164 lines
17 KiB
XML
164 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.96.81737" ID-GBIF-Dataset="d148b8fa-4f4c-4069-80e5-f04f98561bfe" ID-Pensoft-Pub="2747-8106-96-51" ID-Pensoft-UUID="FCEF2A912BC258BBB022681E86486B06" ID-ZooBank="CD8C70BA025345408EFCD430A12F6B35" ModsDocID="2747-8106-96-51" checkinTime="1649795137967" checkinUser="pensoft" docAuthor="Schwarzhans, Werner & Keupp, Helmut" docDate="2022" docId="71A2E839B20B523DBA6D3F93DB99997E" docLanguage="en" docName="Zitteliana 96: 51-67" docOrigin="Zitteliana 96" docPubDate="2022-04-12" docSource="http://dx.doi.org/10.3897/zitteliana.96.81737" docTitle="Leptolepis macrocephalus" docType="treatment" docVersion="2" id="FCEF2A912BC258BBB022681E86486B06" lastPageNumber="51" masterDocId="FCEF2A912BC258BBB022681E86486B06" masterDocTitle="Early teleost otolith morphogenesis observed in the Jurassic of Franconia, Bavaria, southern Germany" masterLastPageNumber="67" masterPageNumber="51" pageId="0" pageNumber="51" updateTime="1649795760618" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Early teleost otolith morphogenesis observed in the Jurassic of Franconia, Bavaria, southern Germany</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Schwarzhans, Werner</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4842-7989</mods:nameIdentifier>
|
||
<mods:affiliation>Zoological Museum, Natural History Museum of Denmark, Universitetsparken 15, 2100 Kobenhavn, Denmark & Ahrensburger Weg 103, 22359 Hamburg, Germany</mods:affiliation>
|
||
<mods:nameIdentifier type="email">wwschwarz@aol.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Keupp, Helmut</mods:namePart>
|
||
<mods:affiliation>Freie Universitaet Berlin, Institut fuer Geologische Wissenschaften, Fachrichtung Palaeontologie, Malteser Strasse 74 - 100, Haus D, 12249 Berlin, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Zitteliana</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2022</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2022-04-12</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>96</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>51</mods:start>
|
||
<mods:end>67</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.3897/zitteliana.96.81737</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zitteliana.96.81737</mods:identifier>
|
||
<mods:identifier type="Pensoft-Pub">2747-8106-96-51</mods:identifier>
|
||
<mods:identifier type="ZooBank">CD8C70BA025345408EFCD430A12F6B35</mods:identifier>
|
||
<mods:identifier type="Pensoft-UUID">FCEF2A912BC258BBB022681E86486B06</mods:identifier>
|
||
</mods:mods>
|
||
<treatment LSID="urn:lsid:plazi:treatment:71A2E839B20B523DBA6D3F93DB99997E" httpUri="http://treatment.plazi.org/id/71A2E839B20B523DBA6D3F93DB99997E" lastPageNumber="51" pageId="0" pageNumber="51">
|
||
<subSubSection pageId="0" pageNumber="51" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="51">
|
||
<taxonomicName LSID="71A2E839-B20B-523D-BA6D-3F93DB99997E" authority="(Schroeder, 1956)" baseAuthorityName="Schroeder" baseAuthorityYear="1956" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Leptolepis macrocephalus" order="Leptolepidiformes" pageId="0" pageNumber="51" rank="species" species="macrocephalus">
|
||
Leptolepis macrocephalus (
|
||
<normalizedToken originalValue="Schröder">Schroeder</normalizedToken>
|
||
, 1956)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="51" type="description">
|
||
<paragraph pageId="0" pageNumber="51">
|
||
<figureCitation captionStart="Plate 3" captionStartId="F5" captionText="Plate 3. Otoliths of Leptolepis kremmeldorfensis, L. inaequalis, L. macrocephalus, and Leptoelops rhenanus. figs 1 - 6: Leptolepis kremmeldorfensis (Schroeder, 1956); 1: holotype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 35); 2: paratype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 36); 3: holotype of Ot. amygdalinus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 14); 4: paratype of Ot. bambergensis Schroeder, 1956 from Freiahorn (SNSB-BSPG 2022 III 10); 5: holotype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 24); 6: paratype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 27). figs 7, 8: Leptolepis cf. kremmeldorfensis (Schroeder, 1956), from the late Pliensbachian of Buttenheim, SNSB-BSPG 2022 IV 15. figs 9 - 11: Leptolepis inaequalis (Weiler, 1954); 9: holotype of Ot. puenzendorfensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 39); 10: holotype of Ot. calloviensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 8); 11: holotype of Ot. inaequalis Weiler, 1954 from the Callovian to Oxfordian of Kandern, Baden-Wuertemberg (SMF P. 3067). figs 12 - 13: Leptolepis macrocephalus (Schroeder, 1956); 12: holotype of Ot. guttaeformis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 6); 13: holotype of Ot. macrocephalus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 1). figs 14 - 16: Leptoelops rhenanus (Weiler, 1954); 14: holotype of Ot. rhenanus Weiler, 1954 from the Callovian of well Bruchsal D 205 (core from 482 to 487 m) near Weingarten, Baden-Wuerttemberg (SMF P. 2953); 15: holotype of Ot. rectisulcatus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 37); 16: holotype of Ot. scissus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 40)." figureDoi="10.3897/zitteliana.96.81737.plate3" httpUri="https://binary.pensoft.net/fig/671633" pageId="0" pageNumber="51">Plate 3, figs 12-13</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="51" type="reference_group">
|
||
<paragraph pageId="0" pageNumber="51">
|
||
<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus macrocephali" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="macrocephali">Otolithus macrocephali</taxonomicName>
|
||
1956
|
||
<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus macrocephali" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="macrocephali">Otolithus macrocephali</taxonomicName>
|
||
-
|
||
<normalizedToken originalValue="Schröder">Schroeder</normalizedToken>
|
||
: pl. 7, fig. 40.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="51">
|
||
<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus guttaeformis" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="guttaeformis">Otolithus guttaeformis</taxonomicName>
|
||
1956
|
||
<taxonomicName authority="Koken, 1884" authorityName="Koken" authorityYear="1884" class="Actinopterygii" family="Leptolepididae" genus="Otolithus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Otolithus guttaeformis" order="Leptolepidiformes" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="guttaeformis">Otolithus guttaeformis</taxonomicName>
|
||
-
|
||
<normalizedToken originalValue="Schröder">Schroeder</normalizedToken>
|
||
: pl. 6, figs 15-16.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="51" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="51">Material.</paragraph>
|
||
<paragraph pageId="0" pageNumber="51">
|
||
<materialsCitation collectingDate="2022-03-01" collectingDateMax="2022-03-06" collectingDateMin="2022-03-01" location="Schroeder's" municipality="Callovian of Puenzendorf" specimenCount="3" typeStatus="holotype">
|
||
<specimenCount type="generic">3 specimens</specimenCount>
|
||
from
|
||
<normalizedToken originalValue="Schröder’s">
|
||
<location LSID="urn:lsid:plazi:treatment:71A2E839B20B523DBA6D3F93DB99997E:FDB30D961BB20AEC312AB54569BC26A2" municipality="Callovian of Puenzendorf" name="Schroeder's">Schroeder's</location>
|
||
</normalizedToken>
|
||
collection from the early
|
||
<collectingMunicipality>
|
||
Callovian of
|
||
<normalizedToken originalValue="Pünzendorf">Puenzendorf</normalizedToken>
|
||
</collectingMunicipality>
|
||
:
|
||
<typeStatus>holotype</typeStatus>
|
||
of
|
||
<taxonomicName genus="Ot." lsidName="Ot. macrocephalus" pageId="0" pageNumber="51" rank="species" species="macrocephalus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">Ot. macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
(emended spelling, SNSB-BSPG
|
||
<collectingDate value="2022-03-01">2022 III 1</collectingDate>
|
||
, Plate
|
||
<figureCitation captionStart="Plate 3" captionStartId="F5" captionText="Plate 3. Otoliths of Leptolepis kremmeldorfensis, L. inaequalis, L. macrocephalus, and Leptoelops rhenanus. figs 1 - 6: Leptolepis kremmeldorfensis (Schroeder, 1956); 1: holotype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 35); 2: paratype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 36); 3: holotype of Ot. amygdalinus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 14); 4: paratype of Ot. bambergensis Schroeder, 1956 from Freiahorn (SNSB-BSPG 2022 III 10); 5: holotype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 24); 6: paratype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 27). figs 7, 8: Leptolepis cf. kremmeldorfensis (Schroeder, 1956), from the late Pliensbachian of Buttenheim, SNSB-BSPG 2022 IV 15. figs 9 - 11: Leptolepis inaequalis (Weiler, 1954); 9: holotype of Ot. puenzendorfensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 39); 10: holotype of Ot. calloviensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 8); 11: holotype of Ot. inaequalis Weiler, 1954 from the Callovian to Oxfordian of Kandern, Baden-Wuertemberg (SMF P. 3067). figs 12 - 13: Leptolepis macrocephalus (Schroeder, 1956); 12: holotype of Ot. guttaeformis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 6); 13: holotype of Ot. macrocephalus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 1). figs 14 - 16: Leptoelops rhenanus (Weiler, 1954); 14: holotype of Ot. rhenanus Weiler, 1954 from the Callovian of well Bruchsal D 205 (core from 482 to 487 m) near Weingarten, Baden-Wuerttemberg (SMF P. 2953); 15: holotype of Ot. rectisulcatus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 37); 16: holotype of Ot. scissus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 40)." figureDoi="10.3897/zitteliana.96.81737.plate3" httpUri="https://binary.pensoft.net/fig/671633" pageId="0" pageNumber="51">3</figureCitation>
|
||
, fig. 13),
|
||
<typeStatus>holotype</typeStatus>
|
||
of
|
||
<taxonomicName genus="Ot." lsidName="Ot. guttaeformis" pageId="0" pageNumber="51" rank="species" species="guttaeformis">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">Ot. guttaeformis</emphasis>
|
||
</taxonomicName>
|
||
(SNSB-BSPG
|
||
<collectingDate value="2022-03-06">2022 III 6</collectingDate>
|
||
, Plate
|
||
<figureCitation captionStart="Plate 3" captionStartId="F5" captionText="Plate 3. Otoliths of Leptolepis kremmeldorfensis, L. inaequalis, L. macrocephalus, and Leptoelops rhenanus. figs 1 - 6: Leptolepis kremmeldorfensis (Schroeder, 1956); 1: holotype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 35); 2: paratype of Ot. kremmeldorfensis Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 36); 3: holotype of Ot. amygdalinus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 14); 4: paratype of Ot. bambergensis Schroeder, 1956 from Freiahorn (SNSB-BSPG 2022 III 10); 5: holotype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 24); 6: paratype of Ot. franconicus Schroeder, 1956 from Kremmeldorf (SNSB-BSPG 2022 III 27). figs 7, 8: Leptolepis cf. kremmeldorfensis (Schroeder, 1956), from the late Pliensbachian of Buttenheim, SNSB-BSPG 2022 IV 15. figs 9 - 11: Leptolepis inaequalis (Weiler, 1954); 9: holotype of Ot. puenzendorfensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 39); 10: holotype of Ot. calloviensis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 8); 11: holotype of Ot. inaequalis Weiler, 1954 from the Callovian to Oxfordian of Kandern, Baden-Wuertemberg (SMF P. 3067). figs 12 - 13: Leptolepis macrocephalus (Schroeder, 1956); 12: holotype of Ot. guttaeformis Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 6); 13: holotype of Ot. macrocephalus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 1). figs 14 - 16: Leptoelops rhenanus (Weiler, 1954); 14: holotype of Ot. rhenanus Weiler, 1954 from the Callovian of well Bruchsal D 205 (core from 482 to 487 m) near Weingarten, Baden-Wuerttemberg (SMF P. 2953); 15: holotype of Ot. rectisulcatus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 37); 16: holotype of Ot. scissus Schroeder, 1956 from Puenzendorf (SNSB-BSPG 2022 III 40)." figureDoi="10.3897/zitteliana.96.81737.plate3" httpUri="https://binary.pensoft.net/fig/671633" pageId="0" pageNumber="51">3</figureCitation>
|
||
, fig. 12) and
|
||
</materialsCitation>
|
||
<specimenCount type="generic" typeStatus="paratype">1 paratype</specimenCount>
|
||
(SNSB-BSPG
|
||
<date value="2022-03-07">2022 III 7</date>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="51" type="differential diagnosis">
|
||
<paragraph pageId="0" pageNumber="51">Differential diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="51">
|
||
<taxonomicName baseAuthorityName="Schroeder" baseAuthorityYear="1956" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" kingdom="Animalia" lsidName="Leptolepis macrocephalus" order="Elopiformes" pageId="0" pageNumber="51" rank="species" species="macrocephalus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">Leptolepis macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
differs from the coeval
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. inaequalis" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="inaequalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. inaequalis</emphasis>
|
||
</taxonomicName>
|
||
in its more compressed shape, which is exhibited by a deeper ventral rim and a more expanded postdorsal rim and is expressed in the OL:OH ratio of 1.35-1.4 (vs. 1.5). It shares with
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. inaequalis" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="inaequalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. inaequalis</emphasis>
|
||
</taxonomicName>
|
||
completely flat inner and outer faces, features that distinguish both species from all other known
|
||
<taxonomicName class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" kingdom="Animalia" lsidName="Leptolepis" order="Elopiformes" pageId="0" pageNumber="51" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">Leptolepis</emphasis>
|
||
</taxonomicName>
|
||
otoliths.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="51" type="discussion">
|
||
<paragraph pageId="0" pageNumber="51">Discussion.</paragraph>
|
||
<paragraph pageId="0" pageNumber="51">
|
||
<taxonomicName baseAuthorityName="Schroeder" baseAuthorityYear="1956" class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" kingdom="Animalia" lsidName="Leptolepis macrocephalus" order="Elopiformes" pageId="0" pageNumber="51" rank="species" species="macrocephalus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">Leptolepis macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Actinopterygii" genus="L." kingdom="Animalia" lsidName="L. inaequalis" pageId="0" pageNumber="51" phylum="Chordata" rank="species" species="inaequalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">L. inaequalis</emphasis>
|
||
</taxonomicName>
|
||
form a small group of otoliths with a special trait of having flat inner and outer faces. They occur during the Callovian and possibly the early Oxfordian. Their attribution to the genus is therefore less certain than that of
|
||
<taxonomicName class="Actinopterygii" family="Leptolepididae" genus="Leptolepis" kingdom="Animalia" lsidName="Leptolepis" order="Elopiformes" pageId="0" pageNumber="51" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="51">Leptolepis</emphasis>
|
||
</taxonomicName>
|
||
otoliths from the Early Jurassic (see above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |