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<document id="FFF8949445BB647E06BC8F917B49E19A" ID-CLB-Dataset="172790" ID-DOI="10.11646/zootaxa.5198.1.1" ID-GBIF-Dataset="17babbe7-1645-47c8-ac5f-2299f1ad436c" ID-ISSN="1175-5326" ID-Zenodo-Dep="7251602" ID-ZooBank="019FE488-4263-4BC2-8606-446E599E226A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666769542164" checkinUser="plazi" docAuthor="Hou, Yanmeng, Gao, Zhizhong &amp; Zhang, Feng" docDate="2022" docId="8A6487A5FFFCA91CFF11FBB7FE63FE42" docLanguage="en" docName="zootaxa.5198.1.1.pdf" docOrigin="Zootaxa 5198 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5198.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Lagynochthonius xinjiaoensis Hou &amp; Gao &amp; Zhang 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="56" masterDocId="765DFFDDFFCEA92BFF86FFEAFFEBFFCB" masterDocTitle="Diversity of cave-dwelling pseudoscorpions from eastern Yunnan in China, with the description of eleven new species of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="65" masterPageNumber="1" pageNumber="51" updateTime="1701997268366" updateUser="plazi">
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<mods:title id="E78FC7FB1260F07D6B82F34B03F8365D">Diversity of cave-dwelling pseudoscorpions from eastern Yunnan in China, with the description of eleven new species of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:namePart id="EB9D8CF5D7456D2D2C63069FBC55B646">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="C6EAC59985EC2849B74C7A8034ED0527">Gao, Zhizhong</mods:namePart>
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<mods:namePart id="22AB965CF53E884232D9FE48A6DE9AFD">Zhang, Feng</mods:namePart>
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<treatment id="8A6487A5FFFCA91CFF11FBB7FE63FE42" ID-DOI="http://doi.org/10.5281/zenodo.7251634" ID-GBIF-Taxon="203411082" ID-Zenodo-Dep="7251634" LSID="urn:lsid:plazi:treatment:8A6487A5FFFCA91CFF11FBB7FE63FE42" httpUri="http://treatment.plazi.org/id/8A6487A5FFFCA91CFF11FBB7FE63FE42" lastPageId="55" lastPageNumber="56" pageId="50" pageNumber="51">
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<paragraph id="027236B3FFFCA919FF11FBB7FCFDFBBC" blockId="50.[151,790,1117,1179]" box="[151,790,1117,1144]" pageId="50" pageNumber="51">
<heading id="593A81DFFFFCA919FF11FBB7FCFDFBBC" bold="true" box="[151,790,1117,1144]" fontSize="11" level="2" pageId="50" pageNumber="51" reason="4">
<emphasis id="30B9EAA1FFFCA919FF11FBB7FCFDFBBC" bold="true" box="[151,790,1117,1144]" pageId="50" pageNumber="51">
<taxonomicName id="C5CD4D30FFFCA919FF11FBB7FDE8FBBC" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[151,515,1117,1144]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="xinjiaoensis" status="sp. nov.">
<emphasis id="30B9EAA1FFFCA919FF11FBB7FDE8FBBC" bold="true" box="[151,515,1117,1144]" italics="true" pageId="50" pageNumber="51">Lagynochthonius xinjiaoensis</emphasis>
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<taxonomicNameLabel id="2B8A57DAFFFCA919FD8DFBB7FD83FBBC" box="[523,616,1117,1143]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
(ṀfflIJ伪oi)
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</paragraph>
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<subSubSection id="4AD76538FFFCA919FF11FB6BFEFFFB50" box="[151,276,1153,1179]" pageId="50" pageNumber="51" type="description">
<paragraph id="027236B3FFFCA919FF11FB6BFEFFFB50" blockId="50.[151,790,1117,1179]" box="[151,276,1153,1179]" pageId="50" pageNumber="51">
<figureCitation id="9AF62A36FFFCA919FF11FB6BFEFFFB50" box="[151,276,1153,1179]" captionStart-0="FIGURE 38" captionStart-1="FIGURE 39" captionStart-2="FIGURE 40" captionStart-3="FIGURE 41" captionStartId-0="51.[151,250,1671,1695]" captionStartId-1="52.[151,250,2008,2032]" captionStartId-2="53.[151,250,966,990]" captionStartId-3="54.[151,250,1845,1869]" captionTargetBox-0="[151,1436,181,1647]" captionTargetBox-1="[173,1414,814,1985]" captionTargetBox-2="[151,1436,181,943]" captionTargetBox-3="[176,1410,181,1821]" captionTargetId-0="figure-19@51.[151,1436,181,1647]" captionTargetId-1="figure-346@52.[173,1414,813,1985]" captionTargetId-2="figure-19@53.[151,1436,181,943]" captionTargetId-3="figure-20@54.[176,1410,181,1821]" captionTargetPageId-0="51" captionTargetPageId-1="52" captionTargetPageId-2="53" captionTargetPageId-3="54" captionText-0="FIGURE 38. Yueliang Cave, type locality of Lagynochthonius xinjiaoensis sp. nov., A. Entrance; B. Inside the cave entrance; C. Ascent tunnel to the deepest part of the cave (red arrow); D. Area where L. xinjiaoensis sp. nov. specimen was collected; E. Live male of L. xinjiaoensis sp. nov., in its natural environment." captionText-1="FIGURE 39. Lagynochthonius xinjiaoensis sp. nov., holotype male, habitus (dorsal view)." captionText-2="FIGURE 40. Lagynochthonius xinjiaoensis sp. nov., holotype male (AD): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view)." captionText-3="FIGURE 41. Lagynochthonius xinjiaoensis sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern and finger tips (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi-0="http://doi.org/10.5281/zenodo.7251724" figureDoi-1="http://doi.org/10.5281/zenodo.7251726" figureDoi-2="http://doi.org/10.5281/zenodo.7251728" figureDoi-3="http://doi.org/10.5281/zenodo.7251730" httpUri-0="https://zenodo.org/record/7251724/files/figure.png" httpUri-1="https://zenodo.org/record/7251726/files/figure.png" httpUri-2="https://zenodo.org/record/7251728/files/figure.png" httpUri-3="https://zenodo.org/record/7251730/files/figure.png" pageId="50" pageNumber="51">Figs 3841</figureCitation>
</paragraph>
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<subSubSection id="4AD76538FFFCA919FF11FB2EFBE4FAED" pageId="50" pageNumber="51" type="materials_examined">
<paragraph id="027236B3FFFCA919FF11FB2EFBE4FAED" blockId="50.[151,1437,1220,2039]" pageId="50" pageNumber="51">
<emphasis id="30B9EAA1FFFCA919FF11FB2EFEA7FB15" bold="true" box="[151,332,1220,1246]" pageId="50" pageNumber="51">Type material.</emphasis>
<materialsCitation id="B2A53CEEFFFCA919FEDCFB2EFBE4FAED" ID-GBIF-Occurrence="3946861315" collectingDate="2021-10-08" collectionCode="MHBU-HBUARA" collectorName="Zegang Feng &amp; Yanmeng Hou &amp; Lu Zhang &amp; Liu Fu" country="China" county="Fuyuan County" elevation="2017" latitude="25.213259" location="Xinjiao Village" longLatPrecision="1" longitude="104.48424" municipality="Laochang Town" pageId="50" pageNumber="51" specimenCode="MHBU-HBUARA#2021-43001" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="DD768811FFFCA919FEDCFB2EFE29FB15" box="[346,450,1220,1246]" pageId="50" pageNumber="51" type="holotype">Holotype</typeStatus>
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(Ps.-
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):
<collectingCountry id="7ADA7623FFFCA919FC29FB2EFC1EFB15" box="[943,1013,1220,1246]" name="China" pageId="50" pageNumber="51">China</collectingCountry>
,
<collectingRegion id="C009F851FFFCA919FB80FB2FFB3BFB15" box="[1030,1232,1220,1247]" country="China" name="Yunnan" pageId="50" pageNumber="51">Yunnan Province</collectingRegion>
,
<collectingCounty id="EB134E3FFFFCA919FB65FB2FFA72FB15" box="[1251,1433,1220,1247]" pageId="50" pageNumber="51">Fuyuan County</collectingCounty>
,
<collectingMunicipality id="E216ACC9FFFCA919FF11FB02FEA6FAC8" box="[151,333,1256,1283]" pageId="50" pageNumber="51">Laochang Town</collectingMunicipality>
,
<location id="07126068FFFCA919FEDEFB02FDEEFAC9" LSID="urn:lsid:plazi:treatment:8A6487A5FFFCA91CFF11FBB7FE63FE42:07126068FFFCA919FEDEFB02FDEEFAC9" box="[344,517,1256,1282]" country="China" county="Fuyuan County" latitude="25.213259" longLatPrecision="1" longitude="104.48424" municipality="Laochang Town" name="Xinjiao Village" pageId="50" pageNumber="51" stateProvince="Yunnan">Xinjiao Village</location>
,
<location id="07126068FFFCA919FD88FB02FD5FFAC9" LSID="urn:lsid:plazi:treatment:8A6487A5FFFCA91CFF11FBB7FE63FE42:07126068FFFCA919FD88FB02FD5FFAC9" box="[526,692,1256,1282]" country="China" county="Fuyuan County" latitude="25.213259" longLatPrecision="1" longitude="104.48424" municipality="Laochang Town" name="Yueliang Cave" pageId="50" pageNumber="51" stateProvince="Yunnan">Yueliang Cave</location>
, under a stone in deep zone [
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,
<geoCoordinate id="67F95074FFFCA919FB2FFB03FAB8FACF" box="[1193,1363,1257,1284]" degrees="104" direction="east" minutes="29" orientation="longitude" pageId="50" pageNumber="51" precision="1" seconds="3.26" value="104.48424">104°293.26″E</geoCoordinate>
],
<elevation id="89E0D180FFFCA919FAE2FB02FF01FAED" metricMagnitude="3" metricUnit="m" metricValue="2.017" pageId="50" pageNumber="51" unit="m" value="2017.0">
<quantity id="C5359B56FFFCA919FAE2FB02FF46FAEC" metricMagnitude="3" metricUnit="m" metricValue="2.017" pageId="50" pageNumber="51" unit="m" value="2017.0">2017 m</quantity>
a.s.l.
</elevation>
,
<date id="76731073FFFCA919FF70FAE6FE4CFAED" box="[246,423,1292,1318]" pageId="50" pageNumber="51" value="2021-10-08">
<collectingDate id="6637E99BFFFCA919FF70FAE6FE4CFAED" box="[246,423,1292,1318]" pageId="50" pageNumber="51" value="2021-10-08">8 October 2021</collectingDate>
</date>
,
<collectorName id="AF385365FFFCA919FE34FAE7FDA3FAEC" box="[434,584,1293,1319]" pageId="50" pageNumber="51">Zegang Feng</collectorName>
,
<collectorName id="AF385365FFFCA919FDD4FAE7FD1DFAEC" box="[594,758,1293,1319]" pageId="50" pageNumber="51">Yanmeng Hou</collectorName>
,
<collectorName id="AF385365FFFCA919FC87FAE7FC9BFAED" box="[769,880,1292,1319]" pageId="50" pageNumber="51">Lu Zhang</collectorName>
&amp;
<collectorName id="AF385365FFFCA919FC15FAE6FC34FAEC" box="[915,991,1292,1319]" pageId="50" pageNumber="51">Liu Fu</collectorName>
leg.
</materialsCitation>
</paragraph>
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<subSubSection id="4AD76538FFFCA919FF41FADAFC3EFA81" box="[199,981,1328,1355]" pageId="50" pageNumber="51" type="etymology">
<paragraph id="027236B3FFFCA919FF41FADAFC3EFA81" blockId="50.[151,1437,1220,2039]" box="[199,981,1328,1355]" pageId="50" pageNumber="51">
<emphasis id="30B9EAA1FFFCA919FF41FADAFEA5FA81" bold="true" box="[199,334,1328,1354]" pageId="50" pageNumber="51">Etymology.</emphasis>
Named after the village of Xinjiao, near the
<typeStatus id="DD768811FFFCA919FCC2FADBFC9FFA80" box="[836,884,1329,1355]" pageId="50" pageNumber="51">type</typeStatus>
locality.
</paragraph>
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<subSubSection id="4AD76538FFFCA919FF41FABEFB23F98D" pageId="50" pageNumber="51" type="diagnosis">
<paragraph id="027236B3FFFCA919FF41FABEFB23F98D" blockId="50.[151,1437,1220,2039]" pageId="50" pageNumber="51">
<emphasis id="30B9EAA1FFFCA919FF41FABEFE9BFABA" bold="true" box="[199,368,1364,1393]" pageId="50" pageNumber="51">
Diagnosis (
<specimenCount id="14CBFD3AFFFCA919FECAFABFFE8AFABA" box="[332,353,1365,1393]" pageId="50" pageNumber="51" type="male"></specimenCount>
).
</emphasis>
Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, with 4 setae (including preocular setae) only, epistome pointed, small and triangular, without flanking basal setae; posterior margin of carapace with 2 setae; tergites IIV each with 2 setae. Pedipalps slender, femur 8.59 times longer than broad; chela robust, 7.03 times longer than broad; both chelal fingers without intercalary teeth; fixed chelal finger with a row of teeth that are distinctly larger than the teeth on the movable chelal finger, pointed and slightly retrorse and without a modified accessory tooth (
<emphasis id="30B9EAA1FFFCA919FADCF9E2FA9BF9E9" box="[1370,1392,1544,1570]" italics="true" pageId="50" pageNumber="51">td</emphasis>
) on dorso-antiaxial face; chemosensory setae (
<emphasis id="30B9EAA1FFFCA919FDF5F9C7FD61F98D" box="[627,650,1581,1606]" italics="true" pageId="50" pageNumber="51">sc</emphasis>
) present on dorsum of chelal hand; sensilla absent.
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFFCA91EFF41F9BAFBEDF947" lastPageId="53" lastPageNumber="54" pageId="50" pageNumber="51" type="description">
<paragraph id="027236B3FFFCA919FF41F9BAFC53F9A1" blockId="50.[151,1437,1220,2039]" box="[199,952,1616,1642]" pageId="50" pageNumber="51">
<emphasis id="30B9EAA1FFFCA919FF41F9BAFEB2F9A1" bold="true" box="[199,345,1616,1642]" pageId="50" pageNumber="51">Description.</emphasis>
Male (
<typeStatus id="DD768811FFFCA919FE2EF9BAFDFBF9A1" box="[424,528,1616,1642]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
), female unknown (
<figureCitation id="9AF62A36FFFCA919FD6BF9BAFCBDF9A1" box="[749,854,1616,1642]" captionStart="FIGURE 38" captionStartId="51.[151,250,1671,1695]" captionTargetBox="[151,1436,181,1647]" captionTargetId="figure-19@51.[151,1436,181,1647]" captionTargetPageId="51" captionText="FIGURE 38. Yueliang Cave, type locality of Lagynochthonius xinjiaoensis sp. nov., A. Entrance; B. Inside the cave entrance; C. Ascent tunnel to the deepest part of the cave (red arrow); D. Area where L. xinjiaoensis sp. nov. specimen was collected; E. Live male of L. xinjiaoensis sp. nov., in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251724" httpUri="https://zenodo.org/record/7251724/files/figure.png" pageId="50" pageNumber="51">Figs 38E</figureCitation>
,
<figureCitation id="9AF62A36FFFCA919FCE7F9BAFC47F9A1" box="[865,940,1616,1642]" captionStart-0="FIGURE 39" captionStart-1="FIGURE 40" captionStart-2="FIGURE 41" captionStartId-0="52.[151,250,2008,2032]" captionStartId-1="53.[151,250,966,990]" captionStartId-2="54.[151,250,1845,1869]" captionTargetBox-0="[173,1414,814,1985]" captionTargetBox-1="[151,1436,181,943]" captionTargetBox-2="[176,1410,181,1821]" captionTargetId-0="figure-346@52.[173,1414,813,1985]" captionTargetId-1="figure-19@53.[151,1436,181,943]" captionTargetId-2="figure-20@54.[176,1410,181,1821]" captionTargetPageId-0="52" captionTargetPageId-1="53" captionTargetPageId-2="54" captionText-0="FIGURE 39. Lagynochthonius xinjiaoensis sp. nov., holotype male, habitus (dorsal view)." captionText-1="FIGURE 40. Lagynochthonius xinjiaoensis sp. nov., holotype male (AD): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view)." captionText-2="FIGURE 41. Lagynochthonius xinjiaoensis sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern and finger tips (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi-0="http://doi.org/10.5281/zenodo.7251726" figureDoi-1="http://doi.org/10.5281/zenodo.7251728" figureDoi-2="http://doi.org/10.5281/zenodo.7251730" httpUri-0="https://zenodo.org/record/7251726/files/figure.png" httpUri-1="https://zenodo.org/record/7251728/files/figure.png" httpUri-2="https://zenodo.org/record/7251730/files/figure.png" pageId="50" pageNumber="51">3941</figureCitation>
).
</paragraph>
<paragraph id="027236B3FFFCA919FF41F99EFF26F979" blockId="50.[151,1437,1220,2039]" pageId="50" pageNumber="51">
Colour (
<figureCitation id="9AF62A36FFFCA919FEA3F99EFE7BF945" box="[293,400,1652,1678]" captionStart="FIGURE 38" captionStartId="51.[151,250,1671,1695]" captionTargetBox="[151,1436,181,1647]" captionTargetId="figure-19@51.[151,1436,181,1647]" captionTargetPageId="51" captionText="FIGURE 38. Yueliang Cave, type locality of Lagynochthonius xinjiaoensis sp. nov., A. Entrance; B. Inside the cave entrance; C. Ascent tunnel to the deepest part of the cave (red arrow); D. Area where L. xinjiaoensis sp. nov. specimen was collected; E. Live male of L. xinjiaoensis sp. nov., in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251724" httpUri="https://zenodo.org/record/7251724/files/figure.png" pageId="50" pageNumber="51">Figs 38E</figureCitation>
,
<figureCitation id="9AF62A36FFFCA919FE1BF99EFE50F945" box="[413,443,1652,1678]" captionStart="FIGURE 39" captionStartId="52.[151,250,2008,2032]" captionTargetBox="[173,1414,814,1985]" captionTargetId="figure-346@52.[173,1414,813,1985]" captionTargetPageId="52" captionText="FIGURE 39. Lagynochthonius xinjiaoensis sp. nov., holotype male, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7251726" httpUri="https://zenodo.org/record/7251726/files/figure.png" pageId="50" pageNumber="51">39</figureCitation>
,
<figureCitation id="9AF62A36FFFCA919FE4EF99EFE02F945" box="[456,489,1652,1678]" captionStart="FIGURE 40" captionStartId="53.[151,250,966,990]" captionTargetBox="[151,1436,181,943]" captionTargetId="figure-19@53.[151,1436,181,943]" captionTargetPageId="53" captionText="FIGURE 40. Lagynochthonius xinjiaoensis sp. nov., holotype male (AD): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251728" httpUri="https://zenodo.org/record/7251728/files/figure.png" pageId="50" pageNumber="51">40</figureCitation>
): generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.
</paragraph>
<paragraph id="027236B3FFFCA919FF41F956FC1EF83C" blockId="50.[151,1437,1220,2039]" pageId="50" pageNumber="51">
Cephalothorax (
<figureCitation id="9AF62A36FFFCA919FEFDF956FE0EF91D" box="[379,485,1724,1750]" captionStart="FIGURE 40" captionStartId="53.[151,250,966,990]" captionTargetBox="[151,1436,181,943]" captionTargetId="figure-19@53.[151,1436,181,943]" captionTargetPageId="53" captionText="FIGURE 40. Lagynochthonius xinjiaoensis sp. nov., holotype male (AD): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251728" httpUri="https://zenodo.org/record/7251728/files/figure.png" pageId="50" pageNumber="51">Figs 40B</figureCitation>
,
<figureCitation id="9AF62A36FFFCA919FE77F956FDCEF91D" box="[497,549,1724,1750]" captionStart="FIGURE 41" captionStartId="54.[151,250,1845,1869]" captionTargetBox="[176,1410,181,1821]" captionTargetId="figure-20@54.[176,1410,181,1821]" captionTargetPageId="54" captionText="FIGURE 41. Lagynochthonius xinjiaoensis sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern and finger tips (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251730" httpUri="https://zenodo.org/record/7251730/files/figure.png" pageId="50" pageNumber="51">41A</figureCitation>
): carapace 0.98 times longer than broad, squarish, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular; with 16 setae arranged s2s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 2, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 1011 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
<figureCitation id="9AF62A36FFFCA919FEDFF836FE2AF83D" box="[345,449,2012,2038]" captionStart="FIGURE 41" captionStartId="54.[151,250,1845,1869]" captionTargetBox="[176,1410,181,1821]" captionTargetId="figure-20@54.[176,1410,181,1821]" captionTargetPageId="54" captionText="FIGURE 41. Lagynochthonius xinjiaoensis sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern and finger tips (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251730" httpUri="https://zenodo.org/record/7251730/files/figure.png" pageId="50" pageNumber="51">Fig. 41C</figureCitation>
); intercoxal tubercle absent; without sub-oral seta.
</paragraph>
<caption id="56B2663BFFFDA918FF11F96DFCC8F92C" ID-DOI="http://doi.org/10.5281/zenodo.7251724" ID-Zenodo-Dep="7251724" httpUri="https://zenodo.org/record/7251724/files/figure.png" pageId="51" pageNumber="52" startId="51.[151,250,1671,1695]" targetBox="[151,1436,181,1647]" targetPageId="51">
<paragraph id="027236B3FFFDA918FF11F96DFCC8F92C" blockId="51.[151,1437,1671,1767]" pageId="51" pageNumber="52">
<emphasis id="30B9EAA1FFFDA918FF11F96DFECBF954" bold="true" box="[151,288,1671,1695]" pageId="51" pageNumber="52">FIGURE 38.</emphasis>
Yueliang Cave, type locality of
<taxonomicName id="C5CD4D30FFFDA918FDEFF96DFC7CF954" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[617,919,1671,1695]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="xinjiaoensis" status="sp. nov.">
<emphasis id="30B9EAA1FFFDA918FDEFF96DFC7CF954" box="[617,919,1671,1695]" italics="true" pageId="51" pageNumber="52">Lagynochthonius xinjiaoensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFFDA918FC18F96DFC04F954" bold="true" box="[926,1007,1671,1695]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="2B8A57DAFFFDA918FC18F96DFC04F954" box="[926,1007,1671,1695]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Entrance; B. Inside the cave entrance; C. Ascent tunnel to the deepest part of the cave (red arrow); D. Area where
<taxonomicName id="C5CD4D30FFFDA918FC11F946FBC0F908" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[919,1067,1708,1731]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="xinjiaoensis" status="sp. nov.">
<emphasis id="30B9EAA1FFFDA918FC11F946FBC0F908" box="[919,1067,1708,1731]" italics="true" pageId="51" pageNumber="52">L. xinjiaoensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFFDA918FBB4F941FB69F908" bold="true" box="[1074,1154,1707,1731]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="2B8A57DAFFFDA918FBB4F941FB69F908" box="[1074,1154,1707,1731]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimen was collected; E. Live male of
<taxonomicName id="C5CD4D30FFFDA918FE99F93AFE58F92C" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[287,435,1744,1767]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="xinjiaoensis" status="sp. nov.">
<emphasis id="30B9EAA1FFFDA918FE99F93AFE58F92C" box="[287,435,1744,1767]" italics="true" pageId="51" pageNumber="52">L. xinjiaoensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFFDA918FE3FF925FDE1F92C" bold="true" box="[441,522,1743,1767]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="2B8A57DAFFFDA918FE3FF925FDE1F92C" box="[441,522,1743,1767]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in its natural environment.
</paragraph>
</caption>
<paragraph id="027236B3FFFDA91FFF41F8FFFD16FF1E" blockId="51.[151,1436,1813,2020]" lastBlockId="52.[151,1437,151,789]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
Chelicera (
<figureCitation id="9AF62A36FFFDA918FEC0F8FFFE5FF8E4" box="[326,436,1813,1839]" captionStart="FIGURE 40" captionStartId="53.[151,250,966,990]" captionTargetBox="[151,1436,181,943]" captionTargetId="figure-19@53.[151,1436,181,943]" captionTargetPageId="53" captionText="FIGURE 40. Lagynochthonius xinjiaoensis sp. nov., holotype male (AD): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251728" httpUri="https://zenodo.org/record/7251728/files/figure.png" pageId="51" pageNumber="52">Figs 40C</figureCitation>
,
<figureCitation id="9AF62A36FFFDA918FE42F8FFFE1DF8E4" box="[452,502,1813,1839]" captionStart="FIGURE 41" captionStartId="54.[151,250,1845,1869]" captionTargetBox="[176,1410,181,1821]" captionTargetId="figure-20@54.[176,1410,181,1821]" captionTargetPageId="54" captionText="FIGURE 41. Lagynochthonius xinjiaoensis sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern and finger tips (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251730" httpUri="https://zenodo.org/record/7251730/files/figure.png" pageId="51" pageNumber="52">41B</figureCitation>
): large, about as long as carapace, 2.45 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 22 teeth, distal one largest; movable finger with 16 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (
<figureCitation id="9AF62A36FFFDA918FF19F823FEEDF828" box="[159,262,1993,2019]" captionStart="FIGURE 41" captionStartId="54.[151,250,1845,1869]" captionTargetBox="[176,1410,181,1821]" captionTargetId="figure-20@54.[176,1410,181,1821]" captionTargetPageId="54" captionText="FIGURE 41. Lagynochthonius xinjiaoensis sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern and finger tips (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251730" httpUri="https://zenodo.org/record/7251730/files/figure.png" pageId="51" pageNumber="52">Fig. 41B</figureCitation>
). Serrula exterior with 27 and serrula interior with 14 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
<figureCitation id="9AF62A36FFFAA91FFD0EFF51FD1AFF1E" box="[648,753,187,213]" captionStart="FIGURE 41" captionStartId="54.[151,250,1845,1869]" captionTargetBox="[176,1410,181,1821]" captionTargetId="figure-20@54.[176,1410,181,1821]" captionTargetPageId="54" captionText="FIGURE 41. Lagynochthonius xinjiaoensis sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern and finger tips (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251730" httpUri="https://zenodo.org/record/7251730/files/figure.png" pageId="52" pageNumber="53">Fig. 41D</figureCitation>
).
</paragraph>
<paragraph id="027236B3FFFAA91FFF41FF35FBA5FCDE" blockId="52.[151,1437,151,789]" pageId="52" pageNumber="53">
Pedipalp (
<figureCitation id="9AF62A36FFFAA91FFEBEFF35FE48FF32" box="[312,419,223,249]" captionStart="FIGURE 40" captionStartId="53.[151,250,966,990]" captionTargetBox="[151,1436,181,943]" captionTargetId="figure-19@53.[151,1436,181,943]" captionTargetPageId="53" captionText="FIGURE 40. Lagynochthonius xinjiaoensis sp. nov., holotype male (AD): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251728" httpUri="https://zenodo.org/record/7251728/files/figure.png" pageId="52" pageNumber="53">Figs 40A</figureCitation>
,
<figureCitation id="9AF62A36FFFAA91FFE29FF35FDEFFF32" box="[431,516,223,249]" captionStart="FIGURE 41" captionStartId="54.[151,250,1845,1869]" captionTargetBox="[176,1410,181,1821]" captionTargetId="figure-20@54.[176,1410,181,1821]" captionTargetPageId="54" captionText="FIGURE 41. Lagynochthonius xinjiaoensis sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern and finger tips (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251730" httpUri="https://zenodo.org/record/7251730/files/figure.png" pageId="52" pageNumber="53">41EG</figureCitation>
): long and slender, trochanter 1.90, femur 8.59, patella 2.22, chela 7.03, hand 2.62 times longer than broad; femur 2.86 times longer than patella; movable chelal finger 1.71 times longer than hand and 0.64 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (
<figureCitation id="9AF62A36FFFAA91FFAACFECDFA7BFE8A" box="[1322,1424,295,321]" captionStart="FIGURE 41" captionStartId="54.[151,250,1845,1869]" captionTargetBox="[176,1410,181,1821]" captionTargetId="figure-20@54.[176,1410,181,1821]" captionTargetPageId="54" captionText="FIGURE 41. Lagynochthonius xinjiaoensis sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern and finger tips (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251730" httpUri="https://zenodo.org/record/7251730/files/figure.png" pageId="52" pageNumber="53">Fig. 41E</figureCitation>
). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria,
<emphasis id="30B9EAA1FFFAA91FFD14FE79FD43FE66" box="[658,680,403,429]" italics="true" pageId="52" pageNumber="53">ib</emphasis>
and
<emphasis id="30B9EAA1FFFAA91FFD63FE79FCEDFE66" box="[741,774,403,429]" italics="true" pageId="52" pageNumber="53">isb</emphasis>
situated close together, submedially on dorsum of chelal hand;
<emphasis id="30B9EAA1FFFAA91FFF58FE5DFF13FE1A" box="[222,248,439,465]" italics="true" pageId="52" pageNumber="53">eb</emphasis>
,
<emphasis id="30B9EAA1FFFAA91FFE81FE5DFEC7FE1A" box="[263,300,439,465]" italics="true" pageId="52" pageNumber="53">esb</emphasis>
and
<emphasis id="30B9EAA1FFFAA91FFEE3FE5DFE94FE1B" box="[357,383,439,464]" italics="true" pageId="52" pageNumber="53">ist</emphasis>
forming a straight oblique row at base of fixed chelal finger;
<emphasis id="30B9EAA1FFFAA91FFBC6FE5DFBBBFE1B" box="[1088,1104,439,464]" italics="true" pageId="52" pageNumber="53">it</emphasis>
slightly distal to
<emphasis id="30B9EAA1FFFAA91FFA91FE5DFADDFE1B" box="[1303,1334,439,464]" italics="true" pageId="52" pageNumber="53">est</emphasis>
, situated subdistally;
<emphasis id="30B9EAA1FFFAA91FFE99FE31FED8FE3F" box="[287,307,475,500]" italics="true" pageId="52" pageNumber="53">et</emphasis>
slightly near to tip of fixed chelal finger, very close to chelal teeth;
<emphasis id="30B9EAA1FFFAA91FFBA1FE31FBAAFE3E" box="[1063,1089,475,501]" italics="true" pageId="52" pageNumber="53">dx</emphasis>
situated distal to
<emphasis id="30B9EAA1FFFAA91FFA80FE31FAF1FE3F" box="[1286,1306,475,500]" italics="true" pageId="52" pageNumber="53">et</emphasis>
;
<emphasis id="30B9EAA1FFFAA91FFAAFFE31FAA9FE3E" box="[1321,1346,475,501]" italics="true" pageId="52" pageNumber="53">sb</emphasis>
slightly closer to
<emphasis id="30B9EAA1FFFAA91FFE85FE15FEFAFDD2" box="[259,273,511,537]" italics="true" pageId="52" pageNumber="53">b</emphasis>
than to
<emphasis id="30B9EAA1FFFAA91FFEF5FE15FE6DFDD3" box="[371,390,511,536]" italics="true" pageId="52" pageNumber="53">st</emphasis>
;
<emphasis id="30B9EAA1FFFAA91FFE11FE15FE4EFDD2" box="[407,421,511,537]" italics="true" pageId="52" pageNumber="53">b</emphasis>
and
<emphasis id="30B9EAA1FFFAA91FFE66FE15FE03FDD3" box="[480,488,511,536]" italics="true" pageId="52" pageNumber="53">t</emphasis>
situated subdistally,
<emphasis id="30B9EAA1FFFAA91FFD5FFE15FD0CFDD2" box="[729,743,511,537]" italics="true" pageId="52" pageNumber="53">b</emphasis>
situated at same level as
<emphasis id="30B9EAA1FFFAA91FFB94FE15FBDAFDD3" box="[1042,1073,511,536]" italics="true" pageId="52" pageNumber="53">est</emphasis>
;
<emphasis id="30B9EAA1FFFAA91FFBC4FE15FBA1FDD3" box="[1090,1098,511,536]" italics="true" pageId="52" pageNumber="53">t</emphasis>
distal to
<emphasis id="30B9EAA1FFFAA91FFB3FFE15FB22FDD3" box="[1209,1225,511,536]" italics="true" pageId="52" pageNumber="53">it</emphasis>
(
<figureCitation id="9AF62A36FFFAA91FFB5CFE15FAAAFDD2" box="[1242,1345,511,537]" captionStart="FIGURE 41" captionStartId="54.[151,250,1845,1869]" captionTargetBox="[176,1410,181,1821]" captionTargetId="figure-20@54.[176,1410,181,1821]" captionTargetPageId="54" captionText="FIGURE 41. Lagynochthonius xinjiaoensis sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern and finger tips (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251730" httpUri="https://zenodo.org/record/7251730/files/figure.png" pageId="52" pageNumber="53">Fig. 41F</figureCitation>
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to
<emphasis id="30B9EAA1FFFAA91FFC68FDC9FBE3FDF7" box="[1006,1032,547,572]" italics="true" pageId="52" pageNumber="53">ist</emphasis>
). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 35 macrodenticles, pointed and gently retrorse, without a modified accessory tooth (
<emphasis id="30B9EAA1FFFAA91FFD7CFD65FCFBFD62" box="[762,784,655,681]" italics="true" pageId="52" pageNumber="53">td</emphasis>
) on dorso-antiaxial face; movable chelal finger with 28 macrodenticles (markedly smaller than the teeth on fixed chelal finger), pointed and gently retrorse, plus 4 vestigial, rounded and contiguous basal teeth,
<quantity id="C5359B56FFFAA91FFDB1FD3DFD99FD3A" box="[567,626,727,753]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" pageId="52" pageNumber="53" unit="in" value="33.0">33 in</quantity>
total (
<figureCitation id="9AF62A36FFFAA91FFD3BFD3DFCFFFD3A" box="[701,788,726,753]" captionStart="FIGURE 5" captionStartId="9.[151,250,1829,1853]" captionTargetBox="[186,1403,183,1797]" captionTargetId="figure-19@9.[185,1403,181,1805]" captionTargetPageId="9" captionText="FIGURE 5. Lagynochthonius crassus sp. nov., holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251616" httpUri="https://zenodo.org/record/7251616/files/figure.png" pageId="52" pageNumber="53">Fig. 5F</figureCitation>
). Sensilla absent. Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
<figureCitation id="9AF62A36FFFAA91FFC5FFD11FBA9FCDE" box="[985,1090,763,789]" captionStart="FIGURE 41" captionStartId="54.[151,250,1845,1869]" captionTargetBox="[176,1410,181,1821]" captionTargetId="figure-20@54.[176,1410,181,1821]" captionTargetPageId="54" captionText="FIGURE 41. Lagynochthonius xinjiaoensis sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern and finger tips (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251730" httpUri="https://zenodo.org/record/7251730/files/figure.png" pageId="52" pageNumber="53">Fig. 41G</figureCitation>
).
</paragraph>
<caption id="56B2663BFFFAA91FFF11F832FBD9F83B" ID-DOI="http://doi.org/10.5281/zenodo.7251726" ID-Zenodo-Dep="7251726" box="[151,1074,2008,2032]" httpUri="https://zenodo.org/record/7251726/files/figure.png" pageId="52" pageNumber="53" startId="52.[151,250,2008,2032]" targetBox="[173,1414,814,1985]" targetPageId="52">
<paragraph id="027236B3FFFAA91FFF11F832FBD9F83B" blockId="52.[151,1074,2008,2032]" box="[151,1074,2008,2032]" pageId="52" pageNumber="53">
<emphasis id="30B9EAA1FFFAA91FFF11F832FECBF83B" bold="true" box="[151,288,2008,2032]" pageId="52" pageNumber="53">FIGURE 39.</emphasis>
<taxonomicName id="C5CD4D30FFFAA91FFEA0F832FDBFF83B" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[294,596,2008,2032]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="xinjiaoensis" status="sp. nov.">
<emphasis id="30B9EAA1FFFAA91FFEA0F832FDBFF83B" box="[294,596,2008,2032]" italics="true" pageId="52" pageNumber="53">Lagynochthonius xinjiaoensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFFAA91FFDDCF832FD40F83B" bold="true" box="[602,683,2008,2032]" pageId="52" pageNumber="53">
<taxonomicNameLabel id="2B8A57DAFFFAA91FFDDCF832FD40F83B" box="[602,683,2008,2032]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male, habitus (dorsal view).
</paragraph>
</caption>
<caption id="56B2663BFFFBA91EFF11FC2CFC66FBC8" ID-DOI="http://doi.org/10.5281/zenodo.7251728" ID-Zenodo-Dep="7251728" httpUri="https://zenodo.org/record/7251728/files/figure.png" pageId="53" pageNumber="54" startId="53.[151,250,966,990]" targetBox="[151,1436,181,943]" targetPageId="53">
<paragraph id="027236B3FFFBA91EFF11FC2CFC66FBC8" blockId="53.[151,1436,966,1027]" pageId="53" pageNumber="54">
<emphasis id="30B9EAA1FFFBA91EFF11FC2CFECAFC14" bold="true" box="[151,289,966,991]" pageId="53" pageNumber="54">FIGURE 40.</emphasis>
<taxonomicName id="C5CD4D30FFFBA91EFEAFFC2DFDB3FC15" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[297,600,967,991]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="xinjiaoensis" status="sp. nov.">
<emphasis id="30B9EAA1FFFBA91EFEAFFC2DFDB3FC15" box="[297,600,967,991]" italics="true" pageId="53" pageNumber="54">Lagynochthonius xinjiaoensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFFBA91EFDE6FC2DFD59FC14" bold="true" box="[608,690,967,991]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="2B8A57DAFFFBA91EFDE6FC2DFD59FC14" box="[608,690,967,991]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (AD): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view).
</paragraph>
</caption>
<paragraph id="027236B3FFFBA91EFF41FBDBFB80FB7C" blockId="53.[151,1437,1073,2035]" pageId="53" pageNumber="54">
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy IXII: 2: 2: 2: 2: 4: 4: 4: 4: 5: 2: T2T: 0, tergites IX with an unpaired median seta. Sternal chaetotaxy IIIXII: 7: 10: 10: 9: 9: 9: 9: 7: 0: 2. Anterior genital operculum with 12 setae, genital opening slit-like, with 1112 marginal setae on each side,
<quantity id="C5359B56FFFBA91EFCF3FB77FC45FB7C" box="[885,942,1181,1208]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="53" pageNumber="54" unit="in" value="35.0">35 in</quantity>
total (
<figureCitation id="9AF62A36FFFBA91EFC70FB77FB8BFB7C" box="[1014,1120,1181,1207]" captionStart="FIGURE 40" captionStartId="53.[151,250,966,990]" captionTargetBox="[151,1436,181,943]" captionTargetId="figure-19@53.[151,1436,181,943]" captionTargetPageId="53" captionText="FIGURE 40. Lagynochthonius xinjiaoensis sp. nov., holotype male (AD): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251728" httpUri="https://zenodo.org/record/7251728/files/figure.png" pageId="53" pageNumber="54">Fig. 40D</figureCitation>
).
</paragraph>
<paragraph id="027236B3FFFBA91EFF41FB2BFEDBFA78" blockId="53.[151,1437,1073,2035]" pageId="53" pageNumber="54">
Legs (
<figureCitation id="9AF62A36FFFBA91EFE8BFB2BFE60FB10" box="[269,395,1217,1243]" captionStart="FIGURE 41" captionStartId="54.[151,250,1845,1869]" captionTargetBox="[176,1410,181,1821]" captionTargetId="figure-20@54.[176,1410,181,1821]" captionTargetPageId="54" captionText="FIGURE 41. Lagynochthonius xinjiaoensis sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern and finger tips (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD)." figureDoi="http://doi.org/10.5281/zenodo.7251730" httpUri="https://zenodo.org/record/7251730/files/figure.png" pageId="53" pageNumber="54">Fig. 41H, I</figureCitation>
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur I, IV and patella IV. Femur of leg I 1.89 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.69 times longer than tibia. Femoropatella of leg IV 4.85 times longer than deep; tibia 7.17 times longer than deep; with basal tactile setae on both tarsal segments: basitarsus 4.10 times longer than deep (TS= 0.24), telotarsus 15.14 times longer than deep and 2.59 times longer than basitarsus (TS= 0.33). Setae of leg I (trochanter to tibia) 4: 12: 11: 11, setae of leg IV (trochanter to basitarsus) 2: 3: 6: 11: 13. Arolium slightly shorter than the claws, not divided; claws simple.
</paragraph>
<paragraph id="027236B3FFFBA91EFF41FA57FBEDF947" blockId="53.[151,1437,1073,2035]" pageId="53" pageNumber="54">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Male: body length 2.15. Pedipalps: trochanter 0.40/0.21 (1.90), femur 1.46/0.17 (8.59), patella 0.51/0.23 (2.22), chela 2.04/0.29 (7.03), hand 0.76/0.29 (2.62), movable chelal finger length 1.30. Chelicera 0.76/0.31 (2.45), movable finger length 0.41. Carapace 0.65/0.66 (0.98). Leg I: trochanter 0.24/0.16 (1.50), femur 0.85/0.10 (8.50), patella 0.45/0.09 (5.00), tibia 0.35/0.06 (5.83), tarsus 0.94/0.07 (13.43). Leg IV: trochanter 0.33/0.18 (1.83), femoropatella 1.26/0.26 (4.85), tibia 0.86/0.12 (7.17), basitarsus 0.41/0.10 (4.10), telotarsus 1.06/0.07 (15.14).</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFFBA91EFF41F97FFDBDF838" pageId="53" pageNumber="54" type="discussion">
<paragraph id="027236B3FFFBA91EFF41F97FFAE6F8A8" blockId="53.[151,1437,1073,2035]" pageId="53" pageNumber="54">
<emphasis id="30B9EAA1FFFBA91EFF41F97FFED0F964" bold="true" box="[199,315,1685,1711]" pageId="53" pageNumber="54">Remarks.</emphasis>
Female unknown.
<taxonomicName id="C5CD4D30FFFBA91EFD8BF97FFCB2F964" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[525,857,1685,1711]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="xinjiaoensis" status="sp. nov.">
<emphasis id="30B9EAA1FFFBA91EFD8BF97FFCB2F964" box="[525,857,1685,1711]" italics="true" pageId="53" pageNumber="54">Lagynochthonius xinjiaoensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFFBA91EFCD8F97FFC5DF964" bold="true" box="[862,950,1685,1711]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="2B8A57DAFFFBA91EFCD8F97FFC5DF964" box="[862,950,1685,1711]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="C5CD4D30FFFBA91EFBF8F97CFAD4F964" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[1150,1343,1685,1711]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="magnidentatus" status="sp. nov.">
<emphasis id="30B9EAA1FFFBA91EFBF8F97CFAD4F964" box="[1150,1343,1685,1711]" italics="true" pageId="53" pageNumber="54">L. magnidentatus</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFFBA91EFAC2F97FFA77F964" bold="true" box="[1348,1436,1685,1711]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="2B8A57DAFFFBA91EFAC2F97FFA77F964" box="[1348,1436,1685,1711]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having only 2 anterolateral setae on the carapace (except preocular setae), an equal number of setae on tergites IIV, and a similarly-shaped chelal hand, but differs by the presence of a larger chela (e.g. chela 7.03 times as long as broad
<emphasis id="30B9EAA1FFFBA91EFF7BF8E8FEF0F8D0" box="[253,283,1794,1819]" italics="true" pageId="53" pageNumber="54">vs.</emphasis>
6.446.46 times, chela length 2.04
<emphasis id="30B9EAA1FFFBA91EFD2BF8E8FD2FF8D0" box="[685,708,1794,1819]" italics="true" pageId="53" pageNumber="54">vs</emphasis>
.
<quantity id="C5359B56FFFBA91EFD55F8EBFC92F8D0" box="[723,889,1793,1820]" metricMagnitude="-3" metricUnit="m" metricValue="1.58" metricValueMax="1.61" metricValueMin="1.55" pageId="53" pageNumber="54" unit="mm" value="1.58" valueMax="1.61" valueMin="1.55">1.551.61 mm</quantity>
, movable chelal finger length 1.30
<emphasis id="30B9EAA1FFFBA91EFA81F8E8FAF5F8D0" box="[1287,1310,1794,1819]" italics="true" pageId="53" pageNumber="54">vs</emphasis>
.
<quantity id="C5359B56FFFBA91EFAAAF8EBFF2DF8F4" metricMagnitude="-4" metricUnit="m" metricValue="9.75" metricValueMax="9.9" metricValueMin="9.6" pageId="53" pageNumber="54" unit="mm" value="0.975" valueMax="0.99" valueMin="0.96">0.960.99 mm</quantity>
; all in males), a different epistome shape (pointed and triangular vs. rounded and obtuse), and the position of trichobothrium
<emphasis id="30B9EAA1FFFBA91EFEC0F8A3FEB4F8A8" box="[326,351,1865,1891]" italics="true" pageId="53" pageNumber="54">sb</emphasis>
(
<emphasis id="30B9EAA1FFFBA91EFEE9F8A3FE63F8A8" box="[367,392,1865,1891]" italics="true" pageId="53" pageNumber="54">sb</emphasis>
1.07 times as far from
<emphasis id="30B9EAA1FFFBA91EFD0BF8A3FD70F8A8" box="[653,667,1865,1891]" italics="true" pageId="53" pageNumber="54">b</emphasis>
as from
<emphasis id="30B9EAA1FFFBA91EFD78F8A0FCDDF8A8" box="[766,822,1866,1891]" italics="true" pageId="53" pageNumber="54">st vs.</emphasis>
1.281.30 times as far from
<emphasis id="30B9EAA1FFFBA91EFBFFF8A3FB6CF8A8" box="[1145,1159,1865,1891]" italics="true" pageId="53" pageNumber="54">b</emphasis>
as from
<emphasis id="30B9EAA1FFFBA91EFB6CF8A0FB16F8A8" box="[1258,1277,1866,1891]" italics="true" pageId="53" pageNumber="54">st</emphasis>
).
</paragraph>
<paragraph id="027236B3FFFBA91EFF41F887FDBDF838" blockId="53.[151,1437,1073,2035]" pageId="53" pageNumber="54">
The discovery of two new species (
<taxonomicName id="C5CD4D30FFFBA91EFDCBF884FCE5F84C" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[589,782,1901,1927]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="magnidentatus" status="sp. nov.">
<emphasis id="30B9EAA1FFFBA91EFDCBF884FCE5F84C" box="[589,782,1901,1927]" italics="true" pageId="53" pageNumber="54">L. magnidentatus</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFFBA91EFC92F887FC87F84C" bold="true" box="[788,876,1901,1927]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="2B8A57DAFFFBA91EFC92F887FC87F84C" box="[788,876,1901,1927]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="C5CD4D30FFFBA91EFC26F884FBA8F84C" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[928,1091,1902,1927]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="xinjiaoensis" status="sp. nov.">
<emphasis id="30B9EAA1FFFBA91EFC26F884FBA8F84C" box="[928,1091,1902,1927]" italics="true" pageId="53" pageNumber="54">L. xinjiaoensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFFBA91EFBCEF887FB4AF84C" bold="true" box="[1096,1185,1901,1927]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="2B8A57DAFFFBA91EFBCEF887FB4AF84C" box="[1096,1185,1901,1927]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) that lack the anteriormedian setae on the carapace is significant. Indeed, there is no other chthoniid that lacks these two setae.Unfortunately, apart from this character, we failed to find any other evidence not belonging to the genus
<taxonomicName id="C5CD4D30FFFBA91EFBE5F85FFACFF804" authorityName="Beier" authorityYear="1951" box="[1123,1316,1973,1999]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="30B9EAA1FFFBA91EFBE5F85FFACFF804" box="[1123,1316,1973,1999]" italics="true" pageId="53" pageNumber="54">Lagynochthonius</emphasis>
</taxonomicName>
, therefore, it is inappropriate to name a new genus.
</paragraph>
</subSubSection>
<caption id="56B2663BFFF8A91DFF11F8DFFBDBF815" ID-DOI="http://doi.org/10.5281/zenodo.7251730" ID-Zenodo-Dep="7251730" httpUri="https://zenodo.org/record/7251730/files/figure.png" pageId="54" pageNumber="55" startId="54.[151,250,1845,1869]" targetBox="[176,1410,181,1821]" targetPageId="54">
<paragraph id="027236B3FFF8A91DFF11F8DFFBDBF815" blockId="54.[151,1437,1845,2014]" pageId="54" pageNumber="55">
<emphasis id="30B9EAA1FFF8A91DFF11F8DFFEF4F886" bold="true" box="[151,287,1845,1869]" pageId="54" pageNumber="55">FIGURE 41.</emphasis>
<taxonomicName id="C5CD4D30FFF8A91DFEA2F8DFFDBAF886" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[292,593,1845,1869]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="xinjiaoensis" status="sp. nov.">
<emphasis id="30B9EAA1FFF8A91DFEA2F8DFFDBAF886" box="[292,593,1845,1869]" italics="true" pageId="54" pageNumber="55">Lagynochthonius xinjiaoensis</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFF8A91DFDD0F8DCFD4DF885" bold="true" box="[598,678,1846,1870]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="2B8A57DAFFF8A91DFDD0F8DCFD4DF885" box="[598,678,1846,1870]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth and galea; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern and finger tips (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (AB, EI); 0.10 mm (CD).
</paragraph>
</caption>
<subSubSection id="4AD76538FFF9A91CFF41FF7CFE63FE42" pageId="55" pageNumber="56" type="distribution">
<paragraph id="027236B3FFF9A91CFF41FF7CFE63FE42" blockId="55.[151,1437,150,393]" pageId="55" pageNumber="56">
<emphasis id="30B9EAA1FFF9A91CFF41FF7CFE1BFF7B" bold="true" box="[199,496,150,177]" pageId="55" pageNumber="56">Distribution and habitat.</emphasis>
This species is known only from the
<typeStatus id="DD768811FFF9A91CFC17FF7DFC2AFF7A" box="[913,961,151,177]" pageId="55" pageNumber="56">type</typeStatus>
locality, Yueliang Cave (
<figureCitation id="9AF62A36FFF9A91CFB5EFF7DFAC5FF7A" box="[1240,1326,151,177]" captionStart="FIGURE 1" captionStartId="3.[151,250,1542,1566]" captionTargetBox="[152,1433,567,1514]" captionTargetId="figure-169@3.[151,1436,561,1519]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map of the type locality of troglomorphic Lagynochthonius species in Yunnan (China). A. L. retrorsus sp. nov. (Qinglong Cave); B. L. fengi sp. nov. (Unnamed Cave 2); C. L. xiaolinensis sp. nov. (Dayan Cave); D. L. xibaiensis sp. nov. (Hama Cave); E. L. spinulentus sp. nov. (Yingshan Cave); F. L. xinjiaoensis sp. nov. (Yueliang Cave); G. L. magnidentatus sp. nov. (Xiao Cave); H. L. bailongtanensis sp. nov. (Qianfo Cave); I. L. laoxueyanensis sp. nov. (Laoxueyan Cave); J. L. serratus sp. nov. (Unnamed Cave 3); K. L. yaowangguensis sp. nov. (Unnamed Cave 4); L. L. crassus sp. nov. (Unnamed Cave 1); MN. L. minimus sp. nov. (M: Yanfen Cave, N: Cizhu Cave)." figureDoi="http://doi.org/10.5281/zenodo.7251604" httpUri="https://zenodo.org/record/7251604/files/figure.png" pageId="55" pageNumber="56">Figs 1F</figureCitation>
,
<figureCitation id="9AF62A36FFF9A91CFABFFF7DFA7BFF7A" box="[1337,1424,151,177]" captionStart="FIGURE 38" captionStartId="51.[151,250,1671,1695]" captionTargetBox="[151,1436,181,1647]" captionTargetId="figure-19@51.[151,1436,181,1647]" captionTargetPageId="51" captionText="FIGURE 38. Yueliang Cave, type locality of Lagynochthonius xinjiaoensis sp. nov., A. Entrance; B. Inside the cave entrance; C. Ascent tunnel to the deepest part of the cave (red arrow); D. Area where L. xinjiaoensis sp. nov. specimen was collected; E. Live male of L. xinjiaoensis sp. nov., in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251724" httpUri="https://zenodo.org/record/7251724/files/figure.png" pageId="55" pageNumber="56">38AD</figureCitation>
), which is located near the top of a mountain and is about
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north of Xinjiao Village (Fuyuan County). This limestone cave has one small and oval entrance (about
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high and
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wide) and another large circular entrance directly above the cave. The total length of the cave is unknown. There is a certain vertical drop in the cave, which requires a
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descent to a deep pit and a
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ascent along a tunnel to reach the depth of the cave. The specimen was collected under a stone adjacent to a stalactite in the deep zone (temperature around 14°C, humidity is approximate 90%).
</paragraph>
</subSubSection>
</treatment>
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