pageNumber="5" value="4.0">4</number>
:
<collectionCode box="[802,880,1122,1148]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="4" pageNumber="5">INBio</collectionCode>
); Osa Peninsula, Rio Conte,
<quantity box="[1225,1323,1122,1148]" metricMagnitude="4" metricUnit="m" metricValue="1.83" pageId="4" pageNumber="5" unit="km" value="18.3">18.3 km</quantity>
NW of Puerto Jimenez,
<date box="[552,679,1162,1188]" pageId="4" pageNumber="5" value="2003-06-18">
<number box="[552,581,1162,1188]" pageId="4" pageNumber="5" value="18.0">18</number>
.vi.
<number box="[613,679,1162,1188]" pageId="4" pageNumber="5" value="2003.0">2003</number>
</date>
, A.E.Z. Short leg. (
<number box="[914,948,1162,1188]" pageId="4" pageNumber="5" value="22.0">22</number>
:
<collectionCode box="[961,1037,1162,1188]" pageId="4" pageNumber="5">AEZS</collectionCode>
,
<collectionCode box="[1050,1128,1162,1188]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="4" pageNumber="5">INBio</collectionCode>
); Rio Tigre near Puerto Jimenez, black light,
<date box="[655,783,1202,1228]" pageId="4" pageNumber="5" value="2003-06-18">
<number box="[655,684,1202,1228]" pageId="4" pageNumber="5" value="18.0">18</number>
.vi.
<number box="[717,783,1202,1228]" pageId="4" pageNumber="5" value="2003.0">2003</number>
</date>
, A.E.Z. Short leg. (
<number box="[1070,1104,1202,1228]" pageId="4" pageNumber="5" value="40.0">40</number>
:
<collectionCode box="[1129,1205,1202,1228]" pageId="4" pageNumber="5">AEZS</collectionCode>
,
<collectionCode box="[1232,1310,1202,1228]" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="4" pageNumber="5">INBio</collectionCode>
).
<emphasis bold="true" box="[264,441,1241,1267]" pageId="4" pageNumber="5">
<collectingCountry box="[264,436,1241,1267]" name="Honduras" pageId="4" pageNumber="5">HONDURAS</collectingCountry>
:
</emphasis>
Lancetilla, nr. Tela,
<date box="[734,861,1242,1268]" pageId="4" pageNumber="5" value="1979-06-19">
<number box="[734,764,1242,1268]" pageId="4" pageNumber="5" value="19.0">19</number>
.vi.
<number box="[796,861,1242,1268]" pageId="4" pageNumber="5" value="1979.0">1979</number>
</date>
, at light, J.A. Chemasak, A. &amp; M. Michelbacher, &amp; W. Middelkauff leg. (
<number box="[725,743,1281,1307]" pageId="4" pageNumber="5" value="2.0">2</number>
:
<collectionCode box="[755,831,1281,1307]" pageId="4" pageNumber="5">AEZS</collectionCode>
,
<collectionCode LSID="urn:lsid:biocol.org:col:1013" box="[844,931,1281,1307]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="4" pageNumber="5">EMEC</collectionCode>
).
<emphasis bold="true" box="[959,1203,1281,1307]" pageId="4" pageNumber="5">
<collectingCountry box="[959,1088,1281,1307]" name="Mexico" pageId="4" pageNumber="5">MEXICO</collectingCountry>
: Jalisco:
</emphasis>
Chamela Biological Station,
<date box="[510,673,1321,1347]" pageId="4" pageNumber="5" value="1987-10-21" valueMax="1987-10-22" valueMin="1987-10-21">
<number box="[510,588,1321,1347]" pageId="4" pageNumber="5" value="21.5" valueMax="22.0" valueMin="21.0">21–22</number>
.x.
<number box="[611,673,1321,1347]" pageId="4" pageNumber="5" value="1987.0">1987</number>
</date>
, black light, Chemsak &amp; Powell leg. (
<number box="[1187,1221,1321,1347]" pageId="4" pageNumber="5" value="39.0">39</number>
:
<collectionCode box="[1242,1318,1321,1347]" pageId="4" pageNumber="5">AEZS</collectionCode>
,
<collectionCode LSID="urn:lsid:biocol.org:col:1013" box="[264,350,1361,1387]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="4" pageNumber="5">EMEC</collectionCode>
);
<emphasis bold="true" box="[373,475,1361,1387]" pageId="4" pageNumber="5">Sinaloa:</emphasis>
<quantity box="[494,567,1361,1387]" metricMagnitude="4" metricUnit="m" metricValue="3.3796224" pageId="4" pageNumber="5" unit="mi" value="21.0">21 mi</quantity>
. E Villa Union,
<quantity box="[764,840,1361,1387]" metricMagnitude="1" metricUnit="m" metricValue="9.144" pageId="4" pageNumber="5" unit="ft" value="300.0">300 ft</quantity>
.,
<date box="[859,994,1361,1387]" pageId="4" pageNumber="5" value="1964-07-25">
<number box="[859,890,1361,1387]" pageId="4" pageNumber="5" value="25.0">25</number>
.vii.
<number box="[931,994,1361,1387]" pageId="4" pageNumber="5" value="1964.0">1964</number>
</date>
, lights, Chemsak &amp; Powell leg. (
<number box="[331,348,1401,1427]" pageId="4" pageNumber="5" value="3.0">3</number>
:
<collectionCode LSID="urn:lsid:biocol.org:col:1013" box="[366,453,1401,1427]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="4" pageNumber="5">EMEC</collectionCode>
);
<emphasis bold="true" box="[478,577,1400,1426]" pageId="4" pageNumber="5">Sonora:</emphasis>
<quantity box="[603,665,1401,1427]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="4" pageNumber="5" unit="mi" value="5.0">5 mi</quantity>
. W Alamos,
<date box="[835,978,1401,1427]" pageId="4" pageNumber="5" value="1959-08-14">
<number box="[835,865,1401,1427]" pageId="4" pageNumber="5" value="14.0">14</number>
.viii.
<number box="[915,978,1401,1427]" pageId="4" pageNumber="5" value="1959.0">1959</number>
</date>
, Werner &amp; Nutting leg. (
<number box="[1300,1318,1401,1427]" pageId="4" pageNumber="5" value="2.0">2</number>
:
<collectionCode box="[264,342,1441,1467]" pageId="4" pageNumber="5">AEZS</collectionCode>
); Alamos,
<date box="[478,614,1441,1467]" pageId="4" pageNumber="5" value="1963-07-15">
<number box="[478,507,1441,1467]" pageId="4" pageNumber="5" value="15.0">15</number>
.vii.
<number box="[550,614,1441,1467]" pageId="4" pageNumber="5" value="1963.0">1963</number>
</date>
, P.J. Spangler leg. (
<number box="[863,881,1441,1467]" pageId="4" pageNumber="5" value="2.0">2</number>
:
<collectionCode box="[897,975,1441,1467]" pageId="4" pageNumber="5">AEZS</collectionCode>
); Sinaloa State: Culiacan, at black light,
<date box="[405,541,1481,1507]" pageId="4" pageNumber="5" value="1963-07-16">
<number box="[405,435,1481,1507]" pageId="4" pageNumber="5" value="16.0">16</number>
.vii.
<number box="[477,541,1481,1507]" pageId="4" pageNumber="5" value="1963.0">1963</number>
</date>
, P.J. Spangler leg. (
<number box="[778,792,1481,1507]" pageId="4" pageNumber="5" value="1.0">1</number>
:
<collectionCode box="[806,884,1481,1507]" pageId="4" pageNumber="5">AEZS</collectionCode>
).
<emphasis bold="true" box="[913,1084,1480,1506]" pageId="4" pageNumber="5">NICARAGA:</emphasis>
Managua, km
<number box="[1271,1322,1481,1507]" pageId="4" pageNumber="5" value="45.8">45.8</number>
of Pan-Am Highway,
<quantity box="[555,641,1521,1547]" metricMagnitude="2" metricUnit="m" metricValue="1.31064" pageId="4" pageNumber="5" unit="ft" value="430.0">430 ft</quantity>
., stream,
<date box="[772,902,1521,1547]" pageId="4" pageNumber="5" value="2002-08-04">
<number box="[772,787,1521,1547]" pageId="4" pageNumber="5" value="4.0">4</number>
.viii.
<number box="[837,902,1521,1547]" pageId="4" pageNumber="5" value="2002.0">2002</number>
</date>
, W.D. Shepard leg. (
<number box="[1194,1208,1521,1547]" pageId="4" pageNumber="5" value="1.0">1</number>
:
<collectionCode box="[1232,1310,1521,1547]" pageId="4" pageNumber="5">AEZS</collectionCode>
). Representative material deposited in
<collectionCode box="[698,806,1561,1587]" pageId="4" pageNumber="5">MALUZ</collectionCode>
,
<collectionCode LSID="urn:lsid:biocol.org:col:33834" box="[817,895,1561,1587]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="4" pageNumber="5">MIZA</collectionCode>
,
<collectionCode box="[906,984,1561,1587]" pageId="4" pageNumber="5">MFPC</collectionCode>
and
<collectionCode box="[1042,1118,1561,1587]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="5">NMW</collectionCode>
.
</paragraph>
<paragraph blockId="4.[264,1323,284,1906]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[312,593,1600,1626]" pageId="4" pageNumber="5">Differential Diagnosis.</emphasis>
Total length:
<number box="[779,815,1601,1627]" pageId="4" pageNumber="5" value="2.3">2.3</number>
–3.0 mm. The entirely melanic, iridescent pronotum easily distinguishes this species from all others in Central
<collectingCountry box="[1076,1177,1640,1666]" name="United States of America" pageId="4" pageNumber="5">America</collectingCountry>
(
<figureCitation box="[1194,1308,1640,1666]" captionStart="FIGURES 1 – 2" captionStartId="5.[264,382,1180,1204]" captionTargetBox="[431,1172,528,1139]" captionTargetId="figure@5.[396,1192,497,1154]" captionTargetPageId="5" captionText="FIGURES 1 – 2. Dorsal and lateral habitus of Hemiosus maculatus Sharp (specimen from Costa Rica). Scale bar 2.1 mm." httpUri="https://zenodo.org/record/174867/files/figure.png" pageId="4" pageNumber="5">
Figs.
<number box="[1261,1308,1640,1666]" pageId="4" pageNumber="5" value="1.5" valueMax="2.0" valueMin="1.0">1–2</number>
</figureCitation>
). There is no detectable external sexual dimorphism.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="description">
<paragraph blockId="4.[264,1323,284,1906]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[312,433,1720,1746]" pageId="4" pageNumber="5">Variation.</emphasis>
The elytral maculae can vary substantially both in size and intensity both within and between populations. It would not be a surprise to us if this species is found to be composite in the future.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="distribution">
<paragraph blockId="4.[264,1323,284,1906]" pageId="4" pageNumber="5">
<emphasis bold="true" box="[312,472,1839,1865]" pageId="4" pageNumber="5">Distribution.</emphasis>
From southern
<collectingCountry box="[682,772,1840,1866]" name="Mexico" pageId="4" pageNumber="5">Mexico</collectingCountry>
into South
<collectingCountry box="[920,1024,1840,1866]" name="United States of America" pageId="4" pageNumber="5">America</collectingCountry>
, including
<collectingCountry box="[1162,1240,1840,1866]" name="Belize" pageId="4" pageNumber="5">Belize</collectingCountry>
,
<collectingCountry name="Costa Rica" pageId="4" pageNumber="5">Costa Rica</collectingCountry>
,
<collectingCountry box="[332,462,1880,1906]" name="Guatemala" pageId="4" pageNumber="5">Guatemala</collectingCountry>
,
<collectingCountry box="[474,591,1880,1906]" name="Honduras" pageId="4" pageNumber="5">Honduras</collectingCountry>
,
<collectingCountry box="[602,695,1880,1906]" name="Mexico" pageId="4" pageNumber="5">Mexico</collectingCountry>
,
<collectingCountry box="[707,830,1880,1906]" name="Nicaragua" pageId="4" pageNumber="5">Nicaragua</collectingCountry>
,
<collectingCountry box="[842,938,1880,1906]" name="Panama" pageId="4" pageNumber="5">Panama</collectingCountry>
, and
<collectingCountry box="[999,1056,1880,1906]" name="Peru" pageId="4" pageNumber="5">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="discussion">
<paragraph blockId="5.[264,1323,283,470]" pageId="5" pageNumber="6">
<emphasis bold="true" box="[312,433,283,309]" pageId="5" pageNumber="6">Remarks.</emphasis>
A full description and additional illustrations of this species are provided by
<bibRefCitation author="Oliva" box="[302,458,324,350]" pageId="5" pageNumber="16" refString="Oliva, A. (1991). The types of Hemiosus Sharp, 1882 (Coleoptera: Hydrophilidae) in the Orchymont and Mouchamps collections. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 61, 167 - 181." type="journal article" year="1991">
Oliva (
<number box="[385,448,324,350]" pageId="5" pageNumber="6" value="1991.0">1991</number>
)
</bibRefCitation>
. In
<collectingCountry box="[512,645,324,350]" name="Costa Rica" pageId="5" pageNumber="6">Costa Rica</collectingCountry>
, this species was frequently collected in the margins and backwaters of sandy-bottomed streams, as well as shallow, isolated pools with sandy substrate along the margin of streams and rivers. It also is attracted to lights near such habitats.
</paragraph>
</subSubSection>
</treatment>
</document>
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