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<document id="36CA27E1E63D6E048AC611979DDF36AB" ID-CLB-Dataset="172790" ID-DOI="10.11646/zootaxa.5198.1.1" ID-GBIF-Dataset="17babbe7-1645-47c8-ac5f-2299f1ad436c" ID-ISSN="1175-5326" ID-Zenodo-Dep="7251602" ID-ZooBank="019FE488-4263-4BC2-8606-446E599E226A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666769542164" checkinUser="plazi" docAuthor="Hou, Yanmeng, Gao, Zhizhong &amp; Zhang, Feng" docDate="2022" docId="8A6487A5FFD5A90BFF11FB88FD0DFA0A" docLanguage="en" docName="zootaxa.5198.1.1.pdf" docOrigin="Zootaxa 5198 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5198.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Lagynochthonius retrorsus Hou &amp; Gao &amp; Zhang 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="33" masterDocId="765DFFDDFFCEA92BFF86FFEAFFEBFFCB" masterDocTitle="Diversity of cave-dwelling pseudoscorpions from eastern Yunnan in China, with the description of eleven new species of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="65" masterPageNumber="1" pageNumber="28" updateTime="1701997268366" updateUser="plazi">
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<mods:title id="1872DD202C0FEDE3B990D5A45CCAA0F1">Diversity of cave-dwelling pseudoscorpions from eastern Yunnan in China, with the description of eleven new species of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:namePart id="32FC8D524BCAADBD7A4D74E94B74995D">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="7CFE135DA6A52E7418F2A2F0C8F77361">Gao, Zhizhong</mods:namePart>
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<mods:namePart id="0E1CE80F2F17A558207C567919285299">Zhang, Feng</mods:namePart>
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<mods:date id="F59BD86C423854BB287DD92B4B165CF5">2022</mods:date>
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<treatment id="8A6487A5FFD5A90BFF11FB88FD0DFA0A" ID-DOI="http://doi.org/10.5281/zenodo.7251614" ID-GBIF-Taxon="203411089" ID-Zenodo-Dep="7251614" LSID="urn:lsid:plazi:treatment:8A6487A5FFD5A90BFF11FB88FD0DFA0A" httpUri="http://treatment.plazi.org/id/8A6487A5FFD5A90BFF11FB88FD0DFA0A" lastPageId="32" lastPageNumber="33" pageId="27" pageNumber="28">
<subSubSection id="4AD76538FFD5A930FF11FB88FD18FBB7" box="[151,755,1122,1149]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="027236B3FFD5A930FF11FB88FD18FBB7" blockId="27.[151,755,1122,1185]" box="[151,755,1122,1149]" pageId="27" pageNumber="28">
<heading id="593A81DFFFD5A930FF11FB88FD18FBB7" bold="true" box="[151,755,1122,1149]" fontSize="11" level="2" pageId="27" pageNumber="28" reason="4">
<emphasis id="30B9EAA1FFD5A930FF11FB88FD18FBB7" bold="true" box="[151,755,1122,1149]" pageId="27" pageNumber="28">
<taxonomicName id="C5CD4D30FFD5A930FF11FB88FE34FBB6" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[151,479,1122,1149]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="retrorsus" status="sp. nov.">
<emphasis id="30B9EAA1FFD5A930FF11FB88FE34FBB6" bold="true" box="[151,479,1122,1149]" italics="true" pageId="27" pageNumber="28">Lagynochthonius retrorsus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2B8A57DAFFD5A930FE61FB89FDAEFBB6" box="[487,581,1123,1149]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
(ẁffiIJ伪oi)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="4AD76538FFD5A930FF11FB6DFEFEFB6A" box="[151,277,1159,1185]" pageId="27" pageNumber="28" type="description">
<paragraph id="027236B3FFD5A930FF11FB6DFEFEFB6A" blockId="27.[151,755,1122,1185]" box="[151,277,1159,1185]" pageId="27" pageNumber="28">
<figureCitation id="9AF62A36FFD5A930FF11FB6DFEFEFB6A" box="[151,277,1159,1185]" captionStart-0="FIGURE 19" captionStart-1="FIGURE20" captionStart-2="FIGURE 21" captionStart-3="FIGURE 22" captionStartId-0="28.[151,250,1430,1454]" captionStartId-1="29.[151,250,2005,2029]" captionStartId-2="30.[151,250,1894,1918]" captionStartId-3="31.[151,250,1849,1873]" captionTargetBox-0="[151,1436,273,1407]" captionTargetBox-1="[162,1416,1102,1971]" captionTargetBox-2="[151,1436,181,1870]" captionTargetBox-3="[191,1397,181,1825]" captionTargetId-0="figure-71@28.[151,1436,273,1407]" captionTargetId-1="figure-497@29.[151,1437,1101,1982]" captionTargetId-2="figure-20@30.[151,1436,181,1870]" captionTargetId-3="figure-19@31.[191,1397,181,1825]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionTargetPageId-2="30" captionTargetPageId-3="31" captionText-0="FIGURE 19. Qinglong Cave, type locality of Lagynochthonius retrorsus sp. nov., A. Entrance; B. Inside the cave entrance; C–D. Areas where L. retrorsus sp. nov. specimens were collected; E–F. Live individuals of L. retrorsus sp. nov. (E: male, F: female) in their natural environment." captionText-1="FIGURE20.Lagynochthoniusretrorsussp.nov.,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview)" captionText-2="FIGURE 21. Lagynochthonius retrorsus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." captionText-3="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi-0="http://doi.org/10.5281/zenodo.7251663" figureDoi-1="http://doi.org/10.5281/zenodo.7251665" figureDoi-2="http://doi.org/10.5281/zenodo.7251667" figureDoi-3="http://doi.org/10.5281/zenodo.7251671" httpUri-0="https://zenodo.org/record/7251663/files/figure.png" httpUri-1="https://zenodo.org/record/7251665/files/figure.png" httpUri-2="https://zenodo.org/record/7251667/files/figure.png" httpUri-3="https://zenodo.org/record/7251671/files/figure.png" pageId="27" pageNumber="28">Figs 19–22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFD5A930FF11FB22FC8DFAB9" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="027236B3FFD5A930FF11FB22FC8DFAB9" blockId="27.[151,1437,1224,2043]" pageId="27" pageNumber="28">
<emphasis id="30B9EAA1FFD5A930FF11FB22FEABFB29" bold="true" box="[151,320,1224,1250]" pageId="27" pageNumber="28">Type material.</emphasis>
<materialsCitation id="B2A53CEEFFD5A930FEC5FB22FACBFAE1" ID-GBIF-Occurrence="3946861310" collectingDate="2021-10-03" collectionCode="MHBU-HBUARA" collectorName="Zegang Feng &amp; Yanmeng Hou &amp; Lu Zhang &amp; Liu Fu" country="China" county="Daguan County" elevation="125" latitude="27.722004" location="Xingfu Village" longLatPrecision="1" longitude="103.9969" municipality="Tianxing Town" pageId="27" pageNumber="28" specimenCode="MHBU-HBUARA#2021-42101" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="DD768811FFD5A930FEC5FB22FE40FB29" box="[323,427,1224,1250]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
<specimenCount id="14CBFD3AFFD5A930FE2BFB23FE29FB28" box="[429,450,1225,1251]" pageId="27" pageNumber="28" type="male">♂</specimenCount>
(Ps.-
<specimenCode id="526B9EC8FFD5A930FE74FB22FC83FB29" box="[498,872,1224,1250]" collectionCode="MHBU-HBUARA" pageId="27" pageNumber="28">MHBU-HBUARA#2021-42101</specimenCode>
):
<collectingCountry id="7ADA7623FFD5A930FCF3FB22FC50FB29" box="[885,955,1224,1250]" name="China" pageId="27" pageNumber="28">China</collectingCountry>
,
<collectingRegion id="C009F851FFD5A930FC46FB23FB95FB29" box="[960,1150,1224,1251]" country="China" name="Yunnan" pageId="27" pageNumber="28">Yunnan Province</collectingRegion>
,
<collectingCounty id="EB134E3FFFD5A930FB03FB23FAD8FB29" box="[1157,1331,1224,1251]" pageId="27" pageNumber="28">Daguan County</collectingCounty>
,
<collectingMunicipality id="E216ACC9FFD5A930FABEFB22FF32FACC" pageId="27" pageNumber="28">Tianxing Town</collectingMunicipality>
,
<location id="07126068FFD5A930FF61FB06FE78FACD" LSID="urn:lsid:plazi:treatment:8A6487A5FFD5A90BFF11FB88FD0DFA0A:07126068FFD5A930FF61FB06FE78FACD" box="[231,403,1260,1286]" country="China" county="Daguan County" latitude="27.722004" longLatPrecision="1" longitude="103.9969" municipality="Tianxing Town" name="Xingfu Village" pageId="27" pageNumber="28" stateProvince="Yunnan">Xingfu Village</location>
,
<location id="07126068FFD5A930FE27FB06FDBBFACD" LSID="urn:lsid:plazi:treatment:8A6487A5FFD5A90BFF11FB88FD0DFA0A:07126068FFD5A930FE27FB06FDBBFACD" box="[417,592,1260,1286]" country="China" county="Daguan County" latitude="27.722004" longLatPrecision="1" longitude="103.9969" municipality="Tianxing Town" name="Qinglong Cave" pageId="27" pageNumber="28" stateProvince="Yunnan">Qinglong Cave</location>
, under stones within
<quantity id="C5359B56FFD5A930FCCEFB06FC22FACC" box="[840,969,1260,1287]" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="27" pageNumber="28" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">
<elevation id="89E0D180FFD5A930FCCEFB06FC22FACC" box="[840,969,1260,1287]" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="27" pageNumber="28" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">100–150 m</elevation>
</quantity>
from the cave entrance [
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,
<geoCoordinate id="67F95074FFD5A930FF11FAFBFEBBFAE7" box="[151,336,1297,1324]" degrees="103" direction="east" minutes="59" orientation="longitude" pageId="27" pageNumber="28" precision="1" seconds="48.86" value="103.9969">103°59′48.86″E</geoCoordinate>
],
<elevation id="89E0D180FFD5A930FEE5FAFAFE02FAE1" box="[355,489,1296,1323]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="27" pageNumber="28" unit="m" value="816.0">
<quantity id="C5359B56FFD5A930FEE5FAFAFE40FAE0" box="[355,427,1296,1323]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="27" pageNumber="28" unit="m" value="816.0">816 m</quantity>
a.s.l.
</elevation>
,
<date id="76731073FFD5A930FE71FAFAFD40FAE1" box="[503,683,1296,1322]" pageId="27" pageNumber="28" value="2021-10-03">
<collectingDate id="6637E99BFFD5A930FE71FAFAFD40FAE1" box="[503,683,1296,1322]" pageId="27" pageNumber="28" value="2021-10-03">3 October 2021</collectingDate>
</date>
,
<collectorName id="AF385365FFD5A930FD31FAFBFCA5FAE0" box="[695,846,1297,1323]" pageId="27" pageNumber="28">Zegang Feng</collectorName>
,
<collectorName id="AF385365FFD5A930FCDCFAFBFC14FAE0" box="[858,1023,1297,1323]" pageId="27" pageNumber="28">Yanmeng Hou</collectorName>
,
<collectorName id="AF385365FFD5A930FB8AFAFBFB90FAE1" box="[1036,1147,1296,1323]" pageId="27" pageNumber="28">Lu Zhang</collectorName>
&amp;
<collectorName id="AF385365FFD5A930FB27FAFAFB05FAE0" box="[1185,1262,1296,1323]" pageId="27" pageNumber="28">Liu Fu</collectorName>
leg.
</materialsCitation>
<materialsCitation id="B2A53CEEFFD5A930FAAEFAFBFC45FA85" ID-GBIF-Occurrence="3946861320" collectionCode="MHBU-HBUARA" country="China" pageId="27" pageNumber="28" specimenCode="MHBU-HBUARA#2021-42102" specimenCount="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<typeStatus id="DD768811FFD5A930FAAEFAFBFA72FAE0" box="[1320,1433,1297,1323]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="14CBFD3AFFD5A930FF11FADEFF2FFA84" box="[151,196,1332,1359]" pageId="27" pageNumber="28" type="male">2 ♂</specimenCount>
(Ps.-
<specimenCode id="526B9EC8FFD5A930FF7DFADEFD87FA85" box="[251,620,1332,1358]" collectionCode="MHBU-HBUARA" pageId="27" pageNumber="28">MHBU-HBUARA#2021-42102</specimenCode>
&amp; HBUARA#2021-42103)
</materialsCitation>
,
<materialsCitation id="B2A53CEEFFD5A930FC3AFADEFC88FAB9" ID-GBIF-Occurrence="3946861308" collectingDate="2021-10-03" collectionCode="MSWU-HBUARA" collectorName="Zegang Feng &amp; Yanmeng Hou &amp; Lu Zhang &amp; Liu Fu" country="China" county="Daguan County" elevation="125" latitude="27.722004" location="Xingfu Village" longLatPrecision="1" longitude="103.9969" municipality="Tianxing Town" pageId="27" pageNumber="28" specimenCode="MSWU-HBUARA#2021-42104" specimenCount="4" specimenCount-female="4" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="14CBFD3AFFD5A930FC3AFADEFC02FA84" box="[956,1001,1332,1359]" pageId="27" pageNumber="28" type="female">4 ♀</specimenCount>
(Ps.-
<specimenCode id="526B9EC8FFD5A930FB99FADEFA7EFA85" box="[1055,1429,1332,1358]" collectionCode="MSWU-HBUARA" pageId="27" pageNumber="28">MSWU-HBUARA#2021-42104</specimenCode>
- HBUARA#2021-42107), all with the same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFD5A930FF41FA96FDF3FA71" pageId="27" pageNumber="28" type="etymology">
<paragraph id="027236B3FFD5A930FF41FA96FDF3FA71" blockId="27.[151,1437,1224,2043]" pageId="27" pageNumber="28">
<emphasis id="30B9EAA1FFD5A930FF41FA96FEA5FA5D" bold="true" box="[199,334,1404,1430]" pageId="27" pageNumber="28">Etymology.</emphasis>
The specific name is derived from the Latin word “
<taxonomicName id="C5CD4D30FFD5A930FC1AFA97FBEAFA5D" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[924,1025,1405,1430]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="retrorsus">
<emphasis id="30B9EAA1FFD5A930FC1AFA97FBEAFA5D" box="[924,1025,1405,1430]" italics="true" pageId="27" pageNumber="28">retrorsus</emphasis>
</taxonomicName>
” (= retrorse) and refers to the shape of the movable chelal finger teeth.
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFD5A930FF41FA2EFEA0F97C" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="027236B3FFD5A930FF41FA2EFEA0F97C" blockId="27.[151,1437,1224,2043]" pageId="27" pageNumber="28">
<emphasis id="30B9EAA1FFD5A930FF41FA2EFE6FFA2A" bold="true" box="[199,388,1476,1505]" pageId="27" pageNumber="28">
Diagnosis (
<specimenCount id="14CBFD3AFFD5A930FECCFA2FFEB4FA2A" box="[330,351,1477,1505]" pageId="27" pageNumber="28" type="male">♂</specimenCount>
<specimenCount id="14CBFD3AFFD5A930FED9FA2FFE9FFA2A" box="[351,372,1477,1505]" pageId="27" pageNumber="28" type="female">♀</specimenCount>
).
</emphasis>
Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, pointed, triangular; posterior margin of carapace with 2 setae; tergites I–IV each with 2 setae. Pedipalps slender, femur 7.86–8.29 (
<specimenCount id="14CBFD3AFFD5A930FB48F9E6FB0EF9ED" box="[1230,1253,1548,1574]" pageId="27" pageNumber="28" type="male">♂</specimenCount>
), 7.73–8.20 (
<specimenCount id="14CBFD3AFFD5A930FAF9F9E6FA7AF9ED" box="[1407,1425,1548,1574]" pageId="27" pageNumber="28" type="female">♀</specimenCount>
) times longer than broad; chela 6.83–7.32 (
<specimenCount id="14CBFD3AFFD5A930FDF4F9DAFD62F981" box="[626,649,1584,1610]" pageId="27" pageNumber="28" type="male">♂</specimenCount>
), 7.04–7.13 (
<specimenCount id="14CBFD3AFFD5A930FCA7F9DAFCD8F981" box="[801,819,1584,1610]" pageId="27" pageNumber="28" type="female">♀</specimenCount>
) times longer than broad; only fixed chelal finger with intercalary teeth (rounded and obtuse) and with a modified accessory tooth (
<emphasis id="30B9EAA1FFD5A930FB86F9BEFBFDF9A5" box="[1024,1046,1620,1646]" italics="true" pageId="27" pageNumber="28">td</emphasis>
) on dorso-antiaxial face; movable chelal finger teeth markedly retrorse and contiguous; chemosensory setae (
<emphasis id="30B9EAA1FFD5A930FC68F993FBEEF959" box="[1006,1029,1657,1682]" italics="true" pageId="27" pageNumber="28">sc</emphasis>
) present on dorsum of chelal hand; sensilla present.
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFD5A90BFF41F92AFABDFC96" lastPageId="32" lastPageNumber="33" pageId="27" pageNumber="28" type="description">
<paragraph id="027236B3FFD5A930FF41F92AFBE4F911" blockId="27.[151,1437,1224,2043]" box="[199,1039,1728,1754]" pageId="27" pageNumber="28">
<emphasis id="30B9EAA1FFD5A930FF41F92AFEB2F911" bold="true" box="[199,345,1728,1754]" pageId="27" pageNumber="28">Description.</emphasis>
Males (
<typeStatus id="DD768811FFD5A930FE35F92AFDFCF911" box="[435,535,1728,1754]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
and
<typeStatus id="DD768811FFD5A930FDCBF92AFD51F911" box="[589,698,1728,1754]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="9AF62A36FFD5A930FD56F92AFCD3F911" box="[720,824,1728,1754]" captionStart="FIGURE 19" captionStartId="28.[151,250,1430,1454]" captionTargetBox="[151,1436,273,1407]" captionTargetId="figure-71@28.[151,1436,273,1407]" captionTargetPageId="28" captionText="FIGURE 19. Qinglong Cave, type locality of Lagynochthonius retrorsus sp. nov., A. Entrance; B. Inside the cave entrance; C–D. Areas where L. retrorsus sp. nov. specimens were collected; E–F. Live individuals of L. retrorsus sp. nov. (E: male, F: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251663" httpUri="https://zenodo.org/record/7251663/files/figure.png" pageId="27" pageNumber="28">Figs 19E</figureCitation>
,
<figureCitation id="9AF62A36FFD5A930FCC2F92AFC9CF911" box="[836,887,1728,1754]" captionStart="FIGURE20" captionStartId="29.[151,250,2005,2029]" captionTargetBox="[162,1416,1102,1971]" captionTargetId="figure-497@29.[151,1437,1101,1982]" captionTargetPageId="29" captionText="FIGURE20.Lagynochthoniusretrorsussp.nov.,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview)" figureDoi="http://doi.org/10.5281/zenodo.7251665" httpUri="https://zenodo.org/record/7251665/files/figure.png" pageId="27" pageNumber="28">20A</figureCitation>
,
<figureCitation id="9AF62A36FFD5A930FC04F92AFC3CF911" box="[898,983,1728,1754]" captionStart="FIGURE 21" captionStartId="30.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1870]" captionTargetId="figure-20@30.[151,1436,181,1870]" captionTargetPageId="30" captionText="FIGURE 21. Lagynochthonius retrorsus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251667" httpUri="https://zenodo.org/record/7251667/files/figure.png" pageId="27" pageNumber="28">21A–D</figureCitation>
,
<figureCitation id="9AF62A36FFD5A930FC65F92AFBE8F911" box="[995,1027,1728,1754]" captionStart="FIGURE 22" captionStartId="31.[151,250,1849,1873]" captionTargetBox="[191,1397,181,1825]" captionTargetId="figure-19@31.[191,1397,181,1825]" captionTargetPageId="31" captionText="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="27" pageNumber="28">22</figureCitation>
).
</paragraph>
<paragraph id="027236B3FFD5A930FF41F90EFEE0F8E9" blockId="27.[151,1437,1224,2043]" pageId="27" pageNumber="28">
Colour (
<figureCitation id="9AF62A36FFD5A930FEA3F90EFE66F935" box="[293,397,1764,1790]" captionStart="FIGURE 19" captionStartId="28.[151,250,1430,1454]" captionTargetBox="[151,1436,273,1407]" captionTargetId="figure-71@28.[151,1436,273,1407]" captionTargetPageId="28" captionText="FIGURE 19. Qinglong Cave, type locality of Lagynochthonius retrorsus sp. nov., A. Entrance; B. Inside the cave entrance; C–D. Areas where L. retrorsus sp. nov. specimens were collected; E–F. Live individuals of L. retrorsus sp. nov. (E: male, F: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251663" httpUri="https://zenodo.org/record/7251663/files/figure.png" pageId="27" pageNumber="28">Figs 19E</figureCitation>
,
<figureCitation id="9AF62A36FFD5A930FE1CF90EFE27F935" box="[410,460,1764,1790]" captionStart="FIGURE20" captionStartId="29.[151,250,2005,2029]" captionTargetBox="[162,1416,1102,1971]" captionTargetId="figure-497@29.[151,1437,1101,1982]" captionTargetPageId="29" captionText="FIGURE20.Lagynochthoniusretrorsussp.nov.,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview)" figureDoi="http://doi.org/10.5281/zenodo.7251665" httpUri="https://zenodo.org/record/7251665/files/figure.png" pageId="27" pageNumber="28">20A</figureCitation>
,
<figureCitation id="9AF62A36FFD5A930FE5FF90EFDDAF935" box="[473,561,1764,1790]" captionStart="FIGURE 21" captionStartId="30.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1870]" captionTargetId="figure-20@30.[151,1436,181,1870]" captionTargetPageId="30" captionText="FIGURE 21. Lagynochthonius retrorsus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251667" httpUri="https://zenodo.org/record/7251667/files/figure.png" pageId="27" pageNumber="28">21A–D</figureCitation>
): generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.
</paragraph>
<paragraph id="027236B3FFD5A937FF41F8C6FB89FF32" blockId="27.[151,1437,1224,2043]" lastBlockId="28.[151,1436,151,249]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
Cephalothorax (
<figureCitation id="9AF62A36FFD5A930FEF9F8C6FE06F88D" box="[383,493,1836,1862]" captionStart="FIGURE 21" captionStartId="30.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1870]" captionTargetId="figure-20@30.[151,1436,181,1870]" captionTargetPageId="30" captionText="FIGURE 21. Lagynochthonius retrorsus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251667" httpUri="https://zenodo.org/record/7251667/files/figure.png" pageId="27" pageNumber="28">Figs 21B</figureCitation>
,
<figureCitation id="9AF62A36FFD5A930FE7AF8C6FDD9F88D" box="[508,562,1836,1862]" captionStart="FIGURE 22" captionStartId="31.[151,250,1849,1873]" captionTargetBox="[191,1397,181,1825]" captionTargetId="figure-19@31.[191,1397,181,1825]" captionTargetPageId="31" captionText="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="27" pageNumber="28">22A</figureCitation>
): carapace 1.00–1.04 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3–4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/3 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 9–10 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
<figureCitation id="9AF62A36FFD2A937FE40FF35FDC5FF32" box="[454,558,223,249]" captionStart="FIGURE 22" captionStartId="31.[151,250,1849,1873]" captionTargetBox="[191,1397,181,1825]" captionTargetId="figure-19@31.[191,1397,181,1825]" captionTargetPageId="31" captionText="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="28" pageNumber="29">Fig. 22C</figureCitation>
); intercoxal tubercle absent; without sub-oral seta.
</paragraph>
<caption id="56B2663BFFD2A937FF11FA7CFDE5FA3D" ID-DOI="http://doi.org/10.5281/zenodo.7251663" ID-Zenodo-Dep="7251663" httpUri="https://zenodo.org/record/7251663/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,250,1430,1454]" targetBox="[151,1436,273,1407]" targetPageId="28">
<paragraph id="027236B3FFD2A937FF11FA7CFDE5FA3D" blockId="28.[151,1437,1430,1526]" pageId="28" pageNumber="29">
<emphasis id="30B9EAA1FFD2A937FF11FA7CFEC9FA65" bold="true" box="[151,290,1430,1454]" pageId="28" pageNumber="29">FIGURE 19.</emphasis>
Qinglong Cave, type locality of
<taxonomicName id="C5CD4D30FFD2A937FDFFFA7CFC60FA65" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[633,907,1430,1454]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="retrorsus" status="sp. nov.">
<emphasis id="30B9EAA1FFD2A937FDFFFA7CFC60FA65" box="[633,907,1430,1454]" italics="true" pageId="28" pageNumber="29">Lagynochthonius retrorsus</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFD2A937FC15FA7CFC0EFA65" bold="true" box="[915,997,1430,1454]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="2B8A57DAFFD2A937FC15FA7CFC0EFA65" box="[915,997,1430,1454]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Entrance; B. Inside the cave entrance; C–D. Areas where
<taxonomicName id="C5CD4D30FFD2A937FEDAFA51FE38FA19" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[348,467,1467,1490]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="retrorsus" status="sp. nov.">
<emphasis id="30B9EAA1FFD2A937FEDAFA51FE38FA19" box="[348,467,1467,1490]" italics="true" pageId="28" pageNumber="29">L. retrorsus</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFD2A937FE5CFA50FDC7FA19" bold="true" box="[474,556,1466,1490]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="2B8A57DAFFD2A937FE5CFA50FDC7FA19" box="[474,556,1466,1490]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were collected; E–F. Live individuals of
<taxonomicName id="C5CD4D30FFD2A937FBCFFA51FB2BFA19" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[1097,1216,1467,1490]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="retrorsus" status="sp. nov.">
<emphasis id="30B9EAA1FFD2A937FBCFFA51FB2BFA19" box="[1097,1216,1467,1490]" italics="true" pageId="28" pageNumber="29">L. retrorsus</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFD2A937FB4EFA50FAF2FA19" bold="true" box="[1224,1305,1466,1490]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="2B8A57DAFFD2A937FB4EFA50FAF2FA19" box="[1224,1305,1466,1490]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(E: male, F: female) in their natural environment.
</paragraph>
</caption>
<paragraph id="027236B3FFD2A937FF41F9CEFD87F8F0" blockId="28.[151,1437,1572,2031]" pageId="28" pageNumber="29">
Chelicera (
<figureCitation id="9AF62A36FFD2A937FEC4F9CEFE47F9F4" box="[322,428,1572,1599]" captionStart="FIGURE 21" captionStartId="30.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1870]" captionTargetId="figure-20@30.[151,1436,181,1870]" captionTargetPageId="30" captionText="FIGURE 21. Lagynochthonius retrorsus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251667" httpUri="https://zenodo.org/record/7251667/files/figure.png" pageId="28" pageNumber="29">Figs 21C</figureCitation>
,
<figureCitation id="9AF62A36FFD2A937FE3FF9CFFE07F9F4" box="[441,492,1573,1599]" captionStart="FIGURE 22" captionStartId="31.[151,250,1849,1873]" captionTargetBox="[191,1397,181,1825]" captionTargetId="figure-19@31.[191,1397,181,1825]" captionTargetPageId="31" captionText="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="28" pageNumber="29">22B</figureCitation>
): large, about as long as carapace, 2.35–2.37 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 17–19 teeth, distal one largest; movable finger with 15–17 retrorse contiguous small teeth; galea completely vestigial (
<figureCitation id="9AF62A36FFD2A937FBF7F95EFB33F904" box="[1137,1240,1716,1743]" captionStart="FIGURE 22" captionStartId="31.[151,250,1849,1873]" captionTargetBox="[191,1397,181,1825]" captionTargetId="figure-19@31.[191,1397,181,1825]" captionTargetPageId="31" captionText="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="28" pageNumber="29">Fig. 22B</figureCitation>
). Serrula exterior with 22–24 and serrula interior with 13–14 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
<figureCitation id="9AF62A36FFD2A937FE70F8CAFD8BF8F0" box="[502,608,1824,1851]" captionStart="FIGURE 22" captionStartId="31.[151,250,1849,1873]" captionTargetBox="[191,1397,181,1825]" captionTargetId="figure-19@31.[191,1397,181,1825]" captionTargetPageId="31" captionText="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="28" pageNumber="29">Fig. 22D</figureCitation>
).
</paragraph>
<paragraph id="027236B3FFD2A936FF41F8AEFAE1FD4E" blockId="28.[151,1437,1572,2031]" lastBlockId="29.[151,1437,151,1077]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
Pedipalp (
<figureCitation id="9AF62A36FFD2A937FEB1F8AEFE49F894" box="[311,418,1860,1887]" captionStart="FIGURE 21" captionStartId="30.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1870]" captionTargetId="figure-20@30.[151,1436,181,1870]" captionTargetPageId="30" captionText="FIGURE 21. Lagynochthonius retrorsus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251667" httpUri="https://zenodo.org/record/7251667/files/figure.png" pageId="28" pageNumber="29">Figs 21A</figureCitation>
,
<figureCitation id="9AF62A36FFD2A937FE2DF8AFFDEBF894" box="[427,512,1861,1887]" captionStart="FIGURE 22" captionStartId="31.[151,250,1849,1873]" captionTargetBox="[191,1397,181,1825]" captionTargetId="figure-19@31.[191,1397,181,1825]" captionTargetPageId="31" captionText="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="28" pageNumber="29">22E–G</figureCitation>
): long and slender, trochanter 1.71–1.87, femur 7.86–8.29, patella 2.35, chela 6.83– 7.32, hand 2.83–2.91 times longer than broad; femur 2.75–2.90 times longer than patella; movable chelal finger 1.43–1.50 times longer than hand and 0.59–0.60 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (
<figureCitation id="9AF62A36FFD2A937FDA5F85AFD63F800" box="[547,648,1968,1995]" captionStart="FIGURE 22" captionStartId="31.[151,250,1849,1873]" captionTargetBox="[191,1397,181,1825]" captionTargetId="figure-19@31.[191,1397,181,1825]" captionTargetPageId="31" captionText="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="28" pageNumber="29">Fig. 22E</figureCitation>
). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria,
<emphasis id="30B9EAA1FFD3A936FB4AFF7DFB09FF7A" box="[1228,1250,151,177]" italics="true" pageId="29" pageNumber="30">ib</emphasis>
and
<emphasis id="30B9EAA1FFD3A936FA9AFF7DFAD6FF7A" box="[1308,1341,151,177]" italics="true" pageId="29" pageNumber="30">isb</emphasis>
situated close together, submedially on dorsum of chelal hand;
<emphasis id="30B9EAA1FFD3A936FD70FF51FCFBFF1E" box="[758,784,187,213]" italics="true" pageId="29" pageNumber="30">eb</emphasis>
,
<emphasis id="30B9EAA1FFD3A936FC98FF51FCA8FF1E" box="[798,835,187,213]" italics="true" pageId="29" pageNumber="30">esb</emphasis>
and
<emphasis id="30B9EAA1FFD3A936FCFEFF51FC79FF1F" box="[888,914,187,212]" italics="true" pageId="29" pageNumber="30">ist</emphasis>
forming a straight oblique row at base of fixed chelal finger;
<emphasis id="30B9EAA1FFD3A936FEB2FF35FEAFFF33" box="[308,324,223,248]" italics="true" pageId="29" pageNumber="30">it</emphasis>
slightly distal to
<emphasis id="30B9EAA1FFD3A936FD8FFF35FDC3FF33" box="[521,552,223,248]" italics="true" pageId="29" pageNumber="30">est</emphasis>
, situated subdistally;
<emphasis id="30B9EAA1FFD3A936FC98FF35FCD9FF33" box="[798,818,223,248]" italics="true" pageId="29" pageNumber="30">et</emphasis>
slightly near to tip of fixed chelal finger, very close to chelal teeth;
<emphasis id="30B9EAA1FFD3A936FEAFFEE9FEA8FED6" box="[297,323,259,285]" italics="true" pageId="29" pageNumber="30">dx</emphasis>
situated distal to
<emphasis id="30B9EAA1FFD3A936FD88FEE9FDC9FED7" box="[526,546,259,284]" italics="true" pageId="29" pageNumber="30">et</emphasis>
;
<emphasis id="30B9EAA1FFD3A936FDB5FEE9FDA7FED6" box="[563,588,259,285]" italics="true" pageId="29" pageNumber="30">sb</emphasis>
closer to
<emphasis id="30B9EAA1FFD3A936FD3BFEE9FD3BFED7" box="[701,720,259,284]" italics="true" pageId="29" pageNumber="30">st</emphasis>
than to
<emphasis id="30B9EAA1FFD3A936FCA9FEE9FCD6FED6" box="[815,829,259,285]" italics="true" pageId="29" pageNumber="30">b</emphasis>
;
<emphasis id="30B9EAA1FFD3A936FCCBFEE9FCB0FED6" box="[845,859,259,285]" italics="true" pageId="29" pageNumber="30">b</emphasis>
and
<emphasis id="30B9EAA1FFD3A936FC12FEE9FC77FED7" box="[916,924,259,284]" italics="true" pageId="29" pageNumber="30">t</emphasis>
situated subdistally,
<emphasis id="30B9EAA1FFD3A936FB0DFEE9FB78FED7" box="[1163,1171,259,284]" italics="true" pageId="29" pageNumber="30">t</emphasis>
situated distal to
<emphasis id="30B9EAA1FFD3A936FADBFEE9FA86FED7" box="[1373,1389,259,284]" italics="true" pageId="29" pageNumber="30">it</emphasis>
;
<emphasis id="30B9EAA1FFD3A936FAFBFEE9FA77FED7" box="[1405,1436,259,284]" italics="true" pageId="29" pageNumber="30">est</emphasis>
situated distal to
<emphasis id="30B9EAA1FFD3A936FED2FECDFE89FE8A" box="[340,354,295,321]" italics="true" pageId="29" pageNumber="30">b</emphasis>
(
<figureCitation id="9AF62A36FFD3A936FEF7FECDFE3EFE8A" box="[369,469,295,321]" captionStart="FIGURE 22" captionStartId="31.[151,250,1849,1873]" captionTargetBox="[191,1397,181,1825]" captionTargetId="figure-19@31.[191,1397,181,1825]" captionTargetPageId="31" captionText="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="29" pageNumber="30">Fig. 22F</figureCitation>
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to
<emphasis id="30B9EAA1FFD3A936FAF7FECDFA60FE8B" box="[1393,1419,295,320]" italics="true" pageId="29" pageNumber="30">ist</emphasis>
). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 28–30 macrodenticles, slightly retrorse and pointed, plus 12–13 rounded and obtuse intercalary microdenticles, and a modified accessory tooth on dorso-antiaxial face (
<emphasis id="30B9EAA1FFD3A936FD47FE5DFD3CFE1A" box="[705,727,439,465]" italics="true" pageId="29" pageNumber="30">td</emphasis>
, slightly distal to
<emphasis id="30B9EAA1FFD3A936FC1AFE5DFC5DFE1A" box="[924,950,439,465]" italics="true" pageId="29" pageNumber="30">dx</emphasis>
),
<quantity id="C5359B56FFD3A936FC4AFE5DFBC5FE1A" box="[972,1070,439,465]" metricMagnitude="0" metricUnit="m" metricValue="1.0795" metricValueMax="1.0922" metricValueMin="1.0668" pageId="29" pageNumber="30" unit="in" value="42.5" valueMax="43.0" valueMin="42.0">42–43 in</quantity>
total; movable chelal finger with 29–30 macrodenticles (slightly smaller than teeth on fixed chelal finger), markedly retrorse, pointed and contiguous, plus 6–8 vestigial, rounded and contiguous basal teeth,
<quantity id="C5359B56FFD3A936FC84FE15FC8EFDD2" box="[770,869,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="9.271" metricValueMax="9.398" metricValueMin="9.144" pageId="29" pageNumber="30" unit="in" value="36.5" valueMax="37.0" valueMin="36.0">36–37 in</quantity>
total; two tubercles present between first and third teeth (
<figureCitation id="9AF62A36FFD3A936FF5AFDC9FEABFDF6" box="[220,320,547,573]" captionStart="FIGURE 22" captionStartId="31.[151,250,1849,1873]" captionTargetBox="[191,1397,181,1825]" captionTargetId="figure-19@31.[191,1397,181,1825]" captionTargetPageId="31" captionText="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="29" pageNumber="30">Fig. 22F</figureCitation>
). Fixed chelal finger with sensilla
<emphasis id="30B9EAA1FFD3A936FD3BFDC9FD38FDF6" box="[701,723,547,573]" italics="true" pageId="29" pageNumber="30">af</emphasis>
<subScript id="9E4934F6FFD3A936FD55FDDFFD00FD88" attach="left" box="[723,747,565,579]" fontSize="6" pageId="29" pageNumber="30">1–2</subScript>
close together, near tip; movable chelal finger with 4 sensilla:
<emphasis id="30B9EAA1FFD3A936FF11FDADFF52FDAB" box="[151,185,583,608]" italics="true" pageId="29" pageNumber="30">am</emphasis>
<subScript id="9E4934F6FFD3A936FF3FFDB3FF39FDAC" attach="left" box="[185,210,601,615]" fontSize="6" pageId="29" pageNumber="30">1–2</subScript>
near tip,
<emphasis id="30B9EAA1FFD3A936FEBCFDADFEA3FDAB" box="[314,328,583,608]" italics="true" pageId="29" pageNumber="30">p</emphasis>
<subScript id="9E4934F6FFD3A936FECEFDB3FEBBFDAC" attach="left" box="[328,336,601,615]" fontSize="6" pageId="29" pageNumber="30">2</subScript>
slightly distad of
<emphasis id="30B9EAA1FFD3A936FD91FDADFDDBFDAA" box="[535,560,583,609]" italics="true" pageId="29" pageNumber="30">sb</emphasis>
,
<emphasis id="30B9EAA1FFD3A936FDB8FDADFDA7FDAB" box="[574,588,583,608]" italics="true" pageId="29" pageNumber="30">p</emphasis>
<subScript id="9E4934F6FFD3A936FDCAFDB3FDBFFDAC" attach="left" box="[588,596,601,615]" fontSize="6" pageId="29" pageNumber="30">1</subScript>
distad of
<emphasis id="30B9EAA1FFD3A936FD47FDADFD24FDAB" box="[705,719,583,608]" italics="true" pageId="29" pageNumber="30">p</emphasis>
<subScript id="9E4934F6FFD3A936FD49FDB3FD3CFDAC" attach="left" box="[719,727,601,615]" fontSize="6" pageId="29" pageNumber="30">2</subScript>
and very close to chelal teeth (
<figureCitation id="9AF62A36FFD3A936FBA9FDADFB79FDAA" box="[1071,1170,583,609]" captionStart="FIGURE 22" captionStartId="31.[151,250,1849,1873]" captionTargetBox="[191,1397,181,1825]" captionTargetId="figure-19@31.[191,1397,181,1825]" captionTargetPageId="31" captionText="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="29" pageNumber="30">Fig. 22F</figureCitation>
). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
<figureCitation id="9AF62A36FFD3A936FB13FD81FB15FD4E" box="[1173,1278,619,645]" captionStart="FIGURE 22" captionStartId="31.[151,250,1849,1873]" captionTargetBox="[191,1397,181,1825]" captionTargetId="figure-19@31.[191,1397,181,1825]" captionTargetPageId="31" captionText="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="29" pageNumber="30">Fig. 22G</figureCitation>
).
</paragraph>
<paragraph id="027236B3FFD3A936FF41FD65FF3EFCF2" blockId="29.[151,1437,151,1077]" pageId="29" pageNumber="30">
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2: 2: 2: 4: 4: 4: 4–5: 5: 3–4: T2T: 0, tergites VIII–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 12–14: 11–12: 7–8: 8: 8: 9: 9: 7: 0: 2. Anterior genital operculum with 10 setae, genital opening slit-like, with 11–13 marginal setae on each side,
<quantity id="C5359B56FFD3A936FB3AFD11FAC9FCDE" box="[1212,1314,763,789]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" metricValueMax="9.144" metricValueMin="8.636" pageId="29" pageNumber="30" unit="in" value="35.0" valueMax="36.0" valueMin="34.0">34–36 in</quantity>
total (
<figureCitation id="9AF62A36FFD3A936FAE9FD11FF23FCF2" captionStart="FIGURE 21" captionStartId="30.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1870]" captionTargetId="figure-20@30.[151,1436,181,1870]" captionTargetPageId="30" captionText="FIGURE 21. Lagynochthonius retrorsus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251667" httpUri="https://zenodo.org/record/7251667/files/figure.png" pageId="29" pageNumber="30">Fig. 21E</figureCitation>
).
</paragraph>
<paragraph id="027236B3FFD3A936FF41FCA9FAF2FBFE" blockId="29.[151,1437,151,1077]" pageId="29" pageNumber="30">
Legs (
<figureCitation id="9AF62A36FFD3A936FE94FCA9FE71FC96" box="[274,410,835,861]" captionStart="FIGURE 22" captionStartId="31.[151,250,1849,1873]" captionTargetBox="[191,1397,181,1825]" captionTargetId="figure-19@31.[191,1397,181,1825]" captionTargetPageId="31" captionText="FIGURE 22. Lagynochthonius retrorsus sp. nov., holotype male:A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D)." figureDoi="http://doi.org/10.5281/zenodo.7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="29" pageNumber="30">Fig. 22H, I</figureCitation>
): generally typical, long and slender. Fine granulation present on anterodorsal faces of trochanter IV, femur I, IV and patella IV. Femur of leg I 1.90–2.06 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.24–2.32 times longer than tibia. Femoropatella of leg IV 3.74–3.75 times longer than deep; tibia 6.00–6.20 times longer than deep; with basal tactile setae on both tarsal segments: basitarsus 3.63 times longer than deep (TS= 0.31–0.34), telotarsus 12.50–14.00 times longer than deep and 2.41–2.59 times longer than basitarsus (TS= 0.31–0.39). Setae of leg I (trochanter to tibia) 3–4: 10–12: 10–12: 14–16, setae of leg IV (trochanter to basitarsus) 2–3: 3: 6: 9–11: 9–11. Arolium slightly shorter than the claws, not divided; claws simple.
</paragraph>
<caption id="56B2663BFFD3A936FF11F83FFA77F826" ID-DOI="http://doi.org/10.5281/zenodo.7251665" ID-Zenodo-Dep="7251665" box="[151,1436,2005,2030]" httpUri="https://zenodo.org/record/7251665/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,2005,2029]" targetBox="[162,1416,1102,1971]" targetPageId="29">
<paragraph id="027236B3FFD3A936FF11F83FFA77F826" blockId="29.[151,1436,2005,2030]" box="[151,1436,2005,2030]" pageId="29" pageNumber="30">
<emphasis id="30B9EAA1FFD3A936FF11F83FFEF7F826" bold="true" box="[151,284,2005,2029]" pageId="29" pageNumber="30">FIGURE20.</emphasis>
<taxonomicName id="C5CD4D30FFD3A936FE98F83FFDC1F826" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[286,554,2005,2029]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="retrorsus" status="sp. nov.">
<emphasis id="30B9EAA1FFD3A936FE98F83FFDC1F826" box="[286,554,2005,2029]" italics="true" pageId="29" pageNumber="30">Lagynochthonius retrorsus</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFD3A936FDAAF83FFD93F826" bold="true" box="[556,632,2005,2029]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="2B8A57DAFFD3A936FDAAF83FFD93F826" box="[556,632,2005,2029]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,A.Holotype male,habitus (dorsal view); B.Paratype female,habitus (dorsal view)
</paragraph>
</caption>
<caption id="56B2663BFFD0A935FF11F88CFF33F80D" ID-DOI="http://doi.org/10.5281/zenodo.7251667" ID-Zenodo-Dep="7251667" httpUri="https://zenodo.org/record/7251667/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,250,1894,1918]" targetBox="[151,1436,181,1870]" targetPageId="30">
<paragraph id="027236B3FFD0A935FF11F88CFF33F80D" blockId="30.[151,1436,1894,1990]" pageId="30" pageNumber="31">
<emphasis id="30B9EAA1FFD0A935FF11F88CFECAF8B5" bold="true" box="[151,289,1894,1918]" pageId="30" pageNumber="31">FIGURE 21.</emphasis>
<taxonomicName id="C5CD4D30FFD0A935FEAEF88CFDD2F8B5" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[296,569,1894,1918]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="retrorsus" status="sp. nov.">
<emphasis id="30B9EAA1FFD0A935FEAEF88CFDD2F8B5" box="[296,569,1894,1918]" italics="true" pageId="30" pageNumber="31">Lagynochthonius retrorsus</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFD0A935FDC6F88CFD79F8B5" bold="true" box="[576,658,1894,1918]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="2B8A57DAFFD0A935FDC6F88CFD79F8B5" box="[576,658,1894,1918]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view).
</paragraph>
</caption>
<caption id="56B2663BFFD1A934FF11F8D3FAA1F82A" ID-DOI="http://doi.org/10.5281/zenodo.7251671" ID-Zenodo-Dep="7251671" httpUri="https://zenodo.org/record/7251671/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,250,1849,1873]" targetBox="[191,1397,181,1825]" targetPageId="31">
<paragraph id="027236B3FFD1A934FF11F8D3FAA1F82A" blockId="31.[151,1437,1849,2018]" pageId="31" pageNumber="32">
<emphasis id="30B9EAA1FFD1A934FF11F8D3FEF4F89A" bold="true" box="[151,287,1849,1873]" pageId="31" pageNumber="32">FIGURE 22.</emphasis>
<taxonomicName id="C5CD4D30FFD1A934FEA2F8D3FDD8F89A" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[292,563,1849,1873]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="retrorsus" status="sp. nov.">
<emphasis id="30B9EAA1FFD1A934FEA2F8D3FDD8F89A" box="[292,563,1849,1873]" italics="true" pageId="31" pageNumber="32">Lagynochthonius retrorsus</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFD1A934FDBFF8D3FD63F89A" bold="true" box="[569,648,1849,1873]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="2B8A57DAFFD1A934FDBFF8D3FD63F89A" box="[569,648,1849,1873]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male: A. Carapace (dorsal view), with a detail of anterior margin; B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Rallum; E. Left pedipalp (minus chela, dorsal view); F. Left chela (lateral view), with details of teeth and with trichobothrial pattern, finger tips, modified tooth and sensilla (abbreviations explained in Materials and methods); G. Left chela (dorsal view), with details of chemosensory setae and modified tooth; H. Leg I (lateral view); I. Leg IV (lateral view). Scale bars: 0.25 mm (A–B, E–I); 0.10 mm (C–D).
</paragraph>
</caption>
<paragraph id="027236B3FFEEA90BFF41FF7DFAA2FE8A" blockId="32.[151,1437,151,1473]" pageId="32" pageNumber="33">
<materialsCitation id="B2A53CEEFFEEA90BFF41FF7DFAAEFE8A" ID-GBIF-Occurrence="3946861301" collectionCode="T, IV" pageId="32" pageNumber="33" specimenCode="T2" specimenCount="1" typeStatus="paratype">
Adult females (
<typeStatus id="DD768811FFEEA90BFEF4FF7DFE0AFF7A" box="[370,481,151,177]" pageId="32" pageNumber="33" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="9AF62A36FFEEA90BFE70FF7DFDB1FF7A" box="[502,602,151,177]" captionStart="FIGURE 19" captionStartId="28.[151,250,1430,1454]" captionTargetBox="[151,1436,273,1407]" captionTargetId="figure-71@28.[151,1436,273,1407]" captionTargetPageId="28" captionText="FIGURE 19. Qinglong Cave, type locality of Lagynochthonius retrorsus sp. nov., A. Entrance; B. Inside the cave entrance; C–D. Areas where L. retrorsus sp. nov. specimens were collected; E–F. Live individuals of L. retrorsus sp. nov. (E: male, F: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251663" httpUri="https://zenodo.org/record/7251663/files/figure.png" pageId="32" pageNumber="33">Figs 19F</figureCitation>
,
<figureCitation id="9AF62A36FFEEA90BFDE3FF7DFD7CFF7A" box="[613,663,151,177]" captionStart="FIGURE20" captionStartId="29.[151,250,2005,2029]" captionTargetBox="[162,1416,1102,1971]" captionTargetId="figure-497@29.[151,1437,1101,1982]" captionTargetPageId="29" captionText="FIGURE20.Lagynochthoniusretrorsussp.nov.,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview)" figureDoi="http://doi.org/10.5281/zenodo.7251665" httpUri="https://zenodo.org/record/7251665/files/figure.png" pageId="32" pageNumber="33">20B</figureCitation>
,
<figureCitation id="9AF62A36FFEEA90BFD27FF7DFD39FF7A" box="[673,722,151,177]" captionStart="FIGURE 21" captionStartId="30.[151,250,1894,1918]" captionTargetBox="[151,1436,181,1870]" captionTargetId="figure-20@30.[151,1436,181,1870]" captionTargetPageId="30" captionText="FIGURE 21. Lagynochthonius retrorsus sp. nov., holotype male (A–D), paratype female (E): A. Left chela (lateral view); B. Carapace (dorsal view); C. Left chelicera (dorsal view); D. Male genital area (ventral view); E. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7251667" httpUri="https://zenodo.org/record/7251667/files/figure.png" pageId="32" pageNumber="33">21E</figureCitation>
). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy I–XII: 2: 2: 2: 2: 4: 4: 4: 5: 5: 4:
<specimenCode id="526B9EC8FFEEA90BFC22FF51FC2FFF1E" box="[932,964,187,213]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="32" pageNumber="33">T2</specimenCode>
<collectionCode id="64DCAE76FFEEA90BFC43FF51FC3CFF1E" box="[965,983,187,213]" name="Tavera, Department of Geology and Geophysics" pageId="32" pageNumber="33">T</collectionCode>
: 0; sternal chaetotaxy
<collectionCode id="64DCAE76FFEEA90BFB54FF51FB1BFF1E" box="[1234,1264,187,213]" pageId="32" pageNumber="33">IV</collectionCode>
–XII: 11: 7–10: 10: 9: 9: 9: 8: 0: 2; anterior genital operculum with 10 setae, posterior margin with 11 marginal setae,
<quantity id="C5359B56FFEEA90BFB7AFF35FADFFF32" box="[1276,1332,223,249]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="32" pageNumber="33" unit="in" value="22.0">22 in</quantity>
total; leg
<collectionCode id="64DCAE76FFEEA90BFF11FEE9FF5FFED6" box="[151,180,259,285]" pageId="32" pageNumber="33">IV</collectionCode>
with a long tactile seta on both tarsal segments: basitarsus 3.50–3.63 times longer than deep (TS= 0.34–0.36), telotarsus 12.00–13.00 times longer than deep and 2.57–2.69 times longer than basitarsus (TS= 0.36–0.38)
</materialsCitation>
.
</paragraph>
<paragraph id="027236B3FFEEA90BFF41FEA1FB8CFDAA" blockId="32.[151,1437,151,1473]" pageId="32" pageNumber="33">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.94–2.04. Pedipalps: trochanter 0.28–0.29/0.15–0.17 (1.71–1.87), femur 1.10–1.16/0.14 (7.86–8.29), patella 0.40/0.17 (2.35), chela 1.57–1.61/0.22–0.23 (6.83–7.32), hand 0.64–0.65/0.22–0.23 (2.83–2.91), movable chelal finger length 0.93–0.96. Chelicera 0.61–0.64/0.26–0.27 (2.35–2.37), movable finger length 0.33–0.34. Carapace 0.57/0.55–0.57 (1.00–1.04). Leg I: trochanter 0.17–0.20/0.11–0.14 (1.43–1.55), femur 0.59–0.66/0.08 (7.38–8.25), patella 0.31–0.32/0.07 (4.43–4.57), tibia 0.28–0.29/0.05–0.06 (4.67–5.80), tarsus 0.65/0.06 (10.83). Leg IV: trochanter 0.25–0.26/0.15–0.16 (1.56–1.73), femoropatella 0.86–0.90/0.23–0.24 (3.74–3.75), tibia 0.60–0.62/0.10 (6.00–6.20), basitarsus 0.29/0.08 (3.63), telotarsus 0.70–0.75/0.05–0.06 (12.50–14.00).</paragraph>
<paragraph id="027236B3FFEEA90BFF41FD81FABDFC96" blockId="32.[151,1437,151,1473]" pageId="32" pageNumber="33">Females: body length 2.04–2.12. Pedipalps: trochanter 0.28/0.15–0.17 (1.65–1.87), femur 1.16–1.23/0.15 (7.73– 8.20), patella 0.42/0.18–0.19 (2.21–2.33), chela 1.64–1.69/0.23–0.24 (7.04–7.13), hand 0.69/0.23–0.24 (2.88–3.00), movable chelal finger length 0.99–1.01. Chelicera 0.65–0.69/0.27–0.29 (2.38–2.41), movable finger length 0.35– 0.38. Carapace 0.57–0.59/0.57–0.60 (0.98–1.00). Leg I: trochanter 0.19/0.12–0.14 (1.36–1.58), femur 0.66/0.09 (7.33), patella 0.29–0.32/0.07 (4.14–4.57), tibia 0.27–0.31/0.06 (4.50–5.17), tarsus 0.61–0.69/0.06 (10.17–11.50). Leg IV: trochanter 0.25–0.27/0.14–0.17 (1.59–1.79), femoropatella 0.87–0.92/0.22–0.24 (3.83–3.95), tibia 0.60– 0.61/0.09–0.10 (6.10–6.67), basitarsus 0.28–0.29/0.08 (3.50–3.63), telotarsus 0.72–0.78/0.06 (12.00–13.00).</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFEEA90BFF41FC8DFB22FB22" pageId="32" pageNumber="33" type="discussion">
<paragraph id="027236B3FFEEA90BFF41FC8DFCA2FBDA" blockId="32.[151,1437,151,1473]" pageId="32" pageNumber="33">
<emphasis id="30B9EAA1FFEEA90BFF41FC8DFED0FC4A" bold="true" box="[199,315,871,897]" pageId="32" pageNumber="33">Remarks.</emphasis>
Based on the key to the Chinese species of
<taxonomicName id="C5CD4D30FFEEA90BFCA8FC8DFC04FC4A" authorityName="Beier" authorityYear="1951" box="[814,1007,871,897]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="30B9EAA1FFEEA90BFCA8FC8DFC04FC4A" box="[814,1007,871,897]" italics="true" pageId="32" pageNumber="33">Lagynochthonius</emphasis>
</taxonomicName>
below,
<taxonomicName id="C5CD4D30FFEEA90BFBCEFC8DFB20FC4B" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[1096,1227,871,896]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="retrorsus" status="sp. nov.">
<emphasis id="30B9EAA1FFEEA90BFBCEFC8DFB20FC4B" box="[1096,1227,871,896]" italics="true" pageId="32" pageNumber="33">L. retrorsus</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFEEA90BFB54FC8DFAC7FC4A" bold="true" box="[1234,1324,871,897]" pageId="32" pageNumber="33">
<taxonomicNameLabel id="2B8A57DAFFEEA90BFB54FC8DFAC7FC4A" box="[1234,1324,871,897]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="C5CD4D30FFEEA90BFF30FC61FEBEFC6E" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[182,341,907,933]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="spinulentus" status="sp. nov.">
<emphasis id="30B9EAA1FFEEA90BFF30FC61FEBEFC6E" box="[182,341,907,933]" italics="true" pageId="32" pageNumber="33">L. spinulentus</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFEEA90BFEDBFC61FE52FC6E" bold="true" box="[349,441,907,933]" pageId="32" pageNumber="33">
<taxonomicNameLabel id="2B8A57DAFFEEA90BFEDBFC61FE52FC6E" box="[349,441,907,933]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but differs by a shorter chela (e.g. chela length 1.57–1.61
<emphasis id="30B9EAA1FFEEA90BFBE7FC61FB94FC6F" box="[1121,1151,907,932]" italics="true" pageId="32" pageNumber="33">vs.</emphasis>
<quantity id="C5359B56FFEEA90BFB0EFC61FAC5FC6E" box="[1160,1326,907,933]" metricMagnitude="-3" metricUnit="m" metricValue="2.115" metricValueMax="2.16" metricValueMin="2.07" pageId="32" pageNumber="33" unit="mm" value="2.115" valueMax="2.16" valueMin="2.07">2.07–2.16 mm</quantity>
, movable chelal finger length 0.93–0.96
<emphasis id="30B9EAA1FFEEA90BFE69FC45FDE6FC03" box="[495,525,943,968]" italics="true" pageId="32" pageNumber="33">vs.</emphasis>
<quantity id="C5359B56FFEEA90BFD92FC44FD51FC02" box="[532,698,942,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.245" metricValueMax="1.25" metricValueMin="1.24" pageId="32" pageNumber="33" unit="mm" value="1.245" valueMax="1.25" valueMin="1.24">1.24–1.25 mm</quantity>
; all in males), the shape of the chelal fingers (the intercalary teeth round and obtuse
<emphasis id="30B9EAA1FFEEA90BFEE7FC39FE6BFC27" box="[353,384,979,1004]" italics="true" pageId="32" pageNumber="33">vs.</emphasis>
small and pointed, teeth of the movable chelal finger retrorse and contiguous
<emphasis id="30B9EAA1FFEEA90BFB74FC39FAFAFC27" box="[1266,1297,979,1004]" italics="true" pageId="32" pageNumber="33">vs.</emphasis>
pointed and well-spaced), and the number of setae on tergites IV (2
<emphasis id="30B9EAA1FFEEA90BFC83FC1DFCCFFBDB" box="[773,804,1015,1040]" italics="true" pageId="32" pageNumber="33">vs.</emphasis>
4).
</paragraph>
<paragraph id="027236B3FFEEA90BFF41FBF1FBC5FBB6" blockId="32.[151,1437,151,1473]" pageId="32" pageNumber="33">
<taxonomicName id="C5CD4D30FFEEA90BFF41FBF1FE13FBFF" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[199,504,1051,1077]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="retrorsus" status="sp. nov.">
<emphasis id="30B9EAA1FFEEA90BFF41FBF1FE13FBFF" box="[199,504,1051,1077]" italics="true" pageId="32" pageNumber="33">Lagynochthonius retrorsus</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFEEA90BFD85FBF1FD8AFBFE" bold="true" box="[515,609,1051,1077]" pageId="32" pageNumber="33">
<taxonomicNameLabel id="2B8A57DAFFEEA90BFD85FBF1FD8AFBFE" box="[515,609,1051,1077]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="C5CD4D30FFEEA90BFC87FBF1FC3FFBFE" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[769,980,1051,1077]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="bailongtanensis">
<emphasis id="30B9EAA1FFEEA90BFC87FBF1FC3FFBFE" box="[769,980,1051,1077]" italics="true" pageId="32" pageNumber="33">L. bailongtanensis</emphasis>
</taxonomicName>
in the shape of epistome (pointed and triangular
<emphasis id="30B9EAA1FFEEA90BFE8AFBD5FEC8FB93" box="[268,291,1087,1112]" italics="true" pageId="32" pageNumber="33">vs</emphasis>
. rounded and obtuse), the dentation of the chelal fingers (with intercalary teeth on fixed chelal finger
<emphasis id="30B9EAA1FFEEA90BFF11FB89FF45FBB7" box="[151,174,1123,1148]" italics="true" pageId="32" pageNumber="33">vs</emphasis>
. without intercalary teeth), and the number of setae on tergites I–IV (2
<emphasis id="30B9EAA1FFEEA90BFC48FB89FC07FBB7" box="[974,1004,1123,1148]" italics="true" pageId="32" pageNumber="33">vs.</emphasis>
3–4).
</paragraph>
<paragraph id="027236B3FFEEA90BFF41FB6DFB22FB22" blockId="32.[151,1437,151,1473]" pageId="32" pageNumber="33">
<taxonomicName id="C5CD4D30FFEEA90BFF41FB6DFE1FFB6B" authority="Hou &amp; Gao &amp; Zhang, 2022" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[199,500,1159,1185]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="retrorsus" status="sp. nov.">
<emphasis id="30B9EAA1FFEEA90BFF41FB6DFE1FFB6B" box="[199,500,1159,1185]" italics="true" pageId="32" pageNumber="33">Lagynochthonius retrorsus</emphasis>
</taxonomicName>
<emphasis id="30B9EAA1FFEEA90BFE7CFB6DFDBFFB6A" bold="true" box="[506,596,1159,1185]" pageId="32" pageNumber="33">
<taxonomicNameLabel id="2B8A57DAFFEEA90BFE7CFB6DFDBFFB6A" box="[506,596,1159,1185]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="C5CD4D30FFEEA90BFC05FB6DFBA0FB6A" authorityName="Hou &amp; Gao &amp; Zhang" authorityYear="2022" box="[899,1099,1159,1185]" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="laoxueyanensis">
<emphasis id="30B9EAA1FFEEA90BFC05FB6DFBA0FB6A" box="[899,1099,1159,1185]" italics="true" pageId="32" pageNumber="33">L. laoxueyanensis</emphasis>
</taxonomicName>
by the presence of intercalary teeth on fixed chelal finger, the number of setae on tergites III and IV (2 and 2
<emphasis id="30B9EAA1FFEEA90BFB99FB41FBD6FB0F" box="[1055,1085,1195,1220]" italics="true" pageId="32" pageNumber="33">vs.</emphasis>
4 and 4, respectively), and the larger body size (body length 1.94–2.04
<emphasis id="30B9EAA1FFEEA90BFDDBFB25FD90FB23" box="[605,635,1231,1256]" italics="true" pageId="32" pageNumber="33">vs.</emphasis>
<quantity id="C5359B56FFEEA90BFD04FB25FD0DFB22" box="[642,742,1231,1257]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" pageId="32" pageNumber="33" unit="mm" value="1.78">1.78 mm</quantity>
in males) (
<bibRefCitation id="665C4B42FFEEA90BFCE5FB25FC15FB22" author="Li, Y. C. &amp; Liu, H. &amp; Shi, A. M." box="[867,1022,1231,1257]" pageId="32" pageNumber="33" pagination="28 - 34" refId="ref39131" refString="Li, Y. C., Liu, H. &amp; Shi, A. M. (2019) A new cave-dwelling species of Lagynochthonius (Arachnida: Pseudoscorpiones: Chthoniidae) from Yunnan Province, China. Zootaxa, 4571 (1), 28 - 34. https: // doi. org / 10.11646 / zootaxa. 4571.1.2" type="journal article" year="2019">
Li
<emphasis id="30B9EAA1FFEEA90BFC02FB25FC5DFB22" box="[900,950,1231,1257]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
. 2019
</bibRefCitation>
;
<bibRefCitation id="665C4B42FFEEA90BFB8CFB25FB56FB22" author="Hou, Y. M. &amp; Gao, Z. Z. &amp; Zhang, F." box="[1034,1213,1231,1257]" pageId="32" pageNumber="33" pagination="65 - 83" refId="ref38684" refString="Hou, Y. M., Gao, Z. Z. &amp; Zhang, F. (2022) Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from Yunnan, China. ZooKeys, 1097, 65 - 83. https: // doi. org / 10.3897 / zookeys. 1097.82527" type="journal article" year="2022">
Hou
<emphasis id="30B9EAA1FFEEA90BFBC7FB25FB98FB22" box="[1089,1139,1231,1257]" italics="true" pageId="32" pageNumber="33">et al</emphasis>
. 2022
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4AD76538FFEEA90BFF41FB18FD0DFA0A" pageId="32" pageNumber="33" type="distribution">
<paragraph id="027236B3FFEEA90BFF41FB18FD0DFA0A" blockId="32.[151,1437,151,1473]" pageId="32" pageNumber="33">
<emphasis id="30B9EAA1FFEEA90BFF41FB18FE04FAC7" bold="true" box="[199,495,1266,1293]" pageId="32" pageNumber="33">Distribution and habitat.</emphasis>
This species is known only from the
<typeStatus id="DD768811FFEEA90BFC0EFB19FC53FAC6" box="[904,952,1267,1293]" pageId="32" pageNumber="33">type</typeStatus>
locality, Qinglong Cave (
<figureCitation id="9AF62A36FFEEA90BFB55FB19FAC4FAC6" box="[1235,1327,1267,1293]" captionStart="FIGURE 1" captionStartId="3.[151,250,1542,1566]" captionTargetBox="[152,1433,567,1514]" captionTargetId="figure-169@3.[151,1436,561,1519]" captionTargetPageId="3" captionText="FIGURE 1. Distribution map of the type locality of troglomorphic Lagynochthonius species in Yunnan (China). A. L. retrorsus sp. nov. (Qinglong Cave); B. L. fengi sp. nov. (Unnamed Cave 2); C. L. xiaolinensis sp. nov. (Dayan Cave); D. L. xibaiensis sp. nov. (Hama Cave); E. L. spinulentus sp. nov. (Yingshan Cave); F. L. xinjiaoensis sp. nov. (Yueliang Cave); G. L. magnidentatus sp. nov. (Xiao Cave); H. L. bailongtanensis sp. nov. (Qianfo Cave); I. L. laoxueyanensis sp. nov. (Laoxueyan Cave); J. L. serratus sp. nov. (Unnamed Cave 3); K. L. yaowangguensis sp. nov. (Unnamed Cave 4); L. L. crassus sp. nov. (Unnamed Cave 1); M–N. L. minimus sp. nov. (M: Yanfen Cave, N: Cizhu Cave)." figureDoi="http://doi.org/10.5281/zenodo.7251604" httpUri="https://zenodo.org/record/7251604/files/figure.png" pageId="32" pageNumber="33">Figs 1A</figureCitation>
,
<figureCitation id="9AF62A36FFEEA90BFABCFB19FA64FAC6" box="[1338,1423,1267,1293]" captionStart="FIGURE 19" captionStartId="28.[151,250,1430,1454]" captionTargetBox="[151,1436,273,1407]" captionTargetId="figure-71@28.[151,1436,273,1407]" captionTargetPageId="28" captionText="FIGURE 19. Qinglong Cave, type locality of Lagynochthonius retrorsus sp. nov., A. Entrance; B. Inside the cave entrance; C–D. Areas where L. retrorsus sp. nov. specimens were collected; E–F. Live individuals of L. retrorsus sp. nov. (E: male, F: female) in their natural environment." figureDoi="http://doi.org/10.5281/zenodo.7251663" httpUri="https://zenodo.org/record/7251663/files/figure.png" pageId="32" pageNumber="33">19A–D</figureCitation>
), which is located about
<quantity id="C5359B56FFEEA90BFE13FAFDFE09FAFA" box="[405,482,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="32" pageNumber="33" unit="km" value="1.2">1.2 km</quantity>
northwest of Xingfu Village (Daguan County). This limestone cave, located at the top of a mountain and surrounded by residential areas, has one small and triangular entrance (about
<quantity id="C5359B56FFEEA90BFB52FAD1FAEBFA9E" box="[1236,1280,1339,1365]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="32" pageNumber="33" unit="m" value="1.0">1 m</quantity>
high and
<quantity id="C5359B56FFEEA90BFAF7FAD1FA76FA9E" box="[1393,1437,1339,1365]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="m" value="2.0">2 m</quantity>
wide). A winding tunnel (approximately
<quantity id="C5359B56FFEEA90BFDE6FAB5FD4CFAB2" box="[608,679,1375,1401]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="32" pageNumber="33" unit="m" value="190.0">190 m</quantity>
in length) leads to the deepest part of the cave. All specimens were collected under stones within
<quantity id="C5359B56FFEEA90BFE6FFA68FD83FA56" box="[489,616,1410,1437]" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="32" pageNumber="33" unit="m" value="125.0" valueMax="150.0" valueMin="100.0">100–150 m</quantity>
from the entrance. This space range is completely dark, with an average temperature of around 13℃ and humidity over 90%.
</paragraph>
</subSubSection>
</treatment>
</document>
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